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Cephalaspidea

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The vast majority possess a shell, although it may be reduced or internal. They have a well-developed headshield, a characteristic broadening at the head, which is used to plow beneath the surface of the sand. This headshield prevents the sand entering the mantle cavity. There is a muscular foot with
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The taxonomy of the shelled cephalaspideans, the bubble snails, like that of many shelled mollusks, used to be based very simply on shell characteristics. But because there are some similarities in shell morphology throughout this group, more recently taxonomists have taken other anatomical
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Oskars T.R., Bouchet P. & Malaquias M.A. (2015). A new phylogeny of the Cephalaspidea (Gastropoda: Heterobranchia) based on expanded taxon sampling and gene markers. Molecular Phylogenetics and Evolution. 89:
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also have well-developed eyes on the anterior end of the head and bundles of sensory cilia around the mouth. With these cilia they are able to track their prey by following the victim's mucous trail.
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of the Cephalaspidea was confirmed, but the families Cylichnidae, Diaphanidae, Haminoeidae, Philinidae, and Retusidae were found non-monophyletic. This had led to the creation of new families ((
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situated between the foot and the headshield. It plays a role in olfactory and sensory detection. It is visible as a dark brown pit at the base of the right
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is another common name for these families of marine gastropods, some of which have thin bubble-like shells. This clade contains more than 600 species.
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has been included into the new Informal Group "Lower Heterobranchia" and the superfamily Cylindrobulloidea becomes part of the Group Cylindrobullida.
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by Malaquias et al. (2009) is arranged as follows (there are listed genera under molecular analysis; not analyzed families are under "
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Headshield slugs often live just beneath the surface of the sand and can also be seen crawling on rocks. They have well-developed
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in the journal Molecular Phylogenetics and Evolution. 89 came to the following conclusion, with the creation of new families
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Jörger, Katharina M; Stöger, Isabella; Kano, Yasunori; Fukuda, Hiroshi; Knebelsberger, Thomas; Schrödl, Michael (2010).
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Malaquias, Manuel AntĂłnio E; Mackenzie-Dodds, Jacqueline; Bouchet, Philippe; Gosliner, Terrence; Reid, David G (2009).
948:. A new phylogeny of the Cephalaspidea (Gastropoda: Heterobranchia) based on expanded taxon sampling and gene markers. 264:
In 2015 a new study based on molecular phylogenetics has changed significantly the taxonomy of the Cephalaspidea. The
1295:"A new phylogeny of the Cephalaspidea (Gastropoda: Heterobranchia) based on expanded taxon sampling and gene markers" 1272: 1715: 1648: 1072: 1139: 1630: 245: 217: 65: 1539: 1526: 350: 1544: 1431: 1393: 1168: 1697: 1658: 1097: 1720: 1504: 1351: 1743: 1684: 1466: 1038: 1679: 340: 261:
characteristics into consideration, such as the radula, gizzard, penis, and Hancock's organ.
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were accepted. The composition of each of the superfamilies was drastically rearranged.
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Headshield slugs are the most morphologically diverse group of all the opisthobranchs.
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Members of this worldwide clade used to be considered the most ancestral of the
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Wägele, Heike; Klussmann-Kolb, Annette; Verbeek, Eva; Schrödl, Michael (2014).
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This article is about the gastropod taxon. For the prehistoric fish taxon, see
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Shells. The visual guide to over 500 species of seashell from around the world
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Malaquias M. A. E. (2010). "Systematics, phylogeny, and natural history of
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Oskars, Trond R.; Bouchet, Philippe; Malaquias, Manuel AntĂłnio E. (2015).
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The publication by Oskars T.R., Bouchet P. & Malaquias M.A. (2015)
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Malaquias et al. (2009) have rearranged taxonomy of Cephalaspidea
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they did not use superfamilies in the classification scheme.
