20:
400:
385:
357:
306:
326:
338:
258:
356:
138:, but does not adhere to it. In many species, what is called the "seed" is an achene, a fruit containing the seed. The seed-like appearance is owed to the hardening of the fruit wall (pericarp), which encloses the solitary seed so closely as to seem like a seed coat.
305:
384:
399:
325:
192:(aka floral tube), which is a structure where basal portions of the calyx, the corolla, and the stamens unite with the receptacle to form a cup-shaped tube.
337:
374:) disperses in the wind, either as single achenes or as the entire aggregate of achenes. The achenes have long hairy appendages that developed from the
19:
274:
is also so similar to an achene that it is often considered to be one, although it derives from a compound inferior ovary (with one
236:
is a type of fruit that closely resembles an achene, but differs in that the pericarp is fused to the thin seed coat in the grain.
552:
278:). A special term for the Asteraceae fruit is cypsela (plural cypselae or cypselas). For example, the white-gray husks of a
89:
518:
579:
211:
Some achenes have accessory hair-like structures that cause them to tumble in the wind in a manner similar to a
486:
510:
763:
438:
535:
443:
618:
61:
458:
503:
502:
480:
572:
291:
393:
showing 4 stages of inflorescence, with two reddish-brown flower heads containing cypselas
181:
with an aggregate of achenes on its outer surface, and what is eaten is accessory tissue.
8:
721:
127:
475:
221:
737:
524:
514:
363:
216:
68:
640:
532:
37:
432:
331:
A microscopic view of a dandelion "clock" showing the receptacle and the cypselas.
711:
696:
625:
565:
240:
178:
119:
716:
706:
665:
608:
316:
287:
251:
250:
are sometimes considered achenes although their one-locule ovary is a compound
205:
188:
produces an aggregate of achene fruits that are encompassed within an expanded
757:
371:
312:
528:
701:
655:
613:
742:
670:
271:
247:
212:
189:
174:
509:(Second ed.). Spring Lake, Utah: Spring Lake Publishing. p.
630:
375:
283:
279:
229:
151:
147:
95:
82:
74:
680:
479:
167:
163:
135:
557:
675:
257:
155:
650:
645:
409:
282:"seed" are the walls of the cypsela fruit. Many cypselas (e.g.
275:
263:
159:
123:
243:
is like an achene, but the fruit is bladder-like or inflated.
49:
635:
589:
233:
201:
115:
660:
185:
131:
40:
505:
130:(they do not open at maturity). Achenes contain a single
177:
are sometimes mistaken for seeds. The strawberry is an
52:
46:
43:
431:
215:. This type sometimes is called a tumble fruit or
755:
501:Harris, James G.; Harris, Melinda Woolf (2001).
430:
315:) can disperse in the wind due to the hair-like
474:
122:. Achenes are monocarpellate (formed from one
573:
311:Cypselae on a dandelion "clock" (the matured
500:
350:are achenes with large wing-like structures.
26:hypanthium encircling separate achene fruits
408:flower head containing cypselas forming as
580:
566:
478:; Peck, H. T.; Colby, F. M., eds. (1905).
256:
102: 'to gape'), also sometimes called
18:
756:
16:Class of simple non-opening dry fruits
561:
494:
490:(1st ed.). New York: Dodd, Mead.
470:
468:
459:participating institution membership
587:
13:
465:
290:tissue attached that functions in
14:
775:
546:
398:
383:
355:
336:
324:
304:
36:
487:New International Encyclopedia
424:
1:
417:
195:
118:produced by many species of
7:
141:
10:
780:
297:
114:, is a type of simple dry
88:
67:
730:
689:
596:
439:Oxford English Dictionary
319:tissue above each ovary.
270:The fruit of the family
200:A winged achene, as in
536:Kahle/Austin Foundation
444:Oxford University Press
267:
134:that nearly fills the
27:
260:
170:are typical achenes.
22:
690:Categories of fruits
292:biological dispersal
442:(Online ed.).
