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The gracile fasciculus carries sensory information from the lower half of the body entering the spinal cord at the lumbar level. The cuneate fasciculus carries sensory information from the upper half of the body (upper limbs, trunk, and neck) entering the spinal cord at the cervical level. The
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The cuneate fasciculus is triangular on transverse section, and lies between the gracile fasciculus and the posterior column, its base corresponding with the surface of the spinal cord. Its fibers, larger than those of the gracile fasciculus, are mostly derived from the same source, viz., the
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posterior nerve roots. Some ascend for only a short distance in the tract, and, entering the gray matter, come into close relationship with the cells of the dorsal nucleus, while others can be traced as far as the medulla oblongata, where they end in the gracile nucleus and cuneate nucleus.
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in the medulla, they form the medial lemniscus which connects with the thalamus; the axons synapse on neurons in the ventral posterolateral nucleus which then send axons to the postcentral gyrus in the parietal lobe. All of the axons in the DCML pathway are rapidly conducting, large,
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gracile fasciculus is wedge-shaped on transverse section and lies next to the posterior median septum. Its base is at the surface of the spinal cord, and its apex directed toward the posterior gray commissure. The gracile fasciculus increases in size from inferior to superior.
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in the dorsal horn as a chemical mediator of pain signaling. The dorsal horn of the spinal cord transmits pain and non-noxious signals from the periphery to the spinal cord itself. Adenosine is another local molecule that modulates dorsal horn pain transmission
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and travel in the midline section of the column called the gracile fasciculus. Axons from the upper body enter at or above T6 and travel up the posterior column on the outside of the gracile fasciculus in a more
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then splitting into two branches. The body is situated in the dorsal root ganglion, with one axon traveling peripherally to tissue, and one traveling into the dorsal column. On the right is a
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be confused with a true dendrite), and one central branch that functions like a typical axon by carrying information to other neurons (again, both branches are actually part of one axon).
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and transmitted to the somatosensory cortex. The name dorsal-column medial lemniscus comes from the two structures that carry the sensory information: the dorsal columns of the
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At the medulla, the medial lemniscus is orientated perpendicular to the way the fibres travelled in their tracts in the posterior column. For example, in the column,
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Damage to the dorsal column-medial lemniscus pathway below the crossing point of its fibers results in loss of vibration and joint sense (proprioception) on the
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is more lateral. At the medial lemniscus, axons from the leg are more ventral, and axons from the arm are more dorsal. Fibres from the
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757:. The secondary axons from neurons giving sensation to the head, stay at around the same place, while the leg axons move outwards.
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O'Sullivan, S. B., & Schmitz, T. J. (2007). Physical
Rehabilitation (5th Edition ed.). Philadelphia: F.A. Davis Company.
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of the brain. The pathway receives information from sensory receptors throughout the body, and carries this in the
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level of the spinal cord and carried in the cuneate tract, and information from the lower body is received at the
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Giuffrida, R; Rustioni, A (1992). "Dorsal root ganglion neurons projecting to the dorsal column nuclei of rats".
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Damage to either of the dorsal column tracts can result in the permanent loss of sensation in the limbs. See
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of the body as the lesion. Damage above the crossing point result a loss of vibration and joint sense on the
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719:) are second-order neurons. Their axons cross over to the other side of the medulla and are now named as the
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813:(stereognosis), to determine what an unknown object is, using the hands without visual or audio input.
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level and carried in the gracile tract. The gracile tract is medial to the more lateral cuneate tract.
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793:. Those originating from the head and the leg swap their relative positions. The axons synapse in the
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1057:"Lateral motor column axons execute a ternary trajectory choice between limb and body tissues"
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The cuneate fasciculus, fasciculus cuneatus, cuneate tract, tract of
Burdach, was named for
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section called the cuneate fasciculus. These fasciculi are in an area of white matter, the
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The axons of second-order neurons of the gracile and cuneate nuclei are known as the
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Boitano, Scott; Brooks, Heddwen L.; Barman, Susan M.; Barrett, Kim E. (2015-10-28).
