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Jennifer Doudna

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the decision. In 2017, the court decided in favor of the Broad Institute, who claimed that they had initiated the research earliest and had first applied it to human cell engineering thus supporting editing in human cells with evidence but that the UC Berkeley group had only suggested this application. UC Berkeley appealed on grounds that they had clearly discussed and spelled out how to do the application the Broad had pursued. In September 2018, the appeals court decided in favor of the Broad Institute's patent. Meanwhile, UC Berkeley and co-applicants' patent to cover the general technique was also granted. To further cloud the issue, in Europe the claim of the Broad Institute, to have initiated the research first, was disallowed. The rejection was due to a procedural flaw in the application involving a different set of personnel listed in the lawsuit and the patent application, leading to speculation that the UC Berkeley group would prevail in Europe. Doudna cofounded Caribou Biosciences, a company to commercialize CRISPR technology, in 2011. In September 2013, Doudna cofounded
40: 4891: 4819: 2260: 750: 737: 500: 807: 816: 1029:, and other colleagues at the Innovative Genomics Institute (IGI), where they created a testing center. This center processed over 500,000 patient samples from UC Berkeley students, staff and faculty as well as members of the surrounding community and farm workers in the Salinas area. Mammoth Biosciences announced a peer-reviewed validation of a rapid, CRISPR-based point of need COVID-19 diagnostic which is faster and less expensive than 4784: 952: 5166: 766:. They showed that a core of five magnesium ions clustered in one region of the P4-P6 domain of the ribozyme, forming a hydrophobic core around which the rest of the structure could fold. This is analogous but chemically distinct from, the way proteins typically have a core of hydrophobic amino acids. Her group has crystallized other ribozymes, including the 1013:, a San Francisco-based bioengineering tech startup. Initial funding raised $ 23 million, with a series B round of funding in 2020 raising $ 45 million. The business is focused on improving access to bio sensing tests which address "challenges across healthcare, agriculture, environmental monitoring, biodefense, and more." 603:, Doudna's interest in science was nurtured by her 10th-grade chemistry teacher, Ms. Jeanette Wong, whom she has routinely cited as a significant influence in sparking her nascent scientific curiosity. A visiting lecturer on cancer cells further encouraged her pursuit of science as a career choice. She spent a summer working in the 1001:"I have so much optimism about what CRISPR can do to help cure unaddressed genetic diseases and improve sustainable agriculture, but I'm also concerned that the benefits of the technology might not reach those who need it most if we're not thoughtful and deliberate about how we develop the technology," Doudna said. 595:
which was a major inspiration. Doudna also developed her interest in science and mathematics in school. Even though Doudna was told that "Women don't go into science," she knew that she wanted to be a scientist no matter what. Nothing said to her made her doubt it, Doudna said, "When someone tells me
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in 2006 who had found Doudna by way of a Google search, having typed "RNAi and UC Berkeley" into her browser, and Doudna's name came up at the top of the list. In 2012, Doudna and her colleagues made a new discovery that reduces the time and work needed to edit genomic DNA. Their discovery relies on
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at the Broad Institute had shown that CRISPR-Cas9 could edit genes in cultured human cells a few months after Doudna and Charpentier published their method. Before the UC Berkeley patent application was decided, a patent was granted to the Broad investigators and UC Berkeley filed a lawsuit against
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into a true catalytic ribozyme that copied RNA templates. Her focus was on engineering ribozymes and understanding their underlying mechanisms; however, she came to realize that not being able to see the molecular mechanisms of ribozymes was a major problem. Doudna went to the lab of Thomas Cech at
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As CRISPR becomes increasingly used to edit multicellular organisms, Doudna continues to be called upon to serve as a thought-leader on the ethics of changing an organism's function using CRISPR technology. Their discovery has since been further developed by many research groups for applications
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Chancellor's Professorship in Biomedicine and Health, and is the chair of the Chancellor's Advisor Committee on Biology. Her lab now focuses on the structure and function of CRISPR-Cas systems, developing new genome editing technology and delivery mechanisms for CRISPR therapeutics, and novel
580:. Nature built her sense of curiosity and her desire to understand the underlying biological mechanisms of life. This was coupled with the atmosphere of intellectual pursuit that her parents encouraged at home. Her father enjoyed reading about science and filled the home with many books on 630:. During her freshman year, while taking a course in general chemistry, she questioned her own ability to pursue a career in science, and considered switching her major to French as a sophomore. However, her French teacher suggested she stick with science. Chemistry professors 1114:. Cate is a Berkeley professor and works on gene-editing yeast to increase their cellulose fermentation for biofuel production. Doudna and Cate have a son born in 2002 who attends UC Berkeley, studying electrical engineering and computer science. They live in Berkeley. 1352: 911:
bacterial "CRISPR" immune system that cooperates with guide RNA and works like scissors. The protein attacks its prey, the DNA of viruses, and slices it up, preventing it from infecting the bacterium. This system was first discovered by
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Doudna's first marriage was in 1988 to a fellow graduate student at Harvard named Tom Griffin, but his interests were more broad and less focused on research than hers and they divorced a few years later. Griffin wanted to move to
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and Doudna followed him to Boston at Harvard, but in 2002 they both accepted faculty positions at Berkeley and moved there together; Cate preferred the less formal environment on the West Coast from his earlier experiences at the
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for her doctoral study and earned a PhD in Biological Chemistry and Molecular Pharmacology in 1989. Her Ph.D. dissertation was on a system that increased the efficiency of a self-replicating catalytic
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at Pomona had a major impact on her. She started her first scientific research in the lab of professor Sharon Panasenko. She earned her Bachelor of Arts degree in biochemistry in 1985. She chose
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The CRISPR system created a new straightforward way to edit DNA and there was a rush to patent the technique. Doudna and UC Berkeley collaborators applied for a patent and so did a group at the
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Jennifer Doudna was born February 19, 1964, in Washington, D.C., as the daughter of Dorothy Jane (Williams) and Martin Kirk Doudna. Her father received his PhD in English literature from the
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founded by Doudna to develop genome editing technology and apply it to some of society's greatest problems in human health, agriculture and climate change. Doudna holds the
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virus utilizes an unusual strategy to synthesize viral proteins. This work could lead to new drugs to stop infections without causing harm to the tissues of the body.
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Jeff Akst, (2018) "The higher court's decision to uphold the ruling of the Patent Trial and Appeal Board essentially ends the intellectual property battle in theUS"
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She is also the founder and chair of the governance board of the Innovative Genomics Institute, which she co-founded in 2014. Doudna is also a faculty scientist at
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Doudna is on the scientific advisory boards of the companies that she cofounded, such as Caribou, Intellia, Mammoth, and Scribe; as well as others such as
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Group I intron P4-P6 catalytic region. Doudna brought Cate with her to Yale, and they married in Hawaii in 2000. Cate later became a professor at the
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to crystallize and determine the three-dimensional structure of a ribozyme for the first time, so ribozyme structure could be compared with that of
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Professor of Molecular Biophysics and Biochemistry at Yale in 2000. In 2000–2001, she was Robert Burns Woodward Visiting Professor of Chemistry at
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in 2016 alongside other CRISPR researchers. In 2018 and 2023, she received honorary Doctor of Science degrees from USC and Harvard, respectively.
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At Yale, Doudna's group was able to crystallize and solve the three-dimensional structure of the catalytic core of the
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in 1996. Doudna joined Yale's Department of Molecular Biophysics and Biochemistry as an assistant professor in 1994.
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Read "Enhancing Philanthropy's Support of Biomedical Scientists: Proceedings of a Workshop on Evaluation" at NAP.edu
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and Scribe Therapeutics, which pioneered CasX, a more compact, next-generation Cas9 which can efficiently cut DNA.
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with Zhang and others despite their legal battles, but she quit in June 2014; Charpentier then invited her to join
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Cohen, Jon (June 4, 2018). "With prestigious prize, an overshadowed CRISPR researcher wins the spotlight".
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showed for the first time that they could use different RNAs to program it to cut and edit different DNAs.
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Giorgia Guglielmi (June 1, 2018). "Million-dollar Kavli prize recognizes scientist scooped on CRISPR".
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ranging from fundamental cell biology, plant, and animal research to treatments for diseases including
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Beginning in March 2020, Doudna organized an effort to use CRISPR-based technologies to address the
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Early in her scientific career, Doudna worked to uncover the structure and biological function of
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One or more of the preceding sentences incorporates text from the royalsociety.org website where:
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As of 2023, Doudna was located at the University of California, Berkeley, where she directs the
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for CRISPR-Cas9 genome editing technology, with Charpentier. She has been a co-recipient of the
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Shape of a self-splicing intron with two exons (shown in red and blue). DS Goodsell, 2005, PDB
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in 2022 as their chief science advisor, to guide investment decisions related to CRISPR.
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and in genetics at Harvard Medical School. From 1991 to 1994, she was Lucille P. Markey
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I can't do something and I know that I can, it just makes me more resolved to do it."
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revolution" for her fundamental work and leadership in developing CRISPR-mediated
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The Code Breaker: Jennifer Doudna, Gene Editing, and the Future of the Human Race
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A Crack in Creation: Gene Editing and the Unthinkable Power to Control Evolution.
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In 2012, Doudna and Emmanuelle Charpentier were the first to propose that CRISPR-
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A Crack in Creation: Gene Editing and the Unthinkable Power to Control Evolution
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Proceedings of the National Academy of Sciences of the United States of America
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Portrait of Jennifer Doudna by Christopher Michel for the National Academies
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CRISPR Scientist's Biography Explores Ethics Of Rewriting The Code Of Life
4250: 2914: 2786:"CRISPR co-inventor Jennifer Doudna talks ethics and biological frontiers" 2771: 2717: 2709: 2647: 2174: 576:, Doudna was fascinated by the environmental beauty of the island and its 11544:
University of California, Berkeley College of Letters and Science faculty
11363: 11346: 11334: 10838: 10794: 10655: 10521: 10490: 10454: 10387: 10275: 10204: 9946: 9836: 9440: 9342: 9273: 9245: 9239: 9235: 9184: 9178: 9110: 9077: 8875: 8857: 8831: 8805: 8745: 8574: 8516: 8512: 8422: 8293: 8289: 8212: 8177: 8045: 7985: 7960: 7900: 7835: 7781: 7514: 7484: 7389: 7288: 7240: 7107: 6639: 6343: 6195: 6179: 6159: 6155: 5932: 5904: 5900: 5864: 5832: 5784: 5752: 5285: 5273: 5253: 5139: 5100: 5042: 3417: 1224: 1200: 1191:
for Biochemistry and Biophysics. She has also been a co-recipient of the
1094: 1090: 1049:, and an adjunct professor of cellular and molecular pharmacology at the 995: 878: 759: 675: 539: 456: 361: 343: 203: 199: 193: 9046: 1278:
in 2014. In 2015, together with Charpentier, she became a fellow of the
11387: 11375: 11221: 11129: 11098: 11009: 10985: 10752: 10699: 10578: 10269: 9990: 9938: 9907: 9390: 9281: 9215: 9148: 8849: 8698: 8592: 8498: 8480: 8404: 8331: 8085: 7965: 7745: 7621: 7597: 7264: 7252: 7218: 7095: 6789: 6095: 6031: 6011: 5936: 5744: 5649: 5597: 5521: 5402: 5398: 5226: 1588:"Cracking the Code: Jennifer Doudna and Her Amazing Molecular Scissors" 1345:"Pondering 'what it means to be human' on the frontier of gene editing" 1240: 1196: 1172: 1057: 994:
In addition to the CRISPR breakthrough, Doudna has discovered that the
968: 883: 531: 459: 440: 210: 187: 745:
X-ray diffraction-based structure of active site of a ribozyme at Yale
729:. She started this project at the Cech lab in 1991 and finished it at 10852: 10713: 10689: 10138: 9897: 9706: 9130: 9006: 8937: 8446: 8380: 8182: 7573: 7125: 6759: 5716: 5342: 4864: 3675:"Meet Altos Labs, Silicon Valley's latest wild bet on living forever" 2846: 2547:"The CRISPR Pioneers: Their Breakthrough Work Could Change the World" 1171:/Cas9 genome editing technology. In 2016, together with Charpentier, 858:
after two months and returned to Berkeley with the help of colleague
855: 851: 723: 462: 4045:. The Royal Netherlands Academy of Arts and Sciences. Archived from 3802:"The Welch Foundation supports basic research in chemistry in Texas" 3724:"Jennifer Doudna, Crispr scientist, on the ethics of editing humans" 3137:"Caribou Biosciences Announces Co-Founding of Intellia Therapeutics" 2500:
Jill Banfield: How a curious Google search led me to Jennifer Doudna
9702: 9120: 8303: 4446:"Nobel Prize Awarded to ASM Members for Development of CRISPR-Cas9" 3932: 1314: 1286:
in 2016. In 2017, Doudna was awarded the Golden Plate Award of the
1145: 706:. While in the Szostak lab, Doudna re-engineered the self-splicing 703: 608: 549:
most influential people in 2015, and in 2023 was inducted into the
543: 519: 465:
in the department of chemistry and the department of molecular and
455:, "for the development of a method for genome editing." She is the 11569:
Yale Department of Molecular Biophysics & Biochemistry faculty
7143: 3482:"UC Berkeley's Jennifer Doudna wins 2020 Nobel Prize in Chemistry" 2528:(published October 7, 2020, to the UC Berkeley channel on YouTube) 1867:"2023 Inductee Jennifer Doudna | National Inventors Hall of Fame®" 1412:
Wu, Katherine J.; Zimmer, Carl; Peltier, Elian (October 7, 2020).
