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2927:, the commander of the Soviet 1st Tank Army, remarked that the enemy "did not suspect that our well-camouflaged tanks were waiting for him. As we later learned from prisoners, we had managed to move our tanks forward unnoticed". The tanks were concealed in previously prepared defensive emplacements, with only their turrets above ground level. In the air, Second World War fighters were often painted in ground colours above and sky colours below, attempting two different camouflage schemes for observers above and below. Bombers and night fighters were often black, while maritime reconnaissance planes were usually white, to avoid appearing as dark shapes against the sky. For ships, dazzle camouflage was mainly replaced with plain grey in the Second World War, though experimentation with colour schemes continued.
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patterns that specifically involve visible symmetry (such as in some butterflies) reduce survivability and increase predation. Some researchers argue that because wing-shape and color pattern are genetically linked, it is genetically costly to develop asymmetric wing colorations that would enhance the efficacy of disruptive cryptic patterning. Symmetry does not carry a high survival cost for butterflies and moths that their predators views from above on a homogeneous background, such as the bark of a tree. On the other hand, natural selection drives species with variable backgrounds and habitats to move symmetrical patterns away from the centre of the wing and body, disrupting their predators' symmetry recognition.
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about to gain the best view of an approaching predator, relying on their size and ability to defend themselves, even from lions, rather than on camouflage. A different explanation is implied by young giraffes being far more vulnerable to predation than adults. More than half of all giraffe calves die within a year, and giraffe mothers hide their newly born calves, which spend much of the time lying down in cover while their mothers are away feeding. The mothers return once a day to feed their calves with milk. Since the presence of a mother nearby does not affect survival, it is argued that these juvenile giraffes must be very well camouflaged; this is supported by coat markings being strongly
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estimate the prey's speed and direction accurately, giving the prey an improved chance of escape. Motion dazzle distorts speed perception and is most effective at high speeds; stripes can also distort perception of size (and so, perceived range to the target). As of 2011, motion dazzle had been proposed for military vehicles, but never applied. Since motion dazzle patterns would make animals more difficult to locate accurately when moving, but easier to see when stationary, there would be an evolutionary trade-off between motion dazzle and crypsis.
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1002:. Their bodies are flattened, with the sides thinning to an edge; the animals habitually press their bodies to the ground; and their sides are fringed with white scales which effectively hide and disrupt any remaining areas of shadow there may be under the edge of the body. The theory that the body shape of the horned lizards which live in open desert is adapted to minimise shadow is supported by the one species which lacks fringe scales, the
1261:, moving with extreme stealth, both slowly and quietly, watching its prey for any sign they are aware of its presence. As an example of the combination of behaviours and other methods of crypsis involved, young giraffes seek cover, lie down, and keep still, often for hours until their mothers return; their skin pattern blends with the pattern of the vegetation, while the chosen cover and lying position together hide the animals' shadows. The
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3361:, towers carrying cell telephone antennae are sometimes camouflaged as tall trees with plastic branches, in response to "resistance from the community". Since this method is costly (a figure of three times the normal cost is mentioned), alternative forms of camouflage can include using neutral colours or familiar shapes such as cylinders and flagpoles. Conspicuousness can also be reduced by siting masts near, or on, other structures.
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270:; and hawks are guided by eyesight to their prey, so much so, that on parts of the Continent persons are warned not to keep white pigeons, as being the most liable to destruction. Hence I can see no reason to doubt that natural selection might be most effective in giving the proper colour to each kind of grouse, and in keeping that colour, when once acquired, true and constant.
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surfaces, each with the simple function of providing concealment against a specific background, such as a bright water surface or the sky. The body of a shark or the fuselage of an aircraft is not gradated from light to dark to appear flat when seen from the side. The camouflage methods used are the matching of background colour and pattern, and disruption of outlines.
2203:. These particles both absorb most of the light, and are sized and shaped so as to scatter rather than reflect most of the rest. Modelling suggests that this camouflage should reduce the distance at which such a fish can be seen by a factor of 6 compared to a fish with a nominal 2% reflectance. Species with this adaptation are widely dispersed in various orders of the
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603:, utilizes specific cryptic patterning in reef ecosystems. The nudibranch syphons pigments from the consumed coral into the epidermis, adopting the same shade as the consumed coral. This allows the nudibranch to change colour (mostly between black and orange) depending on the coral system that it inhabits. However,
71:'s wings. A third approach, motion dazzle, confuses the observer with a conspicuous pattern, making the object visible but momentarily harder to locate, as well as making general aiming easier. The majority of camouflage methods aim for crypsis, often through a general resemblance to the background, high contrast
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of 2004 attempted to suit all environments, but was withdrawn after a few years of service. Terrain-specific patterns have sometimes been developed but are ineffective in other terrains. The problem of making a pattern that works at different ranges has been solved with multiscale designs, often with
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During the late 19th century camouflage was applied to
British coastal fortifications. The fortifications around Plymouth, England were painted in the late 1880s in "irregular patches of red, brown, yellow and green." From 1891 onwards British coastal artillery was permitted to be painted in suitable
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are less likely to be detected when in that microhabitat, but must spend energy to reach, and sometimes to remain in, such areas. Outside the microhabitat, the organism has a higher chance of detection. Generalized camouflage allows species to avoid predation over a wide range of habitat backgrounds,
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More recently, fashion designers have often used camouflage fabric for its striking designs, its "patterned disorder" and its symbolism. Camouflage clothing can be worn largely for its symbolic significance rather than for fashion, as when, during the late 1960s and early 1970s in the United States,
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The skins of cephalopods such as the octopus contain complex units, each consisting of a chromatophore with surrounding muscle and nerve cells. The cephalopod chromatophore has all its pigment grains in a small elastic sac, which can be stretched or allowed to relax under the control of the brain to
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Automotive manufacturers often use patterns to disguise upcoming products. This camouflage is designed to obfuscate the vehicle's visual lines, and is used along with padding, covers, and decals. The patterns' purpose is to prevent visual observation (and to a lesser degree photography), that would
882:. Another possibility is that some plants have leaves differently coloured on upper and lower surfaces or on parts such as veins and stalks to make green-camouflaged insects conspicuous, and thus benefit the plants by favouring the removal of herbivores by carnivores. These hypotheses are testable.
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use disruptive camouflage to help them approach prey, while potential prey use it to avoid detection by predators. Disruptive patterning is common in military usage, both for uniforms and for military vehicles. Disruptive patterning, however, does not always achieve crypsis on its own, as an animal
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Camouflage can be achieved by different methods, described below. Most of the methods help to hide against a background; but mimesis and motion dazzle protect without hiding. Methods may be applied on their own or in combination. Many mechanisms are visual, but some research has explored the use of
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had dark backs and light bellies. There is fossil evidence of camouflaged insects going back over 100 million years, for example lacewings larvae that stick debris all over their bodies much as their modern descendants do, hiding them from their prey. Dinosaurs appear to have been camouflaged,
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worked, using streaks of boldly contrasting colour, paradoxically making objects less visible by breaking up their outlines. While Cott was more systematic and balanced in his view than Thayer, and did include some experimental evidence on the effectiveness of camouflage, his 500-page textbook was,
426:
corrected Thayer's errors, sometimes sharply: "Thus we find Thayer straining the theory to a fantastic extreme in an endeavour to make it cover almost every type of coloration in the animal kingdom." Cott built on Thayer's discoveries, developing a comprehensive view of camouflage based on "maximum
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began to create dummy air fields to draw the attention of enemy planes to empty land. They created decoy homes and lined fake runways with flares, which were meant to help protect real towns from night raids. This strategy was not common practice and did not succeed at first, but in 1918 it caught
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encouraged the use of methods including countershading, but despite his authority on the subject, failed to persuade the
British authorities. Soldiers often wrongly viewed camouflage netting as a kind of invisibility cloak, and they had to be taught to look at camouflage practically, from an enemy
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have a high contrast pattern that could be disruptive coloration, the adults are very conspicuous when in the open. Some authors have argued that adult giraffes are cryptic, since when standing among trees and bushes they are hard to see at even a few metres' distance. However, adult giraffes move
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and buildings as well as individual soldiers and their positions. Vehicle camouflage methods begin with paint, which offers at best only limited effectiveness. Other methods for stationary land vehicles include covering with improvised materials such as blankets and vegetation, and erecting nets,
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designed to be further camouflaged by decoration with materials such as tufts of grass from the sniper's immediate environment. Such suits were used as early as 1916, the
British army having adopted "coats of motley hue and stripes of paint" for snipers. Cott takes the example of the larva of the
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have deletions of the agouti gene that prevent its expression, meaning no yellow or red color is produced. The evolution, history and widespread scope of the agouti gene shows that different organisms often rely on orthologous or even identical genes to develop a variety of camouflage strategies.
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less obtrusive and helping hunters to approach wary game animals. Patterns derived from military camouflage are frequently used in fashion clothing, exploiting their strong designs and sometimes their symbolism. Camouflage themes recur in modern art, and both figuratively and literally in science
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In fish such as the herring which live in shallower water, the mirrors must reflect a mixture of wavelengths, and the fish accordingly has crystal stacks with a range of different spacings. A further complication for fish with bodies that are rounded in cross-section is that the mirrors would be
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Where transparency cannot be achieved, it can be imitated effectively by silvering to make an animal's body highly reflective. At medium depths at sea, light comes from above, so a mirror oriented vertically makes animals such as fish invisible from the side. Most fish in the upper ocean such as
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The term countershading has a second meaning unrelated to "Thayer's Law". It is that the upper and undersides of animals such as sharks, and of some military aircraft, are different colours to match the different backgrounds when seen from above or from below. Here the camouflage consists of two
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of a wavelength apart to interfere constructively and achieve nearly 100 per cent reflection. In the deep waters that the hatchetfish lives in, only blue light with a wavelength of 500 nanometres percolates down and needs to be reflected, so mirrors 125 nanometres apart provide good camouflage.
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Most forms of camouflage are made ineffective by movement: a deer or grasshopper may be highly cryptic when motionless, but instantly seen when it moves. But one method, motion dazzle, requires rapidly moving bold patterns of contrasting stripes. Motion dazzle may degrade predators' ability to
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which can be dispersed throughout the cell, or aggregated near its centre. When the pigmented organelles are dispersed, the cell makes a patch of the animal's skin appear dark; when they are aggregated, most of the cell, and the animal's skin, appears light. In frogs, the change is controlled
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The costs associated with distractive or disruptive crypsis are more complex than the costs associated with background matching. Disruptive patterns distort the body outline, making it harder to precisely identify and locate. However, disruptive patterns result in higher predation. Disruptive
3341:(also called blinds or shooting butts). Modern hunting clothing makes use of fabrics that provide a disruptive camouflage pattern; for example, in 1986 the hunter Bill Jordan created cryptic clothing for hunters, printed with images of specific kinds of vegetation such as grass and branches.
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at a depth of 650 metres (2,130 ft); better transparency is required for invisibility in shallower water, where the light is brighter and predators can see better. For example, a cod can see prey that are 98 percent transparent in optimal lighting in shallow water. Therefore, sufficient
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Movement catches the eye of prey animals on the lookout for predators, and of predators hunting for prey. Most methods of crypsis therefore also require suitable cryptic behaviour, such as lying down and keeping still to avoid being detected, or in the case of stalking predators such as the
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moth, which fixes a screen of fragments of leaves to its specially hooked bristles, to argue that military camouflage uses the same method, pointing out that the "device is ... essentially the same as one widely practised during the Great War for the concealment, not of caterpillars, but of
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As there is a lack of evidence for camouflage in the fossil record, studying the evolution of camouflage strategies is very difficult. Furthermore, camouflage traits must be both adaptable (provide a fitness gain in a given environment) and heritable (in other words, the trait must undergo
1640:
Countershading uses graded colour to counteract the effect of self-shadowing, creating an illusion of flatness. Self-shadowing makes an animal appear darker below than on top, grading from light to dark; countershading 'paints in' tones which are darkest on top, lightest below, making the
1310:: the hoverflies to approach possible mates, and the dragonflies to approach rivals when defending territories. Motion camouflage is achieved by moving so as to stay on a straight line between the target and a fixed point in the landscape; the pursuer thus appears not to move, but only to
3405:
I very well remember at the beginning of the war being with
Picasso on the boulevard Raspail when the first camouflaged truck passed. It was at night, we had heard of camouflage but we had not seen it and Picasso amazed looked at it and then cried out, yes it is we who made it, that is
2297:. The resemblance is sufficient to make small birds take action to avoid the apparent predator. The female cuckoo then has time to lay her egg in their nest without being seen to do so. The cuckoo's egg itself mimics the eggs of the host species, reducing its chance of being rejected.
1012:, minimise their shadows when perched by closing the wings over their backs, aligning their bodies with the sun, and tilting to one side towards the sun, so that the shadow becomes a thin inconspicuous line rather than a broad patch. Similarly, some ground-nesting birds, including the
2597:(reconnaissance boats) to gather intelligence along the coast of Britain; the ships were painted entirely in bluish-green wax, with sails, ropes and crew the same colour. There is little evidence of military use of camouflage on land before 1800, but two unusual ceramics show men in
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The scheme of decoration for the great fancy dress ball given by the
Chelsea Arts Club at the Albert Hall, the other day, was based on the principles of "Dazzle", the method of "camouflage" used during the war in the painting of ships ... The total effect was brilliant and
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introduced a "drab" uniform in 1848. Hodson wrote that it would be more appropriate for the hot climate, and help make his troops "invisible in a land of dust". Later they improvised by dyeing cloth locally. Other regiments in India soon adopted the khaki uniform, and by 1896
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of Kerr, worked to persuade the
British army to use more effective camouflage methods, including countershading, but, like Kerr and Thayer in the First World War, with limited success. For example, he painted two rail-mounted coastal guns, one in conventional style, one
1852:. The planes were fitted with forward-pointing lamps automatically adjusted to match the brightness of the night sky. This enabled them to approach much closer to a target – within 3,000 yards (2,700 m) – before being seen. Counterillumination was made obsolete by
4342:
Werneck, Jane
Margaret Costa de Frontin; Torres, Lucas; Provance, David Willian; Brugnera, Ricardo; Grazia, Jocelia (3 December 2021). "First Report of Predation by a Stink Bug on a Walking-Stick Insect with Reflections on Evolutionary Mechanisms for Camouflage".
2821:, which paradoxically made ships more visible but harder to target. In Wilkinson's own words, dazzle was designed "not for low visibility, but in such a way as to break up her form and thus confuse a submarine officer as to the course on which she was heading".
310:, he classified different types such as "special protective resemblance" (where an animal looks like another object), or "general aggressive resemblance" (where a predator blends in with the background, enabling it to approach prey). His experiments showed that
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349:. He wrote that "the scattered green spots upon the under surface of the wings might have been intended for a rough sketch of the small flowerets of the plant , so close is their mutual resemblance." He also explained the coloration of sea fish such as the
1131:
Some animals actively seek to hide by decorating themselves with materials such as twigs, sand, or pieces of shell from their environment, to break up their outlines, to conceal the features of their bodies, and to match their backgrounds. For example, a
1006:, which lives in rocky areas and resembles a rock. When this species is threatened, it makes itself look as much like a rock as possible by curving its back, emphasizing its three-dimensional shape. Some species of butterflies, such as the speckled wood,
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earth, we must believe that these tints are of service to these birds and insects in preserving them from danger. Grouse, if not destroyed at some period of their lives, would increase in countless numbers; they are known to suffer largely from
571:, over a period of about 8000 years the single agouti gene developed 9 mutations that each made expression of yellow fur stronger under natural selection, and largely eliminated melanin-coding black fur coloration. On the other hand, all black
1317:
The same method can be used for military purposes, for example by missiles to minimise their risk of detection by an enemy. However, missile engineers, and animals such as bats, use the method mainly for its efficiency rather than camouflage.
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ineffective if laid flat on the skin, as they would fail to reflect horizontally. The overall mirror effect is achieved with many small reflectors, all oriented vertically. Silvering is found in other marine animals as well as fish. The
614:, which limits the geographical range and efficacy in nudibranch nutritional crypsis. Furthermore, the nudibranch colour change is not immediate, and switching between coral hosts when in search for new food or shelter can be costly.
1839:
during the Second World War. It involved projecting light on to the sides of ships to match the faint glow of the night sky, requiring awkward external platforms to support the lamps. The
Canadian concept was refined in the American
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to Darwin of 8 March 1868 mentioned such colour change: "Would you like to see the specimens of pupæ of butterflies whose colours have changed in accordance with the colour of the surrounding objects? They are very curious, and Mr.
3043:
1996:
can be made transparent, provided either they are very thin or organised as regular layers or fibrils that are small compared to the wavelength of visible light. A familiar example is the transparency of the lens of the vertebrate
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and jellyfish obey the rule, often being mainly transparent. Cott suggests this follows the more general rule that animals resemble their background: in a transparent medium like seawater, that means being transparent. The small
2087:, are so transparent as to be "almost invisible"; further, these species appear to select whether to be transparent or more conventionally mottled (disruptively patterned) according to the local background in the environment.
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2798:, artist Solomon J. Solomon and the American artist Abbott Thayer led attempts to introduce scientific principles of countershading and disruptive patterning into military camouflage, with limited success. In early 1916 the
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culture from before 500 AD, hunting birds with blowpipes which are fitted with a kind of shield near the mouth, perhaps to conceal the hunters' hands and faces. Another early source is a 15th-century French manuscript,
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similarly relies on a combination of methods: it is adapted to lie flat in the open desert, relying on stillness, its cryptic coloration, and concealment of its shadow to avoid being noticed by predators. In the ocean, the
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colours "to harmonise with the surroundings" and by 1904 it was standard practice that artillery and mountings should be painted with "large irregular patches of different colours selected to suit local conditions."
877:
plants, where there is a dappled background; leaf mottling is correlated with closed habitats. Disruptive camouflage would have a clear evolutionary advantage in plants: they would tend to escape from being eaten by
2494:. The bold stripes of the zebra have been claimed to be disruptive camouflage, background-blending and countershading. After many years in which the purpose of the coloration was disputed, an experimental study by
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1900:
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may serve as camouflage by distracting the predator's attention from recognising the prey as a whole, for example by keeping the predator from identifying the prey's outline. Experimentally, search times for
1636:
Countershading acts as a form of camouflage by 'painting out' the self-shadowing of the body or object. The result is a 'flat' appearance, instead of the 'solid' appearance of the body before countershading.
1211:
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1824:) scattered all over its underside; these create a sparkling glow that prevents the animal from appearing as a dark shape when seen from below. Counterillumination camouflage is the likely function of the
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393:, arguing that "All patterns and colors whatsoever of all animals that ever preyed or are preyed on are under certain normal circumstances obliterative" (that is, cryptic camouflage), and that "Not one '
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In 1919, the attendants of a "dazzle ball", hosted by the
Chelsea Arts Club, wore dazzle-patterned black and white clothing. The ball influenced fashion and art via postcards and magazine articles. The
1510:
could in theory make use of both dynamic colour change and counterillumination. Simple methods such as changing uniforms and repainting vehicles for winter have been in use since World War II. In 2011,
55:
is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the
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countershaded animal nearly invisible against a suitable background. Thayer observed that "Animals are painted by Nature, darkest on those parts which tend to be most lighted by the sky's light, and
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The belly of the zebra is white, and the dark stripes narrow towards the belly, so the animal is certainly countershaded, but this does not prove that the main function of the stripes is camouflage.
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Counter-illumination means producing light to match a background that is brighter than an animal's body or military vehicle; it is a form of active camouflage. It is notably used by some species of
833:
Disruptive patterns use strongly contrasting, non-repeating markings such as spots or stripes to break up the outlines of an animal or military vehicle, or to conceal telltale features, especially
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511:
Camouflage does not have a single genetic origin. However, studying the genetic components of camouflage in specific organisms illuminates the various ways that crypsis can evolve among lineages.
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2506:. However, a simulation study by Martin How and Johannes Zanker in 2014 suggests that when moving, the stripes may confuse observers, such as mammalian predators and biting insects, by two
990:
Camouflaged animals and vehicles are readily given away by their shapes and shadows. A flange helps to hide the shadow and a pale fringe breaks up and averages out any shadow that remains.
7240:
Moskát, C; Honza, M. (2002). "European Cuckoo
Cuculus canorus parasitism and host's rejection behaviour in a heavily parasitized Great Reed Warbler Acrocephalus arundinaceus population".
3175:-like range of patch sizes so they appear disruptively coloured both at close range and at a distance. The first genuinely digital camouflage pattern was the Canadian Disruptive Pattern (
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Most forms of camouflage are ineffective when the camouflaged animal or object moves, because the motion is easily seen by the observing predator, prey or enemy. However, insects such as
6782:"Long-wave sensitivity in deep-sea stomiid dragonfish with far-red bioluminescence: evidence for a dietary origin of the chlorophyll-derived retinal photosensitizer of Malacosteus niger"
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Hunters of game have long made use of camouflage in the form of materials such as animal skins, mud, foliage, and green or brown clothing to enable them to approach wary game animals.
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1156:) over the body. On these, the nymph spreads an inner layer of fine particles and an outer layer of coarser particles. The camouflage may conceal the bug from both predators and prey.
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While camouflage can increase an organism's fitness, it has genetic and energetic costs. There is a trade-off between detectability and mobility. Species camouflaged to fit a specific
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2901:. In aerial photographs, the countershaded gun was essentially invisible. The power of aerial observation and attack led every warring nation to camouflage targets of all types. The
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but is less effective. The development of generalized or specialized camouflage strategies is highly dependent on the biotic and abiotic composition of the surrounding environment.
405:' color, exists anywhere in the world where there is not every reason to believe it the very best conceivable device for the concealment of its wearer", and using paintings such as
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Some deep sea fishes have very black skin, reflecting under 0.5% of ambient light. This can prevent detection by predators or prey fish which use bioluminescence for illumination.
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The development of military camouflage was driven by the increasing range and accuracy of infantry firearms in the 19th century. In particular the replacement of the inaccurate
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sways mimetically, like the seaweeds amongst which it rests, as if rippled by wind or water currents. Swaying is seen also in some insects, like Macleay's spectre stick insect,
2195:
had a particularly black skin which reflected only 0.044% of 480 nm wavelength light. The ultra-blackness is achieved with a thin but continuous layer of particles in the
1607:
1493:, changing their coat colour (by moulting and growing new fur or feathers) from brown or grey in the summer to white in the winter; the Arctic fox is the only species in the
4277:
van't Hof, Arjen E.; Campagne, Pascal; Rigden, Daniel J.; Yung, Carl J.; Lingley, Jessica; Quail, Michael A.; Hall, Neil; Darby, Alistair C.; Saccheri, Ilik J. (June 2016).
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1276:. The behaviour may be motion crypsis, preventing detection, or motion masquerade, promoting misclassification (as something other than prey), or a combination of the two.
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have the ability to actively camouflage themselves, controlling crypsis through neural activity. For example, the genome of the common cuttlefish includes 16 copies of the
8285:, pp. 200, 258. Volume 6 of Science in World War II; Office of Scientific Research and Development. Editors: Chauncey Guy Suits and George Russell Harrison. Little, Brown.
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gene, which grants the organism remarkable control over coloration and iridescence. The reflectin gene is thought to have originated through transposition from symbiotic
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and for ground vehicles in different theatres of war. The use of radar since the mid-20th century has largely made camouflage for fixed-wing military aircraft obsolete.
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Lindgren, Johan; Sjövall, Peter; Carney, Ryan M.; et al. (February 2014). "Skin pigmentation provides evidence of convergent melanism in extinct marine reptiles".
2293:, provides examples of mimesis both in the adult and in the egg. The female lays her eggs in nests of other, smaller species of bird, one per nest. The female mimics a
687:
Some animals' colours and patterns match a particular natural background. This is an important component of camouflage in all environments. For instance, tree-dwelling
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780:
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Roosevelt attacks Thayer on page 191, arguing that neither zebra nor giraffe are "'adequately obliterated' by countershading or coloration pattern or anything else."
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464:). Thus, studying the evolution of camouflage strategies requires an understanding of the genetic components and various ecological pressures that drive crypsis.
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fish it is common to find the upper surface dark-coloured and the lower surface white, so that the animal is inconspicuous when seen either from above or below."
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began a series of monumental camouflage paintings, which helped to transform camouflage into a popular print pattern. A year later, in 1987, New York designer
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2021:-type eye of vertebrates and cephalopods must be completely opaque. Finally, some structures are visible for a reason, such as to lure prey. For example, the
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Guan, Zhe; Cai, Tiantian; Liu, Zhongmin; Dou, Yunfeng; Hu, Xuesong; Zhang, Peng; Sun, Xin; Li, Hongwei; Kuang, Yao; Zhai, Qiran; Ruan, Hao (September 2017).