905:(formerly the only member of Bullactidae) into the family 1292: 1199: 1029:Pruvot-Fol, 1954 (formerly Laoninae) – type genus: 1017:Oskars, Bouchet, and Malaquias, 2015 – type genus: 1005:Oskars, Bouchet, and Malaquias, 2015 – type genus: 993:Oskars, Bouchet, and Malaquias, 2015 – type genus: 981:Oskars, Bouchet, and Malaquias, 2015 – type genus: 969:Oskars, Bouchet, and Malaquias, 2015 – type genus: 220:to detect prey, which may be other opisthobranchs, 1735: 1421: 1419: 1387: 1385: 1383: 1381: 1379: 353:, 1931 – These are now included in the suborder 1036:Cephalaspidea (temporary name) with the genera 208:or without parapodia (fleshy wing-like flaps). 1416: 1376: 1195: 1193: 1191: 212:Life habits and related anatomical structures 228:. Several species are voracious carnivores. 1188: 36: 1367: 1326: 1235: 1225: 912:Jörger et al. (2010) moved Cephalaspidea 664:Fischer, 1887 – This means Cephalaspidea 1288: 1286: 1284: 323:Suborder Cephalaspidea P. Fischer, 1883 1736: 532:as a valid name outside Cephalaspidea. 406:Cephalaspidea is arranged as follows: 231:Members of the brightly colored genus 1465: 1464: 1299:Molecular Phylogenetics and Evolution 1281: 1258: 899:Subsequently, Malaquias (2010) moved 1698:BF18F633-A9A4-FF41-2B9C-C662FD31F875 1659:5b5e2fd9-efd8-4423-883b-589770fec44e 975:Oskars, Bouchet, and Malaquias, 2015 922:and they confirmed the placement of 317: 1356:Organisms Diversity & Evolution 13: 55:, head end towards the lower left 14: 1760: 1410:10.1111/j.1463-6409.2008.00354.x 1159: 1138: 1117: 1096: 941: 894: 696:H. Adams & A. Adams, 1854 – 645:H. Adams & A. Adams, 1854 – 509: 400:taxonomy of Bouchet & Rocroi 393: 165:of sea slugs and bubble snails, 64: 1343: 1252: 1011:Ortea, Moro and Espinosa, 2013 546:The taxonomy of Cephalaspidea 1: 1181: 621:Philippi, 1853 incertae sedis 1444:10.1080/00222933.2010.487574 7: 1319:10.1016/j.ympev.2015.04.011 987:H. Adams and A. Adams, 1854 890:Thiele, 1826 incertae sedis 846:Odhner, 1968 incertae sedis 628:Thiele, 1931 incertae sedis 255: 10: 1765: 1749:Protostome unranked clades 1438:(33 & 34): 2015-2029. 1432:Journal of Natural History 1169:Siphopteron quadrispinosum 314:) are reinstated as valid 202: 18: 1473: 1369:10.1007/s13127-013-0151-5 1071:was not supported in the 957:Dall, 1913 – type genus: 657:Burn, 1963 incertae sedis 614:Gray, 1850 incertae sedis 138: 133: 61:Scientific classification 59: 44: 35: 30: 1227:10.1186/1471-2148-10-323 1206:BMC Evolutionary Biology 918:and Runcinacea into the 175:within the larger clade 288:) ) and one new genus ( 294:). Two family names ( 1680:Paleobiology Database 1259:Dance, S. P. (1996). 1073:Bayesian phylogenetic 45:A live individual of 1267:. pp. 200–203. 