8050: 1247:
in the category of Welfare Betterment. In 2020, she received the
726: 679: 9877:
Princess of Asturias Award for Technical and Scientific Research
7877: 3601:"Crispr's Next Frontier Is In-Human Treatment, Co-Inventor Says" 3065:"Broad Institute takes a hit in European CRISPR patent struggle" 2578:
Martin Jínek; Krzysztof Chylinski; Ines Fonfara; Michael Hauer;
2273:"Scopus preview – Doudna, Jennifer A. – Author details – Scopus" 1987:"Remembering The Hilo Teacher Who Inspired A Nobel Prize Winner" 820:"CRISPR Biology and Biotechnology: The Future of Genome Editing" 8674: 4856:. University of California Television (UCTV). October 26, 2019. 4151:"Jennifer Doudna To Receive 2018 Pearl Meister Greengard Prize" 3654: 3576:"Jennifer Doudna's first East Bay CRISPR spinout files for IPO" 3557: 2347:"CRISPR pioneer Doudna envisions ending asthma, aiding climate" 1615:"Jennifer Doudna, a Pioneer Who Helped Simplify Genome Editing" 1168: 945: 895: 863: 719: 687: 489: 310: 253: 124: 9512:
Prince of Asturias Award for Technical and Scientific Research
4876:"How CRISPR lets us edit our DNA". TED talk by Jennifer Doudna 4531:"Golden Plate Awardees of the American Academy of Achievement" 4268:. American Association for the Advancement of Science (AAAS). 3503:"Jennifer Doudna Opens Laboratory at the Gladstone Institutes" 2664:; H Shinagawa; K Makino; M Amemura; A Nakata (December 1987). 850:
In 2009, she took a leave of absence from Berkeley to work at
693: 4941: 4850:"Into the Future with CRISPR Technology with Jennifer Doudna" 4182:"Doudna receives Medal of Honor from American Cancer Society" 3290: 1320:
most influential people in 2015, and she was a runner-up for
422: 3418:"CRISPR pioneer Doudna opens COVID-19 testing lab – BioNews" 951: 584:. When Doudna was in the sixth grade, he gave her a copy of 10079: 9915: 9844: 8506: 6609: 3852:"CRISPR Pioneer Doudna Joins Global Investment Firm as CSO" 3751:"US Biotech Firms Made China's Gene-Edited Babies Possible" 3628: 3321:"Mammoth Biosciences Closes on Series A Worth $ 23 Million" 3117: 2813:"Scientists Seek Ban on Method of Editing the Human Genome" 1929:. American Society for Biochemistry and Molecular Biology. 904: 481: 431: 2509:, Jill Banfield (Berkeley News, published October 7, 2020) 387: 11524:
Members of the United States National Academy of Sciences
4868: 4599:"John Simon Guggenheim Foundation | Jennifer Doudna" 3878: 3866: 2519:
Jennifer Doudna's First Reactions to 2020 Nobel Prize Win
1805:"Laureates of the Japan Prize: Jennifer A. Doudna, Ph.D." 1255:). Also in 2020, Doudna and Charpentier were awarded the 847:
for her experiments with high powered x-ray diffraction.
699: 643: 258: 4844:. Innovative Genomics Institute – IGI. November 4, 2017. 1227:
in Nanoscience (jointly with Emmanuelle Charpentier and
3484:. University of California, Berkeley. October 7, 2020. 3373: 2324: 1302:
appointed Doudna, and two other female Nobel laureates
1235:
of the Technion/Israel for the year 2018 (jointly with
889: 588:'s 1968 book on the discovery of the structure of DNA, 4498:
Creative Commons Attribution 4.0 International License
1259:"for the development of a method for genome editing." 522:, as determined by X-ray crystallography and the 2015 2190:"Pomona College alumna wins Nobel Prize in Chemistry" 1124:
List of awards and honors received by Jennifer Doudna
428: 425: 419: 11388:
Dejusticia: The Center for Law, Justice and Society
4565:"Jennifer A. Doudna, Ph.D. Biography and Interview" 4204: 3527:"Interview with Jennifer Doudna (recorded in 2004)" 1650:"Alan T. Waterman Award Recipients, 1976 – present" 1089:, where Doudna was also interested in working with 416: 3776: 3722: 3599: 3574: 2985:Decker, Susan; Cortez, Michelle (April 28, 2018). 2214: 1449:"Press release: The Nobel Prize in Chemistry 2020" 944:on any clinical application of gene editing using 916:and colleagues in 1987 and later characterized by 16:American biochemist and Nobel laureate (born 1964) 4210: 11425: 8263: 1313:She along with Charpentier was named one of the 835:In 2002, she joined her husband, Jamie Cate, at 11549:University of California, San Francisco faculty 11474:Fellows of the American Academy of Microbiology 9845:European Organization for Nuclear Research CERN 1828: 1826: 1411: 6341: 4901: 4503:"Royal Society Terms, conditions and policies" 3957:"Beckman Young Investigators Award Recipients" 3529:. National Academy of Sciences. Archived from 2110: 1407: 1405: 1306:and Emmanuelle Charpentier, as members of the 1284:Foreign Member of the Royal Society (ForMemRS) 11519:Members of the Pontifical Academy of Sciences 11114: 10506: 10065: 9482: 9062: 8249: 7863: 7159: 6595: 6327: 4927: 3219: 1750:"Laureates: Biopharmaceutical Science (2016)" 1553:Jennifer A. Doudna and Samuel H. Sternberg. 774:(IRES) and protein-RNA complexes such as the 11382:Bangladesh Environmental Lawyers Association 2984: 1823: 862:, canceling all of her obligations to study 116:First X-ray based structure of catalytic RNA 11514:Members of the National Academy of Medicine 4832:. iBiology Science Stories. March 23, 2015. 3285: 3283: 2106: 2104: 2102: 2100: 2098: 1920: 1918: 1916: 1914: 1773: 1771: 1606: 1402: 822:, Jennifer Doudna delivers the 29th annual 694:Research on ribozyme structure and function 556: 11121: 11107: 10520: 10513: 10499: 10072: 10058: 9489: 9475: 9069: 9055: 8256: 8242: 7870: 7856: 7166: 7152: 6602: 6588: 6334: 6320: 4934: 4920: 4889: 4863:. Author interview, audio and transcript. 4817: 3800:Seijo, Bibiana Campos (October 26, 2019). 2258: 2096: 2094: 2092: 2090: 2088: 2086: 2084: 2082: 2080: 2078: 1644: 1642: 38: 9076: 8641:The Event Horizon Telescope Collaboration 3500: 3028: 2896: 2699: 2629: 2181: 2156: 2138: 1501:. Lawrence Berkeley National Laboratory. 439:; born February 19, 1964) is an American 11128: 4735: 4257: 3884: 3872: 3824: 3672: 3379: 3280: 3256: 3254: 3123: 2330: 2187: 1984: 1911: 1768: 1207:(2017). In 2018, Doudna was awarded the 950: 748: 735: 498: 473:. She has been an investigator with the 6611:Eli Lilly Award in Biological Chemistry 4509:from the original on September 25, 2015 4420:American Academy of Arts & Sciences 4179: 4161:from the original on September 14, 2018 4131:from the original on September 13, 2018 4061: 3963:from the original on September 11, 2019 3959:. Arnold and Mabel Beckman Foundation. 3896: 3849: 3839:from the original on December 22, 2023. 3764:from the original on February 13, 2024. 3747: 3720: 3673:Regalado, Antonio (September 4, 2021). 3635:from the original on December 10, 2023. 3446: 3239:from the original on September 27, 2020 3105:from the original on September 12, 2018 3062: 2958: 2075: 1960:The Women in Medicine Legacy Foundation 1924: 1744: 1742: 1639: 1612: 1594:from the original on September 20, 2020 1585: 1496:"Curriculum Vitae (Jennifer A. Doudna)" 1378:"Jennifer Doudna – American biochemist" 1150:Eli Lilly Award in Biological Chemistry 1051:University of California, San Francisco 873:, a collaboration between Berkeley and 781:Doudna was promoted to the position of 618:Doudna was an undergraduate student at 518:for her research on the structure of a 301:University of California, San Francisco 11454:21st-century American women scientists 11444:20th-century American women scientists 11426: 4662:from the original on November 30, 2021 4545:from the original on December 12, 2017 4426:from the original on November 13, 2023 4396:from the original on November 13, 2023 4075:from the original on September 5, 2016 4069:"2015 Genetics Prize: Jennifer Doudna" 3911:from the original on February 4, 2023. 3789:from the original on November 2, 2021. 3737:from the original on January 31, 2020. 3622: 3612:from the original on October 20, 2019. 3547: 3501:Langelier, Julie (September 5, 2018). 2959:Netburn, Deborah (February 15, 2017). 2559:from the original on December 25, 2016 2300: 2298: 2223:Towards the Design of an RNA Replicase 2220: 2019:from the original on November 13, 2022 1993:from the original on November 13, 2022 1966:from the original on December 10, 2022 1933:from the original on December 31, 2019 1847:from the original on December 29, 2016 1727:from the original on September 5, 2016 1721:"2015 Genetics Prize: Jennifer Doudna" 1713: 1110:, and Doudna liked that Berkeley is a 1004: 653: 317:Towards the Design of an RNA Replicase 11102: 10494: 10053: 9875: 9510: 9501:for Technical and Scientific Research 9470: 9050: 8237: 7851: 7147: 6583: 6315: 4915: 4900: 4707:"Harvard awards six honorary degrees" 4653: 4452:from the original on October 18, 2021 4448:. American Society for Microbiology. 4366:from the original on January 12, 2020 4310:from the original on November 9, 2019 4263: 4121:"National Academy of Sciences Awards" 4101:from the original on February 3, 2019 3799: 3697: 3645: 3597: 3572: 3251: 3058: 3056: 3029:Paganelli, Jennifer (June 19, 2018). 2954: 2952: 2950: 2540: 2538: 2536: 2534: 2470: 2468: 2341: 2339: 2283:from the original on January 20, 2022 2202:from the original on October 24, 2020 2063:. UH Hilo Stories. September 19, 2018 1950: 1948: 1811:from the original on November 7, 2017 1756:from the original on November 7, 2017 1685: 1683: 1681: 1679: 1677: 1675: 1673: 1671: 1549: 1547: 1545: 1543: 1508:from the original on January 15, 2017 1459:from the original on January 15, 2021 1455:. Nobel Foundation. October 7, 2020. 1443: 1441: 1355:from the original on October 15, 2020 1268:American Academy of Arts and Sciences 1108:Lawrence Berkeley National Laboratory 1099:Massachusetts Institute of Technology 1043:Lawrence Berkeley National Laboratory 965:Massachusetts Institute of Technology 854:to lead discovery research. She left 845:Lawrence Berkeley National Laboratory 11504:Kavli Prize laureates in Nanoscience 11479:Foreign members of the Royal Society 9499:Prince or Princess of Asturias Award 4682:"Past Recipients – Honorary Degrees" 4579:from the original on August 12, 2020 4473:"Professor Jennifer Doudna ForMemRS" 4470: 4180:Sanders, Robert (October 18, 2018). 3897:Kreiger, Lisa M. (October 7, 2020). 3825:Regalado, Antonio (April 26, 2022). 3814:from the original on March 14, 2024. 3748:Molteni, Megan (November 30, 2018). 3721:Kuchler, Hannah (January 31, 2020). 3587:from the original on March 14, 2024. 3488:from the original on October 7, 2020 2810: 2544: 2225:(Ph.D. thesis). Harvard University. 2188:Engineer, Anushe (October 7, 2020). 1833:King, Mary-Claire (April 16, 2015). 1832: 1739: 1627:from the original on January 2, 2022 1581: 1579: 1577: 1575: 1573: 1571: 1569: 1567: 1565: 1563: 1541: 1539: 1537: 1535: 1533: 1531: 1529: 1527: 1525: 1523: 1490: 1488: 1486: 1484: 1482: 1480: 1478: 1476: 1474: 1426:from the original on October 8, 2020 1390:from the original on October 7, 2020 1372: 1370: 1223:. Also in 2018, she was awarded the 1117: 1104:University of California, Santa Cruz 1016: 890:CRISPR-Cas9 genome editing discovery 11494:Howard Hughes Medical Investigators 9627:Instituto Nacional de Biodiversidad 7173: 4635:from the original on March 27, 2023 4629:Association for Molecular Pathology 4605:from the original on April 13, 2020 4483:from the original on April 29, 2016 4192:from the original on August 2, 2020 4023:from the original on April 20, 2022 3987:Arnold and Mabel Beckman Foundation 3862:from the original on April 1, 2022. 