109:
131:
442:
457:, which also sports the same thin hairs. The head is flat and square, with a pair of long, large pincers. When hunting, they cling to grasses or pieces of wood along the bottom, and hold perfectly still until prey passes by, then they lunge, trapping their prey between their front legs and biting down with their pincers. The larvae are also known to partially consume prey and discard the carcass if another potential prey swims nearby. Their usual prey includes
4197:
4185:
427:, are found to be relatively tolerant to recent urbanization. One of the most important limiting factors for diving beetle occurrence is the presence of fish, which predate on the beetles (mostly on larvae), compete for food, and change the structure of the habitat. The presence or absence of fish can also affect habitat use and habitat selection of dytiscids. Some species, such as
581:
469:. After about a week, or longer in some species, they emerge from the mud as adults. Adult diving beetles have been found to oviposit their eggs within frog spawn in highly ephemeral habitats, with their eggs hatching within 24 hours after the frogs and the larvae voraciously predating on the recently hatched tadpoles.
593:
The greatest threat to diving beetles is the degradation and disappearance of their habitats due to anthropogenic activities. For example, urbanisation has led to the decreasing quantity and quality of dytiscid habitats, which consequentially has increased the distance between habitats.; thus,
414:
Diving beetles are the most diverse beetles in the aquatic environment and can be found in almost every kind of freshwater habitat, from small rock pools to big lakes. Some dytiscid species are also found in brackish water. Diving beetles live in water bodies in various landscapes, including
644:, despite their arsenal of chemical defenses. But by far the most important predator of diving beetles are fish, which limit the occurrence of most diving beetle species to fishless ponds, or to margins of aquatic habitats. Although the larvae of a few dytiscid species may become
565:) are bred for human consumption, though as they are cumbersome to raise due to their carnivorous habit and have a fairly bland (though apparently not offensive) taste and little meat, this is decreasing. Dytiscidae are reportedly also eaten in
701:. It is believed that inducing the beetles to bite the nipples will stimulate breast growth. The effect of that habit has not been tested, but it is notable that the defense glands of diving beetles contain many types of bioactive steroids.
612:
exhibited no change. Brownification, which refers to the change in surface water colour towards yellow–brown hues caused by recent climate change and land-use change, can also drive changes in dytiscid communities. As some species, such as
652:, their presence is also often incompatible with fish. Therefore, the main focus of water beetle conservation is the protection of natural, fish-less habitats. In the European Union, two species of diving beetles are protected by the
452:
When still in larval form, the beetles vary in size from about 1 to 5 cm (0.39 to 1.97 in). The larval bodies are shaped like crescents, with the tail long and covered with thin hairs. Six legs protrude from along the
653:
684:
creation story. According to the narrative, upon finding nowhere to rest in the "liquid chaos" the beetle brought up soft mud from the bottom. This mud then spread out to form all of the land on Earth.
594:
dytiscids may be exposed to high predation risks during dispersal. Urbanisation has complex effects on the inter- and intraspecific variation in dytiscid traits. Some flight-related traits of
3285:
433:, occur in exposed areas of waters, whereas many diving beetles species prefer habitats with aquatic plants, especially plants with complex structures, such as sedges and bulrush.
3964:
4118:
3878:"New record of a parasitising species of Hydrachna (Acari, Hydrachnidia) on water beetles Eretes griseus (Fabricius, 1781) (Coleoptera, Dytiscidae, Dytiscinae, Eretini)"
4083:
3745:"A new family of mites (Acari: Prostigmata: Raphignathina), highly specialized subelytral parasites of dytiscid water beetles (Coleoptera: Dytiscidae: Dytiscinae)"
596:
423:
4163:(2000): Predaceous Diving Beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska. NRC Research Press, Ottawa.
529:
602:
3506:"Microhabitats with emergent plants counterbalance the negative effects of fish presence on diving beetle (Coleoptera: Dytiscidae) diversity in urban ponds"
3034:
541:
3266:
Lundkvist, E.; Landin, J.; Karlsson, F. (2002). "Dispersing diving beetles (Dytiscidae) in agricultural and urban landscapes in south-eastern Sweden".
535:
465:, among other smaller water-dwelling creatures. As the larvae mature, they crawl from the water on the sturdy legs, and bury themselves in the mud for
3586:"Diving beetle (Coleoptera: Dytiscidae) community dissimilarity reveals how low landscape connectivity restricts the ecological value of urban ponds"
398:
and immediately upon biting, they deliver digestive enzymes into prey to suck their liquefied remains. The family includes more than 4,000 described
4134:
3823:"New host-parasite association of Acherontacarus rutilans (Acari, Hydrachnidia, Acherontacaridae) on Scarodytes halensis (Coleoptera: Dytiscidae)"
6332:
3627:"Urbanisation drives inter- and intraspecific variation in flight-related morphological traits of aquatic insects at different landscape scales"
3198:
6371:
202:
371:
are the largest, reaching up to 4.5 and 4.75 cm (1.8 and 1.9 in) respectively. In contrast, the smallest is likely the
Australian
1453:
3784:"Attachment sites, phenology, and growth of larvae of Eylais sp. (Acari) on Dytiscus alaskanus J. Balfour-Browne (Coleoptera: Dytiscidae)"
3230:
Yee, D.A. (2014). "An
Introduction to the Dytiscidae: Their Diversity, Historical Importance, Cultural Significance, and Other Musings".
4155:
359:. The adults of most are between 1 and 2.5 cm (0.4–1.0 in) long, though much variation is seen between species. The European
6117:
3122:
G.N. Foster; D.T. Bilton (2014). "The
Conservation of Predaceous Diving Beetles: Knowns, Unknowns and Anecdotes". In D.A. Yee (ed.).
606:, such as body length and hindwing traits, were found to change along the urban gradient at different scales, whereas the traits of
6306:
108:
6345:
3971:
4168:
3724:
3247:
3181:
3131:
6350:
4058:
3407:"Predation risk modifies habitat use and habitat selection of diving beetles (Coleoptera: Dytiscidae) in an Urban Pondscape"
625:, tend to occur in clear water, brownification may threaten dytiscid species that are intolerant to highly coloured waters.
4230:
3440:"Predator related oviposition site selection of aquatic beetles (Hydroporus spp.) and effects on offspring life-history"
3147:
378:
of subterranean waters, which only is about 0.9 mm (0.035 in) long. Most are dark brown, blackish, or dark
6433:
6285:
6376:
6298:
3480:
6358:
6215:
6192:
117:
3351:"Environmental determinants of diving beetle assemblages (Coleoptera: Dytiscidae) in an urban landscape"
3284:
Law, A.; Baker, A.; Sayer, C.; Foster, G.; Gunn, I.D.; Taylor, P.; Blaikie, James; Willby, N.J. (2019).
6220:
3995:
Bouchard, Patrice; Bousquet, Yves; Davies, Anthony E.; Alonso-Zarazaga, Miguel A.; et al. (2011).
3876:
Arjomandi, Elham; Zawal, Andrzej; Hajiqanbar, Hamidreza; Filip, Ewa; Szenejko, Magdalena (2019-07-22).
130:
6425:
6438:
6386:
3822:
6471:
6466:
4223:
3717:
Ecology, Systematics, and the
Natural History of Predaceous Diving Beetles (Coleoptera: Dytiscidae)
3286:"The effectiveness of aquatic plants as surrogates for wider biodiversity in standing fresh waters"
3232:
Ecology, Systematics, and the
Natural History of Predaceous Diving Beetles (Coleoptera: Dytiscidae)
3205:
3124:
Ecology, Systematics, and the
Natural History of Predaceous Diving Beetles (Coleoptera: Dytiscidae)
6412:
355:. They occur in virtually any freshwater habitat around the world, but a few species live among
6132:
4084:"New fossil taxa and notes on the Mesozoic evolution of Liadytidae and Dytiscidae (Coleoptera)"
668:
6407:
6337:
3715:
Konrad
Dettner (2014). "Chemical Ecology and Biochemistry of Dytiscidae". In D.A. Yee (ed.).
1402:
553:
6399:
6254:
6179:
6170:
5369:
4201:
4004:
3885:
3834:
3597:
3517:
3362:
3300:
3057:
217:
3199:"A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga)"
8:
6461:
5588:
4216:
4124:
897:
662:
615:
558:
547:
489:
429:
361:
4008:
3965:
A World
Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga)
3889:
3838:
3689:
3662:
3601:
3521:
3366:
3304:
4035:
3916:
3858:
3663:"Water Colour Shapes Diving Beetle (Coleoptera: Dytiscidae) Assemblages in Urban Ponds"
3326:
3078:
739:
attach to various locations. These mites are parasitic as larvae with the exception of
520:
515:
125:
3783:
6394:
6241:
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3330:
3243:
3177:
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3061:
2780:
417:
395:
367:
4139:
In: The Human Use of
Insects as a Food Resource: A Bibliographic Account in Progress
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5441:
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5101:
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4554:
4522:
3473:
The
Aquatic Adephaga (Coleoptera) of the Fennoscandia and Denmark. II. Dytiscidae
2645:
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3743:
Mortazavi, Abdolazim; Hajiqanbar, Hamidreza; Lindquist, Evert E (2018-10-20).
654:
Berne Convention on the Conservation of European Wildlife and Natural Habitats
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1995:
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has been used as food in certain regions such as Nagano prefecture. In the
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2005:
1985:
1965:
1935:
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41:
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2435:
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2355:
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2085:
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2055:
2045:
2025:
2015:
1975:
1955:
1825:
1785:
1745:
1625:
1565:
1555:
1535:
1483:
1464:
1251:
1139:
1110:
937:
917:
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804:
574:
305:
273:
86:
51:
3562:
Diving Beetles of the World: Systematics and Biology of the Dytiscidae
3321:
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462:
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142:
91:
19:"Water tiger" redirects here. For the mythical Chinese creature, see
6311:
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71:
4082:
Prokin, A.A.; Petrov, P.N.; Wang, B.; Ponomarenko, A.G. (2013).
5295:
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182:
172:
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20:
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4135:
Chapter 26 - Eastern Asia: China, Japan, and other countries
4119:
Commonwealth Scientific and Industrial Research Organisation
3821:
Aykut, Medeni; Esen, Yunus; Taşar, Gani Erhan (2016-07-03).
3742:
1980:
Roughley & Larson in Larson, Alarie & Roughley, 2000
394:
due to their voracious appetite. They have short, but sharp
4081:
649:
629:
510:
415:
agricultural and urban landscapes. Some species, such as
4611:(furniture beetles, death watch beetles, spider beetles)
3564:. Baltimore: Johns Hopkins University Press. p. 20.
3437:
3265:
3121:
733:
attach to the mesosternal regions and those in genus
619:, are tolerant to brown water, whereas some species,
3560:
Miller, Kelly; Bergsten, Johannes (3 October 2016).
