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In one of the exhibition rooms of the Museum of the
Zoological Institute of the Academy of Sciences of the U.S.S.R. in Leningrad there is a stuffed adult domestic horse with stripes. was purchased – while the horse was still alive – from its owner, a Leningrad cabman, and delivered by the latter to
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mode of inheritance. Female horses with this gene have a striped coat pattern; hairs from the stripes have a different texture as well as color, are less straight, and are more unruly. Male horses have sparse manes and tails but do not show a striped coat texture pattern. A Brindle1 test is
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204:. This makes it very difficult to test for, and there are currently no commercially available tests that are able to detect brindle. Brindle dogs will usually test as KBky, and carriers (one dominant black allele, one brindle) cannot be identified without breeding.
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The brindled pattern found in horses could be described as vertical stripes that are found along the neck, back, hindquarters, and upper legs. The horse's head is usually a solid color and is not affected by the striping. The brindled pattern has no effect on dark
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Brindle typically appears as black stripes on a red base. The stripes are eumelanin (black/brown pigment) and the base is phaeomelanin (red/yellow pigment), so the appearance of those pigments can be changed by any of the genes which usually affect them.
344:. It usually does not affect the head and legs as much as the body, with the heaviest concentrations of brindling being on the neck, shoulders and hindquarters. The coloring has been documented in the past. At the
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genes in one horse. These forms are not heritable. In some horses the pattern seems to be inherited, indicating that one or more genes are responsible. One heritable brindle pattern in a family of
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The word brindled is a variant of Middle
English brended (early 15c.), from bren "brown color" (13c.), noun from past participle of brennen "burn" (from Proto-Germanic *brennan "to burn", from
240:, although the latter is not acceptable in the show ring. The "dark" markings are black or the dilutions gray (called blue) or brown (sometimes called red). It is not uncommon for a brindled
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232:). This coloration looks very similar to tricolor, and can be distinguished only at close range. Dogs of this color are often described as "trindle". It can also occur in combination with
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Brindle is caused by a complex gene process and is technically a form of mosaicism, where some cells express one allele (KB) and other cells express a different allele (ky), a little like
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Brindle coloring consists of irregular stripes extending vertically over the horse's body and horizontally around the legs. Brindle horses can also have a dorsal stripe when caused by
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With this coat pattern there is a base coat that covers the entire body of the horse. This base coat color can be any color. Recorded examples have been
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had a brindled coat. He was the companion and household protector of the
Ingalls family in their early travels. He dies of old age at the beginning of
280:. They are difficult to breed to perfection, as the black and red hairs should intermingle evenly all over. Brindle guinea pigs' fur type is
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Eumelanin (the pigment making up the stripes) can be affected by: merle (and harlequin), liver, dilution, greying, and recessive red.
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root *gwher- "to heat, warm"). The etymological sense of the adjective appears to be "marked as though by branding or burning".
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type. closely resembles the striping found in many breeds of cattle and in dogs and known as the "brindling pattern".
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as a base coat. Over top of the base color is either a lighter or darker color, giving the appearance of stripes.
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in the 1880s. FitzPatrick later collected tales of their adventures into a popular book of the same name.
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horse of brindle coloring from the early 19th century was mounted and put on display due to its rarity.
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Brindle coloring in horses is extremely rare and most commonly caused by pigment concentrating along
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the Museum after the horse's death. Judging by the stuffed object, it was a large horse of the
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692:"An Intronic MBTPS2 Variant Results in a Splicing Defect in Horses with Brindle Coat Texture"
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during fetal development. In two confirmed cases it has been linked to spontaneous
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Phaeomelanin (the pigment making up the base) can be affected by: Intensity locus.
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was identified in 2016 and named
Brindle1 (BR1). The Brindle1 phenotype has an
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on horses. Some brindle-colored horses are more eye-catching than others.
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And of course, white markings and ticking can occur on any brindle dog.
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Brindle coloration is less distinct on longer-haired dogs, like the
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Genetics of dog brindling with reference to
Shetland Sheepdogs
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University of
California-Davis Veterinary Genetics Laboratory
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Lusis, J.A. (1942). "Striping
Patterns in Domestic Horses".
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in the points, or as a brindle merle, in breeds such as the
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For rose-moles all in stipple upon trout that swim ...
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388:. Earliest documented cases were said to have red dun or
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to become progressively more black or silver as it ages.
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790:"Lizards 101: Crested Gecko Morphs and Color Variations"
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A European domestic short-haired, female, brindled cat.
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The brindle pattern may also take the place of tan in
67:. Unsourced material may be challenged and removed.
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836:Dog Coat Colour: Genetics & pigmentation
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524:" (1918), the concept occurs in the opening:
690:Murgiano, Leonardo; Waluk, Dominik (2016).