406:Coreopsis tinctoria
391:Coreopsis tinctoria
173:The achenes of the
553:Botanical Glossary
268:
222:Anemone virginiana
28:
751:
750:
457:(Subscription or
348:Acer buergerianum
219:. An example is
106:and occasionally
771:
764:Fruit morphology
582:
575:
568:
559:
558:
540:
539:
533:Internet Archive
508:
498:
492:
491:
483:
472:
463:
462:
454:
452:
450:
435:
428:
402:
387:
359:
340:
328:
308:
120:flowering plants
99:
92:
78:
71:
59:
58:
55:
54:
51:
48:
45:
42:
779:
778:
774:
773:
772:
770:
769:
768:
754:
753:
752:
747:
726:
722:Dehiscent fruit
712:Aggregate fruit
697:Accessory fruit
685:
597:Types of fruits
592:
586:
549:
544:
543:
521:
499:
495:
473:
466:
456:
448:
446:
429:
425:
420:
413:
412:finish blooming
403:
394:
388:
379:
360:
351:
341:
332:
329:
320:
309:
300:
198:
179:accessory fruit
144:
39:
35:
17:
12:
11:
5:
777:
767:
766:
749:
748:
746:
745:
740:
734:
732:
728:
727:
725:
724:
719:
717:Multiple fruit
714:
709:
707:Compound fruit
704:
699:
693:
691:
687:
686:
684:
683:
678:
673:
668:
663:
658:
653:
648:
643:
638:
633:
628:
623:
622:
621:
616:
606:
600:
598:
594:
593:
585:
584:
577:
570:
562:
556:
555:
548:
547:External links
545:
542:
541:
519:
493:
481:"Achene"
464:
422:
421:
419:
416:
415:
414:
404:
397:
395:
389:
382:
380:
378:of the flower.
361:
354:
352:
342:
335:
333:
330:
323:
321:
310:
303:
299:
296:
204:, is called a
197:
194:
146:The fruits of
143:
140:
85:' and
15:
9:
6:
4:
3:
2:
776:
765:
762:
761:
759:
744:
741:
739:
736:
735:
733:
729:
723:
720:
718:
715:
713:
710:
708:
705:
703:
700:
698:
695:
694:
692:
688:
682:
679:
677:
674:
672:
669:
667:
664:
662:
659:
657:
654:
652:
649:
647:
644:
642:
639:
637:
634:
632:
629:
627:
624:
620:
617:
615:
612:
611:
610:
607:
605:
602:
601:
599:
595:
591:
583:
578:
576:
571:
569:
564:
563:
560:
554:
551:
550:
537:
534:
530:
526:
522:
520:0-9640221-6-8
516:
512:
507:
506:
497:
489:
488:
482:
477:
476:Gilman, D. C.
471:
469:
460:
445:
441:
440:
434:
427:
423:
411:
407:
401:
396:
392:
386:
381:
377:
373:
372:Ranunculaceae
369:
365:
358:
353:
349:
345:
339:
334:
327:
322:
318:
314:
307:
302:
301:
295:
294:of the seed.
293:
289:
285:
281:
277:
273:
266:
265:
259:
255:
253:
249:
244:
242:
237:
235:
231:
226:
224:
223:
218:
214:
209:
207:
203:
193:
191:
187:
182:
180:
176:
171:
169:
165:
161:
157:
153:
149:
139:
137:
133:
129:
125:
121:
117:
113:
109:
105:
101:
98:
91:
87:
84:
80:
77:
70:
66:
63:
62:Ancient Greek
57:
33:
25:
21:
702:Simple fruit
603:
531:– via
504:
496:
485:
447:. Retrieved
437:
426:
405:
390:
367:
347:
343:
269:
262:
245:
238:
227:
220:
210:
199:
183:
172:
145:
111:
107:
103:
96:
93:
86:
75:
72:
65:
31:
29:
23:
743:Drift fruit
614:Hesperidium
261:Cypsela of
128:indehiscent
60:; from
671:Schizocarp
461:required.)
418:References
368:Pulsatilla
272:Asteraceae
246:Fruits of
213:tumbleweed
196:Variations
190:hypanthium
175:strawberry
112:achenocarp
631:Caryopsis
588:Types of
313:capitulum
284:dandelion
280:sunflower
230:caryopsis
152:buckwheat
148:buttercup
83:privative
758:Category
738:Diaspore
731:Function
681:Syconium
641:Follicle
529:45951032
433:"achene"
370:(family
364:diaspore
217:diaspore
168:cannabis
164:amaranth
142:Examples
136:pericarp
108:achenium
97:khaínein
676:Silique
626:Capsule
449:30 July
410:florets
344:Samaras
298:Gallery
286:) have
241:utricle
156:caraway
90:χαίνειν
81: '
666:Samara
651:Loment
646:Legume
604:Achene
590:fruits
527:
517:
276:locule
264:Cynara
248:sedges
206:samara
166:, and
160:quinoa
126:) and
124:carpel
32:achene
636:Drupe
609:Berry
455:
376:style
317:calyx
288:calyx
252:ovary
234:grain
202:maple
116:fruit
104:akene
64:
661:Pome
619:Pepo
525:OCLC
515:ISBN
451:2018
362:The
186:rose
132:seed
24:Rosa
656:Nut
511:131
366:of
346:of
239:An
232:or
110:or
30:An
760::
523:.
513:.
484:.
467:^
436:.
254:.
228:A
225:.
208:.
184:A
162:,
158:,
154:,
150:,
50:iː
581:e
574:t
567:v
538:.
453:.
100:)
94:(
79:)
76:a
73:(
69:ἀ
56:/
53:n
47:k
44:ˈ
41:ə
38:/
34:(
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.