1083:. The superfamilies 928:Lower Heterobranchia 224:or bristleworms and 153:, also known as the 1311:2015MolPE..89..130O 1218:2010BMCEE..10..323J 1147:Chelidonura varians 774:Pilsbry, 1895–96 – 48:Chelidonura varians 16:Order of gastropods 1265:Dorling Kindersley 708:G. O Sars, 1878 – 572:d'Orbigny, 1843 – 561:Architectibranchia 539:as a valid family. 524:Architectibranchia 302:) and two genera ( 246:chemosensory organ 218:sensory structures 1731: 1730: 1667:Open Tree of Life 1467:Taxon identifiers 1398:Zoologica Scripta 1126:Haminoea cymbalum 1062: 1053: 1044: 920:Euopisthobranchia 822:Swainson, 1840 – 810:Hertling, 1932 – 689:is incertae sedis 564:Haszprunar, 1985 348:Cylindrobulloidea 318:Linnaean taxonomy 196: 177:Euopisthobranchia 147: 146: 31:Headshield slugs 1756: 1724: 1723: 1711: 1710: 1701: 1700: 1688: 1687: 1675: 1674: 1662: 1661: 1652: 1651: 1639: 1638: 1636:NHMSYS0021056023 1626: 1625: 1613: 1612: 1600: 1599: 1587: 1586: 1574: 1573: 1561: 1560: 1548: 1547: 1535: 1534: 1522: 1521: 1509: 1508: 1507: 1494: 1493: 1492: 1462: 1461: 1447: 1427:Bullacta exarata 1423: 1414: 1413: 1389: 1374: 1373: 1371: 1347: 1341: 1340: 1330: 1290: 1279: 1278: 1256: 1250: 1249: 1239: 1229: 1197: 1163: 1142: 1121: 1100: 1067:The superfamily 1060: 1051: 1042: 902:Bullacta exarata 868:Phanerophthalmus 853:Pilsbry, 1893 – 808:Philinoglossidae 502:The superfamily 475:Philinoglossidae 449:Smaragdinellidae 194: 155:headshield slugs 69: 68: 40: 28: 27: 1764: 1763: 1759: 1758: 1757: 1755: 1754: 1753: 1734: 1733: 1732: 1727: 1719: 1714: 1706: 1704: 1696: 1691: 1683: 1678: 1670: 1665: 1657: 1655: 1647: 1642: 1634: 1629: 1621: 1616: 1608: 1603: 1595: 1590: 1582: 1577: 1569: 1564: 1556: 1551: 1543: 1538: 1530: 1525: 1517: 1512: 1503: 1502: 1497: 1488: 1487: 1482: 1469: 1451: 1450: 1424: 1417: 1390: 1377: 1348: 1344: 1291: 1282: 1275: 1257: 1253: 1198: 1189: 1184: 1177: 1164: 1155: 1143: 1134: 1122: 1113: 1101: 1075:hypothesis and 991:Colpodaspididae 944: 932:Notodiaphanidae 897: 720:Thiele, 1925 – 677:Odhner, 1914 – 626:Notodiaphanidae 512: 430:Notodiaphanidae 396: 320: 278:Colpodaspididae 258: 242:Hancock's organ 214: 205: 63: 24: 17: 12: 11: 5: 1762: 1752: 1751: 1746: 1729: 1728: 1726: 1725: 1712: 1702: 1689: 1676: 1663: 1653: 1640: 1627: 1614: 1601: 1588: 1575: 1562: 1549: 1536: 1523: 1510: 1495: 1479: 1477: 1471: 1470: 1459: 1458: 1449: 1448: 1415: 1375: 1342: 1280: 1273: 1251: 1186: 1185: 1183: 1180: 1179: 1178: 1174:Gastropteridae 1165: 1158: 1156: 1144: 1137: 1135: 1123: 1116: 1114: 1102: 1095: 1065: 1064: 1034: 1033:A. Adams, 1865 1024: 1012: 1000: 988: 976: 964: 943: 940: 896: 893: 892: 891: 884: 883:incertae sedis 847: 840: 831:Sagaminopteron 820:Gastropteridae 816: 804: 768: 756: 744: 732: 714: 702: 690: 659: 658: 651: 630: 629: 622: 615: 608: 590: 554:incertae sedis 544: 543: 540: 533: 526: 511: 508: 500: 499: 485: 471:Gastropteridae 451: 433: 418: 395: 392: 391: 390: 389: 388: 387:Philippi, 1853 381: 371: 364: 357: 344: 334: 319: 316: 257: 254: 226:foraminiferans 213: 210: 204: 201: 195:Spengel, 1881. 