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and allows the detection of fine textures. It also allows for the ability known as
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915:. The gracile fasciculus, the tract of Goll, was named after Swiss neuroanatomist
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310:(body position) from the skin and joints. It transmits this information to the
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The DCML pathway is made up of the axons of first, second, and third-order
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Axons from the lower body enter the posterior column below the level of
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which places them on either side of the posterior intermediate sulcus.
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Sensory information from the upper half of the body is received at the
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in the thalamus and fibres from these ascend to the postcentral gyrus.
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There are three groupings of neurons that are involved in the pathway:
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546:(T6). Ascending tracts typically have three levels of neurons, namely
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The formation of the spinal nerve from the dorsal and ventral roots.
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The axons travel up the rest of the brainstem, and synapse at the
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Spinal cord tracts - tracts of the DCML pathway shown upper right.
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for sensation from the neck, trunk, and extremities, and at the
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The column reaches the junction between the spinal cord and the
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1034:(5th ed.). Sunderland, Mass.: Sinauer. pp. 198–200.
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Transmit sensation of fine touch, vibration and proprioception
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Discriminative sensation is well developed in the fingers of
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of a typical neuron by receiving input (although it should
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The Major
Systems: Sensory Systems - Somatosensory System
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of the body to the lesion. The pathway is tested with
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49:. Unsourced material may be challenged and removed.
789:in the thalamus, ascend the posterior limb of the
727:on each side. This crossing over is known as the
542:The two ascending tracts meet at the level of the
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393:through the two dorsal columns. The first-order
753:The medial lemniscus rotates 90 degrees at the
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442:and when they cross over the midline, at the
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1113:. McGraw-Hill Education. pp. 168–170.
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841:is started. Alternatively, proprioceptive
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785:Axons from the third-order neurons in the
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1405:Posterior column-medial lemniscus pathway
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503:; axons from these neurons ascend as the
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109:Learn how and when to remove this message
2596:Flocculonodular lobe/vestibulocerebellum
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864:The sensory neurons in this pathway are
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58:"Dorsal column–medial lemniscus pathway"
2555:Intermediate hemisphere/spinocerebellum
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671:. They are separated by a partition of
565:Neural connections in the DCML pathway.
330:, tracts that make up the white matter
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2725:Ventral/anterior spinocerebellar tract
2654:Dorsal/posterior spinocerebellar tract
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519:; the axons synapse on neurons in the
268:dorsal column–medial lemniscus pathway
124:Dorsal column-medial lemniscus pathway
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1111:Ganong's Review of Medical Physiology
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715:The neurons in these two nuclei (the
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47:adding citations to reliable sources
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1055:Luria, V; Laufer, E (Jul 2, 2007).
354:and relayed from there through the
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2522:Lateral hemisphere/pontocerebellum
2045:Posterior limb of internal capsule
1861:Posterior limb of internal capsule
362:, and the medial lemniscus in the
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1816:Posterior external arcuate fibers
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479:form the ascending tracts of the
1268:Substantia gelatinosa of Rolando
250:Anatomical terms of neuroanatomy
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507:; the fibers cross over at the
34:needs additional citations for
955:Essentials of Human Physiology
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772:for sensation from the head).
766:ventral posterolateral nucleus
422:ventral posterolateral nucleus
381:. The first-order neurons are
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2751:Rostral spinocerebellar tract
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845:and other skin surface touch
770:ventral posteromedial nucleus
523:which then send axons to the
334:(also known as the posterior
1362:Central gelatinous substance
795:primary somatosensory cortex
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697:First-order neurons secrete
587:(shown left), with a single
583:The first-order neuron is a
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400:second-order neurons of the
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2697:Anterior lobe of cerebellum
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342:, the fibers of the tracts
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2649:Posterior thoracic nucleus
2363:Thalamocortical radiations
2287:Thalamocortical radiations
1306:Posterior thoracic nucleus
147:Originating in peripheral
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2683:Accessory cuneate nucleus
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2601:Vestibulocerebellar tract
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2435:Vestibulocerebellar tract
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1944:Posterior parietal cortex
1911:Anterior white commissure
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1631:Anterior white commissure
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1301:Intermediolateral nucleus
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951:"Section 8/8ch5/s8ch5_22"
787:ventral posterior nucleus
575:Periphery and spinal cord
346:and are continued in the
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155:information to the brain.
