31:
53:
4700:
1905:
1042:
occupying different positions within
Euryarchaeota. It is also debated whether the phylum Altiarchaeota should be classified in DPANN or Euryarchaeota. An alternative location for DPANN in the phylogenetic tree is as follows. The groups marked in quotes are lineages assigned to DPANN, but
3514:
Rinke C, Schwientek P, Sczyrba A, Ivanova NN, Anderson IJ, Cheng JF, Darling A, Malfatti S, Swan BK, Gies EA, Dodsworth JA, Hedlund BP, Tsiamis G, Sievert SM, Liu WT, Eisen JA, Hallam SJ, Kyrpides NC, Stepanauskas R, Rubin EM, Hugenholtz P, Woyke T (July 2013).
4260:
Waters E, Hohn MJ, Ahel I, Graham DE, Adams MD, Barnstead M, Beeson KY, Bibbs L, Bolanos R, Keller M, Kretz K, Lin X, Mathur E, Ni J, Podar M, Richardson T, Sutton GG, Simon M, Soll D, Stetter KO, Short JM, Noordewier M (October 2003).
342:
They are mostly anaerobic and have not been cultivated. They live in extreme environments such as thermophilic, hyperacidophilic, hyperhalophilic or metal-resistant; or also in the temperate environment of marine and lake
3116:
Rinke, Schwientek, Sczyrba, Ivanova, Anderson, Cheng, Darling, Malfatti, Swan, Gies, Dodsworth, Hedlund, Tsiamis, Sievert, Liu, Eisen, Hallam, Kyrpides, Stepanauskas, Rubin, Hugenholtz and Woyke 2024
4103:
Ortiz-Alvarez R, Casamayor EO (April 2016). "High occurrence of
Pacearchaeota and Woesearchaeota (Archaea superphylum DPANN) in the surface waters of oligotrophic high-altitude lakes".
4542:
426:
DPANN may be the first divergent clade of archaea according to some phylogenetic analyses. Recent phylogenetic analyses have found the following phylogeny between phyla.
3936:
Baker BJ, Tyson GW, Webb RI, Flanagan J, Hugenholtz P, Allen EE, Banfield JF (December 2006). "Lineages of acidophilic archaea revealed by community genomic analysis".
39:
3588:
Castelle CJ, Wrighton KC, Thomas BC, Hug LA, Brown CT, Wilkins MJ, Frischkorn KR, Tringe SG, Singh A, Markillie LM, Taylor RC, Williams KH, Banfield JF (March 2015).
4326:"Insights into archaeal evolution and symbiosis from the genomes of a nanoarchaeon and its inferred crenarchaeal host from Obsidian Pool, Yellowstone National Park"
2657:
4520:
1954:
3437:
3772:"Assessing biosynthetic potential of agricultural groundwater through metagenomic sequencing: A diverse anammox community dominates nitrate-rich groundwater"
2862:
2585:
3408:
2259:
262:. DPANN was also detected in nitrate-rich groundwater, on the water surface but not below, indicating that these taxa are still quite difficult to locate.
3370:
5100:
2938:
2142:
3419:
2698:
2333:
2087:
2015:
4988:
4732:
3397:
3390:
3149:
3123:
2955:
2609:
2530:
2316:
2276:
2181:
1448:
564:
321:
142:
107:
3989:"Metatranscriptomic analysis of microbes in an Oceanfront deep-subsurface hot spring reveals novel small RNAs and type-specific tRNA degradation"
2681:
2633:
2451:
3430:
3243:
2513:
1545:
1369:
683:
487:
290:
112:
97:
5590:
4546:
1958:
405:. They were the first discovered (in 2002) in a hydrothermal source next to the coast of Iceland. They live as symbionts of other archaea.
211:
were discovered and proposed within the DPANN superphylum. In 2017, another phylum
Altiarchaeota was placed into this superphylum. The
4594:
4203:
Narasingarao P, Podell S, Ugalde JA, Brochier-Armanet C, Emerson JB, Brocks JJ, Heidelberg KB, Banfield JF, Allen EE (January 2012).
387:. They have been identified both in sediments and in surface waters of aquifers and lakes, abounding especially in saline conditions.
238:
The DPANN groups together different phyla with a variety of environmental distribution and metabolism, ranging from symbiotic and
359:. Found by phylogenetic analysis of the genomes recovered from the groundwater filtration of a gold mine abandoned in the USA.
4725:
4044:
Baker BJ, Comolli LR, Dick GJ, Hauser LJ, Hyatt D, Dill BD, Land ML, Verberkmoes NC, Hettich RL, Banfield JF (May 2010).
1940:
5595:
304:
Limited metabolic capacities are a product of the small genome and are reflected in the fact that many lack central
4770:
4501:
Yutian Feng, Uri Neri, Sean
Gosselin, Artemis S Louyakis, R Thane Papke, Uri Gophna, Johann Peter Gogarten (2021).
4205:"De novo metagenomic assembly reveals abundant novel major lineage of Archaea in hypersaline microbial communities"
297:
or parasitic association with other organisms. Many of their characteristics are similar or analogous to those of
3770:
Ludington WB, Seher TD, Applegate O, Li X, Kliegman JI, Langelier C, Atwill ER, Harter T, DeRisi JL (2017-04-06).
4765:
4718:
3516:
3590:"Genomic expansion of domain archaea highlights roles for organisms from new phyla in anaerobic carbon cycling"
4849:
4839:
4822:
3705:"Multidomain ribosomal protein trees and the planctobacterial origin of neomura (Eukaryotes, archaebacteria)"
289:, they have limited but sufficient catabolic capacities to lead a free life, although many are thought to be
3875:
4502:
1915:
298:
4626:
4599:
4572:
169:
52:
5139:
4854:
4444:"Undinarchaeota illuminate DPANN phylogeny and the impact of gene transfer on archaeal evolution"
16:
A superphylum of
Archaea grouping taxa that display various environmental and metabolic features
5636:
5530:
3683:
3450:
220:
5631:
5608:
5568:
4817:
4704:
3639:"Genomic exploration of the diversity, ecology, and evolution of the archaeal domain of life"
3472:"Major New Microbial Groups Expand Diversity and Alter our Understanding of the Tree of Life"
4442:
Dombrowski N, Williams TA, Sun J, Woodcroft BJ, Lee JH, Minh BQ, et al. (August 2020).
4378:
Williams TA, Szöllősi GJ, Spang A, Foster PG, Heaps SE, Boussau B, et al. (June 2017).
328:
to meet their biological requirements. But those that have the potential to live freely are
5577:
4455:
4391:
4278:
4216:
4159:
4112:
4057:
4000:
3945:
3900:
3783:
3758:
Culture
Independent Genomic Comparisons Reveal Environmental Adaptations for Altiarchaeales
3601:
3531:
370:
3889:"Three-dimensional analysis of the structure and ecology of a novel, ultra-small archaeon"
3756:
Jordan T. Bird, Brett J. Baker, Alexander J. Probst, Mircea Podar, Karen G. Lloyd (2017).
8:
5641:
1830:
954:
4459:
4395:
4282:
4220:
4163:
4116:
4061:
4004:
3949:
3904:
3787:
3757:
3605:
3535:
5055:
4933:
4834:
4812:
4800:
4775:
4476:
4443:
4414:
4379:
4352:
4325:
4237:
4204:
4080:
4045:
4021:
3988:
3969:
3806:
3771:
3729:
3704:
3557:
47:
4710:
4301:
4262:
5603:
5510:
5446:
5326:
5227:
5150:
5073:
4844:
4807:
4795:
4785:
4677:
4669:
4649:
4481:
4419:
4357:
4306:
4242:
4185:
4180:
4171:
4147:
4128:
4085:
4026:
3961:
3918:
3858:
3850:
3830:
3811:
3734:
3660:
3619:
3549:
3493:
3202:
3107:
2879:
2165:
1815:
1294:
939:
608:
396:
224:
200:
127:
4324:
Podar M, Makarova KS, Graham DE, Wolf YI, Koonin EV, Reysenbach AL (December 2013).
3973:
5454:
5384:
4759:
4661:
4471:
4463:
4409:
4399:
4380:"Integrative modeling of gene and genome evolution roots the archaeal tree of life"
4347:
4337:
4296:
4286:
4232:
4224:
4175:
4167:
4120:
4075:
4065:
4016:
4008:
3953:
3908:
3842:
3801:
3791:
3724:
3716:
3650:
3609:
3561:
3539:
3483:
2993:
1718:
842:
333:
5553:
5368:
5312:
5266:
5175:
5050:
5035:
5017:
5002:
4945:
4928:
3796:
3226:
3175:
2978:
2126:
1693:
1470:
1271:
1143:
817:
586:
390:
255:
192:
92:
223:(LBA) where the lineages are grouped basally or artificially at the base of the
183:
DPANN is an acronym formed by the initials of the first five groups discovered,
5284:
5195:
5125:
4975:
4957:
4790:
4467:
4384:
Proceedings of the
National Academy of Sciences of the United States of America
4271:
Proceedings of the
National Academy of Sciences of the United States of America
4050:
Proceedings of the
National Academy of Sciences of the United States of America
3720:
3682:
Nina
Dombrowski, Jun-Hoe Lee, Tom A Williams, Pierre Offre, Anja Spang (2019).
