56:
1709:
1689:
3550:
3551:"Search results - 4[CHR] AND "Homo sapiens"[Organism] AND ( ("genetype miscrna"[Properties] OR "genetype ncrna"[Properties] OR "genetype rrna"[Properties] OR "genetype trna"[Properties] OR "genetype scrna"[Properties] OR "genetype snrna"[Properties] OR "genetype snorna"[Properties]) NOT "genetype protein coding"[Properties] AND alive[prop]) - Gene"
315:
The chromosome is ~193 megabases in length. In a 2012 paper, 775 protein-encoding genes were identified on this chromosome. 211 (27.9%) of these coding sequences did not have any experimental evidence at the protein level, in 2012. 271 appear to be membrane proteins. 54 have been classified as
1714:
G-banding patterns of human chromosome 4 in three different resolutions (400, 550 and 850). Band length in this diagram is based on the ideograms from ISCN (2013). This type of ideogram represents actual relative band length observed under a microscope at the different moments during the
3525:
3575:
40:
3314:
142:
3600:
Lemmers RJ, van der Vliet PJ, Klooster R, Sacconi S, Camaño P, Dauwerse JG, Snider L, Straasheijm KR, van Ommen GJ, Padberg GW, Miller DG, Tapscott SJ, Tawil R, Frants RR, van der Maarel SM (September 2010).
1409:
1694:
G-banding ideogram of human chromosome 4 in resolution 850 bphs. Band length in this diagram is proportional to base-pair length. This type of ideogram is generally used in genome browsers (e.g.
1321:
3385:
Chen LC, Liu MY, Hsiao YC, Choong WK, Wu HY, Hsu WL, Liao PC, Sung TY, Tsai SF, Yu JS, Chen YJ (2012) Decoding the disease-associated proteins encoded in the human chromosome 4. J Proteome Res
813:
1654:
1649:
4753:
3801:
These values (ISCN start/stop) are based on the length of bands/ideograms from the ISCN book, An
International System for Human Cytogenetic Nomenclature (2013).
3526:"Search results - 4[CHR] AND "Homo sapiens"[Organism] AND ("genetype protein coding"[Properties] AND alive[prop]) - Gene"
345:) takes an extremely conservative strategy. So CCDS's gene number prediction represents a lower bound on the total number of human protein-coding genes.
3576:"Search results - 4[CHR] AND "Homo sapiens"[Organism] AND ("genetype pseudo"[Properties] AND alive[prop]) - Gene"
1574:
590:
4254:
461:
214:
173:
608:
3315:"Search results - 4[CHR] AND "Homo sapiens"[Organism] AND ("has ccds"[Properties] AND alive[prop]) - Gene"
1538:
1478:
551:
630:
1614:
1220:
219:
4651:
1028:
4706:
4701:
1465:
499:
3863:
Goldfrank D, Schoenberger E, Gilbert F (2003). "Disease genes and chromosomes: disease maps of the human genome. Chromosome 4".
1548:
620:
4743:
4384:
4008:
3732:
3695:
3446:
3354:
1553:
494:
The following is a partial list of genes on human chromosome 4. For complete list, see the link in the infobox on the right.
4779:
4748:
1517:
1116:
489:
17:
329:
The following are some of the gene count estimates of human chromosome 4. Because researchers use different approaches to
4506:
3969:
1240:
1122:
976:
916:
4247:
1293:
4523:
398:
149:
3946:
1443:
419:
190:
1639:
1543:
1528:
1257:
1056:
880:
804:
3711:
Sethakulvichai, W.; Manitpornsut, S.; Wiboonrat, M.; Lilakiatsakun, W.; Assawamakin, A.; Tongsima, S. (2012).
4369:
1589:
39:
3756:
3671:
3658:
3373:
4774:
4240:
4190:
1666:
1599:
1533:
3475:
1198:
178:
876:
4516:
4001:
1624:
1569:
1375:
1369:
231:
207:
55:
4570:
4354:
4221:
1629:
1579:
788:
3683:
1644:
1564:
1490:
1413:
954:
596:
527:
4476:
4449:
1619:
1594:
1094:
894:
531:
295:. People normally have two copies of this chromosome. Chromosome 4 spans more than 190 million
251:
195:
4511:
4409:
3344:
1609:
377:
342:
270:
238:
137:
100:
544:
Glycerol-3-phosphate acyltransferase 3 a.k.a. 1-acylglycerol-3-phosphate O-acyltransferase 9
4731:
4656:
4641:
4471:
4419:
4394:
4334:
4209:
3994:
3907:
3614:
1110:
1098:
354:
8:
4399:
4216:
3894:
Hillier LW, Graves TA, Fulton RS, Fulton LA, Pepin KH, Minx P, et al. (April 2005).
1708:
1699:
1208:
940:
614:
226:
78:
3911:
3717:
2012 Ninth
International Conference on Computer Science and Software Engineering (JCSSE)
3618:
1474:
3738:
3635:
3602:
3423:
3396:
1688:
1202:
922:
123:
4461:
4373:
4184:
3925:
3880:
3728:
3712:
3691:
3640:
3428:
3350:
1584:
668:
330:
3742:
4550:
4429:
3915:
3872:
3786:
3720:
3630:
3622:
3454:
3418:
3408:
1661:
154:
991:: tyrosine-protein kinase KIT, CD117; mast/stem cell growth factor receptor (SCFR)
4543:
4364:
4312:
3685:
1604:
1507:
The following are some of the diseases related to genes located on chromosome 4:
1309:
1046:
338:
334:
118:
91:
3710:
341:). Among various projects, the collaborative consensus coding sequence project (
4434:
4358:
3876:
3802:
3724:
1523:
1512:
898:
890:
359:
304:
4768:
4414:
4147:
4142:
4137:
4132:
4127:
4122:
4117:
4112:
4107:
4102:
4097:
4092:
4087:
3896:"Generation and annotation of the DNA sequences of human chromosomes 2 and 4"
3413:
1559:
827:
704:
4232:
3687:
ISCN 2013: An
International System for Human Cytogenetic Nomenclature (2013)
3626:
3500:
166:
4389:
4263:
4204:
4170:
4165:
4082:
4077:
4072:
4067:
4062:
4052:
4047:
4042:
4017:
3929:
3884:
3684:
International
Standing Committee on Human Cytogenetic Nomenclature (2013).