3774: 3710:from the original on March 5, 2024. 3687:from the original on March 4, 2024. 3662:from the original on July 25, 2023. 3598:Darie, Tatiana (October 19, 2019). 3459:from the original on April 25, 2020 3397:Innovative Genomics Institute (IGI) 3331:from the original on April 15, 2020 3301:from the original on April 22, 2020 3207:from the original on April 12, 2020 3063:Servick, Kelly (January 18, 2018). 2480:Innovative Genomics Institute (IGI) 2452:Innovative Genomics Institute (IGI) 2402:Innovative Genomics Institute (IGI) 2312:from the original on April 12, 2020 2295: 2111:Melissa Marino (December 1, 2004). 1701:from the original on April 20, 2022 1586:Russell, Sabin (December 8, 2014). 1296:Association for Molecular Pathology 1185:Canada Gairdner International Award 1165:Breakthrough Prize in Life Sciences 1036: 792: 536:Canada Gairdner International Award 524:Breakthrough Prize in Life Sciences 190:in Biopharmaceutical Science (2016) 176:Breakthrough Prize in Life Sciences 13: 4654:Dulle, Colleen (August 18, 2021). 3850:Goodwin, Kate (February 8, 2022). 3623:Fannin, Rebecca (March 12, 2022). 3361:from the original on July 21, 2020 3075:from the original on March 1, 2022 3053: 2947: 2935:from the original on July 30, 2020 2792:from the original on March 8, 2019 2531: 2465: 2336: 2245: 1985:Cataluna, Lee (October 11, 2020). 1945: 1797: 1785:from the original on June 12, 2018 1668: 1656:from the original on March 2, 2015 1438: 471:University of California, Berkeley 291:University of California, Berkeley 14: 11580: 9651:Pedro Miguel Echenique Landiríbar 4775: 3775:Tan, Gillian (November 2, 2021). 3447:Release, Press (April 21, 2020). 3428:from the original on May 10, 2020 3177:from the original on May 16, 2021 3147:from the original on May 16, 2021 2823:from the original on May 31, 2020 2811:Wade, Nicholas (March 19, 2015). 2113:"Biography of Jennifer A. Doudna" 1560: 1520: 1471: 1367: 1187:. Also in 2016, she received the 1134:Beckman Young Investigators Award 882:techniques for precisely editing 542:(2017). She was named one of the 11559:Wolf Prize in Medicine laureates 5164: 4838:"Jennifer Doudna: CRISPR Basics" 4782: 4741:"Introduction – Into the Breach" 4699: 4674: 4647: 4617: 4591: 4557: 4523: 4464: 4438: 3291:"Mammoth Biosciences | About Us" 2997:from the original on May 9, 2018 2684:10.1128/JB.169.12.5429-5433.1987 1613:Pollack, Andrew (May 11, 2015). 1557:Houghton Mifflin Harcourt, 2017. 1280:American Academy of Microbiology 1219:, and a Medal of Honor from the 1079: 814: 805: 412: 10940:/ John Frederick Grassle (2013) 4729: 4573:American Academy of Achievement 4539:American Academy of Achievement 4408: 4378: 4352: 4322: 4296: 4173: 4143: 4113: 4087: 4035: 4005: 3975: 3949: 3915: 3890: 3843: 3818: 3807:Chemical & Engineering News 3793: 3768: 3741: 3714: 3691: 3666: 3639: 3616: 3591: 3566: 3471: 3440: 3410: 3385: 3343: 3313: 3201:Chemical & Engineering News 3189: 3159: 3129: 3087: 3022: 3009: 2978: 2921: 2853:; et al. (April 3, 2015). 2835: 2804: 2778: 2724: 2654: 2571: 2512: 2493: 2440: 2415: 2390: 2364: 2265: 2053: 2031: 2005: 1978: 1885: 1859: 1691:"Laureates: Jennifer A. Doudna" 1652:. National Science Foundation. 1288:American Academy of Achievement 1136:. In 2000, she was awarded the 1045:, a senior investigator at the 551:National Inventors Hall of Fame 475:Howard Hughes Medical Institute 281:University of Colorado, Boulder 146: 11449:21st-century American chemists 11439:20th-century American chemists 3698:Bayer, Max (January 7, 2024). 3554:Jennifer Doudna's publications 2732:Francisco Juan Martínez Mojica 2221:Doudna, Jennifer Anne (1989). 1956:"Becoming a Scientific Leader" 1781:. Canada Gairdner Foundation. 1383:Encyclopædia Britannica Online 1337: 1308:Pontifical Academy of Sciences 1209:NAS Award in Chemical Sciences 768:Hepatitis Delta Virus ribozyme 716:University of Colorado Boulder 672:University of Colorado Boulder 664:Massachusetts General Hospital 611:Don Hemmes and graduated from 1: 11554:Winners of the Heineken Prize 11489:Harvard Medical School alumni 10650:Ernest John Christopher Polge 3646:Knapp, Alex (July 25, 2023). 1330: 1290:. In 2020, she was awarded a 1276:National Academy of Inventors 1068:, Synthego, Tempus Labs, and 871:Innovative Genomics Institute 824:Ullyot Public Affairs Lecture 670:in Biomedical Science at the 505:Innovative Genomics Institute 11529:Nobel laureates in Chemistry 9175:Jørgen Christensen-Dalsgaard 8549:Sudbury Neutrino Observatory 4125:National Academy of Sciences 3581:San Francisco Business Times 2013:"Big Questions, Big Answers" 1807:The Japan Prize Foundation. 1272:National Academy of Medicine 1264:National Academy of Sciences 1231:). In 2019 she received the 1148:. In 2001, she received the 772:internal ribosome entry site 678:. As of 2022, Doudna has an 662:in molecular biology at the 605:University of Hawaii at Hilo 567:University of Hawaii at Hilo 340:Other academic advisors 7: 11183:(Jordan, USA, Saudi Arabia) 8473:Supernova Cosmology Project 7182:Biochemistry and Biophysics 3573:Leuty, Ron (July 1, 2021). 3262:"Jennifer A. Doudna, Ph.D." 1897:. Marquis-Who's Who. 1999. 1205:Albany Medical Center Prize 1142:National Science Foundation 1009:In 2017, Doudna co-founded 776:signal recognition particle 10: 11585: 11469:American crystallographers 11464:American women biochemists 8081:Lakshminarayanan Mahadevan 4903:Awards for Jennifer Doudna 4227:10.1038/D41586-018-05308-5 3095:"Caribou Biosciences, Inc" 2524:November 20, 2021, at the 1590:. Cal Alumni Association. 1121: 443:who has pioneered work in 182:Princess of Asturias Award 11356: 11303: 11191:Biopharmaceutical Science 11190: 11137: 11019: 10880: 10745: 10666:William Hayward Pickering 10610: 10541:Ephraim Katchalski-Katzir 10529: 10437: 10354: 10285: 10218: 10127: 10088: 9882: 9871: 9557:Antonio González González 9535:Luis Antonio Santaló Sors 9517: 9506: 9327: 9198: 9085: 8712: 8365: 8272: 8170: 8154: 7893: 7732: 7673: 7566: 7459: 7346: 7181: 6617: 6512: 6482: 6446: 6415: 6396: 6376: 6351: 5970: 5671: 5412: 5173: 5162: 4953: 4909: 4097:. Albany Medical Center. 4071:. The Gruber Foundation. 2748:10.1016/J.TIM.2016.06.005 2505:October 12, 2020, at the 2423:"Structure and Mechanism" 1925:Mukhopadyay, Rajendrani. 1752:. Tang Prize Foundation. 1723:. The Gruber Foundation. 1167:for her contributions to 1154:American Chemical Society 987:In 2017, she co-authored 894:Doudna was introduced to 828:Science History Institute 813: 804: 799: 382: 378: 349: 339: 327: 309: 273: 241: 234: 170:Jacob Heskel Gabbay Award 156: 133: 109: 78: 49: 37: 23: 11459:American Nobel laureates 9918:Scientific Collaboration 9623:Manuel Losada Villasante 9599:Francisco Bolívar Zapata 6088:Venkatraman Ramakrishnan 5367:Wendell Meredith Stanley 4947:Nobel Prize in Chemistry 4813:publications indexed by 3560:bibliographic database. 3035:Caribou Biosciences, Inc 2254:publications indexed by 1927:"On the same wavelength" 1257:Nobel Prize in Chemistry 1193:Gruber Prize in Genetics 1025:along with Dave Savage, 711:Group I catalytic intron 686:and of 111 according to 674:, where she worked with 658:After her PhD, she held 557:Early life and education 528:Gruber Prize in Genetics 449:Nobel Prize in Chemistry 394:Hughes Institute website 218:Nobel Prize in Chemistry 11539:Pomona College trustees 11163:Veerabhadran Ramanathan 11138:Sustainable Development 10807:/ Keith J. Sainsbury / 10009:Everett Peter Greenberg 9661:Enrique Moreno González 9605:Federico García Moliner 8967:Shankar Balasubramanian 8703:Alexander Zamolodchikov 8635:Peter van Nieuwenhuizen 4686:honorarydegrees.usc.edu 4274:10.1126/science.aau3736 3562:(subscription required) 2881:10.1126/SCIENCE.AAB1028 2671:Journal of Bacteriology 2614:10.1126/SCIENCE.1225829 2140:10.1073/PNAS.0408147101 1989:. Honolulu Civil Beat. 1262:She was elected to the 1221:American Cancer Society 1213:Pearl Meister Greengard 1159:In 2015, together with 11210:Emmanuelle Charpentier 10978:Emmanuelle Charpentier 10597:Frank Sherwood Rowland 10459:Lynne Elizabeth Maquat 10445:Emmanuelle Charpentier 10082:Wolf Prize in Medicine 9888:Emmanuelle Charpentier 9547:David Vázquez Martínez 9541:Antonio Garcia-Bellido 9314:Armand Paul Alivisatos 9260:Emmanuelle Charpentier 8981:Clifford P. Brangwynne 8814:Emmanuelle Charpentier 7479:BirdLife International 7460:Environmental Sciences 6518:Nobel Prize recipients 6448:Physiology or Medicine 6361:Emmanuelle Charpentier 6252:Emmanuelle Charpentier 6240:M. Stanley Whittingham 5211:Hans von Euler-Chelpin 4520: 4155:Rockefeller University 2736:Trends in Microbiology 2584:Emmanuelle Charpentier 1253:Emmanuelle Charpentier 1249:Wolf Prize in Medicine 1237:Emmanuelle Charpentier 1217:Rockefeller University 1161:Emmanuelle Charpentier 1138:Alan T. Waterman Award 1132:and received the 1996 956: 922:Emmanuelle Charpentier 754: 741: 646:and was supervised by 640:Harvard Medical School 563:University of Michigan 516:Alan T. Waterman Award 512:awards and fellowships 507: 453:Emmanuelle Charpentier 164:Alan T. Waterman Award 11564:Women Nobel laureates 11534:Pomona College alumni 11145:Gro Harlem Brundtland 9817:Arturo Álvarez-Buylla 8485:High-Z Supernova Team 6814:Charles C. Richardson 5622:Christian B. Anfinsen 5262:Frédéric Joliot-Curie 4494: 3993:on September 13, 2019 3832:MIT Technology Review 3680:MIT Technology Review 3143:. November 18, 2014. 3041:on September 12, 2018 3019:, September 10, 2018. 2931:. February 28, 2020. 2015:. SBGrid Consortium. 1962:. September 8, 2021. 1894:Who's who in the West 1695:breakthroughprize.org 1292:Guggenheim Fellowship 1074:Sixth Street Partners 1066:Johnson & Johnson 982:Intellia Therapeutics 954: 752: 739: 624:Claremont, California 502: 403:Jennifer Anne Doudna 11317:Wm. Theodore de Bary 11246:Charles A. Dinarello 11083:John Michael Wallace 10297:Alexander Varshavsky 10199:Elizabeth F. Neufeld 10159:Jean-Pierre Changeux 10153:Stanley Norman Cohen 9779:George M. Whitesides 9437:Christopher A. Walsh 9304:George M. Whitesides 8924:Virginia Man-Yee Lee 8788:Alexander Varshavsky 8607:Jocelyn Bell Burnell 8505:and contributors to 7078:Christopher J. Chang 7054:Paul J. Hergenrother 7030:Benjamin Cravatt III 6928:Jacqueline K. Barton 6880:Christopher T. Walsh 6409:World Food Programme 6284:Karl Barry Sharpless 5781:Dudley R. Herschbach 5606:Luis Federico Leloir 4751:Simon & Schuster 4658:. America Magazine. 4422:. October 31, 2023. 4416:"Jennifer A. Doudna" 4386:"Jennifer A. Doudna" 3983:"Jennifer A. Doudna" 3533:on December 18, 2015 3507:Gladstone Institutes 3357:. January 30, 2020. 3264:HHMI. Archived from 3141:www.businesswire.com 2788:. January 25, 2019. 