3488:
3471:
3380:
3009:
2359:
1922:
1582:
1402:
1227:
720:
509:
380:
356:
259:
204:
184:
147:
102:
30:
3614:
3589:
417:
Tom A. Williams et al. 2017, Castelle et al. 2015 and Dombrowski et al. 2020.
5625:
5393:
5302:
5239:
5215:
4870:
4673:
4650:"Valid publication of names of two domains and seven kingdoms of prokaryotes"
4621:
3887:
Comolli LR, Baker BJ, Downing KH, Siegerist CE, Banfield JF (February 2009).
3854:
3831:"Valid publication of names of two domains and seven kingdoms of prokaryotes"
3354:
3294:
3084:
2562:
2490:
2229:
1775:
1760:
1738:
1567:
1514:
1499:
1417:
1079:
1035:
899:
884:
862:
805:
705:
652:
637:
524:
402:
384:
366:
362:
251:
243:
228:
208:
196:
188:
137:
132:
122:
117:
4516:
4404:
4291:
4124:
4070:
3957:
3655:
3638:
227:
without being related. These analyzes instead suggest that DPANN belongs to
5436:
5431:
5414:
5363:
5352:
5012:
4997:
4940:
4879:
4681:
4665:
4485:
4423:
4361:
4310:
4246:
4189:
4132:
4089:
4030:
3965:
3922:
3862:
3846:
3815:
3738:
3664:
3623:
3553:
3517:"Insights into the phylogeny and coding potential of microbial dark matter"
3497:
2808:
1249:
1180:
1114:
1039:
336:
329:
305:
239:
232:
4342:
4228:
4146:
Takai K, Moser DP, DeFlaun M, Onstott TC, Fredrickson JK (December 2001).
3913:
3888:
5562:
5292:
4967:
4952:
4567:
4012:
1310:
1165:
1092:
309:
161:
4202:
3544:
4741:
3684:
Genomic diversity, lifestyles and evolutionary origins of DPANN archaea
313:
1034:
Other phylogenetic analyzes have suggested that DPANN could belong to
1839:
963:
373:
from a US mine. They are of very small size and provisionally called
344:
294:
247:
212:
5524:
5547:
4829:
4699:
1054:
437:
393:. Found in wastewater from mines and in sediments from hot springs.
325:
216:
219:
rate of the included phyla, which can lead to the artifact of the
180:
due to their smaller size (nanometric) compared to other archaea.
5582:
4745:
4654:
International Journal of Systematic and Evolutionary Microbiology
4148:"Archaeal diversity in waters from deep South African gold mines"
3835:
International Journal of Systematic and Evolutionary Microbiology
1068:
450:
282:
165:
64:
4503:
The Evolutionary Origins of Extreme Halophilic Archaeal Lineages
4267:: insights into early archaeal evolution and derived parasitism"
281:
They are characterized by being small in size compared to other
317:
286:
4441:
3513:
374:
3987:
Murakami S, Fujishima K, Tomita M, Kanai A (February 2012).
3986:
3886:
347:. They are rarely found on the ground or in the open ocean.
215:
of DPANN is not yet considered established, due to the high
5275:
4377:
1967:
1706:
830:
420:
Jordan et al. 2017 Cavalier-Smith2020 and Feng et al 2021.
266:
3769:
3587:
168:
first proposed in 2013. Many members show novel signs of
4145:
4740:
4521:
List of Prokaryotic names with Standing in Nomenclature
4323:
1955:
List of Prokaryotic names with Standing in Nomenclature
4102:
4043:
3935:
4259:
3698:
3696:
3694:
3692:
377:(Archaeal Richmond Mine acidophilic nanoorganisms).
3702:
3636:
285:(nanometric size) and in keeping with their small
235:occupying various positions within Euryarchaeota.
4437:
4435:
4433:
3703:Cavalier-Smith, Thomas; Chao, Ema E-Yung (2020).
3689:
399:. Distributed in environments with high salinity.
265:Since the recognition of the kingdom rank by the
5623:
3469:
1953:The currently accepted taxonomy is based on the
4648:Göker, Markus; Oren, Aharon (22 January 2024).
3829:Göker, Markus; Oren, Aharon (22 January 2024).
4430:
3637:Spang A, Caceres EF, Ettema TJ (August 2017).
3583:
3581:
3579:
3577:
3575:
3573:
3571:
269:, the proposed name for this group is kingdom
172:from other domains of life. They are known as
5283:
4726:
4614:
4547:National Center for Biotechnology Information
4046:"Enigmatic, ultrasmall, uncultivated Archaea"
1959:National Center for Biotechnology Information
4196:
4139:
4096:
4037:
3980:
3929:
3880:
3678:
3676:
3674:
4373:
4371:
3568:
3509:
3507:
4733:
4719:
4253:
1043:phylogenetically separated from the rest.
29:
4647:
4475:
4413:
4403:
4351:
4341:
4300:
4290:
4236:
4179:
4079:
4069:
4020:
3912:
3828:
3805:
3795:
3728:
3671:
3654:
3613:
3543:
3487:
1941:Learn how and when to remove this message
4368:
3752:
3750:
3748:
3504:
4514:
4497:
4495:
5624:
4152:Applied and Environmental Microbiology
3993:Applied and Environmental Microbiology
5529:
5528:
4714:
4560:
3745:
4587:
4492:
1898:
320:; hence most DPANN archaea, such as
13:
4540:
4105:Environmental Microbiology Reports
3268:Phylum "Iainarchaeota" (DUSEL-3)
276:
14:
5653:
4692:
3470:Castelle CJ, Banfield JF (2018).
350:
4698:
4172:10.1128/AEM.67.21.5750-5760.2001
3231:corrig. Narasingarao et al. 2012
1903:
51:
4641:
4534:
4508:
4317:
3869:
3323:Order "Fermentimicrarchaeales"
3822:
3763:
3630:
3463:
1912:This section needs editing to
1:
3456:
3797:10.1371/journal.pone.0174930
3342:Order "Norongarragalinales"
409:
7:
3444:
3191:Order "Aenigmatarchaeales"
1978:
1894:
254:and non-extremophiles like
10:
5658:
4468:10.1038/s41467-020-17408-w
3721:10.1007/s00709-019-01442-7
3489:10.1016/j.cell.2018.02.016
3413:Vazquez-Campos et al. 2021
3365:Vazquez-Campos et al. 2021
3344:Vazquez-Campos et al. 2021
3338:Vazquez-Campos et al. 2021
3331:Vazquez-Campos et al. 2021
3318:Vazquez-Campos et al. 2021
3311:Vazquez-Campos et al. 2021
3305:Vazquez-Campos et al. 2021
3279:Probst & Banfield 2017
3260:corrig. Probst et al. 2014
3254:corrig. Probst et al. 2014
3213:corrig.La Cono et al. 2020
3160:corrig. Probst et al. 2019
5537:
5502:
5445:
5392:
5383:
5274:
5265:
5165:
5098:
4986:
4921:
4902:
4878:
4869:
4753:
3615:10.1016/j.cub.2015.01.014
3235:Order "Nanohalarchaeales"
3207:corrig. Rinke et al. 2013
3193:corrig. Rinke et al. 2020
3187:corrig. Rinke et al. 2020
3185:Class "Aenigmatarchaeia"
3180:corrig. Rinke et al. 2013
3007:
2990:
2983:
2952:
2935:
2928:
2920:
2876:
2859:
2852:
2844:
2822:
2805:
2798:
2791:
2784:
2695:
2678:
2671:
2654:
2647:
2630:
2623:
2606:
2599:
2582:
2575:
2567:
2527:
2510:
2503:
2495:
2483:
2448:
2440:
2432:
2425:
2418:
2356:
2330:
2313:
2306:
2299:
2273:
2256:
2249:
2242:
2234:
2222:
2178:
2170:
2139:
2131:
2119:
2084:
2076:
2068:
2061:
2054:
2047:
2012:
2004:
1996:
1989:
1836:
1813:
1806:
1773:
1758:
1751:
1736:
1729:
1722:
1704:
1697:
1580:
1565:
1558:
1543:
1536:
1512:
1497:
1490:
1483:
1468:
1461:
1446:
1439:
1415:
1400:
1393:
1386:
1366:
1359:
1308:
1291:
1284:
1269:
1262:
1247:
1240:
1225:
1218:
1211:
1178:
1163:
1156:
1141:
1134:
1127:
1112:
1105:
1090:
1083:
1073:
1051:
960:
937:
930:
897:
882:
875:
860:
853:
846:
828:
821:
803:
796:
718:
703:
696:
681:
674:
650:
635:
628:
621:
606:
599:
584:
577:
562:
555:
522:
507:
500:
485:
478:
471:
461:
454:
434:
419:
416:
89:
84:
48:Scientific classification
46:
37:
28:
23:
4627:Genome Taxonomy Database
4600:Genome Taxonomy Database
4573:Genome Taxonomy Database
4549:(NCBI) taxonomy database
3329:Order "Gugararchaeales"
3164:Order "Huberarchaeales"
3138:Order "Undinarchaeales"
1914:comply with Knowledge's
369:. Discovered in 2006 in
170:horizontal gene transfer
40:Parvarchaeum acidiphilum
5140:Methanomassiliicoccales
5065:Methanonatronarchaeales
4568:"GTDB release 08-RS214"
4405:10.1073/pnas.1618463114
4292:10.1073/pnas.1735403100
4125:10.1111/1758-2229.12370
4071:10.1073/pnas.0914470107
3958:10.1126/science.1132690
3876:Genomes Online Database
3656:10.1126/science.aaf3883
3336:Order "Micrarchaeales"
3283:Order "Iainarchaeales"
3217:Order "Nanohalobiales"
1038:or that it may even be
5062:Methanonatronarchaeia
4972:"Methanofastidiosales"
4666:10.1099/ijsem.0.006242
3847:10.1099/ijsem.0.006242
3451:List of Archaea genera
3316:Order "Burarchaeales"
3309:Order "Anstonellales"
3258:Order "Altarchaeales"
3158:Class "Huberarchaeia"
3140:Dombrowski et al. 2020
3134:Dombrowski et al. 2020
3132:Class "Undinarchaeia"
3128:Dombrowski et al. 2020
2658:Fermentimicrarchaeales
324:, which rely on other
221:long branch attraction
5344:"Methanomethylicales"
4912:"Hydrothermarchaeles"
4909:"Hydrothermarchaeia"
4903:"Hydrothermarchaeota"
4448:Nature Communications
4343:10.1186/1745-6150-8-9
4265:Nanoarchaeum equitans
4229:10.1038/ismej.2011.78
3914:10.1038/ismej.2008.99
3363:Class "Nanoarchaeia"
3303:Class "Micrarchaeia"
3299:Baker & Dick 2013
3277:Order "Forterreales"
3271:Class "Iainarchaeia"
3211:Class "Nanohalobiia"
1970:phylogeny of "DPANN"
5462:"Heimdallarchaeales"
5341:"Methanomethylicia"
5323:"Geothermarchaeales"
5144:"Natronoplasmatales"
5130:"Aciduliprofundales"
5089:"Syntropharchaeales"
4595:"ar53_r214.sp_label"
4541:Sayers; et al.