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2480:Inferior olivary nucleus
2367:Supplementary motor area
2291:Supplementary motor area
2101:Genu of internal capsule
1953:Spinomesencephalic tract
1139:"Somatosensory Pathways"
663:) that lies between the
515:which connects with the
304:two-point discrimination
2568:Cerebellothalamic tract
2049:Decussation of pyramids
1820:Internal arcuate fibers
1669:Posterior median sulcus
1654:Anterior median fissure
721:internal arcuate fibers
669:posterior median sulcus
544:sixth thoracic vertebra
505:internal arcuate fibers
440:internal arcuate fibers
2502:Deep cerebellar nuclei
2469:Deep cerebellar nuclei
2460:Pontocerebellar fibers
2335:Subthalamic fasciculus
2327:Subthalamic fasciculus
2068:Neuromuscular junction
1074:10.1186/1749-8104-2-13
931:(Anterolateral system)
913:Karl Friedrich Burdach
907:Additional information
901:Brown-Séquard syndrome
686:) with neurons in the
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499:known together as the
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288:central nervous system
16:Sensory spinal pathway
2688:Cuneocerebellar tract
2531:Dentatothalamic tract
2485:Olivocerebellar tract
2382:nigrostriatal pathway
2343:Lenticular fasciculus
2267:Lenticular fasciculus
2170:Vestibulospinal tract
1674:Posterolateral sulcus
1255:Posterior grey column
1030:Purves, Dale (2011).
1001:10.1002/cne.903160206
880:Clinical significance
591:originating from the
585:pseudounipolar neuron
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521:ventral nuclear group
475:. The axons from the
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274:) (also known as the
2200:Reticulospinal tract
2111:Facial motor nucleus
2097:Primary motor cortex
2041:Primary motor cortex
1987:Spinoreticular tract
1828:Trigeminal lemniscus
1767:Meissner's corpuscle
1659:Anterolateral sulcus
1490:Spinoreticular tract
1316:Anterior grey column
1199:Cervical enlargement
717:dorsal column nuclei
706:Second-order neurons
625:dorsal root ganglion
501:dorsal column nuclei
402:dorsal column nuclei
375:second-order neurons
312:somatosensory cortex
188:Somatosensory system
43:improve this article
2563:Reticular formation
2351:Thalamic fasciculus
2331:Subthalamic nucleus
2275:Thalamic fasciculus
2195:Reticular formation
2191:Vestibulocerebellum
2161:Vestibulocerebellum
2106:Corticobulbar tract
2054:Corticospinal tract
1992:Reticular formation
1982:Group C nerve fiber
1958:Superior colliculus
1923:Spinothalamic tract
1810:sensory decussation
1495:Spino-olivary tract
1293:Lateral grey column
929:Spinothalamic tract
859:lamellar corpuscles
819:cutaneous receptors
776:Third-order neurons
729:sensory decussation
657:posterior funiculus
570:First-order neurons
556:third-order neurons
509:sensory decussation
477:first-order neurons
473:dorsal root ganglia
471:, beginning in the
444:sensory decussation
387:dorsal root ganglia
379:third-order neurons
371:first-order neurons
210:Sensorimotor cortex
2790:Spinal cord tracts
2746:Golgi tendon organ
2720:Golgi tendon organ
2551:Interposed nucleus
1781:Cuneate fasciculus
1777:Gracile fasciculus
1763:Pacinian corpuscle
1341:Gamma motor neuron
1329:Alpha motor neuron
1234:Terminal ventricle
1204:Lumbar enlargement
1061:Neural Development
855:bulbous corpuscles
823:tactile corpuscles
781:Thalamus to cortex
619:in shape with its
607:is generated by a
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485:cuneate fasciculus
481:gracile fasciculus
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389:, that send their
328:cuneate fasciculus
324:gracile fasciculus
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2734:Cerebellar vermis
2663:Cerebellar vermis
2606:Vestibular nuclei
2592:Fastigial nucleus
2430:Vestibular nuclei
2409:
2408:
2405:
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2347:Ansa lenticularis
2271:Ansa lenticularis
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2222:→ muscles of neck
2219:Tectospinal tract
2165:Vestibular nuclei
2140:Rubrospinal tract
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1940:Postcentral gyrus
1903:Free nerve ending
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1865:Postcentral gyrus
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1644:External features
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1556:Hypothalamospinal
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949:Nosek, Thomas M.
