608:
1584:
Ethiopian wolf territories are largely stable, but packs can expand whenever the opportunity arises, such as when another pack disappears. The size of each territory correlates with the abundance of rodents, the number of wolves in a pack, and the survival of pups. Ethiopian wolves rest together in the open at night, and congregate for greetings and border patrols at dawn, noon, and evening. They may shelter from rain under overhanging rocks and behind boulders. The species never sleeps in dens, and only uses them for nursing pups. When patrolling their territories, Ethiopian wolves regularly scent-mark, and interact aggressively and vocally with other packs. Such confrontations typically end with the retreat of the smaller group.
1638:
1080:
1036:
1100:
1007:
978:
922:
893:
951:
855:
1688:
1560:
112:
623:
1674:. Solitary wolves hunt for rodents in the midst of the monkeys, ignoring juvenile monkeys, though these are similar in size to some of their prey. The monkeys, in turn, tolerate and largely ignore the wolves, although they take flight if they observe feral dogs, which sometimes prey on them. Within the troops, the wolves enjoy much higher success in capturing rodents than usual, perhaps because the monkeys' activities flush out the rodents, or because the presence of numerous larger animals makes it harder for rodents to spot a threat.
717:, Gojjam, and in every northern Afroalpine region where agriculture is well developed and human pressure acute. This revelation stressed the importance of the Bale Mountains wolf populations for the species' long-term survival, as well as the need to protect other surviving populations. A decade after the rabies outbreak, the Bale populations had fully recovered to pre-epizootic levels, prompting the species' downlisting to endangered in 2004, though it still remains the world's rarest canid, and Africa's most endangered
164:
1786:
2281:
1486:
139:
1654:... they are most amusing to watch, when hunting. The rats, which are brown, with short tails, live in big colonies and dart from burrow to burrow, while the cuberow stands motionless till one of them shows, when he makes a pounce for it. If he is unsuccessful, he seems to lose his temper, and starts digging violently; but this is only lost labour, as the ground is honeycombed with holes, and every rat is yards away before he has thrown up a pawful.
1605:
coat is gradually replaced with the adult colouration. By the age of five weeks, the pups feed on a combination of milk and solid food, and become completely weaned off milk at the age of 10 weeks to six months. All members of the pack contribute to protecting and feeding the pups, with subordinate females sometimes assisting the dominant female by suckling them. Full growth and sexual maturity are attained at the age of two years.
1450:
1183:
834:
510:
2355:) are usually agonistic, with Ethiopian wolves dominating African wolves if the latter enter their territories, and vice versa. Although African golden wolves are inefficient rodent hunters and thus not in direct competition with Ethiopian wolves, it is likely that heavy human persecution prevents the former from attaining numbers large enough to completely displace the latter.
1367:
285:
2228:
in single figures for several years. Habitat loss in the
Ethiopian highlands is directly linked to agricultural expansion into Afroalpine areas. In the northern highlands, human density is among the highest in Africa, with 300 people per km in some localities, with almost all areas below 3,700 m having been converted into
2413:
The overall aim of the EWCP is to protect the wolf's
Afroalpine habitat in Bale, and establish additional conservation areas in Menz and Wollo. The EWCP carries out education campaigns for people outside the wolf's range to improve dog husbandry and manage diseases within and around the park, as well
2267:
farmers, herders, woodcutters, and many others. In Bale there are numerous villages in and around the area, comprising over 8,500 households with more than 12,500 dogs. In 2007, the estimate of households within wolf habitat numbered 1,756. Because of the high number of dogs, the risk of infection in
2232:
fields. Suitable areas of land below this limit are under some level of protection, such as Guassa-Menz and the
Denkoro Reserve, or within the southern highlands, such as the Arsi and Bale Mountains. The most vulnerable wolf populations to habitat loss are those within relatively low-lying Afroalpine
2227:
During the 1990s, wolf populations in Gosh Meda and
Guguftu became extinct. In both cases, the extent of Afroalpine habitat above the limit of agriculture had been reduced to less than 20 km. The EWCP team confirmed the extinction of a wolf population in Mt. Guna in 2011, whose numbers had been
1579:
of six wolves are more common. Packs are formed by dispersing males and a few females, which with the exception of the breeding female, are reproductively suppressed. Each pack has a well-established hierarchy, with dominance and subordination displays being common. Upon dying, a breeding female can
1538:
to rusty red, with dense whitish to pale ginger underfur. The fur of the throat, chest and underparts is white, with a distinct white band occurring around the sides of the neck. There is a sharp boundary between the red coat and white marks. The ears are thickly furred on the edges, though naked on
1202:
Due to the high density of rodents in their new
Afroalpine habitat, the ancestors of the Ethiopian wolf gradually developed into specialised rodent hunters. This specialisation is reflected in the animal's skull morphology, with its very elongated head, long jaw, and widely spaced teeth. During this
2312:
were more available. The extinction of wolves in Mt. Choqa was likely due to persecution. Although people living close to wolves in modern times believe that wolf populations are recovering, negative attitudes towards the species persist due to livestock predation. Wolves were largely unmolested by
2262:
In the Simien
Mountains National Park, human and livestock populations are increasing by 2% annually, with further road construction allowing easy access to peasants into wolf home ranges; 3,171 people in 582 households were found to be living in the park and 1,477 outside the park in October 2005.
1583:
These packs live in communal territories, which encompass 6 km (2.3 sq mi) of land on average. In areas with little food, the species lives in pairs, sometimes accompanied by pups, and defends larger territories averaging 13.4 km (5.2 sq mi). In the absence of disease,
1547:
during the wet season (August–October), and there is no evident seasonal variation in coat colour, though the contrast between the red coat and white markings increases with age and social rank. Females tend to have paler coats than males. During the breeding season, the female's coat turns yellow,
3271:
Gopalakrishnan, Shyam; Sinding, Mikkel-Holger S.; Ramos-Madrigal, Jazmín; Niemann, Jonas; Samaniego
Castruita, Jose A.; Vieira, Filipe G.; Carøe, Christian; Montero, Marc de Manuel; Kuderna, Lukas; Serres, Aitor; González-Basallote, Víctor Manuel; Liu, Yan-Hu; Wang, Guo-Dong; Marques-Bonet, Tomas;
2397:
in 1983, with the Bale
Mountains Research Project being established shortly after. This was followed by a detailed, four-year field study, which prompted the IUCN/SSC Canid Specialist Group to produce an action plan in 1997. The plan called for the education of people in wolf-inhabited areas, wolf
1604:
is 60–62 days, with pups being born between
October and December. Pups are born toothless and with their eyes closed, and are covered in a charcoal-grey coat with a buff patch on the chest and abdomen. Litters consist of two to six pups, which emerge from their den after three weeks, when the dark
3208:
Koepfli, Klaus-Peter; Pollinger, John; Godinho, Raquel; Robinson, Jacqueline; Lea, Amanda; Hendricks, Sarah; Schweizer, Rena M.; Thalmann, Olaf; Silva, Pedro; Fan, Zhenxin; Yurchenko, Andrey A.; Dobrynin, Pavel; Makunin, Alexey; Cahill, James A.; Shapiro, Beth; Álvares, Francisco; Brito, José C.;
1717:
around 3,200 to 4,500 m, with some wolves inhabiting the Bale
Mountains being present in montane grasslands at 3,000 m. Although specimens were collected in Gojjam and northwestern Shoa at 2,500 m in the early 20th century, no recent records exist of the species occurring below 3,000 m. In modern
1539:
the inside. The naked borders of the lips, the gums and palate are black. The lips, a small spot on the cheeks and an ascending crescent below the eyes are white. The thickly furred tail is white underneath, and has a black tip, though, unlike most other canids, there is no dark patch marking the
3032:
Vonholdt, B. M.; Pollinger, J. P.; Lohmueller, K. E.; Han, E.; Parker, H. G.; Quignon, P.; Degenhardt, J. D.; Boyko, A. R.; Earl, D. A.; Auton, A.; Reynolds, A.; Bryc, K.; Brisbin, A.; Knowles, J. C.; Mosher, D. S.; Spady, T. C.; Elkahloun, A.; Geffen, E.; Pilot, M.; Jedrzejewski, W.; Greco, C.;
2389:
Steps taken to ensure the survival of the Ethiopian wolf include dog vaccination campaigns in Bale, Menz, and Simien, sterilization programs for wolf-dog hybrids in Bale, rabies vaccination of wolves in parts of Bale, community and school education programs in Bale and Wollo, contributing to the
1304:
to both. There are two genetically distinct African golden wolf populations that exist in northwestern and eastern Africa. This suggests that Ethiopian wolves – or a close and extinct relative – once had a much larger range within Africa to admix with other canids. There is evidence of gene flow
2325:. It was once hunted by sportsmen, though this is now illegal. Vehicle collisions killed at least four wolves in the Sanetti Plateau since 1988, while two others were left with permanent limps. Similar accidents are a risk in areas where roads cut across wolf habitats, such as in Menz and Arsi.
2291:
Although wolves in Bale have learned to use cattle to conceal their presence when hunting for rodents, the level of grazing in the area can adversely affect the vegetation available for the wolves' prey. Although no declines in wolf populations related to overgrazing have occurred, high grazing
2193:
The Ethiopian wolf has been considered rare since it was first recorded scientifically. The species likely has always been confined to Afroalpine habitats, so it was never widespread. In historical times, all of the Ethiopian wolf's threats are both directly and indirectly human-induced, as the
2333:
Management plans for hybridization with dogs involve sterilization of known hybrids. Incidences of Ethiopian wolf-dog hybridization have been recorded in Bale's Web Valley. At least four hybrids were identified and sterilized in the area. Although hybridization has not been detected elsewhere,
1259:
In 2015, a study of mitochondrial genome sequences and whole genome nuclear sequences of African and Eurasian canids indicated that extant wolf-like canids have colonised Africa from Eurasia at least five times throughout the Pliocene and Pleistocene, which is consistent with fossil evidence
1531:, males are larger than females, having 20% greater body mass. Adults measure 841–1,012 mm (33.1–39.8 in) in body length, and 530–620 mm (21–24 in) in height. Adult males weigh 14.2–19.3 kg (31–43 lb), while females weigh 11.2–14.15 kg (24.7–31.2 lb).
1260:
suggesting that much of African canid fauna diversity resulted from the immigration of Eurasian ancestors, likely coincident with Plio-Pleistocene climatic oscillations between arid and humid conditions. According to a phylogeny derived from nuclear sequences, the Eurasian golden jackal (
2210:
Rabies outbreaks, stemming from infected dogs, have killed many Ethiopian wolves over the 1990s and 2000s. Two well-documented outbreaks in Bale, one in 1991 and another in 2008–2009, resulted in the die-off or disappearance of 75% of known animals. Both incidents prompted reactive
2253:
or reduced fitness exists, the extremely small wolf population sizes, particularly those north of the Rift Valley, raise concerns among conservationists. Elsewhere, the Bale populations are fairly continuous, while those in Simien can still interbreed through habitat corridors.
1580:
be replaced by a resident daughter, though this increases the risk of inbreeding. Such a risk is sometimes circumvented by multiple paternity and extra-pack matings. The dispersal of wolves from their packs is largely restricted by the scarcity of unoccupied habitat.
660:
The Ethiopian wolf was recognised as requiring protection in 1938, and received it in 1974. The first in-depth studies on the species occurred in the 1980s with the onset of the American-sponsored Bale Mountains Research Project. Ethiopian wolf populations in the
3272:
Mirarab, Siavash; Fernandes, Carlos; Gaubert, Philippe; Koepfli, Klaus-Peter; Budd, Jane; Rueness, Eli Knispel; Heide-Jørgensen, Mads Peter; Petersen, Bent; Sicheritz-Ponten, Thomas; Bachmann, Lutz; Wiig, Øystein; Hansen, Anders J.; Gilbert, M. Thomas P. (2018).
1616:
Most females disperse from their natal pack at about two years of age, and some become "floaters" that may successfully immigrate into existing packs. Breeding pairs are most often unrelated to each other, suggesting that female-biased dispersal reduces
2249:, which is likely to increase with current rates of human expansion. The dangers posed by fragmentation include increased contact with humans, dogs, and livestock, and further risk of isolation and inbreeding in wolf populations. Although no evidence of
2510:
Martínez-Navarro, Bienvenido; Gossa, Tegenu; Carotenuto, Francesco; Bartolini-Lucenti, Saverio; Palmqvist, Paul; Asrat, Asfawossen; Figueirido, Borja; Rook, Lorenzo; Niespolo, Elizabeth M.; Renne, Paul R.; Herzlinger, Gadi; Hovers, Erella (2023-05-16).
2321:. However, they are perceived as threats to livestock elsewhere, with cases of retaliatory killings occurring in the Arsi Mountains. The Ethiopian wolf has not been recorded to be exploited for its fur, though in one case, wolf hides were used as
2263:
Although the area of the park has since been expanded, further settlement stopped, and grazing restricted, effective enforcement may take years. As of 2011, about 30,000 people live in 30 villages around and two within the park, including 4,650
1645:
Unlike most social carnivores, the Ethiopian wolf tends to forage and feed on small prey alone. It is most active during the day, the time when rodents are themselves most active, though they have been observed to hunt in groups when targeting
2338:
or a reduction in fitness, though this does not appear to have taken place. Due to the female's strong preference to avoid inbreeding, hybridization could be the result of not finding any males who are not close relatives outside of dogs.
708:
since the early 20th century, and in the Bale Mountains in the late 1950s. The status of the Ethiopian wolf was reassessed in the late 1990s, following improvements in travel conditions into northern Ethiopia. The surveys taken revealed
3868:
A sporting trip through Abyssinia : a narrative of a nine months' journey from the plains of the Hawash to the snows of Simien, with a description of the game, from elephant to ibex, and notes on the manners and customs of the
2082:
The local ericaceous forests and grasslands are under the protection of the Borena Saiynt Regional Park from agriculture as low as 3,200 m. The EWCP and the FZS have been involved in educational programs and wolf monitoring in the
1725:
The Ethiopian wolf uses all Afroalpine habitats, but has a preference for open areas containing short herbaceous and grassland communities inhabited by rodents, which are most abundant along flat or gently sloping areas with poor
3090:
Gottelli, D.; Sillero-Zubiri, C.; Applebaum, G. D.; Roy, M. S.; Girman, D. J.; Garcia-Moreno, J.; Ostrander, E. A.; Wayne, R. K. (1994). "Molecular genetics of the most endangered canid: The Ethiopian wolf Canis simensis".
1305:
between the eastern population and the Ethiopian wolf, which has led to the eastern population being distinct from the northwestern population. The common ancestor of both African golden wolf populations was a genetically
1508:
and has comparatively longer legs. Its skull is very flat, with a long facial region accounting for 58% of the skull's total length. The ears are broad, pointed, and directed forward. The teeth, particularly the
1669:
herds, a tactic thought to aid in ambushing rodents out of their holes by using the cattle to hide their presence. Ethiopian wolves have also been observed forming temporary associations with troops of grazing
642:) by Europeans and Ze Etiyopia by its citizens, wrote that the animal's skin was never worn by natives, as it was popularly believed that the wearer would die should any wolf hairs enter an open wound, while
2622:
1157:
approximately 1.8 million years ago. The Ethiopian wolf has survived numerous climatic changes in its Ethiopian highland habitat, with its range repeatedly expanding and contracting with glacial cycles.
1264:) diverged from the wolf/coyote lineage 1.9 million years ago, and with mitochondrial genome sequences indicating the Ethiopian wolf diverged from this lineage slightly prior to that. Further studies on
2219:
is not necessarily fatal to wolves, though a recent increase in infection has occurred, with outbreaks of canine distemper having been detected in 2005–2006 in Bale and in 2010 across subpopulations.
1153:
and dated to between 1.6 and 1.4 million years ago. The authors of this study state that the ancestors of the Ethiopian wolf arrived in Africa from Eurasia at the same time as the ancestors of the
2367:
appendices, though it is afforded full official protection under Ethiopia's Wildlife Conservation Regulations of 1974, Schedule VI, with the killing of a wolf carrying a two-year jail sentence.
2202:, road construction, and livestock farming), diseases (primarily rabies and canine distemper), conflict with humans (poisoning, persecution, and road kills), and hybridisation with dogs.
2398:
population monitoring, and the stemming of rabies in dog populations. The Ethiopian Wolf Conservation Programme was formed in 1995 by Oxford University, with donors including the
2194:
wolf's highland habitat, with its high annual rainfall and rich fertile soils, is ideal for agricultural activities. Its proximate threats include habitat loss and fragmentation (
1458:
This canid was initially classed as a distinct subspecies on account of its bright red coat, though this characteristic is unreliable as a taxonomic distinction. However, its
2124:
The wolf's range is protected by community resource management, and the Guassa Community Conservation Area. Educational campaigns are undertaken in schools near wolf ranges.
3261:. Master of Science Thesis. Center for Ecological and Evolutionary Synthesis Department of Bioscience Faculty of Mathematics and Natural Science, University of Oslo, Norway
3977:
2414:
as monitoring wolves in Bale, south and north Wollo. The program seeks to vaccinate up to 5,000 dogs a year to reduce rabies and distemper in wolf-inhabited areas.
2386:. Areas of suitable wolf habitat have recently increased to 87%, as a result of boundary extensions in Simien and the creation of the Arsi Mountains National Park.
2417:
In 2016, the Korean company Sooam Biotech was reported to be attempting to clone the Ethiopian wolf using dogs as surrogate mothers to help conserve the species.
3018:
1551:
Animals resulting from Ethiopian wolf-dog hybridisation tend to be more heavily built than pure wolves, and have shorter muzzles and different coat patterns.
1600:
involves the breeding male following the female closely. The breeding female only accepts the advances of the breeding male, or males from other packs. The
1757:, and short grasslands growing in shallow soils. In its northern range, the wolf's habitat is composed of plant communities characterised by a matrix of
2764:
3209:
Geffen, Eli; Leonard, Jennifer A.; Helgen, Kristofer M.; Johnson, Warren E.; o'Brien, Stephen J.; Van Valkenburgh, Blaire; Wayne, Robert K. (2015).