3283:
743:, which are parasitic for their entire life cycle.
6453:
5488:(jewel beetles, or metallic wood-boring beetles)
3438:Brodin, T.; Johansson, F.; Bergsten, J. (2006).
3404:
660:for the protection of natural aquatic habitats:
4127:. Version of 2004-JUL-02. Retrieved 2008-AUG-04
3583:
3503:
3348:
3003:Prokin, Petrov, B. Wang & Ponomarenko, 2013
5211:(true weevils, bark beetles, ambrosia beetles)
4378:
4238:
3820:
3714:
3710:
3708:
3559:
3504:Liao, W.; Venn, Stephen.; Niemelä, J. (2023).
4224:
4147:(2003): Fried water beetles Cantonese style.
3470:
3196:
1670:Balke, Hendrich, Mazzoldi & Biström, 2002
588:
5454:(ironclad beetles, cylindrical bark beetles)
4161:Larson, D.J., Alarie, Y., and Roughley, R.E.
3997:"Family-group names in Coleoptera (Insecta)"
3988:
3574:De Foliart (2002), Jäch (2003), CSIRO (2004)
3171:
3117:
3115:
3113:
3111:
3015:Prokin, Petrov, Wang & Ponomarenko, 2013
477:Adult Dytiscidae, particularly of the genus
3705:
3261:
3259:
3076:Ponomarenko, Coram & Jarzembowski, 2005
3041:), Kzyl-Zhar, Kazakhstan, Late Cretaceous (
751:The following taxonomic sequence gives the
4231:
4217:
497:in a Nevada cave, likely sourced from the
107:
6118:List of subgroups of the order Coleoptera
5198:(straight snout weevils, New York weevil)
4297:
4034:
4016:
3915:
3897:
3749:Zoological Journal of the Linnean Society
3688:
3678:
3642:
3609:
3555:
3553:
3551:
3539:
3529:
3455:
3422:
3405:Liao, W.; Zanca, T.; Niemelä, J. (2024).
3384:
3374:
3320:
3108:
509:is eaten roasted and salted to accompany
436:
5625:(cicada beetle, cicada parasite beetles)
3656:
3654:
3584:Liao, W.; Venn, S.; Niemelä, J. (2022).
3349:Liao, W.; Venn, S.; Niemelä, J. (2020).
3256:
3148:"Dytiscidae - Hurdan, the answer engine"
1184:Miller, Michat, & Ferreira Jr., 2024
1164:Miller, Michat, & Ferreira Jr., 2024
675:
579:
440:
382:in color with golden highlights in some
4063:Integrated Taxonomic Information System
3956:
3624:
3400:
3398:
3396:
3344:
3342:
3340:
6454:
5715:(false fireflies, long-lipped beetles)
4540:
4491:
4123:(2004): Water for a Healthy Country -
3548:
3431:
3279:
3277:
3176:. Dorling Kindersley. pp. 86–87.
1830:K.B. Miller, Gibson & Alarie, 2009
709:Dytiscidae are parasitised by various
6131:
6130:
4539:
4490:
4365:
4364:
4296:
4212:
3781:
3738:
3736:
3651:
3618:
3577:
1690:Hendrich, Toussaint & Balke, 2014
6426:921787F1-F106-994C-B3F2-FBD9FDB4BDF4
6387:e3c6e4f5-658e-44cc-a9e3-e647b2d4bedb
6299:ab35662b-0ff1-4f78-be34-7d0077e9baf7
5741:(false metallic wood-boring beetles)
4051:
3932:
3660:
3393:
3337:
3204:. University of Umeå. Archived from
3081:, United Kingdom, Early Cretaceous (
2430:Jean, Telles & K.B. Miller, 2012
1840:Mazza, Cianferoni & Rocchi, 2013
1530:Biström, Nilsson & Wewalka, 1997
680:The diving beetle plays a role in a
445:Larva of the European diving beetle
343:(δυτικός), "able to dive" – are the
3962:
3471:Nilsson, A. N.; Holmen, M. (1995).
3274:
3229:
1590:Wewalka, Balke & Hendrich, 2001
922:Balke, Hendrich & Wewalka, 1992
557:, and probably also the well-known
13:
3827:International Journal of Acarology
3733:
3475:. Leiden, the Netherlands: Brill.
697:, are collected by young girls in
628:Dytiscid adults are eaten by many
14:
6483:
4176:
3631:Insect Conservation and Diversity
527:, the latter species, as well as
5352:Perimylopidae, or Promecheilidae
4937:(minute brown scavenger beetles)
4195:
4183:
3457:10.1111/j.1365-2427.2006.01563.x
1850:K.B. Miller & Spangler, 2008
1194:Trémouilles & Bachmann, 1980
493:were found in prehistoric human
129:
39:
5567:(variegated mud-loving beetles)
4075:
3869:
3814:
3775:
3568:
3510:Global Ecology and Conservation
3497:
3464:
3411:Global Ecology and Conservation
3097:Yixian Formation, China, Aptian
3017:Yixian Formation, China, Aptian
3005:Yixian Formation, China, Aptian
2993:Yixian Formation, China, Aptian
2981:Yixian Formation, China, Aptian
729:of their hosts, those in genus
688:
3719:. Springer. pp. 235–306.
3223:
3190:
3165:
3140:
3126:. Springer. pp. 437–462.
2721:K.B. Miller & Lubkin, 2001
2550:K.B. Miller & García, 2011
2100:K.B. Miller & García, 2011
746:
1:
5396:(narrow-waisted bark beetles)
5088:(parasitic flat bark beetles)
5039:(short-winged flower beetles)
4895:(lady beetles, or God's cows)
4632:(tooth-necked fungus beetles)
3847:10.1080/01647954.2016.1174304
3355:Biodiversity and Conservation
3101:
3060:, Kazakhstan, Late Jurassic (
693:Adult Dytiscidae, as well as
5908:(sand-loving scarab beetles)
5298:(minute tree-fungus beetles)
5124:(silvanid flat bark beetles)
3037:, Russia, Early Cretaceous (
1463:, Myanmar, Late Cretaceous (
704:
472:
7:
6057:(primitive carrion beetles)
6013:(minute mud-loving beetles)
5876:(bumble bee scarab beetles)
5646:(soft-bodied plant beetles)
5617:(soft bodied plant beetles)
4420:(predaceous diving beetles)
3788:Canadian Journal of Zoology
3782:Aiken, R. B. (1985-02-01).
3531:10.1016/j.gecco.2022.e02361
3424:10.1016/j.gecco.2024.e02801
3240:10.1007/978-94-017-9109-0_1
2965:, China, Early Cretaceous (
2060:Wewalka & Biström, 1994
1880:Wewalka & Biström, 1998
118:Cybister lateralimarginalis
10:
6488:
5892:(scavenger scarab beetles)
5884:(enigmatic scarab beetles)
5057:Lamingtonium binnaberrense
4771:(soft-wing flower beetles)
3625:Liao, W.; Lin, H. (2024).
3611:10.1007/s10980-022-01413-z
3376:10.1007/s10531-020-01977-9
2590:Larson & LaBonte, 1994
2270:Hendrich & Balke, 2009
2130:Roughley & Wolfe, 1987
2110:Hendrich & Balke, 2009
2090:Castro & Delgado, 2001
1084:Shaverdo & Balke, 2008
589:Diving beetle conservation
409:
18:
6139:
6115:
6045:
6031:(water scavenger beetles)
5996:
5962:
5953:
5831:
5822:
5775:
5764:Rhinorhipus tamborinensis
5750:
5634:
5605:
5543:(long-toed water beetles)
5502:
5476:
5467:
5436:(polypore fungus beetles)
5338:(palm and flower beetles)
5322:(tumbling flower beetles)
5269:(ant-like flower beetles)
5249:
5228:
5154:
5047:(lined flat bark beetles)
5021:(pleasing fungus beetles)
4961:
4916:(handsome fungus beetles)
4852:
4708:
4654:
4645:
4620:
4562:
4553:
4549:
4535:
4500:
4486:
4461:(burrowing water beetles)
4374:
4360:
4306:
4292:
4248:
4105:10.11646/zootaxa.3666.2.2
3899:10.3897/zookeys.865.34532
3761:10.1093/zoolinnean/zlx113
3268:Annales Zoologici Fennici
3073:"Palaeodytes" incompletus
3021:Subfamily Incertae sedis
1900:Young & Longley, 1976
1760:Spangler & Barr, 1995
345:predaceous diving beetles
230:
225:
126:Scientific classification
124:
115:
106:
30:
6078:(feather-winged beetles)
5412:(false longhorn beetles)
5359:(jugular-horned beetles)
5306:(false darkling beetles)
5277:(cryptic fungus beetles)
5096:(shining flower beetles)
4436:(crawling water beetles)
4335:Sikhotealinia zhiltzovae
3970:(Report). Archived from
1377:Schaum & White, 1847
1204:Schaum & White, 1847
5559:(forest stream beetles)
5420:(synchroa bark beetles)
5380:(dead log bark beetles)
5372:(fire-coloured beetles)
5261:(ant-like leaf beetles)
4995:(silken fungus beetles)
4929:(well polished beetles)
4903:(minute fungus beetles)
4722:Acanthocnemus nigricans
3680:10.3390/insects15050308
2875:J. Balfour-Browne, 1939
2815:J. Balfour-Browne, 1939
2530:Abeille de Perrin, 1904
2230:J. Balfour-Browne, 1946
1990:J. Balfour-Browne, 1953
1570:Fery & Petrov, 2013
640:, and other vertebrate
6070:(round fungus beetles)
5404:(false flower beetles)
5388:(wedge-shaped beetles)
5346:(false blister beetle)
5330:(hairy fungus beetles)
5285:(conifer bark beetles)
5182:(leaf-rolling weevils)
4843:(bark-gnawing beetles)
4603:(wounded-tree beetles)
4469:(false ground beetles)
4391:(trout-stream beetles)
4018:10.3897/zookeys.88.807
3197:Nilsson, A.N. (2013).
3095:Prokin & Ren, 2010
2991:Prokin & Ren, 2010
2979:Prokin & Ren, 2010
2960:Prokin & Ren, 2010
2948:Prokin & Ren, 2010
982:J.Balfour-Browne, 1943
669:Graphoderus bilineatus
585:
449:
437:Larvae and development
390:are commonly known as
326:Prokin & Ren, 2010
6408:Paleobiology Database
5982:(false clown beetles)
5804:(plate-thigh beetles)
5686:(false click beetles)
5591:(water-penny beetles)
5240:(ship-timber beetles)
5219:(pine flower weevils)
5067:(root-eating beetles)
4887:(minute bark beetles)
4798:Phloiophilus edwardsi
4325:(reticulated beetles)
4149:American Entomologist
3172:G.C. McGavin (2010).