4013:10.1128/AEM.06811-11
3438:Wiannamattarchaeales
3325:Kadnikov et al. 2020
3252:Class "Altarchaeia"
2824:Methanobacteriota_B
371:acidic mine drainage
299:ultra-small bacteria
5509:Alternative views:
5112:"Thermoprofundales"
5083:"Syntropharchaeia"
4922:"Methanobacteriota"
4895:"Persephonarchaeia"
4460:2020NatCo..11.3939D
4396:2017PNAS..114E4602W
4390:(23): E4602–E4611.
4283:2003PNAS..10012984W
4277:(22): 12984–12988.
4221:2012ISMEJ...6...81N
4164:2001ApEnM..67.5750T
4117:2016EnvMR...8..210O
4062:2010PNAS..107.8806B
4005:2012ApEnM..78.1015M
3950:2006Sci...314.1933B
3944:(5807): 1933–1935.
3905:2009ISMEJ...3..159C
3788:2017PLoSO..1274930L
3606:2015CBio...25..690C
3545:10.1038/nature12352
3536:2013Natur.499..431R
3393:" (DHVE-5, DUSEL-1)
3219:La Cono et al. 2020
2926:"Neoeuryarchaeota"
2863:Hydrothermarchaeota
2586:Norongarragalinales
1923:improve the content
178:ultra-small archaea
5320:Conexivisphaerales
5181:"Aenigmatarchaeia"
5147:"Sysuiplasmatales"
5056:Methanomicrobiales
5045:"Methanoliparales"
4934:Methanobacteriales
4505:. Oxford Academic.
3649:(6351): eaaf3883.
3409:Tiddalikarchaeales
3247:Probst et al. 2018
3154:Probst et al. 2019
2260:Tiddalikarchaeales
5619:
5618:
5604:Open Tree of Life
5531:Taxon identifiers
5519:
5518:
5498:
5497:
5494:
5493:
5447:Heimdallarchaeota
5426:"Sigynarchaeales"
5379:
5378:
5347:"Nezhaarchaeales"
5327:Nitrososphaerales
5253:"Undinarchaeota"
5233:"Nanohalarchaeia"
5206:"Huberarchaeota"
5176:Aenigmatarchaeota
5161:
5160:
5151:Thermoplasmatales
5136:"Lunaplasmatales"
5133:"Gimiplasmatales"
5086:"Methanophagales"
5074:Methanosarcinales
5042:"Methanoliparia"
5007:"Mnemosynellales"
4158:(12): 5750–5760.
4056:(19): 8806–8811.
3530:(7459): 431–437.
3414:
3404:(ARMAN 4 & 5)
3403:
3402:Rinke et al. 2020
3386:
3385:Huber et al. 2011
3376:
3371:Jingweiarchaeales
3366:
3360:
3359:Huber et al. 2002
3345:
3339:
3332:
3326:
3319:
3312:
3306:
3300:
3286:
3285:Rinke et al. 2020
3280:
3274:
3273:Rinke et al. 2020
3261:
3255:
3248:
3232:
3220:
3214:
3208:
3194:
3188:
3181:
3176:Aenigmatarchaeota
3167:
3166:Rinke et al. 2020
3161:
3155:
3141:
3135:
3129:
3117:
3105:
3104:Rinke et al. 2013
3097:
3096:
3093:
3092:
3074:
3073:
3065:
3064:
3056:
3055:
3047:
3046:
3038:
3037:
3029:
3028:
3020:
3019:
2967:
2966:
2909:
2908:
2900:
2899:
2891:
2890:
2880:Methanobacteriota
2833:
2832:
2773:
2772:
2764:
2763:
2755:
2754:
2746:
2745:
2737:
2736:
2728:
2727:
2719:
2718:
2710:
2709:
2551:
2550:
2542:
2541:
2472:
2471:
2463:
2462:
2407:
2406:
2398:
2397:
2389:
2388:
2380:
2379:
2371:
2370:
2345:
2344:
2288:
2287:
2211:
2210:
2202:
2201:
2193:
2192:
2154:
2153:
2137:"Aenigmarchaeia"
2108:
2107:
2099:
2098:
2074:"Huberarchaeota"
2036:
2035:
2027:
2026:
2002:"Undinarchaeota"
1951:
1950:
1943:
1892:
1891:
1887:
1886:
1878:
1877:
1869:
1868:
1860:
1859:
1851:
1850:
1833:
1816:Heimdallarchaeota
1795:
1794:
1786:
1785:
1683:
1682:
1674:
1673:
1665:
1664:
1656:
1655:
1647:
1646:
1638:
1637:
1629:
1628:
1620:
1619:
1611:
1610:
1602:
1601:
1593:
1592:
1525:
1524:
1471:Aenigmatarchaeota
1428:
1427:
1348:
1347:
1339:
1338:
1330:
1329:
1321:
1320:
1295:Nanohaloarchaeota
1200:
1199:
1191:
1190:
1029:
1028:
1020:
1019:
1011:
1010:
1002:
1001:
993:
992:
984:
983:
975:
974:
957:
940:Heimdallarchaeota
919:
918:
910:
909:
785:
784:
776:
775:
767:
766:
758:
757:
749:
748:
740:
739:
731:
730:
663:
662:
609:Nanohaloarchaeota
587:Aenigmatarchaeota
544:
543:
535:
534:
293:that depend on a
225:phylogenetic tree
201:Nanohaloarchaeota
155:
154:
93:Aenigmatarchaeota
80:
79:Rinke et al. 2013
5649:
5612:
5611:
5599:
5598:
5586:
5585:
5573:
5572:
5571:
5558:
5557:
5556:
5526:
5525:
5487:"Wukongarchaeia"
5479:"Borrarchaeales"
5468:"Kariarchaeales"
5459:"Gerdarchaeales"
5455:Heimdallarchaeia
5423:"Lokiarchaeales"
5407:"Hermodarchaeia"
5390:
5389:
5360:"Marsarchaeales"
5336:"Culexarchaeles"
5333:"Culexarchaeia"
5317:"Caldarchaeales"
5281:
5280:
5272:
5271:
5228:Nanohalarchaeota
5101:Thermoplasmatota
5078:Methanotrichales
5070:Methanosarcinia
5032:"Methanocellia"
4964:"Theionarchaeia"
4876:
4875:
4782:
4762:
4735:
4728:
4721:
4712:
4711:
4703:Data related to
4702:
4686:
4685:
4645:
4639:
4638:
4636:
4634:
4618:
4612:
4611:
4609:
4607:
4591:
4585:
4584:
4582:
4580:
4564:
4558:
4557:
4555:
4554:
4538:
4532:
4531:
4529:
4528:
4512:
4506:
4499:
4490:
4489:
4479:
4439:
4428:
4427:
4417:
4407:
4375:
4366:
4365:
4355:
4345:
4321:
4315:
4314:
4304:
4294:
4257:
4251:
4250:
4240:
4209:The ISME Journal
4200:
4194:
4193:
4183:
4143:
4137:
4136:
4100:
4094:
4093:
4083:
4073:
4041:
4035:
4034:
4024:
3999:(4): 1015–1022.
3984:
3978:
3977:
3933:
3927:
3926:
3916:
3893:The ISME Journal
3884:
3878:
3873:
3867:
3866:
3826:
3820:
3819:
3809:
3799:
3767:
3761:
3754:
3743:
3742:
3732:
3700:
3687:
3680:
3669:
3668:
3658:
3634:
3628:
3627:
3617:
3585:
3566:
3565:
3547:
3521:
3511:
3502:
3501:
3491:
3482:(6): 1181–1197.