1534:
The Ethiopian wolf has short guard hairs and thick underfur, which provides protection at temperatures as low as −15 °C. Its overall colour is
3944:
3772:
9678:
389:
in size and build, and is distinguished by its long and narrow skull, and its red and white fur. Unlike most large canids, which are widespread,
1662:, with the level of effort varying from scratching lightly at the hole to totally destroying a set of burrows, leaving metre-high earth mounds.
8885:
3999:
3794:
681:
in 1994. The IUCN/SSC Canid Specialist Group advocated a three-front strategy of education, wolf population monitoring, and rabies control in
9756:
9730:
8956:
2697:
3995:
415:
at altitudes of 3,000–4,500 m, with the overall adult population estimated at 360–440 individuals in 2011, more than half of them in the
1521:
are relatively small. The Ethiopian wolf has eight mammae, of which only six are functional. The front paws have five toes, including a
653:. Since then, it was scarcely heard of in Europe up until the early 20th century, when several skins were shipped to England by Major
9879:
9849:
1145:
The earliest known fossil of the Ethiopian wolf is known from the Melka Wakena paleoanthropological site-complex in the Southeastern
3141:
Dalton, R. (2001). The skull morphology of the Ethiopian wolf (Canis simensis). B.Sc.thesis. University of Edinburgh, Edinburgh, UK.
1575:
The Ethiopian wolf is a social animal, living in family groups containing up to 20 adults (individuals older than one year), though
9652:
3899:
700:, the paucity of information stemming from that area indicated that the species was likely declining there, while reports from the
607:
4020:
4007:
9704:
1265:
3412:"The physiology of cooperative breeding in a rare social canid; sex, suppression and pseudopregnancy in female Ethiopian wolves"
2390:
running of national parks, and population monitoring and surveying. A 10-year national action plan was formed in February 2011.
9748:
3912:
2667:
9709:
669:, though the critical state of the species was revealed during the early 1990s after a combination of shooting and a severe
9894:
2445:
This is in reference to the Ethiopian wolf's reported habit of following mares and cows about to give birth to feed on the
523:
9884:
4055:
527:
9889:
9869:
3770:
Foraging ecology and trophic niche overlap between sympatric African wolf and Ethiopian wolf in the Ethiopian Highlands
3033:
Randi, E.; Bannasch, D.; Wilton, A.; Shearman, J.; Musiani, M.; Cargill, M.; Jones, P. G.; Qian, Z.; Huang, W. (2010).
2893:
2379:
1211:, which caused the species' Afroalpine habitat to fragment, thus isolating Ethiopian wolf populations from each other.
5425:
2765:"The effect of the last glacial age on speciation and population genetic structure of the endangered Ethiopian wolf (
390:
143:
9761:
5769:
1821:
3821:
696:. Elsewhere, information on Ethiopian wolves remained scarce; although first described in 1835 as living in the
9864:
2459:
2375:
1982:
3741:
2480:
2334:
scientists are concerned that it could pose a threat to the wolf population's genetic integrity, resulting in
400:
with very specific habitat requirements. It is one of the world's rarest canids, and Africa's most endangered
9826:
7987:
2815:
2407:
2394:
1809:
9874:
3561:"Wolves are better hunters when monkeys are around: An unexpected co-existence in the Ethiopian highlands."
2403:
2383:
2371:
2049:
1637:
1169:
may be an evolutionary relic of a gray wolf-like ancestor's past invasion of northern Africa from Eurasia.
662:
9743:
9691:
9608:
7978:
5181:
3895:
1877:
1801:
1659:
1381:
1323:
3575:"Solitary Ethiopian wolves increase predation success on rodents when among grazing gelada monkey herds"
1165:
analysis showed a closer relationship to the gray wolf and the coyote than to other African canids, and
9443:
9099:
3769:
1841:
431:
7465:
6304:
3742:"Field report : Association between an Arabian wolf and a domestic dog in central Saudi Arabia"
2878:
9581:
9362:
9257:
7996:
6286:
6014:
5981:
5957:
5778:
5030:
2246:
1852:
leaves have occasionally been found in Ethiopian wolf stomachs. The sedge possibly is ingested for
3004:
Life in Abyssinia: Being Notes Collected During Three Years' Residence and Travels in that Country
2736:
2607:
685:. The establishment of the Ethiopian Wolf Conservation Programme in Bale soon followed in 1995 by
163:
9024:
8005:
7736:
7429:
7364:
6746:
6322:
5795:
5317:
4048:
3560:
2195:
2133:
1813:
1719:
1496:
The Ethiopian wolf is similar in size and build to North America's coyote; it is larger than the
1285:
1204:
430:, on account of its small numbers and fragmented range. Threats include increasing pressure from
9800:
1777:
moorlands at 3,200–3,600 m in Simien may provide a refuge for wolves in highly disturbed areas.
1079:
1035:
9683:
9543:
9248:
9000:
8813:
8689:
8301:
8283:
8111:
7805:
7796:
7438:
6176:
4968:
3891:
3003:
2335:
2177:
1999:
As of 2011, no sightings have been made, despite intensive monitoring during two field visits.
1770:
1687:
1099:
1006:
459:
2655:
2636:
2632:
977:
921:
892:
9795:
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8662:
8653:
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8129:
8120:
7492:
6262:
6253:
6244:
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6131:
6113:
6104:
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4256:
3573:
Venkataraman, V. V.; Kerby, J. T.; Nguyen, N.; Ashenafi, Z. T.; Fashing, P. J. (2015-03-27).
2513:"The earliest Ethiopian wolf: implications for the species evolution and its future survival"
2250:
1861:
1622:
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1241:
950:
854:
678:
258:
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3563:
ScienceDaily., 22 June 2015. <www.sciencedaily.com/releases/2015/06/150622085224.htm>.
3211:"Genome-wide Evidence Reveals that African and Eurasian Golden Jackals Are Distinct Species"
1856:
or for parasite control. The species may scavenge on carcasses, but is usually displaced by
1773:, all of which are favoured by the wolf's rodent prey. Although marginal in importance, the
9639:
9590:
9266:
9167:
9126:
8840:
8351:
8079:
7405:
6470:
6220:
6211:
6167:
6122:
5745:
5251:
4986:
4812:
4015:
3985:
3629:
3285:
3222:
3100:
3046:
3035:"Genome-wide SNP and haplotype analyses reveal a rich history underlying dog domestication"
2849:
2783:
2399:
1845:
1805:
1606:
1245:
1229:
1191:
111:
9722:
3462:"Inbreeding is reduced by female-biased dispersal and mating behavior in Ethiopian wolves"
8:
9506:
9452:
9224:
8671:
8557:
8539:
8384:
8233:
8138:
7388:
7334:
7083:
6825:
6816:
6080:
5353:
5335:
5190:
5039:
4542:
3862:
3852:
2426:
2370:
The species is present in several protected areas, including three areas in South Wollo (
2348:
2305:
1789:
1597:
1501:
1497:
1360:
1293:
1269:
1253:
1146:
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654:
435:
378:
128:
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3460:
Randall, DA; Pollinger, JP; Wayne, RK; Tallents, LA; Johnson, PJ; Macdonald, DW (2007).
3289:
3226:
3104:
3050:
2787:
2545:
2512:
1722:, which extends up to 3,700 m, has largely restricted the species to the highest peaks.
1650:
calves. Major Percy-Cotton described the hunting behaviour of Ethiopian wolves as thus:
1621:. Inbreeding is ordinarily avoided because it leads to a reduction in progeny fitness (
9595:
9321:
8548:
8070:
7760:
7483:
7456:
6719:
6461:
6313:
5416:
5308:
5208:
5148:
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4146:
4041:
4004:
3624:
3518:
3442:
3306:
3273:
3124:
3112:
3067:
3034:
2998:
2807:
2728:
2720:
2245:
Some Ethiopian wolf populations, particularly those in North Wollo, show signs of high
1224:
by its larger size, relatively longer legs, distinct reddish coat, and white markings.
467:
297:
158:
1978:
encroachment into rodent habitat, disease, and competition/predation by golden wolves
1969:
The second-largest population, the most genetically diverse, is a tourist attraction.
622:
9782:
9616:
9200:
9117:
8897:
8393:
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7284:
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6513:
6295:
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6023:
5907:
5260:
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3434:
3411:
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2732:
2663:
2550:
2532:
2053:
2014:
An ORDA Biodiversity Conservation Project is active in the area, in conjunction with
1540:
1306:
1162:
736:
732:
686:
455:
3877:
3446:
3128:
2811:
2584:
1372:
631:
274:
9621:
9479:
9312:
8590:
8020:
7310:
7092:
6842:
6032:
5931:
5715:
5706:
5290:
5081:
5072:
4866:
4857:
4415:
3955:
3848:
3586:
3502:
3473:
3426:
3377:
3301:
3293:
3230:
3188:
3108:
3062:
3054:
2886:
Canids: Foxes, Wolves, Jackals, and Dogs:Status Survey and Conservation Action Plan
2791:
2712:
2602:
2540:
2524:
2216:
2105:
1951:
1889:
1881:
1857:
1817:
1692:
1601:
1385:
1225:
1194:
has resulted in a skull similar in shape to that of jackals and the South American
1154:
1031:
710:
697:
639:
555:
451:
3991:
3522:
3430:
2479:
For a full set of supporting references refer to the note (a) in the phylotree at
638:. European writers traveling in Ethiopia during the mid-19th century (then called
284:
9859:
9813:
9769:
9497:
9488:
9416:
9371:
9033:
8983:
8974:
8849:
8795:
8739:
8608:
8581:
8474:
8292:
8251:
8216:
7961:
7024:
6737:
6704:
6695:
6202:
5940:
5389:
4839:
4766:
4724:
4460:
4265:
4199:
4176:
4011:
3776:
1744:
1610:
674:
517:
408:
120:
9735:
2884:. In Sillero-Zubiri, Claudio; Hoffmann, Michael; Macdonald, David Whyte (eds.).
2509:
2147:
The third-largest population, in the second-largest Afroalpine area in Ethiopia
1626:
1203:
period, the species likely attained its highest abundance, and had a relatively
9854:
9470:
9389:
9191:
8938:
8831:
8804:
7857:
7474:
7074:
6621:
6158:
6140:
6071:
5682:
5673:
5275:
5236:
4190:
4102:
3883:
Neue Wirbelthiere zu der Fauna von Abyssinien gehörig, entdeckt und beschrieben
2528:
2322:
2158:
2114:
A core population with ideal habitat, it is increasingly a tourist attraction.
2045:
Isolation, habitat degradation, human-wildlife conflict, and road encroachment
1913:
1833:
1647:
1564:
1470:
643:
635:
580:
416:
412:
3381:
3297:
3235:
3210:
1840:
calves. In areas where the big-headed African mole-rat is absent, the smaller
9843:
9717:
9566:
9434:
9330:
9303:
8929:
8627:
8572:
8094:
8038:
7779:
7420:
6875:
6437:
5602:
5569:
5371:
5362:
5299:
5199:
5157:
5009:
4602:
3591:
3574:
2593:
2536:
2296:
and vegetation deterioration in Afroalpine areas such as Delanta and Simien.
2182:
Most of the species' habitat occurs within the Bale Mountains National Park.
2129:
1972:
Human disturbance, road traffic, extensive agriculture, habitat degradation,
1909:
1785:
1743:
and grasses, with low vegetation cover. Other favourable habitats consist of
1593:
1576:
1297:
946:
705:
626:
Mounted specimen (1902), one of the first post-1835 specimens to reach Europe
447:
148:
62:
6059:
5871:
3478:
3461:
3363:"Scent-marking and territorial behaviour of Ethiopian wolves Canis simensis"
3177:"A review of the family Canidae, with a classification by numerical methods"
2280:
2011:, isolation, and possible competition with abundant golden wolf populations
689:, in conjunction with the Ethiopian Wildlife Conservation Authority (EWCA).
9696:
9461:
9407:
9108:
8876:
8822:
8599:
8498:
8053:
8029:
7848:
7839:
7659:
7626:
7549:
7510:
6920:
6902:
6764:
6636:
6095:
5496:
5459:
5344:
5326:
4739:
4685:
4611:
4289:
4033:
3514:
3493:
Charlesworth D, Willis JH (2009). "The genetics of inbreeding depression".
3438:
3315:
3244:
3076:
2803:
2662:. Vol. 1 (3rd ed.). Johns Hopkins University Press. p. 577.
2554:
2293:
2268:
local wolf populations is high. Furthermore, intentional and unintentional
2212:
2008:
1514:
1505:
1208:
917:
3270:
3193:
3120:
2056:, which is to encompass about a third of the wolf's range in North Wollo.
9774:
9665:
9575:
9075:
8266:
8183:
8156:
7905:
7819:
7751:
7677:
7668:
7635:
7593:
7558:
7275:
7168:
7000:
6540:
6452:
6360:
6005:
5578:
5398:
5063:
4977:
4960:
4937:
4900:
4784:
4748:
4593:
4382:
4237:
4116:
3830:
3829:. IUCN/SSC Canid Specialist Group. Oxford, United Kingdom. Archived from
3739:
2199:
2111:
Although diminishing from disease, the population is healthy and stable.
2076:
1974:
1905:
1753:
1485:
1248:
refuted the separate genus in favour of placing the species in the genus
714:
463:
439:
394:
37:
9631:
3259:
Can hybridization be detected between African wolf and sympatric canids?
3058:
9657:
9284:
8530:
8461:
8448:
8415:
8201:
7939:
7699:
7247:
7224:
7110:
7101:
7050:
7015:
6991:
6755:
6654:
6603:
6549:
6531:
6504:
6411:
6361:
5760:
5733:
5661:
5644:
5407:
5380:
5226:
5130:
5114:
4950:
4830:
4670:
4506:
4373:
4165:
4133:
4117:
2943:
2724:
2446:
2269:
1990:
1901:
1868:, with farmers often leaving herds in wolf-inhabited areas unattended.
1829:
1736:
1699:
1618:
1548:
becomes woolier, and the tail turns brownish, losing much of its hair.
1518:
1459:
1389:
1195:
646:
423:
82:
47:
1175:
9670:
9215:
9150:
8763:
8703:
8489:
8310:
7925:
7917:
7866:
7716:
7602:
7584:
7298:
7132:
6965:
6807:
6773:
6585:
6277:
5551:
5021:
4497:
4065:
3176:
2467:
1865:
1774:
1748:
1714:
1706:
1702:
1466:
1187:
753:
718:
650:
401:
215:
175:
87:
31:
9537:
3506:
2716:
1908:
for several decades. Southeast of the Rift Valley, it occurs in the
1207:. This changed about 15,000 years ago with the onset of the current
692:
Soon after, a further wolf population was discovered in the Central
677:. In response, the IUCN reclassified the species from endangered to
9560:
8912:
8375:
8192:
8168:
7529:
7379:
7325:
7261:
7256:
7209:
7141:
7059:
6929:
6678:
6669:
6567:
6522:
6402:
6235:
5972:
5881:
5850:
5810:
5697:
5629:
5537:
5528:
4848:
4644:
4397:
4280:
4247:
4096:
4084:
2309:
1853:
1837:
1727:
1544:
1510:
1150:
443:
195:
77:
72:
57:
52:
42:
3089:
2022:
governments, and supported by the Guna Highland Water enterprise.
1844:
is targeted. In the Simien Mountains, the Ethiopian wolf preys on
9821:
9294:
9141:
9015:
8730:
8242:
7572:
7038:
6594:
6558:
6425:
6393:
6377:
5890:
5593:
5172:
4653:
4560:
4478:
4406:
4348:
4325:
4218:
3981:
3274:"Interspecific Gene Flow Shaped the Evolution of the Genus Canis"
2660:
Mammal Species of the World: A Taxonomic and Geographic Reference
2234:
2084:
1765:
1759:
1710:
1522:
1449:
749:
374:
225:
92:
67:
3972:
3619:
3617:
2762:
1900:. It has recently become extinct in Gosh Meda in north Shoa and
704:
plateau were a century out of date. Wolves were recorded in the
9644:
9603:
9398:
9345:
9090:
8778:
8512:
7232:
6893:
6728:
6485:
5922:
4694:
4620:
4569:
4551:
4524:
4445:
4307:
4090:
4078:
4027:
3819:
3795:"Inside the cloning factory that creates 500 new animals a day"
3727:
3715:
3703:
3691:
3679:
3667:
3655:
3643:
3608:
3572:
3535:
3397:
3348:
3333:
3207:
3150:
2930:
2836:
2264:
2229:
2073:
After Simien, the second-largest area north of the Rift Valley
2019:
2015:
1955:
1885:
1876:
Six current Ethiopian wolf populations are known. North of the
1828:
and eggs. Ethiopian wolves have twice been observed to feed on
1671:
1666:
1462:
are consistently longer than those of the nominate subspecies.
1221:
1220:
species present in Africa, and is readily distinguishable from
1182:
888:
701:
670:
397:
386:
205:
185:
3901:
The Ethiopian Wolf: Status Survey and Conservation Action Plan
3031:
2578:
2576:
2574:
2572:
2570:
2568:
2566:
2564:
393:
feeders, the Ethiopian wolf is a highly specialised feeder of
9808:
9380:
9239:
9182:
9055:
8858:
8715:
8342:
8324:
7947:
7617:
6866:
6851:
5512:
4803:
4298:
3614:
3459:
2364:
2314:
2150:
Habitat degradation, expanding agriculture, and road traffic
2096:
2028:
1996:
An isolated patch, estimated at less than 20 km by 2004
1897:
1849:
1825:
1800:
In the Bale Mountains, the Ethiopian wolf's primary prey are
1734:. Prime wolf habitat in the Bale Mountains consists of short
1535:
1527:
1397:
1233:
1216:
1002:
833:
478:
Alternative English names for the Ethiopian wolf include the
235:
2582:
2393:
The species' critical situation was first publicised by the
1525:, while the hind paws have four. As is typical in the genus
673:
epidemic decimated most packs studied in the Web Valley and
8754:
7650:
6975:
6911:
4584:
4316:
4174:
2763:
Gottelli, D; Marino, J; Sillero-Zubiri, C; Funk, S (2004).