811:haptic perception
680:medulla oblongata
525:postcentral gyrus
398:make contact with
340:medulla oblongata
316:postcentral gyrus
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149:sensory receptors
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2004:Cingulate cortex
1928:Spinal lemniscus
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1824:Medial lemniscus
1772:Posterior column
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1273:Nucleus proprius
1263:Marginal nucleus
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1209:Conus medullaris
1192:General features
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957:. Archived from
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847:mechanoreceptors
839:action potential
825:that lie in the
791:internal capsule
744:trigeminal nerve
725:medial lemniscus
723:, that form the
637:posterior column
605:action potential
513:medial lemniscus
356:internal capsule
348:medial lemniscus
256:edit on Wikidata
253:
200:Medial lemniscus
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2795:Sensory systems
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2675:muscle spindles
2641:muscle spindles
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2618:Spinocerebellar
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2518:Dentate nucleus
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2214:Midbrain tectum
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1962:Midbrain tectum
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1812:/arcuate fibers
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1789:Cuneate nucleus
1785:Gracile nucleus
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1589:Vestibulospinal
1585:Extrapyramidal
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1433:Spinocerebellar
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1367:Gray commissure
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1283:Rexed lamina VI
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1214:Filum terminale
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843:muscle spindles
817:is detected by
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746:(supplying the
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692:cuneate nucleus
688:gracile nucleus
609:mechanoreceptor
577:
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497:cuneate nucleus
493:gracile nucleus
469:sensory neurons
457:
412:) in the lower
410:cuneate nucleus
406:gracile nucleus
391:afferent fibers
385:located in the
383:sensory neurons
284:sensory pathway
282:) is the major
260:
156:
136:
115:
104:
98:
95:
52:
50:
40:
28:
17:
12:
11:
5:
2813:
2803:
2802:
2797:
2792:
2775:
2774:
2771:
2770:
2767:
2766:
2764:
2763:
2738:
2737:
2712:
2710:
2704:
2703:
2701:
2700:
2667:
2666:
2633:
2631:
2629:proprioception
2620:
2615:Bidirectional:
2612:
2611:
2609:
2608:
2588:
2587:
2547:
2546:
2514:
2512:
2508:
2507:
2505:
2504:
2476:
2475:
2455:Pontine nuclei
2451:
2450:
2426:
2424:
2417:
2411:
2410:
2407:
2406:
2403:
2402:
2400:
2399:
2389:
2387:
2376:
2375:
2312:
2310:
2300:
2299:
2252:
2250:
2237:
2231:
2230:
2227:
2226:
2224:
2223:
2210:
2207:
2206:
2204:
2203:
2187:
2185:
2177:
2176:
2174:
2173:
2157:
2155:
2147:
2146:
2144:
2143:
2131:
2129:
2121:
2120:
2118:
2117:
2115:Facial muscles
2093:
2091:
2080:
2078:Extrapyramidal
2074:
2073:
2071:
2070:
2034:
2032:
2023:
2019:
2018:
2015:
2014:
2011:
2010:
2008:
2007:
1977:
1975:
1968:
1967:
1948:
1947:
1898:
1896:
1886:
1875:
1874:
1871:
1870:
1868:
1867:
1856:
1854:
1847:
1846:
1844:
1843:
1804:
1802:
1795:
1794:
1792:
1791:
1759:
1757:
1747:
1738:
1734:
1733:
1714:
1713:
1706:
1699:
1691:
1682:
1681:
1679:
1678:
1677:
1676:
1671:
1663:
1662:
1661:
1656:
1647:
1645:
1641:
1640:
1637:
1636:
1634:
1633:
1627:
1625:
1621:
1620:
1617:
1616:
1614:
1613:
1612:
1611:
1606:
1601:
1596:
1594:Reticulospinal
1591:
1583:
1582:
1581:
1575:Corticospinal
1572:
1570:
1564:
1563:
1561:
1560:
1559:
1558:
1553:
1551:Reticulospinal
1548:
1543:
1536:Extrapyramidal
1533:
1532:
1531:
1520:
1518:
1509:
1505:
1504:
1501:
1500:
1498:
1497:
1492:
1486:
1484:
1478:
1477:
1475:
1474:
1469:
1467:Posterolateral
1464:
1463:
1462:
1457:
1447:
1446:
1445:
1440:
1429:
1427:
1421:
1420:
1418:
1417:
1412:
1407:
1401:
1399:
1390:
1383:
1377:
1376:
1373:
1372:
1370:
1369:
1364:
1359:
1353:
1351:
1347:
1346:
1344:
1343:
1338:
1337:
1336:
1334:Onuf's nucleus
1326:
1320:
1318:
1312:
1311:
1309:
1308:
1303:
1297:
1295:
1289:
1288:
1286:
1285:
1280:
1278:Rexed lamina V
1275:
1270:
1265:
1259:
1257:
1248:
1242:
1241:
1239:
1238:
1237:
1236:
1226:
1221:
1216:
1211:
1206:
1201:
1195:
1193:
1189:
1188:
1181:
1180:
1173:
1166:
1158:
1152:
1151:
1149:. Apr 9, 2002.
1133:
1132:External links
1130:
1127:
1126:
1119:
1098:
1047:
1040:
1022:
976:
964:
961:on 2016-03-24.
940:
939:
937:
934:
933:
932:
924:
921:
917:Friedrich Goll
908:
905:
894:Romberg's test
881:
878:
866:pseudounipolar
802:
799:
782:
779:
777:
774:
712:
709:
707:
704:
665:posterolateral
633:posterior horn
629:posterior root
617:pseudounipolar
597:bipolar neuron
576:
573:
571:
568:
489:dorsal columns
456:
453:
332:dorsal columns
308:proprioception
262:
261:
252:
246:
245:
242:
238:
237:
232:
226:
225:
221:
220:
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197:
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158:
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153:proprioceptive
146:
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126:
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117:
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31:
29:
22:
15:
9:
6:
4:
3:
2:
2812:
2801:
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2787:
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2761:
2757:
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2740:
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2735:
2731:
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2726:
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2711:
2709:
2705:
2698:
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2689:
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2680:
2676:
2672:
2669:
2668:
2664:
2660:
2656:
2655:
2650:
2646:
2642:
2638:
2635:
2634:
2632:
2630:
2624:
2621:
2619:
2613:
2607:
2603:
2602:
2597:
2593:
2590:
2589:
2586:
2582:
2578:
2574:
2570:
2569:
2564:
2560:
2556:
2552:
2549:
2548:
2545:
2541:
2537:
2533:
2532:
2527:
2523:
2519:
2516:
2515:
2513:
2509:
2503:
2499:
2498:Purkinje cell
2495:
2491:
2487:
2486:
2481:
2478:
2477:
2474:
2470:
2466:
2462:
2461:
2456:
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2449:
2445:
2441:
2437:
2436:
2431:
2428:
2427:
2425:
2421:
2418:
2416:
2412:
2398:
2394:
2393:Pars compacta
2391:
2390:
2388:
2385:
2383:
2377:
2372:
2368:
2364:
2360:
2356:
2352:
2348:
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2308:
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2296:
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2288:
2284:
2280:
2276:
2272:
2268:
2264:
2260:
2256:
2251:
2248:
2247:
2241:
2238:
2236:
2235:Basal ganglia
2232:
2221:
2220:
2215:
2212:
2211:
2208:
2202:
2201:
2196:
2192:
2189:
2188:
2186:
2183:
2178:
2172:
2171:
2166:
2162:
2159:
2158:
2156:
2153:
2148:
2142:
2141:
2136:
2133:
2132:
2130:
2127:
2122:
2116:
2112:
2108:
2107:
2102:
2098:
2095:
2094:
2092:
2089:
2084:
2081:
2079:
2075:
2069:
2065:
2063:
2059:
2055:
2050:
2046:
2042:
2039:
2036:
2035:
2033:
2031:
2027:
2024:
2020:
2005:
2001:
1997:
1993:
1989:
1988:
1983:
1979:
1978:
1976:
1973:
1969:
1965:
1963:
1959:
1955:
1954:
1945:
1941:
1937:
1933:
1929:
1925:
1924:
1921:
1917:
1912:
1908:
1907:A delta fiber
1904:
1900:
1899:
1897:
1894:
1890:
1887:
1885:
1880:
1879:Anterolateral
1876:
1866:
1862:
1858:
1857:
1855:
1852:
1848:
1841:
1837:
1833:
1829:
1825:
1821:
1817:
1813:
1811:
1806:
1805:
1803:
1800:
1796:
1790:
1786:
1782:
1778:
1774:
1773:
1768:
1764:
1761:
1760:
1758:
1755:
1751:
1748:
1746:
1742:
1739:
1735:
1731:
1727:
1726:neural tracts
1723:
1719:
1712:
1707:
1705:
1700:
1698:
1693:
1692:
1689:
1675:
1672:
1670:
1667:
1666:
1664:
1660:
1657:
1655:
1652:
1651:
1649:
1648:
1646:
1642:
1632:
1629:
1628:
1626:
1622:
1610:
1607:
1605:
1602:
1600:
1597:
1595:
1592:
1590:
1587:
1586:
1584:
1580:
1577:
1576:
1574:
1573:
1571:
1569:
1565:
1557:
1554:
1552:
1549:
1547:
1544:
1542:
1539:
1538:
1537:
1534:
1530:
1527:
1526:
1525:
1524:Corticospinal
1522:
1521:
1519:
1517:
1513:
1510:
1506:
1496:
1493:
1491:
1488:
1487:
1485:
1483:
1479:
1473:
1470:
1468:
1465:
1461:
1458:
1456:
1453:
1452:
1451:
1450:Spinothalamic
1448:
1444:
1441:
1439:
1436:
1435:
1434:
1431:
1430:
1428:
1426:
1422:
1416:
1413:
1411:
1408:
1406:
1403:
1402:
1400:
1398:
1394:
1391:
1387:
1384:
1382:
1378:
1368:
1365:
1363:
1360:
1358:
1357:Rexed laminae
1355:
1354:
1352:
1348:
1342:
1339:
1335:
1332:
1331:
1330:
1327:
1325:
1322:
1321:
1319:
1317:
1313:
1307:
1304:
1302:
1299:
1298:
1296:
1294:
1290:
1284:
1281:
1279:
1276:
1274:
1271:
1269:
1266:
1264:
1261:
1260:
1258:
1256:
1252:
1249:
1247:
1243:
1235:
1232:
1231:
1230:
1229:Central canal
1227:
1225:
1222:
1220:
1217:
1215:
1212:
1210:
1207:
1205:
1202:
1200:
1197:
1196:
1194:
1190:
1186:
1179:
1174:
1172:
1167:
1165:
1160:
1159:
1156:
1148:
1144:
1140:
1136:
1135:
1122:
1120:9780071825108
1116:
1112:
1105:
1103:
1094:
1090:
1085:
1080:
1075:
1070:
1066:
1062:
1058:
1051:
1043:
1041:9780878936953
1037:
1033:
1026:
1018:
1014:
1010:
1006:
1002:
998:
995:(2): 206–20.
994:
990:
983:
981:
971:
969:
960:
956:
952:
945:
941:
930:
927:
926:
920:
919:(1829–1903).