2545:Park, Alice (2016). 1282:. She was elected a 1047:Gladstone Institutes 963:affiliated with the 938:Huntington's disease 682:of 141 according to 668:Postdoctoral Scholar 660:research fellowships 626:, where she studied 296:Gladstone Institutes 54:Jennifer Anne Doudna 11258:Tadamitsu Kishimoto 11151:Arthur H. Rosenfeld 11067:/ Kazuo Hagimoto / 10906:Tadamitsu Kishimoto 10636:Jacques-Louis Lions 10428:Jeffrey M. Friedman 10254:Stanley B. Prusiner 10023:Jeffrey M. Friedman 9851:Avelino Corma Canós 9647:Emilio Méndez Pérez 9226:Mildred Dresselhaus 8908:Jeffrey M. Friedman 8798:Charles David Allis 8483:and members of the 8471:and members of the 7036:Dewey G. McCafferty 6964:Stuart L. Schreiber 6898:Harold M. Weintraub 6802:Phillips W. Robbins 6276:Carolyn R. Bertozzi 6188:Jean-Pierre Sauvage 5925:Jens Christian Skou 5893:F. Sherwood Rowland 5805:Charles J. Pedersen 5769:Herbert A. Hauptman 5462:Vincent du Vigneaud 5079:Richard Willstätter 4963:Jacobus van 't Hoff 4340:on January 29, 2020 4306:. October 7, 2019. 4304:"Harvey Prize 2018" 3513:on October 10, 2018 3295:Mammoth Biosciences 3173:. October 7, 2020. 3126:, pp. 208–213. 2873:2015Sci...348...36B 2606:2012Sci...337..816J 2131:2004PNAS..10116987M 2125:(49): 16987–16989. 1873:. December 28, 2023 1349:The Washington Post 1183:, she received the 1163:, she received the 1011:Mammoth Biosciences 1005:Mammoth Biosciences 978:CRISPR Therapeutics 955:CRISPR-Cas9 complex 654:Career and research 599:While she attended 445:CRISPR gene editing 11045:Robert A. Weinberg 11027:Robert G. Gallager 10972:Steven D. Tanksley 10958:Theodore Friedmann 10861:Victor A. McKusick 10819:Masatoshi Takeichi 10767:Timothy R. Parsons 10763:John B. Goodenough 10757:Kimishige Ishizaka 10732:Jack L. Strominger 10728:W. Wesley Peterson 10708:Hiroyuki Yoshikawa 10704:Joseph Engelberger 10680:Edward F. Knipling 10406:Jeffrey V. Ravetch 10327:Alexander Levitzki 10248:Yasutomi Nishizuka 10149:Barbara McClintock 9821:Giacomo Rizzolatti 9751:Juan Ignacio Cirac 9551:Emilio Rosenblueth 9268:Virginijus Šikšnys 9159:Ewine van Dishoeck 9139:Alexei Starobinsky 9097:Donald Lynden-Bell 9001:Masashi Yanagisawa 8912:Franz-Ulrich Hartl 8842:Stephen J. Elledge 8794:Alim Louis Benabid 8681:Charles H. Bennett 8631:Daniel Z. Freedman 8581:Charles L. Bennett 8545:Arthur B. McDonald 8533:Kōichirō Nishikawa 8463:John Henry Schwarz 8453:Alexander Polyakov 8431:Michel Della Negra 8350:Daniel A. Spielman 8265:Breakthrough Prize 8101:Simon Peyton Jones 7060:Scott K. Silverman 7042:Linda Hsieh-Wilson 7024:Andreas Matouschek 6982:Gregory L. Verdine 6946:George L. McLendon 6748:Harold A. Scheraga 6658:Earl A. Evans, Jr. 6236:John B. Goodenough 6164:William E. Moerner 6044:Richard R. Schrock 6016:Roderick MacKinnon 5988:K. Barry Sharpless 5813:Johann Deisenhofer 5642:Geoffrey Wilkinson 5638:Ernst Otto Fischer 5562:Robert S. Mulliken 5498:Jaroslav Heyrovský 5446:Hermann Staudinger 5266:Irène Joliot-Curie 4888:on Nobelprize.org 4886:Jennifer A. Doudna 4017:Breakthrough Prize 3477:Multiple sources: 3422:www.bionews.org.uk 3327:. August 1, 2019. 3099:Bloomberg business 2817:The New York Times 1620:The New York Times 1419:The New York Times 1325:Person of the Year 1229:Virginijus Šikšnys 1181:Rodolphe Barrangou 957: 930:sickle cell anemia 787:Harvard University 755: 742: 508: 486:history of biology 388:Doudna Lab website 213:in Medicine (2020) 96:Harvard University 11421: 11420: 11415: 11414: 11228:Anthony R. Hunter 11096: 11095: 11065:Masataka Nakazawa 11059:Christopher Field 10944:Yasuharu Suematsu 10888:Shun-ichi Iwasaki 10847:Peter Shaw Ashton 10787:Benoit Mandelbrot 10781:Andrzej Tarkowski 10601:Elias James Corey 10488: 10487: 10469:Daniel J. Drucker 10331:Anthony R. Hunter 10303:Ralph L. Brinster 10185:Pedro Cuatrecasas 10173:Donald F. Steiner 10163:Solomon H. Snyder 10118:Oleh Hornykiewicz 10080:Laureates of the 10047: 10046: 10043: 10042: 10019:Daniel J. Drucker 10005:Jeffrey I. Gordon 9943:Ingrid Daubechies 9933:Sandra Myrna Díaz 9867: 9866: 9831:Richard A. Lerner 9793:Raymond Tomlinson 9589:Santiago Grisolía 9497:Laureates of the 9464: 9463: 9429:Jean-Louis Mandel 9423:Ardem Patapoutian 9409:Robert Fettiplace 9405:A. James Hudspeth 9395:Michael Merzenich 9363:Cornelia Bargmann 9278:Maximilian Haider 9189:David Charbonneau 9107:Raymond N. Wilson 9044: 9043: 9023:Fredrick Van Goor 8916:Arthur L. Horwich 8890:Adrian R. Krainer 8784:Richard P. Lifton 8742:Napoleone Ferrara 8722:Cornelia Bargmann 8571:Andrew Strominger 8567:Joseph Polchinski 8377:Nima Arkani-Hamed 8326:Vincent Lafforgue 8316:Christopher Hacon 8231: 8230: 8056:Eugenia Kumacheva 7951:Alastair Compston 7936:Katharine Cashman 7845: 7844: 7674:Cognitive Science 7141: 7140: 7114:Bradley Pentelute 7084:Matthew D. Disney 6892:Roger D. Kornberg 6874:Charles R. Cantor 6850:James E. Dahlberg 6820:Mario R. Capecchi 6796:Gerald M. Edelman 6784:William P. Jencks 6742:Robert A. Alberty 6736:William F. Neuman 6664:Herbert E. Carter 6577: 6576: 6484:Economic Sciences 6309: 6308: 6292:Moungi G. Bawendi 6212:Richard Henderson 6056:Roger D. Kornberg 6024:Aaron Ciechanover 5857:Rudolph A. Marcus 5761:Robert Merrifield 5697:Peter D. Mitchell 5470:Cyril Hinshelwood 5231:Friedrich Bergius 5155:Richard Zsigmondy 5071:Theodore Richards 5019:Ernest Rutherford 4760:978-1-9821-1585-2 4625:"Past Recipients" 4390:www.nasonline.org 4360:"Wolf Prize 2020" 4043:"Jennifer Doudna" 4013:"Jennifer Doudna" 3942:978-0-309-10097-7 3887:, pp. 63–66. 3875:, pp. 54–55. 3424:. April 3, 2020. 3235:. June 17, 2017. 3171:finance.yahoo.com 2965:Los Angeles Times 2600:(6096): 816–821. 2586:(June 28, 2012). 2580:Jennifer A Doudna 2476:"Jennifer Doudna" 2308:. March 8, 2020. 2041:. The Kavli Prize 1779:"Jennifer Doudna" 1245:LUI Che Woo Prize 1118:Awards and honors 1112:public university 1087:Boulder, Colorado 1023:COVID-19 pandemic 1017:COVID-19 response 920:, but Doudna and 833: 832: 514:include the 2000 400: 399: 350:Doctoral students 236:Scientific career 64:February 19, 1964 11576: 11394:The Legal Agenda 11254:(Australia, USA) 11198:James P. Allison 11123: 11116: 11109: 11100: 11099: 10934:C. Grant Willson 10722:Marc Van Montagu 10696:Takashi Sugimura 10630:Xavier Le Pichon 10575:Donald Henderson 10571:Georges Vendryes 10515: 10508: 10501: 10492: 10491: 10479:José-Alain Sahel 10422:James P. Allison 10412:Lewis C. Cantley 10402:Philippa Marrack 10244:Michael Berridge 10074: 10067: 10060: 10051: 10050: 9873: 9872: 9841:François Englert 9699:Lawrence Roberts 9693:Jean Weissenbach 9617:Manuel Patarroyo 9593:Salvador Moncada 9577:Marcos Moshinsky 9529:Manuel Ballester 9508: 9507: 9491: 9484: 9477: 9468: 9467: 9451:Winrich Freiwald 9353:Thomas C. Südhof 9349:Richard Scheller 9310:Robert S. Langer 9250:Christoph Gerber 9125:Michael E. Brown 9071: 9064: 9057: 9048: 9047: 9035:Andrew Singleton 8985:Anthony A. Hyman 8955:Jeffery W. Kelly 8950: 8942:Richard J. Youle 8886:C. Frank Bennett 8880:Don W. Cleveland 8854:Yoshinori Ohsumi 8780:Robert S. Langer 8768:James P. Allison 8730:Lewis C. Cantley 8671:Hidetoshi Katori 8651:Jens H. Gundlach 8561:Super-Kamiokande 8443:Joseph Incandela 8427:Fabiola Gianotti 8389:Maxim Kontsevich 8344:Takuro Mochizuki 8286:Maxim Kontsevich 8258: 8251: 8244: 8235: 8234: 8116:Sriram Ramaswamy 8061:Corinne Le Quéré 7991:Maria Fitzgerald 7976:Marcus du Sautoy 7971:Christl Donnelly 7941:Sarah Cleaveland 7872: 7865: 7858: 7849: 7848: 7539:Sinninghe Damsté 7168: 7161: 7154: 7145: 7144: 6988:Alanna Schepartz 6952:Peter G. Schultz 6922:James E. Rothman 6910:David V. Goeddel 6886:Phillip A. Sharp 6868:Robert G. Roeder 6838:Bruce M. Alberts 6808:Gordon G. Hammes 6724:Nathan O. Kaplan 6712:John M. Buchanan 6694:Dilworth Woodley 6670:Joseph S. Fruton 6652:David Rittenberg 6628:Harold S. Alcott 6622:William M. Allen 6604: 6597: 6590: 6581: 6580: 6520: 6503:Robert B. Wilson 6469:(United Kingdom) 6467:Michael Houghton 6427:(United Kingdom) 6336: 6329: 6322: 6313: 6312: 6302: 6300:Alexei I. Ekimov 6286: 6280:Morten P. Meldal 6270: 6258: 6246: 6230: 6214: 6204:Jacques Dubochet 6198: 6182: 6166: 6150: 6134: 6128:Robert Lefkowitz 6122: 6114: 6098: 6092:Thomas A. Steitz 6082: 6066: 6058: 6050: 6040:Robert H. Grubbs 6034: 6018: 6006: 5990: 5963: 5961:Hideki Shirakawa 5947: 5939: 5927: 5911: 5895: 5879: 5871: 5859: 5851: 5849:Richard R. Ernst 5843: 5835: 5823: 5807: 5791: 5775: 5763: 5755: 5747: 5739: 5727: 5725:Frederick Sanger 5711: 5705:Herbert C. Brown 5699: 5691: 5683: 5681:William Lipscomb 5664: 5652: 5644: 5632: 5616: 5614:Gerhard Herzberg 5608: 5600: 5588: 5580: 5564: 5556: 5548: 5540: 5528: 5516: 5508: 5500: 5492: 5490:Frederick Sanger 5484: 5476: 5474:Nikolay Semyonov 5464: 5456: 5448: 5440: 5428: 5426:Glenn T. Seaborg 5405: 5393: 5385: 5377: 5369: 5353: 5351:Artturi Virtanen 5345: 5337: 5335:George de Hevesy 5329: 5328: 5322: 5321: 5315: 5314: 5308: 5296: 5288: 5276: 5268: 5256: 5248: 5247: 5241: 5233: 5221: 5213: 5201: 5193: 5191:Heinrich Wieland 5185: 5183:Theodor Svedberg 5168: 5167: 5157: 5149: 5148: 5142: 5134: 5126: 5118: 5110: 5109: 5103: 5095: 5094: 5088: 5087: 5081: 5073: 5065: 5057: 5045: 5037: 5029: 5021: 5013: 5005: 4997: 4995:Adolf von Baeyer 4989: 4981: 4979:Svante Arrhenius 4973: 4965: 4936: 4929: 4922: 4913: 4912: 4898: 4897: 4893: 4877: 4871:, March 8, 2021. 4857: 4845: 4833: 4821: 4786: 4785: 4771: 4769: 4767: 4737:Isaacson, Walter 4723: 4722: 4720: 4718: 4711:news.harvard.edu 4703: 4697: 4696: 4694: 4692: 4678: 4672: 4671: 4669: 4667: 4651: 4645: 4644: 4642: 4640: 4621: 4615: 4614: 4612: 4610: 4595: 4589: 4588: 4586: 4584: 4561: 4555: 4554: 4552: 4550: 4527: 4521: 4518: 4516: 4514: 4492: 4490: 4488: 4468: 4462: 4461: 4459: 4457: 4442: 4436: 4435: 4433: 4431: 4412: 4406: 4405: 4403: 4401: 4382: 4376: 4375: 4373: 4371: 4356: 4350: 4349: 4347: 4345: 4336:. Archived from 4326: 4320: 4319: 4317: 4315: 4300: 4294: 4293: 4261: 4255: 4254: 4208: 4202: 4201: 4199: 4197: 4177: 4171: 4170: 4168: 4166: 4147: 4141: 4140: 4138: 4136: 4117: 4111: 4110: 4108: 4106: 4091: 4085: 4084: 4082: 4080: 4065: 4059: 4058: 4056: 4054: 4039: 4033: 4032: 4030: 4028: 4009: 4003: 4002: 4000: 3998: 3989:. Archived from 3979: 3973: 3972: 3970: 3968: 3953: 3947: 3946: 3919: 3913: 3912: 3904:The Mercury News 3894: 3888: 3882: 3876: 3870: 3864: 3863: 3847: 3841: 3840: 3822: 3816: 3815: 3797: 3791: 3790: 3780: 3772: 3766: 3765: 3753: 3745: 3739: 3738: 3726: 3718: 3712: 3711: 3695: 3689: 3688: 3670: 3664: 3663: 3651: 3643: 3637: 3636: 3620: 3614: 3613: 3603: 3595: 3589: 3588: 3578: 3570: 3564: 3563: 3551: 3545: 3542: 3540: 3538: 3522: 3520: 3518: 3509:. 