2561:
2318:
2285:
2092:
1893:
1740:
1731:
1393:
1310:
850:
427:
1658:
The technique described above is commonly used in hunting
1366:
630:
The species was first scientifically described in 1835 by
6884:
6493:
4821:
2463:
2342:
2272:
are frequent in the ericaceous moorlands wolves inhabit.
2173:
The largest population, with the highest density of prey
828:
682:
3174:
2304:
Direct killings of wolves were more frequent during the
1698:
The Ethiopian wolf is restricted to isolated pockets of
1186:
Ethiopian wolf skull: Despite its close relation to the
665:
were negatively affected by the political unrest of the
3996:
IUCN Species Survival Commission#Canid Specialist Group
3890:
3360:
3019:
The Variation of Animals and Plants Under Domestication
2915:
2888:. IUCN-The World Conservation Union. pp. 167–174.
2378:, and Borena Sayint Regional Park), one in north Shoa (
2313:
humans in Bale, as they were not considered threats to
2104:
As of 2010, an estimated 40% have been missing since a
2079:, ploughing, persecution, and local negative attitudes
2695:
2257:
2064:
Patches connected by corridors, totalling 243 km
1960:
Patches connected by corridors, totalling 273 km
3566:
3361:
Sillero-Zubiri, Claudio; Macdonald, David W. (1998).
1275:
1268:
found instances of Ethiopian wolves hybridizing with
616:
Neue Wirbelthiere zu der Fauna von Abyssinien gehörig
7715:
5111:
2876:
2872:
2870:
2299:
1570:
381:. In southeastern Ethiopia, it is also known as the
3886:(in German). Frankfurt am Main : S. Schmerber.
3740:Chris Barichievy; Shayne Clugston; Robert Sheldon.
3175:Clutton-Brock, J.; Corbet, G.G.; Hills, M. (1976).
3025:
2121:encroachment into rodent habitat, and road traffic
1796:), one of the Ethiopian wolf's primary prey animals
1232:originally classified the species under a separate
3823:Strategic Planning for Ethiopian Wolf Conservation
3492:
3264:
2153:Protected within the Arsi Mountains Regional Park
1665:Wolves in Bale have been observed to forage among
3978:Conservation of Ethiopian wolves (Canis simensis)
2867:
2040:Possibly declining, otherwise stable, as of 1998
1596:usually takes place between August and November.
9841:
5511:
2649:
2647:
2645:
1641:Southern Ethiopian wolf feeding, Bale Mountains.
1300:, and with the Ethiopian wolf being genetically
3786:
2034:Patchily distributed in an area of 140 km
1985:, and has been monitored regularly since 2003.
1587:
1244:considered it to be "only an exaggerated fox".
614:by Friedrich Carl Vogel (from Eduard Rüppell's
4000:International Union for Conservation of Nature
2473:
2458:This was later proven incorrect in 2010, when
6375:
4216:
4049:
3861:
3547:
3162:
2986:
2642:
2608:10.2305/IUCN.UK.2011-1.RLTS.T3748A10051312.en
2052:and its associates are working to create the
1629:expression of deleterious recessive alleles.
1321:As of 2005, two subspecies are recognised by
4063:
3973:Ethiopian Wolf Conservation Programme (EWCP)
3486:
3409:
1252:, upon noting cranial similarities with the
462:, which seeks to protect the wolves through
2944:"Ethiopia | International Wolf Center"
2284:Ethiopian wolves resting alongside grazing
2240:
1709:inhabited by Afroalpine rodents. Its ideal
1517:measure 14–22 mm in length, while the
4236:
4056:
4042:
2758:
2756:
2658:. In Wilson, D. E.; Reader, D. M. (eds.).
2653:
2505:
2503:
2501:
2499:
2328:
283:
137:
110:
8459:
4948:
4131:
3820:IUCN; SSC Canid Specialist Group (2011).
3590:
3477:
3410:van Kesteren, Freya; et al. (2013).
3393:
3391:
3305:
3234:
3192:
3066:
2696:Sillero-Zubiri, C.; Gottelli, D. (1994).
2606:
2544:
1288:was used to compare members of the genus
724:
6805:
3847:
3673:
3604:
3602:
3329:
3327:
3325:
2971:
2691:
2689:
2687:
2685:
2683:
2681:
2679:
2363:The Ethiopian wolf is not listed on the
2279:
2054:Abuna Yoseph Community Conservation Area
1871:
1784:
1686:
1636:
1613:have been observed in Ethiopian wolves.
1558:
1484:
1181:
621:
606:
5879:
4642:
4163:
3876:
3792:
3344:
3342:
2982:
2980:
2967:
2965:
2956:
2926:
2924:
2909:
2877:Sillero-Zubiri, C.; Marino, J. (2004).
2753:
2628:
2583:Marino, J.; Sillero-Zubiri, C. (2011).
2496:
2233:ranges, such as those in Aboi Gara and
1816:. Other secondary prey species include
528:question marks, boxes, or other symbols
9842:
8972:
7208:
3529:
3388:
3144:
2832:
2830:
2828:
2481:Evolution of the wolf#Wolf-like canids
2343:Competition with African golden wolves
2141:54 wolves in 9 packs, as of 2007-2010
9542:
9541:
8446:
7903:
7207:
6359:
5494:
4935:
4115:
4037:
3721:
3709:
3697:
3685:
3661:
3649:
3637:
3599:
3322:
2676:
2215:in 2003 and 2008–2009, respectively.
2108:outbreak was detected in local dogs.
1632:
602:
9827:FE981C90-6153-4BC5-89C2-F3A999EF5A11
3728:IUCN/SSC Canid Specialist Group 2011
3716:IUCN/SSC Canid Specialist Group 2011
3704:IUCN/SSC Canid Specialist Group 2011
3692:IUCN/SSC Canid Specialist Group 2011
3680:IUCN/SSC Canid Specialist Group 2011
3668:IUCN/SSC Canid Specialist Group 2011
3656:IUCN/SSC Canid Specialist Group 2011
3644:IUCN/SSC Canid Specialist Group 2011
3609:IUCN/SSC Canid Specialist Group 2011
3541:
3536:IUCN/SSC Canid Specialist Group 2011
3398:IUCN/SSC Canid Specialist Group 2011
3349:IUCN/SSC Canid Specialist Group 2011
3339:
3334:IUCN/SSC Canid Specialist Group 2011
3156:
3151:IUCN/SSC Canid Specialist Group 2011
2977:
2962:
2931:IUCN/SSC Canid Specialist Group 2011
2921:
2837:IUCN/SSC Canid Specialist Group 2011
530: instead of Ethiopic characters.
16:Canine native to Ethiopian Highlands
5426:South African small-spotted genet (
3907:. IUCN/SSC Canid Specialist Group.
2950:
2916:Sillero-Zubiri & MacDonald 1997
2825:
2594:IUCN Red List of Threatened Species
2258:Encroachment within protected areas
2176:Disease (rabies and distemper) and
2170:Declining, but stable in long term
1513:, are small and widely spaced. The
501:
13:
3937:
3113:10.1111/j.1365-294X.1994.tb00070.x
3007:. Vol. II. John Murray. pp. 12–13.
2380:Guassa Community Conservation Area
1864:. It typically poses no threat to
1214:The Ethiopian wolf is one of five
14:
9906:
5770:Broad-striped Malagasy mongoose (
3966:
3793:Zastrow, Mark (8 February 2016).
3625:"Canis simensis (Ethiopian wolf)"
2300:Human persecution and disturbance
2292:intensities are known to lead to
1571:Social and territorial behaviours
1313:and 28% Ethiopian wolf ancestry.
739:with timing in millions of years
657:during his travels in Abyssinia.
9880:Fauna of the Ethiopian Highlands
9850:IUCN Red List endangered species
5495:
2796:10.1111/j.1365-294x.2004.02226.x
1822:yellow-spotted brush-furred rats
1448:
1365:
1098:
1078:
1034:
1005:
976:
949:
920:
891:
853:
832:
508:
454:. Its conservation is headed by
422:The Ethiopian wolf is listed as
162:
35:
7904:
4936:
3992:Ethiopian Wolf (Canis simensis)
3762:
3733:
3553:
3453:
3403:
3354:
3251:
3201:
3168:
3135:
3083:
3010:
2992:
2936:
2452:
2439:
2358:
2222:
2167:About 250 adults and subadults
1981:The entire range is within the
1904:, and has not been reported in
1691:Northern Ethiopian wolf in the
1563:Southern Ethiopian wolf in the
1332:
634:, who provided a skull for the
8447:
3813:
2842:
2376:Simien Mountains National Park
2275:
2101:A single patch of 112 km
1983:Simien Mountains National Park
1810:black-clawed brush-furred rats
1480:
1408:(Erlanger & Neumann, 1900)
649:that the species gave rise to
1:
5182:Malabar large-spotted civet (
3872:. London : Rowland Ward.
3431:10.1016/j.physbeh.2013.08.016
3022:. Vol. I. Orange Judd. p. 48.
2489:
2408:Wildlife Conservation Network
2395:Wildlife Conservation Society
1920:Summary of current status of
1836:calves. It will also prey on
1316:
411:is limited to seven isolated
9444:Indonesian mountain weasel (
9100:North American river otter (
7988:Humboldt's hog-nosed skunk (
5042:Macrogalidia musschenbroekii
3949:National Geographic Magazine
2404:Frankfurt Zoological Society
2384:Arsi Mountains National Park
2372:Bale Mountains National Park
2050:Frankfurt Zoological Society
1880:, the species occurs in the
1802:big-headed African mole-rats
1660:big-headed African mole-rats
1588:Reproduction and development
1554:
1149:. It is the right half of a
663:Bale Mountains National Park
7:
9895:Ethiopian montane moorlands
6305:Somalian slender mongoose (
3857:. London, R. Ward, limited.
3181:Bull. Br. Mus. (Nat. Hist.)
2420:
2117:Human disturbance, rabies,
1324:Mammal Species of the World
432:expanding human populations
10:
9911:
9885:Endangered fauna of Africa
9258:Asian small-clawed otter (
7997:American hog-nosed skunk (
7979:Molina's hog-nosed skunk (
6287:Angolan slender mongoose (
5982:Ethiopian dwarf mongoose (
5031:Small-toothed palm civet (
3854:The Game Animals of Africa
2654:Wozencraft, C. W. (2005).
2529:10.1038/s42003-023-04908-w
2205:
2188:
1934:Population size estimates
1842:Northeast African mole-rat
1804:, though it also feeds on
1794:Tachyoryctes macrocephalus
1747:grasslands, high-altitude
1682:
1677:
310:Erlanger and Neumann, 1900
9890:Mammals described in 1840
9870:Endemic fauna of Ethiopia
9550:
9343:
9292:
9283:
9237:
9213:
9180:
9163:
9139:
9088:
9071:
9054:
9025:Saharan striped polecat (
9013:
8996:
8955:
8910:
8893:
8884:
8875:
8776:
8752:
8728:
8711:
8702:
8625:
8570:
8528:
8511:
8487:
8470:
8455:
8442:
8414:
8373:
8340:
8323:
8264:
8214:
8181:
8164:
8155:
8092:
8051:
8018:
8006:Striped hog-nosed skunk (
7959:
7938:
7916:
7912:
7899:
7835:
7818:
7777:
7749:
7737:Mediterranean monk seal (
7732:
7698:
7648:
7615:
7580:
7571:
7545:
7528:
7509:
7466:Juan Fernández fur seal (
7430:South American fur seal (
7418:
7401:
7377:
7365:South American sea lion (
7360:
7323:
7306:
7297:
7273:
7245:
7223:
7219:
7203:
7167:
7140:
7131:
7072:
7046:
7037:
7013:
6989:
6974:
6963:
6864:
6838:
6790:
6747:South American gray fox (
6717:
6691:
6667:
6650:
6635:
6620:
6483:
6450:
6433:
6424:
6391:
6371:
6355:
6323:Common slender mongoose (
6275:
6233:
6189:
6093:
6067:
6058:
6003:
5970:
5953:
5920:
5903:
5870:
5849:
5808:
5796:Narrow-striped mongoose (
5791:
5758:
5741:
5732:
5695:
5669:
5660:
5643:
5627:
5591:
5565:
5550:
5526:
5507:
5503:
5490:
5458:
5454:
5318:Crested servaline genet (
5273:
5234:
5225:
5170:
5144:
5129:
5107:
5061:
5017:
5008:
4959:
4944:
4931:
4899:
4801:
4737:
4720:
4711:
4683:
4666:
4633:
4582:
4495:
4458:
4441:
4432:
4395:
4371:
4362:
4346:
4278:
4245:
4232:
4212:
4159:
4142:
4127:
4111:
4073:
3419:Physiology & Behavior
3382:10.1017/s0952836998007134
3298:10.1016/j.cub.2018.08.041
3236:10.1016/j.cub.2015.06.060
1292:. The study supports the
1093:
1073:
1066:
1058:
1029:
1000:
973:
944:
915:
886:
848:
826:
819:
812:
805:
797:
789:
781:
773:
765:
757:
473:
303:
296:
291:
282:
264:
257:
159:Scientific classification
157:
135:
126:
118:
109:
23:
9381:Stoat/Beringian ermine (
9249:African clawless otter (
9001:African striped weasel (
8814:Yellow-throated marten (
8690:Formosan ferret-badger (
8394:Western mountain coati (
8385:Eastern mountain coati (
8302:Western lowland olingo (
8284:Eastern lowland olingo (
8112:Southern spotted skunk (
7806:Southern elephant seal (
7797:Northern elephant seal (
7587:Pagophilus groenlandicus
6364:("dog-like" carnivorans)
6177:Stripe-necked mongoose (
5798:Mungotictis decemlineata
4673:Herpailurus yagouaroundi
4120:("cat-like" carnivorans)
3956:Ethiopian wolves on high
3016:Darwin, Charles (1868).
2466:'s sole ancestor is the
2462:studies showed that the
2432:
2241:Population fragmentation
1296:being distinct from the
544:Linguistic group or area
9426:Yellow-bellied weasel (
8681:Burmese ferret-badger (
8663:Chinese ferret-badger (
8654:Bornean ferret badger (
8645:Vietnam ferret-badger (
8130:Eastern spotted skunk (
8121:Western spotted skunk (
7851:Leptonychotes weddellii
7493:Subantarctic fur seal (
7439:Australasian fur seal (
6263:Black-footed mongoose (
6245:Bushy-tailed mongoose (
6194:White-tailed mongoose (
6132:Indian brown mongoose (
6114:Short-tailed mongoose (
6105:Small Indian mongoose (
6042:Flat-headed kusimanse (
6015:Alexander's kusimanse (
5991:Common dwarf mongoose (
5958:Pousargues's mongoose (
5820:Brown-tailed mongoose (
5779:Grandidier's mongoose (
5033:Arctogalidia trivirgata
4257:Sunda clouded leopard (
3892:Sillero-Zubiri, Claudio
3863:Powell-Cotton, P. H. G.
2329:Hybridisation with dogs
2196:subsistence agriculture
1780:
1720:subsistence agriculture
1713:extends from above the
1465:Southeast Rift Valley:
1441:Southern Ethiopian wolf
1352:Northern Ethiopian wolf
1286:whole genome sequencing
1205:continuous distribution
385:. It is similar to the
9267:Congo clawless otter (
9170:Hydrictis maculicollis
9168:Spotted-necked otter (
9127:Southern river otter (
9078:Pteronura brasiliensis
8841:European pine marten (
8352:South American coati (
8080:Palawan stink badger (
7406:New Zealand sea lion (
6471:Japanese raccoon dog (
6168:Crab-eating mongoose (
6123:Indian grey mongoose (
5746:Ring-tailed vontsira (
5557:(bone-crushing hyenas)
5252:Central African oyan (
4813:Chinese mountain cat (
3781:The Rufford Foundation
3592:10.1093/jmammal/gyu013
2879:"6.4 – Ethiopian wolf"
2517:Communications Biology
2336:outbreeding depression
2288:
2178:agricultural expansion
2061:South Wollo highlands
1888:, in the northern and
1797:
1695:
1656:
1642:
1567:
1493:
1199:
725:Taxonomy and evolution
627:
619:
516:This section contains
460:Conservation Programme
407:The species's current
119:Ethiopian wolf on the
9865:Carnivorans of Africa
9796:Paleobiology Database
9507:Back-striped weasel (
9453:Black-footed ferret (
9225:Smooth-coated otter (
9003:Poecilogale albinucha
8672:Javan ferret-badger (
8558:Sumatran hog badger (
8540:Northern hog badger (
8234:Crab-eating raccoon (
8139:Pygmy spotted skunk (
7860:Lobodon carcinophagus
7596:Histriophoca fasciata
7389:Australian sea lion (
7335:California sea lion (
7084:American black bear (
6826:Black-backed jackal (
6817:Side-striped jackal (
6657:Chrysocyon brachyurus
6081:Long-nosed mongoose (
5354:Rusty-spotted genet (
5191:Large-spotted civet (
5040:Sulawesi palm civet (
4969:Owston's palm civet (
4543:Andean mountain cat (
3988:Department of Zoology
3960:BBC Wildlife Magazine
3479:10.1093/beheco/arm010
3194:10.5962/bhl.part.6922
2349:African golden wolves
2283:
2251:inbreeding depression
1872:Range and populations
1862:African golden wolves
1846:Abyssinian grass rats
1788:
1690:
1652:
1640:
1625:) due largely to the
1623:inbreeding depression
1562:
1491:Louis Agassiz Fuertes
1488:
1276:Admixture with other
1270:African golden wolves
1185:
679:critically endangered
625:
610:
536:Indigenous names for
292:Ethiopian wolf range
9322:Long-tailed weasel (
8900:Lyncodon patagonicus
8549:Greater hog badger (
8071:Sunda stink badger (
7761:Hawaiian monk seal (
7484:Guadalupe fur seal (
7457:Antarctic fur seal (
7448:Galápagos fur seal (
7344:Galápagos sea lion (
6462:Common raccoon dog (
6314:Cape gray mongoose (
6254:Jackson's mongoose (
6214:Paracynictis selousi
6205:Cynictis penicillata
5829:Durrell's vontsira (
5417:Giant forest genet (
5209:Large Indian civet (
5149:Small Indian civet (
4758:Sunda leopard cat (
4534:Southern tiger cat (
4470:African golden cat (
4147:African palm civet (
4016:Born Free Foundation
3986:University of Oxford
3630:Animal Diversity Web
3579:Journal of Mammalogy
3284:(21): 3441–3449.e5.