4936:) going undetected by several scent-orientated predator and prey species led us to investigate chemical crypsis in this ambushing species. We trained dogs (
1963:, but it also makes them large for their muscle mass, so they cannot swim fast, making this form of camouflage a costly trade-off with mobility. Gelatinous
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to match the changing colour of the tree trunks on which they rest, from pale and mottled to almost black in polluted areas. This is taken by zoologists as
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The presence of bold skin markings does not in itself prove that an animal relies on camouflage, as that depends on its behaviour. For example, although
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to different kinds of terrain (such as woodland, snow, and desert). The design of a pattern effective in all terrains has proved elusive. The American
2610:, showing a horse pulling a cart which contains a hunter armed with a crossbow under a cover of branches, perhaps serving as a hide for shooting game.
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795:
765:
4639:
4548:
3147:
devices. After the Second World War, radar made camouflage generally less effective, though coastal boats are sometimes painted like land vehicles.
943:
10225:
540:) or gene loss (as with cephalopods with no active camouflage capabilities). This is unique as an instance of camouflage arising as an instance of
8510:
1523:
panels are heated and cooled to match either the vehicle's surroundings (crypsis), or an object such as a car (mimesis), when viewed in infrared.
79:. In the open ocean, where there is no background, the principal methods of camouflage are transparency, silvering, and countershading, while the
8064:
For Solomon, see BBC Radio 4 programme "Warpaint: the story of camouflage" by Patrick Wright, August 2002 (repeated Radio 4 Extra, 17 June 2014).
6935:"The almost invisible league: crypsis and association between minute fishes and shrimps as a possible defence against visually hunting predators"
3022:
2030:
1740:
235:
by providing individual animals with a reproductive advantage, enabling them to leave more offspring, on average, than other members of the same
7975:
7945:
5946:
912:
12044:
11297:
141:
patterns that were highly visible, but designed to confuse enemy submarines as to the target's speed, range, and heading. During and after the
10084:
8681:
8427:
2536:
888:
7491:"Polarotactic tabanids find striped patterns with brightness and/or polarization modulation least attractive: an advantage of zebra stripes"
6339:
3231:
2809:
was introduced in the early 20th century as the range of naval guns increased, with ships painted grey all over. In April 1917, when German
2355:
1967:
animals are between 50 and 90 percent transparent. A transparency of 50 percent is enough to make an animal invisible to a predator such as
13019:
9328:
7604:
Jett, Stephen C. (March 1991). "Further Information on the Geography of the Blowgun and Its Implications for Early Transoceanic Contacts".
9074:
1557:
12188:
10921:
9899:
9276:
1453:
Each chromatophore contains pigment of only one colour. In fish and frogs, colour change is mediated by a type of chromatophore known as
389:
125:
made personal concealment in battle a survival skill. In the 20th century, military camouflage developed rapidly, especially during the
12258:
10064:
10049:
3456:
as drawing "an ironic parallel between this idea of a natural paradise and the camouflage patterns on a tank". The title refers to the
2947:
1676:
Countershading is less often used for military camouflage, despite Second World War experiments that showed its effectiveness. English
1195:
9096:
8326:
3702:
3633:
These distraction markings are sometimes called dazzle markings, but have nothing to do with motion dazzle or wartime dazzle painting.
1470:
vary its opacity. By controlling chromatophores of different colours, cephalopods can rapidly change their skin patterns and colours.
12884:
9814:
8940:
3893:"Cubist Slugs. Review of DPM: Disruptive Pattern Material; An Encyclopedia of Camouflage: Nature – Military – Culture by Roy Behrens"
1466:. In fish, the change is controlled by the brain, which sends signals directly to the chromatophores, as well as producing hormones.
744:
11830:
8611:
12894:
11795:
8954:
2827:
2672:
jacket, while the Line regiments continued to wear scarlet tunics. A contemporary study in 1800 by the English artist and soldier
1179:
548:. However, other methods of horizontal gene transfer are common in the evolution of camouflage strategies in other lineages.
117:
was spurred by the increasing range and accuracy of firearms in the 19th century. In particular the replacement of the inaccurate
10100:
8205:
4128:
2883:
2665:
2637:
2253:
caterpillar mimics a twig, or a grasshopper mimics a dry leaf. It is also found in nest structures; some eusocial wasps, such as
2066:, and fish; whereas the adults of most of these are opaque and pigmented, resembling the seabed or shores where they live. Adult
1721:
205: ... seeks its prey by so changing its colour as to render it like the colour of the stones adjacent to it; it does so also
12268:
11996:
10483:
8984:
5803:
1768:
1016:, select a resting position facing the sun. Eliminating shadow was identified as a principle of military camouflage during the
7470:
Waage, J. K. (1981). "How the zebra got its stripes: biting flies as selective agents in the evolution of zebra colouration".
5742:
3761:
11105:
10645:
10292:
9650:
9635:
9613:
9595:
9577:
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4661:
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1947:, giving them almost perfect camouflage. However, transparency is difficult for bodies made of materials that have different
1762:
Two model birds painted by Thayer: painted in background colours on the left, countershaded and nearly invisible on the right
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3151:
too came to be seen as less important because of radar, and aircraft of different air forces, such as the Royal Air Force's
2734:
12273:
9697:
8463:
3493:. The word is used more figuratively in works of literature such as Thaisa Frank's collection of stories of love and loss,
8027:
5106:"Cuttlefish camouflage: The effects of substrate contrast and size in evoking uniform, mottle or disruptive body patterns"
3537:, of a 280 calibre gun illustrates the interplay of art and war, as artists like Mare contributed their skills as wartime
2245:), the camouflaged object looks like something else which is of no special interest to the observer. Mimesis is common in
13305:
12461:
10423:
5365:
4025:
Sabeti, P. C.; Schaffner, S. F.; Fry, B.; et al. (16 June 2006). "Positive Natural Selection in the Human Lineage".
2687:
army) similarly wore green jackets while other units wore more conspicuous colours. The first British Army unit to adopt
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9925:
7915:
6758:
2755:
12861:
12214:
12037:
11878:
11290:
11224:
10489:
7684:
5485:
2968:. In Australia, artists were also prominent in the Sydney Camouflage Group, formed under the chairmanship of Professor
1836:
1702:
321:. Poulton's "general protective resemblance" was at that time considered to be the main method of camouflage, as when
13394:
11550:
9478:
8391:. Australia in the War of 1939–1945. Series 4 – Civil. Vol. 5. Canberra: Australian War Memorial. p. 538ff.
6419:
3565:
2133:
is extremely flattened laterally, leaving the body just millimetres thick, and the body is so silvery as to resemble
5521:
Givnish, T. J. (1990). "Leaf Mottling: Relation to Growth Form and Leaf Phenology and Possible Role as Camouflage".
1314:
in the target's field of vision. Some insects sway while moving to appear to be blown back and forth by the breeze.
13399:
13343:
13012:
11171:
9747:
4153:
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often ironically wore military clothing during demonstrations against the American involvement in the Vietnam War.
3143:
waves. Some military textiles and vehicle camouflage paints also reflect infrared to help provide concealment from
2575:
703:
are almost all desert coloured in tones of sand, buff, ochre, and brownish grey, whether they are mammals like the
107:, whether for camouflage or for signalling. It is possible that some plants use camouflage to evade being eaten by
12011:
8122:
2303:
12293:
12006:
11873:
11585:
10914:
9974:
2864:
Austro-Hungarian ski patrol in two-part snow uniforms with improvised head camouflage on Italian front, 1915–1918
2692:
2570:
1108:
Many prey animals have conspicuous high-contrast markings which paradoxically attract the predator's gaze. These
563:
genes are orthologous genes involved in camouflage across many lineages. They produce yellow and red coloration (
8722:
7754:
2764:
to create schemes such as tree observation posts and covers for guns. Other armies soon followed them. The term
249:
When we see leaf-eating insects green, and bark-feeders mottled-grey; the alpine ptarmigan white in winter, the
12678:
12551:
10495:
9961:
9233:
6480:
422:
6653:
5757:
4878:
Miller, Ashadee Kay; Maritz, Bryan; McKay, Shannon; Glaudas, Xavier; Alexander, Graham J. (22 December 2015).
699:
are streaked brown and buff; in each case the animal's coloration matches the hues of its habitat. Similarly,
325:
wrote in 1892 that "tree-frequenting animals are often green in colour. Among vertebrates numerous species of
13404:
12724:
12323:
12278:
12030:
11283:
3556:
3438:
2704:
1984:
1944:
341:
are examples". Beddard did however briefly mention other methods, including the "alluring coloration" of the
11810:
8811:
13285:
12156:
10556:
10441:
8583:
Jukkola, E. E.; Cohen, R. (1946). "Color Stability of Olive Drab Infrared-Reflecting Camouflage Finishes".
3167:
2498:
suggested in 2012 that the pattern reduces the attractiveness of stationary models to biting flies such as
1875:
1832:
1504:
The principle of varying coloration either rapidly or with the changing seasons has military applications.
648:
uses several methods of camouflage, including disruptive coloration, lying flat, and concealment of shadow.
567:), and work in competition with other genes that produce black (melanin) and brown (eumelanin) colours. In
10760:
3183:). A pixellated appearance is not essential for this effect, though it is simpler to design and to print.
2546:
13005:
12513:
11404:
11033:
10662:
10501:
10435:
10280:
10268:
8561:
8001:
4834:
Conner, William E. (2014). "Adaptive Sounds and Silences: Acoustic Anti-Predator Strategies in Insects".
4148:
3257:
3218:
3159:
3121:
1845:
1828:
of many marine organisms, though light is also produced to attract or to detect prey and for signalling.
274:
20:
2164:, including squid, octopus and cuttlefish, have multilayer mirrors made of protein rather than guanine.
1440:
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11502:
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10907:
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9779:
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7756:
Twelve Years of a Soldier's Life in India, being extracts from the letters of the late Major WSR Hodson
7336:
7215:
5853:
5628:
5201:
4677:
3965:
3152:
2915:
2684:
2232:
1519:
infrared camouflage technology. It uses about 1,000 hexagonal panels to cover the sides of a tank. The
1413:
627:
6602:
Kiltie, Richard A. (January 1998). "Countershading: Universally deceptive or deceptively universal?".
6553:
3448:
have used camouflage to reflect on war. His 1973 screenprint of a tank camouflaged in a leaf pattern,
2017:
or equivalent light-absorbing structures of eyes – they must absorb light to be able to function. The
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12371:
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12106:
12101:
11780:
11492:
11100:
10750:
10638:
10364:
10358:
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8314:
white has proved to be the most efficient concealing coloration for aircraft on anti-submarine patrol
8167:
7979:
7953:
7646:
3013:
2997:
2814:
11275:
9799:
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5507:
5478:
Defensive coloration in plants: a review of current ideas about anti-herbivore coloration strategies
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13379:
12937:
12749:
12744:
12714:
12518:
11981:
11863:
10888:
9950:
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9219:
9029:"The Great Dazzle Ball at the Albert Hall: The Shower of Bomb Balloons and Some Typical Costumes".
5104:
Barbosa, A.; Mathger, L. M.; Buresch, K. C.; Kelly, J.; Chubb, C.; Chiao, C.; Hanlon R. T. (2008).
3298:
1244:
731:
uniforms are a muddy or dusty colour, originally chosen for service in South Asia. Many moths show
541:
13119:
12022:
11653:
8650:
6735:
4588:"First observation of the nudibranch Tenellia feeding on the scleractinian coral Pavona decussata"
4432:"The Evolutionary History of Nebraska Deer Mice: Local Adaptation in the Face of Strong Gene Flow"
2930:
As in the First World War, artists were pressed into service; for example, the surrealist painter
2633:
480:
period show that some marine reptiles were countershaded. The skins, pigmented with dark-coloured
12911:
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12234:
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409:(1907) to reinforce his argument. Thayer was roundly mocked for these views by critics including
380:
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provided evidence that grey uniforms were less visible than green ones at a range of 150 yards.
451:) were shown to survive according to how well their egg contrast matched the local environment.
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10808:
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9885:
9879:
9769:
9555:
DPM – Disruptive Pattern Material: An Encyclopaedia of Camouflage: Nature, Military and Culture
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7269:
6479:. Newfoundland Labrador Department of Environment and Conservation. p. 7. Archived from
4364:
3757:
3652:
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1990, developed from a 1938 pattern, a non-digital pattern which works at different distances
3144:
2961:
2700:
2439:
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2138:
1920:
1813:
1802:
816:
747:, as well as demonstrating that it changes where necessary to resemble the local background.
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283:
72:
45:
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Camouflage is occasionally used to make built structures less conspicuous: for example, in
3260:
3103:
3068:
2849:
2423:
2294:
2216:
1798:
1788:
1775:
1109:
1103:
866:
84:
9844:
4169:"Pharaoh Cuttlefish, Sepia pharaonis, Genome Reveals Unique Reflectin Camouflage Gene Set"
2185:
is one of several deep-sea fishes camouflaged against very dark water with a black dermis.
1457:
that contain dark pigment. A melanophore is star-shaped; it contains many small pigmented
821:
592:
There are many examples of the tradeoffs between specific and general cryptic patterning.
8:
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12906:
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9215:
8727:. U.S. Army Natick Soldier Research, Development and Engineering Center. 2009. p. 27
8627:
5743:"Anatomy of disguise: camouflaging structures in nymphs of Some Reduviidae (Heteroptera)"
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3445:
3148:
3131:
2875:
2760:
2729:
2532:
2515:
2182:
1925:
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advised soldiers to copy animals' methods, using their instincts for wartime camouflage.
1650:
1535:
Rock ptarmigan, changing colour in springtime. The male is still mostly in winter plumage
1434:
taken a few minutes apart, showing its ability to match its coloration to the environment
1424:
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1218:
1145:
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322:
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146:
114:
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5388:
4979:
4603:
4504:
4392:
4296:
4237:
4089:
4038:
3985:
2954:. Hugh Cott was chief instructor; the artist camouflage officers, who called themselves
13384:
13291:
13191:
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12957:
12916:
12901:
12830:
12779:
12774:
12583:
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11991:
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7310:
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6806:
6781:
6401:
6376:
Cloney, R. A.; Florey, E. (1968). "Ultrastructure of Cephalopod Chromatophore Organs".
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3202:
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in naval camouflage Measure 32, Design 11a, one of many dazzle schemes used on warships
3049:
2989:
2969:
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2255:
2130:
1689:
1349:
568:
536:, with divergence occurred through subsequent gene duplication (such as in the case of
485:
410:
342:
206:
193:
68:
13325:
How Long Is the Coast of Britain? Statistical Self-Similarity and Fractional Dimension
5454:
4513:
4488:
4279:"The industrial melanism mutation in British peppered moths is a transposable element"
2950:
during the 1941–1942 war in the Western Desert, including the successful deception of
654:
427:
disruptive contrast", countershading and hundreds of examples. The book explained how
13318:
13218:
13213:
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5002:
cryptic coloration in British field uniforms was not fully adopted until the Boer War
4993:
4923:
4905:
4857:
4847:
4816:
4763:
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4703:
4657:
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Pfeifer, Susanne P; Laurent, Stefan; Sousa, Vitor C.; et al. (15 January 2018).
4412:
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4308:
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became a lecturer at the newly founded Camouflage Development and Training Centre at
2910:
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2212:
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1856:, and neither diffused lighting camouflage nor Yehudi lights entered active service.
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1506:
1409:
1383:
1307:
1285:
1149:
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529:
301:
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183:(384–322 BC) commented on the colour-changing abilities, both for camouflage and for
138:
104:
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8992:
8698:
6951:
6934:
6405:
6136:
5139:
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2278:. This tactic has occasionally been used in warfare, for example with heavily armed
841:. Disruptive patterns may use more than one method to defeat visual systems such as
739:
which has coloration that blends in with tree bark. The coloration of these insects
585:
528:
bacteria, which provide bioluminescence to its hosts. While not all cephalopods use
13389:
13169:
12673:
12536:
12528:
12446:
12328:
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12146:
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9601:
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7863:
Details of Equipment of Her Majesty's Army Part 2 Section XI B – Garrison Artillery
7662:
The People of the Caribbean: An Encyclopedia of Archaeology and Traditional Culture
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7554:
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7434:
7366:
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6212:
6171:
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6114:
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6016:
Srinivasan, M. V.; Davey, M. (1995). "Strategies for active camouflage of motion".
5981:
5970:"The swaying behavior of Extatosoma tiaratum: motion camouflage in a stick insect?"
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used Warhol's camouflage prints as the basis for his Autumn Winter 1987 collection.
3460:
3000:
with Baron Nicholas Cerkasoff, and went on to create camouflage for the Air Force.
2977:
2889:
2795:
2615:
2611:
1975:
1948:
1909:
raise the average brightness of the plane from a dark shape to the same as the sky.
1594:
1431:
1397:
1267:
1249:
1186:
1169:
1017:
736:
724:
660:
644:
402:
254:
142:
33:
28:
5307:
1446:
Fish and frog melanophore cells change colour by moving pigment-containing bodies.
13092:
12809:
12668:
12638:
12633:
12623:
12556:
12541:
12421:
12401:
12283:
12151:
12057:
11948:
11858:
11800:
11384:
11310:
11028:
10964:
10954:
10878:
10773:
10730:
10605:
10163:
10122:
10069:
9969:
9784:
9403:
8988:
7300:
7183:"A parasite in wolf's clothing: hawk mimicry reduces mobbing of cuckoos by hosts"
6932:
6649:
6505:
6217:
5122:
5105:
4638:; Allen, William L.; Sherratt, Thomas N.; Speed, Michael P. (20 September 2018).
4224:
3706:
3505:
3476:
3373:
3127:
3107:
2943:
2924:
2920:
2919:
for military deception, including the use of camouflage. For example, during the
2806:
2787:
2751:
2713:
2668:
and the 60th Rifle Regiment, were the first to adopt camouflage in the form of a
2657:
2507:
2137:. The mirrors consist of microscopic structures similar to those used to provide
2101:
1931:
1825:
1817:
1728:
1490:
1401:
1387:
1332:
1141:
1126:
1048:"Shape, shine, shadow" make these 'camouflaged' military vehicles easily visible.
1008:
970:
850:
or a military target may be given away by factors like shape, shine, and shadow.
433:
338:
134:
126:
80:
10979:
8474:
4843:
3957:
2766:
2259:, build a nest envelope in patterns that mimic the leaves surrounding the nest.
13280:
13275:
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12952:
12789:
12613:
12566:
12496:
12491:
12386:
12253:
12126:
11933:
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11903:
11706:
11671:
11610:
11487:
11442:
11334:
11151:
11067:
11055:
10959:
10833:
10803:
10798:
10745:
10735:
10471:
10382:
10116:
9945:
9940:
9824:
9727:
9243:
9123:
8980:
8858:"Lost in the Wilderness, the military's misadventures in pixellated camouflage"
8507:
Chapter 20: Vehicle Camouflage And Nuclear, Biological, And Chemical Operations
8295:
7558:
3390:
3382:
3126:
Camouflage has been used to protect military equipment such as vehicles, guns,
2981:
2935:
2931:
2898:
2688:
2290:
2208:
2134:
1709:
1627:
1486:
1152:
fan' to decorate its body with sand or dust. There are two layers of bristles (
949:
842:
500:
495:
384:
224:
171:
76:
8669:
7011:
6986:
5433:
Lev-Yadun, Simcha (2003). "Why do some thorny plants resemble green zebras?".
5408:
5396:
4612:
4587:
4487:
Eizirik, Eduardo; Yuhki, Naoya; Johnson, Warren E.; et al. (March 2003).
4246:
4219:
4185:
4168:
1290:
532:, ancient cephalopods may have inherited the gene horizontally from symbiotic
13368:
13270:
13265:
12814:
11790:
11764:
11721:
11711:
11666:
11633:
11525:
11359:
11314:
11141:
11060:
11038:
11013:
10984:
10974:
10936:
10823:
10705:
10685:
10680:
10562:
10286:
10181:
10175:
10000:
9829:
9415:
9348:
7833:
7340:
7271:
7020:
6648:
5849:
5205:
5166:
4997:
4909:
4861:
4782:
4749:
4721:
4707:
4635:
4586:
Wong, Kwan Ting; Ng, Tsz Yan; Tsang, Ryan Ho Leung; Ang, Put (24 June 2017).
4544:
4455:
4348:
4312:
4278:
4204:
3486:
3398:
3338:
3319:
3179:), issued to the army in 2002, soon followed by the American Marine pattern (
3136:
2590:
2419:
2324:
2309:
2286:
2271:
2250:
1906:
1841:
1520:
1405:
1159:
Similar principles can be applied for military purposes, for instance when a
1077:
995:
906:
Russian T-90 battle tank painted in bold disruptive pattern of sand and green
564:
549:
364:
267:
13129:
10710:
7199:
7182:
6092:
5986:
5969:
5200:
4698:
4681:
4447:
4046:
3963:
3699:
3520:
2743:
2574:
that Mediterranean pirate ships could be painted blue-gray for concealment.
1793:
727:. Military uniforms, too, generally resemble their backgrounds; for example
130:
13181:
13104:
13087:
12942:
12799:
12784:
12441:
12411:
12356:
12239:
12204:
12081:
11580:
11072:
11018:
10868:
10863:
10858:
10793:
10610:
10370:
10262:
10193:
10146:
10140:
10128:
9955:
9229:
7566:
7516:
7456:
7438:
7380:
7361:
7319:
7028:
6985:
Davis, Alexander L.; Thomas, Kate N.; Goetz, Freya E.; et al. (2020).
6815:
6797:
6623:
6309:
6291:
6236:
6185:
6167:
6128:
6037:
5891:
5873:
5715:
5697:
5666:
5648:
5462:
5243:
5225:
5131:
4927:
4900:
4820:
4802:
4767:
4741:
4522:
4473:
4431:
4416:
4320:
4255:
4105:
4054:
4011:
3602:, who bred them, would, I am sure, be delighted to bring them to show you."
3490:
3469:
3465:
3453:
3358:
3330:
3323:
3087:
aircraft hangar built to resemble a street of village houses, Belgium, 1944
2902:
2893:
2661:
2558:
2329:
2072:
2026:
1988:
use partial transparency for camouflage in the dim light of the rainforest.
1666:
1408:
cells to resemble their current background, or, as in most chameleons, for
1202:
1173:
caterpillar-tractors, battery positions, observation posts and so forth."
1164:
1137:
918:
759:
Lion in Kruger National Park, South Africa, blending in with the tall grass
600:
545:
258:
19:
This article is about a form of protective coloration. For other uses, see
7144:
6473:
The Status of Arctic Hare (Lepus arcticus bangsii) in Insular Newfoundland
6423:
6397:
6072:
2113:
The herring's reflectors are nearly vertical for camouflage from the side.
13299:
13257:
13176:
13072:
13067:
12986:
12947:
12091:
11638:
11600:
11575:
11565:
11530:
11477:
11457:
11249:
11131:
11050:
10944:
10853:
10838:
10783:
10778:
10768:
10322:
10274:
10134:
9990:
9912:
9535:
9502:
9484:
9466:
8106:
Sumrall, R. F. (February 1973). "Ship Camouflage (WWII): Deceptive Art".
6933:
Carvalho, Lucélia Nobre; Zuanon, Jansen; Sazima, Ivan (April–June 2006).
6274:
Stuart-Fox, Devi; Moussalli, Adnan; Whiting, Martin J. (23 August 2008).
5682:"Concealed by conspicuousness: distractive prey markings and backgrounds"
5282:
3539:
3501:
3163:
2993:
2965:
2956:
2709:
2669:
2653:
2641:
2586:
2551:
2362:
2161:
2105:, is a typical silvered fish of medium depths, camouflaged by reflection.
1979:
1821:
1512:
1474:
1454:
870:
838:
712:
553:
439:
Experimental evidence that camouflage helps prey avoid being detected by
417:
398:
60:
9489:
Camoupedia: A Compendium of Research on Art, Architecture and Camouflage
8834:
8596:
6201:"Echolocating Bats Use a Nearly Time-Optimal Strategy to Intercept Prey"
5300:
Field Manual Headquarters No. 20-3 − Camouflage, Concealment, and Decoys
4304:
4097:
1894:
showing 4 (of about 60) diffused lighting fittings, 2 lifted, 2 deployed
637:
556:
both have camouflage-related genes that stem from transposition events.
165:
13246:
13241:
13062:
12839:
12804:
12381:
12346:
11938:
11883:
11853:
11759:
11676:
11620:
11497:
11447:
11239:
10848:
10843:
10828:
10813:
10700:
10535:
10352:
10219:
8130:
7625:
7507:
7490:
6389:
5550:
4220:"Origin of the Reflectin Gene and Hierarchical Assembly of Its Protein"
3776:
3611:
Cott explains Beddard's observation as a coincident disruptive pattern.