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Habener 11240:John Mendelsohn 11216:Jennifer Doudna 11186: 11133: 11127: 11097: 11092: 11087:Ronald M. Evans 11073:Karl Deisseroth 11069:Gero Miesenböck 11041:Bert Vogelstein 11037:Martin A. Green 11015: 10996:Max Dale Cooper 10982:Jennifer Doudna 10954:Yutaka Takahasi 10876: 10823:Erkki Ruoslahti 10805:Akira Fujishima 10773:Tim Berners-Lee 10741: 10736:Don Craig Wiley 10622:W. Jason Morgan 10606: 10565:Theodore Maiman 10551:Willem J. Kolff 10525: 10519: 10489: 10484: 10449:Jennifer Doudna 10433: 10384:Nahum Sonenberg 10378:Ronald M. Evans 10372:Rudolf Jaenisch 10368:Shinya Yamanaka 10350: 10317:Robert Weinberg 10311:Oliver Smithies 10281: 10214: 10209:Edward B. Lewis 10143:James L. Gowans 10123: 10104:Jon J. van Rood 10084: 10078: 10048: 10039: 10035:Svetlana Mojsov 10031:Jens Juul Holst 10027:Joel F. Habener 9987:Geoffrey Hinton 9951:Emmanuel Candès 9912:Barry C. Barish 9892:Jennifer Doudna 9878: 9863: 9859:Galen D. Stucky 9745:Antonio Damasio 9739:Robert Weinberg 9735:Bert Vogelstein 9711:Tim Berners-Lee 9685:Francis Collins 9634:Valentín Fuster 9513: 9502: 9495: 9465: 9460: 9455:Doris Ying Tsao 9447:Nancy Kanwisher 9413:Christine Petit 9323: 9300:David L. Allara 9286:Ondrej Krivanek 9264:Jennifer Doudna 9194: 9117:David C. Jewitt 9103:Jerry E. Nelson 9093:Maarten Schmidt 9081: 9075: 9045: 9040: 9031:Ellen Sidransky 9011:Michel Sadelain 8997:Emmanuel Mignot 8971:David Klenerman 8944: 8934:Catherine Dulac 8846:Harry F. Noller 8824:Karl Deisseroth 8810:Jennifer Doudna 8776:Michael N. Hall 8762:Bert Vogelstein 8758:Shinya Yamanaka 8754:Robert Weinberg 8750:Charles Sawyers 8708: 8685:Gilles Brassard 8665:Steven Weinberg 8647:Eric Adelberger 8557:Yōichirō Suzuki 8469:Saul Perlmutter 8435:Tejinder Virdee 8419:Stephen Hawking 8368: 8361: 8282:Simon Donaldson 8268: 8262: 8232: 8227: 8193:Jennifer Doudna 8166: 8150: 8121:Caroline Series 8076:Eleanor Maguire 8041:Ramanujan Hegde 7889: 7876: 7846: 7841: 7728: 7669: 7562: 7455: 7342: 7177: 7175:Heineken Prizes 7172: 7142: 7137: 7132:Jordan L. Meier 7120:Neal K. Devaraj 7102:Elizabeth Nolan 7018:Kevan M. Shokat 7012:Jennifer Doudna 6958:William DeGrado 6916:Gerald M. Rubin 6832:David F. Wilson 6772:Frederick Crane 6718:David M. Bonner 6700:Irving M. Klotz 6688:Sidney Colowick 6613: 6608: 6578: 6573: 6516: 6508: 6505:(United States) 6499:(United States) 6478: 6475:(United States) 6473:Charles M. Rice 6463:(United States) 6461:Harvey J. Alter 6442: 6439:(United States) 6431:Reinhard Genzel 6411: 6392: 6391:(United States) 6372: 6369:(United States) 6367:Jennifer Doudna 6347: 6340: 6310: 6305: 6289: 6273: 6264:David MacMillan 6261: 6256:Jennifer Doudna 6249: 6233: 6217: 6201: 6192:Fraser Stoddart 6185: 6176:Paul L. Modrich 6169: 6153: 6137: 6125: 6117: 6112:Ei-ichi Negishi 6104:Richard F. Heck 6101: 6085: 6072:Osamu Shimomura 6069: 6061: 6053: 6037: 6021: 6009: 5993: 5980:William Knowles 5977: 5966: 5957:Alan MacDiarmid 5950: 5942: 5930: 5914: 5909:Richard Smalley 5898: 5885:Paul J. Crutzen 5882: 5874: 5862: 5854: 5846: 5838: 5826: 5810: 5801:Jean-Marie Lehn 5794: 5778: 5766: 5758: 5750: 5742: 5730: 5714: 5702: 5694: 5686: 5678: 5667: 5662:Vladimir Prelog 5655: 5647: 5635: 5619: 5611: 5603: 5591: 5583: 5567: 5559: 5554:Robert Woodward 5551: 5546:Dorothy Hodgkin 5543: 5531: 5519: 5511: 5503: 5495: 5487: 5479: 5467: 5459: 5451: 5443: 5431: 5419: 5408: 5396: 5391:William Giauque 5388: 5380: 5375:Robert Robinson 5372: 5359:James B. Sumner 5356: 5348: 5340: 5332: 5326: 5325: 5319: 5318: 5312: 5311: 5306:Leopold Ružička 5302:Adolf Butenandt 5299: 5291: 5279: 5271: 5259: 5251: 5245: 5244: 5239:Irving Langmuir 5236: 5224: 5216: 5204: 5196: 5188: 5180: 5169: 5165: 5160: 5152: 5146: 5145: 5137: 5129: 5124:Frederick Soddy 5121: 5113: 5107: 5106: 5098: 5092: 5091: 5085: 5084: 5076: 5068: 5060: 5051:Victor Grignard 5048: 5040: 5032: 5027:Wilhelm Ostwald 5024: 5016: 5008: 5000: 4992: 4984: 4976: 4968: 4960: 4949: 4940: 4905: 4875: 4848: 4836: 4824: 4811:Jennifer Doudna 4807: 4806: 4805: 4800:Jennifer Doudna 4787: 4783: 4778: 4765: 4763: 4761: 4732: 4727: 4726: 4716: 4714: 4705: 4704: 4700: 4690: 4688: 4680: 4679: 4675: 4665: 4663: 4652: 4648: 4638: 4636: 4623: 4622: 4618: 4608: 4606: 4597: 4596: 4592: 4582: 4580: 4569:achievement.org 4563: 4562: 4558: 4548: 4546: 4535:achievement.org 4529: 4528: 4524: 4512: 4510: 4501: 4486: 4484: 4469: 4465: 4455: 4453: 4444: 4443: 4439: 4429: 4427: 4414: 4413: 4409: 4399: 4397: 4384: 4383: 4379: 4369: 4367: 4358: 4357: 4353: 4343: 4341: 4328: 4327: 4323: 4313: 4311: 4302: 4301: 4297: 4262: 4258: 4221:(7708): 17–18. 4209: 4205: 4195: 4193: 4178: 4174: 4164: 4162: 4149: 4148: 4144: 4134: 4132: 4119: 4118: 4114: 4104: 4102: 4093: 4092: 4088: 4078: 4076: 4067: 4066: 4062: 4052: 4050: 4049:on May 13, 2016 4041: 4040: 4036: 4026: 4024: 4011: 4010: 4006: 3996: 3994: 3981: 3980: 3976: 3966: 3964: 3955: 3954: 3950: 3943: 3921: 3920: 3916: 3895: 3891: 3883: 3879: 3871: 3867: 3848: 3844: 3823: 3819: 3798: 3794: 3773: 3769: 3746: 3742: 3730:Financial Times 3719: 3715: 3696: 3692: 3671: 3667: 3644: 3640: 3621: 3617: 3596: 3592: 3571: 3567: 3561: 3556:indexed by the 3552: 3548: 3536: 3534: 3525: 3516: 3514: 3491: 3489: 3480: 3476: 3472: 3462: 3460: 3445: 3441: 3431: 3429: 3416: 3415: 3411: 3401: 3399: 3391: 3390: 3386: 3378: 3374: 3364: 3362: 3349: 3348: 3344: 3334: 3332: 3319: 3318: 3314: 3304: 3302: 3289: 3288: 3281: 3271: 3269: 3268:on May 16, 2013 3260: 3259: 3252: 3242: 3240: 3232:TheGuardian.com 3225: 3224: 3220: 3210: 3208: 3195: 3194: 3190: 3180: 3178: 3165: 3164: 3160: 3150: 3148: 3135: 3134: 3130: 3122: 3118: 3108: 3106: 3093: 3092: 3088: 3078: 3076: 3061: 3054: 3044: 3042: 3027: 3023: 3014: 3010: 3000: 2998: 2983: 2979: 2969: 2967: 2957: 2948: 2938: 2936: 2927: 2926: 2922: 2851:Michael Botchan 2843:David Baltimore 2840: 2836: 2826: 2824: 2809: 2805: 2795: 2793: 2784: 2783: 2779: 2742:(10): 811–820. 2729: 2725: 2678:(12): 5429–33. 2659: 2655: 2576: 2572: 2562: 2560: 2543: 2532: 2526:Wayback Machine 2517: 2513: 2507:Wayback Machine 2498: 2494: 2484: 2482: 2474: 2473: 2466: 2456: 2454: 2446: 2445: 2441: 2431: 2429: 2421: 2420: 2416: 2406: 2404: 2396: 2395: 2391: 2381: 2379: 2370: 2369: 2365: 2355: 2353: 2345: 2344: 2337: 2329: 2325: 2315: 2313: 2304: 2303: 2296: 2286: 2284: 2271: 2270: 2266: 2252:Jennifer Doudna 2250: 2246: 2219: 2215: 2205: 2203: 2186: 2182: 2109: 2076: 2066: 2064: 2059: 2058: 2054: 2044: 2042: 2037: 2036: 2032: 2022: 2020: 2011: 2010: 2006: 1996: 1994: 1983: 1979: 1969: 1967: 1954: 1953: 1946: 1936: 1934: 1923: 1912: 1905: 1891: 1890: 1886: 1876: 1874: 1865: 1864: 1860: 1850: 1848: 1831: 1824: 1814: 1812: 1803: 1802: 1798: 1788: 1786: 1777: 1776: 1769: 1759: 1757: 1748: 1747: 1740: 1730: 1728: 1719: 1718: 1714: 1704: 1702: 1689: 1688: 1669: 1659: 1657: 1648: 1647: 1640: 1630: 1628: 1611: 1607: 1597: 1595: 1584: 1561: 1552: 1521: 1511: 1509: 1505: 1498: 1494: 1493: 1472: 1462: 1460: 1447: 1446: 1439: 1429: 1427: 1410: 1403: 1393: 1391: 1376: 1375: 1368: 1358: 1356: 1351:. May 3, 2016. 1343: 1342: 1338: 1333: 1215:Prize from the 1203:(2017) and the 1126: 1120: 1082: 1062:Isomorphic Labs 1039: 1019: 1007: 974:Editas Medicine 961:Broad Institute 934:cystic fibrosis 892: 815: 800:External videos 795: 747: 731:Yale University 696: 656: 648:Jack W. Szostak 582:popular science 578:flora and fauna 559: 463:Chair Professor 415: 411: 405: 392: 391: 386: 374: 371:Basem Al-Shayeb 357:Rachel Haurwitz 321: 305: 286:Yale University 269: 230: 209: 152: 149: 2000) 144: 140: 129: 105: 74: 68: 65: 59: 57: 56: 55: 45: 33: 28: 26: 25:Jennifer Doudna 17: 12: 11: 5: 11582: 11572: 11571: 11566: 11561: 11556: 11551: 11546: 11541: 11536: 11531: 11526: 11521: 11516: 11511: 11506: 11501: 11496: 11491: 11486: 11484:Genome editing 11481: 11476: 11471: 11466: 11461: 11456: 11451: 11446: 11441: 11436: 11419: 11418: 11413: 11412: 11410: 11409: 11403: 11397: 11391: 11385: 11379: 11373: 11367: 11366:(South Africa) 11360: 11358: 11354: 11353: 11351: 11350: 11344: 11341:Jessica Rawson 11338: 11332: 11326: 11320: 11314: 11307: 11305: 11301: 11300: 11298: 11297: 11291: 11285: 11279: 11273: 11272:(Hungary, USA) 11270:Katalin Karikó 11267: 11261: 11255: 11249: 11243: 11237: 11231: 11225: 11219: 11213: 11207: 11201: 11194: 11192: 11188: 11187: 11185: 11184: 11178: 11172: 11166: 11160: 11154: 11148: 11141: 11139: 11135: 11134: 11126: 11125: 11118: 11111: 11103: 11094: 11093: 11091: 11090: 11076: 11062: 11051:Katalin Karikó 11048: 11034: 11023: 11021: 11017: 11016: 11014: 11013: 11006:Yoshio Okamoto 11003: 11000:Jacques Miller 10989: 10975: 10965: 10951: 10948:C. David Allis 10941: 10931: 10924:Nicholas Lydon 10913: 10902:Dennis Ritchie 10895: 10892:Peter Vitousek 10884: 10882: 10878: 10877: 10875: 10874: 10867:Dennis Meadows 10864: 10857:Robert E. Kahn 10850: 10843:Peter Grünberg 10836: 10826: 10812: 10809:John H. Lawton 10798: 10791:James A. Yorke 10784: 10770: 10760: 10749: 10747: 10743: 10742: 10740: 10739: 10725: 10711: 10693: 10686:Charles K. Kao 10683: 10673: 10670:Arvid Carlsson 10663: 10660:Kary B. Mullis 10653: 10643: 10633: 10614: 10612: 10608: 10607: 10605: 10604: 10594: 10587:Luc Montagnier 10568: 10557:Henry Beachell 10554: 10547:David Turnbull 10544: 10537:John R. Pierce 10533: 10531: 10527: 10526: 10518: 10517: 10510: 10503: 10495: 10486: 10485: 10483: 10482: 10472: 10466: 10463:Adrian Krainer 10452: 10441: 10439: 10435: 10434: 10432: 10431: 10425: 10419: 10416:C. Ronald Kahn 10409: 10395: 10381: 10375: 10365: 10358: 10356: 10352: 10351: 10349: 10348: 10338: 10324: 10321:Roger Y. Tsien 10314: 10307:Mario Capecchi 10300: 10289: 10287: 10283: 10282: 10280: 10279: 10273: 10263: 10257: 10251: 10241: 10235: 10232:Seymour Benzer 10229: 10226:Maclyn McCarty 10222: 10220: 10216: 10215: 10213: 10212: 10202: 10192: 10182: 10179:Osamu Hayaishi 10176: 10170: 10167:James W. Black 10156: 10146: 10135:César Milstein 10131: 10129: 10125: 10124: 10122: 10121: 10114:Arvid Carlsson 10107: 10092: 10090: 10086: 10085: 10077: 10076: 10069: 10062: 10054: 10045: 10044: 10041: 10040: 10038: 10037: 10015: 10013:Bonnie Bassler 10001: 9999:Demis Hassabis 9983: 9965:Philip Felgner 9957:Katalin Karikó 9953: 9935: 9925: 9919: 9900: 9894: 9883: 9880: 9879: 9869: 9868: 9865: 9864: 9862: 9861: 9847: 9833: 9827:Gregory Winter 9823: 9809: 9795: 9785: 9771:Shuji Nakamura 9763: 9757:Peter Lawrence 9753: 9747: 9741: 9719: 9713: 9703:Robert E. Kahn 9695: 9689:Hamilton Smith 9673: 9671:Luc Montagnier 9663: 9657:Ricardo Miledi 9653: 9643: 9636: 9630: 9619: 9613: 9607: 9601: 9595: 9585: 9579: 9573:Manuel Cardona 9569: 9563:Jacinto Convit 9559: 9553: 9543: 9537: 9531: 9525: 9518: 9515: 9514: 9504: 9503: 9494: 9493: 9486: 9479: 9471: 9462: 9461: 9459: 9458: 9444: 9426: 9416: 9402: 9399:Carla J. Shatz 9388: 9385:Marcus Raichle 9374: 9360: 9346: 9339:Thomas Jessell 9331: 9329: 9325: 9324: 9322: 9321: 9318:Chad A. Mirkin 9307: 9296:Ralph G. Nuzzo 9289: 9271: 9257: 9243: 9232:Thomas Ebbesen 9229: 9223: 9220:Nadrian Seeman 9213: 9202: 9200: 9196: 9195: 9193: 9192: 9182: 9168: 9162: 9156: 9142: 9128: 9114: 9100: 9089: 9087: 9083: 9082: 9074: 9073: 9066: 9059: 9051: 9042: 9041: 9039: 9038: 9019:Paul Negulescu 9004: 8989:Demis Hassabis 8978: 8959:Katalin Karikó 8952: 8927: 8905: 8898:Xiaowei Zhuang 8883: 8872:Kazutoshi Mori 8861: 8839: 8817: 8791: 8765: 8738:Titia de Lange 8726:David Botstein 8718: 8716: 8710: 8709: 8707: 8706: 8696: 8678: 8668: 8658: 8644: 8638: 8627:Sergio