3085:) (undescribed genus)
676:Cultural significance
597:Acilius canaliculatus
583:
444:
424:Acilius canaliculatus
339:– based on the Greek
6294:Fauna Europaea (new)
5860:(false stag beetles)
5845:Belohina inexpectata
5707:(net-winged beetles)
5670:(rare click beetles)
5583:(travertine beetles)
5575:(minute mud beetles)
5140:(dry-fungus beetles)
4975:Agapytho foveicollis
4732:(false skin beetles)
4595:(Jacobson's beetles)
4315:Crowsoniella relicta
4192:at Wikimedia Commons
3058:Karabastau Formation
656:, and thus serve as
37:Late Jurassic–Recent
5858:Diphyllostomatidae
5190:(primitive weevils)
5003:(flat bark beetles)
4753:(checkered beetles)
4740:(fruitworm beetles)
4428:(whirligig beetles)
4131:De Foliart, Gene R.
4059:"Dytiscidae Report"
4009:2011ZooK...88....1B
3890:2019ZooK..865...31A
3839:2016IJAca..42..242A
3602:2022LaEco..37.1049L
3522:2023GEcoC..4102361L
3367:2020BiCon..29.2343L
3305:2019FrBio..64.1664L
755:, their associated
663:Dytiscus latissimus
616:Dytiscus marginalis
603:Hydaticus seminiger
559:great diving beetle
447:Dytiscus marginalis
430:Oreodytes sanmarkii
362:Dytiscus latissimus
5723:(glowworm beetles)
5428:(darkling beetles)
5275:Archeocrypticidae
5132:(palmetto beetles)
4879:(dry bark beetles)
4666:(longhorn beetles)
3644:10.1111/icad.12703
3444:Freshwater Biology
3293:Freshwater Biology
3079:Durlston Formation
3035:Doronino Formation
2260:W.J. Macleay, 1871
713:. Those in genera
586:
521:Guangdong Province
450:
6449:
6448:
6395:Open Tree of Life
6133:Taxon identifiers
6124:
6123:
6111:
6110:
6107:
6106:
6103:
6102:
6095:
6087:
6086:(carrion beetles)
6079:
6071:
6058:
6032:
6014:
5983:
5975:
5949:
5948:
5941:
5933:
5925:
5917:
5909:
5901:
5893:
5885:
5877:
5869:
5861:
5848:
5818:
5817:
5805:
5797:
5767:
5742:
5724:
5716:
5708:
5700:
5687:
5679:
5671:
5663:
5662:(soldier beetles)
5655:
5652:Brachypsectridae
5647:
5626:
5618:
5592:
5584:
5576:
5568:
5560:
5552:
5544:
5531:
5523:
5515:
5489:
5463:
5462:
5455:
5437:
5429:
5421:
5413:
5410:Stenotrachelidae
5405:
5397:
5389:
5381:
5373:
5360:
5347:
5339:
5331:
5323:
5315:
5314:(blister beetles)
5307:
5299:
5286:
5278:
5270:
5262:
5241:
5220:
5212:
5199:
5191:
5183:
5175:
5165:
5141:
5133:
5125:
5097:
5089:
5081:
5068:
5060:
5048:
5040:
5022:
5004:
4996:
4978:
4938:
4930:
4917:
4904:
4896:
4888:
4880:
4862:Akalyptoischiidae
4844:
4801:
4784:
4772:
4754:
4741:
4733:
4725:
4675:
4667:
4641:
4640:
4633:
4612:
4604:
4596:
4583:
4575:
4531:
4530:
4508:
4482:
4481:
4478:
4477:
4470:
4462:
4454:
4437:
4429:
4421:
4413:
4405:
4392:
4356:
4355:
4338:
4326:
4318:
4188:Media related to
4169:978-0-660-17967-4
4125:Family Dytiscidae
3726:978-94-017-9109-0
3661:Liao, W. (2024).
3590:Landscape Ecology
3313:10.1111/fwb.13369
3249:978-94-017-9108-3
3234:. pp. 1–16.
3183:978-1-4053-4997-0
3133:978-94-017-9109-0
3096:
3077:
3056:
3055:Ponomarenko, 1987
3033:
3032:Ponomarenko, 1977
3016:
3004:
2992:
2980:
2961:
2949:
2939:
2929:
2919:
2908:
2898:
2887:
2876:
2866:
2856:
2846:
2836:
2826:
2816:
2806:
2796:
2786:
2781:Japanolaccophilus
2776:
2766:
2756:
2744:
2733:
2722:
2711:
2701:
2691:
2681:
2671:
2661:
2651:
2641:
2631:
2621:
2611:
2601:
2591:
2581:
2571:
2561:
2551:
2541:
2531:
2521:
2511:
2510:Omer-Cooper, 1958
2501:
2491:
2481:
2471:
2470:Franciscolo, 1979
2461:
2451:
2441:
2431:
2421:
2411:
2401:
2391:
2381:
2371:
2370:K.B. Miller, 2012
2361:
2351:
2341:
2331:
2321:
2311:
2301:
2291:
2281:
2271:
2261:
2251:
2241:
2231:
2221:
2211:
2201:
2191:
2181:
2171:
2161:
2151:
2141:
2131:
2121:
2111:
2101:
2091:
2081:
2071:
2061:
2051:
2041:
2031:
2030:Motschulsky, 1853
2021:
2011:
2001:
2000:Motschulsky, 1853
1991:
1981:
1971:
1961:
1951:
1941:
1931:
1921:
1911:
1901:
1891:
1881:
1871:
1861:
1851:
1841:
1831:
1821:
1811:
1801:
1791:
1781:
1771:
1770:K.B. Miller, 1997
1761:
1751:
1741:
1731:
1721:
1711:
1701:
1691:
1681:
1671:
1661:
1651:
1641:
1631:
1621:
1611:
1601:
1591:
1581:
1571:
1561:
1551:
1541:
1531:
1521:
1509:
1499:
1489:
1478:
1459:
1448:
1447:Omer-Cooper, 1956
1438:
1428:
1418:
1408:
1398:
1388:
1378:
1368:
1358:
1348:
1338:
1328:
1318:
1308:
1298:
1288:
1278:
1268:
1256:
1245:
1235:
1225:
1215:
1205:
1195:
1185:
1175:
1165:
1155:
1145:
1135:
1116:
1105:
1095:
1085:
1075:
1065:
1064:Motschulsky, 1855
1055:
1045:
1035:
1025:
1014:
1003:
993:
983:
973:
963:
953:
943:
933:
923:
913:
903:
891:
880:
870:
860:
850:
840:
830:
820:
810:
800:
790:
780:
768:
725:live beneath the
418:Agabus uliginosus
368:Megadytes ducalis
333:
332:
327:
318:
310:
302:
294:
286:
278:
270:
262:
254:
246:
238:
221:
16:Family of beetles
6479:
6442:
6441:
6429:
6428:
6416:
6415:
6403:
6402:
6390:
6389:
6380:
6379:
6367:
6366:
6364:NHMSYS0001718020
6354:
6353:
6341:
6340:
6328:
6327:
6315:
6314:
6302:
6301:
6289:
6288:
6276:
6275:
6263:
6262:
6250:
6249:
6237:
6236:
6224:
6223:
6211:
6210:
6201:
6200:
6188:
6187:
6175:
6174:
6173:
6160:
6159:
6158:
6128:
6127:
6093:
6085:
6077:
6069:
6056:
6030:
6012:
5981:
5973:
5960:
5959:
5955:Staphyliniformia
5939:
5931:
5923:
5915:
5907:
5899:
5891:
5883:
5875:
5867:
5859:
5842:
5829:
5828:
5803:
5794:Declinia relicta
5791:
5761:
5740:
5734:Sinopyrophoridae
5729:Rhagophthalmidae
5722:
5714:
5706:
5698:
5685:
5677:
5669:
5661:
5653:
5645:
5644:Artematopodidae
5624:
5616:
5590:
5582:
5574:
5566:
5558:
5551:(riffle beetles)
5550:
5542:
5530:(turtle beetles)
5529:
5521:
5513:
5487:
5474:
5473:
5453:
5435:
5427:
5419:
5411:
5403:
5395:
5387:
5379:
5371:
5358:
5345:
5337:
5329:
5321:
5313:
5305:
5297:
5284:
5276:
5268:
5260:
5239:
5218:
5210:
5197:
5189:
5181:
5174:(fungus weevils)
5173:
5164:
5163:
5159:
5146:Tasmosalpingidae
5139:
5131:
5123:
5095:
5087:
5079:
5066:
5054:
5046:
5038:
5020:
5002:
4994:
4972:
4936:
4928:
4915:
4902:
4894:
4886:
4878:
4842:
4795:
4778:
4770:
4752:
4739:
4731:
4719:
4718:Acanthocnemidae
4673:
4665:
4652:
4651:
4631:
4610:
4602:
4594:
4581:
4573:
4560:
4559:
4551:
4550:
4537:
4536:
4506:
4488:
4487:
4468:
4460:
4448:
4435:
4427:
4419:
4411:
4404:(ground beetles)
4403:
4390:
4376:
4375:
4362:
4361:
4332:
4324:
4312:
4311:Crowsoniellidae
4294:
4293:
4233:
4226:
4219:
4210:
4209:
4200:Data related to
4199:
4187:
4162:
4146:
4145:Jäch, Manfred A.
4132:
4122:
4109:
4108:
4088:
4079:
4073:
4072:
4070:
4069:
4055:
4049:
4048:
4038:
4020:
3992:
3986:
3985:
3983:
3982:
3976:
3969:
3960:
3954:
3953:
3951:
3950:
3936:
3930:
3929:
3919:
3901:
3873:
3867:
3866:
3818:
3812:
3811:
3779:
3773:
3772:
3740:
3731:
3730:
3712:
3703:
3702:
3692:
3682:
3658:
3649:
3648:
3646:
3622:
3616:
3615:
3613:
3596:(4): 1049–1058.
3581:
3575:
3572:
3566:
3565:
3557:
3546:
3545:
3543:
3533:
3501:
3495:
3494:
3468:
3462:
3461:
3459:
3450:(7): 1277–1285.
3435:
3429:
3428:
3426:
3402:
3391:
3390:
3388:
3378:
3361:(7): 2343–2359.
3346:
3335:
3334:
3324:
3299:(9): 1664–1675.