3599:
3199:
2996:
took a certificate course in military and industrial camouflage at the
2973:
2503:
2405:
2200:
2067:
2063:
2059:
2055:
2022:
2002:
1579:
1576:
1548:
1478:
986:
965:
874:
716:
708:
515:
477:
317:
were camouflaged to match the backgrounds on which they were reared as
250:
188:
88:
16:
Concealment in plain sight by any means, e.g. colour, pattern and shape
9570:
Hide and Seek: Camouflage, Photography and the Media of Reconnaissance
6780:
Douglas, R. H.; Mullineaux, C. W.; Partridge, J. C. (September 2000).
6199:
Ghose, K.; Horiuchi, T. K.; Krishnaprasad, P. S.; Moss, C. F. (2006).
5637:
Philosophical Transactions of the Royal Society B: Biological Sciences
4871:
4195:
4167:
Song, Weiwei; Li, Ronghua; Zhao, Yun; et al. (15 February 2021).
3993:
3135:
screens and soft covers which may suitably reflect, scatter or absorb
2178:
1972:
transparency for camouflage is more easily achieved in deeper waters.
133:
designed camouflage schemes and observation posts disguised as trees.
13348:
13251:
12997:
12709:
12663:
12391:
11835:
11805:
11605:
11560:
11535:
11472:
11462:
11437:
11429:
11374:
11305:
11254:
11087:
11043:
11023:
10695:
10346:
10187:
7489:
Egri, Ádám; Blahó, Miklós; Kriska, György; et al. (March 2012).
5765:
4682:"Revealed by Conspicuousness: Distractive Markings Reduce Camouflage"
3282:
3084:
3033:
3029:
2499:
2263:
2191:
1952:
1868:
1685:
1680:
1677:
1458:
1393:
1303:
1136:
larva builds a decorated case and lives almost entirely inside it; a
1133:
934:
879:
826:
672:
609:
594:
519:
481:
334:
232:
180:
153:
108:
96:
44:
A soldier applying camouflage face paint; both helmet and jacket are
10899:
8936:
Telecommunication Mast Management Guidelines for the City of Tshwane
6119:
6094:
5542:
5340:
5327:(1911). "Revealing and concealing coloration in birds and mammals".
4988:
4961:
4547:; Allen, William L.; Sherratt, Thomas N.; Speed, Michael P. (2018).
2109:
2095:
1645:". Accordingly, the principle of countershading is sometimes called
472:
Camouflage is a soft-tissue feature that is rarely preserved in the
360:
13077:
13042:
12764:
12693:
12224:
12052:
11731:
11643:
11590:
11545:
11121:
10429:
9995:
9051:. The Museum at FIT. 9 September – 16 December 2006. Archived from
7734:"Khaki Uniform 1848–49: First Introduction by Lumsden and Hodson".
7418:
3762:"Alfred Russel Wallace Letters and Reminiscences By James Marchant"
3302:
3099:
2906:
2696:
2683:, rifle units such as the 1st United States Sharp Shooters (in the
2495:
2267:
2051:
2038:. Examples of transparent marine animals include a wide variety of
2010:
1964:
1960:
1956:
1563:
Arctic hares in the low arctic change from brown to white in winter
1299:
1153:
1114:
859:
789:
nests in open sand with only its camouflaged plumage to protect it.
700:
692:
688:
489:
448:
440:
383:
formulated what is sometimes called Thayer's Law, the principle of
350:
7877:
Regulations for the equipment of the army. Part 2. section XII (a)
5722:
5680:
Dimitrova, M.; Stobbe, N.; Schaefer, H. M.; Merilaita, S. (2009).
4486:
2813:
were sinking many British ships with torpedoes, the marine artist
1959:
is acellular and highly transparent. This conveniently makes them
825:
Illustration of the principle of "maximum disruptive contrast" by
13203:
13198:
13186:
13164:
13149:
13057:
12981:
12971:
12754:
12561:
12431:
12426:
12053:
12001:
11661:
11306:
11183:
11156:
10568:
10540:
10465:
9834:
9819:
9097:"Ian Hamilton Finlay: Arcadia (collaboration with George Oliver)"
8613:
Stealth Warplanes: Deception, Evasion, and Concealment in the Air
7345:"Motion dazzle and camouflage as distinct anti-predator defenses"
5633:"Defining disruptive coloration and distinguishing its functions"
4838:. Animal Signals and Communication. Vol. 1. pp. 65–79.
3572:
3172:
2810:
2602:
2490:
An animal that is commonly thought to be dazzle-patterned is the
2228:
2142:
2123:
2119:
2043:
2035:
1831:
Counterillumination has rarely been used for military purposes. "
1654:
1632:
1613:
1516:
1494:
1463:
1311:
854:
846:
802:
696:
679:(sound) detection. Methods may also apply to military equipment.
676:
444:
394:
372:
354:
236:
220:
202:
100:
64:
56:
8152:"Obituary: Mr Norman Wilkinson, Inventor of 'dazzle' painting".
7820:
Barrass, S (2018). "British Military Camouflage Prior to 1914".
6093:
Mizutani, A. K.; Chahl, J. S.; Srinivasan, M. V. (5 June 2003).
5679:
4944:) to test whether a canid and a herpestid predator could detect
4680:; Marshall, Kate L. A.; Troscianko, Jolyon; et al. (2013).
4489:"Molecular Genetics and Evolution of Melanism in the Cat Family"
3365:
subsequently enable reproduction of the vehicle's form factors.
3071:
underside 'azure' paint scheme, meant to hide it against the sky
2474:
10447:
10419:
10388:
10376:
10298:
9049:
John Galliano for Christian Dior, silk camouflage evening dress
8356:
4676:
4276:
3621:
3620:
Before 1860, unpolluted tree trunks were often covered in pale
3524:
3457:
3394:
3195:
3180:
3176:
3171:
a pixellated appearance and designed digitally, that provide a
2992:
and Garden Island Dockyard. In the United States, artists like
2880:
World War II ship camouflage measures of the United States Navy
2739:
2649:
2401:
2279:
2275:
2196:
2018:
2014:
2006:
1993:
1670:
1658:
1601:, changes colour mainly in relation to mood and for signalling.
1498:
1497:
to do so. However, Arctic hares which live in the far north of
1160:
999:
998:
of North America, have evolved elaborate measures to eliminate
704:
473:
330:
326:
318:
118:
10623:
6968:
6966:
6964:
6962:
6901:
6899:
6897:
6895:
6893:
6891:
6889:
5170:"Camouflage and perceptual organization in the animal kingdom"
5103:
4787:"Overcoming the Detectability Costs of Symmetrical Coloration"
1955:
have gelatinous bodies, composed mainly of water; their thick
1404:
actively change their skin patterns and colours using special
40:
13236:
11209:
9675:
9663:
7488:
4780:
4341:
3872:
3314:
3140:
2775:
2491:
2479:
2386:
2047:
2039:
1853:
1809:
1482:
1258:
728:
720:
314:
290:
122:
92:
4880:"An ambusher's arsenal: chemical crypsis in the puff adder (
4024:
1080:
is draped away from a military vehicle to reduce its shadow.
36:
can change its pattern and colours to match its environment.
13114:
13052:
13047:
7946:"Art of the First World War: André Mare and Leon Underwood"
6959:
6886:
6759:"Oceanic Bioluminescence: an Overview of General Functions"
5796:"Natural Bling: 6 Amazing Animals That Decorate Themselves"
2614:
are said to have used plant materials as camouflage in the
2598:
2378:
2246:
2013:. Other structures cannot be made transparent, notably the
1033:
745:
evidence that camouflage is influenced by natural selection
286:
262:
8368:
8344:
8178:
7926:
7573:
6779:
6680:
6273:
6015:
5592:
5590:
5426:
5363:
4075:
1551:, 1940, with white camouflage overalls over their uniforms
607:
can only feed and lay eggs on the branches of host-coral,
11005:
10453:
9012:
9010:
8914:
8773:"Army drops universal camouflage after spending billions"
8748:
Camouflage uniforms: international combat dress 1940–2010
7048:(4th ed.). John Wiley, Blackwell. pp. 512–513.
6378:
Zeitschrift für Zellforschung und Mikroskopische Anatomie
6255:
5818:
5758:
10.1206/0003-0082(2006)3542[1:AODCSI]2.0.CO;2
5476:
Lev-Yadun, Simcha (2006). Teixeira da Silva, J.A. (ed.).
4429:
3945:
3386:
3241:
2716:
six years later it was used throughout the British Army.
1998:
1968:
1940:
1616:
infrared camouflage lets an armoured vehicle mimic a car.
1117:
increased when artificial prey had distractive markings.
937:
and five chicks exhibit exceptional disruptive camouflage
572:
8046:
7339:; Searle, W. T. L.; Seymour, J. E.; Marshall, K. L. A.;
5306:. Department of the Army. 30 August 1999. Archived from
5259:
Transportation and Cargo Security: Threats and Solutions
4877:
4634:
4543:
4129:"Oldest insect camouflage behaviour revealed by fossils"
2585:) says that "Venetian blue" (sea green) was used in the
2557:
Ship camouflage was occasionally used in ancient times.
2219:
of ultra-blackness camouflage independently many times.
1234:
Soviet tanks under netting dressed with vegetation, 1938
1064:'s body is flattened and fringed to minimise its shadow.
219:
Camouflage has been a topic of interest and research in
9522:
Camouflage and Art: Design for Deception in World War 2
8582:
7918:(December 1949). "Souvenir de Camouflage (1914–1918)".
7331:
7329:
7162:
6984:
6716:
6580:
6375:
6011:
6009:
5994:
5686:
Proceedings of the Royal Society B: Biological Sciences
5587:
5043:
4888:
Proceedings of the Royal Society B: Biological Sciences
4791:
Proceedings of the Royal Society B: Biological Sciences
4730:
Proceedings of the Royal Society B: Biological Sciences
4726:"The Predation Costs of Symmetrical Cryptic Coloration"
3933:
3909:
3860:
1501:, where summer is very short, remain white year-round.
1222:, masked hunter bug nymph, camouflaged with sand grains
1140:
covers its back with seaweed, sponges, and stones. The
1092:
A caterpillar's fringe of bristles conceals its shadow.
443:
was first provided in 2016, when ground-nesting birds (
9007:
8076:
Camouflage and Art Design for Deception in World War 2
7585:
7399:
7099:
7062:
6692:
5968:
Bian, Xue; Elgar, Mark A.; Peters, Richard A. (2016).
5903:
5901:
3970:"Camouflage predicts survival in ground-nesting birds"
3824:
3728:
3726:
3016:, painted white to minimise visibility against the sky
2518:, where the perceived motion is in a wrong direction.
9377:
Survival of the Beautiful: Art, Science and Evolution
9313:
Camouflage Cultures: Beyond the Art of Disappearance.
9278:
Concealing-Coloration in the Animal Kingdom Macmillan
8877:
7693:
6916:
6914:
6829:
6827:
6825:
6704:
6568:
6243:
5208:; Windsor, A. M. M.; Walker, H. J. (7 October 2006).
5146:
5062:
5060:
5058:
3921:
3812:
3738:
3377:
The "dazzle ball" held by the Chelsea Arts Club, 1919
2984:
were among the members of the group, which worked at
2482:'s bold pattern may induce motion dazzle in observers
1582:
and white-overalled crew and infantry in Russia, 1943
1352:
dragonflies use motion camouflage to approach rivals.
1294:
Comparison of motion camouflage and classical pursuit
869:
was little studied until the late 20th century. Leaf
8810:. Ex-Military Land Rover Association. Archived from
7326:
6095:"Insect behaviour: Motion camouflage in dragonflies"
6006:
5621:
5573:. University of California Press. pp. 117–118.
5366:"On the origin, evolution and phylogeny of giraffes
5019:
5007:
3848:
3800:
3788:
3624:; polluted trunks were bare, and often nearly black.
2754:, the French army formed a camouflage corps, led by
2438:
Cuckoo eggs mimicking smaller eggs, in this case of
2349:, superbly mimics a leaf with a bright orange border
1336:
hoverflies use motion camouflage to approach females
695:
of the forest floor are brown and speckled; reedbed
436:
narrative which illustrated theories with examples.
345:
and the possibility of a different mechanism in the
137:, merchant ships and troop carriers were painted in
10060:
List of countries that prohibit camouflage clothing
9037:
7736:
Journal of the Society for Army Historical Research
7387:
6728:
6668:
5898:
5830:
5673:
5608:
5031:
3723:
2712:uniform was used everywhere outside Europe; by the
2656:made personal concealment in battle essential. Two
974:
have pale markings, possibly disruptive camouflage.
873:with white spots may serve as camouflage in forest
387:. However, he overstated the case in the 1909 book
9374:
9295:Camouflage Australia: Art, Nature, Science and War
8078:. London, England: Unicorn Press. pp. 12–13.
7914:
7774:. Vol. 4. Osprey Publishing. pp. 24–33.
7753:Hodson, W. S. R. (1859). Hodson, George H. (ed.).
7270:Scott-Samuel, N. E.; Baddeley, R.; Palmer, C. E.;
6978:
6911:
6835:"Diffused Lighting and its use in the Chaleur Bay"
6822:
6630:
6357:
5858:"The evolutionary ecology of decorating behaviour"
5329:Bulletin of the American Museum of Natural History
5055:
4962:"How a naturalist found safe colours for soldiers"
4781:Wainwright, J. Benito; Scott-Samuel, Nicholas E.;
4391:Voisey, Joanne; Van Daal, Angela (February 2002).
3836:
2852:camouflaged against observation from the air, 1917
2514:, where the perceived motion is inverted, and the
952:: a disruptively camouflaged invertebrate predator
476:record, but rare fossilised skin samples from the
7606:Annals of the Association of American Geographers
7427:Philosophical Transactions of the Royal Society B
6786:Philosophical Transactions of the Royal Society B
5377:Transactions of the Royal Society of South Africa
3711:
13366:
9645:. First Discovery series. Moonlight Publishing.
9033:. No. 154. 22 March 1919. pp. 414–415.
7265:
7263:
7120:Q-Ships versus U-Boats: America's Secret Project
6738:. Smithsonian National Museum of Natural History
6422:. MarineBio Conservation Society. Archived from
5627:
4720:
4166:
1252:sways like seaweeds to reinforce its camouflage.
145:, a variety of camouflage schemes were used for
105:actively changing their skin pattern and colours
9471:False Colors: Art, Design and Modern Camouflage
8979:
8123:"Schnellboot: An Illustrated Technical History"
7674:
7423:"Contrasting coloration in terrestrial mammals"
7239:
7180:
5967:
5570:Introduction to horned lizards of North America
5359:
5357:
5355:
5353:
5351:
2738:Iron observation post camouflaged as a tree by
2274:mimics a particular kind of flower, such as an
1943:animals that float near the surface are highly
152:Non-military use of camouflage includes making
9408:The Camouflage Story (from Aintree to Alamein)
8406:. University of Minnesota Press. p. 183.
8404:John Vassos: Industrial Design for Modern Life
7041:
4585:
4390:
3682:. IX, 622a: 2–10. Cited in Borrelli, Luciana;
3389:from the time of the First World War onwards.
3162:have been developed to suit the need to match
2426:, giving female time to lay eggs parasitically
1416:does use active colour change for camouflage.
1372:Preying mantises exhibiting motion camouflage.
13013:
12855:
12038:
11291:
10915:
10639:
9691:
9664:Ohio State University: The Camouflage Project
8889:
8835:"Dual Texture – U.S. Army digital camouflage"
8471:Chapter 5: Cover, Concealment, and Camouflage
8199:"Maskirovka: The Soviet System of Camouflage"
7851:. The Royal Engineers Institute. p. 280.
7849:Permanent Fortification for English Engineers
7638:
7260:
6524:
5848:
5562:
5560:
4393:"Agouti: from Mouse to Man, from Skin to Fat"
4217:
3968:; Spottiswoode, Claire N. (29 January 2016).
3688:A catalogue of body patterning in Cephalopoda
2537:List of military clothing camouflage patterns
1377:
1032:Three countershaded and cryptically coloured
9330:Dazzled and Deceived: Mimicry and Camouflage
7532:"Motion camouflage induced by zebra stripes"
7530:How, Martin J.; Zanker, Johannes M. (2014).
7276:"Dazzle Camouflage Affects Speed Perception"
6862:
6860:
6443:
6441:
6276:"Predator-specific camouflage in chameleons"
5348:
4866:adaptive silence, acoustic crypsis, stealth,
3964:Troscianko, Jolyon; Wilson-Aggarwal, Jared;
3256:Main (4-colour woodland) variant of Chinese
2365:hides from predators by mimicking a dry leaf
1844:project, and trialled in aircraft including
894:Leopard: a disruptively camouflaged predator
175:can change colour (and shape) for camouflage
9900:Concealing-Coloration in the Animal Kingdom
9089:
8101:
8099:
8097:
8095:
6987:"Ultra-black Camouflage in Deep-Sea Fishes"
6463:
6149:
6143:
5514:
3674:
3672:
3475:which recurs in Hamilton Finlay's work. In
3368:
1951:from seawater. Some marine animals such as
921:'s bold markings are powerfully disruptive.
493:as a 120 million year old fossil of a
390:Concealing-Coloration in the Animal Kingdom
13020:
13006:
12862:
12848:
12259:Latitudinal gradients in species diversity
12045:
12031:
11298:
11284:
10922:
10908:
10646:
10632:
10065:Middle East Command Camouflage Directorate
10050:Camouflage clothing in Trinidad and Tobago
9698:
9684:
9626:Kalman, Bobbie; Crossingham, John (2001).
9588:Animal Camouflage: Mechanisms and Function
9507:Ship Shape: A Dazzle Camouflage Sourcebook
9369:
8932:
8609:
7994:
6652:; Dobkin, David S.; Wheye, Darryl (1988).
6052:
5566:
5557:
5480:. Global Science Books. pp. 292–299.
5210:"Disruptive contrast in animal camouflage"
5194:
4953:
4827:
4724:; Hiby, Elly; Lloyd, Emily (22 May 2006).
3878:
2948:Middle East Command Camouflage Directorate
2550:Roman ships, depicted on a 3rd-century AD
1684:observer's viewpoint. At the same time in
1657:and grasshoppers; marine animals, such as
304:, especially camouflage. In his 1890 book
12885:Coloration evidence for natural selection
10281:Six-Color Desert Pattern (Chocolate Chip)
9420:Ships and Seamanship in the Ancient World
8941:City of Tshwane Metropolitan Municipality
8770:
8724:Photosimulation Camouflage Detection Test
8294:
8166:
8108:United States Naval Institute Proceedings
7529:
7506:
7446:
7370:
7360:
7309:
7299:
7233:
7198:
7010:
6950:
6857:
6805:
6597:
6595:
6438:
6299:
6226:
6216:
6175:
6118:
5985:
5881:
5705:
5656:
5475:
5432:
5323:
5317:
5233:
5164:
5121:
5081:Collins Wild Guide: Butterflies and Moths
4987:
4917:
4899:
4836:Insect Hearing and Acoustic Communication
4810:
4757:
4697:
4611:
4512:
4463:
4245:
4194:
4184:
4001:
2141:: stacks of between 5 and 10 crystals of
1820:. The latter has light-producing organs (
12869:
12157:Predator–prey (Lotka–Volterra) equations
11796:Tritrophic interactions in plant defense
9121:
8955:"The secrets behind all that camouflage"
8895:
8739:
8691:
8509:. Department of the Army. pp. 1–9.
8473:. Department of the Army. Archived from
8428:"Concealment, Camouflage, and Deception"
8092:
7976:"Art of the First World War: André Mare"
7794:
7769:
7715:. Weider History Group. 20 February 2008
7659:
7213:
7142:
6866:
6506:"Tanks test infrared invisibility cloak"
6086:
5961:
5740:
3669:
3381:Military camouflage patterns influenced
3372:
3348:
3313:
2733:
2632:
2545:
2473:
2177:
2108:
2094:
1974:
1929:Many animals of the open sea, like this
1924:
1792:
1631:
1473:On a longer timescale, animals like the
1289:
1243:
985:
820:
810:
599:, a species of nudibranch that feeds on
359:
273:
164:
39:
27:
12189:Random generalized Lotka–Volterra model
9347:
9257:
9214:
9203:
8745:
8648:
8401:
8258:Fighter Combat: Tactics and Maneuvering
8196:
8105:
8073:
7819:
7042:Gullan, P. J.; Cranston, P. S. (2010).
7035:
6972:
6905:
6869:"Cloak of Light Makes Drone Invisible?"
5520:
5291:
5097:
3830:
3806:
3794:
3782:
3756:
3744:
2884:German World War II camouflage patterns
2628:
1782:
774:is coloured like its desert background.
13367:
13027:
11997:Herbivore adaptations to plant defense
9553:Newman, Alex; Blechman, Hardy (2004).
9436:
9414:
9402:
9396:
9326:
9271:
9242:
9016:
8920:
8900:. Stackpole Books. pp. 116, 125.
8883:
8855:
8651:"Military uniforms and the law of war"
8585:Industrial & Engineering Chemistry
8386:
8374:
8362:
8350:
8184:
8120:
8052:
7932:
7889:
7772:The British Army on Campaign 1816–1902
7752:
7699:
7591:
7579:
7405:
7168:
7111:
7105:
7068:
6757:Young, Richard Edward (October 1983).
6710:
6698:
6686:
6601:
6592:
6586:
6369:
6322:
6316:
6261:
6152:"The mathematics of motion camouflage"
6058:
5824:
5728:
5287:. U.S. War Department. November 2015 .
5275:
5049:
5025:
4833:
4141:
3951:
3890:
3866:
3854:
3842:
3732:
3717:
2972:, a zoologist from Sydney University.
2803:the Germans off guard multiple times.
2404:lured enemy submarines by mimicking a
1036:almost invisible in the Israeli desert
682:
223:for well over a century. According to
13001:
12843:
12026:
11279:
11106:Evolution of color vision in primates
10929:
10903:
10627:
9679:
9292:
9148:
8771:Freedberg, S. J. Jr. (25 June 2012).
8564:. Military Suppliers & News. 2012
8516:from the original on 17 November 2015
8229:
7978:. Memorial-Caen. 1998. Archived from
7952:. Memorial-Caen. 1998. Archived from
7846:
7469:
7117:
6841:. Royal Canadian Navy. Archived from
6756:
6722:
6469:
6192:
5793:
5609:U.S. War Department (November 1943).
5364:Mitchell, G.; Skinner, J. D. (2003).
5284:FM 5–20: Camouflage, Basic Principles
5256:
5250:
5078:
4959:
4894:(1821). The Royal Society: 20152182.
4126:
2327:lures its insect prey by mimicking a
1400:, squid, octopus and even the isopod
981:
12012:Predator avoidance in schooling fish
9228:
9045:"Love and War: The Weaponized Woman"
8687:from the original on 13 August 2011.
8254:
7883:
7603:
7417:
7393:
7181:Welbergen, J; Davies, N. B. (2011).
7080:
6920:
6674:
6636:
6574:
6363:
6249:
6000:
5927:. Horned Lizard Conservation Society
5913:
5907:
5836:
5783:from the original on 16 August 2017.
5596:
5182:from the original on 29 October 2013
5152:
5066:
5037:
5013:
3939:
3927:
3915:
3818:
2081:and the shrimps it associates with,
2025:(stinging cells) of the transparent
1746:Countershaded ship and submarine in
1279:
1239:
13306:The Chemical Basis of Morphogenesis
12462:Intermediate disturbance hypothesis
10424:Marine Corps Combat Utility Uniform
9177:"Stephen Sprouse | Suit | American"
8832:
8804:"Berlin Brigade Urban Paint Scheme"
7797:The British Army in World War I (1)
7495:The Journal of Experimental Biology
7274:(June 2011). Burr, David C. (ed.).
7216:"Cuckoo in egg pattern 'arms race'"
7149:Great War Primary Documents Archive
6532:"Adaptiv – A Cloak of Invisibility"
5939:
5602:
4932:Field observations of puff adders (
4142:Watson, Traci (14 September 2016).
3353:Cellphone tower disguised as a tree
3344:
2869:
2270:to lure their prey. For example, a
1935:jellyfish, are largely transparent.
1649:. Countershading is widely used by
968:plants such as the saw greenbriar,
805:has the colour of fresh vegetation.
13:
12215:Ecological effects of biodiversity
9460:
9286:
9077:. Canadian War Museum. 5 June 2009
8801:
7618:10.1111/j.1467-8306.1991.tb01681.x
6554:"Innovation Adaptiv Car Signature"
6156:Proceedings of the Royal Society B
6018:Proceedings of the Royal Society B
5214:Proceedings of the Royal Society B
4654:10.1093/oso/9780199688678.003.0003
4563:10.1093/oso/9780199688678.003.0002
3301:developed in 1962, widely used in
2782:, and may have been influenced by
2723:
2167:
1189:has covered its body with sponges.
1120:
14:
13416:
11551:Generalist and specialist species
9815:As evidence for natural selection
9657:
9254:Reprinted 1985, Penguin Classics.