3257:Bahlk, S.H. (2015).
2601:: e.T3748A10051312.
2400:Born Free Foundation
1607:Cooperative breeding
1502:side-striped jackals
1337:Trinomial authority
1246:Juliet Clutton-Brock
1230:Glover Morrill Allen
1192:convergent evolution
550:Literal translation
444:disease transference
9875:Mammals of Ethiopia
9365:M. aistoodonnivalis
9201:Hairy-nosed otter (
9118:Neotropical otter (
8898:Patagonian weasel (
8361:White-nosed coati (
7561:Cystophora cristata
7552:Erignathus barbatus
7285:Northern fur seal (
7062:Helarctos malayanus
6698:Atelocynus microtis
6405:U. cinereoargenteus
6296:Egyptian mongoose (
6223:Rhynchogale melleri
6221:Meller's mongoose (
6212:Selous's mongoose (
6196:Ichneumia albicauda
6150:Collared mongoose (
6024:Angolan kusimanse (
5908:Liberian mongoose (
5261:West African oyan (
5160:Civettictis civetta
5091:Golden palm civet (
5049:Masked palm civet (
5024:Arctictis binturong
4998:Hemigalus derbyanus
4996:Banded palm civet (
4987:Hose's palm civet (
4776:Rusty-spotted cat (
4030:, images and videos
4005:Supporting the EWCP
3954:Williams, S. 2004.
3924:on 26 December 2011
3896:MacDonald, David W.
3290:2018CBio...28E3441G
3227:2015CBio...25.2158K
3105:1994MolEc...3..301G
3059:10.1038/nature08837
3051:2010Natur.464..898V
2918:, p. 5 & 8
2788:2004MolEc..13.2275G
2631:, Vol. Säugetiere:
2427:Black-backed jackal
2306:Ethiopian Civil War
2144:Probably declining
2067:16–19 (as of 2000)
2037:19–23 (as of 2000)
1924:
1824:, and occasionally
1790:Big-headed mole rat
1498:black-backed jackal
1489:Painting (1926) by
1361:Nominate subspecies
1294:African golden wolf
1254:side-striped jackal
1147:Ethiopian Highlands
1096:Black-backed jackal
1076:Side-striped jackal
667:Ethiopian Civil War
655:Percy Powell-Cotton
540:
436:habitat degradation
379:Ethiopian Highlands
361:), also called the
129:Conservation status
9480:European polecat (
9313:Colombian weasel (
8963:(African polecats)
8591:Caucasian badger (
8519:(Eurasian badgers)
7969:(hog-nosed skunks)
7706:("southern seals")
7605:Halichoerus grypus
7536:("northern seals")
7408:Phocarctos hookeri
7313:Eumetopias jubatus
7311:Steller sea lion (
7093:Asian black bear (
6843:African wild dog (
6797:(wolf-like canids)
6074:Atilax paludinosus
6033:Common kusimanse (
5960:Dologale dybowskii
5932:Gambian mongoose (
5716:Western falanouc (
5707:Eastern falanouc (
5676:Cryptoprocta ferox
5581:Parahyaena brunnea
5336:Johnston's genet (
5291:Abyssinian genet (
5243:(African linsangs)
5151:Viverricula indica
5082:Brown palm civet (
5073:Asian palm civet (
4980:Cynogale bennettii
4971:Chrotogale owstoni
4867:European wildcat (
4858:Black-footed cat (
4448:Leptailurus serval
4416:Asian golden cat (
4182:(Asiatic linsangs)
4010:2009-02-12 at the
3945:Wolves of Ethiopia
3775:2016-10-06 at the
3559:Dartmouth College.
3548:Powell-Cotton 1902
3466:Behavioral Ecology
3370:Journal of Zoology
3163:Powell-Cotton 1902
2999:Parkyns, Mansfield
2989:, pp. 206–207
2987:Powell-Cotton 1902
2382:), and one in the
2289:
2007:Small population,
1919:
1892:highlands, and in
1798:
1696:
1643:
1633:Hunting behaviours
1568:
1494:
1396:, north and south
1309:canid of 72% grey
1200:
628:
620:
603:Historical account
535:
468:community outreach
458:'s Ethiopian Wolf
9837:
9836:
9783:Open Tree of Life
9544:Taxon identifiers
9535:
9534:
9531:
9530:
9527:
9526:
9523:
9522:
9519:
9518:
9498:Siberian weasel (
9489:American ermine (
9417:Japanese weasel (
9372:Mountain weasel (
9356:
9279:
9278:
9065:
9050:
9049:
9046:
9045:
9034:Striped polecat (
8984:Marbled polecat (
8966:
8951:
8950:
8923:
8871:
8870:
8850:Japanese marten (
8796:American marten (
8789:
8647:M. cucphuongensis
8638:
8621:
8620:
8609:European badger (
8582:Japanese badger (
8522:
8475:American badger (
8438:
8437:
8434:
8433:
8410:
8409:
8406:
8405:
8334:
8293:Northern olingo (
8277:
8252:Cozumel raccoon (
8227:
8151:
8150:
8105:
8064:
7972:
7953:
7895:
7894:
7891:
7890:
7887:
7886:
7883:
7882:
7879:
7878:
7869:Ommatophoca rossi
7842:Hydrurga leptonyx
7829:
7826:(Antarctic seals)
7799:M. angustirostris
7790:
7773:
7772:
7739:Monachus monachus
7726:
7709:
7694:
7693:
7690:
7689:
7539:
7522:
7505:
7504:
7367:Otaria flavescens
7356:
7355:
7267:
7235:Odobenus rosmarus
7199:
7198:
7195:
7194:
7191:
7190:
7187:
7186:
7150:
7127:
7126:
7123:
7122:
7025:Spectacled bear (
6983:
6959:
6958:
6955:
6954:
6951:
6950:
6799:
6786:
6785:
6705:Crab-eating fox (
6696:Short-eared dog (
6644:
6629:
6616:
6615:
6498:
6440:Otocyon megalotis
6384:
6351:
6350:
6347:
6346:
6343:
6342:
6339:
6338:
6335:
6334:
6203:Yellow mongoose (
6054:
6053:
5941:Banded mongoose (
5910:Liberiictus kuhni
5864:
5845:
5844:
5841:
5840:
5728:
5727:
5654:
5623:
5622:
5619:
5618:
5615:
5614:
5559:
5520:
5486:
5485:
5482:
5481:
5478:
5477:
5450:
5449:
5446:
5445:
5442:
5441:
5438:
5437:
5390:Servaline genet (
5309:Bourlon's genet (
5284:
5245:
5221:
5220:
5103:
5102:
5075:P. hermaphroditus
4927:
4926:
4923:
4922:
4919:
4918:
4895:
4894:
4891:
4890:
4887:
4886:
4883:
4882:
4879:
4878:
4840:African wildcat (
4797:
4796:
4767:Flat-headed cat (
4707:
4706:
4491:
4490:
4428:
4427:
4342:
4341:
4338:
4337:
4266:Clouded leopard (
4225:
4200:Spotted linsang (
4184:
4149:Nandinia binotata
3943:Morell, V. 2006.
3914:978-2-8317-0407-4
3849:Lydekker, Richard
3093:Molecular Ecology
3045:(7290): 898–902.
2776:Molecular Ecology
2705:Mammalian Species
2669:978-0-8018-8221-0
2656:"Order Carnivora"
2186:
2185:
1963:102 (as of 2010)
1858:free-ranging dogs
1541:supracaudal gland
1478:
1477:
1435:
1427:
1419:
1409:
1163:mitochondrial DNA
1143:
1142:
1138:
1137:
1129:
1128:
1120:
1119:
1111:
1110:
1047:
1046:
1018:
1017:
989:
988:
962:
961:
933:
932:
904:
903:
875:
874:
866:
865:
733:Phylogenetic tree
711:local extinctions
687:Oxford University
600:
599:
524:rendering support
456:Oxford University
452:free-ranging dogs
349:
348:
343:
335:
327:
319:
311:
152:
9902:
9830:
9829:
9817:
9816:
9804:
9803:
9791:
9790:
9778:
9777:
9765:
9764:
9752:
9751:
9739:
9738:
9726:
9725:
9713:
9712:
9700:
9699:
9687:
9686:
9674:
9673:
9661:
9660:
9648:
9647:
9635:
9634:
9625:
9624:
9612:
9611:
9599:
9598:
9586:
9585:
9584:
9571:
9570:
9569:
9539:
9538:
9471:Malayan weasel (
9390:Steppe polecat (
9363:Sichuan weasel (
9355:
9354:
9350:
9290:
9289:
9227:L. perspicillata
9192:Eurasian otter (
9069:
9068:
9064:
9063:
9059:
8970:
8969:
8965:
8964:
8960:
8939:Greater grison (
8922:
8921:
8917:
8891:
8890:
8882:
8881:
8832:Nilgiri marten (
8805:Pacific marten (
8788:
8787:
8783:
8709:
8708:
8692:M. subaurantiaca
8637:
8636:
8635:(ferret-badgers)
8632:
8526:
8525:
8521:
8520:
8516:
8468:
8467:
8457:
8456:
8444:
8443:
8338:
8337:
8333:
8332:
8328:
8276:
8275:
8271:
8226:
8225:
8221:
8162:
8161:
8104:
8103:
8102:(spotted skunks)
8099:
8063:
8062:
8058:
7971:
7970:
7966:
7957:
7956:
7952:
7951:
7943:
7914:
7913:
7901:
7900:
7858:Crabeater seal (
7833:
7832:
7828:
7827:
7823:
7789:
7788:
7787:(elephant seals)
7784:
7763:N. schauinslandi
7730:
7729:
7725:
7724:
7720:
7713:
7712:
7708:
7707:
7703:
7578:
7577:
7543:
7542:
7538:
7537:
7533:
7526:
7525:
7521:
7520:
7514:
7475:Brown fur seal (
7450:A. galapagoensis
7337:Z. californianus
7304:
7303:
7271:
7270:
7266:
7265:
7251:
7221:
7220:
7213:
7205:
7204:
7149:
7148:
7144:
7138:
7137:
7053:Melursus ursinus
7044:
7043:
6987:
6986:
6982:
6979:
6972:
6971:
6939:Ethiopian wolf (
6803:
6802:
6798:
6795:
6648:
6647:
6643:
6640:
6633:
6632:
6628:
6625:
6514:Blanford's fox (
6497:
6490:
6431:
6430:
6389:
6388:
6383:
6380:
6373:
6372:
6365:
6357:
6356:
6159:Ruddy mongoose (
6141:Javan mongoose (
6072:Marsh mongoose (
6065:
6064:
6044:C. platycephalus
5877:
5876:
5868:
5867:
5863:
5860:
5858:
5739:
5738:
5683:Malagasy civet (
5667:
5666:
5658:
5657:
5653:
5648:
5641:
5640:
5636:
5563:
5562:
5558:
5555:
5524:
5523:
5519:
5516:
5509:
5508:
5505:
5504:
5492:
5491:
5456:
5455:
5283:
5280:
5244:
5241:
5232:
5231:
5142:
5141:
5138:
5127:
5126:
5122:
5109:
5108:
5015:
5014:
4957:
4956:
4946:
4945:
4933:
4932:
4727:Otocolobus manul
4718:
4717:
4640:
4639:
4516:Geoffroy's cat (
4439:
4438:
4369:
4368:
4360:
4359:
4355:
4243:
4242:
4234:
4233:
4230:
4229:
4224:
4221:
4214:
4213:
4191:Banded linsang (
4183:
4180:
4172:
4171:
4161:
4160:
4140:
4139:
4129:
4128:
4121:
4113:
4112:
4058:
4051:
4044:
4035:
4034:
4023:(Canis simensis)
3933:
3931:
3929:
3923:
3917:. Archived from
3906:
3887:
3873:
3858:
3844:
3842:
3841:
3835:
3828:
3807:
3806:
3804:
3802:
3790:
3784:
3768:Gutema, T. M.,
3766:
3760:
3759:
3757:
3755:
3746:
3737:
3731:
3725:
3719:
3713:
3707:
3706:, pp. 27–28
3701:
3695:
3694:, pp. 26–27
3689:
3683:
3682:, pp. 22–26
3677:
3671:
3665:
3659:
3658:, pp. 40–46
3653:
3647:
3641:
3635:
3634:
3621:
3612:
3611:, pp. 19–20
3606:
3597:
3596:
3594:
3570:
3564:
3557:
3551:
3545:
3539:
3533:
3527:
3526:
3490:
3484:
3483:
3481:
3457:
3451:
3450:
3416:
3407:
3401:
3395:
3386:
3385:
3367:
3358:
3352:
3346:
3337:
3331:
3320:
3319:
3309:
3268:
3262:
3255:
3249:
3248:
3238:
3205:
3199:
3198:
3196:
3172:
3166:
3160:
3154:
3148:
3142:
3139:
3133:
3132:
3087:
3081:
3080:
3070:
3029:
3023:
3014:
3008:
2996:
2990:
2984:
2975:
2969:
2960:
2954:
2948:
2947:
2940:
2934:
2928:
2919:
2913:
2907:
2906:
2904:
2902:
2883:
2874:
2865:
2864:
2862:
2860:
2850:"Ethiopian Wolf"
2846:
2840:
2834:
2823:
2822:
2820:
2814:. Archived from
2782:(8): 2275–2286.
2773:
2760:
2751:
2750:
2748:
2747:
2741:
2735:. Archived from
2702:
2698:"Canis simensis"
2693:
2674:
2673:
2651:
2640:
2626:
2620:
2619:
2617:
2615:
2610:
2580:
2559:
2558:
2548:
2507:
2483:
2477:
2471:
2456:
2450:
2443:
2347:Encounters with
2237:in North Wollo.
2217:Canine distemper
2106:canine distemper
1952:Simien Mountains
1925:
1918:
1882:Simien Mountains
1693:Simien Mountains
1602:gestation period
1452:
1433:
1425:
1417:
1415:
1407:
1386:Simien Mountains
1369:
1331:
1330:
1226:John Edward Gray
1155:African wild dog
1102:
1082:
1069:
1068:
1061:
1060:
1038:
1032:African wild dog
1009:
980:
953:
924:
895:
857:
837:
836:
822:
821:
815:
814:
808:
807:
800:
799:
792:
791:
784:
783:
776:
775:
768:
767:
760:
759:
746:
745:
737:wolf-like canids
729:
728:
698:Simien Mountains
541:
534:
512:
511:
502:Indigenous names
492:Ethiopian jackal
341:
333:
325:
317:
309:
287:
270:
250:C. simensis
167:
166:
146:
141:
140:
114:
104:
34:
27:Temporal range:
21:
20:
9910:
9909:
9905:
9904:
9903:
9901:
9900:
9899:
9840:
9839:
9838:
9833:
9825:
9820:
9812:
9807:
9799:
9794:
9786:
9781:
9773:
9770:Observation.org
9768:
9760:
9755:
9747:
9742:
9734:
9729:
9721:
9716:
9708:
9703:
9695:
9690:
9682:
9677:
9669:
9664:
9656:
9651:
9643:
9638:
9630:
9628:
9620:
9615:
9607:
9602:
9594:
9589:
9580:
9579:
9574:
9565:
9564:
9559:
9546:
9536:
9515:
9491:M. richardsonii
9435:European mink (
9352:
9351:
9349:
9339:
9331:American mink (
9304:Amazon weasel (
9275:
9233:
9209:
9176:
9159:
9135:
9084:
9061:
9060:
9058:
9042:
9009:
8992:
8962:
8961:
8959:
8947:
8930:Lesser grison (
8919:
8918:
8916:
8906:
8867:
8785:
8784:
8782:
8772:
8748:
8724:
8698:
8634:
8633:
8631:
8617:
8566:
8518:
8517:
8515:
8507:
8483:
8464:
8451:
8430:
8402:
8369:
8330:
8329:
8327:
8319:
8273:
8272:
8270:
8260:
8223:
8222:
8220:
8210:
8177:
8147:
8114:S. angustifrons
8101:
8100:
8098:
8088:
8061:(stink badgers)
8060:
8059:
8057:
8047:
8039:Striped skunk (
8014:
8008:C. semistriatus
7968:
7967:
7965:
7945:
7944:
7942:
7934:
7928:Ailurus fulgens
7908:
7875:
7825:
7824:
7822:
7814:
7786:
7785:
7783:
7769:
7745:
7722:
7721:
7719:
7705:
7704:
7702:
7686:
7644:
7611:
7567:
7535:
7534:
7532:
7518:
7516:
7515:
7513:
7501:
7414:
7397:
7373:
7352:
7319:
7293:
7260:
7255:
7253:
7252:
7250:
7246:
7241:
7215:
7211:
7183:
7163:
7146:
7145:
7119:
7068:
7033:
7009:
6980:
6978:
6968:
6947:
6876:Golden jackal (
6860:
6834:
6796:
6794:
6782:
6713:
6707:Cerdocyon thous
6687:
6663:
6641:
6639:
6626:
6624:
6612:
6577:Rüppell's fox (
6491:
6489:
6479:
6464:N. procyonoides
6446:
6438:Bat-eared fox (
6420:
6385:
6381:
6367:
6363:
6331:
6316:H. pulverulenta
6271:
6229:
6185:
6152:U. semitorquata
6107:U. auropunctata
6089:
6050:
5999:
5966:
5949:
5916:
5899:
5861:
5859:
5854:
5853:
5837:
5804:
5787:
5754:
5748:Galidia elegans
5724:
5691:
5651:
5649:
5647:
5637:
5632:
5611:
5603:Spotted hyena (
5587:
5570:Striped hyena (
5556:
5554:
5546:
5517:
5515:
5499:
5474:
5434:
5372:Aquatic genet (
5363:Pardine genet (
5300:Angolan genet (
5281:
5279:
5269:
5254:P. richardsonii
5242:
5240:
5217:
5200:Malayan civet (
5166:
5158:African civet (
5134:
5133:
5123:
5118:
5117:
5113:
5099:
5057:
5004:
4989:Diplogale hosei
4953:
4940:
4915:
4875:
4793:
4733:
4713:
4703:
4679:
4662:
4635:
4629:
4603:Eurasian lynx (
4578:
4487:
4454:
4434:
4424:
4391:
4364:
4356:
4351:
4334:
4274:
4226:
4222:
4208:
4181:
4168:
4155:
4136:
4123:
4119:
4107:
4069:
4062:
4021:Ethiopian wolf
4012:Wayback Machine
3969:
3940:
3938:Further reading
3927:
3925:
3921:
3915:
3904:
3878:Rüppell, Eduard
3839:
3837:
3833:
3826:
3816:
3811:
3810:
3800:
3798:
3797:. New Scientist
3791:
3787:
3783:(November 2015)
3777:Wayback Machine
3767:
3763:
3753:
3751:
3744:
3738:
3734:
3726:
3722:
3714:
3710:
3702:
3698:
3690:
3686:
3678:
3674:
3666:
3662:
3654:
3650:
3642:
3638:
3623:
3622:
3615:
3607:
3600:
3571:
3567:
3558:
3554:
3546:
3542:
3534:
3530:
3507:10.1038/nrg2664
3495:Nat. Rev. Genet
3491:
3487:
3458:
3454:
3414:
3408:
3404:
3396:
3389:
3365:
3359:
3355:
3347:
3340:
3332:
3323:
3278:Current Biology
3269:
3265:
3256:
3252:
3221:(16): 2158–65.