3290:
3281:
3272:
3271:
3263:
3254:
3253:
3227:
3221:
3220:
3218:
3216:
3210:
3203:
3194:
3188:
3187:
3169:
3163:
3162:
3160:
3159:
3150:. Archived from
3144:
3138:
3137:
3119:
3094:
3075:
3068:, Russia, Aptian
3054:
3031:
3014:
3002:
2990:
2978:
2963:Yixian Formation
2959:
2947:
2937:
2927:
2917:
2906:
2896:
2885:
2874:
2864:
2854:
2844:
2834:
2824:
2814:
2804:
2794:
2784:
2774:
2764:
2754:
2742:
2731:
2720:
2709:
2699:
2689:
2679:
2669:
2659:
2650:Sanfilippo, 1958
2649:
2639:
2629:
2619:
2609:
2599:
2589:
2580:Zimmermann, 1919
2579:
2569:
2559:
2549:
2539:
2529:
2519:
2509:
2499:
2489:
2479:
2469:
2459:
2449:
2439:
2429:
2419:
2409:
2399:
2389:
2379:
2369:
2359:
2349:
2339:
2329:
2319:
2309:
2299:
2289:
2279:
2269:
2259:
2249:
2239:
2229:
2219:
2209:
2199:
2189:
2179:
2169:
2159:
2149:
2139:
2129:
2119:
2109:
2099:
2089:
2079:
2069:
2059:
2049:
2039:
2029:
2020:Clairville, 1806
2019:
2009:
1999:
1989:
1979:
1969:
1959:
1949:
1939:
1929:
1919:
1910:Zimmermann, 1921
1909:
1899:
1889:
1879:
1869:
1859:
1849:
1839:
1829:
1819:
1809:
1799:
1789:
1779:
1769:
1759:
1749:
1739:
1729:
1719:
1709:
1700:Zimmermann, 1919
1699:
1689:
1680:Zimmermann, 1919
1679:
1669:
1659:
1649:
1639:
1629:
1619:
1609:
1599:
1589:
1579:
1569:
1559:
1550:Zimmermann, 1924
1549:
1539:
1529:
1519:
1507:
1498:K.B.Miller, 2002
1497:
1488:K.B.Miller, 2001
1487:
1477:K.B.Miller, 2001
1476:
1458:Yang et al. 2019
1457:
1446:
1436:
1426:
1416:
1406:
1396:
1386:
1376:
1367:Zimmermann, 1928
1366:
1356:
1346:
1336:
1326:
1316:
1306:
1296:
1286:
1276:
1266:
1254:
1243:
1233:
1223:
1213:
1203:
1193:
1183:
1173:
1163:
1153:
1143:
1133:
1114:
1103:
1093:
1083:
1073:
1063:
1053:
1043:
1033:
1024:Zimmermann, 1921
1023:
1012:
1001:
991:
981:
971:
961:
951:
942:Clairville, 1806
941:
931:
921:
911:
901:
889:
878:
868:
858:
848:
838:
828:
818:
809:Guéorguiev, 1971
808:
798:
788:
778:
766:
658:umbrella species
622:Hyphydrus ovatus
325:
316:
308:
300:
292:
284:
276:
268:
260:
252:
244:
236:
216:
134:
133:
111:
101:
38:
34:Temporal range:
28:
27:
6487:
6486:
6482:
6481:
6480:
6478:
6477:
6476:
6472:Aquatic insects
6467:Beetle families
6452:
6451:
6450:
6445:
6437:
6432:
6424:
6419:
6411:
6406:
6398:
6393:
6385:
6383:
6375:
6370:
6362:
6357:
6349:
6344:
6336:
6331:
6323:
6318:
6310:
6305:
6297:
6292:
6284:
6279:
6271:
6266:
6258:
6253:
6245:
6240:
6232:
6227:
6219:
6214:
6206:
6204:
6196:
6191:
6183:
6178:
6169:
6168:
6163:
6154:
6153:
6148:
6135:
6125:
6120:
6099:
6041:
5992:
5974:(clown beetles)
5945:
5916:(betsy beetles)
5824:Scarabaeiformia
5814:
5771:
5746:
5678:(click beetles)
5654:(Texas beetles)
5630:
5601:
5597:Ptilodactylidae
5522:(cedar beetles)
5498:
5459:
5442:Trictenotomidae
5328:Mycetophagidae
5245:
5224:
5161:
5160:
5158:
5150:
5102:Phloeostichidae
5053:Lamingtoniidae
5045:Laemophloeidae
4993:Cryptophagidae
4957:
4848:
4831:Thanerocleridae
4794:Phloiophilidae
4781:Metaxina ornata
4746:Chaetosomatidae
4704:
4637:
4616:
4574:(auger beetles)
4555:Bostrichiformia
4545:
4527:
4523:Torridincolidae
4507:(skiff beetles)
4505:Hydroscaphidae
4496:
4474:
4467:Trachypachidae
4412:(tiger beetles)
4370:
4352:
4302:
4288:
4244:
4237:
4207:
4179:
4174:
4160:
4144:
4130:
4117:
4113:
4112:
4086:
4080:
4076:
4067:
4065:
4057:
4056:
4052:
3993:
3989:
3980:
3978:
3974:
3967:
3961:
3957:
3948:
3946:
3938:
3937:
3933:
3874:
3870:
3819:
3815:
3800:10.1139/z85-041
3780:
3776:
3741:
3734:
3727:
3713:
3706:
3659:
3652:
3623:
3619:
3582:
3578:
3573:
3569:
3558:
3549:
3502:
3498:
3483:
3469:
3465:
3436:
3432:
3403:
3394:
3347:
3338:
3288:
3282:
3275:
3264:
3257:
3250:
3228:
3224:
3214:
3212:
3211:on 3 April 2018
3208:
3201:
3195:
3191:
3184:
3170:
3166:
3157:
3155:
3146:
3145:
3141:
3134:
3120:
3109:
3104:
2918:Mouchamps, 1964
2879:
2825:Brancucci, 1983
2795:Régimbart, 1895
2736:
2670:Brancucci, 1985
2646:Trogloguignotus
2546:Spanglerodessus
2466:Sanfilippodytes
2330:Régimbart, 1903
2250:Régimbart, 1906
1940:Fairmaire, 1869
1860:Brancucci, 1979
1810:Babington, 1841
1740:Régimbart, 1894
1640:Régimbart, 1895
1630:Régimbart, 1895
1600:Babington, 1841
1560:Régimbart, 1900
1494:Microhydrodytes
1470:
1248:
1006:
883:
879:Régimbart, 1879
749:
707:
691:
678:
591:
554:C. tripunctatus
475:
439:
412:
215:
128:
102:
100:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
36:
35:
32:
24:
17:
12:
11:
5:
6485:
6475:
6474:
6469:
6464:
6447:
6446:
6444:
6443:
6430:
6417:
6404:
6391:
6381:
6368:
6355:
6342:
6329:
6316:
6303:
6290:
6281:Fauna Europaea
6277:
6264:
6251:
6238:
6225:
6212:
6202:
6189:
6176:
6161:
6145:
6143:
6137:
6136:
6122:
6121:
6116:
6113:
6112:
6109:
6108:
6105:
6104:
6101:
6100:
6098:
6097:
6094:(rove beetles)
6092:Staphylinidae
6089:
6081:
6073:
6065:
6060:
6051:
6049:
6047:Staphylinoidea
6043:
6042:
6040:
6039:
6034:
6029:Hydrophilidae
6026:
6021:
6016:
6008:
6002:
6000:
5998:Hydrophiloidea
5994:
5993:
5991:
5990:
5985:
5977:
5968:
5966:
5957:
5951:
5950:
5947:
5946:
5944:
5943:
5940:(hide beetles)
5935:
5927:
5924:(rain beetles)
5919:
5911:
5903:
5900:(stag beetles)
5895:
5887:
5879:
5871:
5863:
5855:
5853:Bolboceratidae
5850:
5837:
5835:
5826:
5820:
5819:
5816:
5815:
5813:
5812:
5807:
5799:
5787:
5781:
5779:
5773:
5772:
5770:
5769:
5760:Rhinorhipidae
5756:
5754:
5752:Rhinorhipoidea
5748:
5747:
5745:
5744:
5736:
5731:
5726:
5718:
5710:
5702:
5694:
5689:
5681:
5673:
5665:
5657:
5649:
5640:
5638:
5632:
5631:
5629:
5628:
5620:
5611:
5609:
5603:
5602:
5600:
5599:
5594:
5586:
5578:
5570:
5565:Heteroceridae
5562:
5554:
5546:
5538:
5533:
5528:Chelonariidae
5525:
5520:Callirhipidae
5517:
5514:(pill beetles)
5508:
5506:
5500:
5499:
5497:
5496:
5491:
5482:
5480:
5471:
5465:
5464:
5461:
5460:
5458:
5457:
5449:
5444:
5439:
5431:
5426:Tenebrionidae
5423:
5415:
5407:
5399:
5391:
5383:
5375:
5367:
5362:
5354:
5349:
5341:
5333:
5325:
5317:
5309:
5301:
5293:
5288:
5280:
5272:
5264:
5255:
5253:
5251:Tenebrionoidea
5247:
5246:
5244:
5243:
5234:
5232:
5226:
5225:
5223:
5222:
5214:
5209:Curculionidae
5206:
5201:
5193:
5185:
5177:
5168:
5166:
5156:Curculionoidea
5152:
5151:
5149:
5148:
5143:
5135:
5127:
5119:
5117:Protocucujidae
5114:
5109:
5104:
5099:
5091:
5083:
5075:
5070:
5062:
5050:
5042:
5034:
5029:
5024:
5016:
5011:
5009:Cybocephalidae
5006:
4998:
4990:
4985:
4980:
4967:
4965:
4959:
4958:
4956:
4955:
4950:
4945:
4940:
4932:
4924:
4919:
4911:
4909:Discolomatidae
4906:
4898:
4893:Coccinellidae
4890:
4882:
4877:Bothrideridae
4874:
4869:
4864:
4858:
4856:
4854:Coccinelloidea
4850:
4849:
4847:
4846:
4841:Trogossitidae
4838:
4833:
4828:
4823:
4818:
4813:
4808:
4803:
4791:
4786:
4774:
4766:
4761:
4759:Lophocateridae
4756:
4748:
4743:
4735:
4727:
4714:
4712:
4706:
4705:
4703:
4702:
4697:
4692:
4687:
4682:
4677:
4674:(leaf beetles)
4672:Chrysomelidae
4669:
4660:
4658:
4656:Chrysomeloidea
4649:
4643:
4642:
4639:
4638:
4636:
4635:
4626:
4624:
4618:
4617:
4615:
4614:
4606:
4601:Nosodendridae
4598:
4593:Jacobsoniidae
4590:
4585:
4582:(skin beetles)
4577:
4568:
4566:
4557:
4547:
4546:
4533:
4532:
4529:
4528:
4526:
4525:
4520:
4515:
4510:
4501:
4498:
4497:
4484:
4483:
4480:
4479:
4476:
4475:
4473:
4472:
4464:
4456:
4451:Meru phyllisae
4444:
4439:
4431:
4423:
4415:
4407:
4399:
4394:
4385:
4383:
4372:
4371:
4358:
4357:
4354:
4353:
4351:
4350:
4345:
4343:Micromalthidae
4340:
4328:
4320:
4307:
4304:
4303:
4290:
4289:
4287:
4286:
4280:
4274:
4268:
4262:
4256:
4249:
4246:
4245:
4236:
4235:
4228:
4221:
4213:
4206:
4205:
4204:at Wikispecies
4193:
4180:
4178:
4177:External links
4175:
4173:
4172:
4158:
4142:
4128:
4114:
4111:
4110:
4099:(2): 137–159.