9208:
8444:from the original on 14 June 2013
7916:Guirand de Scévola, Lucien-Victor
7799:. Osprey Publishing. p. 37.
7679:. Osprey Publishing. p. 20.
6736:"Midwater Squid, Abralia veranyi"
6604:Trends in Ecology & Evolution
6470:Hearn, Brian (20 February 2012).
6345:from the original on 22 July 2011
6061:"Dragonfly flight tricks the eye"
2608:The Hunting Book of Gaston Phebus
2262:Mimesis is also employed by some
1878:, 1942, set to maximum brightness
1621:
467:
289:with camouflage they acquired as
157:fiction and works of literature.
13128:
12274:Occupancy–abundance relationship
10995:
10671:Lucien-Victor Guirand de Scévola
9926:Lucien-Victor Guirand de Scévola
9798:
9748:Coincident disruptive coloration
9628:What are Camouflage and Mimicry?
9169:
9142:
9115:
9067:
9022:
8973:
8947:
8926:
8849:
8826:
8795:
8764:
8715:
8700:FM 21–76 US Army Survival Manual
8642:
8616:. MBI Publishing, Zenith Press.
8603:
8576:
8554:
8528:
8492:
8456:
8420:
8395:
8389:The Role of Science and Industry
8380:
8319:
8288:
8281:Stephenson, Hubert Kirk (1948).
8275:
8248:
8234:. Headline Review. p. 278.
8223:
8211:from the original on 19 May 2014
8190:
8160:
8145:
8114:
8067:
8058:
8020:
7968:
7938:
7908:
7890:Wright, Patrick (23 June 2005).
7869:
7855:
7840:
7813:
7788:
7763:
7746:
7727:
7705:
7668:
7653:
7632:
7597:
7523:
7482:
7463:
7411:
7254:10.1046/j.1474-919X.2002.00085.x
7214:Brennand, Emma (24 March 2011).
7207:
7174:
7136:
7074:
6926:
6773:
6750:
6642:
4409:10.1034/j.1600-0749.2002.00039.x
3891:Wright, Patrick (23 June 2005).
3785:, p. Fold-out after p. 339.
3645:
3636:
3564:
3548:
3513:
3290:
3274:
3249:
3230:
3210:
3188:
3092:
3076:
3060:
3042:
3021:
3005:
2857:
2842:
2826:
2756:Lucien-Victor Guirand de Scévola
2462:
2446:
2431:
2412:
2393:
2370:
2354:
2338:
2317:
2302:
2046:(coelenterates), siphonophores,
1899:
1883:
1861:
1767:
1755:
1739:
1720:
1701:
1606:
1587:
1568:
1556:
1547:Norwegian volunteer soldiers in
1540:
1528:
1439:
1423:
1357:
1341:
1323:
1227:
1210:
1194:
1178:
1085:
1069:
1053:
1041:
1025:
957:
942:
926:
911:
899:
887:
794:
779:
764:
752:
653:
636:
12294:Relative abundance distribution
12007:Plant defense against herbivory
11874:Competitive exclusion principle
11586:Mesopredator release hypothesis
10653:
9620:
9198:
8204:. U.S. Army Russian Institute.
6952:10.1590/S1679-62252006000200008
6546:
6498:
6420:"Day Octopuses, Octopus cyanea"
6412:
6267:
6059:Hopkin, Michael (5 June 2003).
5842:
5787:
5734:
5469:
5158:
5072:
4774:
4714:
4670:
4628:
4579:
4537:
4480:
4436:Molecular Biology and Evolution
4423:
4384:
4335:
4270:
4211:
4160:
4144:"This Dinosaur Wore Camouflage"
4135:
4120:
4069:
4018:
3884:
3627:
3614:
3605:
3225:in 1963 and universally by 1968
2940:Home Guard Manual of Camouflage
2521:
2001:, which is made of the protein
1914:
83:is among other things used for
11879:Consumer–resource interactions
10496:Operational Camouflage Pattern
9962:Adaptive Coloration in Animals
9705:
9666:– interplay of science and art
9590:. Cambridge University Press.
9267:. Kegan Paul, Trench, Trübner.
9235:Adaptive Coloration in Animals
9181:The Metropolitan Museum of Art
8933:du Plessis, A. (3 July 2002).
7085:. Cornell Press. p. 233.
6867:Hambling, David (9 May 2008).
5435:Journal of Theoretical Biology
5261:. Prentice Hall. p. 219.
5083:. HarperCollins. p. 158.
3750:
3587:
3237:2007 2-colour snow variant of
3205:pattern to be issued, in 2002.
2541:
2126:are camouflaged by silvering.
1097:
423:Adaptive Coloration in Animals
231:, features such as camouflage
191:including the octopus, in his
1:
12725:Biological data visualization
12552:Environmental niche modelling
12279:Population viability analysis
10293:Australian Disruptive Pattern
9606:Mimicry in plants and animals
9353:The Biology of the Deep Ocean
9152:A Brief History of Camouflage
8174:. Seeley Service. p. 79.
7639:Payne-Gallwey, Ralph (1903).
6451:. Churchill Polar Bears. 2011
5615:Tactical and Technical Trends
5455:10.1016/s0022-5193(03)00196-6
4514:10.1016/S0960-9822(03)00128-3
4127:Pavid, Katie (28 June 2016).
3662:
3531:
3495:A Brief History of Camouflage
3264:
3115:
2892:, the zoologist Hugh Cott, a
2792:smoke blown in someone's face
2758:, employing artists known as
2619:
2579:
2562:
2282:disguised as merchant ships.
2009:which is made of the protein
1985:Hyalinobatrachium uranoscopum
1462:relatively slowly, mainly by
1396:, frog, flatfish such as the
741:evolved between 1860 and 1940
621:
12210:Density-dependent inhibition
10557:Diffused lighting camouflage
10442:Universal Camouflage Pattern
10075:USN WWII camouflage measures
9075:"Camouflage: The Exhibition"
8649:Pfanner, Toni (March 2004).
8197:Keating, Kenneth C. (1981).
7675:Haythornthwaite, P. (2002).
7301:10.1371/journal.pone.0020233
6616:10.1016/0169-5347(88)90079-1
6218:10.1371/journal.pbio.0040108
5794:Bates, Mary (10 June 2015).
5731:, pp. 50–51 and passim.
5123:10.1016/j.visres.2008.02.011
3690:. Firenze University Press.
3686:; Fiorito, Graziano (2006).
3401:'s reaction in around 1915:
3168:Universal Camouflage Pattern
3160:camouflaged textile patterns
3155:, were often uncamouflaged.
2249:animals, for example when a
2090:
1876:diffused lighting camouflage
1833:Diffused lighting camouflage
454:
7:
12679:Liebig's law of the minimum
12514:Resource selection function
11405:Metabolic theory of ecology
11034:Simple eye in invertebrates
10502:Netherlands Fractal Pattern
10436:Tactical Assault Camouflage
10269:Disruptive Pattern Material
9520:Goodden, Henrietta (2009).
9355:. Oxford University Press.
9297:. Sydney University Press.
8896:Blakeley, Peter F. (2012).
8856:Engber, D. (5 July 2012) .
8074:Goodden, Henrietta (2007).
8006:Online Etymology Dictionary
7660:Saunders, Nicholas (2005).
7645:. Longmans, Green. p.
7472:J. Entom. Soc. South Africa
7083:The Social Biology of Wasps
4844:10.1007/978-3-642-40462-7_5
4173:Frontiers in Marine Science
4149:National Geographic Society
3463:of poetry and art, and the
3219:Disruptive Pattern Material
3122:List of camouflage patterns
2703:and his second in command,
2526:
1835:" was trialled by Canada's
506:
129:. On land, artists such as
21:Camouflage (disambiguation)
10:
13421:
12579:Niche apportionment models
12299:Relative species abundance
11503:Primary nutritional groups
11400:List of feeding behaviours
11230:Infrared sensing in snakes
9671:A Chronology of Camouflage
9586:; Merilaita, Sami (2011).
9568:Shell, Hanna Rose (2012).
9441:. Thames and Hudson, with
8746:Brayley, Martin J (2009).
8402:Shapiro, Danielle (2016).
8232:Kursk: the greatest battle
8156:. 1 June 1971. p. 12.
7770:Barthorp, Michael (1988).
7677:British Rifleman 1797–1815
7559:10.1016/j.zool.2013.10.004
7118:Beyer, Kenneth M. (1999).
6763:Bulletin of Marine Science
5921:"What is a Horned Lizard?"
5611:"Principles of Camouflage"
5567:Sherbrooke, W. C. (2003).
3555:Camouflage clothing in an
3309:
3119:
2909:created the comprehensive
2873:
2727:
2530:
2466:
2233:Cryptic aggressive mimicry
2226:
2222:
2171:
1918:
1786:
1733:Carcharhinus amblyrhynchos
1625:
1381:
1378:Changeable skin coloration
1283:
1148:uses its hind legs and a '
1124:
1101:
994:Some animals, such as the
814:
663:resembles a broken branch.
628:List of camouflage methods
625:
579:
375:as if it were camouflaged.
160:
75:, eliminating shadow, and
18:
13336:
13286:D'Arcy Wentworth Thompson
13229:
13137:
13126:
13035:
12925:
12877:
12828:
12760:Ecosystem based fisheries
12702:
12602:
12527:
12400:
12372:Interspecific competition
12337:
12264:Minimum viable population
12197:
12122:Maximum sustainable yield
12107:Intraspecific competition
12102:Effective population size
12065:
11982:Anti-predator adaptations
11967:
11846:
11773:
11730:
11652:
11619:
11516:
11493:Photosynthetic efficiency
11428:
11322:
11202:
11114:
11101:Evolution of color vision
11086:
11004:
10993:
10935:
10759:
10661:
10586:
10549:
10528:
10521:
10410:
10365:Camouflage Central-Europe
10359:Desert Camouflage Pattern
10235:
10206:
10156:
10099:
10092:
10083:
10032:
10023:
9983:
9911:
9872:
9865:
9807:
9796:
9775:Multi-spectral camouflage
9713:
9333:. Yale University Press.
9315:Sydney University Press.
8991:Scribners. Archived from
8670:10.1017/s1560775500180113
8610:Richardson, Doug (2001).
8261:. Naval Institute Press.
8034:. Oxford University Press
8032:Oxford English Dictionary
7759:. John W. Parker and Son.
7122:. Naval Institute Press.
7081:Ross, Kenneth G. (1991).
7012:10.1016/j.cub.2020.06.044
5750:American Museum Novitates
5397:10.1080/00359190309519935
4613:10.1007/s00338-017-1603-8
4247:10.1016/j.cub.2017.07.061
4186:10.3389/fmars.2021.639670
2998:American School of Design
2660:skirmishing units of the
2652:with weapons such as the
2078:Microphilypnus amazonicus
1874:by night with incomplete
1837:National Research Council
13395:Antipredator adaptations
12938:Anti-predator adaptation
12750:Ecological stoichiometry
12715:Alternative stable state
10889:Wilfred Clement Von Berg
9951:Johann Georg Otto Schick
9273:Thayer, Abbott Handerson
9249:On the Origin of Species
8703:. Department of the Army
8542:. Department of the Army
8500:"FM 21-305/AFMAN 24-306"
8365:, pp. 154, 186–188.
8300:The Herring Gull's World
8255:Shaw, Robert L. (1985).
7742:(Winter): 341–347. 2004.
7713:"Killers in Green Coats"
7143:McMullen, Chris (2001).
5631:; Merilaita, S. (2009).
4960:Costa, James T. (2007).
3580:
3571:A camouflage skirt as a
3369:Fashion, art and society
3299:Six-Color Desert Pattern
3258:People's Liberation Army
2938:, writing the practical
2794:. The English zoologist
2467:Not to be confused with
2312:caterpillars mimic twigs
2084:Pseudopalaemon gouldingi
1575:Snow-camouflaged German
1430:Four frames of the same
542:horizontal gene transfer
499:has been preserved with
432:like Thayer's, mainly a
95:. Some animals, such as
81:ability to produce light
13400:Biological interactions
12912:Paradox of the plankton
12594:Ontogenetic niche shift
12457:Ideal free distribution
12367:Ecological facilitation
12117:Malthusian growth model
12087:Consumer-resource model
11944:Paradox of the plankton
11909:Energy systems language
11629:Chemoorganoheterotrophy
11596:Optimal foraging theory
11571:Heterotrophic nutrition
10726:Abbott Handerson Thayer
10341:Desert Night Camouflage
9894:Abbott Handerson Thayer
9630:. Crabtree Publishing.
9155:. Black Sparrow Press.
9031:Illustrated London News
8302:. Collins. p. 14.
6939:Neotropical Ichthyology
6150:Glendinning, P (2004).
4646:Oxford University Press
4555:Oxford University Press
4047:10.1126/science.1124309
3705:6 February 2018 at the
3678:Aristotle (c. 350 BC).
3653:Ian Hamilton Finlay#Art
3444:Modern artists such as
3424:Illustrated London News
3028:1937 summer variant of
2800:Royal Naval Air Service
2638:Green-jacketed rifleman
2062:worms, many shrimplike
1750:1902 patent application
1414:Smith's dwarf chameleon
1263:flat-tail horned lizard
1062:flat-tail horned lizard
1004:roundtail horned lizard
719:, or reptiles like the
381:Abbott Handerson Thayer
12740:Ecological forecasting
12684:Marginal value theorem
12482:Landscape epidemiology
12417:Cross-boundary subsidy
12352:Biological interaction
11702:Microbial intelligence
11390:Green world hypothesis
10809:Stanley William Hayter
10721:Solomon Joseph Solomon
9886:The Colours of Animals
9880:Edward Bagnall Poulton
9770:Multi-scale camouflage
9641:Mettler, Rene (2001).
9327:Forbes, Peter (2009).
9264:The Colours of Animals
9149:Frank, Thaisa (1992).
9122:Haldeman, Joe (2004).
8540:Appendix D: Camouflage
8387:Mellor, D. P. (1958).
7897:London Review of Books
7439:10.1098/rstb.2008.0221
7362:10.1186/1741-7007-9-81
7145:"Royal Navy 'Q' Ships"
6839:Naval Museum of Quebec
6798:10.1098/rstb.2000.0681
6292:10.1098/rsbl.2008.0173
6168:10.1098/rspb.2003.2622
6038:10.1098/rspb.1995.0004
5874:10.1098/rsbl.2015.0325
5698:10.1098/rspb.2009.0052
5649:10.1098/rstb.2008.0216
5368:Giraffa camelopardalis
5335:(Article 8): 119–231.
5226:10.1098/rspb.2006.3614
4901:10.1098/rspb.2015.2182
4803:10.1098/rspb.2019.2664
4742:10.1098/rspb.2005.3438
4372:Cite journal requires
4349:10.21203/rs.2.10812/v1
3898:London Review of Books
3758:Wallace, Alfred Russel
3452:, is described by the
3434:
3418:
3378:
3354:
3335:driven grouse shooting
3326:
3239:Finnish Defence Forces
3110:overalls, January 1943
2747:
2674:Charles Hamilton Smith
2645:
2583: 360–400 AD
2554:
2483:
2186:
2114:
2106:
1989:
1936:
1805:
1637:
1295:
1253:
991:
830:
772:Black-faced sandgrouse
416:The English zoologist
376:
307:The Colours of Animals
298:Edward Bagnall Poulton
296:The English zoologist
293:
272:
217:
176:
49:
46:disruptively patterned
37:
13120:Widmanstätten pattern
12878:Patterns of evolution
12745:Ecological humanities
12644:Ecological energetics
12589:Niche differentiation
12452:Habitat fragmentation
12220:Ecological extinction
12167:Small population size
11919:Feed conversion ratio
11899:Ecological succession
11831:San Francisco Estuary
11745:Ecological efficiency
11687:Microbial cooperation
11235:Monocular deprivation
11194:Underwater camouflage
11189:Structural coloration
11167:Disruptive coloration
10478:Multi-Terrain Pattern
10460:Airman Battle Uniform
10257:Rhodesian Brushstroke
9857:Underwater camouflage
9743:Disruptive coloration
8230:Clark, Lloyd (2011).
7950:The Elm at Vermezeele
7920:Revue des Deux Mondes
7795:Chappell, M. (2003).
7200:10.1093/beheco/arr008
6656:. Stanford University
6073:10.1038/news030602-10
5987:10.1093/beheco/arv125
5741:Wierauch, C. (2006).
5257:Sweet, K. M. (2006).
4699:10.1093/beheco/ars156
4641:Disruptive camouflage
4448:10.1093/molbev/msy004
4397:Pigment Cell Research
3595:Alfred Russel Wallace
3429:
3403:
3376:
3352:
3317:
3120:Further information:
2874:Further information:
2737:
2728:Further information:
2636:
2549:
2477:
2347:Teratodus monticollis
2227:Further information:
2183:Blackdevil anglerfish
2181:
2174:Underwater camouflage
2172:Further information:
2139:structural coloration
2112:
2098:
2005:, and the vertebrate
1992:Some tissues such as
1978:
1928:
1921:Underwater camouflage
1919:Further information:
1796:
1665:; and birds, such as
1635:
1599:Chamaeleo calyptratus
1382:Further information:
1293:
1247:
989:
845:. Predators like the
824:
817:disruptive coloration
811:Disruptive coloration
626:Further information:
554:walking stick insects
429:disruptive camouflage
363:
277:
247:
199:
168:
154:cell telephone towers
87:on the undersides of
73:disruptive coloration
59:'s spotted coat, the
43:
31:
13405:Evolutionary ecology
12890:Convergent evolution
12871:Evolutionary ecology
12770:Evolutionary ecology
12735:Ecological footprint
12730:Ecological economics
12654:Ecological threshold
12649:Ecological indicator
12519:Source–sink dynamics
12472:Land change modeling
12467:Insular biogeography
12319:Species distribution
12058:Modelling ecosystems
11717:Microbial metabolism
11556:Intraguild predation
11345:Biogeochemical cycle
11311:Modelling ecosystems
11215:Blindness in animals
11147:Counter-illumination
11096:Evolution of the eye
10595:Dazzled and Deceived
9760:Distractive markings
9738:Counter-illumination
9437:Newark, Tim (2007).
9224:. Swan Sonnenschein.
9216:Beddard, Frank Evers
9204:Camouflage in nature
8480:on 26 September 2021
8133:on 19 September 2016
7826:Fortress Study Group
7343:(25 November 2011).
6997:(17): 3470–3476.e3.
6654:"The Color of Birds"
6332:Bioscience Explained
5414:on 23 September 2015
5313:on 17 November 2021.
4232:(18): 2833–2842.e6.
3297:US "Chocolate Chip"
3104:Battle of Stalingrad
2837:in dazzle camouflage
2770:probably comes from
2629:19th-century origins
2595:speculatoria navigia
2217:convergent evolution
1799:counter-illumination
1789:Counter-illumination
1783:Counter-illumination
1776:Focke-Wulf Fw 190D-9
1205:with plant materials
1110:distractive markings
1104:Distractive markings
867:camouflage in plants
711:, birds such as the
407:Peacock in the Woods
369:Peacock in the Woods
347:orange tip butterfly
169:Octopuses like this
85:counter-illumination
13354:Mathematics and art
13344:Pattern recognition
13314:Aristid Lindenmayer
12963:Distraction display
12907:Divergent evolution
12820:Theoretical ecology
12795:Natural environment
12659:Ecosystem diversity
12629:Ecological collapse
12619:Bateman's principle
12574:Limiting similarity
12487:Landscape limnology
12309:Species homogeneity
12147:Population modeling
12142:Population dynamics
11959:Trophic state index
10484:Australian Multicam
10287:U.S. "M81" Woodland
10045:Aircraft camouflage
10040:Military camouflage
9753:Disruptive eye mask
9443:Imperial War Museum
9397:Military camouflage
9311:Elias, Ann (2015).
9293:Elias, Ann (2011).
9055:on 12 December 2012
8597:10.1021/ie50441a019
8377:, pp. 156–166.
8353:, pp. 151–152.
8187:, pp. 149–150.
7935:, pp. 104–105.
7879:. War Office. 1904.
7865:. War Office. 1891.
7582:, pp. 211–212.
7551:2014Zool..117..163H
7292:2011PLoSO...620233S
7003:2020CBio...30E3470D
6975:, pp. 192–195.
6908:, pp. 190–191.
6792:(1401): 1269–1272.
6689:, pp. 146–150.
6556:. BAE Systems. 2012
6534:. BAE Systems. 2011
6323:Wallin, M. (2002).
6264:, pp. 52, 236.
6111:2003Natur.423..604M
6030:1995RSPSB.259...19S
6003:, pp. 141–143.
5827:, pp. 102–103.
5800:National Geographic
5692:(1663): 1905–1910.
5599:, pp. 104–105.
5535:1990FuEco...4..463G
5447:2003JThBi.224..483L
5389:2003TRSSA..58...51M
5325:Roosevelt, Theodore
5220:(1600): 2433–2436.
4980:2007Natur.448..408C
4785:(15 January 2020).
4736:(1591): 1267–1271.
4604:2017CorRe..36.1121W
4550:Background matching
4505:2003CBio...13..448E
4305:10.1038/nature17951
4297:2016Natur.534..102H
4238:2017CBio...27E2833G
4156:on 6 November 2019.
4098:10.1038/nature12899
4090:2014Natur.506..484L
4039:2006Sci...312.1614S
4033:(5780): 1614–1620.
3986:2016NatSR...619966T
3954:, pp. 153–155.
3942:, pp. 174–186.
3918:, pp. 172–173.
3881:, pp. 132–133.
3684:Gherardi, Francesca
3446:Ian Hamilton Finlay
3439:anti-war protestors
3337:conceal hunters in
3149:Aircraft camouflage
2876:list of camoufleurs
2778:slang term meaning
2730:list of camoufleurs
2666:95th Rifle Regiment
2533:Military camouflage
2516:barberpole illusion
2453:Wrap-around spider
2345:Hooded grasshopper
2099:The adult herring,
1651:terrestrial animals
1273:Extatosoma tiaratum
1219:Reduvius personatus
865:The possibility of
835:by masking the eyes
733:industrial melanism
683:Background matching
671:techniques against
525:Aliivibrio fischeri
486:leatherback turtles
484:, reveal that both
401:'... nor any '
323:Frank Evers Beddard
312:swallow-tailed moth
179:In ancient Greece,
115:Military camouflage
13292:On Growth and Form
13192:Logarithmic spiral
13029:Patterns in nature
12958:Deimatic behaviour
12917:Predator satiation
12902:Parallel evolution
12831:Outline of ecology
12780:Industrial ecology
12775:Functional ecology
12639:Ecological deficit
12584:Niche construction
12547:Ecosystem engineer
12324:Species–area curve
12245:Introduced species
12060:: Other components
11992:Deimatic behaviour
11894:Ecological network
11826:North Pacific Gyre
11811:hydrothermal vents
11750:Ecological pyramid
11697:Microbial food web
11508:Primary production
11453:Foundation species
11162:Deimatic behaviour
10789:Bainbridge Copnall
10601:Stealth technology
10111:Splittertarnmuster
10006:Thomas N. Sherratt
9509:. Bobolink Books.
9491:. Bobolink Books.
9473:. Bobolink Books.
9259:Poulton, Edward B.
8995:on 1 February 2014
8923:, pp. 48, 50.
7847:Lewis, JF (1890).
7508:10.1242/jeb.065540
7187:Behavioral Ecology
6512:. 5 September 2011
6390:10.1007/BF00347297
6325:"Nature's Palette"
5974:Behavioral Ecology
5947:"Leafy Sea Dragon"
5523:Functional Ecology
5079:Still, J. (1996).
4942:Suricata suricatta
4797:(1918): 20192664.