918:
914:
904:
902:
897:
895:
891:
890:opposite side
887:
877:
875:
871:
867:
862:
860:
856:
852:
848:
844:
840:
836:
833:close to the
832:
828:
824:
820:
816:
812:
808:
798:
796:
792:
788:
773:
771:
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763:
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749:
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741:
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718:
703:
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689:
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670:
666:
662:
658:
654:
649:
644:
642:
638:
635:, and up the
634:
630:
626:
622:
618:
614:
610:
606:
598:
594:
590:
586:
581:
563:
559:
557:
553:
549:
545:
540:
536:
532:
530:
529:parietal lobe
526:
522:
518:
514:
511:and form the
510:
506:
502:
498:
494:
490:
486:
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478:
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372:
367:
365:
361:
357:
353:
349:
345:
341:
337:
333:
329:
325:
321:
320:parietal lobe
317:
313:
309:
305:
301:
297:
293:
290:that conveys
289:
285:
281:
277:
273:
269:
257:
251:
247:
243:
239:
236:
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227:
222:
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144:
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127:
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113:
110:
102:
91:
88:
84:
81:
77:
74:
70:
67:
63:
60: –
59:
55:
54:Find sources:
48:
44:
38:
37:
32:This article
30:
26:
21:
20:
2749:
2723:
2686:
2652:
2599:
2585:Motor cortex
2566:
2544:Motor cortex
2529:
2483:
2473:Granule cell
2458:
2448:Granule cell
2433:
2380:
2371:Motor cortex
2315:Motor cortex
2305:
2295:Motor cortex
2255:Motor cortex
2245:
2217:
2198:
2181:
2168:
2151:
2138:
2125:
2104:
2087:
2052:
2037:
1985:
1951:
1949:
1914:
1808:
1770:
1381:White matter
1246:Grey columns
1219:Cauda equina
1142:
1110:
1064:
1060:
1050:
1032:Neuroscience
1031:
1025:
992:
988:
959:the original
954:
944:
910:
898:
883:
873:
863:
851:Merkel cells
804:
784:
759:
752:
733:
714:
696:
677:
656:
645:
602:
552:second-order
541:
537:
533:
484:
480:
466:
437:
426:
368:
350:, on to the
279:
275:
271:
267:
265:
234:
105:
96:
86:
79:
72:
65:
53:
41:Please help
36:verification
33:
2627:Unconscious
2573:Red nucleus
2135:Red nucleus
1722:spinal cord
1609:Olivospinal
1604:Raphespinal
1599:Tectospinal
1546:Raphespinal
1541:Rubrospinal
1472:Spinotectal
1324:Interneuron
1185:Spinal cord
738:is medial,
699:substance P
673:glial cells
641:spinal cord
631:, into the
548:first-order
360:spinal cord
224:Identifiers
195:Decussation
172:Neural tube
99:August 2024
2784:Categories
2760:Cerebellum
2742:upper limb
2716:lower limb
2708:Reflex arc
2671:upper limb
2637:lower limb
2494:Hemisphere
2444:Cerebellum
2415:Cerebellar
2293:) → 5° (
2182:extension:
2152:extension:
936:References
815:Fine touch
740:upper limb
736:lower limb
483:, and the
449:myelinated
296:fine touch
292:sensations
241:Acronym(s)
69:newspapers
2369:) → 7° (
2306:indirect:
2030:Pyramidal
1730:fasciculi
1397:Posterior
1147:U Alberta
886:same side
835:epidermis
711:Brainstem
661:funiculus
593:cell body
455:Structure
364:brainstem
344:decussate
300:vibration
167:Precursor
2800:Thalamus
2748:) → 2° (
2722:) → 2° (
2681:) → 2° (
2647:) → 2° (
2577:Thalamus
2536:Thalamus
2511:Efferent
2423:Afferent
2397:Striatum
2361:) → 6° (
2359:Thalamus
2341:) → 5° (
2333:) → 4° (
2325:) → 3° (
2321:) → 2° (
2319:Striatum
2285:) → 4° (
2283:Thalamus
2265:) → 3° (
2261:) → 2° (
2259:Striatum
2126:flexion:
2088:flexion:
2062:Anterior
2038:flexion:
2002:) → 3° (
2000:Thalamus
1994:) → 2° (
1942:) → 4° (
1938:) → 3° (
1936:Thalamus
1920:Anterior
1909:) → 2° (
1895:/lateral
1832:Thalamus
1650:Ventral
1579:Anterior
1568:Anterior
1482:Anterior
1460:anterior
1224:Meninges
1093:17605791
1017:20445258
923:See also
870:dendrite
849:such as
801:Function
764:(at the
762:thalamus
667:and the
603:When an
517:thalamus
495:and the
451:fibers.
