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Deltochilini

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31: 205:, but in 2006, Smith suggested that the name Canthonini, though junior, might be preserved under Code Article 35.5, which can be used to preserve junior names if they are higher in taxonomic rank than the senior names of constituent taxa. However, this Article does not apply when two names are competing for the same taxonomic rank (as in this case, both being used at the rank of tribe), so this conclusion has been reversed by subsequent classifications, restoring Deltochilini as the valid name. 54: 1042: 1030: 208:
The tribe comprises about 800 species in 120 genera: They constitute a high proportion of the dung beetle diversity in many parts of the world; more than 35% of the genera in the Americas, Australia, Madagascar and many islands. However, in the Afro-Eurasian regions, they make up less than 20% of the
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Smith, A.B.T. (2006) A Review of the Family-group Names for the Superfamily Scarabaeoidea (Coleoptera) with Corrections to Nomenclature and a Current Classification. The Coleopterists Bulletin, 60:144-204.
185:). Members of this group vary widely in size (2–33 mm long) and shape, but were thought to be derived from an ancient ball-rolling lineage. The outer edges of the front 39: 983:
Bouchard, P.; Bousquet, Y.; Davies, A.; Alonso-Zarazaga, M.; Lawrence, J.; Lyal, C.; Newton, A.; Reid, C.; Schmitt, M.; Ślipiński, A.; Smith, A. (2011).
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There has been some controversy regarding the valid name for this tribe. Deltochilini is the senior name, and has precedence under the
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have less than four teeth. The grouping based on these characteristics has, however, been found to have little
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validity, and the placement of several genera in the tribe is likely to change.
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Scholtz, Clarke H.; Davis, Adrian L. V.; Kryger, Ute (2009).
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Evolutionary biology and conservation of dung beetles
917:, and the smaller species may utilize leaf litter ( 976: 948: 1224: 1207: 944: 942: 940: 938: 985:"Family-group names in Coleoptera (Insecta)" 1214: 1200: 935: 29: 1006: 996: 1225: 1057: 1056: 1173: 1157:a46c1b4d-9ee8-4a6a-9b1d-655f3a59a84e 13: 14: 1254: 1040: 1028: 52: 967: 1: 953:. Sofia-Moscow: Pensoft Pub. 928: 1186:. You can help Knowledge by 7: 196: 10: 1259: 1172: 900: 1065: 212: 150: 145: 49:Scientific classification 47: 37: 28: 23: 998:10.3897/zookeys.88.807 1037:at Wikimedia Commons 921:). They are largely 909:, but some feed on 40:Canthon smaragdulus 1243:Scarabaeinae stubs 1195: 1194: 1167: 1166: 1059:Taxon identifiers 1033:Media related to 905:Most species are 157: 156: 141: 1250: 1216: 1209: 1202: 1174: 1160: 1159: 1150: 1149: 1137: 1136: 1124: 1123: 1111: 1110: 1101: 1100: 1099: 1086: 1085: 1084: 1054: 1053: 