3215:Current Biology
3206:
3202:
3173:
3169:
3161:
3157:
3149:
3145:
3140:
3136:
3088:
3084:
3030:
3026:
3015:
3011:
2997:
2993:
2985:
2978:
2970:
2963:
2955:
2951:
2946:. 28 July 2012.
2942:
2941:
2937:
2929:
2922:
2914:
2910:
2900:
2898:
2896:
2881:
2875:
2868:
2858:
2856:
2848:
2847:
2843:
2835:
2826:
2818:
2771:
2761:
2754:
2745:
2743:
2739:
2717:10.2307/3504136
2700:
2694:
2677:
2670:
2652:
2643:
2627:
2623:
2613:
2611:
2581:
2562:
2508:
2497:
2492:
2487:
2486:
2478:
2474:
2457:
2453:
2444:
2440:
2435:
2423:
2361:
2345:
2331:
2302:
2278:
2260:
2243:
2225:
2208:
2191:
1993:, South Gondar
1874:
1783:
1685:
1680:
1635:
1611:pseudopregnancy
1590:
1573:
1557:
1504:as well as the
1483:
1455:de Beaux, 1922
1447:
1443:
1434:(Heuglin, 1862)
1432:
1428:
1424:
1420:
1418:(Heuglin, 1862)
1416:
1413:C. s. semiensis
1411:
1410:
1406:
1404:C. s. crinensis
1364:
1358:
1354:
1319:
1282:
1176:Canis evolution
1139:
1130:
1121:
1112:
1048:
1019:
990:
975:Ethiopian wolf
963:
934:
905:
876:
867:
831:
752: 3.5
727:
675:Sanetti Plateau
605:
595:
588:
574:
565:
547:Indigenous name
533:
532:
531:
522:Without proper
513:
509:
504:
496:Abyssinian wolf
476:
434:, resulting in
413:mountain ranges
278:
272:
266:
253:
161:
153:
142:
138:
131:
121:Sanetti Plateau
105:
103:
102:
101:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
40:
29:
28:
25:
17:
12:
11:
5:
9908:
9898:
9897:
9892:
9887:
9882:
9877:
9872:
9867:
9862:
9857:
9852:
9835:
9834:
9832:
9831:
9818:
9805:
9792:
9779:
9766:
9753:
9740:
9727:
9714:
9701:
9688:
9675:
9662:
9649:
9636:
9626:
9613:
9609:canis-simensis
9600:
9596:Canis_simensis
9587:
9582:Canis simensis
9572:
9556:
9554:
9552:Canis simensis
9548:
9547:
9533:
9532:
9529:
9528:
9525:
9524:
9521:
9520:
9517:
9516:
9514:
9513:
9509:M. strigidorsa
9504:
9495:
9486:
9477:
9468:
9462:Least weasel (
9459:
9450:
9441:
9432:
9423:
9414:
9408:Haida ermine (
9405:
9396:
9392:M. eversmannii
9387:
9378:
9369:
9359:
9357:
9341:
9340:
9338:
9337:
9328:
9319:
9310:
9300:
9298:
9287:
9281:
9280:
9277:
9276:
9274:
9273:
9264:
9255:
9245:
9243:
9235:
9234:
9232:
9231:
9221:
9219:
9211:
9210:
9208:
9207:
9198:
9188:
9186:
9178:
9177:
9175:
9174:
9164:
9161:
9160:
9158:
9157:
9147:
9145:
9137:
9136:
9134:
9133:
9124:
9120:L. longicaudis
9115:
9109:Marine otter (
9106:
9096:
9094:
9086:
9085:
9083:
9082:
9072:
9066:
9052:
9051:
9048:
9047:
9044:
9043:
9041:
9040:
9031:
9021:
9019:
9011:
9010:
9008:
9007:
8997:
8994:
8993:
8991:
8990:
8980:
8978:
8967:
8953:
8952:
8949:
8948:
8946:
8945:
8936:
8926:
8924:
8908:
8907:
8905:
8904:
8894:
8888:
8879:
8873:
8872:
8869:
8868:
8866:
8865:
8856:
8847:
8838:
8829:
8823:Beech marten (
8820:
8811:
8802:
8792:
8790:
8774:
8773:
8771:
8770:
8760:
8758:
8750:
8749:
8747:
8746:
8736:
8734:
8726:
8725:
8723:
8722:
8712:
8706:
8700:
8699:
8697:
8696:
8687:
8678:
8669:
8660:
8651:
8641:
8639:
8623:
8622:
8619:
8618:
8616:
8615:
8606:
8600:Asian badger (
8597:
8588:
8578:
8576:
8568:
8567:
8565:
8564:
8555:
8546:
8542:A. albogularis
8536:
8534:
8523:
8509:
8508:
8506:
8505:
8499:Honey badger (
8495:
8493:
8485:
8484:
8482:
8481:
8471:
8465:
8460:
8453:
8452:
8440:
8439:
8436:
8435:
8432:
8431:
8429:
8428:
8427:
8426:
8420:
8418:
8412:
8411:
8408:
8407:
8404:
8403:
8401:
8400:
8391:
8381:
8379:
8371:
8370:
8368:
8367:
8358:
8348:
8346:
8335:
8321:
8320:
8318:
8317:
8308:
8299:
8290:
8280:
8278:
8262:
8261:
8259:
8258:
8249:
8240:
8236:P. cancrivorus
8230:
8228:
8212:
8211:
8209:
8208:
8204:B. sumichrasti
8199:
8189:
8187:
8179:
8178:
8176:
8175:
8165:
8159:
8153:
8152:
8149:
8148:
8146:
8145:
8136:
8127:
8118:
8108:
8106:
8090:
8089:
8087:
8086:
8077:
8067:
8065:
8049:
8048:
8046:
8045:
8036:
8030:Hooded skunk (
8026:
8024:
8016:
8015:
8013:
8012:
8003:
7994:
7985:
7975:
7973:
7954:
7936:
7935:
7933:
7932:
7922:
7920:
7910:
7909:
7897:
7896:
7893:
7892:
7889:
7888:
7885:
7884:
7881:
7880:
7877:
7876:
7874:
7873:
7864:
7855:
7849:Weddell seal (
7846:
7840:Leopard seal (
7836:
7830:
7816:
7815:
7813:
7812:
7803:
7793:
7791:
7775:
7774:
7771:
7770:
7768:
7767:
7757:
7755:
7747:
7746:
7744:
7743:
7733:
7727:
7710:
7696:
7695:
7692:
7691:
7688:
7687:
7685:
7684:
7675:
7666:
7660:Caspian seal (
7656:
7654:
7646:
7645:
7643:
7642:
7633:
7627:Spotted seal (
7623:
7621:
7613:
7612:
7610:
7609:
7600:
7591:
7581:
7575:
7569:
7568:
7566:
7565:
7556:
7550:Bearded seal (
7546:
7540:
7523:
7519:or true seals)
7517:(earless seals
7507:
7506:
7503:
7502:
7500:
7499:
7490:
7481:
7472:
7463:
7454:
7445:
7436:
7426:
7424:
7416:
7415:
7413:
7412:
7402:
7399:
7398:
7396:
7395:
7385:
7383:
7375:
7374:
7372:
7371:
7361:
7358:
7357:
7354:
7353:
7351:
7350:
7341:
7331:
7329:
7321:
7320:
7318:
7317:
7307:
7301:
7295:
7294:
7292:
7291:
7281:
7279:
7268:
7243:
7242:
7240:
7239:
7229:
7227:
7217:
7216:
7201:
7200:
7197:
7196:
7193:
7192:
7189:
7188:
7185:
7184:
7182:
7181:
7180:
7179:
7173:
7171:
7165:
7164:
7162:
7161:
7160:
7159:
7153:
7151:
7135:
7129:
7128:
7125:
7124:
7121:
7120:
7118:
7117:
7108:
7099:
7090:
7080:
7078:
7070:
7069:
7067:
7066:
7057:
7047:
7041:
7035:
7034:
7032:
7031:
7021:
7019:
7011:
7010:
7008:
7007:
7003:A. melanoleuca
6997:
6995:
6984:
6969:
6964:
6961:
6960:
6957:
6956:
6953:
6952:
6949:
6948:
6946:
6945:
6936:
6927:
6921:Eastern wolf (
6918:
6909:
6903:African wolf (
6900:
6891:
6885:Domestic dog (
6882:
6872:
6870:
6862:
6861:
6859:
6858:
6849:
6839:
6836:
6835:
6833:
6832:
6823:
6813:
6811:
6800:
6788:
6787:
6784:
6783:
6781:
6780:
6771:
6765:Sechuran fox (
6762:
6758:L. gymnocercus
6753:
6744:
6738:Darwin's fox (
6735:
6725:
6723:
6715:
6714:
6712:
6711:
6702:
6692:
6689:
6688:
6686:
6685:
6675:
6673:
6665:
6664:
6662:
6661:
6651:
6645:
6630:
6618:
6617:
6614:
6613:
6611:
6610:
6601:
6592:
6583:
6574:
6565:
6556:
6547:
6538:
6529:
6520:
6511:
6507:V. bengalensis
6501:
6499:
6481:
6480:
6478:
6477:
6468:
6458:
6456:
6448:
6447:
6445:
6444:
6434:
6428:
6422:
6421:
6419:
6418:
6409:
6399:
6397:
6386:
6376:
6369:
6368:
6353:
6352:
6349:
6348:
6345:
6344:
6341:
6340:
6337:
6336:
6333:
6332:
6330:
6329:
6320:
6311:
6302:
6293:
6283:
6281:
6273:
6272:
6270:
6269:
6260:
6251:
6247:B. crassicauda
6241:
6239:
6231:
6230:
6228:
6227:
6218:
6209:
6200:
6190:
6187:
6186:
6184:
6183:
6174:
6165:
6156:
6147:
6138:
6129:
6120:
6111:
6101:
6099:
6091:
6090:
6088:
6087:
6078:
6068:
6062:
6056:
6055:
6052:
6051:
6049:
6048:
6039:
6030:
6021:
6011:
6009:
6001:
6000:
5998:
5997:
5988:
5978:
5976:
5968:
5967:
5965:
5964:
5954:
5951:
5950:
5948:
5947:
5938:
5928:
5926:
5918:
5917:
5915:
5914:
5904:
5901:
5900:
5898:
5897:
5887:
5885:
5874:
5865:
5847:
5846:
5843:
5842:
5839:
5838:
5836:
5835:
5826:
5816:
5814:
5806:
5805:
5803:
5802:
5792:
5789:
5788:
5786:
5785:
5781:G. grandidieri
5776:
5766:
5764:
5756:
5755:
5753:
5752:
5742:
5736:
5730:
5729:
5726:
5725:
5723:
5722:
5713:
5703:
5701:
5693:
5692:
5690:
5689:
5680:
5670:
5664:
5655:
5638:
5628:
5625:
5624:
5621:
5620:
5617:
5616:
5613:
5612:
5610:
5609:
5599:
5597:
5589:
5588:
5586:
5585:
5576:
5566:
5560:
5548:
5547:
5545:
5544:
5534:
5532:
5521:
5501:
5500:
5488:
5487:
5484:
5483:
5480:
5479:
5476:
5475:
5473:
5472:
5471:
5470:
5464:
5462:
5452:
5451:
5448:
5447:
5444:
5443:
5440:
5439:
5436:
5435:
5433:
5432:
5423:
5414:
5405:
5396:
5387:
5378:
5369:
5360:
5351:
5345:Letaba genet (
5342:
5333:
5327:Common genet (
5324:
5315:
5306:
5297:
5287:
5285:
5271:
5270:
5268:
5267:
5258:
5248:
5246:
5229:
5223:
5222:
5219:
5218:
5216:
5215:
5206:
5197:
5188:
5178:
5176:
5168:
5167:
5165:
5164:
5155:
5145:
5139:
5124:
5112:
5105:
5104:
5101:
5100:
5098:
5097:
5093:P. zeylonensis
5088:
5079:
5069:
5067:
5059:
5058:
5056:
5055:
5051:Paguma larvata
5046:
5037:
5028:
5018:
5012:
5006:
5005:
5003:
5002:
4993:
4984:
4975:
4965:
4963:
4954:
4949:
4942:
4941:
4929:
4928:
4925:
4924:
4921:
4920:
4917:
4916:
4914:
4913:
4912:
4911:
4905:
4903:
4897:
4896:
4893:
4892:
4889:
4888:
4885:
4884:
4881:
4880:
4877:
4876:
4874:
4873:
4864:
4855:
4846:
4837:
4828:
4822:Domestic cat (
4819:
4809:
4807:
4799:
4798:
4795:
4794:
4792:
4791:
4782:
4778:P. rubiginosus
4773:
4764:
4755:
4751:P. bengalensis
4745:
4743:
4735:
4734:
4732:
4731:
4725:Pallas's cat (
4721:
4715:
4709:
4708:
4705:
4704:
4702:
4701:
4691:
4689:
4681:
4680:
4678:
4677:
4667:
4664:
4663:
4661:
4660:
4650:
4648:
4637:
4631:
4630:
4628:
4627:
4618:
4612:Iberian lynx (
4609:
4600:
4590:
4588:
4580:
4579:
4577:
4576:
4567:
4558:
4549:
4540:
4531:
4522:
4513:
4503:
4501:
4493:
4492:
4489:
4488:
4486:
4485:
4476:
4466:
4464:
4456:
4455:
4453:
4452:
4442:
4436:
4430:
4429:
4426:
4425:
4423:
4422:
4413:
4403:
4401:
4393:
4392:
4390:
4389:
4379:
4377:
4366:
4357:
4347:
4344:
4343:
4340:
4339:
4336:
4335:
4333:
4332:
4323:
4314:
4305:
4296:
4290:Snow leopard (
4286:
4284:
4276:
4275:
4273:
4272:
4263:
4253:
4251:
4240:
4227:
4217:
4210:
4209:
4207:
4206:
4197:
4187:
4185:
4169:
4164:
4157:
4156:
4154:
4153:
4143:
4137:
4132:
4125:
4124:
4109:
4108:
4106:
4105:
4103:Laurasiatheria
4099:
4093:
4087:
4081:
4074:
4071:
4070:
4061:
4060:
4053:
4046:
4038:
4032:
4031:
4018:
4002:
3989:
3975:
3968:
3967:External links
3965:
3964:
3963:
3952:
3939:
3936:
3935:
3934:
3913:
3888:
3874:
3859:
3845:
3815:
3812:
3809:
3808:
3785:
3761:
3732:
3720:
3708:
3696:
3684:
3672:
3660:
3648:
3636:
3613:
3598:
3585:(1): 129–137.
3565:
3552:
3540:
3538:, pp. 3–4
3528:
3501:(11): 783–96.
3485:
3452:
3402:
3400:, pp. 4–5
3387:
3376:(3): 351–361.
3353:
3338:
3321:
3263:
3250:
3200:
3167:
3155:
3153:, pp. 2–3
3143:
3134:
3082:
3024:
3009:
2991:
2976:
2961:
2949:
2935:
2920:
2908:
2895:978-2831707860
2894:
2866:
2841:
2839:, pp. 7–8
2824:
2821:on 2013-05-16.