3644:
3432:
1399:
1138:
1012:
866:
274:
255:
4608:
4603:
4466:
4444:
4439:
4307:
4025:
1254:: encoding protein RNA (guanine-9-) methyltransferase domain containing 2
1182:
3920:
3895:
4628:
4618:
4613:
4598:
4538:
4528:
4404:
4378:
4329:
4319:
4267:
3837:
3830:
3822:
988:
972:
960:
950:
946:
800:
364:
288:
3954:
1107:: NAD-dependent methylenetetrahydrofolate dehydrogenase 2-like protein
4580:
4324:
4302:
4283:
3818:
3603:"A unifying genetic model for facioscapulohumeral muscular dystrophy"
3501:"Human chromosome 4: entries, gene names and cross-references to MIM"
3397:"Between a chicken and a grape: estimating the number of human genes"
1727:
1387:
1381:
1313:
823:
521:
296:
63:
45:
4566:
4350:
4157:
4034:
3599:
1419:
1072:
982:
832:
602:
555:
111:
4736:
4590:
4501:
4496:
3713:"Estimation of band level resolutions of human chromosome images"
1716:
1695:
1494:
1447:
1437:
1363:
1357:
1303:
1251:
1230:
1166:
1162:
1132:
1104:
1066:
1050:
1040:
1016:
860:
854:
850:
844:
838:
755:
593:: encoding protein B cell scaffold protein with ankyrin repeats 1
564:
440:
264:
161:
4646:
4533:
4297:
4293:
4288:
3986:
1634:
1327:
1299:
1287:
1281:
1275:
1269:
1226:
1188:
1178:
1084:
966:
937:: encoding Huntington-like neurodegenerative disorder 2 protein
904:
817:
782:: Thought to be inactive but 2010 research shows a key role in
740:
731:
722:
611:: encoding protein Cytokine dependent hematopoietic cell linker
584:
574:
541:
537:
515:
303:) and represents between 6 and 6.5 percent of the total DNA in
292:
245:
202:
3757:
Ideogram data for Homo sapience (850 bphs, Assembly GRCh38.p3)
3704:
3672:
Ideogram data for Homo sapience (550 bphs, Assembly GRCh38.p3)
3659:
Ideogram data for Homo sapience (400 bphs, Assembly GRCh38.p3)
3374:
Ideogram data for Homo sapience (850 bphs, Assembly GRCh38.p3)
4711:
4696:
4691:
4686:
4681:
4676:
4671:
4666:
4661:
4636:
4574:
3829:: Region which is negatively stained by G banding, generally
1484:
1453:
1431:
1405:
1393:
1351:
1345:
1339:
1333:
1317:
1263:
1245:
1172:
1156:
1150:
1144:
1078:
1062:
994:
963:: linker protein for immunoglobulin alpha and mu polypeptides
886:
773:
761:
749:
713:
695:
686:
677:
671:: chemokine (C-X-C motif) ligand 4, Platelet factor-4, PF-4,
659:
650:
641:
626:
580:
560:
511:
505:
3862:
1223:: encoding protein Protein phosphatase with ef-hand domain 2
1470:
1459:
1425:
1236:
1214:
1194:
1128:
1090:
1034:
1031:: encoding protein Oral cancer overexpressed 1 pseudogene 1
1022:
1006:
1000:
934:
928:
910:
872:
809:
794:
783:
779:
767:
635:
570:
567:
Amyloid beta A4 precursor protein-binding family B member 2
547:
707:: chemokine (C-X-C motif) ligand 8, interleukin 8 (IL-8),
3944:
3893:
623:: encoding protein Zinc finger, cchc domain containing 23
300:
4754:
International System for Human
Cytogenetic Nomenclature
3342:
1119:: encoding protein Nucleosome assembly protein 1 like 5
758:: Defective in cullin neddylation 1 domain containing 4
1342:: Transforming acidic coiled-coil-containing protein 3
337:
on each chromosome varies (for technical details, see
3394:
1125:: encoding protein Neuron derived neurotrophic factor
919:: encoding protein FRY like transcription coactivator
629:: Collapsin response mediator protein 1, a member of
1390:: encoding protein Transmembrane serine protease 11F
997:: Ligand dependent nuclear receptor corepressor like
3974:Human Genome Project Information Archive 1990–2003
3785:For cytogenetic banding nomenclature, see article
1175:: Prostate androgen-regulated mucin-like protein 1
3476:"Chromosome 4: Chromosome summary - Homo sapiens"
3468:
1260:: encoding protein Stimulator of chondrogenesis 1
1069:Major facilitator superfamily domain containing 7
4766:
3677:
1402:: encoding protein TNFAIP3 interacting protein 3
835:: Family with sequence similarity 114, member A1
534:syndrome (Finnish heritage disease) related gene
3759:. Last update 2014-06-03. Retrieved 2017-04-26.
3674:. Last update 2015-08-11. Retrieved 2017-04-26.
3661:. Last update 2014-03-04. Retrieved 2017-04-26.
3376:. Last update 2014-06-03. Retrieved 2017-04-26.
3336:
863:: Family with sequence similarity 221, member B
847:: Family with sequence similarity 193, member A
841:: Family with sequence similarity 149, member A
3543:
1575:Hereditary benign intraepithelial dyskeratosis
4262:
4248:
4002:
3447:"Statistics & Downloads for chromosome 4"
1730:of human chromosome 4 in resolution 850 bphs
3690:. Karger Medical and Scientific Publishers.
1378:: encoding protein Transmembrane protein 243
1372:: encoding protein Transmembrane protein 175
1366:: encoding protein Transmembrane protein 165
1360:: encoding protein Transmembrane protein 155
776:: encoding protein Transmembrane protein 261
3568:
3439:
3388:
1539:Congenital central hypoventilation syndrome
1191:: Phosphatidylinositol 4-kinase type 2-beta
1169:Poly(A) binding protein, cytoplasmic 4-like
587:N-acylethanolamine-hydrolyzing acid amidase
4255:
4241:
4009:
3995:
3518:
3343:Tom Strachan; Andrew Read (2 April 2010).
1037:: Cell growth-regulating nucleolar protein
907:: fibroblast growth factor receptor-like 1
698:: chemokine (C-X-C motif) ligand 7, PPBP,
599:: encoding protein BEN domain containing 4
54:
38:
3919:
3634:
3422:
3412:
3368:
3366:
1681:G-banding ideograms of human chromosome 4
1615:Nonsyndromic deafness, autosomal dominant
1502:
1009:: Leucine-rich repeat LGI family member 2
502:: aminoadipate-semialdehyde dehydrogenase
48:. One is from mother, one is from father.
3817:: Region which is positively stained by
3593:
524:: encoding protein AFAP1 antisense RNA 1
3797:
3795:
3664:
3651:
3379:
1422:: encoding protein UGDH antisense RNA 1
1410:ubiquitin carboxyl-terminal esterase L1
889:: fibroblast growth factor receptor 3 (
14:
4767:
3493:
3363:
3309:
3307:
1549:Facioscapulohumeral muscular dystrophy
1330:: Spermatogenesis-associated protein 5
1217:: Serine/threonine-protein kinase PLK4
1211:: encoding protein Placenta specific 8
943:: encoding protein Helicase, POLQ-like
4744:List of organisms by chromosome count
4236:
3990:
1554:Fibrodysplasia ossificans progressiva
1322:non A4 component of amyloid precursor
1025:: U6 snRNA-associated Sm-like protein
925:: encoding protein Follistatin like 5
743:: chemokine (C-X-C motif) ligand 13,
734:: chemokine (C-X-C motif) ligand 11,
725:: chemokine (C-X-C motif) ligand 10,
518:Peroxisomal acyl-coenzyme A oxidase 3
3792:
1518:Autosomal dominant polycystic kidney
1462:: small subunit processome component
1384:: Transmembrane protease, serine 11D
716:: chemokine (C-X-C motif) ligand 9,
689:: chemokine (C-X-C motif) ligand 6,
680:: chemokine (C-X-C motif) ligand 5,
662:: chemokine (C-X-C motif) ligand 3,
653:: chemokine (C-X-C motif) ligand 2,
644:: chemokine (C-X-C motif) ligand 1,
605:: Coiled-coil domain containing 109B
490:Category:Genes on human chromosome 4
3304:
1673:
1487:: zinc-finger GRF-type containing 1
1296:: solute carrier family 10 member 4
1290:: solute carrier family 9 member B2
1233:Prosaposin-like 1 (gene/pseudogene)
24:
3855:
1440:: encoding UPF0602 Protein C4orf47
1354:: THAP domain-containing protein 6
1348:: Teneurin transmembrane protein 3
1306:Small integral membrane protein 20
1241:quinoid dihydropteridine reductase
985:: UV stimulated scaffold protein A
977:Vascular endothelial growth factor
324:
25:
4791:
3938:
1456:: ubiquitin specific peptidase 53
1159:: OTU domain-containing protein 4
975:: Kinase insert domain receptor (
4016:
3755:Genome Decoration Page, NCBI.
3670:Genome Decoration Page, NCBI.
3451:HUGO Gene Nomenclature Committee
3372:Genome Decoration Page, NCBI.
1707:
1687:
4365:Macrochromosome/Microchromosome
3945:National Institutes of Health.
3808:
3779:
3762:
3749:
3349:. Garland Science. p. 45.
1450:Ubiquitin specific peptidase 38
1396:: TNFAIP3-interaction protein 2
1141:: encoding protein Nephronectin
969:: Integrator complex subunit 12
508:: activin-like kinase 2 (ALK-2)
3657:Genome Decoration Page, NCBI.
3395:Pertea M, Salzberg SL (2010).
1640:Tetrahydrobiopterin deficiency
1529:Crouzonodermoskeletal syndrome
1248:: RNA binding motif protein 47
1003:: LIM domain-binding protein 2
881:basic fibroblast growth factor
92:
44:Human chromosome 4 pair after
13:
1:
3297:
1590:Mast cell activation syndrome
577:Ecto-ADP-ribosyltransferase 3
3453:. 2017-05-12. Archived from
1600:Mucopolysaccharidosis type I
1534:Chronic lymphocytic leukemia
1081:: non-coding RNA MicroRNA 95
1019:Uncharacterized LOC100505912
814:Ellis–Van Creveld syndrome 2
770:: Dickkopf-related protein 2
483:
316:cancer-associated proteins.