1045:Data related to 1044: 1032: 1021: 1020: 1010: 1000: 980: 974: 971: 965: 964: 946: 235:Aliuscanthoniola 140:Lacordaire, 1856 139: 57: 56: 43:Fabricius, 1781 33: 21: 20: 16:Tribe of beetles 1258: 1257: 1253: 1252: 1251: 1249: 1248: 1247: 1223: 1222: 1221: 1220: 1170: 1168: 1163: 1155: 1153: 1145: 1140: 1132: 1127: 1119: 1114: 1106: 1104: 1095: 1094: 1089: 1080: 1079: 1074: 1061: 1025: 1024: 981: 977: 972: 968: 961: 947: 936: 931: 903: 898: 865:Tesserodoniella 753:Pseudophacosoma 466:Dwesasilvasedis 319:Baloghonthobium 215: 199: 138: 51: 17: 12: 11: 5: 1256: 1246: 1245: 1240: 1235: 1219: 1218: 1211: 1204: 1196: 1193: 1192: 1165: 1164: 1162: 1161: 1151: 1138: 1125: 1112: 1102: 1087: 1071: 1069: 1063: 1062: 1051: 1050: 1049:at Wikispecies 1038: 1023: 1022: 975: 966: 959: 933: 932: 930: 927: 902: 899: 897: 896: 889: 882: 879:Vulcanocanthon 875: 868: 861: 854: 847: 840: 833: 826: 823:Sphaerocanthon 819: 812: 805: 798: 791: 788:Scybalocanthon 784: 781:Sauvagesinella 777: 770: 767:Saphobiamorpha 763: 756: 749: 746:Pseudonthobium 742: 735: 728: 721: 714: 707: 704:Parvuhowdenius 700: 693: 686: 679: 672: 665: 658: 651: 644: 637: 630: 623: 616: 609: 602: 595: 588: 581: 574: 567: 560: 553: 546: 539: 532: 525: 518: 511: 504: 497: 490: 483: 476: 469: 462: 455: 448: 441: 434: 427: 420: 413: 410:Cephalodesmius 406: 399: 392: 385: 378: 371: 368:Canthodimorpha 364: 357: 354:Cambefortantus 350: 343: 336: 329: 322: 315: 308: 301: 298:Apterepilissus 294: 287: 280: 273: 266: 259: 252: 245: 238: 231: 224: 216: 214: 211: 198: 195: 172:scarab beetles 155: 154: 148: 147: 143: 142: 133: 129: 128: 123: 119: 118: 113: 109: 108: 103: 99: 98: 93: 89: 88: 83: 79: 78: 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733: 732:Pseudignambia 729: 727: 726: 725:Phacosomoides 722: 720: 719: 715: 713: 712: 708: 706: 705: 701: 699: 698: 694: 692: 691: 687: 685: 684: 680: 678: 677: 673: 671: 670: 666: 664: 663: 659: 657: 656: 652: 650: 649: 645: 643: 642: 641:Nebulasilvius 638: 636: 635: 631: 629: 628: 624: 622: 621: 617: 615: 614: 610: 608: 607: 606:Melanocanthon 603: 601: 600: 599:Megathoposoma 596: 594: 593: 589: 587: 586: 582: 580: 579: 578:Madaphacosoma 575: 573: 572: 568: 566: 565: 561: 559: 558: 554: 552: 551: 550:Janssensantus 547: 545: 544: 540: 538: 537: 533: 531: 530: 526: 524: 523: 519: 517: 516: 512: 510: 509: 505: 503: 502: 498: 496: 495: 491: 489: 488: 484: 482: 481: 477: 475: 474: 470: 468: 467: 463: 461: 460: 456: 454: 453: 449: 447: 446: 442: 440: 439: 438:Deltepilissus 435: 433: 432: 431:Cryptocanthon 428: 426: 425: 421: 419: 418: 414: 412: 411: 407: 405: 404: 400: 398: 397: 393: 391: 390: 386: 384: 383: 379: 377: 376: 372: 370: 369: 365: 363: 362: 358: 356: 355: 351: 349: 348: 347:Caeconthobium 344: 342: 341: 337: 335: 334: 333:Boletoscapter 330: 328: 327: 323: 321: 320: 316: 314: 313: 309: 307: 306: 302: 300: 299: 295: 293: 292: 291:Aptenocanthon 288: 286: 285: 281: 279: 278: 274: 272: 271: 267: 265: 264: 260: 258: 257: 253: 251: 250: 246: 244: 243: 239: 237: 236: 232: 230: 229: 225: 223: 222: 218: 217: 210: 206: 204: 194: 192: 188: 184: 180: 177: 