4686:Behavioral Ecology
3974:Scientific Reports
3680:Historia Animalium
3411:Gertrude Stein in
3379:
3355:
3327:
3203:digital camouflage
3037:Plane tree pattern
2990:RAAF Base Richmond
2970:William John Dakin
2748:
2681:American Civil War
2646:
2555:
2512:wagon-wheel effect
2484:
2256:Leipomeles dorsata
2187:
2131:marine hatchetfish
2115:
2107:
2056:gastropod molluscs
1990:
1949:refractive indices
1937:
1806:
1690:William John Dakin
1638:
1350:Australian Emperor
1296:
1254:
992:
982:Eliminating shadow
831:
691:are mainly green;
462:positive selection
377:
294:
284:swallowtailed moth
227:'s 1859 theory of
194:Historia animalium
177:
69:leaf-mimic katydid
50:
38:
13362:
13361:
13319:Benoît Mandelbrot
13219:Self-organization
13155:Natural selection
13145:Pattern formation
12995:
12994:
12933:Signalling theory
12837:
12836:
12720:Balance of nature
12477:Landscape ecology
12362:Community ecology
12304:Species diversity
12240:Indicator species
12235:Gradient analysis
12112:Logistic function
12020:
12019:
11977:Animal coloration
11954:Trophic mutualism
11692:Microbial ecology
11483:Photoheterotrophs
11468:Myco-heterotrophy
11380:Ecosystem ecology
11365:Carrying capacity
11330:Abiotic component
11273:
11272:
11265:Visual perception
11260:Underwater vision
11225:Feature detection
11220:Eyespot apparatus
11179:Eyespot (mimicry)
11127:Animal coloration
10930:Vision in animals
10897:
10896:
10676:Jean-Louis Forain
10621:
10620:
10582:
10581:
10578:
10577:
10517:
10516:
10329:Camouflage Daguet
10202:
10201:
10055:Dazzle camouflage
10019:
10018:
9921:Mary Taylor Brush
9765:Motion camouflage
9733:Active camouflage
9651:978-1-85103-298-3
9643:Animal Camouflage
9636:978-0-86505-962-7
9614:978-0-07-070100-7
9602:Wickler, Wolfgang
9596:978-0-521-15257-0
9578:978-1-935-40822-2
9563:978-0-9543404-0-7
9548:978-1-58567-381-0
9530:978-0-906290-87-3
9524:. Unicorn Press.
9515:978-0-9713244-7-3
9505:(editor) (2012).
9497:978-0-9713244-6-6
9452:978-0-500-51347-7
9429:978-0-8018-5130-8
9388:978-1-60819-216-8
9371:Rothenberg, David
9362:978-0-19-854956-7
9340:978-0-300-17896-8
9321:978-1-743324-25-7
9304:978-1-920899-73-8
9221:Animal Coloration
9162:978-0-87685-857-8
9135:978-0-441-01161-2
9103:. Tate. July 2008
8907:978-0-8117-0566-0
8837:. United Dynamics
8783:on 31 August 2012
8757:978-1-84797-137-1
8623:978-0-7603-1051-9
8413:978-0-8166-9341-2
8309:978-0-00-219444-0
8268:978-0-87021-059-4
8241:978-0-7553-3639-5
8172:A Brush with Life
8168:Wilkinson, Norman
8085:978-0-906290-87-3
8055:, pp. 85–89.
7806:978-1-84176-399-6
7781:978-0-85045-849-7
7433:(1516): 537–548.
7341:Ruxton, Graeme D.
7272:Cuthill, Innes C.
7129:978-1-55750-044-1
7092:978-0-801-49906-7
7055:978-1-4443-3036-6
6725:, pp. 57–66.
6589:, pp. 72–73.
6577:, pp. 35–46.
6449:"Arctic Wildlife"
6252:, pp. 30–31.
6162:(1538): 477–481.
5925:hornedlizards.org
5850:Ruxton, Graeme D.
5643:(1516): 481–488.
5580:978-0-520-22825-2
5268:978-0-13-170356-8
5206:Cuthill, Innes C.
5167:Cuthill, Innes C.
5155:, pp. 83–91.
5116:(10): 1242–1253.
5090:978-0-00-220010-3
5052:, pp. 45–46.
4853:978-3-642-40461-0
4783:Cuthill, Innes C.
4722:Cuthill, Innes C.
4663:978-0-19-968867-8
4636:Ruxton, Graeme D.
4572:978-0-19-968867-8
4545:Ruxton, Graeme D.
4291:(7605): 102–105.
4084:(7489): 484–488.
3994:10.1038/srep19966
3930:, pp. 47–67.
3869:, pp. 5, 16.
3821:, pp. 74–75.
3696:978-88-8453-377-7
3485:is a novel about
3473:Et in Arcadia ego
2986:Bankstown Airport
2952:Operation Bertram
2942:. The film-maker
2819:dazzle camouflage
2701:Sir Harry Lumsden
2691:uniforms was the
2566: 172–250 AD
2469:dazzle camouflage
2457:mimicking a stick
2383:Operation Bertram
2213:natural selection
2211:), implying that
2205:phylogenetic tree
1507:Active camouflage
1384:Active camouflage
1367:
1308:motion camouflage
1286:Motion camouflage
1280:Motion camouflage
1240:Cryptic behaviour
1144:of the predatory
1014:European nightjar
787:Egyptian nightjar
573:domesticated cats
569:eastern deer mice
538:Sepia officinalis
530:active camouflage
403:sexually selected
397:' mark, not one '
367:'s 1907 painting
302:animal coloration
242:Origin of Species
229:natural selection
103:, are capable of
13412:
13170:Sexual selection
13132:
13022:
13015:
13008:
12999:
12998:
12864:
12857:
12850:
12841:
12840:
12537:Ecological niche
12509:selection theory
12329:Umbrella species
12314:Species richness
12250:Invasive species
12230:Flagship species
12137:Population cycle
12132:Overexploitation
12097:Ecological yield
12047:
12040:
12033:
12024:
12023:
11929:Mesotrophic soil
11869:Climax community
11801:Marine food webs
11740:Biomagnification
11541:Chemoorganotroph
11395:Keystone species
11355:Biotic component
11300:
11293:
11286:
11277:
11276:
11245:Palpebral (bone)
11078:Schizochroal eye
10999:
10924:
10917:
10910:
10901:
10900:
10884:Julian Trevelyan
10819:Jasper Maskelyne
10751:Norman Wilkinson
10741:Edward Wadsworth
10716:Kimon Nicolaïdes
10691:John Graham Kerr
10648:
10641:
10634:
10625:
10624:
10526:
10525:
10097:
10096:
10090:
10089:
10030:
10029:
9936:Norman Wilkinson
9931:John Graham Kerr
9870:
9869:
9802:
9790:Urban camouflage
9700:
9693:
9686:
9677:
9676:
9540:Deception in War
9456:
9433:
9411:
9404:Barkas, Geoffrey
9392:
9380:
9366:
9344:
9308:
9282:
9268:
9253:
9239:
9225:
9192:
9191:
9189:
9187:
9173:
9167:
9166:
9146:
9140:
9139:
9119:
9113:
9112:
9110:
9108:
9093:
9087:
9086:
9084:
9082:
9071:
9065:
9064:
9062:
9060:
9041:
9035:
9034:
9026:
9020:
9014:
9005:
9004:
9002:
9000:
8989:Toklas, Alice B.
8987:. Translated by
8977:
8971:
8970:
8968:
8966:
8951:
8945:
8944:
8930:
8924:
8918:
8912:
8911:
8893:
8887:
8881:
8875:
8874:
8872:
8870:
8853:
8847:
8846:
8844:
8842:
8830:
8824:
8823:
8821:
8819:
8814:on 12 March 2013
8799:
8793:
8792:
8790:
8788:
8779:. Archived from
8768:
8762:
8761:
8743:
8737:
8736:
8734:
8732:
8719:
8713:
8712:
8710:
8708:
8695:
8689:
8688:
8686:
8655:
8646:
8640:
8639:
8637:
8635:
8626:. Archived from
8607:
8601:
8600:
8580:
8574:
8573:
8571:
8569:
8562:"SSZ Camouflage"
8558:
8552:
8551:
8549:
8547:
8532:
8526:
8525:
8523:
8521:
8515:
8504:
8496:
8490:
8489:
8487:
8485:
8479:
8468:
8460:
8454:
8453:
8451:
8449:
8443:
8437:. pp. 1–4.
8432:
8424:
8418:
8417:
8399:
8393:
8392:
8384:
8378:
8372:
8366:
8360:
8354:
8348:
8342:
8341:
8339:
8337:
8323:
8317:
8316:
8292:
8286:
8279:
8273:
8272:
8252:
8246:
8245:
8227:
8221:
8220:
8218:
8216:
8210:
8203:
8194:
8188:
8182:
8176:
8175:
8164:
8158:
8157:
8149:
8143:
8142:
8140:
8138:
8129:. Archived from
8118:
8112:
8111:
8103:
8090:
8089:
8071:
8065:
8062:
8056:
8050:
8044:
8043:
8041:
8039:
8024:
8018:
8017:
8015:
8013:
7998:
7992:
7991:
7989:
7987:
7972:
7966:
7965:
7963:
7961:
7942:
7936:
7930:
7924:
7923:
7912:
7906:
7905:
7887:
7881:
7880:
7873:
7867:
7866:
7859:
7853:
7852:
7844:
7838:
7837:
7817:
7811:
7810:
7792:
7786:
7785:
7767:
7761:
7760:
7750:
7744:
7743:
7731:
7725:
7724:
7722:
7720:
7709:
7703:
7697:
7691:
7690:
7672:
7666:
7665:
7657:
7651:
7650:
7636:
7630:
7629:
7601:
7595:
7589:
7583:
7577:
7571:
7570:
7536:
7527:
7521:
7520:
7510:
7486:
7480:
7479:
7467:
7461:
7460:
7450:
7415:
7409:
7403:
7397:
7391:
7385:
7384:
7374:
7364:
7333:
7324:
7323:
7313:
7303:
7267:
7258:
7257:
7237:
7231:
7230:
7228:
7226:
7211:
7205:
7204:
7202:
7178:
7172:
7171:, pp. 6–42.
7166:
7160:
7159:
7157:
7155:
7140:
7134:
7133:
7115:
7109:
7103:
7097:
7096:
7078:
7072:
7066:
7060:
7059:
7039:
7033:
7032:
7014:
6982:
6976:
6970:
6957:
6956:
6954:
6930:
6924:
6918:
6909:
6903:
6884:
6883:
6881:
6879:
6864:
6855:
6854:
6852:
6850:
6831:
6820:
6819:
6809:
6777:
6771:
6770:
6754:
6748:
6747:
6745:
6743:
6732:
6726:
6720:
6714:
6708:
6702:
6696:
6690:
6684:
6678:
6672:
6666:
6665:
6663:
6661:
6650:Ehrlich, Paul R.
6646:
6640:
6634:
6628:
6627:
6599:
6590:
6584:
6578:
6572:
6566:
6565:
6563:
6561:
6550:
6544:
6543:
6541:
6539:
6528:
6522:
6521:
6519:
6517:
6502:
6496:
6495:
6493:
6491:
6485:
6478:
6467:
6461:
6460:
6458:
6456:
6445:
6436:
6435:
6433:
6431:
6426:on 20 March 2016
6416:
6410:
6409:
6373:
6367:
6361:
6355:
6354:
6352:
6350:
6344:
6329:
6320:
6314:
6313:
6303:
6271:
6265:
6259:
6253:
6247:
6241:
6240:
6230:
6220:
6196:
6190:
6189:
6179:
6147:
6141:
6140:
6122:
6090:
6084:
6083:
6081:
6079:
6056:
6050:
6049:
6013:
6004:
5998:
5992:
5991:
5989:
5965:
5959:
5958:
5956:
5954:
5943:
5937:
5936:
5934:
5932:
5917:
5911:
5905:
5896:
5895:
5885:
5846:
5840:
5834:
5828:
5822:
5816:
5815:
5813:
5811:
5802:. Archived from
5791:
5785:
5784:
5782:
5747:
5738:
5732:
5726:
5720:
5719:
5709:
5677:
5671:
5670:
5660:
5625:
5619:
5618:
5606:
5600:
5594:
5585:
5584:
5564:
5555:
5554:
5518:
5512:
5511:
5505:
5501:
5499:
5491:
5473:
5467:
5466:
5430:
5424:
5423:
5421:
5419:
5413:
5407:. Archived from
5374:
5361:
5346:
5344:
5321:
5315:
5314:
5312:
5305:
5295:
5289:
5288:
5279:
5273:
5272:
5254:
5248:
5247:
5237:
5198:
5192:
5191:
5189:
5187:
5181:
5174:
5165:Osorio, Daniel;
5162:
5156:
5150:
5144:
5143:
5125:
5101:
5095:
5094:
5076:
5070:
5064:
5053:
5047:
5041:
5035:
5029:
5023:
5017:
5016:, pp. 5–19.
5011:
5005:
5004:
4991:
4957:
4951:
4950:
4948:using olfaction.
4940:) and meerkats (
4938:Canis familiaris
4921:
4903:
4875:
4869:
4868:
4831:
4825:
4824:
4814:
4778:
4772:
4771:
4761:
4718:
4712:
4711:
4701:
4674:
4668:
4667:
4632:
4626:
4625:
4615:
4583:
4577:
4576:
4541:
4535:
4534:
4516:
4484:
4478:
4477:
4467:
4427:
4421:
4420:
4388:
4382:
4381:
4375:
4370:
4368:
4360:
4339:
4333:
4332:
4274:
4268:
4267:
4249:
4215:
4209:
4208:
4198:
4188:
4164:
4158:
4157:
4152:. Archived from
4139:
4133:
4132:
4124:
4118:
4117:
4073:
4067:
4066:
4022:
4016:
4015:
4005:
3961:
3955:
3949:
3943:
3937:
3931:
3925:
3919:
3913:
3907:
3906:
3888:
3882:
3876:
3870:
3864:
3858:
3852:
3846:
3840:
3834:
3828:
3822:
3816:
3810:
3804:
3798:
3792:
3786:
3780:
3774:
3773:
3771:
3769:
3760:(8 March 1868).
3754:
3748:
3742:
3736:
3730:
3721:
3715:
3709:
3676:
3656:
3649:
3643:
3640:
3634:
3631:
3625:
3618:
3612:
3609:
3603:
3591:
3568:
3557:anti-war protest
3552:
3536:
3533:
3517:
3489:alien beings by
3416:
3345:Civil structures
3294:
3278:
3269:
3266:
3253:
3234:
3214:
3192:
3102:soldiers in the
3096:
3080:
3064:
3046:
3025:
3012:Maritime patrol
3009:
2978:Sydney Ure Smith
2890:Second World War
2870:Second World War
2861:
2846:
2830:
2815:Norman Wilkinson
2796:John Graham Kerr
2624:
2623: 1655–1740
2621:
2616:First Maroon War
2612:Jamaican Maroons
2584:
2581:
2567:
2564:
2508:visual illusions
2450:
2435:
2416:
2397:
2374:
2358:
2342:
2321:
2306:
2207:of bony fishes (
2154:
2153:
2149:
1903:
1887:
1865:
1771:
1759:
1743:
1724:
1705:
1610:
1595:Veiled chameleon
1591:
1572:
1560:
1544:
1532:
1515:announced their
1443:
1432:peacock flounder
1427:
1398:peacock flounder
1392:Animals such as
1369:
1368:
1345:
1327:
1268:leafy sea dragon
1250:leafy sea dragon
1231:
1214:
1198:
1182:
1170:blotched emerald
1089:
1073:
1057:
1045:
1029:
1018:Second World War
961:
946:
930:
915:
903:
891:
798:
783:
768:
756:
735:, including the
661:Papuan frogmouth
657:
645:Draco dussumieri
640:
605:P. melanocrachia
339:green tree-snake
245:, Darwin wrote:
215:
143:Second World War
34:peacock flounder
13420:
13419:
13415:
13414:
13413:
13411:
13410:
13409:
13380:Survival skills
13365:
13364:
13363:
13358:
13332:
13225:
13133:
13124:
13031:
13026:
12996:
12991:
12921:
12873:
12868:
12838:
12833:
12824:
12810:Systems ecology
12698:
12669:Extinction debt
12634:Ecological debt
12624:Bioluminescence
12605:
12598:
12567:marine habitats
12542:Ecological trap
12523:
12403:
12396:
12339:
12333:
12289:Rapoport's rule
12284:Priority effect
12225:Endemic species
12193:
12152:Population size
12068:
12061:
12051:
12021:
12016:
11969:
11963:
11949:Trophic cascade
11859:Bioaccumulation
11842:
11769:
11726:
11648:
11615:
11512:
11424:
11385:Ecosystem model
11318:
11304:
11274:
11269:
11198:
11110:
11082:
11000:
10991:
10931:
10928:
10898:
10893:
10879:Ernest Townsend
10774:Geoffrey Barkas
10755:
10731:Maximilian Toch
10657:
10652:
10622:
10617:
10606:Cloaking device
10574:
10545:
10513:
10412:
10406:
10317:Type 87 (China)
10237:
10231:
10198:
10172:(1917 aircraft)
10164:Camouflage tree
10152:
10123:Rauchtarnmuster
10102:
10079:
10070:Ship camouflage
10015:
9979:
9975:Timothy O'Neill
9970:Geoffrey Barkas
9907:
9861:
9803:
9794:
9785:Snow camouflage
9780:Self-decoration
9709:
9704:
9660:
9623:
9608:. McGraw-Hill.
9584:Stevens, Martin
9542:. John Murray.
9503:Behrens, Roy R.
9485:Behrens, Roy R.
9467:Behrens, Roy R.
9463:
9461:Further reading
9453:
9430:
9399:
9389:
9363:
9341:
9305:
9289:
9287:General reading
9244:Darwin, Charles
9211:
9206:
9201:
9196:
9195:
9185:
9183:
9175:
9174:
9170:
9163:
9147:
9143:
9136:
9120:
9116:
9106:
9104:
9095:
9094:
9090:
9080:
9078:
9073:
9072:
9068:
9058:
9056:
9043:
9042:
9038:
9028:
9027:
9023:
9015:
9008:
8998:
8996:
8981:Stein, Gertrude
8978:
8974:
8964:
8962:
8959:Automotive News
8953:
8952:
8948:
8931:
8927:
8919:
8915:
8908:
8894:
8890:
8882:
8878:
8868:
8866:
8854:
8850:
8840:
8838:
8831:
8827:
8817:
8815:
8800:
8796:
8786:
8784:
8769:
8765:
8758:
8744:
8740:
8730:
8728:
8721:
8720:
8716:
8706:
8704:
8697:
8696:
8692:
8684:
8664:(853): 99–100.
8653:
8647:
8643:
8633:
8631:
8630:on 7 April 2015
8624:
8608:
8604:
8581:
8577:
8567:
8565:
8560:
8559:
8555:
8545:
8543:
8534:
8533:
8529:
8519:
8517:
8513:
8502:
8498:
8497:
8493:
8483:
8481:
8477:
8466:
8462:
8461:
8457:
8447:
8445:
8441:
8430:
8426:
8425:
8421:
8414:
8400:
8396:
8385:
8381:
8373:
8369:
8361:
8357:
8349:
8345:
8335:
8333:
8325:
8324:
8320:
8310:
8296:Tinbergen, Niko
8293:
8289:
8283:Applied Physics
8280:
8276:
8269:
8253:
8249:
8242:
8228:
8224:
8214:
8212:
8208:
8201:
8195:
8191:
8183:
8179:
8165:
8161:
8151:
8150:
8146:
8136:
8134:
8119:
8115:
8104:
8093:
8086:
8072:
8068:
8063:
8059:
8051:
8047:
8037:
8035:
8028:"Camouflage, n"
8026:
8025:
8021:
8011:
8009:
8000:
7999:
7995:
7985:
7983:
7974:
7973:
7969:
7959:
7957:
7944:
7943:
7939:
7931:
7927:
7913:
7909:
7888:
7884:
7875:
7874:
7870:
7861:
7860:
7856:
7845:
7841:
7818:
7814:
7807:
7793:
7789:
7782:
7768:
7764:
7751:
7747:
7733:
7732:
7728:
7718:
7716:
7711:
7710:
7706:
7698:
7694:
7687:
7673:
7669:
7658:
7654:
7637:
7633:
7602:
7598:
7590:
7586:
7578:
7574:
7534:
7528:
7524:
7487:
7483:
7468:
7464:
7416:
7412:
7404:
7400:
7392:
7388:
7337:Stevens, Martin
7334:
7327:
7268:
7261:
7238:
7234:
7224:
7222:
7212:
7208:
7179:
7175:
7167:
7163:
7153:
7151:
7141:
7137:
7130:
7116:
7112:
7104:
7100:
7093:
7079:
7075:
7067:
7063:
7056:
7040:
7036:
6991:Current Biology
6983:
6979:
6971:
6960:
6931:
6927:
6919:
6912:
6904:
6887:
6877:
6875:
6865:
6858:
6848:
6846:
6833:
6832:
6823:
6778:
6774:
6755:
6751:
6741:
6739:
6734:
6733:
6729:
6721:
6717:
6709:
6705:
6697:
6693:
6685:
6681:
6673:
6669:
6659:
6657:
6647:
6643:
6635:
6631:
6600:
6593:
6585:
6581:
6573:
6569:
6559:
6557:
6552:
6551:
6547:
6537:
6535:
6530:
6529:
6525:
6515:
6513:
6504:
6503:
6499:
6489:
6487:
6486:on 4 March 2016
6483:
6476:
6468:
6464:
6454:
6452:
6447:
6446:
6439:
6429:
6427:
6418:
6417:
6413:
6374:
6370:
6362:
6358:
6348:
6346:
6342:
6327:
6321:
6317:
6280:Biology Letters
6272:
6268:
6260:
6256:
6248:
6244:
6197:
6193:
6148:
6144:
6120:10.1038/423604a
6091:
6087:
6077:
6075:
6057:
6053:
6024:(1354): 19–25.
6014:
6007:
5999:
5995:
5966:
5962:
5952:
5950:
5945:
5944:
5940:
5930:
5928:
5919:
5918:
5914:
5906:
5899:
5868:(6): 20150325.
5862:Biology Letters
5856:(1 June 2015).
5854:Stevens, Martin
5847:
5843:
5835:
5831:
5823:
5819:
5809:
5807:
5806:on 11 June 2015
5792:
5788:
5780:
5745:
5739:
5735:
5727:
5723:
5678:
5674:
5629:Stevens, Martin
5626:
5622:
5607:
5603:
5595:
5588:
5581:
5565:
5558:
5543:10.2307/2389314
5519:
5515:
5503:
5502:
5493:
5492:
5488:
5474:
5470:
5431:
5427:
5417:
5415:
5411:
5372:
5362:
5349:
5322:
5318:
5310:
5303:
5297:
5296:
5292:
5281:
5280:
5276:
5269:
5255:
5251:
5202:Stevens, Martin
5199:
5195:
5185:
5183:
5179:
5172:
5163:
5159:
5151:
5147:
5110:Vision Research
5102:
5098:
5091:
5077:
5073:
5065:
5056:
5048:
5044:
5040:, pp. 5–6.
5036:
5032:
5024:
5020:
5012:
5008:
4989:10.1038/448408c
4958:
4954:
4876:
4872:
4854:
4832:
4828:
4779:
4775:
4719:
4715:
4678:Stevens, Martin
4675:
4671:
4664:
4644:. Vol. 1.
4633:
4629:
4584:
4580:
4573:
4553:. Vol. 1.