536:For skies of couple-color as a brinded cow;
442:, the term "brindle" is used to describe a
346:Zoological Museum of the Academy of Science
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467:A brindle horse was mentioned in the book
316:, resulting in an animal with two sets of
158:coloring pattern in animals, particularly
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260:Brindled domestic cats are also known as
127:Learn how and when to remove this message
486:in his travels around the South African
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304:Brindle horse with dark bay base coat
27:Coat coloring pattern in some animals
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533:Glory be to God for dappled things —
65:adding citations to reliable sources
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846:Genetics of brindling in the Corgi
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571:Thrice the brinded cat hath mew'd.
228:coats of some dog breeds (such as
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817:Muir, Kenneth (1962). "Macbeth".
635:"The Abyssinian Guinea Pig Guide"
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52:needs additional citations for
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446:with darker stripes of color.
419:is a species of brindled gnu.
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147:with the brindle color pattern
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276:Brindle is an old variety in
696:G3: Genes, Genomes, Genetics
641:. 2015-09-09. Archived from
510:By the Shores of Silver Lake
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576:However, in this context –
500:Little House on the Prairie
408:Brindle coloring exists in
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1388:Category:Horse coat colors
1357:Endothelin receptor type B
1322:Equine coat color genetics
480:Staffordshire Bull Terrier
296:A reverse brindle chestnut
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32:Brindle (disambiguation)
1377:Melanocortin 1 receptor
851:Cat coat color genetics
432:Tswalu Kalahari Reserve
326:American Quarter Horses
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804:"Etymology Dictionary"
667:"Brindle Coat Texture"
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1315:Genetics and breeding
1156:Base color variations
1085:Lethal white syndrome
819:The Arden Shakespeare
708:10.1534/g3.116.032433
518:Gerard Manley Hopkins
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264:, and in some cases,
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505:Laura Ingalls Wilder
476:Jock of the Bushveld
238:Cardigan Welsh Corgi
61:improve this article
30:For other uses, see
578:Elizabethan English
558:William Shakespeare
1417:Animal coat colors
1286:Primitive markings
887:Equine coat colors
753:10.1007/bf01763802
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262:tortoiseshell cats
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792:. 22 August 2018.
615:Tortoiseshell cat
605:Equine coat color
484:Percy FitzPatrick
322:equine coat color
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821:. p. 108.
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702:(9): 2963–70.
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404:A brindled cow
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72:Find sources:
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50:This article
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19:
1372:STX17 (Gray)
1332:Horse genome
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1214:Knabstrupper
1207:
1204:Varnish roan
1098:and patterns
1080:Sabino-white
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1044:Blue dun or
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674:. Retrieved
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647:. Retrieved
643:the original
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428:brindled gnu
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352:, a Russian
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334:semidominant
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284:(rosetted).
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117:October 2009
114:
104:
97:
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83:
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59:Please help
54:verification
51:
1412:Dog anatomy
1327:Color breed
1182:Tricoloured
1159:(primarily
1019:Smoky cream
996:Smoky black
552:Shakespeare
522:Pied Beauty
360:Description
337:available.
278:guinea pigs
272:Guinea pigs
168:guinea pigs
1401:Categories
1161:UK English
1042:(see also
989:Isabelline
925:Seal brown
676:2017-02-10
649:2016-01-11
621:References
282:Abyssinian
266:tabby cats
145:Great Dane
87:newspapers
1306:Tiger eye
1210:Appaloosa
1208:See also
1034:Champagne
923:included
747:: 31–62.
610:Tabby cat
503:books by
493:Jack the
450:Etymology
314:chimerism
218:Akita Inu
182:'s coat.
76:"Brindle"
1352:KIT gene
1337:Wildtype
1301:Mushroom
1236:Rabicano
1177:Skewbald
1009:Cremello
984:Palomino
979:Buckskin
908:Chestnut
761:19047145
741:Genetica
726:27449517
600:Dog coat
594:See also
586:, as in
560:'s play
520:' poem "
382:palomino
378:chestnut
330:X-linked
230:Basenjis
226:tricolor
18:Brindled
1342:Melanin
1276:Pangaré
1266:Cropout
1261:Brindle
1172:Piebald
1141:Tobiano
1014:Perlino
773:in the
768:trotter
717:5015953
583:branded
563:Macbeth
495:bulldog
430:at the
152:Brindle
101:scholar
1347:Agouti
1296:Flaxen
1146:Tovero
1134:Sabino
1046:Grullo
913:Sorrel
759:
724:
714:
410:cattle
390:grulla
384:, and
367:points
288:Horses
176:horses
164:cattle
103:
96:
89:
82:
74:
1291:Sooty
1271:Liver
1249:Other
1129:Frame
1119:Overo
1068:White
1054:Pearl
966:Cream
903:Black
757:S2CID
488:veldt
444:morph
426:Male
234:merle
180:tiger
154:is a
108:JSTOR
94:books
1367:PAX3
1362:MITF
1231:Roan
1212:and
949:Gray
944:Gray
779:site
722:PMID
548:But
516:In
438:For
415:The
248:Cats
208:Dogs
172:cats
160:dogs
156:coat
80:news
1039:Dun
920:Bay
749:doi
712:PMC
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556:In
550:not
456:PIE
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354:cab
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