4074:
4050:
3987:
3963:Nilsson, A.N.
3955:
3931:
3884:(865): 31–38.
3868:
3833:(5): 242–246.
3813:
3794:(2): 267–271.
3774:
3755:(3): 695–749.
3732:
3725:
3704:
3650:
3637:(2): 287–303.
3617:
3576:
3567:
3547:
3496:
3481:
3463:
3430:
3392:
3336:
3273:
3255:
3248:
3222:
3189:
3182:
3164:
3139:
3132:
3106:
3105:
3103:
3100:
3099:
3098:
3086:
3069:
3066:Zaza Formation
3046:
3019:
3018:
3006:
2994:
2982:
2970:
2941:
2940:
2930:
2920:
2900:
2899:
2878:
2877:
2867:
2857:
2847:
2837:
2827:
2817:
2807:
2797:
2787:
2777:
2767:
2757:
2746:
2735:
2734:
2732:Théobald, 1937
2723:
2717:Calicovatellus
2712:
2702:
2692:
2682:
2672:
2662:
2652:
2642:
2632:
2622:
2612:
2610:Spangler, 1981
2602:
2592:
2582:
2572:
2562:
2552:
2542:
2540:Spangler, 1996
2532:
2522:
2520:Spangler, 1996
2512:
2502:
2492:
2482:
2472:
2462:
2452:
2442:
2432:
2426:Psychopomporus
2422:
2412:
2402:
2392:
2382:
2372:
2362:
2352:
2342:
2332:
2322:
2312:
2310:Seidlitz, 1887
2302:
2292:
2286:Neoclypeodytes
2282:
2272:
2262:
2252:
2242:
2232:
2222:
2212:
2202:
2192:
2182:
2172:
2162:
2152:
2142:
2132:
2122:
2112:
2102:
2096:Incomptodessus
2092:
2082:
2072:
2062:
2052:
2042:
2040:Stephens, 1828
2032:
2022:
2012:
2002:
1992:
1982:
1972:
1962:
1952:
1946:Heterosternuta
1942:
1932:
1922:
1912:
1902:
1892:
1890:Seidlitz, 1887
1882:
1872:
1862:
1852:
1842:
1832:
1822:
1812:
1802:
1792:
1782:
1772:
1762:
1752:
1742:
1732:
1722:
1712:
1702:
1692:
1682:
1672:
1662:
1652:
1642:
1632:
1622:
1612:
1602:
1592:
1582:
1572:
1562:
1552:
1542:
1532:
1522:
1511:
1501:
1500:
1490:
1469:
1468:
1449:
1439:
1429:
1419:
1409:
1399:
1389:
1387:Chatanay, 1911
1379:
1369:
1359:
1349:
1339:
1329:
1319:
1309:
1299:
1297:Linnaeus, 1758
1289:
1279:
1269:
1258:
1247:
1246:
1236:
1226:
1216:
1214:Chatanay, 1911
1206:
1196:
1186:
1176:
1166:
1156:
1146:
1136:
1125:
1118:
1117:
1097:
1096:
1086:
1076:
1066:
1056:
1046:
1044:Erichson, 1832
1036:
1026:
1020:Agaporomorphus
1005:
1004:
994:
984:
978:Neoscutopterus
974:
972:Seidlitz, 1887
964:
954:
944:
934:
932:Spangler, 1972
924:
914:
904:
893:
890:Erichson, 1837
882:
881:
871:
861:
851:
849:Erichson, 1832
841:
831:
821:
819:Jakovlev, 1897
811:
801:
791:
781:
770:
748:
745:
731:Acherontacarus
706:
703:
690:
687:
677:
674:
646:apex predators
590:
587:
584:Dytiscidae sp.
530:C. bengalensis
487:. Remnants of
474:
471:
438:
435:
411:
408:
365:and Brazilian
331:
330:
329:
328:
319:
311:
303:
295:
287:
279:
271:
263:
255:
247:
245:Erichson, 1837
239:
228:
227:
223:
222:
210:
206:
205:
200:
196:
195:
190:
186:
185:
180:
176:
175:
170:
166:
165:
160:
156:
155:
150:
146:
145:
140:
136:
135:
122:
121:
113:
112:
104:
103:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
40:
33:
15:
9:
6:
4:
3:
2:
6484:
6473:
6470:
6468:
6465:
6463:
6460:
6459:
6457:
6440:
6435:
6431:
6427:
6422:
6418:
6414:
6409:
6405:
6401:
6396:
6392:
6388:
6382:
6378:
6373:
6369:
6365:
6360:
6356:
6352:
6347:
6343:
6339:
6334:
6330:
6326:
6321:
6317:
6313:
6308:
6304:
6300:
6295:
6291:
6287:
6282:
6278:
6274:
6269:
6265:
6261:
6256:
6252:
6248:
6243:
6239:
6235:
6230:
6226:
6222:
6217:
6213:
6209:
6203:
6199:
6194:
6190:
6186:
6181:
6177:
6172:
6166:
6162:
6157:
6151:
6147:
6146:
6144:
6142:
6138:
6134:
6129:
6119:
6114:
6096:
6090:
6088:
6082:
6080:
6074:
6072:
6066:
6064:
6061:
6059:
6053:
6052:
6050:
6048:
6044:
6038:
6035:
6033:
6027:
6025:
6022:
6020:
6017:
6015:
6009:
6007:
6004:
6003:
6001:
5999:
5995:
5989:
5986:
5984:
5980:Sphaeritidae
5978:
5976:
5970:
5969:
5967:
5965:
5961:
5958:
5956:
5952:
5942:
5936:
5934:
5930:Scarabaeidae
5928:
5926:
5920:
5918:
5912:
5910:
5904:
5902:
5896:
5894:
5888:
5886:
5880:
5878:
5872:
5870:
5868:(dor beetles)
5864:
5862:
5856:
5854:
5851:
5849:
5846:
5839:
5838:
5836:
5834:
5833:Scarabaeoidea
5830:
5827:
5825:
5821:
5811:
5808:
5806:
5800:
5798:
5795:
5788:
5786:
5783:
5782:
5780:
5778:
5774:
5768:
5765:
5758:
5757:
5755:
5753:
5749:
5743:
5737:
5735:
5732:
5730:
5727:
5725:
5719:
5717:
5711:
5709:
5703:
5701:
5695:
5693:
5690:
5688:
5682:
5680:
5674:
5672:
5668:Cerophytidae
5666:
5664:
5658:
5656:
5650:
5648:
5642:
5641:
5639:
5637:
5633:
5627:
5623:Rhipiceridae
5621:
5619:
5613:
5612:
5610:
5608:
5604:
5598:
5595:
5593:
5587:
5585:
5579:
5577:
5571:
5569:
5563:
5561:
5557:Eulichadidae
5555:
5553:
5547:
5545:
5539:
5537:
5536:Cneoglossidae
5534:
5532:
5526:
5524:
5518:
5516:
5510:
5509:
5507:
5505:
5501:
5495:
5494:Schizopodidae
5492:
5490:
5484:
5483:
5481:
5479:
5475:
5472:
5470:
5469:Elateriformia
5466:
5456:
5450:
5448:
5445:
5443:
5440:
5438:
5434:Tetratomidae
5432:
5430:
5424:
5422:
5416:
5414:
5408:
5406:
5400:
5398:
5392:
5390:
5386:Ripiphoridae
5384:
5382:
5376:
5374:
5370:Pyrochroidae
5368:
5366:
5365:Pterogeniidae
5363:
5361:
5355:
5353:
5350:
5348:
5342:
5340:
5334:
5332:
5326:
5324:
5318:
5316:
5310:
5308:
5304:Melandryidae
5302:
5300:
5294:
5292:
5291:Chalcodryidae
5289:
5287:
5281:
5279:
5273:
5271:
5265:
5263:
5257:
5256:
5254:
5252:
5248:
5242:
5236:
5235:
5233:
5231:
5227:
5221:
5217:Nemonychidae
5215:
5213:
5207:
5205:
5202:
5200:
5194:
5192:
5186:
5184:
5178:
5176:
5170:
5169:
5167:
5157:
5153:
5147:
5144:
5142:
5136:
5134:
5128:
5126:
5120:
5118:
5115:
5113:
5112:Propalticidae
5110:
5108:
5107:Priasilphidae
5105:
5103:
5100:
5098:
5092:
5090:
5086:Passandridae
5084:
5082:
5080:(sap beetles)
5076:
5074:
5071:
5069:
5063:
5061:
5058:
5051:
5049:
5043:
5041:
5035:
5033:
5030:
5028:
5025:
5023:
5017:
5015:
5012:
5010:
5007:
5005:
4999:
4997:
4991:
4989:
4988:Cavognathidae
4986:
4984:
4981:
4979:
4976:
4969:
4968:
4966:
4964:
4960:
4954:
4951:
4949:
4946:
4944:
4941:
4939:
4933:
4931:
4925:
4923:
4922:Eupsilobiidae
4920:
4918:
4914:Endomychidae
4912:
4910:
4907:
4905:
4901:Corylophidae
4899:
4897:
4891:
4889:
4883:
4881:
4875:
4873:
4870:
4868:
4865:
4863:
4860:
4859:
4857:
4855:
4851:
4845:
4839:
4837:
4834:
4832:
4829:
4827:
4824:
4822:
4819:
4817:
4816:Protopeltidae
4814:
4812:
4811:Prionoceridae
4809:
4807:
4804:
4802:
4799:
4792:
4790:
4787:
4785:
4782:
4775:
4773:
4767:
4765:
4764:Mauroniscidae
4762:
4760:
4757:
4755:
4749:
4747:
4744:
4742:
4736:
4734:
4728:
4726:
4723:
4716:
4715:
4713:
4711:
4707:
4701:
4698:
4696:
4693:
4691:
4688:
4686:
4685:Megalopodidae
4683:
4681:
4678:
4676:
4670:
4668:
4664:Cerambycidae
4662:
4661:
4659:
4657:
4653:
4650:
4648:
4644:
4634:
4630:Derodontidae
4628:
4627:
4625:
4623:
4622:Derodontoidea
4619:
4613:
4607:
4605:
4599:
4597:
4591:
4589:
4588:Endecatomidae
4586:
4584:
4578:
4576:
4572:Bostrichidae
4570:
4569:
4567:
4565:
4564:Bostrichoidea
4561:
4558:
4556:
4552:
4548:
4544:
4538:
4534:
4524:
4521:
4519:
4518:Sphaeriusidae
4516:
4514:
4511:
4509:
4503:
4502:
4499:
4495:
4489:
4485:
4471:
4465:
4463:
4457:
4455:
4452:
4445:
4443:
4440:
4438:
4432:
4430:
4424:
4422:
4416:
4414:
4410:Cicindelidae
4408:
4406:
4400:
4398:
4395:
4393:
4387:
4386:
4384:
4381:
4377:
4373:
4369:
4363:
4359:
4349:
4346:
4344:
4341:
4339:
4336:
4329:
4327:
4321:
4319:
4316:
4309:
4308:
4305:
4301:
4295:
4291:
4285:
4281:
4279:
4275:
4273:
4269:
4267:
4263:
4261:
4257:
4255:
4251:
4250:
4247:
4242:
4234:
4229:
4227:
4222:
4220:
4215:
4214:
4211:
4203:
4198:
4194:
4191:
4186:
4182:
4181:
4170:
4166:
4159:
4157:
4153:
4150:
4143:
4140:
4136:
4129:
4126:
4120:
4116:
4115:
4106:
4102:
4098:
4094:
4093:
4085:
4078:
4064:
4060:
4054:
4046:
4042:
4037:
4032:
4028:
4024:
4019:
4014:
4010:
4006:
4003:(88): 1–972.