4542:
4538:
4493:Current Biology
4485:
4481:
4428:
4424:
4389:
4385:
4373:
4371:
4362:
4361:
4340:
4336:
4275:
4271:
4225:Current Biology
4216:
4212:
4165:
4161:
4140:
4136:
4125:
4121:
4074:
4070:
4023:
4019:
3966:Stevens, Martin
3962:
3958:
3950:
3946:
3938:
3934:
3926:
3922:
3914:
3910:
3889:
3885:
3879:Rothenberg 2011
3877:
3873:
3865:
3861:
3853:
3849:
3841:
3837:
3829:
3825:
3817:
3813:
3805:
3801:
3793:
3789:
3781:
3777:
3767:
3765:
3764:. Darwin Online
3755:
3751:
3743:
3739:
3731:
3724:
3716:
3712:
3707:Wayback Machine
3677:
3670:
3665:
3660:
3659:
3650:
3646:
3641:
3637:
3632:
3628:
3619:
3615:
3610:
3606:
3592:
3588:
3583:
3576:
3569:
3560:
3553:
3544:
3534:
3518:
3506:Stephen Sprouse
3477:science fiction
3417:
3410:
3371:
3347:
3312:
3305:
3295:
3286:
3279:
3270:
3267:
3254:
3245:
3235:
3226:
3215:
3206:
3193:
3164:combat clothing
3124:
3118:
3111:
3108:snow camouflage
3097:
3088:
3081:
3072:
3065:
3056:
3047:
3038:
3026:
3017:
3010:
2944:Geoffrey Barkas
2925:General Katukov
2921:Battle of Kursk
2886:
2872:
2865:
2862:
2853:
2847:
2838:
2831:
2807:Ship camouflage
2752:First World War
2732:
2726:
2724:First World War
2714:Second Boer War
2693:Corps of Guides
2631:
2622:
2582:
2568:) wrote in his
2565:
2544:
2539:
2531:Main articles:
2529:
2524:
2472:
2465:
2458:
2451:
2442:
2436:
2427:
2417:
2408:
2398:
2389:
2385:by mimicking a
2375:
2366:
2359:
2350:
2343:
2334:
2322:
2313:
2307:
2235:
2225:
2215:has driven the
2176:
2170:
2168:Ultra-blackness
2151:
2147:
2146:
2102:Clupea harengus
2093:
2034:resemble small
1932:Aurelia labiata
1923:
1917:
1910:
1904:
1895:
1890:Bulwark of HMS
1888:
1879:
1866:
1846:B-24 Liberators
1826:bioluminescence
1791:
1785:
1778:
1772:
1763:
1760:
1751:
1744:
1735:
1729:grey reef shark
1725:
1716:
1706:
1630:
1624:
1617:
1611:
1602:
1592:
1583:
1573:
1564:
1561:
1552:
1545:
1536:
1533:
1491:snow camouflage
1451:
1450:
1449:
1448:
1447:
1444:
1436:
1435:
1428:
1402:idotea balthica
1390:
1388:Snow camouflage
1380:
1373:
1370:
1358:
1353:
1346:
1337:
1333:Syritta pipiens
1328:
1288:
1282:
1242:
1235:
1232:
1223:
1215:
1206:
1199:
1190:
1183:
1129:
1127:Self-decoration
1123:
1121:Self-decoration
1106:
1100:
1093:
1090:
1081:
1074:
1065:
1058:
1049:
1046:
1037:
1030:
1009:Pararge aegeria
984:
975:
971:Smilax bona-nox
962:
953:
947:
938:
931:
922:
916:
907:
904:
895:
892:
819:
813:
806:
799:
790:
784:
775:
769:
760:
757:
685:
668:
667:
666:
665:
664:
658:
650:
649:
641:
630:
624:
582:
509:
470:
457:
434:natural history
411:Teddy Roosevelt
216:
213:
163:
127:First World War
24:
17:
12:
11:
5:
13418:
13408:
13407:
13402:
13397:
13392:
13387:
13382:
13377:
13360:
13359:
13357:
13356:
13351:
13346:
13340:
13338:
13334:
13333:
13331:
13330:
13329:
13328:
13316:
13311:
13310:
13309:
13297:
13296:
13295:
13283:
13281:Wilson Bentley
13278:
13276:Joseph Plateau
13273:
13268:
13263:
13262:
13261:
13249:
13244:
13239:
13233:
13231:
13227:
13226:
13224:
13223:
13222:
13221:
13216:
13214:Plateau's laws
13211:
13209:Fluid dynamics
13206:
13196:
13195:
13194:
13189:
13184:
13174:
13173:
13172:
13167:
13162:
13157:
13147:
13141:
13139:
13135:
13134:
13127:
13125:
13123:
13122:
13117:
13112:
13107:
13102:
13101:
13100:
13095:
13090:
13085:
13075:
13070:
13065:
13060:
13055:
13050:
13045:
13039:
13037:
13033:
13032:
13025:
13024:
13017:
13010:
13002:
12993:
12992:
12990:
12989:
12984:
12979:
12974:
12969:
12967:
12966:
12965:
12960:
12955:
12953:Apparent death
12950:
12945:
12935:
12929:
12927:
12923:
12922:
12920:
12919:
12914:
12909:
12904:
12899:
12898:
12897:
12887:
12881:
12879:
12875:
12874:
12867:
12866:
12859:
12852:
12844:
12835:
12834:
12829:
12826:
12825:
12823:
12822:
12817:
12812:
12807:
12802:
12797:
12792:
12790:Microecosystem
12787:
12782:
12777:
12772:
12767:
12762:
12757:
12752:
12747:
12742:
12737:
12732:
12727:
12722:
12717:
12712:
12706:
12704:
12700:
12699:
12697:
12696:
12691:
12689:Thorson's rule
12686:
12681:
12676:
12671:
12666:
12661:
12656:
12651:
12646:
12641:
12636:
12631:
12626:
12621:
12616:
12614:Assembly rules
12610:
12608:
12600:
12599:
12597:
12596:
12591:
12586:
12581:
12576:
12571:
12570:
12569:
12559:
12554:
12549:
12544:
12539:
12533:
12531:
12525:
12524:
12522:
12521:
12516:
12511:
12499:
12497:Patch dynamics
12494:
12492:Metapopulation
12489:
12484:
12479:
12474:
12469:
12464:
12459:
12454:
12449:
12444:
12439:
12434:
12429:
12424:
12419:
12414:
12408:
12406:
12398:
12397:
12395:
12394:
12389:
12387:Storage effect
12384:
12379:
12374:
12369:
12364:
12359:
12354:
12349:
12343:
12341:
12335:
12334:
12332:
12331:
12326:
12321:
12316:
12311:
12306:
12301:
12296:
12291:
12286:
12281:
12276:
12271:
12269:Neutral theory
12266:
12261:
12256:
12254:Native species
12247:
12242:
12237:
12232:
12227:
12222:
12217:
12212:
12207:
12201:
12199:
12195:
12194:
12192:
12191:
12186:
12185:
12184:
12179:
12169:
12164:
12159:
12154:
12149:
12144:
12139:
12134:
12129:
12127:Overpopulation
12124:
12119:
12114:
12109:
12104:
12099:
12094:
12089:
12084:
12079:
12073:
12071:
12063:
12062:
12050:
12049:
12042:
12035:
12027:
12018:
12017:
12015:
12014:
12009:
12004:
11999:
11994:
11989:
11984:
11979:
11973:
11971:
11965:
11964:
11962:
11961:
11956:
11951:
11946:
11941:
11936:
11934:Nutrient cycle
11931:
11926:
11924:Feeding frenzy
11921:
11916:
11911:
11906:
11904:Energy quality
11901:
11896:
11891:
11886:
11881:
11876:
11871:
11866:
11864:Cascade effect
11861:
11856:
11850:
11848:
11844:
11843:
11841:
11840:
11839:
11838:
11833:
11828:
11823:
11818:
11813:
11808:
11798:
11793:
11788:
11783:
11777:
11775:
11771:
11770:
11768:
11767:
11762:
11757:
11752:
11747:
11742:
11736:
11734:
11728:
11727:
11725:
11724:
11719:
11714:
11709:
11707:Microbial loop
11704:
11699:
11694:
11689:
11684:
11679:
11674:
11672:Lithoautotroph
11669:
11664:
11658:
11656:
11654:Microorganisms
11650:
11649:
11647:
11646:
11641:
11636:
11631:
11625:
11623:
11617:
11616:
11614:
11613:
11611:Prey switching
11608:
11603:
11598:
11593:
11588:
11583:
11578:
11573:
11568:
11563:
11558:
11553:
11548:
11543:
11538:
11533:
11528:
11522:
11520:
11514:
11513:
11511:
11510:
11505:
11500:
11495:
11490:
11488:Photosynthesis
11485:
11480:
11475:
11470:
11465:
11460:
11455:
11450:
11445:
11443:Chemosynthesis
11440:
11434:
11432:
11426:
11425:
11423:
11422:
11417:
11412:
11407:
11402:
11397:
11392:
11387:
11382:
11377:
11372:
11367:
11362:
11357:
11352:
11347:
11342:
11337:
11335:Abiotic stress
11332:
11326:
11324:
11320:
11319:
11303:
11302:
11295:
11288:
11280:
11271:
11270:
11268:
11267:
11262:
11257:
11252:
11247:
11242:
11237:
11232:
11227:
11222:
11217:
11212:
11206:
11204:
11203:Related topics
11200:
11199:
11197:
11196:
11191:
11186:
11181:
11176:
11175:
11174:
11164:
11159:
11154:
11152:Countershading
11149:
11144:
11139:
11134:
11129:
11124:
11118:
11116:
11112:
11111:
11109:
11108:
11103:
11098:
11092:
11090:
11084:
11083:
11081:
11080:
11075:
11070:
11068:Holochroal eye
11065:
11064:
11063:
11058:
11048:
11047:
11046:
11036:
11031:
11026:
11021:
11016:
11010:
11008:
11002:
11001:
10994:
10992:
10990:
10989:
10988:
10987:
10982:
10977:
10967:
10962:
10957:
10952:
10947:
10941:
10939:
10933:
10932:
10927:
10926:
10919:
10912:
10904:
10895:
10894:
10892:
10891:
10886:
10881:
10876:
10871:
10866:
10861:
10856:
10851:
10846:
10841:
10836:
10834:Roland Penrose
10831:
10826:
10821:
10816:
10811:
10806:
10804:Frederick Gore
10801:
10799:Victorine Foot
10796:
10791:
10786:
10781:
10776:
10771:
10765:
10763:
10757:
10756:
10754:
10753:
10748:
10746:Everett Warner
10743:
10738:
10736:Leon Underwood
10733:
10728:
10723:
10718:
10713:
10708:
10703:
10698:
10693:
10688:
10683:
10678:
10673:
10667:
10665:
10659:
10658:
10651:
10650:
10643:
10636:
10628:
10619:
10618:
10616:
10615:
10614:
10613:
10608:
10598:
10590:
10588:
10584:
10583:
10580:
10579:
10576:
10575:
10573:
10572:
10566:
10560:
10553:
10551:
10547:
10546:
10544:
10543:
10538:
10532:
10530:
10523:
10519:
10518:
10515:
10514:
10512:
10511:
10505:
10499:
10493:
10487:
10481:
10475:
10469:
10463:
10457:
10451:
10445:
10439:
10433:
10427:
10416:
10414:
10408:
10407:
10405:
10404:
10398:
10392:
10386:
10383:wz. 93 Pantera
10380:
10374:
10368:
10362:
10356:
10350:
10344:
10338:
10332:
10326:
10320:
10314:
10308:
10302:
10296:
10290:
10284:
10278:
10272:
10266:
10260:
10254:
10248:
10241:
10239:
10233:
10232:
10230:
10229:
10223:
10217:
10210:
10208:
10204:
10203:
10200:
10199:
10197:
10196:
10191:
10185:
10179:
10173:
10167:
10160:
10158:
10154:
10153:
10151:
10150:
10144:
10138:
10132:
10126:
10120:
10117:Platanenmuster
10114:
10107:
10105:
10094:
10087:
10081:
10080:
10078:
10077:
10072:
10067:
10062:
10057:
10052:
10047:
10042:
10036:
10034:
10027:
10021:
10020:
10017:
10016:
10014:
10013:
10011:Martin Stevens
10008:
10003:
9998:
9993:
9987:
9985:
9981:
9980:
9978:
9977:
9972:
9967:
9966:
9965:
9953:
9948:
9946:Leon Underwood
9943:
9941:Everett Warner
9938:
9933:
9928:
9923:
9917:
9915:
9909:
9908:
9906:
9905:
9904:
9903:
9891:
9890:
9889:
9876:
9874:
9867:
9863:
9862:
9860:
9859:
9854:
9853:
9852:
9847:
9842:
9832:
9827:
9825:Decorator crab
9822:
9817:
9811:
9809:
9805:
9804:
9797:
9795:
9793:
9792:
9787:
9782:
9777:
9772:
9767:
9762:
9757:
9756:
9755:
9750:
9740:
9735:
9730:
9728:Countershading
9725:
9719:
9717:
9711:
9710:
9703:
9702:
9695:
9688:
9680:
9674:
9673:
9669:Behrens, Roy.
9667:
9659:
9658:External links
9656:
9655:
9654:
9639:
9622:
9619:
9618:
9617:
9599:
9581:
9572:. Zone Books.
9566:
9551:
9533:
9518:
9500:
9482:
9462:
9459:
9458:
9457:
9451:
9434:
9428:
9416:Casson, Lionel
9412:
9398:
9395:
9394:
9393:
9387:
9381:. Bloomsbury.
9367:
9361:
9349:Herring, Peter
9345:
9339:
9324:
9309:
9303:
9288:
9285:
9284:
9283:
9269:
9255:
9252:. John Murray.
9240:
9226:
9210:
9209:Early research
9207:
9205:
9202:
9200:
9197:
9194:
9193:
9168:
9161:
9141:
9134:
9114:
9088:
9066:
9036:
9021:
9019:, p. 100.
9006:
8972:
8946:
8925:
8913:
8906:
8888:
8876:
8848:
8833:Craemer, Guy.
8825:
8794:
8763:
8756:
8738:
8714:
8690:
8641:
8622:
8602:
8591:(9): 927–930.
8575:
8553:
8527:
8491:
8455:
8419:
8412:
8394:
8379:
8367:
8355:
8343:
8331:Farnham Castle
8327:"World War II"
8318:
8308:
8287:
8274:
8267:
8247:
8240:
8222:
8189:
8177:
8159:
8144:
8113:
8091:
8084:
8066:
8057:
8045:
8019:
7993:
7982:on 28 May 2013
7967:
7956:on 29 May 2013
7937:
7925:
7907:
7892:"Cubist Slugs"
7882:
7868:
7854:
7839:
7812:
7805:
7787:
7780:
7762:
7745:
7726:
7704:
7692:
7686:978-1841761770
7685:
7667:
7652:
7631:
7596:
7594:, p. 235.
7584:
7572:
7545:(3): 163–170.
7522:
7501:(5): 736–745.
7481:
7462:
7410:
7408:, p. 136.
7398:
7386:
7325:
7259:
7248:(4): 614–622.
7232:
7206:
7193:(3): 574–579.
7173:
7161:
7135:
7128:
7110:
7108:, p. 134.
7098:
7091:
7073:
7071:, p. 151.
7061:
7054:
7034:
6977:
6958:
6945:(2): 219–224.
6925:
6910:
6885:
6856:
6845:on 22 May 2013
6821:
6772:
6749:
6727:
6715:
6703:
6701:, p. 152.
6691:
6679:
6667:
6641:
6629:
6591:
6579:
6567:
6545:
6523:
6497:
6462:
6437:
6411:
6384:(2): 250–280.
6368:
6356:
6315:
6266:
6254:
6242:
6191:
6142:
6085:
6051:
6005:
5993:
5960:
5938:
5912:
5910:, p. 141.
5897:
5841:
5839:, p. 360.
5829:
5817:
5786:
5752:(3542): 1–18.
5733:
5721:
5672:
5620:
5601:
5586:
5579:
5556:
5529:(4): 463–474.
5513:
5487:978-4903313092
5486:
5468:
5441:(4): 483–489.
5425:
5347:
5316:
5290:
5274:
5267:
5249:
5193:
5157:
5145:
5096:
5089:
5071:
5054:
5042:
5030:
5018:
5006:
4952:
4934:Bitis arietans
4882:Bitis arietans
4870:
4852:
4826:
4773:
4713:
4692:(1): 213–222.
4669:
4662:
4627:
4578:
4571:
4536:
4499:(5): 448–453.
4479:
4442:(4): 792–806.
4422:
4383:
4374:|journal=
4334:
4269:
4210:
4159:
4134:
4119:
4068:
4017:
3956:
3944:
3932:
3920:
3908:
3883:
3871:
3859:
3847:
3835:
3833:, p. 122.
3823:
3811:
3799:
3787:
3775:
3749:
3747:, p. 111.
3737:
3722:
3710:
3667:
3666:
3664:
3661:
3658:
3657:
3644:
3635:
3626:
3613:
3604:
3593:A letter from
3585:
3584:
3582:
3579:
3578:
3577:
3570:
3563:
3561:
3554:
3547:
3545:
3519:
3512:
3408:
3391:Gertrude Stein
3370:
3367:
3346:
3343:
3311:
3308:
3307:
3306:
3296:
3289:
3287:
3281:Modern German
3280:
3273:
3271:
3255:
3248:
3246:
3236:
3229:
3227:
3223:special forces
3216:
3209:
3207:
3198:was the first
3194:
3187:
3117:
3114:
3113:
3112:
3098:
3091:
3089:
3082:
3075:
3073:
3066:
3059:
3057:
3048:
3041:
3039:
3027:
3020:
3018:
3011:
3004:
2982:William Dobell
2936:Farnham Castle
2932:Roland Penrose
2871:
2868:
2867:
2866:
2863:
2856:
2854:
2850:Siege howitzer
2848:
2841:
2839:
2832:
2825:
2725:
2722:
2705:William Hodson
2658:Napoleonic War
2630:
2627:
2543:
2540:
2528:
2525:
2523:
2520:
2464:
2461:
2460:
2459:
2452:
2445:
2443:
2437:
2430:
2428:
2418:
2411:
2409:
2399:
2392:
2390:
2376:
2369:
2367:
2360:
2353:
2351:
2344:
2337:
2335:
2333:orchid blossom
2323:
2316:
2314:
2308:
2301:
2291:brood parasite
2224:
2221:
2209:Actinopterygii
2169:
2166:
2135:aluminium foil
2092:
2089:
1916:
1913:
1912:
1911:
1905:
1898:
1896:
1889:
1882:
1880:
1867:
1860:
1818:midwater squid
1812:, such as the
1787:Main article:
1784:
1781:
1780:
1779:
1774:Countershaded
1773:
1766:
1764:
1761:
1754:
1752:
1745:
1738:
1736:
1727:Countershaded
1726:
1719:
1717:
1714:Gazella dorcas
1710:Dorcas gazelle
1708:Countershaded
1707:
1700:
1628:Countershading
1626:Main article:
1623:
1622:Countershading
1620:
1619:
1618:
1612:
1605:
1603:
1593:
1586:
1584:
1574:
1567:
1565:
1562:
1555:
1553:
1546:
1539:
1537:
1534:
1527:
1487:rock ptarmigan
1445:
1438:
1437:
1429:
1422:
1421:
1420:
1419:
1418:
1379:
1376:
1375:
1374:
1371:
1356:
1354:
1347:
1340:
1338:
1329:
1322:
1284:Main article:
1281:
1278:
1241:
1238:
1237:
1236:
1233:
1226:
1224:
1216:
1209:
1207:
1200:
1193:
1191:
1187:decorator crab
1184:
1177:
1138:decorator crab
1125:Main article:
1122:
1119:
1102:Main article:
1099:
1096:
1095:
1094:
1091:
1084:
1082:
1075:
1068:
1066:
1059:
1052:
1050:
1047:
1040:
1038:
1031:
1024:
996:horned lizards
983:
980:
977:
976:
963:
956:
954:
950:Jumping spider
948:
941:
939:
932:
925:
923:
917:
910:
908:
905:
898:
896:
893:
886:
843:edge detection
815:Main article:
812:
809:
808:
807:
800:
793:
791:
785:
778:
776:
770:
763:
761:
758:
751:
701:desert animals
684:
681:
659:
652:
651:
642:
635:
634:
633:
632:
631:
623:
620:
581:
578:
550:Peppered moths
508:
505:
501:countershading
496:Psittacosaurus
469:
468:Fossil history
466:
456:
453:
385:countershading
278:Experiment by
253:the colour of
225:Charles Darwin
211:
172:Octopus cyanea
162:
159:
77:countershading
15:
9:
6:
4:
3:
2:
13417:
13406:
13403:
13401:
13398:
13396:
13393:
13391:
13388:
13386:
13383:
13381:
13378:
13376:
13373:
13372:
13370:
13355:
13352:
13350:
13347:
13345:
13342:
13341:
13339:
13335:
13327:
13326:
13322:
13321:
13320:
13317:
13315:
13312:
13308:
13307:
13303:
13302:
13301:
13298:
13294:
13293:
13289:
13288:
13287:
13284:
13282:
13279:
13277:
13274:
13272:
13271:Ernst Haeckel
13269:
13267:
13266:Adolf Zeising
13264:
13260:
13259:
13255:
13254:
13253:
13250:
13248:
13245:
13243:
13240:
13238:
13235:
13234:
13232:
13228:
13220:
13217:
13215:
13212:
13210:
13207:
13205:
13202:
13201:
13200:
13197:
13193:
13190:
13188:
13185:
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13180:
13179:
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13175:
13171:
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13148:
13146:
13143:
13142:
13140:
13136:
13131:
13121:
13118:
13116:
13113:
13111:
13110:Vortex street
13108:
13106:
13103:
13099:
13096:
13094:
13091:
13089:
13088:Quasicrystals
13086:
13084:
13081:
13080:
13079:
13076:
13074:
13071:
13069:
13066:
13064:
13061:
13059:
13056:
13054:
13051:
13049:
13046:
13044:
13041:
13040:
13038:
13034:
13030:
13023:
13018:
13016:
13011:
13009:
13004:
13003:
13000:
12988:
12985:
12983:
12980:
12978:
12975:
12973:
12970:
12968:
12964:
12961:
12959:
12956:
12954:
12951:
12949:
12946:
12944:
12941:
12940:
12939:
12936:
12934:
12931:
12930:
12928:
12924:
12918:
12915:
12913:
12910:
12908:
12905:
12903:
12900:
12896:
12893:
12892:
12891:
12888:
12886:
12883:
12882:
12880:
12876:
12872:
12865:
12860:
12858:
12853:
12851:
12846:
12845:
12842:
12832:
12827:
12821:
12818:
12816:
12815:Urban ecology
12813:
12811:
12808:
12806:
12803:
12801:
12798:
12796:
12793:
12791:
12788:
12786:
12783:
12781:
12778:
12776:
12773:
12771:
12768:
12766:
12763:
12761:
12758:
12756:
12753:
12751:
12748:
12746:
12743:
12741:
12738:
12736:
12733:
12731:
12728:
12726:
12723:
12721:
12718:
12716:
12713:
12711:
12708:
12707:
12705:
12701:
12695:
12692:
12690:
12687:
12685:
12682:
12680:
12677:
12675:
12674:Kleiber's law
12672:
12670:
12667:
12665:
12662:
12660:
12657:
12655:
12652:
12650:
12647:
12645:
12642:
12640:
12637:
12635:
12632:
12630:
12627:
12625:
12622:
12620:
12617:
12615:
12612:
12611:
12609:
12607:
12601:
12595:
12592:
12590:
12587:
12585:
12582:
12580:
12577:
12575:
12572:
12568:
12565:
12564:
12563:
12560:
12558:
12555:
12553:
12550:
12548:
12545:
12543:
12540:
12538:
12535:
12534:
12532:
12530:
12526:
12520:
12517:
12515:
12512:
12510:
12508:
12504:
12500:
12498:
12495:
12493:
12490:
12488:
12485:
12483:
12480:
12478:
12475:
12473:
12470:
12468:
12465:
12463:
12460:
12458:
12455:
12453:
12450:
12448:
12447:Foster's rule
12445:
12443:
12440:
12438:
12435:
12433:
12430:
12428:
12425:
12423:
12420:
12418:
12415:
12413:
12410:
12409:
12407:
12405:
12399:
12393:
12390:
12388:
12385:
12383:
12380:
12378:
12375:
12373:
12370:
12368:
12365:
12363:
12360:
12358:
12355:
12353:
12350:
12348:
12345:
12344:
12342:
12336:
12330:
12327:
12325:
12322:
12320:
12317:
12315:
12312:
12310:
12307:
12305:
12302:
12300:
12297:
12295:
12292:
12290:
12287:
12285:
12282:
12280:
12277:
12275:
12272:
12270:
12267:
12265:
12262:
12260:
12257:
12255:
12251:
12248:
12246:
12243:
12241:
12238:
12236:
12233:
12231:
12228:
12226:
12223:
12221:
12218:
12216:
12213:
12211:
12208:
12206:
12203:
12202:
12200:
12196:
12190:
12187:
12183:
12180:
12178:
12175:
12174:
12173:
12170:
12168:
12165:
12163:
12160:
12158:
12155:
12153:
12150:
12148:
12145:
12143:
12140:
12138:
12135:
12133:
12130:
12128:
12125:
12123:
12120:
12118:
12115:
12113:
12110:
12108:
12105:
12103:
12100:
12098:
12095:
12093:
12090:
12088:
12085:
12083:
12080:
12078:
12075:
12074:
12072:
12070:
12064:
12059:
12055:
12048:
12043:
12041:
12036:
12034:
12029:
12028:
12025:
12013:
12010:
12008:
12005:
12003:
12000:
11998:
11995:
11993:
11990:
11988:
11985:
11983:
11980:
11978:
11975:
11974:
11972:
11966:
11960:
11957:
11955:
11952:
11950:
11947:
11945:
11942:
11940:
11937:
11935:
11932:
11930:
11927:
11925:
11922:
11920:
11917:
11915:
11912:
11910:
11907:
11905:
11902:
11900:
11897:
11895:
11892:
11890:
11887:
11885:
11882:
11880:
11877:
11875:
11872:
11870:
11867:
11865:
11862:
11860:
11857:
11855:
11852:
11851:
11849:
11845:
11837:
11834:
11832:
11829:
11827:
11824:
11822:
11819:
11817:
11814:
11812:
11809:
11807:
11804:
11803:
11802:
11799:
11797:
11794:
11792:
11789:
11787:
11784:
11782:
11779:
11778:
11776:
11772:
11766:
11765:Trophic level
11763:
11761:
11758:
11756:
11753:
11751:
11748:
11746:
11743:
11741:
11738:
11737:
11735:
11733:
11729:
11723:
11722:Phage ecology
11720:
11718:
11715:
11713:
11712:Microbial mat
11710:
11708:
11705:
11703:
11700:
11698:
11695:
11693:
11690:
11688:
11685:
11683:
11680:
11678:
11675:
11673:
11670:
11668:
11667:Bacteriophage
11665:
11663:
11660:
11659:
11657:
11655:
11651:
11645:
11642:
11640:
11637:
11635:
11634:Decomposition
11632:
11630:
11627:
11626:
11624:
11622:
11618:
11612:
11609:
11607:
11604:
11602:
11599:
11597:
11594:
11592:
11589:
11587:
11584:
11582:
11581:Mesopredators
11579:
11577:
11574:
11572:
11569:
11567:
11564:
11562:
11559:
11557:
11554:
11552:
11549:
11547:
11544:
11542:
11539:
11537:
11534:
11532:
11529:
11527:
11526:Apex predator
11524:
11523:
11521:
11519:
11515:
11509:
11506:
11504:
11501:
11499:
11496:
11494:
11491:
11489:
11486:
11484:
11481:
11479:
11476:
11474:
11471:
11469:
11466:
11464:
11461:
11459:
11456:
11454:
11451:
11449:
11446:
11444:
11441:
11439:
11436:
11435:
11433:
11431:
11427:
11421:
11418:
11416:
11413:
11411:
11408:
11406:
11403:
11401:
11398:
11396:
11393:
11391:
11388:
11386:
11383:
11381:
11378:
11376:
11373:
11371:
11368:
11366:
11363:
11361:
11360:Biotic stress
11358:
11356:
11353:
11351:
11348:
11346:
11343:
11341:
11338:
11336:
11333:
11331:
11328:
11327:
11325:
11321:
11316:
11312:
11308:
11301:
11296:
11294:
11289:
11287:
11282:
11281:
11278:
11266:
11263:
11261:
11258:
11256:
11253:
11251:
11248:
11246:
11243:
11241:
11238:
11236:
11233:
11231:
11228:
11226:
11223:
11221:
11218:
11216:
11213:
11211:
11210:Animal senses
11208:
11207:
11205:
11201:
11195:
11192:
11190:
11187:
11185:
11182:
11180:
11177:
11173:
11170:
11169:
11168:
11165:
11163:
11160:
11158:
11155:
11153:
11150:
11148:
11145:
11143:
11142:Chromatophore
11140:
11138:
11135:
11133:
11130:
11128:
11125:
11123:
11120:
11119:
11117:
11113:
11107:
11104:
11102:
11099:
11097:
11094:
11093:
11091:
11089:
11085:
11079:
11076:
11074:
11071:
11069:
11066:
11062:
11059:
11057:
11054:
11053:
11052:
11049:
11045:
11042:
11041:
11040:
11039:Mammalian eye
11037:
11035:
11032:
11030:
11027:
11025:
11022:
11020:
11017:
11015:
11014:Arthropod eye
11012:
11011:
11009:
11007:
11003:
10998:
10986:
10983:
10981:
10978:
10976:
10973:
10972:
10971:
10968:
10966:
10963:
10961:
10958:
10956:
10953:
10951:
10948:
10946:
10943:
10942:
10940:
10938:
10934:
10925:
10920:
10918:
10913:
10911:
10906:
10905:
10902:
10890:
10887:
10885:
10882:
10880:
10877:
10875:
10872:
10870:
10867:
10865:
10862:
10860:
10857:
10855:
10852:
10850:
10847:
10845:
10842:
10840:
10837:
10835:
10832:
10830:
10827:
10825:
10824:Oliver Messel
10822:
10820:
10817:
10815:
10812:
10810:
10807:
10805:
10802:
10800:
10797:
10795:
10792:
10790:
10787:
10785:
10782:
10780:
10777:
10775:
10772:
10770:
10767:
10766:
10764:
10762:
10758:
10752:
10749:
10747:
10744:
10742:
10739:
10737:
10734:
10732:
10729:
10727:
10724:
10722:
10719:
10717:
10714:
10712:
10709:
10707:
10706:Alister Hardy
10704:
10702:
10699:
10697:
10694:
10692:
10689:
10687:
10686:Loyd A. Jones
10684:
10682:
10681:Louis Guingot
10679:
10677:
10674:
10672:
10669:
10668:
10666:
10664:
10660:
10656:
10649:
10644:
10642:
10637:
10635:
10630:
10629:
10626:
10612:
10609:
10607:
10604:
10603:
10602:
10599:
10597:
10596:
10592:
10591:
10589:
10585:
10570:
10567:
10564:
10563:Yehudi lights
10561:
10558:
10555:
10554:
10552:
10548:
10542:
10539:
10537:
10534:
10533:
10531:
10527:
10524:
10520:
10509:
10506:
10503:
10500:
10497:
10494:
10491:
10488:
10485:
10482:
10479:
10476:
10473:
10470:
10467:
10464:
10461:
10458:
10455:
10452:
10449:
10446:
10443:
10440:
10437:
10434:
10431:
10428:
10425:
10421:
10418:
10417:
10415:
10409:
10402:
10399:
10396:
10393:
10390:
10387:
10384:
10381:
10378:
10375:
10372:
10369:
10366:
10363:
10360:
10357:
10354:
10351:
10348:
10345:
10342:
10339:
10336:
10333:
10330:
10327:
10324:
10321:
10318:
10315:
10312:
10309:
10306:
10303:
10300:
10297:
10294:
10291:
10288:
10285:
10282:
10279:
10276:
10273:
10270:
10267:
10264:
10261:
10258:
10255:
10252:
10249:
10246:
10243:
10242:
10240:
10234:
10227:
10224:
10221:
10218:
10215:
10212:
10211:
10209:
10205:
10195:
10192:
10189:
10186:
10183:
10182:Denison smock
10180:
10177:
10176:Telo mimetico
10174:
10171:
10168:
10165:
10162:
10161:
10159:
10155:
10148:
10145:
10142:
10139:
10136:
10133:
10130:
10127:
10124:
10121:
10118:
10115:
10112:
10109:
10108:
10106:
10104:
10098:
10095:
10091:
10088:
10086:
10082:
10076:
10073:
10071:
10068:
10066:
10063:
10061:
10058:
10056:
10053:
10051:
10048:
10046:
10043:
10041:
10038:
10037:
10035:
10031:
10028:
10026:
10022:
10012:
10009:
10007:
10004:
10002:
10001:Innes Cuthill
9999:
9997:
9994:
9992:
9989:
9988:
9986:
9982:
9976:
9973:
9971:
9968:
9964:
9963:
9959:
9958:
9957:
9954:
9952:
9949:
9947:
9944:
9942:
9939:
9937:
9934:
9932:
9929:
9927:
9924:
9922:
9919:
9918:
9916:
9914:
9910:
9902:
9901:
9897:
9896:
9895:
9892:
9888:
9887:
9883:
9882:
9881:
9878:
9877:
9875:
9871:
9868:
9864:
9858:
9855:
9851:
9848:
9846:
9843:
9841:
9838:
9837:
9836:
9833:
9831:
9830:Flower mantis
9828:
9826:
9823:
9821:
9818:
9816:
9813:
9812:
9810:
9806:
9801:
9791:
9788:
9786:
9783:
9781:
9778:
9776:
9773:
9771:
9768:
9766:
9763:
9761:
9758:
9754:
9751:
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9745:
9744:
9741:
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9716:
9712:
9708:
9701:
9696:
9694:
9689:
9687:
9682:
9681:
9678:
9672:
9668:
9665:
9662:
9661:
9652:
9648:
9644:
9640:
9637:
9633:
9629:
9625:
9624:
9615:
9611:
9607:
9603:
9600:
9597:
9593:
9589:
9585:
9582:
9579:
9575:
9571:
9567:
9564:
9560:
9556:
9552:
9549:
9545:
9541:
9537:
9534:
9531:
9527:
9523:
9519:
9516:
9512:
9508:
9504:
9501:
9498:
9494:
9490:
9486:
9483:
9480:
9479:0-9713244-0-9
9476:
9472:
9468:
9465:
9464:
9454:
9448:
9444:
9440:
9435:
9431:
9425:
9422:. JHU Press.
9421:
9417:
9413:
9409:
9405:
9401:
9400:
9390:
9384:
9379:
9378:
9372:
9368:
9364:
9358:
9354:
9350:
9346:
9342:
9336:
9332:
9331:
9325:
9322:
9318:
9314:
9310:
9306:
9300:
9296:
9291:
9290:
9280:
9279:
9274:
9270:
9266:
9265:
9260:
9256:
9251:
9250:
9245:
9241:
9237:
9236:
9231:
9230:Cott, Hugh B.
9227:
9223:
9222:
9217:
9213:
9212:
9182:
9178:
9172:
9164:
9158:
9154:
9153:
9145:
9137:
9131:
9128:. Ace Books.
9127:
9126:
9118:
9102:
9101:Arcadia, 1973
9098:
9092:
9076:
9070:
9054:
9050:
9046:
9040:
9032:
9025:
9018:
9013:
9011:
8994:
8990:
8986:
8982:
8976:
8961:. 12 May 2015
8960:
8956:
8950:
8942:
8938:
8937:
8929:
8922:
8917:
8909:
8903:
8899:
8892:
8886:, p. 38.
8885:
8880:
8865:
8864:
8859:
8852:
8836:
8829:
8813:
8809:
8805:
8798:
8782:
8778:
8774:
8767:
8759:
8753:
8749:
8742:
8726:
8725:
8718:
8702:
8701:
8694:
8683:
8679:
8675:
8671:
8667:
8663:
8659:
8652:
8645:
8629:
8625:
8619:
8615:
8614:
8606:
8598:
8594:
8590:
8586:
8579:
8563:
8557:
8541:
8537:
8531:
8512:
8508:
8501:
8495:
8476:
8472:
8465:
8459:
8440:
8436:
8429:
8423:
8415:
8409:
8405:
8398:
8390:
8383:
8376:
8371:
8364:
8359:
8352:
8347:
8332:
8328:
8322:
8315:
8311:
8305:
8301:
8297:
8291:
8284:
8278:
8270:
8264:
8260:
8259:
8251:
8243:
8237:
8233:
8226:
8207:
8200:
8193:
8186:
8181:
8173:
8169:
8163:
8155:
8148:
8132:
8128:
8124:
8117:
8109:
8102:
8100:
8098:
8096:
8087:
8081:
8077:
8070:
8061:
8054:
8049:
8033:
8029:
8023:
8007:
8003:
7997:
7981:
7977:
7971:
7955:
7951:
7947:
7941:
7934:
7929:
7921:
7917:
7911:
7903:
7899:
7898:
7893:
7886:
7878:
7872:
7864:
7858:
7850:
7843:
7835:
7831:
7827:
7823:
7816:
7808:
7802:
7798:
7791:
7783:
7777:
7773:
7766:
7758:
7757:
7749:
7741:
7737:
7730:
7714:
7708:
7702:, p. 43.
7701:
7696:
7688:
7682:
7678:
7671:
7663:
7656:
7648:
7644:
7643:
7635:
7627:
7623:
7619:
7615:
7612:(1): 89–102.
7611:
7607:
7600:
7593:
7588:
7581:
7576:
7568:
7564:
7560:
7556:
7552:
7548:
7544:
7540:
7533:
7526:
7518:
7514:
7509:
7504:
7500:
7496:
7492:
7485:
7477:
7473:
7466:
7458:
7454:
7449:
7444:
7440:
7436:
7432:
7428:
7424:
7420:
7414:
7407:
7402:
7396:, p. 94.
7395:
7390:
7382:
7378:
7373:
7368:
7363:
7358:
7354:
7350:
7346:
7342:
7338:
7332:
7330:
7321:
7317:
7312:
7307:
7302:
7297:
7293:
7289:
7286:(6): e20233.
7285:
7281:
7277:
7273:
7266:
7264:
7255:
7251:
7247:
7243:
7236:
7221:
7217:
7210:
7201:
7196:
7192:
7188:
7184:
7177:
7170:
7165:
7150:
7146:
7139:
7131:
7125:
7121:
7114:
7107:
7102:
7094:
7088:
7084:
7077:
7070:
7065:
7057:
7051:
7047:
7046:
7038:
7030:
7026:
7022:
7018:
7013:
7008:
7004:
7000:
6996:
6992:
6988:
6981:
6974:
6969:
6967:
6965:
6963:
6953:
6948:
6944:
6940:
6936:
6929:
6922:
6917:
6915:
6907:
6902:
6900:
6898:
6896:
6894:
6892:
6890:
6874:
6870:
6863:
6861:
6844:
6840:
6836:
6830:
6828:
6826:
6817:
6813:
6808:
6803:
6799:
6795:
6791:
6787:
6783:
6776:
6769:(4): 829–845.
6768:
6764:
6760:
6753:
6737:
6731:
6724:
6719:
6713:, p. 36.
6712:
6707:
6700:
6695:
6688:
6683:
6677:, p. 40.
6676:
6671:
6655:
6651:
6645:
6639:, p. 41.
6638:
6633:
6625:
6621:
6617:
6613:
6609:
6605:
6598:
6596:
6588:
6583:
6576:
6571:
6555:
6549:
6533:
6527:
6511:
6507:
6501:
6482:
6475:
6474:
6466:
6450:
6444:
6442:
6425:
6421:
6415:
6407:
6403:
6399:
6395:
6391:
6387:
6383:
6379:
6372:
6366:, p. 32.
6365:
6360:
6341:
6337:
6333:
6326:
6319:
6311:
6307:
6302:
6297:
6293:
6289:
6285:
6281:
6277:
6270:
6263:
6258:
6251:
6246:
6238:
6234:
6229:
6224:
6219:
6214:
6210:
6206:
6202:
6195:
6187:
6183:
6178:
6173:
6169:
6165:
6161:
6157:
6153:
6146:
6138:
6134:
6130:
6126:
6121:
6116:
6112:
6108:
6104:
6100:
6096:
6089:
6074:
6070:
6066:
6062:
6055:
6047:
6043:
6039:
6035:
6031:
6027:
6023:
6019:
6012:
6010:
6002:
5997:
5988:
5983:
5979:
5975:
5971:
5964:
5948:
5942:
5926:
5922:
5916:
5909:
5904:
5902:
5893:
5889:
5884:
5879:
5875:
5871:
5867:
5863:
5859:
5855:
5851:
5845:
5838:
5833:
5826:
5821:
5805:
5801:
5797:
5790:
5779:
5775:
5771:
5767:
5763:
5759:
5755:
5751:
5744:
5737:
5730:
5725:
5717:
5713:
5708:
5703:
5699:
5695:
5691:
5687:
5683:
5676:
5668:
5664:
5659:
5654:
5650:
5646:
5642:
5638:
5634:
5630:
5624:
5616:
5612:
5605:
5598:
5593:
5591:
5582:
5576:
5572:
5571:
5563:
5561:
5552:
5548:
5544:
5540:
5536:
5532:
5528:
5524:
5517:
5509:
5497:
5489:
5483:
5479:
5472:
5464:
5460:
5456:
5452:
5448:
5444:
5440:
5436:
5429:
5410:
5406:
5402:
5398:
5394:
5390:
5386:
5382:
5378:
5371:
5369:
5360:
5358:
5356:
5354:
5352:
5342:
5338:
5334:
5330:
5326:
5320:
5309:
5302:
5301:
5294:
5286:
5285:
5278:
5270:
5264:
5260:
5253:
5245:
5241:
5236:
5231:
5227:
5223:
5219:
5215:
5211:
5207:
5203:
5197:
5178:
5171:
5168:
5161:
5154:
5149:
5141:
5137:
5133:
5129:
5124:
5119:
5115:
5111:
5107:
5100:
5092:
5086:
5082:
5075:
5069:, p. 17.
5068:
5063:
5061:
5059:
5051:
5046:
5039:
5034:
5028:, p. 51.
5027:
5022:
5015:
5010:
5003:
4999:
4995:
4990:
4985:
4981:
4977:
4974:(7152): 408.
4973:
4969:
4968:
4963:
4956:
4949:
4947:
4943:
4939:
4935:
4929:
4925:
4920:
4915:
4911:
4907:
4902:
4897:
4893:
4889:
4885:
4883:
4874:
4867:
4863:
4859:
4855:
4849:
4845:
4841:
4837:
4830:
4822:
4818:
4813:
4808:
4804:
4800:
4796:
4792:
4788:
4784:
4777:
4769:
4765:
4760:
4755:
4751:
4747:
4743:
4739:
4735:
4731:
4727:
4723:
4717:
4709:
4705:
4700:
4695:
4691:
4687:
4683:
4679:
4673:
4665:
4659:
4655:
4651:
4647:
4643:
4642:
4637:
4631:
4623:
4619:
4614:
4609:
4605:
4601:
4597:
4593:
4589:
4582:
4574:
4568:
4564:
4560:
4556:
4552:
4551:
4546:
4540:
4532:
4528:
4524:
4520:
4515:
4510:
4506:
4502:
4498:
4494:
4490:
4483:
4475:
4471:
4466:
4461:
4457:
4453:
4449:
4445:
4441:
4437:
4433:
4426:
4418:
4414:
4410:
4406:
4402:
4398:
4394:
4387:
4379:
4366:
4358:
4354:
4350:
4346:
4338:
4330:
4326:
4322:
4318:
4314:
4310:
4306:
4302:
4298:
4294:
4290:
4286:
4285:
4280:
4273:
4265:
4261:
4257:
4253:
4248:
4243:
4239:
4235:
4231:
4227:
4226:
4221:
4214:
4206:
4202:
4197:
4192:
4187:
4182:
4178:
4174:
4170:
4163:
4155:
4151:
4150:
4145:
4138:
4130:
4123:
4115:
4111:
4107:
4103:
4099:
4095:
4091:
4087:
4083:
4079:
4072:
4064:
4060:
4056:
4052:
4048:
4044:
4040:
4036:
4032:
4028:
4021:
4013:
4009:
4004:
3999:
3995:
3991:
3987:
3983:
3979:
3975:
3971:
3967:
3960:
3953:
3948:
3941:
3936:
3929:
3924:
3917:
3912:
3904:
3900:
3899:
3894:
3887:
3880:
3875:
3868:
3863:
3857:, p. 77.
3856:
3851:
3844:
3839:
3832:
3827:
3820:
3815:
3809:, p. 87.
3808:
3803:
3797:, p. 83.
3796:
3791:
3784:
3779:
3763:
3759:
3753:
3746:
3741:
3735:, p. 84.
3734:
3729:
3727:
3719:
3714:
3708:
3704:
3701:
3697:
3693:
3689:
3685:
3681:
3675:
3673:
3668:
3654:
3648:
3639:
3630:
3623:
3617:
3608:
3601:
3596:
3590:
3586:
3574:
3567:
3562:
3558:
3551:
3546:
3542:
3541:
3529:
3526:
3522:
3516:
3511:
3510:
3509:
3507:
3503:
3498:
3496:
3492:
3488:
3487:shapeshifting
3484:
3483:
3478:
3474:
3471:
3468:
3467:
3462:
3459:
3455:
3451:
3447:
3442:
3440:
3433:
3428:
3426:
3425:
3414:
3407:
3402:
3400:
3399:Pablo Picasso
3396:
3393:recalled the
3392:
3388:
3384:
3375:
3366:
3362:
3360:
3351:
3342:
3340:
3336:
3332:
3325:
3321:
3316:
3304:
3300:
3293:
3288:
3284:
3277:
3272:
3262:
3259:
3252:
3247:
3243:
3240:
3233:
3228:
3224:
3220:
3213:
3208:
3204:
3201:
3197:
3191:
3186:
3185:
3184:
3182:
3178:
3174:
3169:
3165:
3161:
3156:
3154:
3150:
3146:
3142:
3138:
3137:near infrared
3133:
3129:
3123:
3109:
3105:
3101:
3095:
3090:
3086:
3079:
3074:
3070:
3063:
3058:
3054:
3053:
3045:
3040:
3036:
3035:
3031:
3024:
3019:
3015:
3008:
3003:
3002:
3001:
2999:
2995:
2991:
2987:
2983:
2979:
2975:
2971:
2967:
2963:
2959:
2958:
2953:
2949:
2945:
2941:
2937:
2933:
2928:
2926:
2922:
2918:
2917:
2912:
2908:
2904:
2900:
2899:countershaded
2895:
2891:
2885:
2881:
2877:
2860:
2855:
2851:
2845:
2840:
2836:
2829:
2824:
2823:
2822:
2820:
2816:
2812:
2808:
2804:
2801:
2797:
2793:
2790:term meaning
2789:
2785:
2781:
2777:
2773:
2769:
2768:
2763:
2762:
2757:
2753:
2745:
2741:
2736:
2731:
2721:
2717:
2715:
2711:
2706:
2702:
2698:
2694:
2690:
2686:
2682:
2677:
2675:
2671:
2667:
2663:
2659:
2655:
2651:
2643:
2639:
2635:
2626:
2617:
2613:
2609:
2604:
2600:
2596:
2592:
2591:Julius Caesar
2588:
2577:
2573:
2572:
2560:
2553:
2548:
2538:
2534:
2519:
2517:
2513:
2509:
2505:
2501:
2497:
2493:
2488:
2481:
2476:
2470:
2463:Motion dazzle
2456:
2449:
2444:
2441:
2434:
2429:
2425:
2422:adult mimics
2421:
2415:
2410:
2407:
2403:
2396:
2391:
2388:
2384:
2381:concealed in
2380:
2373:
2368:
2364:
2357:
2352:
2348:
2341:
2336:
2332:
2331:
2326:
2325:Flower mantis
2320:
2315:
2311:
2310:Peppered moth
2305:
2300:
2299:
2298:
2296:
2292:
2288:
2287:common cuckoo
2283:
2281:
2277:
2273:
2272:flower mantis
2269:
2265:
2260:
2258:
2257:
2252:
2251:peppered moth
2248:
2244:
2241:(also called
2240:
2234:
2230:
2220:
2218:
2214:
2210:
2206:
2202:
2198:
2194:
2193:
2184:
2180:
2175:
2165:
2163:
2157:
2145:spaced about
2144:
2140:
2136:
2132:
2127:
2125:
2121:
2111:
2104:
2103:
2097:
2088:
2086:
2085:
2080:
2079:
2074:
2069:
2065:
2061:
2057:
2053:
2049:
2045:
2041:
2037:
2033:
2032:
2031:Agalma okenii
2028:
2024:
2020:
2016:
2012:
2008:
2004:
2000:
1995:
1987:
1986:
1981:
1977:
1973:
1970:
1966:
1962:
1958:
1954:
1950:
1946:
1942:
1934:
1933:
1927:
1922:
1908:
1907:Yehudi Lights
1902:
1897:
1893:
1886:
1881:
1877:
1873:
1872:
1864:
1859:
1858:
1857:
1855:
1851:
1847:
1843:
1842:Yehudi lights
1838:
1834:
1829:
1827:
1823:
1819:
1815:
1814:firefly squid
1811:
1804:
1803:firefly squid
1800:
1797:Principle of
1795:
1790:
1777:
1770:
1765:
1758:
1753:
1749:
1742:
1737:
1734:
1730:
1723:
1718:
1715:
1711:
1704:
1699:
1698:
1697:
1693:
1691:
1687:
1682:
1679:
1674:
1672:
1668:
1664:
1660:
1656:
1652:
1648:
1644:
1634:
1629:
1615:
1609:
1604:
1600:
1596:
1590:
1585:
1581:
1578:
1571:
1566:
1559:
1554:
1550:
1543:
1538:
1531:
1526:
1525:
1524:
1522:
1521:Peltier plate
1518:
1514:
1509:
1508:
1502:
1500:
1496:
1492:
1488:
1484:
1480:
1476:
1471:
1467:
1465:
1460:
1456:
1442:
1433:
1426:
1417:
1415:
1411:
1407:
1406:chromatophore
1403:
1399:
1395:
1389:
1385:
1355:
1351:
1344:
1339:
1335:
1334:
1326:
1321:
1320:
1319:
1315:
1313:
1309:
1305:
1301:
1292:
1287:
1277:
1275:
1274:
1269:
1264:
1260:
1251:
1246:
1230:
1225:
1221:
1220:
1213:
1208:
1204:
1197:
1192:
1188:
1181:
1176:
1175:
1174:
1171:
1166:
1162:
1157:
1155:
1151:
1147:
1143:
1139:
1135:
1128:
1118:
1116:
1111:
1105:
1088:
1083:
1079:
1072:
1067:
1063:
1056:
1051:
1044:
1039:
1035:
1028:
1023:
1022:
1021:
1019:
1015:
1011:
1010:
1005:
1001:
997:
988:
979:
973:
972:
967:
960:
955:
951:
945:
940:
936:
929:
924:
920:
914:
909:
902:
897:
890:
885:
884:
883:
881:
876:
872:
868:
863:
861:
856:
851:
848:
844:
840:
836:
828:
823:
818:
804:
801:Bright green
797:
792:
788:
782:
777:
773:
767:
762:
755:
750:
749:
748:
746:
742:
738:
737:peppered moth
734:
730:
726:
722:
718:
714:
710:
706:
702:
698:
694:
690:
680:
678:
674:
662:
656:
647:
646:
639:
629:
619:
615:
613:
611:
606:
602:
598:
597:melanocrachia
596:
590:
587:
577:
574:
570:
566:
562:
557:
555:
551:
547:
543:
539:
535:
531:
527:
526:
521:
517:
512:
504:
502:
498:
497:
491:
487:
483:
479:
475:
465:
463:
452:
450:
446:
442:
437:
435:
430:
425:
424:
420:'s 1940 book
419:
414:
412:
408:
404:
400:
399:warning color
396:
392:
391:
386:
382:
374:
370:
366:
365:Abbott Thayer
362:
358:
356:
352:
348:
344:
343:flower mantis
340:
336:
332:
328:
324:
320:
316:
313:
309:
308:
303:
299:
292:
288:
285:
281:
276:
271:
269:
268:birds of prey
264:
260:
256:
252:
246:
244:
243:
238:
234:
230:
226:
222:
210:
208:
204:
198:
196:
195:
190:
186:
182:
174:
173:
167:
158:
155:
150:
148:
144:
140:
136:
132:
128:
124:
120:
116:
112:
110:
106:
102:
98:
94:
90:
86:
82:
78:
74:
70:
66:
62:
58:
54:
47:
42:
35:
30:
26:
22:
13323:
13304:
13290:
13256:
13182:Chaos theory
13159:
13105:Tessellation
12976:
12943:Alarm signal
12800:Regime shift
12785:Macroecology
12506:
12502:
12442:Edge effects
12412:Biogeography
12357:Commensalism
12205:Biodiversity
12082:Allee effect
11986:
11821:kelp forests
11774:Example webs
11639:Detritivores
11478:Organotrophs
11458:Kinetotrophs
11410:Productivity
11136:
11073:Parietal eye
11019:Compound eye
10874:Steven Sykes
10869:Basil Spence
10864:Alan Sorrell
10859:Edward Seago
10794:Hugh B. Cott
10761:World War II
10611:Invisibility
10593:
10371:Soldier 2000
10251:Tiger stripe
10194:Ghillie suit
10147:Leibermuster
10141:Erbsenmuster
10129:Palmenmuster
9960:
9898:
9884:
9722:
9706:
9653:. (ages 4–8)
9642:
9638:. (ages 4–8)
9627:
9621:For children
9605:
9587:
9569:
9554:
9539:
9536:Latimer, Jon
9521:
9506:
9488:
9470:
9438:
9419:
9407:
9376:
9352:
9329:
9312:
9294:
9277:
9263:
9248:
9234:
9220:
9199:Bibliography
9184:. Retrieved
9180:
9171:
9151:
9144:
9124:
9117:
9105:. Retrieved
9100:
9091:
9079:. Retrieved
9069:
9057:. Retrieved
9053:the original
9048:
9039:
9030:
9024:
8997:. Retrieved
8993:the original
8975:
8963:. Retrieved
8958:
8949:
8935:
8928:
8916:
8898:Wingshooting
8897:
8891:
8879:
8869:27 September
8867:. Retrieved
8861:
8851:
8841:27 September
8839:. Retrieved
8828:
8818:25 September
8816:. Retrieved
8812:the original
8807:
8797:
8787:27 September
8785:. Retrieved
8781:the original
8776:
8766:
8747:
8741:
8729:. Retrieved
8723:
8717:
8705:. Retrieved
8699:
8693:
8661:
8657:
8644:
8632:. Retrieved
8628:the original
8612:
8605:
8588:
8584:
8578:
8566:. Retrieved
8556:
8544:. Retrieved
8539:
8530:
8518:. Retrieved
8506:
8494:
8482:. Retrieved
8475:the original
8470:
8458:
8446:. Retrieved
8434:
8422:
8403:
8397:
8388:
8382:
8370:
8358:
8346:
8334:. Retrieved
8330:
8321:
8313:
8299:
8290:
8282:
8277:
8257:
8250:
8231:
8225:
8213:. Retrieved
8192:
8180:
8171:
8162:
8153:
8147:
8135:. Retrieved
8131:the original
8126:
8121:Prinzeugen.