2767:Canis simensis
2752:
2675:
2668:
2641:
2621:
2587:Canis simensis
2560:
2494:
2493:
2491:
2488:
2485:
2484:
2472:
2451:
2437:
2436:
2434:
2431:
2430:
2429:
2422:
2419:
2360:
2357:
2353:Canis lupaster
2344:
2341:
2330:
2327:
2301:
2298:
2277:
2274:
2259:
2256:
2242:
2239:
2224:
2221:
2207:
2204:
2190:
2187:
2184:
2183:
2180:
2174:
2171:
2168:
2165:
2164:1,141 km
2162:
2159:Bale Mountains
2155:
2154:
2151:
2148:
2145:
2142:
2139:
2136:
2130:Arsi Mountains
2126:
2125:
2122:
2115:
2112:
2109:
2102:
2099:
2089:
2088:
2080:
2074:
2071:
2068:
2065:
2062:
2058:
2057:
2046:
2043:
2041:
2038:
2035:
2032:
2024:
2023:
2012:
2005:
2003:
2000:
1997:
1994:
1987:
1986:
1979:
1970:
1967:
1964:
1961:
1958:
1948:
1947:
1944:
1941:
1938:
1935:
1932:
1929:
1922:Canis simensis
1914:Bale Mountains
1890:southern Wollo
1873:
1870:
1834:mountain nyala
1814:highland hares
1782:
1779:
1771:giant lobelias
1684:
1681:
1679:
1676:
1648:mountain nyala
1634:
1631:
1589:
1586:
1572:
1569:
1565:Bale Mountains
1556:
1553:
1482:
1479:
1476:
1475:
1473:
1471:Bale Mountains
1463:
1456:
1453:
1445:C. s. citernii
1437:
1436:
1422:C. s. simensis
1401:
1378:
1376:
1370:
1356:C. s. simensis
1348:
1347:
1344:
1341:
1338:
1335:
1318:
1315:
1281:
1274:
1180:
1179:
1141:
1140:
1136:
1135:
1132:
1131:
1127:
1126:
1123:
1122:
1118:
1117:
1114:
1113:
1109:
1108:
1105:
1104:
1092:
1089:
1088:
1085:
1084:
1072:
1067:
1065:
1059:
1057:
1054:
1053:
1050:
1049:
1045:
1044:
1041:
1040:
1028:
1025:
1024:
1021:
1020:
1016:
1015:
1012:
1011:
999:
996:
995:
992:
991:
987:
986:
983:
982:
972:
969:
968:
965:
964:
960:
959:
956:
955:
943:
940:
939:
936:
935:
931:
930:
927:
926:
914:
911:
910:
907:
906:
902:
901:
898:
897:
885:
882:
881:
878:
877:
873:
872:
869:
868:
864:
863:
860:
859:
847:
844:
843:
840:
839:
825:
820:
818:
813:
811:
806:
804:
798:
796:
790:
788:
782:
780:
774:
772:
766:
764:
758:
756:
744:
741:
740:
726:
723:
706:Arsi Mountains
644:Charles Darwin
636:British Museum
632:Eduard Rüppell
612:Canis simensis
604:
601:
598:
597:
594:Horse's jackal
592:
586:Jeedala fardaa
583:
577:
576:
571:
558:
552:
551:
548:
545:
538:Canis simensis
526:, you may see
514:
507:
506:
505:
503:
500:
475:
472:
417:Bale Mountains
377:native to the
358:Canis simensis
353:Ethiopian wolf
347:
346:
345:
344:
342:de Beaux, 1922
336:
328:
320:
312:
301:
300:
294:
293:
289:
288:
280:
279:
273:
268:Canis simensis
262:
261:
255:
254:
247:
245:
241:
240:
233:
229:
228:
223:
219:
218:
213:
209:
208:
203:
199:
198:
193:
189:
188:
183:
179:
178:
173:
169:
168:
155:
154:
136:
133:
132:
127:
124:
123:
116:
115:
107:
106:
98:
97:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
36:
26:
24:Ethiopian wolf
15:
9:
6:
4:
3:
2:
9907:
9896:
9893:
9891:
9888:
9886:
9883:
9881:
9878:
9876:
9873:
9871:
9868:
9866:
9863:
9861:
9858:
9856:
9853:
9851:
9848:
9847:
9845:
9828:
9823:
9819:
9815:
9810:
9806:
9802:
9797:
9793:
9789:
9784:
9780:
9776:
9771:
9767:
9763:
9758:
9754:
9750:
9745:
9741:
9737:
9732:
9728:
9724:
9719:
9715:
9711:
9706:
9702:
9698:
9693:
9689:
9685:
9680:
9676:
9672:
9667:
9663:
9659:
9654:
9650:
9646:
9641:
9637:
9633:
9627:
9623:
9618:
9614:
9610:
9605:
9601:
9597:
9592:
9588:
9583:
9577:
9573:
9568:
9562:
9558:
9557:
9555:
9553:
9549:
9545:
9540:
9512:
9510:
9505:
9503:
9501:
9496:
9494:
9492:
9487:
9485:
9483:
9478:
9476:
9474:
9469:
9467:
9465:
9460:
9458:
9456:
9451:
9449:
9447:
9446:M. lutreolina
9442:
9440:
9438:
9433:
9431:
9429:
9424:
9422:
9420:
9415:
9413:
9411:
9406:
9404:
9402:
9397:
9395:
9393:
9388:
9386:
9384:
9379:
9377:
9375:
9370:
9368:
9366:
9361:
9360:
9358:
9348:
9347:
9342:
9336:
9334:
9329:
9327:
9325:
9320:
9318:
9316:
9311:
9309:
9307:
9302:
9301:
9299:
9297:
9296:
9291:
9288:
9286:
9282:
9272:
9270:
9265:
9263:
9261:
9256:
9254:
9252:
9247:
9246:
9244:
9242:
9241:
9236:
9230:
9228:
9223:
9222:
9220:
9218:
9217:
9212:
9206:
9204:
9199:
9197:
9195:
9190:
9189:
9187:
9185:
9184:
9179:
9173:
9171:
9166:
9165:
9162:
9156:
9154:
9149:
9148:
9146:
9144:
9143:
9138:
9132:
9130:
9125:
9123:
9121:
9116:
9114:
9112:
9107:
9105:
9103:
9102:L. canadensis
9098:
9097:
9095:
9093:
9092:
9087:
9081:
9079:
9076:Giant otter (
9074:
9073:
9070:
9067:
9057:
9053:
9039:
9037:
9032:
9030:
9028:
9023:
9022:
9020:
9018:
9017:
9012:
9006:
9004:
8999:
8998:
8995:
8989:
8987:
8982:
8981:
8979:
8977:
8976:
8971:
8968:
8958:
8954:
8944:
8942:
8937:
8935:
8933:
8928:
8927:
8925:
8915:
8914:
8909:
8903:
8901:
8896:
8895:
8892:
8889:
8887:
8883:
8880:
8878:
8874:
8864:
8862:
8857:
8855:
8853:
8848:
8846:
8844:
8839:
8837:
8835:
8834:M. gwatkinsii
8830:
8828:
8826:
8821:
8819:
8817:
8812:
8810:
8808:
8803:
8801:
8799:
8794:
8793:
8791:
8781:
8780:
8775:
8769:
8767:
8762:
8761:
8759:
8757:
8756:
8751:
8745:
8743:
8738:
8737:
8735:
8733:
8732:
8727:
8721:
8719:
8714:
8713:
8710:
8707:
8705:
8701:
8695:
8693:
8688:
8686:
8684:
8679:
8677:
8675:
8674:M. orientalis
8670:
8668:
8666:
8661:
8659:
8657:
8652:
8650:
8648:
8643:
8642:
8640:
8630:
8629:
8624:
8614:
8612:
8607:
8605:
8603:
8598:
8596:
8594:
8589:
8587:
8585:
8580:
8579:
8577:
8575:
8574:
8569:
8563:
8561:
8556:
8554:
8552:
8547:
8545:
8543:
8538:
8537:
8535:
8533:
8532:
8527:
8524:
8514:
8510:
8504:
8502:
8497:
8496:
8494:
8492:
8491:
8486:
8480:
8478:
8477:Taxidea taxus
8473:
8472:
8469:
8466:
8463:
8458:
8454:
8450:
8445:
8441:
8424:
8423:
8422:
8421:
8419:
8417:
8413:
8399:
8397:
8392:
8390:
8388:
8387:N. meridensis
8383:
8382:
8380:
8378:
8377:
8372:
8366:
8364:
8359:
8357:
8355:
8350:
8349:
8347:
8345:
8344:
8339:
8336:
8326:
8322:
8316:
8314:
8309:
8307:
8305:
8300:
8298:
8296:
8291:
8289:
8287:
8282:
8281:
8279:
8269:
8268:
8263:
8257:
8255:
8250:
8248:
8246:
8241:
8239:
8237:
8232:
8231:
8229:
8219:
8218:
8213:
8207:
8205:
8200:
8198:
8196:
8191:
8190:
8188:
8186:
8185:
8180:
8174:
8172:
8167:
8166:
8163:
8160:
8158:
8154:
8144:
8142:
8137:
8135:
8133:
8128:
8126:
8124:
8119:
8117:
8115:
8110:
8109:
8107:
8097:
8096:
8091:
8085:
8083:
8078:
8076:
8074:
8073:M. javanensis
8069:
8068:
8066:
8056:
8055:
8050:
8044:
8042:
8037:
8035:
8033:
8028:
8027:
8025:
8023:
8022:
8017:
8011:
8009:
8004:
8002:
8000:
7999:C. leuconotus
7995:
7993:
7991:
7990:C. humboldtii
7986:
7984:
7982:
7977:
7976:
7974:
7964:
7963:
7958:
7955:
7949:
7941:
7937:
7931:
7929:
7924:
7923:
7921:
7919:
7915:
7911:
7907:
7902:
7898:
7872:
7870:
7865:
7863:
7861:
7856:
7854:
7852:
7847:
7845:
7843:
7838:
7837:
7834:
7831:
7821:
7817:
7811:
7809:
7804:
7802:
7800:
7795:
7794:
7792:
7782:
7781:
7776:
7766:
7764:
7759:
7758:
7756:
7754:
7753:
7748:
7742:
7740:
7735:
7734:
7731:
7728:
7718:
7714:
7711:
7701:
7697:
7683:
7681:
7678:Baikal seal (
7676:
7674:
7672:
7669:Ringed seal (
7667:
7665:
7663:
7658:
7657:
7655:
7653:
7652:
7647:
7641:
7639:
7636:Harbor seal (
7634:
7632:
7630:
7625:
7624:
7622:
7620:
7619:
7614:
7608:
7606:
7601:
7599:
7597:
7594:Ribbon seal (
7592:
7590:
7588:
7583:
7582:
7579:
7576:
7574:
7570:
7564:
7562:
7559:Hooded seal (
7557:
7555:
7553:
7548:
7547:
7544:
7541:
7531:
7527:
7524:
7512:
7508:
7498:
7496:
7495:A. tropicalis
7491:
7489:
7487:
7482:
7480:
7478:
7473:
7471:
7469:
7464:
7462:
7460:
7455:
7453:
7451:
7446:
7444:
7442:
7437:
7435:
7433:
7428:
7427:
7425:
7423:
7422:
7421:Arctocephalus
7417:
7411:
7409:
7404:
7403:
7400:
7394:
7392:
7387:
7386:
7384:
7382:
7381:
7376:
7370:
7368:
7363:
7362:
7359:
7349:
7347:
7346:Z. wollebaeki
7342:
7340:
7338:
7333:
7332:
7330:
7328:
7327:
7322:
7316:
7314:
7309:
7308:
7305:
7302:
7300:
7296:
7290:
7288:
7283:
7282:
7280:
7278:
7277:
7272:
7269:
7263:
7258:
7254:(eared seals:
7249:
7244:
7238:
7236:
7231:
7230:
7228:
7226:
7222:
7218:
7214:
7206:
7202:
7177:
7176:
7175:
7174:
7172:
7170:
7166:
7157:
7156:
7155:
7154:
7152:
7143:
7139:
7136:
7134:
7130:
7116:
7114:
7109:
7107:
7105:
7100:
7098:
7096:
7095:U. thibetanus
7091:
7089:
7087:
7086:U. americanus
7082:
7081:
7079:
7077:
7076:
7071:
7065:
7063:
7058:
7056:
7054:
7049:
7048:
7045:
7042:
7040:
7036:
7030:
7028:
7023:
7022:
7020:
7018:
7017:
7012:
7006:
7004:
7001:Giant panda (
6999:
6998:
6996:
6994:
6993:
6988:
6985:
6977:
6973:
6970:
6967:
6962:
6944:
6942:
6937:
6935:
6933:
6928:
6926:
6924:
6919:
6917:
6915:
6910:
6908:
6906:
6901:
6899:
6897:
6892:
6890:
6888:
6887:C. familiaris
6883:
6881:
6879:
6874:
6873:
6871:
6869:
6868:
6863:
6857:
6855:
6850:
6848:
6846:
6845:Lycaon pictus
6841:
6840:
6837:
6831:
6829:
6824:
6822:
6820:
6815:
6814:
6812:
6810:
6809:
6804:
6801:
6793:
6789:
6779:
6777:
6772:
6770:
6768:
6763:
6761:
6759:
6754:
6752:
6750:
6745:
6743:
6741:
6736:
6734:
6732:
6727:
6726:
6724:
6722:
6721:
6716:
6710:
6708:
6703:
6701:
6699:
6694:
6693:
6690:
6684:
6682:
6677:
6676:
6674:
6672:
6671:
6666:
6660:
6658:
6653:
6652:
6649:
6646:
6638:
6634:
6631:
6623:
6619:
6609:
6607:
6602:
6600:
6598:
6593:
6591:
6589:
6584:
6582:
6580:
6575:
6573:
6571:
6566:
6564:
6562:
6557:
6555:
6553:
6548:
6546:
6544:
6541:Tibetan fox (
6539:
6537:
6535:
6530:
6528:
6526:
6521:
6519:
6517:
6512:
6510:
6508:
6503:
6502:
6500:
6495:
6488:
6487:
6482:
6476:
6474:
6473:N. viverrinus
6469:
6467:
6465:
6460:
6459:
6457:
6455:
6454:
6449:
6443:
6441:
6436:
6435:
6432:
6429:
6427:
6423:
6417:
6415:
6414:U. littoralis
6410:
6408:
6406:
6401:
6400:
6398:
6396:
6395:
6390:
6387:
6379:
6374:
6370:
6366:
6358:
6354:
6328:
6326:
6321:
6319:
6317:
6312:
6310:
6308:
6303:
6301:
6299:
6294:
6292:
6290:
6289:H. flavescens
6285:
6284:
6282:
6280:
6279:
6274:
6268:
6266:
6261:
6259:
6257:
6252:
6250:
6248:
6243:
6242:
6240:
6238:
6237:
6232:
6226:
6224:
6219:
6217:
6215:
6210:
6208:
6206:
6201:
6199:
6197:
6192:
6191:
6188:
6182:
6180:
6179:U. vitticolla
6175:
6173:
6171:
6166:
6164:
6162:
6157:
6155:
6153:
6148:
6146:
6144:
6139:
6137:
6135:
6130:
6128:
6126:
6121:
6119:
6117:
6112:
6110:
6108:
6103:
6102:
6100:
6098:
6097:
6092:
6086:
6084:
6083:Xenogale naso
6079:
6077:
6075:
6070:
6069:
6066:
6063:
6061:
6057:
6047:
6045:
6040:
6038:
6036:
6031:
6029:
6027:
6022:
6020:
6018:
6013:
6012:
6010:
6008:
6007:
6002:
5996:
5994:
5989:
5987:
5985:
5980:
5979:
5977:
5975:
5974:
5969:
5963:
5961:
5956:
5955:
5952:
5946:
5944:
5939:
5937:
5935:
5930:
5929:
5927:
5925:
5924:
5919:
5913:
5911:
5906:
5905:
5902:
5896:
5894:
5889:
5888:
5886:
5884:
5883:
5878:
5875:
5873:
5869:
5866:
5857:
5856:sensu stricto
5852:
5848:
5834:
5832:
5827:
5825:
5823:
5818:
5817:
5815:
5813:
5812:
5807:
5801:
5799:
5794:
5793:
5790:
5784:
5782:
5777:
5775:
5773:
5768:
5767:
5765:
5763:
5762:
5757:
5751:
5749:
5744:
5743:
5740:
5737:
5735:
5731:
5721:
5719:
5714:
5712:
5710:
5705:
5704:
5702:
5700:
5699:
5694:
5688:
5686:
5685:Fossa fossana
5681:
5679:
5677:
5672:
5671:
5668:
5665:
5663:
5659:
5656:
5646:
5642:
5639:
5635:
5631:
5626:
5608:
5606:
5601:
5600:
5598:
5596:
5595:
5590:
5584:
5582:
5579:Brown hyena (
5577:
5575:
5573:
5572:Hyaena hyaena
5568:
5567:
5564:
5561:
5553:
5549:
5543:
5541:
5536:
5535:
5533:
5531:
5530:
5525:
5522:
5514:
5510:
5506:
5502:
5498:
5493:
5489:
5468:
5467:
5466:
5465:
5463:
5461:
5457:
5453:
5431:
5429:
5424:
5422:
5420:
5415:
5413:
5411:
5406:
5404:
5402:
5399:Hausa genet (
5397:
5395:
5393:
5388:
5386:
5384:
5379:
5377:
5375:
5370:
5368:
5366:
5361:
5359:
5357:
5352:
5350:
5348:
5343:
5341:
5339:
5334:
5332:
5330:
5325:
5323:
5321:
5316:
5314:
5312:
5307:
5305:
5303:
5302:G. angolensis
5298:
5296:
5294:
5293:G. abyssinica
5289:
5288:
5286:
5278:
5277:
5272:
5266:
5264:
5259:
5257:
5255:
5250:
5249:
5247:
5239:
5238:
5233:
5230:
5228:
5224:
5214:
5212:
5207:
5205:
5203:
5202:V. tangalunga
5198:
5196:
5194:
5189:
5187:
5185:
5180:
5179:
5177:
5175:
5174:
5169:
5163:
5161:
5156:
5154:
5152:
5147:
5146:
5143:
5140:
5137:
5136:sensu stricto
5132:
5128:
5125:
5121:
5116:
5110:
5106:
5096:
5094:
5089:
5087:
5085:
5080:
5078:
5076:
5071:
5070:
5068:
5066:
5065:
5060:
5054:
5052:
5047:
5045:
5043:
5038:
5036:
5034:
5029:
5027:
5025:
5020:
5019:
5016:
5013:
5011:
5010:Paradoxurinae
5007:
5001:
4999:
4994:
4992:
4990:
4985:
4983:
4981:
4978:Otter civet (
4976:
4974:
4972:
4967:
4966:
4964:
4962:
4958:
4955:
4952:
4947:
4943:
4939:
4934:
4930:
4909:
4908:
4907:
4906:
4904:
4902:
4898:
4872:
4870:
4869:F. silvestris
4865:
4863:
4861:
4856:
4854:
4852:
4847:
4845:
4843:
4838:
4836:
4834:
4829:
4827:
4825:
4820:
4818:
4816:
4811:
4810:
4808:
4806:
4805:
4800:
4790:
4788:
4787:P. viverrinus
4785:Fishing cat (
4783:
4781:
4779:
4774:
4772:
4770:
4765:
4763:
4761:
4760:P. javanensis
4756:
4754:
4752:
4749:Leopard cat (
4747:
4746:
4744:
4742:
4741:
4736:
4730:
4728:
4723:
4722:
4719:
4716:
4710:
4700:
4698:
4693:
4692:
4690:
4688:
4687:
4682:
4676:
4674:
4669:
4668:
4665:
4659:
4657:
4652:
4651:
4649:
4647:
4646:
4641:
4638:
4632:
4626:
4624:
4619:
4617:
4615:
4610:
4608:
4606:
4601:
4599:
4597:
4596:L. canadensis
4594:Canada lynx (
4592:
4591:
4589:
4587:
4586:
4581:
4575:
4573:
4568:
4566:
4564:
4559:
4557:
4555:
4550:
4548:
4546:
4541:
4539:
4537:
4532:
4530:
4528:
4523:
4521:
4519:
4514:
4512:
4510:
4505:
4504:
4502:
4500:
4499:
4494:
4484:
4482:
4477:
4475:
4473:
4468:
4467:
4465:
4463:
4462:
4457:
4451:
4449:
4444:
4443:
4440:
4437:
4431:
4421:
4419:
4418:C. temminckii
4414:
4412:
4410:
4405:
4404:
4402:
4400:
4399:
4394:
4388:
4386:
4383:Marbled cat (
4381:
4380:
4378:
4376:
4375:
4370:
4367:
4361:
4358:
4354:
4353:sensu stricto
4350:
4345:
4331:
4329:
4324:
4322:
4320:
4315:
4313:
4311:
4306:
4304:
4302:
4297:
4295:
4293:
4288:
4287:
4285:
4283:
4282:
4277:
4271:
4269:
4264:
4262:
4260:
4255:
4254:
4252:
4250:
4249:
4244:
4241:
4239:
4235:
4231:
4228:
4220:
4215:
4211:
4205:
4203:
4202:P. pardicolor
4198:
4196:
4194:
4189:
4188:
4186:
4179:
4178:
4173:
4170:
4167:
4162:
4158:
4152:
4150:
4145:
4144:
4141:
4138:
4135:
4130:
4126:
4122:
4114:
4110:
4104:
4100:
4098:
4094:
4092:
4088:
4086:
4082:
4080:
4076:
4075:
4072:
4067:
4059:
4054:
4052:
4047:
4045:
4040:
4039:
4036:
4029:
4025:
4024:
4019:
4017:
4013:
4009:
4006:
4003:
4001:
3997:
3993:
3990:
3987:
3983:
3979:
3976:
3974:
3971:
3970:
3961:
3957:
3953:
3951:, March 2006.