429:cervical
418:thalamus
408:and the
352:thalamus
336:funiculi
326:and the
216:Function
2565:, or →
2246:direct:
2058:Lateral
1974:/medial
1916:Lateral
1737:Sensory
1665:Dorsal
1529:Lateral
1516:Lateral
1455:lateral
1443:ventral
1425:Lateral
1415:Cuneate
1410:Gracile
1389:Sensory
1084:1949814
1009:1374085
829:of the
821:called
684:synapse
653:lateral
639:of the
623:in the
611:in the
527:in the
414:medulla
318:in the
314:of the
286:of the
161:Details
83:scholar
2744:→ 1° (
2718:→ 1° (
2673:→ 1° (
2639:→ 1° (
1438:dorsal
1117:
1091:
1081:
1067:: 13.
1038:
1015:
1007:
827:dermis
807:humans
613:tissue
554:, and
433:lumbar
377:, and
306:, and
183:System
85:
78:
71:
64:
56:
2022:Motor
1718:Brain
1508:Motor
1350:Other
1013:S2CID
487:(the
404:(the
395:axons
254:[
230:Latin
176:crest
90:JSTOR
76:books
2583:) →
2542:) →
2313:1° (
2253:1° (
1980:1° (
1972:Slow
1950:2° (
1918:and
1901:1° (
1893:Fast
1884:pain
1822:) →
1783:) →
1745:DCML
1728:and
1720:and
1624:Both
1115:ISBN
1089:PMID
1036:ISBN
1005:PMID
831:skin
755:pons
748:head
621:body
589:axon
280:PCML
272:DCML
266:The
244:DCML
174:and
62:news
2756:ICP
2730:SCP
2693:ICP
2679:DRG
2659:ICP
2645:DRG
2594:in
2559:SCP
2553:in
2526:SCP
2520:in
2490:ICP
2482:→
2465:MCP
2440:ICP
2357:of
2339:GPi
2323:GPe
2281:of
2263:GPi
2163:→
1998:of
1960:of
1934:of
1932:VPL
1840:VPM
1836:VPL
1079:PMC
1069:doi
997:doi
993:316
874:not
659:(a
294:of
45:by
2786::
2758:→
2754:→
2732:→
2728:→
2695:→
2691:→
2685:→
2677:→
2661:→
2657:→
2651:→
2643:→
2604:→
2598:→
2581:VL
2575:→
2571:→
2561:→
2557:→
2540:VL
2534:→
2528:→
2524:→
2500:→
2496:→
2492:→
2488:→
2471:→
2467:→
2463:→
2457:→
2446:→
2442:→
2438:→
2432:→
2395:→
2365:→
2355:VL
2353:→
2349:→
2337:→
2329:→
2317:→
2289:→
2279:VL
2277:→
2273:→
2257:→
2216:→
2197:→
2193:→
2167:→
2137:→
2113:→
2109:→
2103:→
2099:→
2066:→
2060:,
2051:→
2047:→
2043:→
1996:MD
1990:→
1984:→
1964:)
1956:→
1930:→
1926:→
1913:→
1905:→
1863:→
1859:→
1851:3°
1838:,
1830:→
1818:,
1807:→
1799:2°
1769:→
1754:1°
1724::
1145:.
1141:.
1101:^
1087:.
1077:.
1063:.
1059:.
1011:.
1003:.
991:.
979:^
967:^
953:.
903:.
896:.
857:,
853:,
731:.
694:.
648:T6
643:.
550:,
531:.
373:,
366:.
302:,
298:,
206:To
2762:)
2736:)
2699:)
2665:)
2579:(
2538:(
2384::
2373:)
2345:/
2297:)
2269:/
2064:)
2056:(
2006:)
1946:)
1881:/
1853::
1842:)
1834:(
1826:/
1814:(
1801::
1787:/
1779:/
1775:(
1765:/
1756::
1710:e
1703:t
1696:v
1177:e
1170:t
1163:v
1123:.
1095:.
1071::
1065:2
1044:.
1019:.
999::
599:.
278:(
270:(
258:]
112:)
106:(
101:)
97:(
87:·
80:·
73:·
66:·
39:.
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