Ferrara 8620: 8610: 8600: 8593:Lyman Page Jr. 8589:Norman Jarosik 8578: 8564: 8553:Takaaki Kajita 8510: 8509:project (2016) 8488: 8466: 8456: 8450: 8412: 8401:Nathan Seiberg 8397:Juan Maldacena 8373: 8371: 8363: 8362: 8360: 8359: 8353: 8347: 8341: 8335: 8329: 8323: 8320:James McKernan 8313: 8307: 8301: 8298:Richard Taylor 8278: 8276: 8270: 8269: 8261: 8260: 8253: 8246: 8238: 8229: 8228: 8226: 8225: 8223:Ellen Williams 8220: 8215: 8210: 8205: 8200: 8195: 8190: 8185: 8180: 8174: 8172: 8168: 8167: 8165: 8164: 8158: 8156: 8152: 8151: 8149: 8148: 8143: 8141:Philip Withers 8138: 8133: 8128: 8123: 8118: 8113: 8108: 8106:Jonathon Pines 8103: 8098: 8093: 8091:Russell Morris 8088: 8083: 8078: 8073: 8068: 8063: 8058: 8053: 8048: 8043: 8038: 8033: 8031:Richard Harvey 8028: 8023: 8018: 8013: 8008: 8003: 7998: 7996:Antony Galione 7993: 7988: 7983: 7978: 7973: 7968: 7963: 7958: 7953: 7948: 7943: 7938: 7933: 7928: 7926:Martin Bridson 7923: 7918: 7913: 7908: 7906:Jas Pal Badyal 7903: 7897: 7895: 7891: 7890: 7875: 7874: 7867: 7860: 7852: 7843: 7842: 7840: 7839: 7833: 7827: 7821: 7818:Van Oldenborgh 7815: 7809: 7803: 7797: 7791: 7785: 7779: 7773: 7767: 7761: 7755: 7749: 7743: 7736: 7734: 7730: 7729: 7727: 7726: 7720: 7714: 7708: 7702: 7696: 7690: 7684: 7677: 7675: 7671: 7670: 7668: 7667: 7661: 7655: 7649: 7643: 7637: 7631: 7625: 7619: 7613: 7607: 7601: 7595: 7589: 7583: 7577: 7570: 7568: 7564: 7563: 7561: 7560: 7554: 7548: 7542: 7536: 7530: 7524: 7518: 7512: 7506: 7500: 7494: 7488: 7482: 7476: 7470: 7463: 7461: 7457: 7456: 7454: 7453: 7447: 7441: 7435: 7429: 7423: 7417: 7411: 7405: 7399: 7393: 7387: 7381: 7375: 7369: 7363: 7357: 7350: 7348: 7344: 7343: 7341: 7340: 7334: 7328: 7322: 7316: 7310: 7304: 7298: 7292: 7286: 7280: 7274: 7268: 7262: 7256: 7250: 7244: 7238: 7228: 7222: 7216: 7210: 7204: 7198: 7192: 7185: 7183: 7179: 7178: 7171: 7170: 7163: 7156: 7148: 7139: 7138: 7136: 7135: 7129: 7123: 7117: 7111: 7105: 7099: 7093: 7087: 7081: 7075: 7072:Nathanael Gray 7069: 7063: 7057: 7051: 7045: 7039: 7033: 7027: 7021: 7015: 7009: 7003: 7000:Chaitan Khosla 6997: 6991: 6985: 6979: 6976:Jeremy M. Berg 6973: 6967: 6961: 6955: 6949: 6943: 6940:Michael M. Cox 6937: 6931: 6925: 6919: 6913: 6907: 6901: 6895: 6889: 6883: 6877: 6871: 6865: 6862:Joan A. Steitz 6859: 6853: 6847: 6841: 6835: 6829: 6823: 6817: 6811: 6805: 6799: 6793: 6787: 6781: 6778:Jerard Hurwitz 6775: 6769: 6763: 6757: 6754:Lester J. Reed 6751: 6745: 6739: 6733: 6727: 6721: 6715: 6709: 6703: 6697: 6691: 6685: 6679: 6676:Max A. Lauffer 6673: 6667: 6661: 6655: 6649: 6643: 6637: 6631: 6625: 6618: 6615: 6614: 6607: 6606: 6599: 6592: 6584: 6575: 6574: 6572: 6571: 6566: 6561: 6556: 6551: 6546: 6541: 6536: 6531: 6526: 6521: 6513: 6510: 6509: 6507: 6506: 6500: 6493: 6491: 6480: 6479: 6477: 6476: 6470: 6464: 6457: 6455: 6444: 6443: 6441: 6440: 6437:Andrea M. Ghez 6434: 6428: 6421: 6419: 6413: 6412: 6407: 6405: 6394: 6393: 6387: 6385: 6374: 6373: 6371: 6370: 6364: 6357: 6355: 6349: 6348: 6339: 6338: 6331: 6324: 6316: 6307: 6306: 6304: 6303: 6287: 6271: 6259: 6247: 6231: 6224:Gregory Winter 6220:Frances Arnold 6215: 6199: 6183: 6167: 6151: 6144:Michael Levitt 6140:Martin Karplus 6135: 6123: 6115: 6099: 6083: 6080:Roger Y. Tsien 6076:Martin Chalfie 6067: 6059: 6051: 6035: 6019: 6007: 5991: 5974: 5972: 5968: 5967: 5965: 5964: 5953:Alan J. Heeger 5948: 5940: 5928: 5921:John E. Walker 5912: 5896: 5880: 5872: 5860: 5852: 5844: 5836: 5824: 5821:Hartmut Michel 5808: 5797:Donald J. Cram 5792: 5776: 5764: 5756: 5748: 5740: 5737:Roald Hoffmann 5728: 5721:Walter Gilbert 5712: 5700: 5692: 5689:Ilya Prigogine 5684: 5675: 5673: 5669: 5668: 5666: 5665: 5658:John Cornforth 5653: 5645: 5633: 5626:Stanford Moore 5617: 5609: 5601: 5589: 5581: 5574:Ronald Norrish 5565: 5557: 5549: 5541: 5529: 5517: 5509: 5501: 5493: 5485: 5482:Alexander Todd 5477: 5465: 5457: 5449: 5441: 5429: 5422:Edwin McMillan 5416: 5414: 5410: 5409: 5407: 5406: 5394: 5386: 5378: 5370: 5354: 5346: 5338: 5330: 5323: 5316: 5309: 5297: 5289: 5282:Norman Haworth 5277: 5269: 5257: 5249: 5242: 5234: 5222: 5214: 5202: 5194: 5186: 5177: 5175: 5171: 5170: 5163: 5161: 5159: 5158: 5150: 5143: 5135: 5127: 5119: 5116:Walther Nernst 5111: 5104: 5096: 5089: 5082: 5074: 5066: 5058: 5046: 5038: 5030: 5022: 5014: 5011:Eduard Buchner 5006: 4998: 4990: 4987:William Ramsay 4982: 4974: 4966: 4957: 4955: 4951: 4950: 4939: 4938: 4931: 4924: 4916: 4910: 4907: 4906: 4895: 4894: 4883: 4872: 4858: 4846: 4834: 4822: 4815:Google Scholar 4788: 4781: 4780: 4779: 4777: 4776:External links 4774: 4773: 4772: 4759: 4731: 4728: 4725: 4724: 4713:. 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July 1, 2022 2363: 2335: 2333:, p. 101. 2323: 2294: 2277:www.scopus.com 2264: 2256:Google Scholar 2244: 2213: 2180: 2074: 2052: 2030: 2004: 1977: 1944: 1910: 1903: 1884: 1871:www.invent.org 1858: 1822: 1796: 1767: 1738: 1712: 1667: 1638: 1605: 1559: 1519: 1470: 1453:nobelprize.org 1437: 1401: 1366: 1335: 1334: 1332: 1329: 1251:(jointly with 1189:Heineken Prize 1130:Searle Scholar 1122:Main article: 1119: 1116: 1081: 1078: 1038: 1035: 1018: 1015: 1006: 1003: 891: 888: 831: 830: 811: 810: 802: 801: 794: 791: 746: 743: 695: 692: 684:Google Scholar 655: 652: 620:Pomona College 572:Growing up in 558: 555: 503:Doudna at the 494:genome editing 398: 397: 384: 380: 379: 376: 375: 373: 372: 369: 367:Lei Stanley Qi 364: 359: 353: 351: 347: 346: 341: 337: 336: 331: 325: 324: 313: 307: 306: 304: 303: 298: 293: 288: 283: 277: 275: 271: 270: 268: 267: 262: 256: 251: 245: 243: 239: 238: 232: 231: 229: 228: 221: 215: 207: 197: 191: 185: 179: 173: 167: 160: 158: 154: 153: 142: 138: 137: 135: 131: 130: 128: 127: 122: 117: 113: 111: 110:Known for 107: 106: 104: 103: 93: 86:Pomona College 82: 80: 76: 75: 69: 53: 51: 47: 46: 44:Doudna in 2023 43: 35: 34: 27: 24: 15: 9: 6: 4: 3: 2: 11581: 11570: 11567: 11565: 11562: 11560: 11557: 11555: 11552: 11550: 11547: 11545: 11542: 11540: 11537: 11535: 11532: 11530: 11527: 11525: 11522: 11520: 11517: 11515: 11512: 11510: 11509:Living people 11507: 11505: 11502: 11500: 11497: 11495: 11492: 11490: 11487: 11485: 11482: 11480: 11477: 11475: 11472: 11470: 11467: 11465: 11462: 11460: 11457: 11455: 11452: 11450: 11447: 11445: 11442: 11440: 11437: 11435: 11432: 11431: 11429: 11407: 11406:Mary Robinson 11404: 11401: 11398: 11395: 11392: 11389: 11386: 11383: 11380: 11377: 11374: 11371: 11370:Louise Arbour 11368: 11365: 11362: 11361: 11359: 11355: 11349:(Taiwan, USA) 11348: 11345: 11342: 11339: 11336: 11333: 11330: 11327: 11324: 11321: 11318: 11315: 11313:(Taiwan, USA) 11312: 11309: 11308: 11306: 11302: 11295: 11292: 11289: 11286: 11283: 11280: 11277: 11276:Pieter Cullis 11274: 11271: 11268: 11265: 11264:Drew Weissman 11262: 11259: 11256: 11253: 11252:Marc Feldmann 11250: 11247: 11244: 11241: 11238: 11235: 11232: 11229: 11226: 11223: 11220: 11217: 11214: 11211: 11208: 11205: 11202: 11199: 11196: 11195: 11193: 11189: 11182: 11181:Omar M. Yaghi 11179: 11176: 11175:Jeffrey Sachs 11173: 11170: 11167: 11164: 11161: 11158: 11155: 11152: 11149: 11146: 11143: 11142: 11140: 11136: 11131: 11124: 11119: 11117: 11112: 11110: 11105: 11104: 11101: 11088: 11084: 11080: 11079:Brian Hoskins 11077: 11074: 11070: 11066: 11063: 11060: 11056: 11055:Drew Weissman 11052: 11049: 11046: 11042: 11038: 11035: 11032: 11028: 11025: 11024: 11022: 11018: 11011: 11007: 11004: 11001: 10997: 10993: 10992:Akira Yoshino 10990: 10987: 10983: 10979: 10976: 10973: 10969: 10966: 10963: 10962:Alain Fischer 10959: 10955: 10952: 10949: 10945: 10942: 10939: 10935: 10932: 10929: 10928:Masato Sagawa 10925: 10921: 10917: 10914: 10911: 10910:Toshio Hirano 10907: 10903: 10899: 10896: 10893: 10889: 10886: 10885: 10883: 10879: 10872: 10871:David E. Kuhl 10868: 10865: 10862: 10858: 10854: 10851: 10848: 10844: 10840: 10837: 10834: 10830: 10829:John Houghton 10827: 10824: 10820: 10816: 10813: 10810: 10806: 10802: 10801:Kenichi Honda 10799: 10796: 10792: 10788: 10785: 10782: 10778: 10774: 10771: 10768: 10764: 10761: 10758: 10754: 10751: 10750: 10748: 10744: 10737: 10733: 10729: 10726: 10723: 10719: 10715: 10712: 10709: 10705: 10701: 10697: 10694: 10691: 10687: 10684: 10681: 10677: 10676:Nick Holonyak 10674: 10671: 10667: 10664: 10661: 10657: 10654: 10651: 10647: 10644: 10641: 10637: 10634: 10631: 10627: 10623: 10619: 10618:Marvin Minsky 10616: 10615: 10613: 10609: 10602: 10598: 10595: 10592: 10588: 10584: 10580: 10576: 10572: 10569: 10566: 10562: 10558: 10555: 10552: 10548: 10545: 10542: 10538: 10535: 10534: 10532: 10528: 10523: 10516: 10511: 10509: 10504: 10502: 10497: 10496: 10493: 10480: 10476: 10473: 10470: 10467: 10464: 10460: 10456: 10453: 10450: 10446: 10443: 10442: 10440: 10436: 10429: 10426: 10423: 10420: 10417: 10413: 10410: 10407: 10403: 10399: 10396: 10393: 10392:Victor Ambros 10389: 10385: 10382: 10379: 10376: 10373: 10369: 10366: 10363: 10360: 10359: 10357: 10353: 10346: 10342: 10339: 10336: 10332: 10328: 10325: 10322: 10318: 10315: 10312: 10308: 10304: 10301: 10298: 10294: 10293:Avram Hershko 10291: 10290: 10288: 10284: 10277: 10274: 10271: 10267: 10264: 10261: 10258: 10255: 10252: 10249: 10245: 10242: 10239: 10238:Judah Folkman 10236: 10233: 10230: 10227: 10224: 10223: 10221: 10217: 10210: 10206: 10203: 10200: 10196: 10195:Henri G. Hers 10193: 10190: 10186: 10183: 10180: 10177: 10174: 10171: 10168: 10164: 10160: 10157: 10154: 10150: 10147: 10144: 10140: 10136: 10133: 10132: 10130: 10126: 10119: 10115: 10111: 10108: 10105: 10101: 10097: 10094: 10093: 10091: 10087: 10083: 10075: 10070: 10068: 10063: 10061: 10056: 10055: 10052: 10036: 10032: 10028: 10024: 10020: 10016: 10014: 10010: 10006: 10002: 10000: 9996: 9995:Yoshua Bengio 9992: 9988: 9984: 9982: 9981:Sarah Gilbert 9978: 9977:Derrick Rossi 9974: 9970: 9966: 9962: 9961:Drew Weissman 9958: 9954: 9952: 9948: 9944: 9940: 9936: 9934: 9930: 9926: 9924: 9920: 9917: 9913: 9909: 9908:Kip S. Thorne 9905: 9901: 9899: 9895: 9893: 9889: 9885: 9884: 9881: 9874: 9870: 9860: 9856: 9855:Mark E. Davis 9852: 9848: 9846: 9842: 9838: 9834: 9832: 9828: 9824: 9822: 9818: 9814: 9813:Joseph Altman 9810: 9808: 9807:Linda Watkins 9804: 9800: 9796: 9794: 9790: 9789:Martin Cooper 9786: 9784: 9780: 9776: 9775:Robert Langer 9772: 9768: 9764: 9762: 9758: 9754: 9752: 9748: 9746: 9742: 9740: 9736: 9732: 9731:Joan Massagué 9728: 9724: 9723:Judah Folkman 9720: 9718: 9714: 9712: 9708: 9704: 9700: 9696: 9694: 9690: 9686: 9682: 9678: 9674: 9672: 9668: 9664: 9662: 9658: 9654: 9652: 9648: 9644: 9642:research team 9641: 9637: 9635: 9631: 9629:of Costa Rica 9628: 9624: 9620: 9618: 9614: 9612: 9608: 9606: 9602: 9600: 9596: 9594: 9590: 9586: 9584: 9580: 9578: 9574: 9570: 9568: 9567:Pablo Rudomín 9564: 9560: 9558: 9554: 9552: 9548: 9544: 9542: 9538: 9536: 9532: 9530: 9526: 9524: 9520: 9519: 9516: 9509: 9505: 9500: 9492: 9487: 9485: 9480: 9478: 9473: 9472: 9469: 9456: 9452: 9448: 9445: 9442: 9438: 9434: 9430: 9427: 9424: 9420: 9417: 9414: 9410: 9406: 9403: 9400: 9396: 9392: 9389: 9386: 9382: 9378: 9377:Brenda Milner 9375: 9372: 9368: 9367:Winfried Denk 9364: 9361: 9358: 