3467:
3412:
3401:
3384:
3374:
3364:
3358:
3343:
3337:
3330:
3324:
3317:
3310:
3304:
3298:
3284:
3278:
3272:
3259:
3253:
3246:
3230:
3218:
3212:
3206:
3203:Nanohalarchaeota
3192:
3186:
3179:
3165:
3159:
3153:
3139:
3133:
3127:
3115:
3103:
2986:
2985:
2939:Thermoplasmatota
2931:
2930:
2923:
2922:
2855:
2854:
2847:
2846:
2801:
2800:
2794:
2793:
2787:
2786:
2674:
2673:
2650:
2649:
2626:
2625:
2602:
2601:
2578:
2577:
2570:
2569:
2506:
2505:
2498:
2497:
2486:
2485:
2443:
2442:
2435:
2434:
2428:
2427:
2421:
2420:
2309:
2308:
2302:
2301:
2252:
2251:
2245:
2244:
2237:
2236:
2225:
2224:
2173:
2172:
2166:Nanohalarchaeota
2143:Aenigmarchaeales
2134:
2133:
2122:
2121:
2082:"Huberarchaeia"
2079:
2078:
2071:
2070:
2064:
2063:
2057:
2056:
2050:
2049:
2010:"Undinarchaeia"
2007:
2006:
1999:
1998:
1992:
1991:
1984:
1983:
1979:
1964:
1963:
1946:
1939:
1935:
1932:
1926:
1907:
1906:
1899:
1831:α─Proteobacteria
1828:
1809:
1808:
1754:
1753:
1732:
1731:
1725:
1724:
1700:
1699:
1561:
1560:
1539:
1538:
1493:
1492:
1486:
1485:
1464:
1463:
1442:
1441:
1396:
1395:
1389:
1388:
1362:
1361:
1287:
1286:
1265:
1264:
1243:
1242:
1221:
1220:
1214:
1213:
1159:
1158:
1137:
1136:
1130:
1129:
1108:
1107:
1086:
1085:
1076:
1075:
1047:
1046:
955:α─Proteobacteria
952:
933:
932:
878:
877:
856:
855:
849:
848:
824:
823:
799:
798:
699:
698:
677:
676:
631:
630:
624:
623:
602:
601:
580:
579:
558:
557:
503:
502:
481:
480:
474:
473:
464:
463:
457:
456:
430:
429:
414:
413:
397:Nanohalarchaeota
128:Nanohalarchaeota
78:
56:
55:
33:
21:
20:
5657:
5656:
5652:
5651:
5650:
5648:
5647:
5646:
5622:
5621:
5620:
5615:
5607:
5602:
5594:
5589:
5581:
5576:
5567:
5566:
5561:
5552:
5551:
5546:
5533:
5522:
5520:
5515:
5490:
5482:"Sifarchaeales"
5473:"Njordarchaeia"
5465:"Hodarchaeales"
5441:
5420:"Helarchaeales"
5404:"Freyrarchaeia"
5401:"Baldrarchaeia"
5375:
5369:Thermoproteales
5313:Nitrososphaeria
5297:"Korarchaeales"
5267:Proteoarchaeota
5261:
5256:"Undinarchaeia"
5209:"Huberarchaeia"
5186:"Altarchaeota"
5157:
5120:"Poseidoniales"
5094:
5051:Methanomicrobia
5036:Methanocellales
5027:"Hikarchaeales"
5018:Halobacteriales
5003:Archaeoglobales
4982:
4946:Methanococcales
4929:Methanobacteria
4917:
4898:
4890:"Hadarchaeales"
4865:
4864:
4780:
4758:
4749:
4739:
4695:
4690:
4689:
4646:
4642:
4632:
4630:
4622:"Taxon History"
4620:
4619:
4615:
4605:
4603:
4593:
4592:
4588:
4578:
4576:
4566:
4565:
4561:
4552:
4550:
4543:"Parvarchaeota"
4539:
4535:
4526:
4524:
4517:"Parvarchaeota"
4513:
4509:
4500:
4493:
4440:
4431:
4376:
4369:
4322:
4318:
4263:"The genome of
4258:
4254:
4201:
4197:
4144:
4140:
4101:
4097:
4042:
4038:
3985:
3981:
3934:
3930:
3885:
3881:
3874:
3870:
3827:
3823:
3782:(4): e0174930.
3768:
3764:
3755:
3746:
3701:
3690:
3681:
3672:
3635:
3631:
3594:Current Biology
3586:
3569:
3519:
3512:
3505:
3468:
3464:
3459:
3447:
3420:Woesearchaeales
3375:Rao et al. 2023
3227:Nanohalarchaeia
3182:(DSEG, DUSEL2)
3089:
3088:
3081:
3075:
3066:
3057:
3048:
3039:
3030:
3021:
2979:Proteoarchaeota
2968:
2910:
2901:
2892:
2834:
2774:
2765:
2756:
2747:
2738:
2729:
2720:
2711:
2699:Gugararchaeales
2573:"Micrarchaeia"
2552:
2543:
2501:"Iainarchaeia"
2473:
2464:
2438:"Altarchaeota"
2408:
2399:
2390:
2381:
2372:
2346:
2334:Woesearchaeales
2289:
2240:"Nanoarchaeia"
2212:
2203:
2194:
2176:"Nanohalobiia"
2155:
2127:Aenigmarchaeota
2109:
2100:
2088:Huberarchaeales
2037:
2028:
2016:Undinarchaeales
1976:
1947:
1936:
1930:
1927:
1920:
1916:Manual of Style
1908:
1904:
1897:
1888:
1879:
1870:
1861:
1852:
1796:
1787:
1694:Proteoarchaeota
1684:
1675:
1666:
1657:
1648:
1639:
1630:
1621:
1612:
1603:
1594:
1526:
1429:
1349:
1340:
1331:
1322:
1272:Methanomicrobia
1201:
1192:
1144:Methanobacteria
1030:
1021:
1012:
1003:
994:
985:
976:
920:
911:
818:Proteoarchaeota
786:
777:
768:
759:
750:
741:
732:
664:
545:
536:
412:
391:Aenigmarchaeota
353:
279:
277:Characteristics
256:Aenigmarchaeota
193:Aenigmarchaeota
77:
50:
17:
12:
11:
5:
5655:
5645:
5644:
5639:
5634:
5617:
5616:
5614:
5613:
5600:
5587:
5574:
5559:
5543:
5541:
5535:
5534:
5517:
5516:
5514:
5513:
5507:
5503:
5500:
5499:
5496:
5495:
5492:
5491:
5489:
5488:
5485:
5484:
5483:
5480:
5476:"Sifarchaeia"
5474:
5471:
5470:
5469:
5466:
5463:
5460:
5451:
5449:
5443:
5442:
5440:
5439:
5434:
5429:
5428:
5427:
5424:
5421:
5411:
5410:"Jordarchaeia"
5408:
5405:
5402:
5398:
5396:
5387:
5381:
5380:
5377:
5376:
5374:
5373:
5372:
5371:
5366:
5361:
5358:
5357:"Gearchaeales"
5350:
5349:
5348:
5345:
5339:
5338:
5337:
5331:
5330:
5329:
5324:
5321:
5318:
5310:
5309:
5308:
5307:"Hecatellales"
5300:
5299:
5298:
5289:
5287:
5285:Thermoproteota
5278:
5269:
5263:
5262:
5260:
5259:
5258:
5257:
5251:
5250:
5249:
5248:"Nanohalobiia"
5246:
5245:"Nanoarchaeia"
5236:
5235:
5234:
5224:
5223:
5222:
5221:"Micrarchaeia"
5212:
5211:
5210:
5204:
5203:
5202:
5201:"Iainarchaeia"
5192:
5191:
5190:
5184:
5183:
5182:
5171:
5169:
5163:
5162:
5159:
5158:
5156:
5155:
5154:
5153:
5148:
5145:
5142:
5137:
5134:
5131:
5126:Thermoplasmata
5123:
5122:
5121:
5117:"Poseidoniia"
5115:
5114:
5113:
5109:"Izemarchaea"
5106:
5104:
5096:
5095:
5093:
5092:
5091:
5090:
5087:
5081:
5080:
5079:
5076:
5068:
5067:
5066:
5060:
5059:
5058:
5048:
5047:
5046:
5040:
5039:
5038:
5030:
5029:
5028:
5024:"Hikarchaeia"
5022:
5021:
5020:
5010:
5009:
5008:
5005:
4994:
4992:
4989:Halobacteriota
4984:
4983:
4981:
4980:
4979:
4978:
4976:Thermococcales
4973:
4965:
4962:
4961:
4960:
4958:Methanopyrales
4950:
4949:
4948:
4938:
4937:
4936:
4925:
4923:
4919:
4918:
4916:
4915:
4914:
4913:
4906:
4904:
4900:
4899:
4897:
4896:
4893:
4892:
4891:
4887:"Hadarchaeia"
4884:
4882:
4873:
4867:
4866:
4863:
4862:
4861:
4860:
4859:
4858:
4855:Mesomycetozoea
4852:
4847:
4837:
4827:
4826:
4825:
4820:
4815:
4810:
4805:
4804:
4803:
4791:Diaphoretickes
4788:
4783:
4778:
4773:
4768:
4763:
4755:
4754:
4751:
4750:
4748:classification
4738:
4737:
4730:
4723:
4715:
4709:
4708:
4707:at Wikispecies
4694:
4693:External links
4691:
4688:
4687:
4640:
4613:
4586:
4559:
4533:
4507:
4491:
4429:
4367:
4330:Biology Direct
4316:
4252:
4195:
4138:
4111:(2): 210–217.