4002:
3998:
3991:
3977:on 2019-07-26
3973:
3966:
3959:
3945:
3941:
3935:
3927:
3923:
3918:
3913:
3909:
3905:
3900:
3895:
3891:
3887:
3883:
3879:
3872:
3864:
3860:
3856:
3852:
3848:
3844:
3840:
3836:
3832:
3828:
3824:
3817:
3809:
3805:
3801:
3797:
3793:
3789:
3785:
3778:
3770:
3766:
3762:
3758:
3754:
3750:
3746:
3739:
3737:
3728:
3722:
3718:
3711:
3709:
3700:
3696:
3691:
3686:
3681:
3676:
3672:
3668:
3664:
3657:
3655:
3645:
3640:
3636:
3632:
3628:
3621:
3612:
3607:
3603:
3599:
3595:
3591:
3587:
3580:
3571:
3563:
3556:
3554:
3552:
3542:
3537:
3532:
3527:
3523:
3519:
3515:
3511:
3507:
3500:
3492:
3488:
3484:
3478:
3474:
3467:
3458:
3453:
3449:
3445:
3441:
3434:
3425:
3420:
3416:
3412:
3408:
3401:
3399:
3397:
3387:
3382:
3377:
3372:
3368:
3364:
3360:
3356:
3352:
3345:
3343:
3341:
3332:
3328:
3323:
3318:
3314:
3310:
3306:
3302:
3298:
3294:
3287:
3280:
3278:
3269:
3262:
3260:
3251:
3245:
3241:
3237:
3233:
3226:
3207:
3200:
3193:
3185:
3179:
3175:
3168:
3154:on 2015-05-21
3153:
3149:
3143:
3135:
3129:
3125:
3118:
3116:
3114:
3112:
3107:
3093:
3092:
3087:
3084:
3080:
3074:
3070:
3067:
3063:
3059:
3053:
3052:
3047:
3044:
3040:
3036:
3030:
3029:
3024:
3023:
3022:
3013:
3012:
3007:
3001:
3000:
2995:
2989:
2988:
2983:
2977:
2976:
2971:
2968:
2964:
2958:
2957:
2952:
2951:
2950:
2946:
2945:Liadytiscinae
2936:
2935:
2931:
2926:
2925:
2924:Batrachomatus
2921:
2916:
2915:
2911:
2910:
2909:
2907:Branden, 1885
2905:
2895:
2894:
2890:
2889:
2888:
2886:Branden, 1885
2884:
2873:
2872:
2868:
2865:Guignot, 1937
2863:
2862:
2858:
2855:Guignot, 1937
2853:
2852:
2851:Philaccolilus
2848:
2843:
2842:
2838:
2835:Steiner, 1981
2833:
2832:
2828:
2823:
2822:
2818:
2813:
2812:
2808:
2803:
2802:
2798:
2793:
2792:
2788:
2783:
2782:
2778:
2773:
2772:
2771:Australphilus
2768:
2763:
2762:
2758:
2755:Guignot, 1948
2753:
2752:
2748:
2747:
2745:
2741:
2740:Laccophilinae
2730:
2729:
2724:
2719:
2718:
2713:
2710:Guignot, 1936
2708:
2707:
2703:
2698:
2697:
2693:
2688:
2687:
2683:
2680:Guignot, 1939
2678:
2677:
2673:
2668:
2667:
2663:
2658:
2657:
2656:Tyndallhydrus
2653:
2648:
2647:
2643:
2640:Guignot, 1941
2638:
2637:
2633:
2628:
2627:
2623:
2618:
2617:
2613:
2608:
2607:
2603:
2598:
2597:
2593:
2588:
2587:
2583:
2578:
2577:
2573:
2568:
2567:
2563:
2558:
2557:
2556:Sternopriscus
2553:
2548:
2547:
2543:
2538:
2537:
2533:
2528:
2527:
2523:
2518:
2517:
2513:
2508:
2507:
2503:
2498:
2497:
2493:
2488:
2487:
2483:
2478:
2477:
2473:
2468:
2467:
2463:
2458:
2457:
2453:
2448:
2447:
2443:
2440:Guignot, 1933
2438:
2437:
2433:
2428:
2427:
2423:
2420:Biström, 1988
2418:
2417:
2413:
2408:
2407:
2403:
2400:Guignot, 1945
2398:
2397:
2393:
2390:Biström, 1988
2388:
2387:
2383:
2378:
2377:
2376:Phreatodessus
2373:
2368:
2367:
2363:
2360:Guignot, 1942
2358:
2357:
2353:
2348:
2347:
2343:
2338:
2337:
2333:
2328:
2327:
2323:
2318:
2317:
2313:
2308:
2307:
2303:
2300:Guignot, 1931
2298:
2297:
2293:
2288:
2287:
2283:
2278:
2277:
2273:
2268:
2267:
2266:Neobidessodes
2263:
2258:
2257:
2253:
2248:
2247:
2243:
2238:
2237:
2233:
2228:
2227:
2223:
2218:
2217:
2213:
2208:
2207:
2203:
2200:Guignot, 1945
2198:
2197:
2193:
2188:
2187:
2183:
2180:Guignot, 1950
2178:
2177:
2173:
2170:Guignot, 1939
2168:
2167:
2163:
2160:Guignot, 1939
2158:
2157:
2153:
2150:Guignot, 1936
2148:
2147:
2143:
2138:
2137:
2133:
2128:
2127:
2123:
2118:
2117:
2113:
2108:
2107:
2103:
2098:
2097:
2093:
2088:
2087:
2083:
2080:Guignot, 1939
2078:
2077:
2073:
2070:Illiger, 1802
2068:
2067:
2063:
2058:
2057:
2053:
2048:
2047:
2043:
2038:
2037:
2033:
2028:
2027:
2023:
2018:
2017:
2013:
2008:
2007:
2003:
1998:
1997:
1993:
1988:
1987:
1983:
1978:
1977:
1973:
1968:
1967:
1963:
1960:Biström, 1982
1958:
1957:
1953:
1948:
1947:
1943:
1938:
1937:
1933:
1928:
1927:
1923:
1920:Guignot, 1949
1918:
1917:
1913:
1908:
1907:
1903:
1898:
1897:
1893:
1888:
1887:
1883:
1878:
1877:
1873:
1868:
1867:
1863:
1858:
1857:
1853:
1848:
1847:
1843:
1838:
1837:
1833:
1828:
1827:
1823:
1818:
1817:
1813:
1808:
1807:
1803:
1798:
1797:
1793:
1788:
1787:
1783:
1778:
1777:
1773:
1768:
1767:
1763:
1758:
1757:
1753:
1748:
1747:
1743:
1738:
1737:
1733:
1728:
1727:
1723:
1718:
1717:
1713:
1708:
1707:
1703:
1698:
1697:
1693:
1688:
1687:
1683:
1678:
1677:
1673:
1668:
1667:
1663:
1658:
1657:
1653:
1648:
1647:
1643:
1638:
1637:
1633:
1628:
1627:
1623:
1618:
1617:
1616:Barretthydrus
1613:
1608:
1607:
1603:
1598:
1597:
1593:
1588:
1587:
1586:Anginopachria
1583:
1578:
1577:
1573:
1568:
1567:
1563:
1558:
1557:
1553:
1548:
1547:
1543:
1540:Guignot, 1953
1538:
1537:
1533:
1528:
1527:
1523:
1520:Biström, 1988
1518:
1517:
1513:
1512:
1510:
1506:
1496:
1495:
1491:
1486:
1485:
1481:
1480:
1479:
1475:
1466:
1462:
1461:Burmese amber
1456:
1455:
1450:
1445:
1444:
1440:
1435:
1434:
1430:
1425:
1424:
1420:
1415:
1414:
1413:Spencerhydrus
1410:
1405:
1404:
1400:
1395:
1394:
1390:
1385:
1384:
1380:
1375:
1374:
1370:
1365:
1364:
1360:
1357:Wickham, 1911
1355:
1354:
1350:
1345:
1344:
1340:
1335:
1334:
1330:
1325:
1324:
1320:
1315:
1314:
1310:
1307:Laporte, 1833
1305:
1304:
1300:
1295:
1294:
1290:
1285:
1284:
1280:
1275:
1274:
1270:
1265:
1264:
1260:
1259:
1257:
1253:
1242:
1241:
1237:
1232:
1231:
1227:
1222:
1221:
1220:Spencerhydrus
1217:
1212:
1211:
1207:
1202:
1201:
1197:
1192:
1191:
1190:Paramegadytes
1187:
1182:
1181:
1177:
1172:
1171:
1167:
1162:
1161:
1157:
1152:
1151:
1147:
1142:
1141:
1137:
1132:
1131:
1127:
1126:
1124:
1123:
1113:
1112:
1108:
1107:
1106:
1104:Branden, 1885
1102:
1092:
1091:
1087:
1082:
1081:
1077:
1072:
1071:
1067:
1062:
1061:
1057:
1052:
1051:
1047:
1042:
1041:
1037:
1032:
1031:
1027:
1022:
1021:
1017:
1016:
1015:
1013:Branden, 1885
1011:
1000:
999:
995:
990:
989:
985:
980:
979:
975:
970:
969:
965:
962:Laporte, 1835
960:
959:
955:
950:
949:
945:
940:
939:
935:
930:
929:
925:
920:
919:
915:
910:
909:
905:
900:
899:
895:
894:
892:
888:
877:
876:
872:
869:Thomson, 1859
867:
866:
862:
857:
856:
852:
847:
846:
842:
837:
836:
832:
827:
826:
822:
817:
816:
812:
807:
806:
802:
797:
796:
792:
787:
786:
782:
777:
776:
772:
771:
769:
767:Thomson, 1867
765:
760:
758:
754:
744:
742:
738:
737:
732:
728:
724:
723:
718:
717:
712:
702:
700:
696:
686:
683:
673:
671:
670:
665:
664:
659:
655:
651:
647:
643:
639:
635:
631:
626:
624:
623:
618:
617:
611:
610:
605:
604:
599:
598:
582:
578:
576:
572:
568:
564:
563:D. marginalis
560:
556:
555:
550:
549:
548:C. sugillatus
544:
543:
538:
537:
532:
531:
526:
522:
518:
517:
512:
508:
507:C. explanatus
504:
500:
499:Humboldt Sink
496:
492:
491:
490:C. explanatus
486:
482:
481:
470:
468:
464:
460:
456:
448:
443:
434:
432:
431:
426:
425:
420:
419:
407:
405:
401:
397:
393:
389:
385:
381:
377:
375:
370:
369:
364:
363:
358:
354:
353:water beetles
350:
346:
342:
338:
324:
323:Liadytiscinae
320:
317:Branden, 1885
315:
312:
309:Branden, 1885
307:
304:
299:
298:Laccophilinae
296:
291:
288:
283:
280:
275:
272:
267:
264:
261:Branden, 1885
259:
256:
253:Branden, 1885
251:
248:
243:
240:
237:Thomson, 1867
235:
232:
231:
229:
224:
219:
214:
211:
208:
207:
204:
201:
199:Superfamily:
198:
197:
194:
191:
188:
187:
184:
181:
178:
177:
174:
171:
168:
167:
164:
161:
158:
157:
154:
151:
148:
147:
144:
141:
138:
137:
132:
127:
123:
120:
119:
114:
110:
105:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
29:
26:
22:
6140:
6019:Helophoridae
6011:Georissidae
6006:Epimetopidae
5922:Pleocomidae
5906:Ochodaeidae
5890:Hybosoridae
5874:Glaphyridae
5866:Geotrupidae
5844:
5841:Belohinidae
5802:Eucinetidae
5793:
5790:Decliniidae
5763:
5721:Phengodidae
5660:Cantharidae
5615:Dascillidae
5607:Dascilloidea
5589:Psephenidae
5581:Lutrochidae
5573:Limnichidae
5486:Buprestidae
5478:Buprestoidea
5418:Synchroidae
5402:Scraptiidae
5394:Salpingidae
5357:Prostomidae
5344:Oedemeridae
5320:Mordellidae
5238:Lymexylidae
5230:Lymexyloidea
5180:Attelabidae
5172:Anthribidae
5130:Smicripidae
5094:Phalacridae
5078:Nitidulidae
5073:Myraboliidae
5065:Monotomidae
5056:
5037:Kateretidae
5014:Cyclaxyridae
4974:
4971:Agapythidae
4935:Latridiidae
4885:Cerylonidae
4872:Anamorphidae
4806:Phycosecidae
4797:
4780:
4777:Metaxinidae
4730:Biphyllidae
4721:
4690:Orsodacnidae
4647:Cucujiformia
4580:Dermestidae
4450:
4417:
4389:Amphizoidae
4334:
4314:
4300:Archostemata
4284:Holometabola
4282:Superorder:
4276:Infraclass:
4156:PDF fulltext
4154:(1): 34–37.
4151:
4148:
4138:
4096:
4090:
4077:
4066:. Retrieved
4062:
4053:
4000:
3990:
3979:. Retrieved
3972:the original
3958:
3947:. Retrieved
3943:
3940:"Dytiscidae"
3934:
3881:
3871:
3830:
3826:
3816:
3791:
3787:
3777:
3752:
3748:
3716:
3670:
3666:
3634:
3630:
3620:
3593:
3589:
3579:
3570:
3561:
3541:10138/352745
3513:
3509:
3499:
3472:
3466:
3447:
3443:
3433:
3414:
3410:
3386:10138/315037
3358:
3354:
3296:
3292:
3267:
3231:
3225:
3213:. Retrieved
3206:the original
3192:
3173:
3167:
3156:. Retrieved
3152:the original
3142:
3123:
3089:
3072:
3049:
3026:
3020:
3009:
2997:
2985:
2973:
2954:
2942:
2932:
2922:
2912:
2901:
2891:
2880:
2869:
2859:
2849:
2841:Neptosternus
2839:
2829:
2821:Laccosternus
2819:
2809:
2799:
2789:
2779:
2769:
2765:Crotch, 1873
2759:
2751:Africophilus
2749:
2743:Gistel, 1856
2737:
2728:Procoelambus
2726:
2715:
2704:
2694:
2684:
2674:
2666:Typhlodessus
2664:
2654:
2644:
2636:Trichonectes
2634:
2624:
2614:
2604:
2594:
2584:
2576:Stictotarsus
2574:
2570:Brinck, 1943
2566:Stictonectes
2564:
2554:
2544:
2534:
2524:
2514:
2504:
2494:
2490:Palmer, 1957
2486:Schistomerus
2484:
2474:
2464:
2460:Bameul, 1989
2456:Rhithrodytes
2454:
2444:
2434:
2424:
2414:
2404:
2394:
2384:
2380:Ordish, 1976
2374:
2364:
2354:
2344:
2334:
2324:
2314:
2304:
2294:
2284:
2274:
2264:
2254:
2244:
2234:
2224:
2214:
2204:
2194:
2190:Brinck, 1943
2184:
2174:
2164:
2154:
2144:
2134:
2126:Laccornellus
2124:
2120:Ordish, 1976
2116:Kuschelydrus
2114:
2106:Kakadudessus
2104:
2094:
2084:
2074:
2064:
2054:
2044:
2034:
2024:
2014:
2004:
1996:Hydroglyphus
1994:
1984:
1974:
1964:
1954:
1950:Strand, 1935
1944:
1934:
1924:
1914:
1906:Hemibidessus
1904:
1894:
1884:
1876:Glareadessus
1874:
1864:
1854:
1844:
1836:Etruscodytes
1834:
1824:
1816:Dimitshydrus
1814:
1806:Desmopachria
1804:
1796:Derovatellus
1794:
1784:
1776:Darwinhydrus
1774:
1764:
1754:
1744:
1734:
1726:Chostonectes
1724:
1714:
1704:
1694:
1684:
1674:
1666:Borneodessus
1664:
1654:
1644:
1636:Bidessonotus
1634:
1624:
1614:
1604:
1596:Anodocheilus
1594:
1584:
1574:
1564:
1554:
1544:
1534:
1524:
1514:
1505:Hydroporinae
1502:
1492:
1482:
1474:Hydrodytinae
1471:
1452:
1443:Tikoloshanes
1441:
1437:Dejean, 1833
1433:Thermonectus
1431:
1427:Brinck, 1945
1421:
1411:
1403:Sandracottus
1401:
1391:
1383:Regimbartina
1381:
1373:Onychohydrus
1371:
1361:
1351:
1341:
1331:
1321:
1317:Dejean, 1833
1311:
1301:
1291:
1281:
1273:Aethionectes
1271:
1261:
1249:
1244:Brinck, 1945
1238:
1234:Brinck, 1945
1228:
1218:
1210:Regimbartina
1208:
1200:Onychohydrus
1198:
1188:
1178:
1168:
1160:Nilssondytes
1158:
1154:Curtis, 1827
1148:
1144:Brinck, 1945
1138:
1128:
1119:
1109:
1101:Coptotominae
1098:
1094:García, 2001
1088:
1078:
1074:Dejean, 1833
1068:
1058:
1048:
1038:
1028:
1018:
1007:
1002:García, 2001
996:
992:Dejean, 1833
986:
976:
966:
956:
946:
936:
926:
916:
912:Brinck, 1948
906:
902:Brinck, 1943
896:
887:Colymbetinae
884:
873:
863:
853:
843:
839:Crotch, 1873
833:
823:
815:Hydronebrius
813:
803:
793:
783:
779:Crotch, 1873
773:
761:
750:
741:Dytiscacarus
740:
734:
730:
720:
716:Dytiscacarus
714:
708:
692:
689:Ethnobiology
679:
667:
661:
627:
620:
614:
609:Ilybius ater
607:
601:
595:
592:
562:
552:
546:
540:
534:
528:
516:C. japonicus
514:
513:. In Japan,
506:
488:
478:
476:
451:
446:
428:
422:
416:
413:
402:in numerous
392:water tigers
391:
372:
366:
360:
344:
340:
336:
334:
301:Gistel, 1856
290:Hydroporinae
285:Miller, 2001
282:Hydrodytinae
258:Coptotominae
242:Colymbetinae
226:Subfamilies
212:
116:
25:
6320:iNaturalist
6165:Wikispecies
6063:Hydraenidae
6037:Spercheidae
6024:Hydrochidae
5988:Synteliidae
5972:Histeridae
5964:Histeroidea
5914:Passalidae
5882:Glaresidae
5739:Throscidae
5699:(fireflies)
5697:Lampyridae
5684:Eucnemidae
5676:Elateridae
5636:Elateroidea
5452:Zopheridae
5336:Mycteridae
5267:Anthicidae
5138:Sphindidae
5122:Silvanidae
5032:Hobartiidae
5019:Erotylidae
4948:Mycetaeidae
4943:Murmidiidae
4927:Euxestidae
4821:Rentoniidae
4695:Oxypeltidae
4680:Disteniidae
4513:Lepiceridae
4442:Hygrobiidae
4434:Haliplidae
4418:Dytiscidae
4397:Aspidytidae
3051:Palaeodytes
2999:Liadyxianus
2975:Liadytiscus
2956:Liadroporus
2943:Subfamily †
2928:Clark, 1863
2897:Sharp, 1882
2861:Philaccolus
2845:Sharp, 1882
2805:Leach, 1815
2801:Laccophilus
2775:Watts, 1978
2700:Gozis, 1886
2660:Sharp, 1882
2630:Watts, 1985
2620:Watts, 1982
2616:Terradessus
2606:Tepuidessus
2600:Gozis, 1914
2596:Suphrodytes
2560:Sharp, 1880
2506:Sharphydrus
2500:Watts, 1997
2496:Sekaliporus
2480:Gozis, 1914
2450:Sharp, 1882
2416:Pseuduvarus
2410:Sharp, 1882
2366:Petrodessus
2350:Sharp, 1882
2340:Balke, 2001
2336:Papuadessus
2326:Pachynectes
2320:Sharp, 1882
2290:Young, 1967
2280:Young, 1967
2276:Neobidessus
2256:Necterosoma
2246:Nebrioporus
2220:Young, 1967
2216:Microdessus
2210:Sharp, 1882
2156:Limbodessus
2140:Gozis, 1914
2050:Sharp, 1880
2010:Sharp, 1882
2006:Hydropeplus
1986:Hydrodessus
1970:Sharp, 1882
1966:Huxelhydrus
1936:Heterhydrus
1930:Sharp, 1880
1926:Herophydrus
1896:Haideoporus
1886:Graptodytes
1870:Watts, 1978
1866:Gibbidessus
1846:Fontidessus
1800:Sharp, 1882