3950:
3946:
3942:
3941:
3920:
3916:
3910:
3903:
3902:
3897:
3893:
3889:
3885:
3884:
3879:
3875:
3871:
3870:
3864:
3860:
3856:
3855:
3850:
3846:
3836:on 2014-09-08
3832:
3825:
3824:
3818:
3817:
3796:
3789:
3782:
3778:
3774:
3771:
3765:
3750:
3743:
3736:
3729:
3724:
3717:
3712:
3705:
3700:
3693:
3688:
3681:
3676:
3669:
3664:
3657:
3652:
3645:
3640:
3632:
3631:
3626:
3620:
3618:
3610:
3605:
3603:
3593:
3588:
3584:
3580:
3576:
3569:
3562:
3556:
3550:, p. 207
3549:
3544:
3537:
3532:
3524:
3520:
3516:
3512:
3508:
3504:
3500:
3496:
3489:
3480:
3475:
3472:(3): 579–89.
3471:
3467:
3463:
3456:
3448:
3444:
3440:
3436:
3432:
3428:
3424:
3420:
3413:
3406:
3399:
3394:
3392:
3383:
3379:
3375:
3371:
3364:
3357:
3350:
3345:
3343:
3335:
3330:
3328:
3326:
3317:
3313:
3308:
3303:
3299:
3295:
3291:
3287:
3283:
3279:
3275:
3267:
3260:
3254:
3246:
3242:
3237:
3232:
3228:
3224:
3220:
3216:
3212:
3204:
3195:
3190:
3186:
3182:
3178:
3171:
3165:, p. 459
3164:
3159:
3152:
3147:
3138:
3130:
3126:
3122:
3118:
3114:
3110:
3106:
3102:
3099:(4): 301–12.
3098:
3094:
3086:
3078:
3074:
3069:
3064:
3060:
3056:
3052:
3048:
3044:
3040:
3036:
3028:
3021:
3020:
3013:
3006:
3005:
3000:
2995:
2988:
2983:
2981:
2974:, p. 462
2973:
2972:Lydekker 1908
2968:
2966:
2958:
2953:
2945:
2939:
2932:
2927:
2925:
2917:
2912:
2897:
2891:
2887:
2880:
2873:
2871:
2855:
2851:
2845:
2838:
2833:
2831:
2829:
2817:
2813:
2809:
2805:
2801:
2797:
2793:
2789:
2785:
2781:
2777:
2770:
2768:
2759:
2757:
2742:on 2015-09-24
2738:
2734:
2730:
2726:
2722:
2718:
2714:
2710:
2706:
2699:
2692:
2690:
2688:
2686:
2684:
2682:
2680:
2671:
2665:
2661:
2657:
2650:
2648:
2646:
2638:
2634:
2630:
2625:
2609:
2604:
2600:
2596:
2595:
2590:
2588:
2579:
2577:
2575:
2573:
2571:
2569:
2567:
2565:
2556:
2552:
2547:
2542:
2538:
2534:
2530:
2526:
2522:
2518:
2514:
2506:
2504:
2502:
2500:
2495:
2482:
2476:
2469:
2465:
2461:
2455:
2448:
2442:
2438:
2428:
2425:
2424:
2418:
2415:
2411:
2409:
2405:
2401:
2396:
2391:
2387:
2385:
2381:
2377:
2373:
2368:
2366:
2356:
2354:
2350:
2340:
2337:
2326:
2324:
2320:
2316:
2311:
2307:
2297:
2295:
2287:
2282:
2273:
2271:
2266:
2255:
2252:
2248:
2247:fragmentation
2238:
2236:
2231:
2220:
2218:
2214:
2203:
2201:
2197:
2181:
2179:
2175:
2172:
2169:
2166:
2163:
2160:
2157:
2156:
2152:
2149:
2146:
2143:
2140:
2137:
2135:
2131:
2128:
2127:
2123:
2120:
2116:
2113:
2110:
2107:
2103:
2100:
2098:
2094:
2091:
2090:
2086:
2081:
2078:
2075:
2072:
2069:
2066:
2063:
2060:
2059:
2055:
2051:
2047:
2044:
2042:
2039:
2036:
2033:
2030:
2026:
2025:
2021:
2017:
2013:
2010:
2006:
2004:
2001:
1998:
1995:
1992:
1989:
1988:
1984:
1980:
1977:
1976:
1971:
1968:
1965:
1962:
1959:
1957:
1953:
1950:
1949:
1946:Conservation
1945:
1942:
1939:
1936:
1933:
1930:
1927:
1926:
1923:
1917:
1915:
1911:
1907:
1903:
1899:
1895:
1891:
1887:
1883:
1879:
1869:
1867:
1863:
1859:
1855:
1851:
1848:. Undigested
1847:
1843:
1839:
1835:
1831:
1827:
1823:
1819:
1815:
1811:
1807:
1803:
1795:
1791:
1787:
1778:
1776:
1772:
1768:
1767:
1762:
1761:
1756:
1755:
1750:
1746:
1742:
1739:
1738:
1733:
1729:
1723:
1721:
1716:
1712:
1708:
1704:
1701:
1694:
1689:
1675:
1673:
1668:
1663:
1661:
1655:
1651:
1649:
1639:
1630:
1628:
1624:
1620:
1614:
1612:
1608:
1603:
1599:
1595:
1594:mating season
1585:
1581:
1578:
1566:
1561:
1552:
1549:
1546:
1542:
1537:
1532:
1530:
1529:
1524:
1520:
1516:
1512:
1507:
1503:
1499:
1492:
1487:
1474:
1472:
1468:
1464:
1461:
1457:
1454:
1451:
1446:
1442:
1439:
1438:
1431:
1423:
1414:
1405:
1402:
1399:
1395:
1391:
1387:
1383:
1379:
1377:
1374:
1371:
1368:
1362:
1357:
1353:
1350:
1349:
1345:
1342:
1339:
1336:
1333:
1329:
1327:
1325:
1314:
1312:
1308:
1303:
1299:
1298:golden jackal
1295:
1291:
1287:
1279:
1273:
1271:
1267:
1266:RAD sequences
1263:
1257:
1255:
1251:
1247:
1243:
1242:Oscar Neumann
1239:
1235:
1231:
1227:
1223:
1219:
1218:
1212:
1210:
1206:
1197:
1193:
1189:
1184:
1178:
1177:
1174:See further:
1172:
1171:
1170:
1168:
1164:
1159:
1156:
1152:
1148:
1134:
1133:
1125:
1124:
1116:
1115:
1107:
1106:
1103:
1101:
1097:
1091:
1090:
1087:
1086:
1083:
1081:
1077:
1071:
1070:
1063:
1062:
1056:
1055:
1052:
1051:
1043:
1042:
1039:
1037:
1033:
1027:
1026:
1023:
1022:
1014:
1013:
1010:
1008:
1004:
998:
997:
994:
993:
985:
984:
981:
979:
971:
970:
967:
966:
958:
957:
954:
952:
948:
947:Golden jackal
942:
941:
938:
937:
929:
928:
925:
923:
919:
913:
912:
909:
908:
900:
899:
896:
894:
890:
884:
883:
880:
879:
871:
870:
862:
861:
858:
856:
852:
846:
845:
842:
841:
838:
835:
830:
824:
823:
817:
816:
810:
809:
802:
801:
794:
793:
786:
785:
778:
777:
770:
769:
762:
761:
755:
751:
748:
747:
743:
742:
738:
734:
731:
730:
722:
720:
716:
712:
707:
703:
699:
695:
690:
688:
684:
683:domestic dogs
680:
676:
672:
668:
664:
658:
656:
652:
648:
645:
641:
637:
633:
624:
617:
613:
609:
593:
591:
587:
584:
582:
579:
578:
572:
569:
563:
559:
557:
554:
553:
549:
546:
543:
542:
539:
529:
525:
521:
519:
518:Ethiopic text
499:
497:
493:
489:
488:Simien jackal
485:
481:
471:
469:
465:
461:
457:
453:
449:
448:interbreeding
445:
441:
437:
433:
429:
425:
420:
418:
414:
410:
405:
403:
399:
396:
392:
388:
384:
380:
376:
372:
368:
367:Simien jackal
364:
360:
359:
354:
340:
337:
334:Heuglin, 1862
332:
329:
324:
321:
318:Heuglin, 1862
316:
313:
308:
305:
304:
302:
299:
295:
290:
286:
281:
276:
271:
269:
263:
260:
259:Binomial name
256:
252:
251:
246:
243:
242:
239:
238:
234:
231:
230:
227:
224:
221:
220:
217:
214:
211:
210:
207:
204:
201:
200:
197:
194:
191:
190:
187:
184:
181:
180:
177:
174:
171:
170:
165:
160:
156:
150:
145:
134:
130:
125:
122:
117:
113:
108:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
39:
33:
22:
19:
9551:
9508:
9499:
9490:
9481:
9472:
9463:
9454:
9445:
9436:
9427:
9418:
9409:
9400:
9391:
9382:
9373:
9364:
9344:
9332:
9323:
9314:
9305:
9293:
9268:
9259:
9250:
9238:
9226:
9214:
9203:L. sumatrana
9202:
9193:
9181:
9169:
9152:
9140:
9128:
9119:
9110:
9101:
9089:
9077:
9035:
9026:
9014:
9002:
8986:V. peregusna
8985:
8973:
8940:
8931:
8911:
8899:
8886:Lyncodontini
8877:Ictonychinae
8861:M. zibellina
8860:
8851:
8842:
8833:
8824:
8816:M. flavigula
8815:
8806:
8798:M. americana
8797:
8777:
8765:
8753:
8741:
8729:
8718:Eira barbara
8717:
8691:
8683:M. personata
8682:
8673:
8664:
8655:
8646:
8626:
8610:
8601:
8593:M. canescens
8592:
8583:
8571:
8559:
8550:
8541:
8529:
8500:
8488:
8476:
8395:
8386:
8374:
8362:
8353:
8341:
8312:
8303:
8294:
8285:
8265:
8253:
8244:
8235:
8215:
8203:
8202:Cacomistle (
8194:
8182:
8171:Potos flavus
8170:
8140:
8131:
8122:
8113:
8093:
8081:
8072:
8052:
8040:
8031:
8019:
8007:
7998:
7989:
7980:
7960:
7927:
7868:
7859:
7850:
7841:
7807:
7798:
7778:
7762:
7750:
7738:
7723:(monk seals)
7679:
7670:
7661:
7649:
7637:
7628:
7616:
7604:
7595:
7586:
7560:
7551:
7494:
7486:A. townsendi
7485:
7476:
7468:A. philippii
7467:
7458:
7449:
7440:
7432:A. australis
7431:
7419:
7407:
7390:
7378:
7366:
7345:
7336:
7324:
7312:
7286:
7274:
7234:
7113:U. maritimus
7112:
7111:Polar bear (
7103:
7102:Brown bear (
7094:
7085:
7073:
7061:
7052:
7051:Sloth bear (
7026:
7014:
7002:
6990:
6940:
6938:
6931:
6922:
6913:
6904:
6895:
6886:
6877:
6865:
6854:Cuon alpinus
6853:
6844:
6828:L. mesomelas
6827:
6818:
6806:
6775:
6766:
6757:
6756:Pampas fox (
6748:
6739:
6730:
6718:
6706:
6697:
6681:S. venaticus
6680:
6668:
6656:
6655:Maned wolf (
6637:Cerdocyonina
6605:
6604:Fennec fox (
6596:
6587:
6579:V. rueppelli
6578:
6569:
6560:
6551:
6550:Arctic fox (
6543:V. ferrilata
6542:
6533:
6532:Corsac fox (
6524:
6515:
6506:
6505:Bengal fox (
6484:
6472:
6463:
6451:
6439:
6413:
6412:Island fox (
6404:
6392:
6325:H. sanguinea
6324:
6315:
6306:
6298:H. ichneumon
6297:
6288:
6276:
6264:
6255:
6246:
6234:
6222:
6213:
6204:
6195:
6178:
6169:
6160:
6151:
6142:
6133:
6125:U. edwardsii
6124:
6116:U. brachyura
6115:
6106:
6094:
6082:
6073:
6043:
6034:
6025:
6017:C. alexandri
6016:
6004:
5992:
5983:
5971:
5959:
5942:
5934:M. gambianus
5933:
5921:
5909:
5893:S. suricatta
5892:
5880:
5855:
5830:
5821:
5809:
5797:
5780:
5771:
5759:
5747:
5717:
5708:
5696:
5684:
5675:
5652:carnivorans)
5633:
5604:
5592:
5580:
5571:
5539:
5527:
5497:Herpestoidea
5460:Herpestoidea
5427:
5419:G. victoriae
5418:
5409:
5408:Cape genet (
5400:
5392:G. servalina
5391:
5382:
5381:King genet (
5374:G. piscivora
5373:
5364:
5355:
5346:
5338:G. johnstoni
5337:
5328:
5319:
5310:
5301:
5292:
5274:
5263:P. leightoni
5262:
5253:
5235:
5210:
5201:
5193:V. megaspila
5192:
5184:V. civettina
5183:
5171:
5159:
5150:
5135:
5119:
5092:
5083:
5074:
5062:
5050:
5041:
5032:
5023:
4997:
4988:
4979:
4970:
4868:
4859:
4851:F. margarita
4850:
4841:
4832:
4831:Jungle cat (
4823:
4814:
4802:
4786:
4777:
4769:P. planiceps
4768:
4759:
4750:
4740:Prionailurus
4738:
4726:
4696:
4684:
4672:
4671:Jaguarundi (
4655:
4643:
4622:
4613:
4604:
4595:
4583:
4571:
4562:
4553:
4544:
4535:
4526:
4518:L. geoffroyi
4517:
4508:
4507:Pampas cat (
4496:
4480:
4471:
4459:
4447:
4417:
4408:
4396:
4385:P. marmorata
4384:
4372:
4352:
4327:
4318:
4309:
4300:
4291:
4279:
4267:
4258:
4246:
4201:
4192:
4175:
4148:
4101:Superorder:
4095:Infraclass:
4022:
3962:, July 2004.
3959:
3948:
3926:. Retrieved
3919:the original
3900:
3882:
3867:
3853:
3838:. Retrieved
3831:the original
3822:
3799:. Retrieved
3788:
3780:
3764:
3752:. Retrieved
3748:
3735:
3730:, p. 30
3723:
3718:, p. 29
3711:
3699:
3687:
3675:
3670:, p. 22
3663:
3651:
3646:, p. 10
3639:
3628:
3582:
3578:
3568:
3555:
3543:
3531:
3498:
3494:
3488:
3469:
3465:
3455:
3422:
3418:
3405:
3373:
3369:
3356:
3336:, p. 32
3281:
3277:
3266:
3258:
3253:
3218:
3214:
3203:
3184:
3180:
3170:
3158:
3146:
3137:
3096:
3092:
3085:
3042:
3038:
3027:
3017:
3012:
3002:
2994:
2959:, p. 39
2957:Rüppell 1835
2952:
2938:
2911:
2899:. Retrieved
2885:
2857:. Retrieved
2853:
2844:
2816:the original
2779:
2775:
2766:
2744:. Retrieved
2737:the original
2711:(485): 1–6.