4095:
4036:
3979:
3928:
3899:(2): 159–167.
3879:
3868:
3821:
3762:
3744:
3715:(3): 621–753.
3688:
3670:
3629:
3600:(6): 690–701.
3567:
3503:
3461:
3460:
3458:
3455:
3454:
3453:
3446:
3443:
3442:
3441:
3434:
3429:Phylum ?"
3427:
3426:
3425:
3424:
3423:
3416:
3405:
3398:Parvarchaeales
3394:
3391:Pacearchaeales
3387:
3381:Nanoarchaeales
3377:
3351:
3350:
3349:
3348:
3347:
3340:
3334:
3327:
3321:
3314:
3291:
3290:
3289:
3288:
3287:
3281:
3266:
3265:
3264:
3263:
3262:
3240:
3239:
3238:
3237:
3236:
3223:
3222:
3221:
3199:
3198:
3197:
3196:
3195:
3172:
3171:
3170:
3169:
3168:
3150:Huberarchaeota
3146:
3145:
3144:
3143:
3142:
3124:Undinarchaeota
3095:
3094:
3091:
3090:
3082:
3079:
3078:
3076:
3072:
3071:
3068:
3067:
3063:
3062:
3059:
3058:
3054:
3053:
3050:
3049:
3045:
3044:
3041:
3040:
3036:
3035:
3032:
3031:
3027:
3026:
3023:
3022:
3018:
3017:
3014:
3013:
3010:Thermoproteota
3006:
3003:
3002:
2999:
2998:
2989:
2984:
2982:
2974:
2973:
2970:
2969:
2965:
2964:
2961:
2960:
2956:Halobacteriota
2951:
2948:
2947:
2944:
2943:
2934:
2929:
2927:
2921:
2919:
2916:
2915:
2912:
2911:
2907:
2906:
2903:
2902:
2898:
2897:
2894:
2893:
2889:
2888:
2885:
2884:
2875:
2872:
2871:
2868:
2867:
2858:
2853:
2851:
2850:"Methanomada"
2845:
2843:
2840:
2839:
2836:
2835:
2831:
2830:
2827:
2826:
2821:
2818:
2817:
2814:
2813:
2804:
2799:
2797:
2792:
2790:
2785:
2783:
2780:
2779:
2776:
2775:
2771:
2770:
2767:
2766:
2762:
2761:
2758:
2757:
2753:
2752:
2749:
2748:
2744:
2743:
2740:
2739:
2735:
2734:
2731:
2730:
2726:
2725:
2722:
2721:
2717:
2716:
2713:
2712:
2708:
2707:
2704:
2703:
2694:
2691:
2690:
2687:
2686:
2677:
2672:
2670:
2667:
2666:
2663:
2662:
2653:
2648:
2646:
2643:
2642:
2639:
2638:
2629:
2624:
2622:
2619:
2618:
2615:
2614:
2610:Micrarchaeales
2605:
2600:
2598:
2595:
2594:
2591:
2590:
2581:
2576:
2574:
2568:
2566:
2558:
2557:
2554:
2553:
2549:
2548:
2545:
2544:
2540:
2539:
2536:
2535:
2531:Iainarchaeales
2526:
2523:
2522:
2519:
2518:
2509:
2504:
2502:
2496:
2494:
2484:
2482:
2479:
2478:
2475:
2474:
2470:
2469:
2466:
2465:
2461:
2460:
2457:
2456:
2447:
2446:"Altarchaeia"
2441:
2439:
2433:
2431:
2426:
2424:
2419:
2417:
2414:
2413:
2410:
2409:
2405:
2404:
2401:
2400:
2396:
2395:
2392:
2391:
2387:
2386:
2383:
2382:
2378:
2377:
2374:
2373:
2369:
2368:
2365:
2364:
2360:Nanoarchaeales
2355:
2352:
2351:
2348:
2347:
2343:
2342:
2339:
2338:
2329:
2326:
2325:
2322:
2321:
2317:Pacearchaeales
2312:
2307:
2305:
2300:
2298:
2295:
2294:
2291:
2290:
2286:
2285:
2282:
2281:
2277:Parvarchaeales
2272:
2269:
2268:
2265:
2264:
2255:
2250:
2248:
2243:
2241:
2235:
2233:
2223:
2221:
2218:
2217:
2214:
2213:
2209:
2208:
2205:
2204:
2200:
2199:
2196:
2195:
2191:
2190:
2187:
2186:
2182:Nanohalobiales
2177:
2171:
2169:
2161:
2160:
2157:
2156:
2152:
2151:
2148:
2147:
2138:
2132:
2130:
2120:
2118:
2115:
2114:
2111:
2110:
2106:
2105:
2102:
2101:
2097:
2096:
2093:
2092:
2083:
2077:
2075:
2069:
2067:
2062:
2060:
2055:
2053:
2048:
2046:
2043:
2042:
2039:
2038:
2034:
2033:
2030:
2029:
2025:
2024:
2021:
2020:
2011:
2005:
2003:
1997:
1995:
1990:
1988:
1982:
1972:
1971:
1949:
1948:
1911:
1909:
1902:
1896:
1893:
1890:
1889:
1885:
1884:
1881:
1880:
1876:
1875:
1872:
1871:
1867:
1866:
1863:
1862:
1858:
1857:
1854:
1853:
1849:
1848:
1845:
1844:
1835:
1824:
1823:
1820:
1819:
1812:
1807:
1805:
1802:
1801:
1798:
1797:
1793:
1792:
1789:
1788:
1784:
1783:
1780:
1779:
1772:
1769:
1768:
1765:
1764:
1757:
1752:
1750:
1747:
1746:
1743:
1742:
1735:
1730:
1728:
1723:
1721:
1715:
1714:
1711:
1710:
1703:
1698:
1696:
1690:
1689:
1686:
1685:
1681:
1680:
1677:
1676:
1672:
1671:
1668:
1667:
1663:
1662:
1659:
1658:
1654:
1653:
1650:
1649:
1645:
1644:
1641:
1640:
1636:
1635:
1632:
1631:
1627:
1626:
1623:
1622:
1618:
1617:
1614:
1613:
1609:
1608:
1605:
1604:
1600:
1599:
1596:
1595:
1591:
1590:
1587:
1586:
1583:Woesearchaeota
1579:
1576:
1575:
1572:
1571:
1564:
1559:
1557:
1554:
1553:
1550:
1549:
1542:
1537:
1535:
1532:
1531:
1528:
1527:
1523:
1522:
1519:
1518:
1511:
1508:
1507:
1504:
1503:
1496:
1491:
1489:
1484:
1482:
1479:
1478:
1475:
1474:
1467:
1462:
1460:
1457:
1456:
1453:
1452:
1449:Undinarchaeota
1445:
1440:
1438:
1435:
1434:
1431:
1430:
1426:
1425:
1422:
1421:
1414:
1411:
1410:
1407:
1406:
1403:Diapherotrites
1399:
1394:
1392:
1387:
1385:
1379:
1378:
1375:
1374:
1365:
1360:
1358:
1355:
1354:
1351:
1350:
1346:
1345:
1342:
1341:
1337:
1336:
1333:
1332:
1328:
1327:
1324:
1323:
1319:
1318:
1315:
1314:
1307:
1304:
1303:
1300:
1299:
1290:
1285:
1283:
1280:
1279:
1276:
1275:
1268:
1263:
1261:
1258:
1257:
1254:
1253:
1246:
1241:
1239:
1236:
1235:
1232:
1231:
1228:Thermoplasmata
1224:
1219:
1217:
1212:
1210:
1207:
1206:
1203:
1202:
1198:
1197:
1194:
1193:
1189:
1188:
1185:
1184:
1177:
1174:
1173:
1170:
1169:
1162:
1157:
1155:
1152:
1151:
1148:
1147:
1140:
1135:
1133:
1128:
1126:
1123:
1122:
1119:
1118:
1111:
1106:
1104:
1101:
1100:
1097:
1096:
1089:
1084:
1082:
1074:
1072:
1064:
1063:
1060:
1059:
1050:
1045:
1031:
1027:
1026:
1023:
1022:
1018:
1017:
1014:
1013:
1009:
1008:
1005:
1004:
1000:
999:
996:
995:
991:
990:
987:
986:
982:
981:
978:
977:
973:
972:
969:
968:
959:
948:
947:
944:
943:
936:
931:
929:
926:
925:
922:
921:
917:
916:
913:
912:
908:
907:
904:
903:
896:
893:
892:
889:
888:
881:
876:
874:
871:
870:
867:
866:
859:
854:
852:
847:
845:
839:
838:
835:
834:
827:
822:
820:
814:
813:
810:
809:
802:
797:
795:
792:
791:
788:
787:
783:
782:
779:
778:
774:
773:
770:
769:
765:
764:
761:
760:
756:
755:
752:
751:
747:
746:
743:
742:
738:
737:
734:
733:
729:
728:
725:
724:
721:Woesearchaeota
717:
714:
713:
710:
709:
702:
697:
695:
692:
691:
688:
687:
680:
675:
673:
670:
669:
666:
665:
661:
660:
657:
656:
649:
646:
645:
642:
641:
634:
629:
627:
622:
620:
617:
616:
613:
612:
605:
600:
598:
595:
594:
591:
590:
583:
578:
576:
573:
572:
569:
568:
565:Undinarchaeota
561:
556:
554:
551:
550:
547:
546:
542:
541:
538:
537:
533:
532:
529:
528:
521:
518:
517:
514:
513:
510:Diapherotrites
506:
501:
499:
496:
495:
492:
491:
484:
479:
477:
472:
470:
462:
460:
455:
453:
447:
446:
443:
442:
433:
428:
422:
421:
418:
411:
408:
407:
406:
400:
394:
388:
381:Woesearchaeota
378:
360:
357:Diapherotrites
352:
351:Classification
349:
278:
275:
260:Diapherotrites
242:forms such as
205:Woesearchaeota
185:Diapherotrites
153:
152:
151:
150:
148:Woesearchaeota
145:
143:Undinarchaeota
140:
135:
130:
125:
120:
115:
110:
108:Huberarchaeota
105:
103:Diapherotrites
100:
95:
87:
86:
82:
81:
72:
68:
67:
62:
58:
57:
44:
43:
35:
34:
26:
25:
15:
9:
6:
4:
3:
2:
5654:
5643:
5640:
5638:
5637:Extremophiles
5635:
5633:
5630:
5629:
5627:
5610:
5605:
5601:
5597:
5592:
5588:
5584:
5579:
5575:
5570:
5564:
5560:
5555:
5549:
5545:
5544:
5542:
5540:
5536:
5532:
5527:
5523:
5512:
5508:
5505:
5504:
5501:
5486:
5481:
5478:
5477:
5475:
5472:
5467:
5464:
5461:
5458:
5457:
5456:
5453:
5452:
5450:
5448:
5444:
5438:
5435:
5433:
5430:
5425:
5422:
5419:
5418:
5416:
5412:
5409:
5406:
5403:
5400:
5399:
5397:
5395:
5394:Lokiarchaeota
5391:
5388:
5386:
5382:
5370:
5367:
5365:
5362:
5359:
5356:
5355:
5354:
5353:Thermoproteia
5351:
5346:
5343:
5342:
5340:
5335:
5334:
5332:
5328:
5325:
5322:
5319:
5316:
5315:
5314:
5311:
5306:
5305:
5304:
5303:Bathyarchaeia
5301:
5296:
5295:
5294:
5291:
5290:
5288:
5286:
5282:
5279:
5277:
5273:
5270:
5268:
5264:
5255:
5254:
5252:
5247:
5244:
5243:
5241:
5240:Nanoarchaeota
5237:
5232:
5231:
5229:
5225:
5220:
5219:
5217:
5216:Micrarchaeota
5213:
5208:
5207:
5205:
5200:
5199:
5197:
5196:Iainarchaeota
5193:
5189:"Altarchaeia"
5188:
5187:
5185:
5180:
5179:
5177:
5173:
5172:
5170:
5168:
5164:
5152:
5149:
5146:
5143:
5141:
5138:
5135:
5132:
5129:
5128:
5127:
5124:
5119:
5118:
5116:
5111:
5110:
5108:
5107:
5105:
5102:
5097:
5088:
5085:
5084:
5082:
5077:
5075:
5072:
5071:
5069:
5064:
5063:
5061:
5057:
5054:
5053:
5052:
5049:
5044:
5043:
5041:
5037:
5034:
5033:
5031:
5026:
5025:
5023:
5019:
5016:
5015:
5014:
5011:
5006:
5004:
5001:
5000:
4999:
4998:Archaeoglobia
4996:
4995:
4993:
4990:
4985:
4977:
4974:
4971:
4970:
4969:
4966:
4963:
4959:
4956:
4955:
4954:
4951:
4947:
4944:
4943:
4942:
4939:
4935:
4932:
4931:
4930:
4927:
4926:
4924:
4920:
4911:
4910:
4908:
4907:
4905:
4901:
4894:
4889:
4888:
4886:
4885:
4883:
4881:
4877:
4874:
4872:
4871:Euryarchaeota
4868:
4856:
4853:
4851:
4848:
4846:
4843:
4842:
4841:
4838:
4836:
4833:
4832:
4831:
4828:
4824:
4821:
4819:
4818:Stramenopiles
4816:
4814:
4811:
4809:
4806:
4802:
4799:
4798:
4797:
4794:
4793:
4792:
4789:
4787:
4784:
4781:(major groups
4779:
4777:
4774:
4772:
4769:
4767:
4764:
4761:
4757:
4756:
4752:
4747:
4743:
4736:
4731:
4729:
4724:
4722:
4717:
4716:
4713:
4706:
4701:
4697:
4696:
4683:
4679:
4675:
4671:
4667:
4663:
4659:
4655:
4651:
4644:
4629:
4628:
4623:
4617:
4602:
4601:
4596:
4590:
4575:
4574:
4569:
4563:
4548:
4544:
4537:
4522:
4518:
4515:J.P. Euzéby.
4511:
4504:
4498:
4496:
4487:
4483:
4478:
4473:
4469:
4465:
4461:
4457:
4453:
4449:
4445:
4438:
4436:
4434:
4425:
4421:
4416:
4411:
4406:
4401:
4397:
4393:
4389:
4385:
4381:
4374:
4372:
4363:
4359:
4354:
4349:
4344:
4339:
4335:
4331:
4327:
4320:
4312:
4308:
4303:
4298:
4293:
4288:
4284:
4280:
4276:
4272:
4268:
4266:
4256:
4248:
4244:
4239:
4234:
4230:
4226:
4222:
4218:
4214:
4210:
4206:
4199:
4191:
4187:
4182:
4177:
4173:
4169:
4165:
4161:
4157:
4153:
4149:
4142:
4134:
4130:
4126:
4122:
4118:
4114:
4110:
4106:
4099:
4091:
4087:
4082:
4077:
4072:
4067:
4063:
4059:
4055:
4051:
4047:
4040:
4032:
4028:
4023:
4018:
4014:
4010:
4006:
4002:
3998:
3994:
3990:
3983:
3975:
3971:
3967:
3963:
3959:
3955:
3951:
3947:
3943:
3939:
3932:
3924:
3920:
3915:
3910:
3906:
3902:
3898:
3894:
3890:
3883:
3877:
3872:
3864:
3860:
3856:
3852:
3848:
3844:
3840:
3836:
3832:
3825:
3817:
3813:
3808:
3803:
3798:
3793:
3789:
3785:
3781:
3777:
3773:
3766:
3759:
3753:
3751:
3749:
3740:
3736:
3731:
3726:
3722:
3718:
3714:
3710:
3706:
3699:
3697:
3695:
3693:
3685:
3679:
3677:
3675:
3666:
3662:
3657:
3652:
3648:
3644:
3640:
3633:
3625:
3621:
3616:
3611:
3607:
3603:
3599:
3595:
3591:
3584:
3582:
3580:
3578:
3576:
3574:
3572:
3563:
3559:
3555:
3551:
3546:
3541:
3537:
3533:
3529:
3525:
3518:
3510:
3508:
3499:
3495:
3490:
3485:
3481:
3477:
3473:
3466:
3462:
3452:
3449:
3448:
3439:
3436:Order ?"
3435:
3432:
3428:
3421:
3417:
3410:
3406:
3399:
3395:
3392:
3388:
3382:
3378:
3372:
3368:
3367:
3362:
3361:
3356:
3355:Nanoarchaeota
3352:
3341:
3335:
3328:
3322:
3315:
3308:
3307:
3302:
3301:
3296:
3295:Micrarchaeota
3292:
3282:
3276:
3275:
3270:
3269:
3267:
3257:
3256:
3251:
3250:
3245:
3241:
3234:
3233:
3228:
3225:Class ?"