8116:
8107:
8075:
8069:
8060:
8048:
8036:. Retrieved
8031:
8022:
8010:. Retrieved
8005:
8002:"Camouflage"
7996:
7984:. Retrieved
7980:the original
7970:
7958:. Retrieved
7954:the original
7949:
7940:
7928:
7922:(in French).
7919:
7910:
7904:(12): 16–20.
7901:
7895:
7885:
7876:
7871:
7862:
7857:
7848:
7842:
7821:
7815:
7796:
7790:
7771:
7765:
7755:
7748:
7739:
7735:
7729:
7717:. Retrieved
7707:
7695:
7676:
7670:
7661:
7655:
7642:The Crossbow
7641:
7634:
7609:
7605:
7599:
7587:
7575:
7542:
7538:
7525:
7498:
7494:
7484:
7475:
7471:
7465:
7430:
7426:
7413:
7401:
7389:
7352:
7348:
7283:
7279:
7245:
7241:
7235:
7223:. Retrieved
7219:
7209:
7190:
7186:
7176:
7164:
7152:. Retrieved
7148:
7138:
7119:
7113:
7101:
7082:
7076:
7064:
7044:
7037:
6994:
6990:
6980:
6973:Herring 2002
6942:
6938:
6928:
6923:, p. 6.
6906:Herring 2002
6876:. Retrieved
6872:
6847:. Retrieved
6843:the original
6838:
6789:
6785:
6775:
6766:
6762:
6752:
6740:. Retrieved
6730:
6718:
6706:
6694:
6682:
6670:
6658:. Retrieved
6644:
6632:
6610:(1): 21–23.
6607:
6603:
6582:
6570:
6558:. Retrieved
6548:
6536:. Retrieved
6526:
6514:. Retrieved
6509:
6500:
6488:. Retrieved
6481:the original
6472:
6465:
6453:. Retrieved
6428:. Retrieved
6424:the original
6414:
6381:
6377:
6371:
6359:
6347:. Retrieved
6335:
6331:
6318:
6286:(4): 326–9.
6283:
6279:
6269:
6257:
6245:
6208:
6205:PLOS Biology
6204:
6194:
6159:
6155:
6145:
6105:(423): 604.
6102:
6098:
6088:
6076:. Retrieved
6064:
6054:
6021:
6017:
5996:
5980:(1): 83–92.
5977:
5973:
5963:
5951:. Retrieved
5941:
5929:. Retrieved
5924:
5915:
5865:
5861:
5844:
5832:
5820:
5808:. Retrieved
5804:the original
5799:
5789:
5749:
5736:
5724:
5689:
5685:
5675:
5640:
5636:
5623:
5614:
5604:
5569:
5526:
5522:
5516:
5477:
5471:
5438:
5434:
5428:
5416:. Retrieved
5409:the original
5383:(1): 51–73.
5380:
5376:
5367:
5332:
5328:
5319:
5308:the original
5299:
5293:
5283:
5277:
5258:
5252:
5217:
5213:
5196:
5184:. Retrieved
5160:
5148:
5113:
5109:
5099:
5080:
5074:
5045:
5033:
5021:
5009:
5001:
4971:
4965:
4955:
4945:
4941:
4937:
4933:
4931:
4891:
4887:
4881:
4873:
4865:
4835:
4829:
4794:
4790:
4776:
4733:
4729:
4716:
4689:
4685:
4672:
4640:
4630:
4595:
4591:
4581:
4549:
4539:
4496:
4492:
4482:
4439:
4435:
4425:
4403:(1): 10–18.
4400:
4396:
4386:
4365:cite journal
4337:
4288:
4282:
4272:
4229:
4223:
4213:
4176:
4172:
4162:
4154:the original
4147:
4137:
4122:
4081:
4077:
4071:
4030:
4026:
4020:
3980:(1): 19966.
3977:
3973:
3959:
3947:
3935:
3923:
3911:
3905:(12): 16–20.
3902:
3896:
3886:
3874:
3862:
3850:
3838:
3831:Beddard 1892
3826:
3814:
3807:Beddard 1892
3802:
3795:Beddard 1892
3790:
3783:Poulton 1890
3778:
3766:. Retrieved
3752:
3745:Poulton 1890
3740:
3713:
3687:
3679:
3647:
3638:
3629:
3616:
3607:
3589:
3538:
3499:
3494:
3491:Joe Haldeman
3480:
3472:
3470:Latin phrase
3466:memento mori
3464:
3449:
3443:
3435:
3430:
3422:
3419:
3413:From Picasso
3412:
3404:
3380:
3363:
3359:South Africa
3356:
3331:Field sports
3328:
3324:field sports
3221:, issued to
3157:
3145:night vision
3125:
3051:
3032:
2962:Steven Sykes
2955:
2939:
2929:
2914:
2903:Soviet Union
2887:
2835:West Mahomet
2834:
2805:
2791:
2783:
2779:
2771:
2765:
2759:
2749:
2718:
2678:
2662:British Army
2647:
2607:
2594:
2569:
2559:Philostratus
2556:
2522:Applications
2504:tsetse flies
2489:
2485:
2454:
2440:reed warbler
2346:
2330:Phalaenopsis
2328:
2284:
2261:
2254:
2242:
2236:
2190:
2188:
2158:
2128:
2116:
2100:
2082:
2076:
2073:Amazon River
2068:comb jellies
2042:, including
2029:
2027:siphonophore
1991:
1983:
1938:
1930:
1915:Transparency
1891:
1870:
1830:
1807:
1732:
1713:
1694:
1688:, zoologist
1675:
1647:Thayer's Law
1646:
1642:
1639:
1598:
1505:
1503:
1472:
1468:
1455:melanophores
1452:
1391:
1331:
1316:
1297:
1271:
1255:
1217:
1203:Ghillie suit
1201:Sniper in a
1165:ghillie suit
1158:
1130:
1107:
1007:
993:
978:
969:
919:Gaboon viper
864:
852:
837:, as in the
832:
725:horned viper
686:
675:(scent) and
669:
643:
616:
608:
604:
593:
591:
586:microhabitat
583:
565:phaeomelanin
558:
546:endosymbiont
537:
533:
523:
513:
510:
494:
471:
458:
438:
421:
415:
406:
388:
378:
368:
305:
295:
259:black-grouse
248:
240:
218:
207:when alarmed
200:
192:
178:
170:
151:
113:
63:of a modern
52:
51:
25:
13300:Alan Turing
13258:Liber Abaci
13177:Mathematics
13083:in crystals
13073:Soap bubble
13068:Phyllotaxis
12987:Unkenreflex
12948:Aposematism
12437:Disturbance
12340:interaction
12162:Recruitment
12092:Depensation
11884:Copiotrophs
11755:Energy flow
11677:Lithotrophy
11621:Decomposers
11601:Planktivore
11576:Insectivore
11566:Heterotroph
11531:Bacterivore
11498:Phototrophs
11448:Chemotrophs
11420:Restoration
11370:Competition
11250:Pseudopupil
11132:Aposematism
11051:Mollusc eye
10854:Peter Scott
10839:Peter Proud
10784:John Codner
10779:Hugh Casson
10769:Tony Ayrton
10663:World War I
10655:Camoufleurs
10323:wz. 89 Puma
10275:wz. 68 Moro
10178:(1929 tent)
10135:Sumpfmuster
9991:Roy Behrens
9984:Researchers
9913:Camoufleurs
9081:14 November
9017:Forbes 2009
8921:Newark 2007
8884:Newark 2007
8802:Davies, W.
8777:AOL Defence
8750:. Crowood.
8634:9 September
8435:Smithsonian
8375:Forbes 2009
8363:Barkas 1952
8351:Forbes 2009
8185:Forbes 2009
8127:Prinz Eugen
8053:Forbes 2009
7933:Forbes 2009
7700:Newark 2007
7664:. ABC-CLIO.
7592:Casson 1995
7580:Casson 1995
7406:Thayer 1909
7349:BMC Biology
7169:Forbes 2009
7106:Forbes 2009
7069:Forbes 2009
7045:The Insects
6742:28 November
6711:Barkas 1952
6699:Forbes 2009
6687:Forbes 2009
6587:Forbes 2009
6560:14 November
6538:14 November
6455:22 December
6349:17 November
6338:(2): 1–12.
6262:Forbes 2009
6211:(5): e108.
5953:21 December
5931:14 November
5825:Forbes 2009
5729:Forbes 2009
5504:|work=
5050:Newark 2007
5026:Forbes 2009
4946:B. arietans
4598:(4): 1121.
4592:Coral Reefs
3952:Forbes 2009
3867:Thayer 1909
3855:Forbes 2009
3843:Thayer 1909
3733:Darwin 1859
3718:Darwin 1859
3540:camoufleurs
3535: 1917
3502:Andy Warhol
3427:announced:
3268: 2006
2994:John Vassos
2966:Tony Ayrton
2960:, included
2957:camoufleurs
2780:to disguise
2761:camoufleurs
2710:khaki drill
2670:rifle green
2654:Baker rifle
2642:Baker rifle
2587:Gallic Wars
2552:sarcophagus
2542:Before 1800
2424:sparrowhawk
2406:merchantman
2363:grasshopper
2295:sparrowhawk
2201:melanosomes
2162:cephalopods
2064:crustaceans
2023:nematocysts
1980:Glass frogs
1945:transparent
1822:photophores
1513:BAE Systems
1475:Arctic hare
1412:. However,
1312:loom larger
1304:dragonflies
1098:Distraction
1076:Camouflage
871:variegation
839:common frog
713:desert lark
601:stony coral
534:A. fischeri
516:cephalopods
379:The artist
371:depicted a
189:cephalopods
89:cephalopods
61:battledress
13375:Camouflage
13369:Categories
13247:Empedocles
13242:Pythagoras
13160:Camouflage
13098:in biology
13093:in flowers
13063:Parastichy
12977:Camouflage
12805:Sexecology
12382:Parasitism
12347:Antibiosis
12182:Resistance
12177:Resilience
12067:Population
11987:Camouflage
11939:Oligotroph
11854:Ascendency
11816:intertidal
11806:cold seeps
11760:Food chain
11561:Herbivores
11536:Carnivores
11463:Mixotrophs
11438:Autotrophs
11317:components
11240:Ommatidium
11172:coincident
11137:Camouflage
11115:Coloration
11056:cephalopod
10950:Chameleons
10849:Brian Robb
10844:Fred Pusey
10829:Colin Moss
10814:Ivan Konev
10711:André Mare
10701:Franz Marc
10550:Prototypes
10536:Berberys-R
10522:Technology
10353:Tropentarn
10220:Strichtarn
10093:Up to WWII
9850:Aggressive
9723:Camouflage
9707:Camouflage
9439:Camouflage
9410:. Cassell.
9238:. Methuen.
9125:Camouflage
9059:1 December
8999:31 January
8808:Newsletter
8464:"FM 21–75"
8336:8 February
8038:8 February
8012:8 February
7986:8 February
7960:8 February
7478:: 351–358.
6849:3 February
6723:Elias 2011
6660:1 February
6490:3 February
6430:31 January
6078:16 January
5186:25 October
4196:1893/32292
4179:: 639670.
3663:References
3600:T. W. Wood
3575:item, 2007
3521:André Mare
3482:Camouflage
3432:fantastic.
3200:pixellated
3116:After 1945
3069:Spitfire's
2974:Max Dupain
2916:Maskirovka
2767:camouflage
2744:André Mare
2500:horseflies
2455:Dolophones
2400:Armed WW1
2243:masquerade
2060:polychaete
2050:(floating
2003:crystallin
1965:planktonic
1848:and naval
1653:, such as
1643:vice versa
1580:jagdpanzer
1577:Marder III
1549:Winter War
1495:dog family
1479:Arctic fox
1459:organelles
1410:signalling
1300:hoverflies
1146:masked bug
966:understory
880:herbivores
875:understory
717:sandgrouse
709:fennec fox
622:Principles
478:Cretaceous
337:, and the
335:tree-frogs
257:, and the
251:red-grouse
185:signalling
131:André Mare
109:herbivores
97:chameleons
67:, and the
53:Camouflage
13385:Deception
13349:Emergence
13252:Fibonacci
12710:Allometry
12664:Emergence
12392:Symbiosis
12377:Mutualism
12172:Stability
12077:Abundance
11889:Dominance
11847:Processes
11836:tide pool
11732:Food webs
11606:Predation
11591:Omnivores
11518:Consumers
11473:Mycotroph
11430:Producers
11375:Ecosystem
11340:Behaviour
11255:Rhopalium
11088:Evolution
11061:gastropod
11029:Eye shine
11024:Eagle eye
10955:Dinosaurs
10696:Paul Klee
10347:Flecktarn
10236:Late 20th
10188:Frog Skin
9956:Hugh Cott
9845:Müllerian
9808:In nature
8985:"Picasso"
8731:5 October
8707:8 January
8678:144589400
8154:The Times
7834:1367-5907
7828:: 34–42.
7419:Caro, Tim
7394:Cott 1940
7225:22 August
7021:0960-9822
6921:Cott 1940
6675:Cott 1940
6637:Cott 1940
6575:Cott 1940
6364:Cott 1940
6250:Cott 1940
6046:131341953
6001:Cott 1940
5908:Cott 1940
5837:Cott 1940
5766:2246/5820
5597:Cott 1940
5506:ignored (
5496:cite book
5153:Cott 1940
5067:Cott 1940
5038:Cott 1940
5014:Cott 1940
4998:0028-0836
4910:0962-8452
4862:2197-7305
4750:0962-8452
4708:1465-7279
4456:0737-4038
4357:240967012
4313:0028-0836
4205:2296-7745
3940:Cott 1940
3928:Cott 1940
3916:Cott 1940
3819:Cott 1940
3500:In 1986,
3283:Flecktarn
3263:pattern,
3153:Lightning
3085:Luftwaffe
3034:Flecktarn
3030:Waffen SS
2784:camouflet
2772:camoufler
2593:sent his
2268:parasites
2264:predators
2192:Oneirodes
2091:Silvering
2052:tunicates
1957:mesogloea
1953:jellyfish
1686:Australia
1681:Hugh Cott
1678:zoologist
1394:chameleon
1154:trichomes
1134:caddisfly
1115:blue tits
935:ptarmigan
860:inherited
827:Hugh Cott
693:woodcocks
689:parakeets
673:olfactory
610:Platygyra
595:Phestilla
520:reflectin
490:mosasaurs
482:eumelanin
455:Evolution
441:predators
418:Hugh Cott
353:: "Among
239:. In his
214:Aristotle
181:Aristotle
121:with the
101:octopuses
13078:Symmetry
13036:Patterns
12895:examples
12765:Endolith
12694:Xerosere
12606:networks
12422:Ecocline
11968:Defense,
11644:Detritus
11546:Foraging
11415:Resource
11122:Albinism
10529:Deployed
10508:Xingkong
10430:MultiCam
10422:(2001) (
10207:Post-war
10131:(c 1941)
10085:Patterns
10025:Military
9996:Tim Caro
9840:Batesian
9604:(1968).
9538:(2001).
9487:(2009).
9469:(2002).
9418:(1995).
9406:(1952).
9373:(2011).
9351:(2002).
9275:(1909).
9261:(1890).
9246:(1859).
9232:(1940).
9218:(1892).
8983:(1939).
8682:Archived
8511:Archived
8439:Archived
8298:(1953).
8206:Archived
8170:(1969).
8110:: 67–81.
7822:Casemate
7567:24368147
7517:22323196
7457:18990666
7421:(2009).
7381:22117898
7355:: 9–81.
7320:21673797
7280:PLOS ONE
7220:BBC News
7029:32679102
6816:11079412
6624:21227055
6510:BBC News
6406:26566732
6340:Archived
6310:18492645
6237:16605303
6186:15129957
6137:52871328
6129:12789327
5892:26041868
5778:Archived
5716:19324754
5667:18990673
5463:12957121
5418:26 April
5341:2246/470
5244:16959632
5177:Archived
5140:16287514
5132:18395241
4928:26674950
4821:31937221
4768:16720401
4622:33882835
4531:19021807
4523:12620197
4474:29346646
4417:11837451
4321:27251284
4256:28889973
4106:24402224
4063:10809290
4055:16778047
4012:26822039
3768:29 March
3703:Archived
3700:Abstract
3409:—
3333:such as
3322:used in
3303:Gulf War
3217:British
3132:aircraft
3100:Red Army
3014:Catalina
2946:ran the
2911:doctrine
2907:Red Army
2817:devised
2776:Parisian
2742:painter
2697:Peshawar
2576:Vegetius
2571:Imagines
2527:Military
2496:Tim Caro
2036:copepods
2011:collagen
1850:Avengers
1816:and the
1748:Thayer's
1663:dolphins
1655:gazelles
1464:hormones
1163:wears a
855:giraffes
697:bitterns
677:acoustic
544:from an
507:Genetics
449:coursers
351:mackerel
300:studied
282:, 1890:
261:that of
212:—
147:aircraft
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13337:Related
13204:Crystal
13199:Physics
13187:Fractal
13165:Mimicry
13150:Biology
13058:Meander
12982:Mimicry
12972:Crypsis
12926:Signals
12755:Ecopath
12562:Habitat
12432:Ecotype
12427:Ecotone
12404:ecology
12402:Spatial
12338:Species
12198:Species
12069:ecology
12054:Ecology
12002:Mimicry
11970:counter
11914:f-ratio
11662:Archaea
11350:Biomass
11323:General
11315:Trophic
11307:Ecology
11184:Mimicry
11157:Crypsis
10970:Mammals
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10569:Adaptiv
10541:Nakidka
10466:Type 07
10426:(2002))
10413:century
10238:century
10170:Lozenge
9835:Mimicry
9820:Crypsis
9715:Methods
9557:. DPM.
9186:13 July
8965:28 July
8568:17 June
8546:17 June
8536:"5–103"
8520:16 June
8484:11 June
8448:16 June
8137:5 March
7824:(111).
7626:2563673
7547:Bibcode
7539:Zoology
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6807:1692851
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4329:3989607
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4234:Bibcode
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4035:Bibcode
4027:Science
4003:4731810
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3622:lichens
3573:fashion
3461:Arcadia
3458:Utopian
3450:Arcadia
3406:cubism.
3397:artist
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3261:Type 99
3244:pattern
3173:fractal
2894:protégé
2888:In the
2811:U-boats
2750:In the
2699:, when
2685:Federal
2679:In the
2640:firing
2603:Mochica
2589:, when
2280:Q-ships
2239:mimesis
2229:Mimicry
2223:Mimesis
2150:⁄
2143:guanine
2124:herring
2120:sardine
2044:radiata
2015:retinas
1994:muscles
1961:buoyant
1801:in the
1614:Adaptiv
1517:Adaptiv
1078:netting
847:leopard
803:katydid
612:carnosa
580:Ecology
445:plovers
395:mimicry
373:peacock
355:pelagic
331:iguanas
327:parrots
280:Poulton
255:heather
237:species
233:evolved
221:zoology
203:octopus
161:History
65:soldier
57:leopard
13230:People
13138:Causes
11786:Rivers
11682:Marine
10975:horses
10937:Vision
10571:(2011)
10565:(1943)
10559:(1941)
10510:(2019)
10504:(2019)
10498:(2015)
10492:(2015)
10490:HunCam
10486:(2014)
10480:(2010)
10474:(2008)
10468:(2007)
10462:(2007)
10456:(2007)
10450:(2006)
10448:ESTDCU
10444:(2004)
10438:(2004)
10432:(2002)
10420:MARPAT
10403:(1998)
10397:(1998)
10391:(1997)
10389:CADPAT
10385:(1993)
10379:(1993)
10377:TAZ 90
10373:(1993)
10367:(1991)
10361:(1990)
10355:(1990)
10349:(1990)
10343:(1990)
10337:(1990)
10331:(1989)
10325:(1989)
10319:(1987)
10313:(1984)
10307:(1984)
10301:(1983)
10299:TAZ 83
10295:(1982)
10289:(1981)
10283:(1981)
10277:(1969)
10271:(1969)
10265:(1967)
10259:(1965)
10253:(1962)
10247:(1958)
10245:Jigsaw
10228:(1968)
10222:(1960)
10216:(1947)
10214:Lizard
10190:(1942)
10184:(1941)
10166:(1915)
10149:(1945)
10143:(1944)
10137:(1943)
10125:(1939)
10119:(1937)
10113:(1931)
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3415:(1938)
3395:cubist
3196:CADPAT
3181:MARPAT
3177:CADPAT
3052:Duluth
2980:, and
2882:, and
2788:French
2746:, 1916
2740:Cubist
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2019:camera
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705:gerbil
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1939:Many
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9188:2024
9157:ISBN
9130:ISBN
9109:2012
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9061:2011
9001:2014
8967:2015
8902:ISBN
8871:2012
8843:2012
8820:2012
8789:2012
8752:ISBN
8733:2012
8709:2013
8658:IRRC
8636:2017
8618:ISBN
8570:2012
8548:2012
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8450:2012
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8338:2013
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7830:ISSN
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7721:2012
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6562:2015
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6394:PMID
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