9357:James Rothman 9354: 9350: 9347: 9344: 9340: 9336: 9335:Sten Grillner 9333: 9332: 9330: 9326: 9319: 9315: 9311: 9308: 9305: 9301: 9297: 9293: 9290: 9287: 9283: 9279: 9275: 9272: 9269: 9265: 9261: 9258: 9255: 9251: 9247: 9244: 9241: 9237: 9233: 9230: 9227: 9224: 9221: 9217: 9216:Donald Eigler 9214: 9211: 9207: 9206:Louis E. Brus 9204: 9203: 9201: 9197: 9190: 9186: 9183: 9180: 9176: 9172: 9169: 9166: 9165:Andrew Fabian 9163: 9160: 9157: 9154: 9150: 9146: 9145:Ronald Drever 9143: 9140: 9136: 9132: 9129: 9126: 9122: 9118: 9115: 9112: 9108: 9104: 9101: 9098: 9094: 9091: 9090: 9088: 9084: 9079: 9072: 9067: 9065: 9060: 9058: 9053: 9052: 9049: 9036: 9032: 9028: 9027:Thomas Gasser 9024: 9020: 9016: 9015:Sabine Hadida 9012: 9008: 9005: 9002: 8998: 8994: 8990: 8986: 8982: 8979: 8976: 8972: 8968: 8964: 8963:Drew Weissman 8960: 8956: 8953: 8948: 8943: 8939: 8935: 8931: 8928: 8925: 8921: 8917: 8913: 8909: 8906: 8903: 8899: 8895: 8894:Angelika Amon 8891: 8887: 8884: 8881: 8877: 8873: 8869: 8865: 8862: 8859: 8855: 8851: 8850:Roeland Nusse 8847: 8843: 8840: 8837: 8833: 8829: 8825: 8821: 8820:Edward Boyden 8818: 8815: 8811: 8807: 8803: 8802:Victor Ambros 8799: 8795: 8792: 8789: 8785: 8781: 8777: 8773: 8772:Mahlon DeLong 8769: 8766: 8763: 8759: 8755: 8751: 8747: 8743: 8739: 8735: 8731: 8727: 8723: 8720: 8719: 8717: 8715: 8714:Life sciences 8711: 8704: 8700: 8697: 8694: 8693:Peter W. Shor 8690: 8689:David Deutsch 8686: 8682: 8679: 8676: 8672: 8669: 8666: 8662: 8659: 8656: 8655:Blayne Heckel 8652: 8648: 8645: 8642: 8639: 8636: 8632: 8628: 8624: 8621: 8618: 8614: 8611: 8608: 8604: 8601: 8598: 8597:David Spergel 8594: 8590: 8586: 8582: 8579: 8576: 8572: 8568: 8565: 8562: 8558: 8554: 8550: 8546: 8542: 8538: 8534: 8530: 8526: 8525:Atsuto Suzuki 8522: 8521:Daya Bay team 8518: 8514: 8511: 8508: 8504: 8500: 8496: 8495:Ronald Drever 8492: 8489: 8486: 8482: 8478: 8477:Brian Schmidt 8474: 8470: 8467: 8464: 8460: 8459:Michael Green 8457: 8454: 8451: 8448: 8444: 8440: 8439:Guido Tonelli 8436: 8432: 8428: 8424: 8420: 8416: 8413: 8410: 8409:Edward Witten 8406: 8402: 8398: 8394: 8390: 8386: 8385:Alexei Kitaev 8382: 8378: 8375: 8374: 8372: 8370: 8364: 8357: 8356:Simon Brendle 8354: 8351: 8348: 8345: 8342: 8339: 8338:Martin Hairer 8336: 8333: 8330: 8327: 8324: 8321: 8317: 8314: 8311: 8310:Jean Bourgain 8308: 8305: 8302: 8299: 8295: 8291: 8287: 8283: 8280: 8279: 8277: 8275: 8271: 8266: 8259: 8254: 8252: 8247: 8245: 8240: 8239: 8236: 8224: 8221: 8219: 8218:Rino Rappuoli 8216: 8214: 8211: 8209: 8206: 8204: 8201: 8199: 8198:Gerd Faltings 8196: 8194: 8191: 8189: 8186: 8184: 8181: 8179: 8176: 8175: 8173: 8169: 8163: 8160: 8159: 8157: 8153: 8147: 8144: 8142: 8139: 8137: 8134: 8132: 8129: 8127: 8124: 8122: 8119: 8117: 8114: 8112: 8109: 8107: 8104: 8102: 8099: 8097: 8094: 8092: 8089: 8087: 8084: 8082: 8079: 8077: 8074: 8072: 8069: 8067: 8064: 8062: 8059: 8057: 8054: 8052: 8049: 8047: 8044: 8042: 8039: 8037: 8036:Adrian Hayday 8034: 8032: 8029: 8027: 8024: 8022: 8019: 8017: 8014: 8012: 8009: 8007: 8006:Harry Gilbert 8004: 8002: 7999: 7997: 7994: 7992: 7989: 7987: 7984: 7982: 7979: 7977: 7974: 7972: 7969: 7967: 7964: 7962: 7959: 7957: 7954: 7952: 7949: 7947: 7946:James Collier 7944: 7942: 7939: 7937: 7934: 7932: 7929: 7927: 7924: 7922: 7919: 7917: 7916:Polina Bayvel 7914: 7912: 7911:Steven Balbus 7909: 7907: 7904: 7902: 7899: 7898: 7896: 7892: 7888: 7884: 7883:Royal Society 7880: 7873: 7868: 7866: 7861: 7859: 7854: 7853: 7850: 7837: 7834: 7831: 7828: 7825: 7822: 7819: 7816: 7813: 7810: 7807: 7804: 7801: 7798: 7795: 7792: 7789: 7786: 7783: 7780: 7777: 7774: 7771: 7770:Van de Pavert 7768: 7765: 7762: 7759: 7756: 7753: 7750: 7747: 7744: 7741: 7738: 7737: 7735: 7731: 7724: 7721: 7718: 7715: 7712: 7709: 7706: 7703: 7700: 7697: 7694: 7691: 7688: 7685: 7682: 7679: 7678: 7676: 7672: 7665: 7662: 7659: 7656: 7653: 7650: 7647: 7644: 7641: 7638: 7635: 7632: 7629: 7626: 7623: 7620: 7617: 7614: 7611: 7608: 7605: 7602: 7599: 7596: 7593: 7590: 7587: 7584: 7581: 7578: 7575: 7572: 7571: 7569: 7565: 7558: 7555: 7552: 7549: 7546: 7543: 7540: 7537: 7534: 7531: 7528: 7525: 7522: 7519: 7516: 7513: 7510: 7507: 7504: 7501: 7498: 7495: 7492: 7489: 7486: 7483: 7480: 7477: 7474: 7471: 7468: 7465: 7464: 7462: 7458: 7451: 7448: 7445: 7442: 7439: 7436: 7433: 7430: 7427: 7424: 7421: 7418: 7415: 7412: 7409: 7406: 7403: 7400: 7397: 7394: 7391: 7388: 7385: 7382: 7379: 7376: 7373: 7370: 7367: 7364: 7361: 7358: 7355: 7352: 7351: 7349: 7345: 7338: 7335: 7332: 7329: 7326: 7323: 7320: 7317: 7314: 7311: 7308: 7305: 7302: 7299: 7296: 7293: 7290: 7287: 7284: 7281: 7278: 7275: 7272: 7269: 7266: 7263: 7260: 7257: 7254: 7251: 7248: 7245: 7242: 7239: 7236: 7232: 7229: 7226: 7223: 7220: 7217: 7214: 7211: 7208: 7205: 7202: 7199: 7196: 7193: 7190: 7187: 7186: 7184: 7180: 7176: 7169: 7164: 7162: 7157: 7155: 7150: 7149: 7146: 7133: 7130: 7127: 7124: 7121: 7118: 7115: 7112: 7109: 7106: 7103: 7100: 7097: 7094: 7091: 7088: 7085: 7082: 7079: 7076: 7073: 7070: 7067: 7066:Alice Y. Ting 7064: 7061: 7058: 7055: 7052: 7049: 7046: 7043: 7040: 7037: 7034: 7031: 7028: 7025: 7022: 7019: 7016: 7013: 7010: 7007: 7006:Xiaodong Wang 7004: 7001: 6998: 6995: 6992: 6989: 6986: 6983: 6980: 6977: 6974: 6971: 6968: 6965: 6962: 6959: 6956: 6953: 6950: 6947: 6944: 6941: 6938: 6935: 6932: 6929: 6926: 6923: 6920: 6917: 6914: 6911: 6908: 6905: 6902: 6899: 6896: 6893: 6890: 6887: 6884: 6881: 6878: 6875: 6872: 6869: 6866: 6863: 6860: 6857: 6854: 6851: 6848: 6845: 6842: 6839: 6836: 6833: 6830: 6827: 6826:Lubert Stryer 6824: 6821: 6818: 6815: 6812: 6809: 6806: 6803: 6800: 6797: 6794: 6791: 6788: 6785: 6782: 6779: 6776: 6773: 6770: 6767: 6764: 6761: 6758: 6755: 6752: 6749: 6746: 6743: 6740: 6737: 6734: 6731: 6728: 6725: 6722: 6719: 6716: 6713: 6710: 6707: 6706:William Shive 6704: 6701: 6698: 6695: 6692: 6689: 6686: 6683: 6682:John D. Ferry 6680: 6677: 6674: 6671: 6668: 6665: 6662: 6659: 6656: 6653: 6650: 6647: 6644: 6641: 6638: 6635: 6634:Abraham White 6632: 6629: 6626: 6623: 6620: 6619: 6616: 6612: 6605: 6600: 6598: 6593: 6591: 6586: 6585: 6582: 6570: 6567: 6565: 6562: 6560: 6557: 6555: 6552: 6550: 6547: 6545: 6542: 6540: 6537: 6535: 6532: 6530: 6527: 6525: 6522: 6519: 6515: 6514: 6511: 6504: 6501: 6498: 6495: 6494: 6492: 6489: 6485: 6481: 6474: 6471: 6468: 6465: 6462: 6459: 6458: 6456: 6453: 6449: 6445: 6438: 6435: 6432: 6429: 6426: 6425:Roger Penrose 6423: 6422: 6420: 6418: 6414: 6410: 6406: 6403: 6399: 6395: 6390: 6386: 6383: 6379: 6375: 6368: 6365: 6362: 6359: 6358: 6356: 6354: 6350: 6345: 6337: 6332: 6330: 6325: 6323: 6318: 6317: 6314: 6301: 6297: 6296:Louis E. Brus 6293: 6288: 6285: 6281: 6277: 6272: 6269: 6268:Benjamin List 6265: 6260: 6257: 6253: 6248: 6245: 6244:Akira Yoshino 6241: 6237: 6232: 6229: 6225: 6221: 6216: 6213: 6209: 6208:Joachim Frank 6205: 6200: 6197: 6193: 6189: 6184: 6181: 6177: 6173: 6172:Tomas Lindahl 6168: 6165: 6161: 6157: 6152: 6149: 6148:Arieh Warshel 6145: 6141: 6136: 6133: 6132:Brian Kobilka 6129: 6124: 6121: 6120:Dan Shechtman 6116: 6113: 6109: 6105: 6100: 6097: 6096:Ada E. Yonath 6093: 6089: 6084: 6081: 6077: 6073: 6068: 6065: 6060: 6057: 6052: 6049: 6045: 6041: 6036: 6033: 6029: 6028:Avram Hershko 6025: 6020: 6017: 6013: 6008: 6005: 6004:Kurt Wüthrich 6001: 6000:Koichi Tanaka 5997: 5992: 5989: 5985: 5981: 5976: 5975: 5973: 5969: 5962: 5958: 5954: 5949: 5946: 5941: 5938: 5934: 5929: 5926: 5922: 5918: 5917:Paul D. Boyer 5913: 5910: 5906: 5902: 5897: 5894: 5890: 5886: 5881: 5878: 5873: 5870: 5869:Michael Smith 5866: 5861: 5858: 5853: 5850: 5845: 5842: 5837: 5834: 5830: 5829:Sidney Altman 5825: 5822: 5818: 5814: 5809: 5806: 5802: 5798: 5793: 5790: 5786: 5782: 5777: 5774: 5770: 5765: 5762: 5757: 5754: 5749: 5746: 5741: 5738: 5734: 5733:Kenichi Fukui 5729: 5726: 5722: 5718: 5713: 5710: 5706: 5701: 5698: 5693: 5690: 5685: 5682: 5677: 5676: 5674: 5670: 5663: 5659: 5654: 5651: 5646: 5643: 5639: 5634: 5631: 5630:William Stein 5627: 5623: 5618: 5615: 5610: 5607: 5602: 5599: 5595: 5590: 5587: 5582: 5579: 5578:George Porter 5575: 5571: 5570:Manfred Eigen 5566: 5563: 5558: 5555: 5550: 5547: 5542: 5539: 5535: 5530: 5527: 5523: 5518: 5515: 5514:Melvin Calvin 5510: 5507: 5506:Willard Libby 5502: 5499: 5494: 5491: 5486: 5483: 5478: 5475: 5471: 5466: 5463: 5458: 5455: 5454:Linus Pauling 5450: 5447: 5442: 5439: 5438:Richard Synge 5435: 5434:Archer Martin 5430: 5427: 5423: 5418: 5417: 5415: 5411: 5404: 5400: 5395: 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1951: 1949: 1932: 1928: 1921: 1919: 1917: 1915: 1906: 1904:9780837909240 1900: 1896: 1895: 1888: 1872: 1868: 1862: 1846: 1842: 1841: 1836: 1829: 1827: 1810: 1806: 1800: 1784: 1780: 1774: 1772: 1755: 1751: 1745: 1743: 1726: 1722: 1716: 1700: 1696: 1692: 1686: 1684: 1682: 1680: 1678: 1676: 1674: 1672: 1655: 1651: 1645: 1643: 1626: 1622: 1621: 1616: 1609: 1593: 1589: 1582: 1580: 1578: 1576: 1574: 1572: 1570: 1568: 1566: 1564: 1556: 1550: 1548: 1546: 1544: 1542: 1540: 1538: 1536: 1534: 1532: 1530: 1528: 1526: 1524: 1504: 1497: 1491: 1489: 1487: 1485: 1483: 1481: 1479: 1477: 1475: 1458: 1454: 1450: 1444: 1442: 1425: 1421: 1420: 1415: 1408: 1406: 1389: 1385: 1384: 1379: 1373: 1371: 1354: 1350: 1346: 1340: 1336: 1328: 1326: 1324: 1319: 1317: 1311: 1309: 1305: 1301: 1297: 1293: 1289: 1285: 1281: 1277: 1273: 1270:in 2003, the 1269: 1266:in 2002, the 1265: 1260: 1258: 1254: 1250: 1246: 1242: 1238: 1234: 1230: 1226: 1222: 1218: 1214: 1210: 1206: 1202: 1198: 1194: 1190: 1186: 1182: 1178: 1174: 1170: 1166: 1162: 1157: 1155: 1151: 1147: 1143: 1139: 1135: 1131: 1128:Doudna was a 1125: 1115: 1113: 1109: 1105: 1100: 1096: 1092: 1088: 1080:Personal life 1077: 1075: 1072:. She joined 1071: 1067: 1063: 1059: 1054: 1052: 1048: 1044: 1034: 1033:based tests. 1032: 1028: 1024: 1014: 1012: 1002: 999: 997: 992: 990: 985: 983: 979: 975: 970: 967:and Harvard. 