7:
4780:Genes on human chromosome 4
1428:: UDP glycosyltransferase 8
1336:: gene with protein product
1199:polycystic kidney disease 2
310:
10:
4796:
4405:Dinoflagellate chromosomes
3877:10.1089/109065703322783752
3725:10.1109/JCSSE.2012.6261965
1544:Ellis–Van Creveld syndrome
877:Fibroblast growth factor 2
869:: FAM47E-STBD1 readthrough
805:Ellis–Van Creveld syndrome
791:: Elmo domain-containing 2
487:
299:(the building material of
287:is one of the 23 pairs of
4749:List of sequenced genomes
4724:
4627:
4589:
4559:
4517:Chromosomal translocation
4487:
4390:A chromosome/B chromosome
4381:(or accessory chromosome)
4343:
4274:
4197:
4183:
4156:
4033:
4024:
1625:Polycystic kidney disease
1570:Hemolytic uremic syndrome
1135:Nitric oxide associated 1
333:their predictions of the
263:
244:
237:
225:
213:
201:
189:
184:
172:
160:
148:
136:
131:
117:
107:
90:
76:
71:
53:
37:
32:
4571:Telomere-binding protein
4385:Supernumerary chromosome
3414:10.1186/gb-2010-11-5-206
3346:Human Molecular Genetics
1667:Wolf–Hirschhorn syndrome
319:
4191:Human mitochondrial DNA
3951:Genetics Home Reference
3627:10.1126/science.1189044
1645:Thanatophoric dysplasia
1497:Zinc finger protein 621
1434:: netrin receptor UNC5C
1414:ubiquitin thiolesterase
4507:Structural alterations
3321:. CCDS Release 20 for
1595:Methylmalonic acidemia
1503:Diseases and disorders
1266:: protein SDA1 homolog
1113:: Myosin light chain 5
1095:methylmalonic aciduria
1087:: Centromere protein U
931:: GUF1 homolog, GTPase
895:thanatophoric dwarfism
764:: DEAH-box helicase 15
4524:Numerical alterations
4512:Chromosomal inversion
4410:Homologous chromosome
1610:Nonsyndromic deafness
1580:Hirschprung's disease
617:: Complement Factor I
4732:Extrachromosomal DNA
4420:Satellite chromosome
4395:Lampbrush chromosome
4335:Nuclear organization
4210:Human Genome Project
4185:Mitochondrial genome
3719:. pp. 276–282.
1630:Romano–Ward syndrome
1565:Huntington's disease
1185:transcription factor
1099:cobalamin deficiency
955:Huntington's disease
913:: FSHD region gene 1
360:Non-coding RNA genes
355:Protein-coding genes
185:External map viewers
18:Chromosome 4 (human)
4775:Chromosomes (human)
4425:Centromere position
4400:Polytene chromosome
4370:Circular chromosome
4217:List of human genes
3921:10.1038/nature03466
3912:2005Natur.434..724H
3844:: Variable region;
3619:2010Sci...329.1650L
1731:
1700:UCSC Genome Browser
1620:Parkinson's disease
1153:: nudix hydrolase 9
1147:: nudix hydrolase 6
1059:: Mastermind-like 3
615:Complement Factor I
132:Complete gene lists
119:Centromere position
3480:Ensembl Release 88
1726:
1475:Wolfram syndrome 1
1203:autosomal dominant
826:: Mutations cause
239:Full DNA sequences
4762:
4761:
4720:
4719:
4457:Centromere number
4374:Linear chromosome
4230:
4229:
4222:Human archaeology
4179:
4178:
3957:on August 3, 2004
3734:978-1-4673-1921-8
3697:978-3-318-02253-7
3356:978-1-136-84407-2
3295:
3294:
1585:Hypochondroplasia
1278:: Sec24 homolog D
1272:: Sec24 homolog B
752:: Cytokine-like 1
481:
480:
331:genome annotation
282:
281:
60:Chromosome 4 pair
16:(Redirected from
4787:
4587:
4586:
4551:Polyploidization
4379:Extra chromosome
4294:Genetic material
4257:
4250:
4243:
4234:
4233:
4031:
4030:
4011:
4004:
3997:
3988:
3987:
3983:
3981:
3980:
3965:
3963:
3962:
3953:. Archived from
3933:
3923:
3906:(7034): 724–31.
3888:
3849:
3812:
3806:
3799:
3790:
3783:
3777:
3766:
3760:
3753:
3747:
3746:
3708:
3702:
3701:
3681:
3675:
3668:
3662:
3655:
3649:
3648:
3638:
3613:(5999): 1650–3.
3597:
3591:
3590:
3588:
3587:
3572:
3566:
3565:
3563:
3562:
3547:
3541:
3540:
3538:
3537:
3522:
3516:
3515:
3513:
3512:
3497:
3491:
3490:
3488:
3487:
3472:
3466:
3465:
3463:
3462:
3443:
3437:
3436:
3426:
3416:
3392:
3386:
3383:
3377:
3370:
3361:
3360:
3340:
3334:
3333:
3331:
3330:
3311:
3285:
3280:
3253:
3248:
3220:
3215:
3188:
3183:
3155:
3150:
3123:
3118:
3090:
3085:
3058:
3053:
3025:
3020:
2993:
2988:
2960:
2955:
2928:
2923:
2895:
2890:
2863:
2858:
2830:
2825:
2798:
2793:
2765:
2760:
2733:
2728:
2700:
2695:
2668:
2663:
2635:
2630:
2603:
2598:
2570:
2565:
2538:
2533:
2505:
2500:
2473:
2468:
2440:
2435:
2408:
2403:
2375:
2370:
2343:
2338:
2310:
2305:
2278:
2273:
2245:
2240:
2213:
2208:
2181:
2176:
2149:
2144:
2117:
2112:
2084:
2079:
2052:
2047:
2019:
2014:
1987:
1982:
1954:
1949:
1922:
1917:
1889:
1884:
1857:
1852:
1824:
1819:
1792:
1732:
1725:
1711:
1691:
1674:Cytogenetic band
1662:Wolfram syndrome
1075:: microRNA 1269a
348:
347:
278:
259:
94:
58:
42:
30:
29:
27:Human chromosome
21:
4795:
4794:
4790:
4789:
4788:
4786:
4785:
4784:
4765:
4764:
4763:
4758:
4716:
4623:
4585:
4555:
4544:Paleopolyploidy
4489:
4483:
4339:
4313:Heterochromatin
4276:
4270:
4261:
4231:
4226:
4193:
4175:
4152:
4020:
4015:
3978:
3976:
3968:
3960:
3958:
3941:
3936:
3858:
3856:Further reading
3853:
3852:
3833:and gene rich;
3825:and gene poor;
3813:
3809:
3800:
3793:
3784:
3780:
3772:": Short arm; "
3767:
3763:
3754:
3750:
3735:
3709:
3705:
3698:
3682:
3678:
3669:
3665:
3656:
3652:
3598:
3594:
3585:
3583:
3574:
3573:
3569:
3560:
3558:
3549:
3548:
3544:
3535:
3533:
3524:
3523:
3519:
3510:
3508:
3499:
3498:
3494:
3485:
3483:
3474:
3473:
3469:
3460:
3458:
3445:
3444:
3440:
3393:
3389:
3384:
3380:
3371:
3364:
3357:
3341:
3337:
3328:
3326:
3313:
3312:
3305:
3300:
3283:
3278:
3251:
3246:
3218:
3213:
3186:
3181:
3153:
3148:
3121:
3116:
3088:
3083:
3056:
3051:
3023:
3018:
2991:
2986:
2958:
2953:
2926:
2921:
2893:
2888:
2861:
2856:
2828:
2823:
2796:
2791:
2763:
2758:
2731:
2726:
2698:
2693:
2666:
2661:
2633:
2628:
2601:
2596:
2568:
2563:
2536:
2531:
2503:
2498:
2471:
2466:
2438:
2433:
2406:
2401:
2373:
2368:
2341:
2336:
2308:
2303:
2276:
2271:
2243:
2238:
2211:
2206:
2179:
2174:
2147:
2142:
2115:
2110:
2082:
2077:
2050:
2045:
2017:
2012:
1985:
1980:
1952:
1947:
1920:
1915:
1887:
1882:
1855:
1850:
1822:
1817:
1790:
1760:
1755:
1750:
1745:
1724:
1723:
1722:
1721:
1720:
1717:mitotic process
1712:
1704:
1703:
1692:
1683:
1682:
1676:
1671:
1605:Muenke syndrome
1520:disease (PKD-2)
1505:
1500:
1043:: Mab-21-like 2
492:
486:
339:gene prediction
335:number of genes
327:
325:Number of genes
322:
313:
269:
250:
126:
85:
67:
61:
49:
28:
23:
22:
15:
12:
11:
5:
4793:
4783:
4782:
4777:
4760:
4759:
4757:
4756:
4751:
4746:
4741:
4740:
4739:
4728:
4726:
4722:
4721:
4718:
4717:
4715:
4714:
4709:
4704:
4699:
4694:
4689:
4684:
4679:
4674:
4669:
4664:
4659:
4654:
4649:
4644:
4639:
4633:
4631:
4625:
4624:
4622:
4621:
4616:
4611:
4606:
4601:
4595:
4593:
4584:
4583:
4578:
4563:
4561:
4557:
4556:
4554:
4553:
4548:
4547:
4546:
4541:
4536:
4531:
4521:
4520:
4519:
4514:
4504:
4499:
4493:
4491:
4485:
4484:
4482:
4481:
4480:
4479:
4474:
4469:
4464:
4454:
4453:
4452:
4447:
4442:
4437:
4435:Submetacentric
4432:
4422:
4417:
4412:
4407:
4402:
4397:
4392:
4387:
4382:
4376:
4367:
4362:
4361:or heterosome)
4355:Sex chromosome
4347:
4345:
4341:
4340:
4338:
4337:
4332:
4327:
4322:
4317:
4316:
4315:
4310:
4300:
4291:
4286:
4280:
4278:
4272:
4271:
4260:
4259:
4252:
4245:
4237:
4228:
4227:
4225:
4224:
4219:
4214:
4213:
4212:
4201:
4199:
4198:Related topics
4195:
4194:
4189:
4187:
4181:
4180:
4177:
4176:
4174:
4173:
4168:
4162:
4160:
4158:Sex chromosome
4154:
4153:
4151:
4150:
4145:
4140:
4135:
4130:
4125:
4120:
4115:
4110:
4105:
4100:
4095:
4090:
4085:
4080:
4075:
4070:
4065:
4060:
4055:
4050:
4045:
4039:
4037:
4028:
4026:Nuclear genome
4022:
4021:
4014:
4013:
4006:
3999:
3991:
3985:
3984:
3970:"Chromosome 4"
3966:
3947:"Chromosome 4"
3940:
3939:External links
3937:
3935:
3934:
3890:
3889:
3859:
3857:
3854:
3851:
3850:
3807:
3803:Arbitrary unit
3791:
3778:
3761:
3748:
3733:
3703:
3696:
3676:
3663:
3650:
3592:
3567:
3542:
3517:
3492:
3467:
3438:
3387:
3378:
3362:
3355:
3335:
3302:
3301:
3299:
3296:
3293:
3292:
3289:
3286:
3281:
3276:
3273:
3270:
3267:
3264:
3260:
3259:
3257:
3254:
3249:
3244:
3241:
3238:
3235:
3232:
3228:
3227:
3224:
3221:
3216:
3211:
3208:
3205:
3202:
3199:
3195:
3194:
3192:
3189:
3184:
3179:
3176:
3173:
3170:
3167:
3163:
3162:
3159:
3156:
3151:
3146:
3143:
3140:
3137:
3134:
3130:
3129:
3127:
3124:
3119:
3114:
3111:
3108:
3105:
3102:
3098:
3097:
3094:
3091:
3086:
3081:
3078:
3075:
3072:
3069:
3065:
3064:
3062:
3059:
3054:
3049:
3046:
3043:
3040:
3037:
3033:
3032:
3029:
3026:
3021:
3016:
3013:
3010:
3007:
3004:
3000:
2999:
2997:
2994:
2989:
2984:
2981:
2978:
2975:
2972:
2968:
2967:
2964:
2961:
2956:
2951:
2948:
2945:
2942:
2939:
2935:
2934:
2932:
2929:
2924:
2919:
2916:
2913:
2910:
2907:
2903:
2902:
2899:
2896:
2891:
2886:
2883:
2880:
2877:
2874:
2870:
2869:
2867:
2864:
2859:
2854:
2851:
2848:
2845:
2842:
2838:
2837:
2834:
2831:
2826:
2821:
2818:
2815:
2812:
2809:
2805:
2804:
2802:
2799:
2794:
2789:
2786:
2783:
2780:
2777:
2773:
2772:
2769:
2766:
2761:
2756:
2753:
2750:
2747:
2744:
2740:
2739:
2737:
2734:
2729:
2724:
2721:
2718:
2715:
2712:
2708:
2707:
2704:
2701:
2696:
2691:
2688:
2685:
2682:
2679:
2675:
2674:
2672:
2669:
2664:
2659:
2656:
2653:
2650:
2647:
2643:
2642:
2639:
2636:
2631:
2626:
2623:
2620:
2617:
2614:
2610:
2609:
2607:
2604:
2599:
2594:
2591:
2588:
2585:
2582:
2578:
2577:
2574:
2571:
2566:
2561:
2558:
2555:
2552:
2549:
2545:
2544:
2542:
2539:
2534:
2529:
2526:
2523:
2520:
2517:
2513:
2512:
2509:
2506:
2501:
2496:
2493:
2490:
2487:
2484:
2480:
2479:
2477:
2474:
2469:
2464:
2461:
2458:
2455:
2452:
2448:
2447:
2444:
2441:
2436:
2431:
2428:
2425:
2422:
2419:
2415:
2414:
2412:
2409:
2404:
2399:
2396:
2393:
2390:
2387:
2383:
2382:
2379:
2376:
2371:
2366:
2363:
2360:
2357:
2354:
2350:
2349:
2347:
2344:
2339:
2334:
2331:
2328:
2325:
2322:
2318:
2317:
2314:
2311:
2306:
2301:
2298:
2295:
2292:
2289:
2285:
2284:
2282:
2279:
2274:
2269:
2266:
2263:
2260:
2257:
2253:
2252:
2249:
2246:
2241:
2236:
2233:
2230:
2227:
2224:
2220:
2219:
2217:
2214:
2209:
2204:
2201:
2198:
2195:
2192:
2188:
2187:
2185:
2182:
2177:
2172:
2169:
2166:
2163:
2160:
2156:
2155:
2153:
2150:
2145:
2140:
2137:
2134:
2131:
2128:
2124:
2123:
2121:
2118:
2113:
2108:
2105:
2102:
2099:
2096:
2092:
2091:
2088:
2085:
2080:
2075:
2072:
2069:
2066:
2063:
2059:
2058:
2056:
2053:
2048:
2043:
2040:
2037:
2034:
2031:
2027:
2026:
2023:
2020:
2015:
2010:
2007:
2004:
2001:
1998:
1994:
1993:
1991:
1988:
1983:
1978:
1975:
1972:
1969:
1966:
1962:
1961:
1958:
1955:
1950:
1945:
1942:
1939:
1936:
1933:
1929:
1928:
1926:
1923:
1918:
1913:
1910:
1907:
1904:
1901:
1897:
1896:
1893:
1890:
1885:
1880:
1877:
1874:
1871:
1868:
1864:
1863:
1861:
1858:
1853:
1848:
1845:
1842:
1839:
1836:
1832:
1831:
1828:
1825:
1820:
1815:
1812:
1809:
1806:
1803:
1799:
1798:
1796:
1793:
1788:
1785:
1782:
1779:
1776:
1773:
1769:
1768:
1765:
1762:
1757:
1752:
1747:
1742:
1739:
1736:
1713:
1706:
1705:
1693:
1686:
1685:
1684:
1680:
1679:
1678:
1677:
1675:
1672:
1670:
1669:
1664:
1659:
1658:
1657:
1652:
1642:
1637:
1632:
1627:
1622:
1617:
1612:
1607:
1602:
1597:
1592:
1587:
1582:
1577:
1572:
1567:
1562:
1557:
1551:
1546:
1541:
1536:
1531:
1526:
1524:Bladder cancer
1521:
1515:
1513:Achondroplasia
1509:
1504:
1501:
1499:
1498:
1488:
1482:
1468:
1463:
1457:
1451:
1441:
1435:
1429:
1423:
1417:
1403:
1397:
1391:
1385:
1379:
1373:
1367:
1361:
1355:
1349:
1343:
1337:
1331:
1325:
1307:
1297:
1291:
1285:
1279:
1273:
1267:
1261:
1255:
1249:
1243:
1234:
1224:
1218:
1212:
1206:
1192:
1186:
1181:: codes for a
1176:
1170:
1160:
1154:
1148:
1142:
1136:
1126:
1120:
1114:
1108:
1102:
1088:
1082:
1076:
1070:
1060:
1054:
1044:
1038:
1032:
1026:
1020:
1010:
1004:
998:
992:
986:
980:
970:
964:
958:
949:(Huntingtin):
944:
938:
932:
926:
920:
914:
908:
902:
899:bladder cancer
891:achondroplasia
884:
870:
864:
858:
848:
842:
836:
830:
821:
807:
798:
792:
786:
777:
771:
765:
759:
753:
747:
738:
729:
720:
711:
702:
693:
684:
675:
666:
657:
648:
639:
633:
624:
618:
612:
606:
600:
594:
588:
578:
568:
558:
545:
535:
525:
519:
509:
503:
496:
485:
482:
479:
478:
475:
473:
470:
467:
464:
458:
457:
454:
452:
449:
446:
443:
437:
436:
433:
431:
428:
425:
422:
416:
415:
412:
410:
407:
404:
401:
395:
394:
391:
389:
386:
383:
380:
374:
373:
370:
367:
362:
357:
352:
326:
323:
321:
318:
312:
309:
280:
279:
267:
261:
260:
248:
242:
241:
235:
234:
229:
223:
222:
217:
211:
210:
205:
199:
198:
193:
187:
186:
182:
181:
176:
170:
169:
164:
158:
157:
152:
146:
145:
140:
134:
133:
129:
128:
124:Submetacentric
121:
115:
114:
109:
105:
104:
97:
88:
87:
84:193,574,945 bp
82:
74:
73:
69:
68:
62:in human male
59:
51:
50:
43:
35:
34:
26:
9:
6:
4:
3:
2:
4792:
4781:
4778:
4776:
4773:
4772:
4770:
4755:
4752:
4750:
4747:
4745:
4742:
4738:
4735:
4734:
4733:
4730:
4729:
4727:
4723:
4713:
4710:
4708:
4705:
4703:
4700:
4698:
4695:
4693:
4690:
4688:
4685:
4683:
4680:
4678:
4675:
4673:
4670:
4668:
4665:
4663:
4660:
4658:
4655:
4653:
4650:
4648:
4645:
4643:
4640:
4638:
4635:
4634:
4632:
4630:
4626:
4620:
4617:
4615:
4612:
4610:
4607:
4605:
4602:
4600:
4597:
4596:
4594:
4592:
4588:
4582:
4579:
4576:
4572:
4568:
4565:
4564:
4562:
4558:
4552:
4549:
4545:
4542:
4540:
4537:
4535:
4532:
4530:
4527:
4526:
4525:
4522:
4518:
4515:
4513:
4510:
4509:
4508:
4505:
4503:
4500:
4498:
4495:
4494:
4492:
4490:and evolution
4486:
4478:
4475:
4473:
4470:
4468:
4465:
4463:
4460:
4459:
4458:
4455:
4451:
4448:
4446:
4443:
4441:
4438:
4436:
4433:
4431:
4428:
4427:
4426:
4423:
4421:
4418:
4416:
4415:Isochromosome
4413:
4411:
4408:
4406:
4403:
4401:
4398:
4396:
4393:
4391:
4388:
4386:
4383:
4380:
4377:
4375:
4371:
4368:
4366:
4363:
4360:
4356:
4352:
4349:
4348:
4346:
4342:
4336:
4333:
4331:
4328:
4326:
4323:
4321:
4318:
4314:
4311:
4309:
4306:
4305:
4304:
4301:
4299:
4295:
4292:
4290:
4287:
4285:
4282:
4281:
4279:
4273:
4269:
4265:
4258:
4253:
4251:
4246:
4244:
4239:
4238:
4235:
4223:
4220:
4218:
4215:
4211:
4208:
4207:
4206:
4203:
4202:
4200:
4196:
4192:
4188:
4186:
4182:
4172:
4169:
4167:
4164:
4163:
4161:
4159:
4155:
4149:
4146:
4144:
4141:
4139:
4136:
4134:
4131:
4129:
4126:
4124:
4121:
4119:
4116:
4114:
4111:
4109:
4106:
4104:
4101:
4099:
4096:
4094:
4091:
4089:
4086:
4084:
4081:
4079:
4076:
4074:
4071:
4069:
4066:
4064:
4061:
4059:
4056:
4054:
4051:
4049:
4046:
4044:
4041:
4040:
4038:
4036:
4032:
4029:
4027:
4023:
4019:
4012:
4007:
4005:
4000:
3998:
3993:
3992:
3989:
3975:
3971:
3967:
3956:
3952:
3948:
3943:
3942:
3931:
3927:
3922:
3917:
3913:
3909:
3905:
3901:
3897:
3892:
3891:
3886:
3882:
3878:
3874:
3871:(4): 351–72.
3870:
3866:
3861:
3860:
3847:
3843:
3839:
3836:
3832:
3828:
3824:
3820:
3816:
3811:
3804:
3798:
3796:
3788:
3782:
3775:
3771:
3765:
3758:
3752:
3744:
3740:
3736:
3730:
3726:
3722:
3718:
3714:
3707:
3699:
3693:
3689:
3688:
3680:
3673:
3667:
3660:
3654:
3646:
3642:
3637:
3632:
3628:
3624:
3620:
3616:
3612:
3608:
3604:
3596:
3581:
3577:
3571:
3556:
3552:
3546:
3531:
3527:
3521:
3506:
3502:
3496:
3481:
3477:
3471:
3457:on 2017-06-29
3456:
3452:
3448:
3442:
3434:
3430:
3425:
3420:
3415:
3410:
3406:
3402:
3398:
3391:
3382:
3375:
3369:
3367:
3358:
3352:
3348:
3347:
3339:
3324:
3320:
3316:
3310:
3308:
3303:
3290:
3287:
3282:
3277:
3274:
3271:
3268:
3265:
3262:
3261:
3258:
3255:
3250:
3245:
3242:
3239:
3236:
3233:
3230:
3229:
3225:
3222:
3217:
3212:
3209:
3206:
3203:
3200:
3197:
3196:
3193:
3190:
3185:
3180:
3177:
3174:
3171:
3168:
3165:
3164:
3160:
3157:
3152:
3147:
3144:
3141:
3138:
3135:
3132:
3131:
3128:
3125:
3120:
3115:
3112:
3109:
3106:
3103:
3100:
3099:
3095:
3092:
3087:
3082:
3079:
3076:
3073:
3070:
3067:
3066:
3063:
3060:
3055:
3050:
3047:
3044:
3041:
3038:
3035:
3034:
3030:
3027:
3022:
3017:
3014:
3011:
3008:
3005:
3002:
3001:
2998:
2995:
2990:
2985:
2982:
2979:
2976:
2973:
2970:
2969:
2965:
2962:
2957:
2952:
2949:
2946:
2943:
2940:
2937:
2936:
2933:
2930:
2925:
2920:
2917:
2914:
2911:
2908:
2905:
2904:
2900:
2897:
2892:
2887:
2884:
2881:
2878:
2875:
2872:
2871:
2868:
2865:
2860:
2855:
2852:
2849:
2846:
2843:
2840:
2839:
2835:
2832:
2827:
2822:
2819:
2816:
2813:
2810:
2807:
2806:
2803:
2800:
2795:
2790:
2787:
2784:
2781:
2778:
2775:
2774:
2770:
2767:
2762:
2757:
2754:
2751:
2748:
2745:
2742:
2741:
2738:
2735:
2730:
2725:
2722:
2719:
2716:
2713:
2710:
2709:
2705:
2702:
2697:
2692:
2689:
2686:
2683:
2680:
2677:
2676:
2673:
2670:
2665:
2660:
2657:
2654:
2651:
2648:
2645:
2644:
2640:
2637:
2632:
2627:
2624:
2621:
2618:
2615:
2612:
2611:
2608:
2605:
2600:
2595:
2592:
2589:
2586:
2583:
2580:
2579:
2575:
2572:
2567:
2562:
2559:
2556:
2553:
2550:
2547:
2546:
2543:
2540:
2535:
2530:
2527:
2524:
2521:
2518:
2515:
2514:
2510:
2507:
2502:
2497:
2494:
2491:
2488:
2485:
2482:
2481:
2478:
2475:
2470:
2465:
2462:
2459:
2456:
2453:
2450:
2449:
2445:
2442:
2437:
2432:
2429:
2426:
2423:
2420:
2417:
2416:
2413:
2410:
2405:
2400:
2397:
2394:
2391:
2388:
2385:
2384:
2380:
2377:
2372:
2367:
2364:
2361:
2358:
2355:
2352:
2351:
2348:
2345:
2340:
2335:
2332:
2329:
2326:
2323:
2320:
2319:
2315:
2312:
2307:
2302:
2299:
2296:
2293:
2290:
2287:
2286:
2283:
2280:
2275:
2270:
2267:
2264:
2261:
2258:
2255:
2254:
2250:
2247:
2242:
2237:
2234:
2231:
2228:
2225:
2222:
2221:
2218:
2215:
2210:
2205:
2202:
2199:
2196:
2193:
2190:
2189:
2186:
2183:
2178:
2173:
2170:
2167:
2164:
2161:
2158:
2157:
2154:
2151:
2146:
2141:
2138:
2135:
2132:
2129:
2126:
2125:
2122:
2119:
2114:
2109:
2106:
2103:
2100:
2097:
2094:
2093:
2089:
2086:
2081:
2076:
2073:
2070:
2067:
2064:
2061:
2060:
2057:
2054:
2049:
2044:
2041:
2038:
2035:
2032:
2029:
2028:
2024:
2021:
2016:
2011:
2008:
2005:
2002:
1999:
1996:
1995:
1992:
1989:
1984:
1979:
1976:
1973:
1970:
1967:
1964:
1963:
1959:
1956:
1951:
1946:
1943:
1940:
1937:
1934:
1931:
1930:
1927:
1924:
1919:
1914:
1911:
1908:
1905:
1902:
1899:
1898:
1894:
1891:
1886:
1881:
1878:
1875:
1872:
1869:
1866:
1865:
1862:
1859:
1854:
1849:
1846:
1843:
1840:
1837:
1834:
1833:
1829:
1826:
1821:
1816:
1813:
1810:
1807:
1804:
1801:
1800:
1797:
1794:
1789:
1786:
1783:
1780:
1777:
1774:
1771:
1770:
1766:
1763:
1758:
1753:
1748:
1743:
1740:
1737:
1734:
1733:
1729:
1718:
1710:
1701:
1697:
1690:
1668:
1665:
1663:
1660:
1656:
1653:
1651:
1648:
1647:
1646:
1643:
1641:
1638:
1636:
1633:
1631:
1628:
1626:
1623:
1621:
1618:
1616:
1613:
1611:
1608:
1606:
1603:
1601:
1598:
1596:
1593:
1591:
1588:
1586:
1583:
1581:
1578:
1576:
1573:
1571:
1568:
1566:
1563:
1561:
1560:Haemophilia C
1558:
1555:
1552:
1550:
1547:
1545:
1542:
1540:
1537:
1535:
1532:
1530:
1527:
1525:
1522:
1519:
1516:
1514:
1511:
1510:
1508:
1496:
1492:
1489:
1486:
1483:
1480:
1476:
1472:
1469:
1467:
1464:
1461:
1458:
1455:
1452:
1449:
1445:
1442:
1439:
1436:
1433:
1430:
1427:
1424:
1421:
1418:
1415:
1411:
1407:
1404:
1401:
1398:
1395:
1392:
1389:
1386:
1383:
1380:
1377:
1374:
1371:
1368:
1365:
1362:
1359:
1356:
1353:
1350:
1347:
1344:
1341:
1338:
1335:
1332:
1329:
1326:
1323:
1319:
1315:
1311:
1308:
1305:
1301:
1298:
1295:
1292:
1289:
1286:
1283:
1280:
1277:
1274:
1271:
1268:
1265:
1262:
1259:
1256:
1253:
1250:
1247:
1244:
1242:
1238:
1235:
1232:
1228:
1225:
1222:
1219:
1216:
1213:
1210:
1207:
1204:
1200:
1196:
1193:
1190:
1187:
1184:
1180:
1177:
1174:
1171:
1168:
1164:
1161:
1158:
1155:
1152:
1149:
1146:
1143:
1140:
1137:
1134:
1130:
1127:
1124:
1121:
1118:
1115:
1112:
1109:
1106:
1103:
1100:
1096:
1092:
1089:
1086:
1083:
1080:
1077:
1074:
1071:
1068:
1064:
1061:
1058:
1055:
1053:MARCKS-like 1
1052:
1048:
1045:
1042:
1039:
1036:
1033:
1030:
1027:
1024:
1021:
1018:
1014:
1011:
1008:
1005:
1002:
999:
996:
993:
990:
987:
984:
981:
978:
974:
971:
968:
965:
962:
959:
956:
952:
948:
945:
942:
939:
936:
933:
930:
927:
924:
921:
918:
915:
912:
909:
906:
903:
900:
896:
892:
888:
885:
882:
878:
874:
871:
868:
865:
862:
859:
856:
852:
849:
846:
843:
840:
837:
834:
831:
829:
828:Haemophilia C
825:
822:
819:
815:
811:
808:
806:
802:
799:
796:
793:
790:
787:
785:
781:
778:
775:
772:
769:
766:
763:
760:
757:
754:
751:
748:
746:
742:
739:
737:
733:
730:
728:
724:
721:
719:
715:
712:
710:
706:
703:
701:
697:
694:
692:
688:
685:
683:
679:
676:
674:
670:
667:
665:
661:
658:
656:
652:
649:
647:
643:
640:
638:: Beta-casein
637:
634:
632:
628:
625:
622:
619:
616:
613:
610:
607:
604:
601:
598:
595:
592:
589:
586:
582:
579:
576:
572:
569:
566:
562:
559:
557:
553:
549:
546:
543:
539:
536:
533:
529:
526:
523:
520:
517:
513:
510:
507:
504:
501:
498:
497:
495:
491:
476:
474:
471:
468:
465:
463:
460:
459:
455:
453:
450:
447:
444:
442:
439:
438:
434:
432:
429:
426:
423:
421:
418:
417:
413:
411:
408:
405:
402:
400:
397:
396:
392:
390:
387:
384:
381:
379:
376:
375:
372:Release date
371:
368:
366:
363:
361:
358:
356:
353:
351:Estimated by
350:
349:
346:
344:
340:
336:
332:
317:
308:
306:
302:
298:
294:
290:
286:
276:
272:
268:
266:
262:
257:
253:
249:
247:
243:
240:
236:
233:
230:
228:
224:
221:
218:
216:
212:
209:
206:
204:
200:
197:
194:
192:
188:
183:
180:
177:
175:
171:
168:
165:
163:
159:
156:
153:
151:
147:
144:
141:
139:
135:
130:
125:
122:
120:
116:
113:
110:
106:
102:
98:
96:
89:
83:
80:
75:
70:
65:
57:
52:
47:
41:
36:
31:
19:
4456:
4424:
4264:Cytogenetics
4205:Human genome
4057:
4018:Human genome
3977:. Retrieved
3973:
3959:. Retrieved
3955:the original
3950:
3903:
3899:
3868:
3864:
3845:
3841:
3834:
3826:
3821:, generally
3814:
3810:
3781:
3776:": Long arm.