188:opisthobranchs 145: 144: 136: 135: 134:Superfamilies 131: 130: 125: 121: 120: 115: 111: 110: 105: 101: 100: 95: 91: 90: 85: 81: 80: 75: 71: 70: 57: 56: 51:in the family 42: 41: 33: 32: 15: 9: 6: 4: 3: 2: 1761: 1750: 1747: 1745: 1744:Cephalaspidea 1742: 1741: 1739: 1722: 1717: 1713: 1709: 1703: 1699: 1694: 1690: 1686: 1681: 1677: 1673: 1668: 1664: 1660: 1654: 1650: 1645: 1641: 1637: 1632: 1628: 1624: 1619: 1615: 1611: 1606: 1602: 1598: 1593: 1589: 1585: 1580: 1576: 1572: 1567: 1563: 1559: 1554: 1550: 1546: 1541: 1537: 1533: 1532:Cephalaspidea 1528: 1524: 1520: 1519:Cephalaspidea 1515: 1511: 1506: 1505:Cephalaspidea 1500: 1496: 1491: 1485: 1481: 1480: 1478: 1476: 1475:Cephalaspidea 1472: 1468: 1463: 1457: 1453: 1452: 1445: 1441: 1437: 1434: 1433: 1428: 1422: 1420: 1411: 1407: 1403: 1399: 1395: 1388: 1386: 1384: 1382: 1380: 1370: 1365: 1361: 1357: 1353: 1346: 1338: 1334: 1329: 1324: 1320: 1316: 1312: 1308: 1304: 1300: 1296: 1289: 1287: 1285: 1276: 1274:0-86318-811-7 1270: 1266: 1262: 1255: 1247: 1243: 1238: 1233: 1228: 1223: 1219: 1215: 1211: 1207: 1203: 1196: 1194: 1192: 1187: 1175: 1171: 1170: 1162: 1157: 1153: 1149: 1148: 1141: 1136: 1132: 1128: 1127: 1120: 1115: 1111: 1107: 1106: 1105:Bulla ampulla 1099: 1094: 1093: 1092: 1090: 1086: 1082: 1078: 1074: 1070: 1061:Iredale, 1936 1059: 1058: 1050: 1049: 1041: 1040: 1035: 1032: 1028: 1025: 1022: 1021: 1016: 1015:Philinorbidae 1013: 1010: 1009: 1004: 1003:Colinatydidae 1001: 999:M. Sars, 1870 998: 997: 992: 989: 986: 985: 980: 977: 974: 973: 968: 965: 962: 961: 956: 953: 952: 951: 949: 942:2015 taxonomy 939: 937: 933: 929: 925: 921: 917: 916: 915:sensu stricto 910: 908: 904: 903: 895:2010 taxonomy 889: 885: 882: 881: 876: 875: 874:Smaragdinella 870: 869: 864: 863: 858: 857: 852: 848: 845: 841: 839: 838: 833: 832: 827: 826: 821: 817: 815: 814: 813:Philinoglossa 809: 805: 803: 802: 797: 796: 791: 790: 785: 784: 779: 778: 773: 769: 767: 766: 762:Gray, 1850 – 761: 757: 755: 754: 750:Gray, 1827 – 749: 745: 743: 742: 738:Dell, 1952 – 737: 733: 731: 730: 725: 724: 719: 715: 713: 712: 707: 706:Scaphandridae 703: 701: 700: 695: 691: 688: 687: 682: 681: 676: 672: 671: 670: 669: 668: 667:sensu stricto 663: 662:Cephalaspidea 656: 652: 650: 649: 644: 640: 639: 638: 636: 635: 627: 623: 620: 616: 613: 609: 607: 606: 601: 600: 596:Gray, 1847 – 595: 591: 589: 588: 583: 582: 577: 576: 571: 567: 566: 565: 563: 562: 557: 555: 551: 550: 541: 538: 537:Scaphandridae 534: 531: 527: 525: 521: 520: 519: 517: 510:2009 taxonomy 507: 505: 498: 494: 490: 486: 484: 480: 476: 472: 468: 464: 460: 456: 452: 450: 446: 442: 438: 434: 431: 427: 423: 419: 417: 413: 409: 408: 407: 405: 401: 394:2005 taxonomy 386: 385:Ringiculoidea 