3224:
3216:
3215:
3210:
3209:
3204:
3200:
3190:
3189:
3184:
3183:
3177:
3173:
3163:
3162:
3157:
3156:
3151:
3147:
3137:
3136:
3131:
3130:
3125:
3121:
3120:
3119:
3114:
3113:
3109:
3102:
3086:
3085:Euryarchaeota
3077:
3070:
3069:
3061:
3060:
3052:
3051:
3043:
3042:
3034:
3033:
3025:
3024:
3016:
3015:
3012:
3011:
3005:
3004:
3001:
3000:
2997:
2995:
2988:
2987:
2980:
2976:
2975:
2972:
2971:
2963:
2962:
2959:
2957:
2950:
2949:
2946:
2945:
2942:
2940:
2933:
2932:
2925:
2924:
2918:
2917:
2914:
2913:
2905:
2904:
2896:
2895:
2887:
2886:
2883:
2881:
2874:
2873:
2870:
2869:
2866:
2864:
2857:
2856:
2849:
2848:
2842:
2841:
2838:
2837:
2829:
2828:
2825:
2820:
2819:
2816:
2815:
2812:
2810:
2803:
2802:
2796:
2795:
2789:
2788:
2782:
2781:
2778:
2777:
2769:
2768:
2760:
2759:
2751:
2750:
2742:
2741:
2733:
2732:
2724:
2723:
2715:
2714:
2706:
2705:
2702:
2700:
2693:
2692:
2689:
2688:
2685:
2683:
2682:Burarchaeales
2676:
2675:
2669:
2668:
2665:
2664:
2661:
2659:
2652:
2651:
2645:
2644:
2641:
2640:
2637:
2635:
2634:Anstonellales
2628:
2627:
2621:
2620:
2617:
2616:
2613:
2611:
2604:
2603:
2597:
2596:
2593:
2592:
2589:
2587:
2580:
2579:
2572:
2571:
2564:
2563:Micrarchaeota
2560:
2559:
2556:
2555:
2547:
2546:
2538:
2537:
2534:
2532:
2525:
2524:
2521:
2520:
2517:
2515:
2508:
2507:
2500:
2499:
2492:
2491:Iainarchaeota
2488:
2487:
2481:
2480:
2477:
2476:
2468:
2467:
2459:
2458:
2455:
2453:
2452:Altarchaeales
2445:
2444:
2437:
2436:
2430:
2429:
2423:
2422:
2416:
2415:
2412:
2411:
2403:
2402:
2394:
2393:
2385:
2384:
2376:
2375:
2367:
2366:
2363:
2361:
2354:
2353:
2350:
2349:
2341:
2340:
2337:
2335:
2328:
2327:
2324:
2323:
2320:
2318:
2311:
2310:
2304:
2303:
2297:
2296:
2293:
2292:
2284:
2283:
2280:
2278:
2271:
2270:
2267:
2266:
2263:
2261:
2254:
2253:
2247:
2246:
2239:
2238:
2231:
2230:Nanoarchaeota
2227:
2226:
2220:
2219:
2216:
2215:
2207:
2206:
2198:
2197:
2189:
2188:
2185:
2183:
2175:
2174:
2167:
2163:
2162:
2159:
2158:
2150:
2149:
2146:
2144:
2136:
2135:
2128:
2124:
2123:
2117:
2116:
2113:
2112:
2104:
2103:
2095:
2094:
2091:
2089:
2081:
2080:
2073:
2072:
2066:
2065:
2059:
2058:
2052:
2051:
2045:
2044:
2041:
2040:
2032:
2031:
2023:
2022:
2019:
2017:
2009:
2008:
2001:
2000:
1994:
1993:
1986:
1985:
1981:
1980:
1977:
1974:
1973:
1969:
1966:
1965:
1962:
1960:
1956:
1945:
1942:
1934:
1924:
1919:
1917:
1910:
1901:
1900:
1883:
1882:
1874:
1873:
1865:
1864:
1856:
1855:
1847:
1846:
1843:
1842:
1841:
1832:
1826:
1825:
1822:
1821:
1818:
1817:
1811:
1810:
1804:
1803:
1800:
1799:
1791:
1790:
1782:
1781:
1778:
1777:
1776:Thorarchaeota
1771:
1770:
1767:
1766:
1763:
1762:
1761:Odinarchaeota
1756:
1755:
1749:
1748:
1745:
1744:
1741:
1740:
1739:Lokiarchaeota
1734:
1733:
1727:
1726:
1720:
1717:
1716:
1713:
1712:
1709:
1708:
1702:
1701:
1695:
1692:
1691:
1688:
1687:
1679:
1678:
1670:
1669:
1661:
1660:
1652:
1651:
1643:
1642:
1634:
1633:
1625:
1624:
1616:
1615:
1607:
1606:
1598:
1597:
1589:
1588:
1585:
1584:
1578:
1577:
1574:
1573:
1570:
1569:
1568:Pacearchaeota
1563:
1562:
1556:
1555:
1552:
1551:
1548:
1547:
1541:
1540:
1534:
1533:
1530:
1529:
1521:
1520:
1517:
1516:
1515:Parvarchaeota
1510:
1509:
1506:
1505:
1502:
1501:
1500:Nanoarchaeota
1495:
1494:
1488:
1487:
1481:
1480:
1477:
1476:
1473:
1472:
1466:
1465:
1459:
1458:
1455:
1454:
1451:
1450:
1444:
1443:
1437:
1436:
1433:
1432:
1424:
1423:
1420:
1419:
1418:Micrarchaeota
1413:
1412:
1409:
1408:
1405:
1404:
1398:
1397:
1391:
1390:
1384:
1381:
1380:
1377:
1376:
1373:
1371:
1364:
1363:
1357:
1356:
1353:
1352:
1344:
1343:
1335:
1334:
1326:
1325:
1317:
1316:
1313:
1312:
1306:
1305:
1302:
1301:
1298:
1296:
1289:
1288:
1282:
1281:
1278:
1277:
1274:
1273:
1267:
1266:
1260:
1259:
1256:
1255:
1252:
1251:
1245:
1244:
1238:
1237:
1234:
1233:
1230:
1229:
1223:
1222:
1216:
1215:
1209:
1208:
1205:
1204:
1196:
1195:
1187:
1186:
1183:
1182:
1176:
1175:
1172:
1171:
1168:
1167:
1161:
1160:
1154:
1153:
1150:
1149:
1146:
1145:
1139:
1138:
1132:
1131:
1125:
1124:
1121:
1120:
1117:
1116:
1110:
1109:
1103:
1102:
1099:
1098:
1095:
1094:
1088:
1087:
1081:
1080:Euryarchaeota
1078:
1077:
1071:
1070:
1066:
1065:
1062:
1061:
1058:
1057:
1056:
1049:
1048:
1044:
1041:
1037:
1036:Euryarchaeota
1032:
1025:
1024:
1016:
1015:
1007:
1006:
998:
997:
989:
988:
980:
979:
971:
970:
967:
966:
965:
956:
950:
949:
946:
945:
942:
941:
935:
934:
928:
927:
924:
923:
915:
914:
906:
905:
902:
901:
900:Thorarchaeota
895:
894:
891:
890:
887:
886:
885:Odinarchaeota
880:
879:
873:
872:
869:
868:
865:
864:
863:Lokiarchaeota
858:
857:
851:
850:
844:
841:
840:
837:
836:
833:
832:
826:
825:
819:
816:
815:
812:
811:
808:
807:
806:Euryarchaeota
801:
800:
794:
793:
790:
789:
781:
780:
772:
771:
763:
762:
754:
753:
745:
744:
736:
735:
727:
726:
723:
722:
716:
715:
712:
711:
708:
707:
706:Pacearchaeota
701:
700:
694:
693:
690:
689:
686:
685:
679:
678:
672:
671:
668:
667:
659:
658:
655:
654:
653:Parvarchaeota
648:
647:
644:
643:
640:
639:
638:Nanoarchaeota
633:
632:
626:
625:
619:
618:
615:
614:
611:
610:
604:
603:
597:
596:
593:
592:
589:
588:
582:
581:
575:
574:
571:
570:
567:
566:
560:
559:
553:
552:
549:
548:
540:
539:
531:
530:
527:
526:
525:Micrarchaeota
520:
519:
516:
515:
512:
511:
505:
504:
498:
497:
494:
493:
490:
489:
483:
482:
476:
475:
469:
466:
465:
459:
458:
452:
449:
448:
445:
444:
441:
440:
439:
432:
431:
427:
424:
423:
415:
404:
403:Nanoarchaeota
401:
398:
395:
392:
389:
386:
385:Pacearchaeota
382:
379:
376:
372:
368:
367:Micrarchaeota
364:
363:Parvarchaeota
361:
358:
355:
354:
348:
346:
340:
338:
335:
331:
327:
323:
322:ARMAN archaea
319:
315:
311:
308:pathways for
307:
302:
301:(CPR group).
300:
296:
292:
288:
284:
274:
272:
268:
263:
261:
257:
253:
252:Parvarchaeota
249:
245:
244:Nanoarchaeota
241:
236:
234:
230:
229:Euryarchaeota
226:
222:
218:
214:
210:
209:Pacearchaeota
206:
202:
198:
197:Nanoarchaeota
194:
190:
189:Parvarchaeota
186:
181:
179:
175:
171:
167:
163:
159:
149:
146:
144:
141:
139:
138:Parvarchaeota
136:
134:
133:Pacearchaeota
131:
129:
126:
124:
123:Nanoarchaeota
121:
119:
118:Micrarchaeota
116:
114:
111:
109:
106:
104:
101:
99:
96:
94:
91:
90:
88:
83:
76:
73:
71:Superphylum:
70:
69:
66:
63:
60:
59:
54:
49:
45:
42:
41:
36:
32:
27:
22:
19:
5632:Archaea taxa
5538:
5521:
5437:Thorarchaeia
5432:Odinarchaeia
5415:Lokiarchaeia
5364:Sulfolobales
5166:
5013:Halobacteria
4941:Methanococci
4880:Hadarchaeota
4840:Opisthokonta
4657:
4653:
4643:
4631:. Retrieved
4625:
4616:
4604:. Retrieved
4598:
4589:
4577:. Retrieved
4571:
4562:
4551:. Retrieved
4536:
4525:. Retrieved
4510:
4451:
4447:
4387:
4383:
4333:
4329:
4319:
4274:
4270:
4264:
4255:
4215:(1): 81–93.
4212:
4208:
4198:
4155:
4151:
4141:
4108:
4104:
4098:
4053:
4049:
4039:
3996:
3992:
3982:
3941:
3937:
3931:
3896:
3892:
3882:
3871:
3838:
3834:
3824:
3779:
3775:
3765:
3760:. Frontiers.