3773:
3769:
3764:
3751:
3716:
3706:
3686:
3679:
3666:
3653:
3610:
3606:
3595:
3584:. Retrieved
3582:. 2017-05-19
3579:
3570:
3559:. Retrieved
3557:. 2017-05-19
3554:
3545:
3534:. Retrieved
3532:. 2017-05-19
3529:
3520:
3509:. Retrieved
3507:. 2018-02-28
3504:
3495:
3484:. Retrieved
3482:. 2017-03-29
3479:
3470:
3459:. Retrieved
3455:the original
3450:
3441:
3404:
3400:
3390:
3381:
3345:
3338:
3327:. Retrieved
3325:. 2016-09-08
3323:Homo sapiens
3322:
3318:
1506:
1013:LOC100505912
867:FAM47E-STBD1
857:Protein ENED
744:
735:
726:
717:
708:
699:
690:
681:
672:
663:
654:
645:
493:
328:
314:
285:Chromosome 4
284:
283:
232:Chromosome 4
220:Chromosome 4
208:Chromosome 4
196:Chromosome 4
33:Chromosome 4
4477:Polycentric
4467:Monocentric
4450:Holocentric
4445:Acrocentric
4440:Telocentric
4430:Metacentric
4308:Euchromatin
4268:chromosomes
3401:Genome Biol
3284:190,214,555
3279:186,200,001
3252:186,200,000
3247:182,300,001
3219:182,300,000
3214:176,600,001
3187:176,600,000
3182:175,400,001
3154:175,400,000
3149:171,000,001
3122:171,000,000
3117:169,200,001
3089:169,200,000
3084:163,600,001
3057:163,600,000
3052:160,800,001
3024:160,800,000
3019:154,600,001
2992:154,600,000
2987:150,200,001
2959:150,200,000
2954:147,500,001
2927:147,500,000
2922:145,900,001
2894:145,900,000
2889:140,600,001
2862:140,600,000
2857:138,500,001
2829:138,500,000
2824:130,100,001
2797:130,100,000
2792:127,900,001
2764:127,900,000
2759:122,800,001
2732:122,800,000
2727:119,900,001
2699:119,900,000
2694:113,200,001
2667:113,200,000
2662:106,700,001
2634:106,700,000
2629:100,100,001
2602:100,100,000
1493:: encoding
1446:: encoding
1302:: encoding
1284:: Septin-11
1229:: encoding
1183:homeodomain
1165:: encoding
1131:: encoding
1101:) cblA type
1065:: encoding
1049:: encoding
979:receptor 2)
853:: encoding
797:: Endomucin
631:CRMP family
583:: encoding
573:: encoding
563:: encoding
540:: encoding
514:: encoding
477:2017-05-19
456:2018-02-28
435:2017-03-29
414:2017-05-12
393:2016-09-08
365:Pseudogenes
289:chromosomes
4769:Categories
4629:Centromere
4560:Structures
4539:Polyploidy
4529:Aneuploidy
4330:Nucleosome
4320:Chromosome
3979:2017-05-06
3961:2017-05-06
3865:Genet Test
3838:Centromere
3586:2017-05-20
3561:2017-05-20
3536:2017-05-20
3511:2018-03-16
3486:2017-05-19
3461:2017-05-19
3407:(5): 206.
3329:2017-05-28
3298:References
2597:97,900,001
2569:97,900,000
2564:94,200,001
2537:94,200,000
2532:92,800,001
2504:92,800,000
2499:87,100,001
2472:87,100,000
2467:86,000,001
2439:86,000,000
2434:83,200,001
2407:83,200,000
2402:81,500,001
2374:81,500,000
2369:78,000,001
2342:78,000,000
2337:75,300,001
2309:75,300,000
2304:69,400,001
2277:69,400,000
2272:65,500,001
2244:65,500,000
2239:58,500,001
2212:58,500,000
2207:51,800,001
2180:51,800,000
2175:50,000,001
2148:50,000,000
2143:48,200,001
2116:48,200,000
2111:44,600,001
2083:44,600,000
2078:41,200,001
2051:41,200,000
2046:35,800,001
2018:35,800,000
2013:27,700,001
1986:27,700,000
1981:21,300,001
1953:21,300,000
1948:17,700,001
1921:17,700,000
1916:15,000,001
1888:15,000,000
1883:11,300,001
1856:11,300,000
951:huntingtin
621:COL25A1-DT
488:See also:
297:base pairs
127:(50.0 Mbp)
4581:Protamine
4488:Processes
4472:Dicentric
4325:Chromatid
4303:Chromatin
4284:Karyotype
3819:G banding
1851:6,000,001
1823:6,000,000
1818:4,500,001
1791:4,500,000
1479:wolframin
1388:TMPRSS11F
1382:TMPRSS11D
1314:synuclein
1015:encoding
953:protein (
824:Factor XI
552:ankyrin 2
522:AFAP1-AS1
484:Gene list
252:NC_000004
179:Gene list
167:Gene list
155:Gene list
143:Gene list
64:karyogram
46:G-banding
4725:See also
4567:Telomere
4534:Euploidy
4462:Acentric
4359:allosome
4351:Autosome
4277:concepts
4035:Autosome
3930:15815621
3885:15000816
3848:: Stalk.
3743:16666470
3645:20724583
3433:20441615
1767:Density
1759:Basepair
1754:Basepair
1420:UGDH-AS1
1073:MIR1269A
1047:Marcksl1
983:KIAA1530
833:FAM114A1
603:CCDC109B
556:neuronal
311:Genomics
271:CM000666
112:Autosome
95:of genes
77:Length (
72:Features
4737:Plasmid
4591:Histone
4502:Meiosis
4497:Mitosis
3908:Bibcode
3831:CG-rich
3823:AT-rich
3636:4677822
3615:Bibcode
3607:Science
3505:UniProt
3424:2898077
1728:G-bands
1696:Ensembl
1495:protein
1448:protein
1438:UPF0602
1376:TMEM243
1370:TMEM175
1364:TMEM165
1358:TMEM155
1304:protein
1294:SLC10A4
1252:RG9MTD2
1231:protein
1167:protein
1163:PABPC4L
1133:protein
1105:MTHFD2L
1067:protein
1051:protein
1041:MAB21L2
1017:protein
861:FAM221B
855:protein
851:FAM198B
845:FAM193A
839:FAM149A
756:DCUN1D4
565:protein
441:UniProt
420:Ensembl
369:Source
265:GenBank
191:Ensembl
162:UniProt
86:(CHM13)
4298:Genome
4289:Ploidy
3928:
3900:Nature
3883:
3741:
3731:
3694:
3643:
3633:
3431:
3421:
3353:
1764:Stain
1756:start
1746:start
1655:Type 2
1650:Type 1
1635:SADDAN
1491:ZNF621
1328:SPATA5
1300:SMIM20
1288:SLC9B2
1282:SEPT11
1276:SEC24D
1270:SEC24B
1227:PSAPL1
1189:PI4K2B
1179:PHOX2B
1117:NAP1L5
1085:MLF1IP
1029:LTO1P1
967:INTS12
905:FGFRL1
818:limbin
789:ELMOD2
745:scyb13
741:CXCL13
736:scyb11
732:CXCL11
727:scyb10
723:CXCL10
585:enzyme
575:enzyme
542:enzyme
538:AGPAT9
516:enzyme
293:humans
246:RefSeq
203:Entrez
4575:TINF2
4344:Types
4275:Basic
3846:stalk
3787:locus
3739:S2CID
3275:11170
3272:10967
3243:10967
3240:10628
3210:10628
3207:10408
3178:10408
3175:10341
3145:10341
3142:10087
3113:10087
2944:31.23
2912:31.22
2879:31.21
2424:21.23
2392:21.22
2359:21.21
1938:15.31
1906:15.32
1873:15.33
1761:stop
1751:stop
1741:Band
1735:Chr.