966: 962: 953: 949: 947: 943: 939: 935: 931: 925: 923: 919: 915: 910: 909:Streptococcus 907:found in the 906: 901: 897: 887: 885: 880: 876: 872: 867: 865: 861: 857: 853: 848: 846: 842: 838: 829: 825: 821: 812: 808: 803: 798: 790: 788: 784: 783:Henry Ford II 779: 777: 773: 769: 765: 762: 761: 751: 738: 734: 732: 728: 725: 721: 717: 712: 709: 705: 701: 691: 689: 685: 681: 677: 673: 669: 665: 661: 651: 649: 645: 641: 637: 636:Corwin Hansch 633: 629: 625: 621: 616: 614: 610: 607:lab of noted 606: 602: 597: 594: 592: 587: 583: 579: 575: 570: 568: 564: 554: 552: 548: 546: 541: 537: 533: 529: 525: 521: 517: 513: 506: 501: 497: 495: 491: 487: 483: 478: 476: 472: 468: 464: 461: 458: 454: 450: 446: 442: 436: 408: 404: 395: 389: 385: 381: 377: 370: 368: 365: 363: 360: 358: 355: 354: 352: 348: 345: 342: 338: 335: 332: 330: 326: 319: 318: 314: 312: 308: 302: 299: 297: 294: 292: 289: 287: 284: 282: 279: 278: 276: 272: 266: 263: 260: 257: 255: 252: 250: 247: 246: 244: 240: 237: 233: 227: 226: 222: 219: 216: 212: 208: 205: 201: 198: 195: 192: 189: 186: 183: 180: 177: 174: 171: 168: 165: 162: 161: 159: 155: 136: 132: 126: 123: 121: 118: 115: 114: 112: 108: 101: 97: 94: 91: 87: 84: 83: 81: 77: 72: 66:(age 60) 52: 48: 41: 36: 31: 22: 19: 11384:(Bangladesh) 11378:(UK, Israel) 11323:Stephen Owen 11311:Yu Ying-shih 11234:Brian Druker 11215: 11204:Tasuku Honjo 11169:Jane Goodall 11157:James Hansen 11031:Svante Pääbo 10981: 10968:Hideo Hosono 10938:Jean Fréchet 10920:Brian Druker 10916:Janet Rowley 10898:Ken Thompson 10815:Makoto Nagao 10777:Anne McLaren 10718:Jozef Schell 10646:Gerhard Ertl 10640:John J. Wild 10626:Dan McKenzie 10591:Robert Gallo 10583:Frank Fenner 10561:Gurdev Khush 10475:Botond Roska 10448: 10398:John Kappler 10362:Axel Ullrich 10345:Aharon Razin 10341:Howard Cedar 10266:Michael Sela 10260:Mary F. Lyon 10189:Meir Wilchek 10110:Roger Sperry 10100:Jean Dausset 10096:George Snell 9973:Özlem Türeci 9929:Joanne Chory 9923:Svante Pääbo 9904:Rainer Weiss 9891: 9803:Baruch Minke 9799:David Julius 9767:Sumio Iijima 9761:Ginés Morata 9717:Jane Goodall 9681:John Sulston 9677:Craig Venter 9667:Robert Gallo 9611:Amable Liñán 9523:Alberto Sols 9433:Harry T. Orr 9419:David Julius 9381:John O'Keefe 9371:Ann Graybiel 9328:Neuroscience 9263: 9254:Calvin Quate 9210:Sumio Iijima 9171:Roger Ulrich 9153:Rainer Weiss 9135:Andrei Linde 9086:Astrophysics 8975:Pascal Mayer 8920:David Julius 8902:Zhijian Chen 8868:Peter Walter 8864:Joanne Chory 8836:Svante Pääbo 8809: 8734:Hans Clevers 8660: 8622: 8613:Charles Kane 8602: 8585:Gary Hinshaw 8503:Rainer Weiss 8490: 8449:(LHC) (2013) 8414: 8393:Andrei Linde 8208:Svante Pääbo 8192: 8162:Adair Turner 8146:Paul Workman 8131:Alison Smith 8126:Ted Shepherd 8096:Luke O'Neill 8011:Patrick Gill 8001:Pratibha Gai 7931:John Burrows 7830:Van Lieshout 7824:Dröge Wendel 7324: 7048:Anna K. Mapp 7011: 6994:John Kuriyan 6970:Peter S. Kim 6934:Peter Walter 6904:Richard Axel 6856:Mark Ptashne 6766:James Watson 6730:Harvey Itano 6646:Eric G. Ball 6548: 6497:Paul Milgrom 6389:Louise Glück 6366: 6255: 6228:George Smith 6108:Akira Suzuki 6064:Gerhard Ertl 6048:Yves Chauvin 5996:John B. Fenn 5984:Ryoji Noyori 5971:2001–present 5945:Ahmed Zewail 5905:Harold Kroto 5889:Mario Molina 5817:Robert Huber 5789:John Polanyi 5773:Jerome Karle 5709:Georg Wittig 5594:Derek Barton 5586:Lars Onsager 5538:Giulio Natta 5534:Karl Ziegler 5526:John Kendrew 5294:Richard Kuhn 5219:Hans Fischer 5035:Otto Wallach 4971:Emil Fischer 4853: 4841: 4829: 4798: 4797: 4796:profile for 4793: 4764:. Retrieved 4745: 4730:Bibliography 4715:. Retrieved 4710: 4701: 4689:. Retrieved 4685: 4676: 4664:. Retrieved 4649: 4637:. Retrieved 4628: 4619: 4607:. Retrieved 4593: 4581:. Retrieved 4568: 4559: 4547:. Retrieved 4534: 4525: 4511:. Retrieved 4495: 4485:. Retrieved 4466: 4454:. Retrieved 4440: 4430:November 13, 4428:. Retrieved 4419: 4410: 4400:November 13, 4398:. Retrieved 4389: 4380: 4368:. Retrieved 4354: 4344:November 12, 4342:. Retrieved 4338:the original 4334:luiprize.org 4333: 4324: 4312:. Retrieved 4298: 4265: 4259: 4218: 4212: 4206: 4194:. Retrieved 4185: 4175: 4163:. 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Retrieved 1348: 1339: 1322: 1315: 1312: 1300:Pope Francis 1261: 1233:Harvey Prize 1199:(2016), the 1195:(2015), the 1158: 1127: 1083: 1055: 1040: 1027:Robert Tjian 1020: 1008: 1000: 993: 988: 986: 958: 926: 908: 893: 868: 849: 834: 780: 758: 756: 707: 697: 657: 632:Fred Grieman 628:biochemistry 617: 598: 589: 586:James Watson 574:Hilo, Hawaii 571: 560: 544: 534:(2016), the 530:(2015), the 509: 479: 477:since 1997. 467:cell biology 460:Chancellor's 402: 401: 334:Jack Szostak 315: 274:Institutions 265:Gene editing 249:Biochemistry 235: 223: 18: 11434:1964 births 11402:(Australia) 11364:Albie Sachs 11357:Rule of Law 11347:Hsu Cho-yun 11337:(Australia) 11335:Wang Gungwu 10839:Albert Fert 10795:Seiji Ogawa 10656:Frank Press 10522:Japan Prize 10455:Joan Steitz 10388:Gary Ruvkun 10335:Tony Pawson 10276:Eric Kandel 10205:John Gurdon 9947:Terence Tao 9837:Peter Higgs 9783:Tobin Marks 9727:Tony Hunter 9707:Vinton Cerf 9583:Guido Münch 9441:Huda Zoghbi 9343:Pasko Rakic 9292:Jacob Sagiv 9274:Harald Rose 9246:Gerd Binnig 9240:John Pendry 9236:Stefan Hell 9199:Nanoscience 9185:Sara Seager 9179:Conny Aerts 9111:Roger Angel 9078:Kavli Prize 8993:John Jumper 8945: [ 8930:David Baker 8876:Kim Nasmyth 8858:Huda Zoghbi 8832:Helen Hobbs 8806:Gary Ruvkun 8746:Eric Lander 8617:Eugene Mele 8575:Cumrun Vafa 8563:team (2016) 8517:Kam-Biu Luk 8513:Yifang Wang 8423:Peter Jenni 8367:Fundamental 8294:Terence Tao 8290:Jacob Lurie 8274:Mathematics 8213:Pasko Rakic 8178:Robert Cava 8136:David Wales 8111:Jim Prosser 8071:David Lodge 8066:Mark Lemmon 8046:David Hight 8016:Anne Glover 7986:Artur Ekert 7961:Jack Cuzick 7921:Graham Bell 7901:Chris Abell 7634:McKitterick 7580:Van der Wee 7108:Howard Hang 6844:C. Fred Fox 6640:George Wald 6344:Nobel Prize 6196:Ben Feringa 6180:Aziz Sancar 6160:Stefan Hell 6156:Eric Betzig 5933:Walter Kohn 5901:Robert Curl 5877:George Olah 5865:Kary Mullis 5841:Elias Corey 5833:Thomas Cech 5785:Yuan T. Lee 5753:Henry Taube 5286:Paul Karrer 5274:Peter Debye 5254:Harold Urey 5140:Fritz Pregl 5101:Fritz Haber 5043:Marie Curie 4666:October 15, 4609:October 15, 4456:October 18, 4370:January 13, 4314:November 9, 4105:November 1, 4079:October 24, 4053:November 6, 3967:November 6, 3537:November 8, 3492:October 10, 2351:Marketplace 2287:October 15, 2045:October 22, 2023:October 22, 1997:October 22, 1937:October 24, 1815:November 1, 1789:November 2, 1760:November 1, 1731:October 24, 1705:October 31, 1660:October 31, 1512:October 24, 1359:October 14, 1298:. In 2021, 1225:Kavli Prize 1201:Japan Prize 1095:Tetrahymena 1091:Thomas Cech 996:hepatitis C 884:microbiomes 879:Li Ka Shing 841:synchrotron 760:Tetrahymena 708:Tetrahymena 702:enzymes or 676:Thomas Cech 540:Japan Prize 457:Li Ka Shing 362:Janice Chen 344:Thomas Cech 204:Nanoscience 200:Kavli Prize 194:Japan Prize 11428:Categories 11390:(Colombia) 11376:Joseph Raz 11222:Feng Zhang 11130:Tang Prize 11010:Rattan Lal 10986:Adi Shamir 10833:Akira Endo 10753:Ian McHarg 10700:Bruce Ames 10579:Isao Arita 10524:recipients 10270:Ruth Arnon 9991:Yann LeCun 9969:Uğur Şahin 9939:Yves Meyer 9914:, and the 9391:Eve Marder 9282:Knut Urban 9149:Kip Thorne 8828:John Hardy 8699:John Cardy 8499:Kip Thorne 8481:Adam Riess 8445:(CMS) and 8405:Ashoke Sen 8332:Alex Eskin 8203:John Hayes 8188:Mark Davis 8086:Gil McVean 8026:Ian Graham 7981:Jim Dunlop 7966:Bill David 7788:Van Golden 7705:McClelland 7521:Brunekreef 7450:Deisseroth 7372:Montagnier 7213:Van Deenen 7096:Minkui Luo 6790:Bruce Ames 6378:Literature 6032:Irwin Rose 6012:Peter Agre 5937:John Pople 5745:Aaron Klug 5650:Paul Flory 5598:Odd Hassel 5522:Max Perutz 5403:Kurt Alder 5399:Otto Diels 5227:Carl Bosch 4475:. London: 3517:October 9, 3453:SynBioBeta 3272:August 26, 3243:October 8, 2427:Doudna Lab 2398:"About Us" 2206:October 8, 2067:October 7, 1463:October 7, 1430:October 7, 1331:References 1243:) and the 1241:Feng Zhang 1197:Tang Prize 1173:Feng Zhang 1058:Altos Labs 969:Feng Zhang 942:moratorium 609:mycologist 532:Tang Prize 441:biochemist 211:Wolf Prize 188:Tang Prize 139:Jamie Cate 60:1964-02-19 11408:(Ireland) 11396:(Lebanon) 11296:(Denmark) 11132:laureates 10853:Vint Cerf 10714:Leo Esaki 10690:Masao Ito 10139:Leo Sachs 9898:Hugh Herr 9640:Atapuerca 9131:Alan Guth 9080:laureates 9007:Carl June 8938:Dennis Lo 8447:Lyn Evans 8429:(ATLAS), 8381:Alan Guth 8267:laureates 8183:Vint Cerf 7956:Brian Cox 7812:Struycken 7800:B. Visser 7794:Koelewijn 7752:C. Visser 7717:Kanwisher 7693:Tomasello 7610:Schilling 7497:Harremoës 7444:Carmeliet 7402:Blackburn 7354:Lauterbur 7235:Reichardt 7225:Weissmann 7126:Yimon Aye 6760:Paul Berg 6433:(Germany) 6353:Chemistry 6346:laureates 5717:Paul Berg 5672:1976–2000 5413:1951–1975 5343:Otto Hahn 5174:1926–1950 4954:1901–1925 4943:Laureates 4865:Fresh Air 4717:April 28, 4639:April 12, 4487:April 29, 4290:158427487 4282:0036-8075 4251:Q74441382 4235:1476-4687 4196:March 16, 3997:August 1, 3365:April 14, 3335:April 14, 3305:April 14, 2991:Bloomberg 2915:Q22065510 2889:0036-8075 2847:Paul Berg 2796:March 16, 2772:Q34533807 2756:0966-842X 2718:Q24679073 2692:0021-9193 2648:Q24669850 2622:0036-8075 2239:303754572 2175:Q34553023 2149:0027-8424 856:Genentech 852:Genentech 724:catalytic 704:ribozymes 615:in 1981. 510:Doudna's 225:Full list 79:Education 11372:(Canada) 11304:Sinology 11278:(Canada) 11212:(France) 11147:(Norway) 10313:(2002/3) 10262:(1996/7) 10256:(1995/6) 10250:(1994/5) 10175:(1984/5) 9121:Jane Luu 8559:and the 8547:and the 8535:and the 8527:and the 8519:and the 8304:Ian Agol 8155:Honorary 8021:Neil Gow 7885:elected 7681:Anderson 7604:De Vries 7557:Le Quéré 7533:Laurance 7503:Thompson 7467:Lovelock 7420:Steinman 7384:Marshall 7360:Van Rood 7347:Medicine 7337:Stillman 7313:De Lange 7295:Jeffreys 7259:Berridge 7189:Chargaff 6363:(France) 4766:March 8, 4739:(2021). 4660:Archived 4633:Archived 4603:Archived 4583:June 26, 4577:Archived 4549:June 26, 4543:Archived 4513:March 9, 4507:Archived 4481:Archived 4450:Archived 4424:Archived 4394:Archived 4364:Archived 4308:Archived 4247:Wikidata 4243:29872189 4190:Archived 4159:Archived 4129:Archived 4099:Archived 4073:Archived 4021:Archived 3961:Archived 3927:. 2006. 3909:Archived 3860:Archived 3856:BioSpace 3837:Archived 3812:Archived 3787:Archived 3762:Archived 3735:Archived 3708:Archived 3685:Archived 3660:Archived 3633:Archived 3610:Archived 3585:Archived 3486:Archived 3457:Archived 3426:Archived 3359:Archived 3355:BioSpace 3329:Archived 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7879:Fellows 7806:Manders 7764:Martens 7740:Verhoef 7723:Zatorre 7687:Dehaene 7658:McNeill 7646:Assmann 7616:Le Goff 7592:Oberman 7567:History 7491:Ehrlich 7473:Branica 7438:Jackson 7432:Alitalo 7426:Clevers 7378:De Wied 7366:Moncada 7301:Szostak 7277:Rothman 7207:De Duve 7195:Brachet 7090:Yi Tang 6417:Physics 4945:of the 4881:YouTube 4854:YouTube 4842:YouTube 4830:YouTube 4792:has an 4790:Scholia 4691:June 6, 4266:Science 3181:May 16, 3151:May 16, 3069:Science 2898:4394183 2869:Bibcode 2860:Science 2710:3316184 2631:6286148 2602:Bibcode 2593:Science 2127:Bibcode 1631:May 12, 1152:of the 1031:qRT-PCR 720:Enzymes 680:h-index 469:at the 451:, with 383:Website 261:biology 151:​ 143:​ 11089:(2024) 11075:(2023) 11061:(2022) 11047:(2021) 11033:(2020) 11012:(2019) 11002:(2018) 10988:(2017) 10974:(2016) 10964:(2015) 10950:(2014) 10930:(2012) 10912:(2011) 10894:(2010) 10873:(2009) 10863:(2008) 10849:(2007) 10835:(2006) 10825:(2005) 10811:(2004) 10797:(2003) 10783:(2002) 10769:(2001) 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Index

ForMemRS

Washington, D.C.
Pomona College
BA
Harvard University
MA
RNA interference
CRISPR
Alan T. Waterman Award
Jacob Heskel Gabbay Award
Breakthrough Prize in Life Sciences
Princess of Asturias Award
Tang Prize
Japan Prize
Kavli Prize
Nanoscience
Wolf Prize
Nobel Prize in Chemistry
Full list
Biochemistry
CRISPR
RNA
Gene editing
University of Colorado, Boulder
Yale University
University of California, Berkeley
Gladstone Institutes
University of California, San Francisco
Thesis

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