382: 379: 376: 372: 370:Pilsbry, 1895 369: 365: 362: 358: 356: 352: 349: 345: 342: 339: 335: 332: 329: 325: 324: 322: 321: 315: 313: 312: 307: 306: 301: 297: 293: 292: 287: 286:Philinorbidae 283: 279: 275: 274:Colinatydidae 271: 267: 262: 253: 251: 247: 243: 238: 236: 235: 229: 227: 223: 219: 209: 200: 197: 193: 189: 184: 182: 181:Bubble shells 178: 174: 171: 168: 164: 161:, is a major 160: 159:bubble snails 156: 152: 151:Cephalaspidea 143: 142: 137: 132: 129: 128:Cephalaspidea 126: 123: 122: 119: 116: 113: 112: 109: 106: 103: 102: 99: 96: 93: 92: 89: 86: 83: 82: 79: 76: 73: 72: 67: 62: 58: 54: 50: 49: 43: 39: 34: 29: 26: 22: 1474: 1435: 1430: 1426: 1404:(1): 23–41. 1401: 1397: 1359: 1355: 1345: 1302: 1298: 1260: 1254: 1209: 1205: 1167: 1145: 1124: 1103: 1085:Haminoeoidea 1081:polyphyletic 1077:Diaphanoidea 1066: 1055: 1046: 1037: 1018: 1006: 994: 982: 970: 958: 955:Acteocinidae 947: 945: 936:monophyletic 913: 911: 900: 898: 880:Ventomnestia 878: 872: 866: 860: 854: 835: 829: 825:Gastropteron 823: 811: 799: 793: 787: 781: 775: 763: 751: 739: 727: 721: 709: 697: 684: 678: 665: 661: 660: 646: 632: 631: 619:Ringiculidae 603: 597: 585: 579: 573: 559: 558: 547: 545: 515: 513: 501: 487:Superfamily 453:Superfamily 437:Haminoeoidea 435:Superfamily 422:Diaphanoidea 420:Superfamily 410:Superfamily 402:(2005), the 397: 383:Superfamily 373:Superfamily 368:Haminoeoidea 366:Superfamily 363:Odhner, 1914 361:Diaphanoidea 359:Superfamily 346:Superfamily 336:Superfamily 326:Superfamily 309: 303: 296:Acteocinidae 289: 263: 259: 239: 232: 230: 215: 206: 198: 185: 180: 158: 154: 150: 148: 140: 139: 127: 46: 25: 21:Cephalaspida 1592:iNaturalist 1499:Wikispecies 1362:: 133–149. 1305:: 130–150. 1131:Haminoeidae 1089:Philinoidea 1020:Philinorbis 996:Colpodaspis 967:Alacuppidae 926:within the 924:Acteonoidea 907:Haminoeidae 888:Bullactidae 851:Haminoeidae 844:Plusculidae 837:Siphopteron 801:Philinopsis 795:Odontoglaja 783:Chelidonura 694:Cylichnidae 686:Colpodaspis 675:Diaphanidae 637:Burn, 1963 594:Aplustridae 535:reinstated 528:reinstated 522:reinstated 504:Acteonoidea 491:: families 489:Runcinoidea 479:Plusculidae 467:Cylichnidae 457:: families 455:Philinoidea 445:Bullactidae 441:Haminoeidae 439:: families 426:Diaphanidae 424:: families 375:Philinoidea 328:Acteonoidea 311:Philinorbis 270:Alacuppidae 234:Chelidonura 222:polychaetes 118:Tectipleura 1738:Categories 1328:1956/10087 1212:(1): 323. 