2708:
2704:
2659:
2629:Rüppell 1835
2624:
2612:. Retrieved
2598:
2592:
2586:
2520:
2516:
2475:
2454:
2441:
2416:
2412:
2392:
2388:
2369:
2362:
2359:Conservation
2352:
2346:
2332:
2303:
2294:soil erosion
2290:
2261:
2244:
2226:
2223:Habitat loss
2213:vaccinations
2209:
2192:
2138:870 km
2118:
2009:habitat loss
1973:
1921:
1875:
1830:rock hyraxes
1799:
1793:
1769:bushes, and
1764:
1758:
1752:
1735:
1724:
1697:
1664:
1657:
1653:
1644:
1615:
1591:
1582:
1574:
1550:
1533:
1526:
1515:canine teeth
1506:African wolf
1495:
1444:
1440:
1429:
1426:(Gray, 1869)
1421:
1412:
1403:
1355:
1351:
1340:Description
1326:Volume Three
1322:
1320:
1289:
1283:
1277:
1262:Canis aureus
1261:
1258:
1249:
1237:
1215:
1213:
1209:interglacial
1201:
1173:
1166:
1160:
1144:
1094:
1074:
1030:
1001:
974:
945:
918:African wolf
916:
887:
849:
829:Domestic dog
827:
691:
659:
647:hypothesised
629:
615:
611:
589:
585:
567:
561:
537:
515:
495:
491:
487:
484:Simenian fox
483:
479:
477:
421:
406:
383:horse jackal
382:
370:
366:
362:
357:
356:
352:
350:
338:
330:
322:
315:C. semiensis
314:
307:C. crinensis
306:
267:
265:
249:
248:
236:
18:
9666:iNaturalist
9576:Wikispecies
9500:M. sibirica
9482:M. putorius
9455:M. nigripes
9437:M. lutreola
9410:M. haidarum
9306:N. africana
9269:A. congicus
9260:A. cinereus
9251:A. capensis
9151:Sea otter (
9129:L. provocax
9036:I. striatus
8957:Ictonychini
8852:M. melampus
8764:Wolverine (
8742:P. pennanti
8665:M. moschata
8656:M. everetti
8602:M. leucurus
8560:A. hoevenii
8551:A. collaris
8501:M. capensis
8396:N. olivacea
8311:Olinguito (
8267:Bassaricyon
8254:P. pygmaeus
8184:Bassariscus
8157:Procyonidae
8132:S. putorius
8123:S. gracilis
8041:M. mephitis
8032:M. macroura
7926:Red panda (
7906:Musteloidea
7867:Ross seal (
7820:Lobodontini
7752:Neomonachus
7680:P. sibirica
7638:P. vitulina
7603:Grey seal (
7585:Harp seal (
7477:A. pusillus
7441:A. forsteri
7276:Callorhinus
7210:Pinnipedia
7169:Musteloidea
6941:C. simensis
6905:C. lupaster
6774:Hoary fox (
6767:L. sechurae
6740:L. fulvipes
6731:L. culpaeus
6627:(true dogs)
6586:Swift fox (
6561:V. macrotis
6453:Nyctereutes
6362:Caniformia
6307:H. ochracea
6265:B. nigripes
6256:B. jacksoni
6143:U. javanica
6060:Herpestinae
6035:C. obscurus
6026:C. ansorgei
6006:Crossarchus
5862:(mongooses)
5851:Herpestidae
5831:S. durrelli
5822:S. concolor
5772:G. fasciata
5709:E. goudotii
5630:Herpestidae
5540:P. cristata
5401:G. thierryi
5356:G. maculata
5320:G. cristata
5311:G. bourloni
5064:Paradoxurus
5022:Binturong (
4961:Hemigalinae
4938:Viverroidea
4901:Viverroidea
4860:F. nigripes
4712:Leopard cat
4697:P. concolor
4614:L. pardinus
4563:L. tigrinus
4554:L. pardalis
4545:L. jacobita
4536:L. guttulus
4509:L. colocola
4268:N. nebulosa
4238:Pantherinae
4118:Feliformia
3928:15 December
3814:Cited works
3801:23 February
3351:, p. 4
3187:: 119–199.
2933:, p. 2
2614:19 November
2323:saddle pads
2276:Overgrazing
2270:brush fires
2200:overgrazing
2119:Helichrysum
2093:Guassa Menz
2077:Overgrazing
1975:Helichrysum
1940:Importance
1906:Mount Choqa
1894:Guassa Menz
1878:Rift Valley
1754:Helichrysum
1519:carnassials
1481:Description
1460:nasal bones
1430:C. s. walgi
1394:Guassa Menz
1382:Rift Valley
1334:Subspecies
1167:C. simensis
1161:In 1994, a
715:Mount Choqa
464:vaccination
440:overgrazing
339:C. citernii
323:C. simensis
30:1.6–0
9844:Categories
9473:M. nudipes
9464:M. nivalis
9428:M. kathiah
9383:M. erminea
9374:M. altaica
9324:N. frenata
9315:N. felipei
9285:Mustelinae
9027:I. libycus
8941:G. vittata
8807:M. caurina
8584:M. anakuma
8462:Mustelidae
8449:Mustelidae
8425:see below↓
8416:Mustelidae
8313:B. neblina
8224:(raccoons)
8195:B. astutus
8193:Ringtail (
8169:Kinkajou (
8141:S. pygmaea
8082:M. marchei
7940:Mephitidae
7808:M. leonina
7700:Monachinae
7671:P. hispida
7662:P. caspica
7459:A. gazella
7391:N. cinerea
7287:C. ursinus
7225:Odobenidae
7178:see below↓
7158:see below↓
7142:Pinnipedia
7060:Sun bear (
7027:T. ornatus
7016:Tremarctos
6992:Ailuropoda
6930:Red wolf (
6896:C. latrans
6819:L. adustus
6776:L. vetulus
6749:L. griseus
6679:Bush dog (
6570:V. pallida
6568:Pale fox (
6552:V. lagopus
6523:Cape fox (
6403:Gray fox (
6161:U. smithii
5993:H. parvula
5984:H. hirtula
5872:Mungotinae
5761:Galidictis
5734:Galidiinae
5662:Euplerinae
5645:Eupleridae
5634:sensu lato
5605:C. crocuta
5538:Aardwolf (
5469:see below↓
5410:G. tigrina
5383:G. poensis
5365:G. pardina
5347:G. letabae
5329:G. genetta
5227:Genettinae
5211:V. zibetha
5131:Viverrinae
5120:sensu lato
5115:Viverrinae
5084:P. jerdoni
4951:Viverridae
4910:see below↓
4849:Sand cat (
4656:A. jubatus
4481:C. caracal
4374:Pardofelis
4193:P. linsang
4134:Feliformia
3840:2012-12-15
3749:Canids.org
2746:2011-09-22
2523:(1): 530.
2490:References
2447:afterbirth
2406:, and the
2031:Highlands
1902:Mount Guna
1806:grass rats
1775:ericaceous
1763:tussocks,
1737:Alchemilla
1707:heathlands
1703:grasslands
1700:Afroalpine
1627:homozygous
1619:inbreeding
1400:highlands
1390:Mount Guna
1380:Northwest
1317:Subspecies
1196:maned wolf
651:greyhounds
575:Trickster
573:Red jackal
480:red jackal
470:programs.
424:endangered
395:Afroalpine
391:generalist
371:Simien fox
363:red jackal
326:Gray, 1869
144:Endangered
9419:M. itatsi
9353:(weasels)
9216:Lutrogale
9153:E. lutris
9111:L. felina
8920:(grisons)
8843:M. martes
8786:(martens)
8704:Guloninae
8490:Mellivora
8363:N. narica
8304:B. medius
8295:B. gabbii
8286:B. alleni
8274:(olingos)
8243:Raccoon (
8095:Spilogale
7981:C. chinga
7962:Conepatus
7918:Ailuridae
7717:Monachini
7629:P. largha
7299:Otariinae
7262:sea lions
7257:fur seals
7248:Otariidae
7133:Mustelida
7104:U. arctos
6966:Arctoidea
6923:C. lycaon
6878:C. aureus
6808:Lupulella
6720:Lycalopex
6597:V. vulpes
6595:Red fox (
6559:Kit fox (
6534:V. corsac
6278:Herpestes
5891:Meerkat (
5650:(Malagasy
5552:Hyaeninae
5513:Hyaenidae
5428:G. felina
4842:F. lybica
4654:Cheetah (
4572:L. wiedii
4561:Oncilla (
4527:L. guigna
4498:Leopardus
4479:Caracal (
4472:C. aurata
4407:Bay cat (
4328:P. pardus
4326:Leopard (
4301:P. tigris
4259:N. diardi
4177:Prionodon
4077:Kingdom:
4066:Carnivora
3994:from the
3425:: 39–45.
2733:253988540
2537:2399-3642
2468:grey wolf
1896:in north
1866:livestock
1818:vlei rats
1730:and deep
1715:tree line
1598:Courtship
1555:Behaviour
1511:premolars
1346:Synonyms
1284:In 2018,
1188:grey wolf
851:Gray wolf
719:carnivore
694:Highlands
640:Abyssinia
562:Ky kebero
402:carnivore
331:C. walgié
244:Species:
216:Carnivora
182:Kingdom:
176:Eukaryota
9809:Species+
9749:14000779
9684:10762843
9561:Wikidata
9399:Ferret (
9333:N. vison
9194:L. lutra
9062:(otters)
9056:Lutrinae
8913:Galictis
8825:M. foina
8740:Fisher (
8628:Melogale
8611:M. meles
8531:Arctonyx
8376:Nasuella
8354:N. nasua
8331:(coatis)
8245:P. lotor
8021:Mephitis
7780:Mirounga
7530:Phocinae
7511:Phocidae
7380:Neophoca
7326:Zalophus
7233:Walrus (
6932:C. rufus
6914:C. lupus
6894:Coyote (
6729:Culpeo (
6670:Speothos
6606:V. zerda
6588:V. velox
6525:V. chama
6236:Bdeogale
6134:U. fusca
5973:Helogale
5943:M. mungo
5882:Suricata
5811:Salanoia
5718:E. major
5698:Eupleres
5529:Proteles
5518:(hyenas)
5282:(genets)
4833:F. chaus
4824:F. catus
4815:F. bieti
4695:Cougar (
4645:Acinonyx
4623:L. rufus
4621:Bobcat (
4570:Margay (
4552:Ocelot (
4525:Kodkod (
4446:Serval (
4409:C. badia
4398:Catopuma
4308:Jaguar (
4292:P. uncia
4281:Panthera
4248:Neofelis
4166:Feloidea
4097:Eutheria
4091:Mammalia
4085:Chordata
4083:Phylum:
4079:Animalia
4008:Archived
3898:(1997).
3880:(1835).
3865:(1902).
3851:(1908).
3773:Archived
3754:19 March
3515:19834483
3447:46671897
3439:23994497
3316:30344120
3245:26234211
3129:25976335
3077:20237475
3001:(1853).
2859:19 March
2812:40027094
2804:15245401
2555:37193884
2546:10187515
2421:See also
2310:firearms
2095:, North
2070:Stable?
2002:Extinct
1991:Mt. Guna
1966:Stable?
1954:, North
1943:Threats
1931:Habitat
1854:roughage
1838:reedbuck
1826:goslings
1751:rich in
1728:drainage
1328:(MSW3).
1151:mandible
596:Reddish
560:ቀይ ቀበሮ (
438:through
298:Synonyms
222:Family:
206:Mammalia
196:Chordata
192:Phylum:
186:Animalia
172:Domain:
149:IUCN 3.1
9822:ZooBank
9736:1005946
9658:5219143
9567:Q188957
9401:M. furo
9346:Mustela
9295:Neogale
9142:Enhydra
9016:Ictonyx
8975:Vormela
8932:G. cuja
8859:Sable (
8766:G. gulo
8731:Pekania
8716:Tayra (
8513:Melinae
8325:Nasuina
8217:Procyon
7573:Phocini
7212:(seals)
7147:(seals)
7039:Ursinae
6981:(bears)
6976:Ursidae
6852:Dhole (
6642:(zorro)
6516:V. cana
6426:Vulpini
6394:Urocyon
6378:Canidae
6170:U. urva
5674:Fossa (
5594:Crocuta
5276:Genetta
5173:Viverra
4714:lineage
4636:lineage
4605:L. lynx
4461:Caracal
4435:lineage
4433:Caracal
4365:lineage
4363:Bay cat
4349:Felinae
4310:P. onca
4299:Tiger (
4219:Felidae
4089:Class:
4068:species
4064:Extant
4014:at the
3998:of the
3984:of the
3982:WildCRU
3869:natives
3307:6224481
3286:Bibcode
3223:Bibcode
3121:7921357
3101:Bibcode
3068:3494089
3047:Bibcode
2854:Awf.org
2784:Bibcode
2725:3504136
2308:, when
2235:Delanta
2206:Disease
2189:Threats
2161:, Bale
2085:Denkoro
1937:Status
1766:Euryops
1760:Festuca
1745:tussock
1718:times,
1711:habitat
1683:Habitat
1678:Ecology
1672:geladas
1523:dewclaw
1375:, 1840
1373:Rüppell
1307:admixed
1280:species
1238:Simenia
1222:jackals
750:Caninae
735:of the
590:Arouayé
556:Amharic
426:by the
398:rodents
373:, is a
275:Ruppell
232:Genus:
226:Canidae
212:Order:
202:Class:
147: (
99:↓
9860:Wolves
9801:433177
9788:752755
9710:621851
9645:327993
9629:ECOS:
9604:ARKive
9091:Lontra
8779:Martes
8054:Mydaus
7948:skunks
6912:Wolf (
6792:Canina
6622:Canini
6486:Vulpes
6382:(dogs)
5923:Mungos
5237:Poiana
4319:P. leo
4317:Lion (
4223:(cats)
4028:ARKive
3911:
3523:771357
3521:
3513:
3445:
3437:
3314:
3304:
3243:
3127:
3119:
3075:
3065:
3039:Nature
2901:4 July
2892:
2810:
2802:
2731:
2723:
2666:
2637:pl. 14
2553:
2543:
2535:
2265:cereal
2230:barley
2087:area.
2027:North
2020:kebele
2016:woreda
1956:Gondar
1886:Gondar
1832:, and
1812:, and
1749:scrubs
1667:cattle
1545:moults
1543:. It
1343:Range
1240:, and
889:Coyote
702:Gojjam
671:rabies
568:Walgie
494:, and
486:, the
482:, the
474:Naming
442:, and
387:coyote
375:canine
365:, the
277:, 1840
9855:Canis
9814:10871
9775:82242
9762:32534
9697:90294
9679:IRMNG
9671:42045
9622:5WXTW
9240:Aonyx
9183:Lutra
8573:Meles
8343:Nasua
7618:Phoca
7075:Ursus
6867:Canis
6494:foxes
4804:Felis
3922:(PDF)
3905:(PDF)
3834:(PDF)
3827:(PDF)
3745:(PDF)
3519:S2CID
3443:S2CID
3415:(PDF)
3366:(PDF)
3125:S2CID
2882:(PDF)
2819:(PDF)
2808:S2CID
2772:(PDF)
2740:(PDF)
2729:S2CID
2721:JSTOR
2701:(PDF)
2633:p. 39
2433:Notes
2365:CITES
2319:goats
2315:sheep
2029:Wollo
1928:Area
1850:sedge
1741:herbs
1732:soils
1577:packs
1536:ochre
1528:Canis
1398:Wollo
1302:basal
1290:Canis
1278:Canis
1250:Canis
1234:genus
1217:Canis
1003:Dhole
803:0.38
787:0.96
581:Oromo
566:ዋልጌ (
450:from
409:range
237:Canis
9757:NCBI
9723:3748
9718:IUCN
9705:ITIS
9653:GBIF
9632:1640
8755:Gulo
7651:Pusa
6096:Urva
4686:Puma
4634:Puma
4585:Lynx
3930:2012
3909:ISBN
3803:2016
3756:2022
3511:PMID
3435:PMID
3312:PMID
3241:PMID
3117:PMID
3073:PMID
2903:2019
2890:ISBN
2861:2022
2800:PMID
2664:ISBN
2616:2021
2599:2011
2551:PMID
2533:ISSN
2317:and
2286:zebu
2134:Bale
2097:Shoa
2048:The
2018:and
1912:and
1910:Arsi
1898:Shoa
1860:and
1781:Diet
1705:and
1609:and
1592:The
1500:and
1469:and
1467:Arsi
1311:wolf
1228:and
1064:2.6
795:0.6
779:2.0
771:2.5
763:3.0
466:and
446:and
428:IUCN
351:The
38:PreꞒ
9744:MSW
9731:MDD
9692:ISC
9640:EoL
9617:CoL
9591:ADW
4026:at
3980:at
3587:doi
3503:doi
3474:doi
3427:doi
3423:122
3378:doi
3374:245
3302:PMC
3294:doi
3231:doi
3189:doi
3109:doi
3063:PMC
3055:doi
3043:464
2792:doi
2713:doi
2709:385
2603:doi
2541:PMC
2525:doi
2464:dog
2460:SNP
1884:in
713:in
490:,
369:or
9846::
9824::
9811::
9798::
9785::
9772::
9759::
9746::
9733::
9720::
9707::
9694::
9681::
9668::
9655::
9642::
9619::
9606::
9593::
9578::
9563::
3958:.
3947:.
3894:;
3779:,
3747:.
3627:.
3616:^
3601:^
3583:96
3581:.
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3509:.
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3497:.
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3183:.
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3095:.
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3037:.
2979:^
2964:^
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2852:.
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2806:.
2798:.
2790:.
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2778:.
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2769:)"
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2644:^
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2597:.
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1272:.
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754:Ma
721:.
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498:.
419:.
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88:Pg
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