173: 169: 165: 161: 153: 149: 144: 137: 134: 131: 130: 127: 124: 121: 120: 117: 114: 111: 110: 107: 104: 101: 100: 97: 94: 91: 90: 87: 84: 81: 80: 77: 74: 71: 70: 67: 64: 61: 60: 55: 50: 46: 42: 41: 36: 32: 27: 24:Deltochilini 22: 19: 1233:Deltochilini 1188:expanding it 1180:Scarabaeinae 1177: 1169: 1066: 1047:Deltochilini 1035:Deltochilini 988: 978: 969: 950: 919:saprophagous 913:, others on 907:coprophagous 904: 891: 884: 877: 870: 863: 856: 849: 842: 837:Sylvicanthon 835: 828: 821: 814: 807: 800: 793: 786: 779: 772: 765: 760:Pycnopanelus 758: 751: 744: 737: 730: 723: 716: 709: 702: 695: 688: 683:Outenikwanus 681: 674: 667: 660: 653: 648:Nesovinsonia 646: 639: 632: 625: 618: 611: 604: 597: 590: 585:Malagoniella 583: 576: 571:Macropanelus 569: 562: 555: 548: 541: 534: 527: 522:Hammondantus 520: 513: 508:Falsignambia 506: 499: 492: 485: 478: 473:Endroedyolus 471: 464: 457: 450: 443: 436: 429: 422: 415: 408: 403:Canthotrypes 401: 396:Canthonosoma 394: 387: 380: 373: 366: 361:Canthochilum 359: 352: 345: 338: 331: 326:Bohepilissus 324: 317: 310: 303: 296: 289: 284:Apotolamprus 282: 275: 268: 261: 256:Anisocanthon 254: 247: 240: 233: 226: 219: 207: 200: 191:phylogenetic 183:Scarabaeinae 163: 160:Deltochilini 159: 158: 151: 136:Deltochilini 135: 126:Scarabaeinae 116:Scarabaeidae 38: 18: 1129:iNaturalist 1091:Wikispecies 886:Xenocanthon 809:Silvaphilus 697:Paracanthon 655:Ochicanthon 620:Monoplistes 613:Mentophilus 536:Holocanthon 445:Deltochilum 389:Canthonidia 382:Canthonella 277:Aphengoecus 270:Anonthobium 242:Amphistomus 176:dung beetle 122:Subfamily: 1227:Categories 1097:Canthonini 1067:Canthonini 929:References 893:Zonocopris 872:Tetraechma 858:Tesserodon 844:Tanzanolus 830:Streblopus 816:Sinapisoma 802:Sikorantus 669:Oficanthon 662:Odontoloma 627:Namakwanus 480:Epactoides 452:Diorygopyx 417:Circellium 340:Byrrhidium 312:Aulacopris 305:Arachnodes 249:Anachalcos 164:Canthonini 106:Coleoptera 86:Arthropoda 774:Saphobius 676:Onthobium 592:Megathopa 515:Gyronotus 501:Eudinopus 487:Epilissus 424:Coproecus 263:Anomiopus 228:Aleiantus 179:subfamily 174:, in the 152:See text 72:Kingdom: 66:Eukaryota 1116:BugGuide 1105:BioLib: 1082:Q4035846 1076:Wikidata 1017:21594053 711:Peckolus 557:Lambroma 543:Ignambia 529:Hansreia 494:Epirinus 221:Agamopus 209:genera. 197:Taxonomy 112:Family: 82:Phylum: 76:Animalia 62:Domain: 1008:3088472 989:ZooKeys 923:diurnal 911:carrion 901:Ecology 718:Penalus 690:Panelus 564:Lepanus 375:Canthon 166:) is a 146:Genera 132:Tribe: 102:Order: 96:Insecta 92:Class: 1154:NZOR: 1147:926261 1134:205338 1015:  1005:  957:  213:Genera 187:tibiae 1178:This 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Index


Canthon smaragdulus
Scientific classification
Edit this classification
Eukaryota
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeidae
Scarabaeinae
Deltochilini
tribe
scarab beetles
dung beetle
subfamily
Scarabaeinae
tibiae
phylogenetic
ICZN
Agamopus
Aleiantus
Aliuscanthoniola
Amphistomus
Anachalcos
Anisocanthon
Anomiopus
Anonthobium
Aphengoecus
Apotolamprus

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