1182:References 1166:Couple of 1079:was found 1052:Habe, 1952 1043:Dall, 1908 1039:Cylichnium 1023:Habe, 1950 979:Mnestiidae 963:Gray, 1847 760:Philinidae 741:Volvulella 736:Rhizoridae 729:Pyrunculus 711:Scaphander 643:Runcinidae 634:Runcinacea 612:Bullinidae 570:Acteonidae 549:sensu lato 530:Runcinacea 516:sensu lato 493:Runcinidae 459:Philinidae 355:Sacoglossa 282:Mnestiidae 250:rhinophore 192:Euthyneura 149:The order 108:Gastropoda 1152:Aglajidae 1069:Bulloidea 1008:Colinatys 960:Acteocina 772:Aglajidae 718:Retusidae 605:Micromelo 581:Mexacteon 483:Retusidae 463:Aglajidae 414:: family 412:Bulloidea 378:J.E. Gray 338:Bulloidea 331:D'Orbigny 266:monophyly 170:gastropod 84:Kingdom: 78:Eukaryota 53:Aglajidae 1571:46447636 1490:Q1371035 1484:Wikidata 1337:25916189 1246:20973994 1110:Bullidae 1048:Micratys 1027:Laonidae 972:Alacuppa 862:Haminoea 748:Bullidae 699:Cylichna 683:, genus 680:Diaphana 655:Ilbiidae 599:Hydatina 497:Ilbiidae 416:Bullidae 300:Laonidae 291:Alacuppa 256:Taxonomy 173:mollusks 141:See text 114:Cohort: 98:Mollusca 94:Phylum: 88:Animalia 74:Domain: 1456:130-150 1307:Bibcode 1237:3087543 1214:Bibcode 984:Mnestia 886:Family 849:Family 842:Family 818:Family 806:Family 789:Navanax 770:Family 765:Philine 758:Family 746:Family 734:Family 716:Family 704:Family 692:Family 673:Family 653:Family 648:Runcina 641:Family 624:Family 617:Family 610:Family 592:Family 568:Family 398:In the 341:Lamarck 203:Anatomy 124:Order: 104:Class: 1708:425098 1705:uBio: 1672:272320 1656:NZOR: 1335:  1271:  1244:  1234:  1057:Noalda 777:Aglaja 723:Retusa 575:Acteon 380:, 1850 351:Thiele 343:, 1801 333:, 1835 167:marine 1716:WoRMS 1693:Plazi 1685:59609 1649:69554 1623:76047 1610:10689 1605:IRMNG 1597:49784 1558:7NF8J 1545:85093 1031:Laona 753:Bulla 404:clade 305:Laona 244:is a 163:taxon 1644:NCBI 1618:ITIS 1579:GBIF 1540:BOLD 1333:PMID 1269:ISBN 1242:PMID 1087:and 1054:and 856:Atys 587:Pupa 556:"): 495:and 481:and 447:and 428:and 240:The 157:and 1721:154 1631:NBN 1584:454 1566:EoL 1553:CoL 1527:AFD 1514:ADW 1440:doi 1406:doi 1364:doi 1323:hdl 1315:doi 1232:PMC 1222:doi 1176:). 1154:). 1133:). 1112:). 1740:: 1718:: 1695:: 1682:: 1669:: 1646:: 1633:: 1620:: 1607:: 1594:: 1581:: 1568:: 1555:: 1542:: 1529:: 1516:: 1501:: 1486:: 1436:44 1418:^ 1402:38 1400:. 1396:. 1378:^ 1360:14 1358:. 1354:. 1331:. 1321:. 1313:. 1303:89 1301:. 1297:. 1283:^ 1263:. 1240:. 1230:. 1220:. 1210:10 1208:. 1204:. 1190:^ 1045:, 938:. 909:. 877:, 871:, 865:, 859:, 834:, 828:, 798:, 792:, 786:, 780:, 726:, 602:, 584:, 578:, 518:: 477:, 473:, 469:, 465:, 461:, 443:, 308:, 298:, 284:, 280:, 276:, 272:, 252:. 179:. 1446:. 1442:: 1412:. 1408:: 1372:. 1366:: 1339:. 1325:: 1317:: 1309:: 1277:. 1248:. 1224:: 1216:: 1172:( 1150:( 1129:( 1108:( 1063:. 432:, 23:.

Index

Cephalaspida

Chelidonura varians
Aglajidae
Scientific classification
Edit this classification
Eukaryota
Animalia
Mollusca
Gastropoda
Tectipleura
Cephalaspidea
taxon
marine
gastropod
mollusks
Euopisthobranchia
opisthobranchs
Euthyneura
sensory structures
polychaetes
foraminiferans
Chelidonura
Hancock's organ
chemosensory organ
rhinophore
monophyly
Alacuppidae
Colinatydidae
Colpodaspididae

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