1790:Sharp, 1882
1780:Sharp, 1882
1766:Crinodessus
1756:Comaldessus
1750:Sharp, 1882
1736:Clypeodytes
1730:Sharp, 1880
1710:Watts, 1978
1706:Carabhydrus
1696:Canthyporus
1686:Brancuporus
1676:Brachyvatus
1660:Angus, 2010
1656:Boreonectes
1650:Sharp, 1882
1610:Sharp, 1882
1580:Sharp, 1882
1546:Allopachria
1526:Agnoshydrus
1516:Africodytes
1423:Sternhydrus
1417:Sharp, 1882
1407:Sharp, 1882
1397:Sharp, 1880
1393:Rhantaticus
1353:Miodytiscus
1347:Sharp, 1882
1327:Leach, 1817
1313:Graphoderus
1287:Watts, 1978
1283:Austrodytes
1277:Sharp, 1882
1267:Leach, 1817
1255:Leach, 1815
1240:Trifurcitus
1230:Sternhydrus
1224:Sharp, 1882
1180:Metaxydytes
1174:Sharp, 1882
1134:Watts, 1978
1130:Austrodytes
1122:Cybistrinae
1080:Madaglymbus
1060:Lacconectus
1054:Broun, 1886
1034:Sharp, 1880
1010:Copelatinae
968:Melanodytes
875:Platynectes
859:Sharp, 1882
855:Leuronectes
829:Sharp, 1882
825:Hydrotrupes
799:Sharp, 1882
789:Leach, 1817
753:subfamilies
747:Systematics
699:East Africa
542:C. limbatus
384:subfamilies
374:Limbodessus
357:leaf litter
277:Leach, 1815
269:Sharp, 1880
266:Cybistrinae
250:Copelatinae
203:Dytiscoidea
6462:Dytiscidae
6456:Categories
6198:Dytiscidae
6185:Dytiscidae
6171:Dytiscidae
6141:Dytiscidae
6084:Silphidae
6076:Ptiliidae
6068:Leiodidae
6055:Agyrtidae
5898:Lucanidae
5777:Scirtoidea
5713:Omethidae
5692:Jurasaidae
5541:Dryopidae
5512:Byrrhidae
5504:Byrrhoidea
5196:Brentidae
5001:Cucujidae
4983:Boganiidae
4963:Cucujoidea
4836:Thymalidae
4826:Rhadalidae
4769:Melyridae
4738:Byturidae
4700:Vesperidae
4609:Ptiniidae
4459:Noteridae
4426:Gyrinidae
4402:Carabidae
4331:Jurodidae
4323:Cupedidae
4270:Subclass:
4260:Arthropoda
4241:Coleoptera
4202:Dytiscidae
4190:Dytiscidae
4068:2019-06-17
3981:2019-06-18
3949:2019-06-17
3673:(4): 308.
3516:: e02361.
3482:9004104569
3417:: e02801.
3322:1893/30068
3158:2015-05-19
3102:References
3083:Berriasian
3028:Cretodytes
2938:Aubé, 1836
2902:Subfamily
2883:Lancetinae
2881:Subfamily
2871:Philodytes
2811:Laccoporus
2791:Laccodytes
2785:Satô, 1972
2738:Subfamily
2690:Aubé, 1837
2586:Stygoporus
2516:Siamoporus
2476:Scarodytes
2436:Pteroporus
2386:Platydytes
2356:Peschetius
2240:Uéno, 1957
2226:Microdytes
2086:Iberoporus
2076:Hypodessus
2056:Hyphovatus
2046:Hyphoporus
2026:Hydrovatus
2016:Hydroporus
1976:Hydrocolus
1956:Hovahydrus
1826:Ereboporus
1820:Uéno, 1996
1786:Deronectes
1746:Coelhydrus
1720:Aubé, 1837
1626:Bidessodes
1566:Amurodytes
1556:Amarodytes
1536:Allodessus
1508:Aubé, 1836
1503:Subfamily
1484:Hydrodytes
1472:Subfamily
1465:Cenomanian
1454:Ambarticus
1337:Hope, 1838
1252:Dytiscinae
1250:Subfamily
1140:Bifurcitus
1120:Subfamily
1111:Coptotomus
1099:Subfamily
1008:Subfamily
952:Fall, 1927
938:Colymbetes
918:Carabdytes
898:Anisomeria
885:Subfamily
835:Ilybiosoma
805:Andonectes
762:Subfamily
575:New Guinea
536:C. guerini
495:coprolites
463:glassworms
337:Dytiscidae
306:Lancetinae
293:Aubé, 1836
274:Dytiscinae
213:Dytiscidae
189:Suborder:
183:Coleoptera
163:Arthropoda
31:Dytiscidae
5938:Trogidae
5932:(scarabs)
5810:Scirtidae
5785:Clambidae
5378:Pythidae
5312:Meloidae
5259:Aderidae
5162:(weevils)
5027:Helotidae
4953:Teredidae
4867:Alexiidae
4751:Cleridae
4710:Cleroidea
4543:Polyphaga
4541:Suborder
4494:Myxophaga
4492:Suborder
4447:Meruidae
4366:Suborder
4348:Ommatidae
4298:Suborder
4272:Pterygota
4252:Kingdom:
4027:1313-2989
3908:1313-2970
3855:0164-7954
3808:0008-4301
3769:0024-4082
3491:0106-8377
3331:202032378
3091:Sinoporus
3062:Oxfordian
3039:Barremian
3011:Mesodytes
2987:Mesoderus
2914:Allomatus
2831:Napodytes
2536:Sinodytes
2526:Siettitia
2406:Primospes
2396:Porhydrus
2316:Pachydrus
2306:Oreodytes
2236:Morimotoa
2196:Metaporus
2186:Megaporus
2176:Lioporeus
2166:Liodessus
2146:Leiodytes
2136:Laccornis
2066:Hyphydrus
1916:Heroceras
1856:Geodessus
1620:Lea, 1927
1606:Antiporus
1363:Notaticus
1343:Megadytes
1333:Hyderodes
1323:Hydaticus
1170:Megadytes
1115:Say, 1830
1070:Liopterus
1050:Exocelina
1040:Copelatus
908:Senilites
865:Platambus
795:Agametrus
736:Hydrachna
705:Parasites
695:Gyrinidae
648:in small
642:predators
473:Edibility
396:mandibles
376:atypicali
149:Kingdom:
143:Eukaryota
6229:BugGuide
6205:BioLib:
6150:Wikidata
5705:Lycidae
5549:Elmidae
5447:Ulodidae
5283:Boridae
5188:Belidae
4789:Peltidae
4382:families
4368:Adephaga
4278:Neoptera
4258:Phylum:
4254:Animalia
4243:families
4133:(2002):
4045:21594053
3926:31379442
3863:88103217
3699:38786864
3690:11122460
3215:10 April
3043:Turonian
2893:Lancetes
2761:Agabetes
2686:Vatellus
2346:Paroster
2296:Neoporus
2036:Hygrotus
1646:Bidessus
1293:Dytiscus
1150:Cybister
1030:Aglymbus
958:Meladema
948:Hoperius
775:Agabinus
764:Agabinae
682:Cherokee
638:reptiles
571:Thailand
480:Cybister
467:pupation
459:tadpoles
234:Agabinae
209:Family:
193:Adephaga
159:Phylum:
153:Animalia
139:Domain:
6156:Q327149
5296:Ciidae
5204:Caridae
4266:Insecta
4264:Class:
4239:Extant
4121:(CSIRO)
4092:Zootaxa
4036:3088472
4005:Bibcode
4001:ZooKeys
3917:6663934
3886:Bibcode
3882:ZooKeys
3835:Bibcode
3667:Insects
3598:Bibcode
3518:Bibcode
3363:Bibcode
3301:Bibcode
3174:Insects
2904:Matinae
2626:Tiporus
2206:Methles
1263:Acilius
1090:Rugosus
998:Rugosus
988:Rhantus
928:Bunites
845:Ilybius
634:mammals
410:Habitat
400:species
341:dytikos
314:Matinae
179:Order:
173:Insecta
169:Class:
6439:150718
6413:346388
6384:NZOR:
6351:111963
6338:100157
6273:1DTSCF
4380:Extant
4167:
4043:
4033:
4025:
3924:
3914:
3906:
3861:
3853:
3806:
3767:
3723:
3697:
3687:
3489:
3479:
3329:
3246:
3180:
3130:
2967:Aptian
2706:Yolina
2676:Uvarus
1716:Celina
1303:Eretes
785:Agabus
757:genera
727:elytra
722:Eylais
573:, and
567:Taiwan
503:Mexico
485:edible
483:, are
455:thorax
404:genera
388:larvae
386:. The
349:family
220:, 1815
21:shuihu
6434:WoRMS
6421:Plazi
6400:66507
6377:50515
6333:IRMNG
6325:57659
6286:11011
4087:(PDF)
3975:(PDF)
3968:(PDF)
3859:S2CID
3327:S2CID
3289:(PDF)
3209:(PDF)
3202:(PDF)
2934:Matus
2446:Queda
1576:Andex
711:mites
650:ponds
630:birds
525:China
511:tacos
501:. In
380:olive
218:Leach
6372:NCBI
6346:ITIS
6312:7824
6307:GBIF
6268:EPPO
6260:7447
6221:1168
6216:BOLD
6208:4591
4165:ISBN
4097:3666
4041:PMID
4023:ISSN
3944:GBIF
3922:PMID
3904:ISSN
3851:ISSN
3804:ISSN
3765:ISSN
3721:ISBN
3695:PMID
3487:ISSN
3477:ISBN
3244:ISBN
3217:2018
3178:ISBN
3128:ISBN
2696:Yola
719:and
666:and
600:and
461:and
421:and
347:, a
335:The
42:PreꞒ
6359:NBN
6255:EoL
6247:9JJ
6242:CoL
6234:195
6193:AFD
6180:ADW
4101:doi
4031:PMC
4013:doi
3912:PMC
3894:doi
3843:doi
3796:doi
3757:doi
3753:184
3685:PMC
3675:doi
3639:doi
3606:doi
3536:hdl
3526:doi
3452:doi
3419:doi
3381:hdl
3371:doi
3317:hdl
3309:doi
3236:doi
3064:),
523:of
351:of
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