1556:(FOP)
1485:ZGRF1
1466:VPS54
1454:USP53
1444:USP38
1432:UNC5C
1406:UCHL1
1400:TNIP3
1394:TNIP2
1352:THAP6
1346:TENM3
1340:TACC3
1334:STATH
1318:alpha
1264:SDAD1
1258:SCRG1
1246:RBM47
1221:PPEF2
1209:PLAC8
1173:PARM1
1157:OTUD4
1151:NUDT9
1145:NUDT6
1079:MIR95
1063:MFSD7
1057:MAML3
995:LCORL
923:FSTL5
887:FGFR3
774:DMAC1
762:DHX15
750:CYTL1
718:scyb9
714:CXCL9
709:scyb8
705:CXCL8
700:scyb7
696:CXCL7
691:scyb6
687:CXCL6
682:scyb5
678:CXCL5
673:scyb4
669:CXCL4
664:scyb3
660:CXCL3
655:scyb2
651:CXCL2
646:scyb1
642:CXCL1
627:CRMP1
597:BEND4
591:BANK1
581:ASAHL
561:APBB2
512:ACOX3
506:ACVR1
500:AASDH
320:Genes
305:cells
275:FASTA
256:FASTA
99:727 (
4357:(or
3926:PMID
3881:PMID
3835:acen
3827:gneg
3815:gpos
3729:ISBN
3692:ISBN
3641:PMID
3580:NCBI
3555:NCBI
3530:NCBI
3429:PMID
3351:ISBN
3319:NCBI
3288:gpos
3269:35.2
3256:gneg
3237:35.1
3226:100
3223:gpos
3204:34.3
3191:gneg
3172:34.2
3158:gpos
3139:34.1
3126:gneg
3110:9985
3096:100
3093:gpos
3080:9985
3077:9681
3074:32.3
3061:gneg
3048:9681
3045:9545
3042:32.2
3031:100
3028:gpos
3015:9545
3012:9207
3009:32.1
2996:gneg
2983:9207
2980:9004
2977:31.3
2963:gpos
2950:9004
2947:8851
2931:gneg
2918:8851
2915:8733
2898:gpos
2885:8733
2882:8581
2866:gneg
2853:8581
2850:8259
2847:31.1
2836:100
2833:gpos
2820:8259
2817:7734
2814:28.3
2801:gneg
2788:7734
2785:7565
2782:28.2
2768:gpos
2755:7565
2752:7277
2749:28.1
2736:gneg
2723:7277
2720:7040
2703:gpos
2690:7040
2687:6685
2671:gneg
2658:6685
2655:6347
2638:gpos
2625:6347
2622:6059
2606:gneg
2593:6059
2590:5890
2573:gpos
2560:5890
2557:5636
2554:22.3
2541:gneg
2528:5636
2525:5517
2522:22.2
2508:gpos
2495:5517
2492:5145
2489:22.1
2476:gneg
2463:5145
2460:4874
2457:21.3
2443:gpos
2430:4874
2427:4739
2411:gneg
2398:4739
2395:4671
2378:gpos
2365:4671
2362:4502
2346:gneg
2333:4502
2330:4333
2327:21.1
2313:gpos
2300:4333
2297:4062
2294:13.3
2281:gneg
2268:4062
2265:3910
2262:13.2
2251:100
2248:gpos
2235:3910
2232:3571
2229:13.1
2216:gneg
2203:3571
2200:3249
2184:acen
2171:3249
2168:3046
2152:acen
2139:3046
2136:2877
2120:gneg
2107:2877
2104:2742
2087:gpos
2074:2742
2071:2505
2055:gneg
2042:2505
2039:2166
2025:100
2022:gpos
2009:2166
2006:1811
2003:15.1
1990:gneg
1977:1811
1974:1557
1971:15.2
1957:gpos
1944:1557
1941:1286
1925:gneg
1912:1286
1909:1066
1892:gpos
1879:1066
1860:gneg
1841:16.1
1827:gpos
1808:16.2
1795:gneg
1778:16.3
1749:ISCN
1744:ISCN
1738:Arm
1471:WFS1
1460:UTP3
1426:UGT8
1310:SNCA
1237:QDPR
1215:PLK4
1195:PKD2
1139:NPNT
1129:NOA1
1123:NDNF
1111:MYL5
1091:MMAA
1035:LYAR
1023:LSM6
1007:LGI2
1001:LDB2
941:HELQ
935:HDL3
929:GUF1
917:FRYL
911:FRG1
873:FGF2
810:EVC2
795:EMCN
784:FSHD
780:DUX4
768:DKK2
636:CSN2
609:CLNK
571:ART3
548:ANK2
472:819
462:NCBI
430:727
409:633
399:HGNC
378:CCDS
343:CCDS
227:UCSC
215:NCBI
174:NCBI
150:HGNC
138:CCDS
108:Type
101:CCDS
4609:H2B
4604:H2A
3916:doi
3904:434
3873:doi
3842:var
3721:doi
3631:PMC
3623:doi
3611:329
3419:PMC
3409:doi
3291:25
3161:75
2966:25
2901:25
2771:50
2706:75
2641:50
2576:75
2511:75
2446:25
2381:50
2316:75
2090:50
1960:75
1895:50
1876:779
1847:779
1844:389
1830:25
1814:389
1811:220
1784:220
989:KIT
973:KDR
961:IGJ
947:HTT
801:EVC
532:AGU
528:AGA
469:934
466:769
445:765
427:993
424:746
406:277
403:731
382:727
301:DNA
291:in
93:No.
4771::
4652:C2
4647:C1
4619:H4
4614:H3
4599:H1
4569::
4266::
4148:22
4143:21
4138:20
4133:19
4128:18
4123:17
4118:16
4113:15
4108:14
4103:13
4098:12
4093:11
4088:10
3972:.
3949:.
3924:.
3914:.
3902:.
3898:.
3879:.
3867:.
3840:.
3794:^
3737:.
3727:.
3715:.
3639:.
3629:.
3621:.
3609:.
3605:.
3578:.
3553:.
3528:.
3503:.
3478:.
3449:.
3427:.
3417:.
3405:11
3403:.
3399:.
3365:^
3317:.
3306:^
3266:q
3234:q
3201:q
3169:q
3136:q
3107:33
3104:q
3071:q
3039:q
3006:q
2974:q
2941:q
2909:q
2876:q
2844:q
2811:q
2779:q
2746:q
2717:27
2714:q
2684:26
2681:q
2652:25
2649:q
2619:24
2616:q
2587:23
2584:q
2551:q
2519:q
2486:q
2454:q
2421:q
2389:q
2356:q
2324:q
2291:q
2259:q
2226:q
2197:12
2194:q
2165:11
2162:q
2133:11
2130:p
2101:12
2098:p
2068:13
2065:p
2036:14
2033:p
2000:p
1968:p
1935:p
1903:p
1870:p
1838:p
1805:p
1775:p
1702:).
1698:,
1473::
1408::
1316:,
1312::
1239::
1197::
1093::
897:,
893:,
875::
812::
803::
554:,
550::
530::
451:—
388:—
307:.
277:)
258:)
79:bp
4712:T
4707:Q
4702:P
4697:O
4692:N
4687:M
4682:K
4677:J
4672:I
4667:H
4662:F
4657:E
4642:B
4637:A
4577:)
4573:(
4372:/
4353:/
4296:/
4256:e
4249:t
4242:v
4171:Y
4166:X
4083:9
4078:8
4073:7
4068:6
4063:5
4058:4
4053:3
4048:2
4043:1
4010:e
4003:t
3996:v
3982:.
3964:.
3932:.
3918::
3910::
3887:.
3875::
3869:7
3805:.
3789:.
3774:q
3770:p
3768:"
3745:.
3723::
3700:.
3647:.
3625::
3617::
3589:.
3564:.
3539:.
3514:.
3489:.
3464:.
3435:.
3411::
3359:.
3332:.
3263:4
3231:4
3198:4
3166:4
3133:4
3101:4
3068:4
3036:4
3003:4
2971:4
2938:4
2906:4
2873:4
2841:4
2808:4
2776:4
2743:4
2711:4
2678:4
2646:4
2613:4
2581:4
2548:4
2516:4
2483:4
2451:4
2418:4
2386:4
2353:4
2321:4
2288:4
2256:4
2223:4
2191:4
2159:4
2127:4
2095:4
2062:4
2030:4
1997:4
1965:4
1932:4
1900:4
1867:4
1835:4
1802:4
1787:1
1781:0
1772:4
1719:.
1481:)
1477:(
1416:)
1412:(
1324:)
1320:(
1205:)
1201:(
1097:(
957:)
901:)
883:)
879:(
820:)
816:(
448:—
385:—
273:(
254:(
103:)
81:)
66:.
20:)
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