3712:
3708:
3646:
3642:
3632:
3597:
3593:
3527:
3523:
3479:
3475:
3465:
3431:Mamarchaeota
3244:Altarchaeota
3112:Nanobdellati
3111:
3110:
3106:(= proposed
3100:
3099:Superphylum
3098:
3008:
2991:
2953:
2936:
2877:
2860:
2823:
2809:Hadarchaeota
2806:
2696:
2679:
2655:
2631:
2607:
2583:
2528:
2514:Forterreales
2511:
2449:
2357:
2331:
2314:
2274:
2257:
2179:
2140:
2085:
2013:
1975:
1952:
1937:
1928:
1921:Please help
1913:
1838:
1837:
1814:
1774:
1759:
1737:
1705:
1581:
1566:
1546:Mamarchaeota
1544:
1513:
1498:
1469:
1447:
1416:
1401:
1382:
1370:Altarchaeota
1367:
1309:
1292:
1270:
1250:Archaeoglobi
1248:
1226:
1181:Methanococci
1179:
1164:
1142:
1115:Hadesarchaea
1113:
1091:
1067:
1053:
1052:
1040:polyphyletic
1033:
962:
961:
938:
898:
883:
861:
829:
804:
719:
704:
684:Mamarchaeota
682:
651:
636:
607:
585:
563:
523:
508:
488:Altarchaeota
486:
467:
436:
435:
425:
341:
337:heterotrophs
330:fermentative
306:biosynthetic
303:
291:episymbionts
280:
271:Nanobdellati
270:
264:
240:thermophilic
237:
233:polyphyletic
182:
177:
173:
157:
156:
113:Mamarchaeota
98:Altarchaeota
74:
38:
18:
5569:DPANN group
5563:Wikispecies
5511:Wikispecies
5293:Korarchaeia
4968:Thermococci
4953:Methanopyri
4742:Prokaryotes
4705:DPANN group
4454:(1): 3939.
3709:Protoplasma
3415:(LFW-252_1)
3333:(LFWA-IIIa)
3320:(LFWA-IIIb)
3313:(LFWA-IIIc)
2994:Asgardaeota
1957:(LPSN) and
1931:August 2024
1311:Haloarchaea
1166:Methanopyri
1093:Thermococci
310:nucleotides
248:acidophiles
174:nanoarchaea
162:superphylum
5642:Superphyla
5626:Categories
4633:6 December
4579:6 December
4553:2021-03-20
4527:2021-06-27
3457:References
3422:" (DHVE-6)
314:aminoacids
5554:Q24862848
4835:Amoebozoa
4813:Alveolata
4801:Cryptista
4776:Eukaryota
4674:1466-5026
3855:1466-5026
3686:. Nature.
3346:(LFWA-II)
1840:Eukaryota
964:Eukaryota
410:Phylogeny
345:sediments
295:symbiotic
213:monophyly
5583:51375420
5548:Wikidata
4830:Amorphea
4808:Rhizaria
4796:Hacrobia
4786:Excavata
4771:Bacteria
4682:38252124
4486:32770105
4424:28533395
4362:23607440
4336:(1): 9.
4311:14566062
4247:21716304
4190:11722932
4133:26711582
4090:20421484
4031:22156430
3974:26033384
3966:17185602
3923:18946497
3863:38252124
3816:28384184
3776:PLOS ONE
3739:31900730
3665:28798101
3624:25702576
3554:23851394
3498:29522741
3445:See also
3353:Phylum "
3293:Phylum "
3201:Phylum "
3174:Phylum "
3148:Phylum "
3122:Phylum "
1961:(NCBI).
1895:Taxonomy
1055:Bacteria
438:Bacteria
326:microbes
217:mutation
203:. Later
61:Domain:
5609:5877687
5596:1783276
5506:Source:
4845:Animals
4766:Archaea
4746:Archaea
4477:7414124
4456:Bibcode
4415:5468678
4392:Bibcode
4353:3655853
4279:Bibcode
4238:3246234
4217:Bibcode
4160:Bibcode
4113:Bibcode
4081:2889320
4058:Bibcode
4022:3272989
4001:Bibcode
3946:Bibcode
3938:Science
3901:Bibcode
3807:5383146
3784:Bibcode
3730:7203096
3643:Science
3602:Bibcode
3562:4394530
3532:Bibcode
3418:Order "
3407:Order "
3396:Order "
3389:Order "
3379:Order "
3369:Order "
3242:Phylum
3108:kingdom
3101:"DPANN"
1069:Archaea
451:Archaea
334:aerobic
283:archaea
166:Archaea
65:Archaea
5385:Asgard
4823:Plants
4760:Domain
4680:
4672:
4606:10 May
4523:(LPSN)
4484:
4474:
4422:
4412:
4360:
4350:
4309:
4302:240731
4299:
4245:
4235:
4188:
4178:
4131:
4088:
4078:
4029:
4019:
3972:
3964:
3921:
3861:
3853:
3814:
3804:
3737:
3727:
3663:
3622:
3560:
3552:
3524:Nature
3496:
3249:(SM1)
1987:DPANN
1719:Asgard
843:Asgard
318:lipids
316:, and
287:genome
231:or is
85:Phyla
24:DPANN
5539:DPANN
5167:DPANN
4850:Fungi
4660:(1).
4181:93369
3970:S2CID
3841:(1).
3558:S2CID
3520:(PDF)
3080:DPANN
1383:DPANN
468:DPANN
375:ARMAN
250:like
160:is a
158:DPANN
75:DPANN
5591:NCBI
5276:TACK
4678:PMID
4670:ISSN
4635:2021
4608:2023
4581:2021
4482:PMID
4420:PMID
4358:PMID
4307:PMID
4243:PMID
4186:PMID
4129:PMID
4086:PMID
4027:PMID
3962:PMID
3919:PMID
3859:PMID
3851:ISSN
3812:PMID
3735:PMID
3661:PMID
3620:PMID
3550:PMID
3494:PMID
3476:Cell
1968:GTDB
1707:TACK
831:TACK
383:and
365:and
332:and
267:ICNP
258:and
207:and
199:and
5578:EoL
4662:doi
4472:PMC
4464:doi
4410:PMC
4400:doi
4388:114
4348:PMC
4338:doi
4297:PMC
4287:doi
4275:100
4233:PMC
4225:doi
4176:PMC
4168:doi
4121:doi
4076:PMC
4066:doi
4054:107
4017:PMC
4009:doi
3954:doi
3942:314
3909:doi
3843:doi
3802:PMC
3792:doi
3725:PMC
3717:doi
3713:257
3651:doi
3647:357
3610:doi
3540:doi
3528:499
3484:doi
3480:172
176:or
164:of
5628::
5606::
5593::
5580::
5565::
5550::
5417:"
5242:"
5230:"
5218:"
5198:"
5178:"
4744::
4676:.
4668:.
4658:74
4656:.
4652:.
4624:.
4597:.
4570:.
4545:.
4519:.
4494:^
4480:.
4470:.
4462:.
4452:11
4450:.
4446:.
4432:^
4418:.
4408:.
4398:.
4386:.
4382:.
4370:^
4356:.
4346:.
4332:.
4328:.
4305:.
4295:.
4285:.
4273:.
4269:.
4241:.
4231:.
4223:.
4211:.
4207:.
4184:.
4174:.
4166:.
4156:67
4154:.
4150:.
4127:.
4119:.
4107:.
4084:.
4074:.
4064:.
4052:.
4048:.
4025:.
4015:.
4007:.
3997:78
3995:.
3991:.
3968:.
3960:.
3952:.
3940:.
3917:.
3907:.
3895:.
3891:.
3857:.
3849:.
3839:74
3837:.
3833:.
3810:.
3800:.
3790:.
3780:12
3778:.
3774:.
3747:^
3733:.
3723:.
3711:.
3707:.
3691:^
3673:^
3659:.
3645:.
3641:.
3618:.
3608:.
3598:25
3596:.
3592:.
3570:^
3556:.
3548:.
3538:.
3526:.
3522:.
3506:^
3492:.
3478:.
3474:.
3411:"
3400:"
3383:"
3373:"
3357:"
3297:"
3229:"
3205:"
3178:"
3152:"
3126:"
3118:)
2996:"
2981:"
2958:"
2941:"
2882:"
2865:"
2811:"
2701:"
2684:"
2660:"
2636:"
2612:"
2588:"
2565:"
2533:"
2516:"
2493:"
2454:"
2362:"
2336:"
2319:"
2279:"
2262:"
2232:"
2184:"
2168:"
2145:"
2129:"
2090:"
2018:"
1834:)
1372:"
1297:"
958:)
339:.
312:,
273:.
246:,
195:,
191:,
187:,
5413:"
5238:"
5226:"
5214:"
5194:"
5174:"
5103:"
5099:"
4991:"
4987:"
4857:)
4734:e
4727:t
4720:v
4684:.
4664::
4637:.
4610:.
4583:.
4556:.
4530:.
4488:.
4466::
4458::
4426:.
4402::
4394::
4364:.
4340::
4334:8
4313:.
4289::
4281::
4249:.
4227::
4219::
4213:6
4192:.
4170::
4162::
4135:.
4123::
4115::
4109:8
4092:.
4068::
4060::
4033:.
4011::
4003::
3976:.
3956::
3948::
3925:.
3911::
3903::
3897:3
3865:.
3845::
3818:.
3794::
3786::
3741:.
3719::
3667:.
3653::
3626:.
3612::
3604::
3564:.
3542::
3534::
3500:.
3486::
3440:"
3433:"
3087:"
3083:"
2992:"
2977:"
2954:"
2937:"
2878:"
2861:"
2807:"
2697:"
2680:"
2656:"
2632:"
2608:"
2584:"
2561:"
2529:"
2512:"
2489:"
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2014:"
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1918:.
1829:+
1827:(
1368:"
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953:+
951:(
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