602:. Branches of the dorsal nerve extend through the glans ventrolaterally displaying a three-dimensional innervation pattern. The main branches form smaller bundles of nerves that expand outwards into the tissue of the glans. The rich innervation of the glans penis reveals its function as a primary anatomical source of male sexual pleasure. Yang & Bradley argue; "the distinct pattern of innervation of the glans emphasizes its role as a sensory structure". While Yang & Bradley's (1998) report "showed no areas in the glans to be more densely innervated than others.", Halata & Munger (1986) report that the density of several nerve terminals is greatest in the corona glandis.
781:
538:
penis can vary among males. While the shape of the glans is typically acorn-like, in some men it might be wider in circumference than the shaft, giving the penis a mushroom-like appearance, while in others it might be narrower and more akin to a probe in shape. Some researchers have suggested that the glans has evolved to become acorn-, mushroom- or cone-shaped so that during copulation it acts to remove any semen still there from previous sex partners, but this is not supported when looking at primate relatives who have different mating behaviors.
421:
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702:. The degree of automatic foreskin retraction varies considerably depending on the foreskin length. The foreskin can be characterized as long when the preputial orifice extends beyond the glans during erection or medium when the orifice is located around the meatus. The primary purpose of the foreskin is considered to be the covering of the glans and the
863:, the complete external differentiation of the organs begins about eight or nine weeks after conception. Some sources state that the process will be completed by the twelfth week, while others state that it is clearly evident by the thirteenth week and that the sex organs are fully developed by the sixteenth week.
637:) are not present". The genital end bulbs, that are present throughout the glans, are most numerous in the corona and near the frenulum. Simple, Pacinian and Ruffinian corpuscles are identified predominantly in the corona glandis. The most numerous nerve terminals are free nerve endings present in almost every
673:
with the axial arteries through perforating branches before they end in the glans. Branches of the dorsal artery curve around each side of the distal shaft to enter the glans and the frenulum ventrally. Venous drainage of the penis begins at the base of the glans. Small tributaries deriving from the
761:
or were circumcised as infants, a procedure where the foreskin is partially or completely removed from the penis. The glans of circumcised men remains fully exposed and dry. Several studies have suggested the glans is generally equally sensitive in both circumcised and uncircumcised penises, though
537:
fills the erectile tissue with blood causing the glans to grow in size and sensitivity. While the penis is rigid when erect, the glans itself remains slightly softer. The soft cushiony texture of the glans absorbs impact during rigorous instances of copulation. The proportional size of the glans
577:
elastic band of tissue located on the underside of the glans that connects the foreskin to the head of the penis. The frenulum is supple enough to allow the retraction of the foreskin over the glans and pull it back when the erection is gone. In flaccid state, it tightens to narrow the foreskin
988:, a condition mostly seen as a late complication of circumcision. It occurs in about 2–20% of circumcised boys and it is rarely seen in uncircumcised men. It is characterized by a narrowing of the meatus, which might cause sudden or often urges to urinate and burning during the process.
1119:, the glans penis is rounded, short, and small and is superficially divided into left and right halves by a trough at the top and a ridge at the bottom. Most of the glans is covered with spines, except for an area near the tip. The glans penis of a male
1801:
973:. Symptoms of balanitis may appear suddenly or develop gradually. They might include pain, irritation, redness or red patches on the glans penis. Careful identification of the cause with the aid of patient history, physical
2399:"Preputial variability and preferential association of long phimotic foreskins with penile cancer: An anatomic comparative study of types of foreskin in a general population and cancer patients"
1798:
2773:
Clifford B., Bleustein; Fogarty, James D.; Eckholdt, Haftan; Arezzo, Joseph C.; Melman, Arnold (April 2005). "Effect of neonatal circumcision on penile neurologic sensation".
3465:
965:. It is a treatable condition that occurs in about 3–11% of males (up to 35% of diabetic males). Edwards reported that it is generally more common in males who have poor
918:. The secretion of testosterone during this phase plays a decisive role in the final shaping of the penis. After birth, testosterone levels drop significantly until
2961:
2857:
Payne, Kimberley; Thaler, Lea; Kukkonen, Tuuli; Carrier, Serge; Binik, Yitzchak (May 2007). "Sensation and sexual arousal in circumcised and uncircumcised men".
223:
2717:... the incidence of non-retractable physiological phimosis was 50% in grade 1 boys and decreased to 35% in grade 4 and 8% in grade 7 boys
1384:
Keenan-Lindsay, Lisa; Sams, Cheryl; O'Connor, Constance; Perry, Shannon; Hockenberry, Marilyn; Leonard
Lowdermilk, Deitra; Wilson, David (December 17, 2021).
565:. Behind the corona is the neck of the penis, which separates the glans and the penile shaft. Ventrally, the two glans wings merge on the midline forming the
3697:
Equine
Clinical Medicine, Surgery and Reproduction - Graham Munroe BVSc (Hons) PhD Cert EO DESM Dip ECVS FRCVS, Scott Weese DVM DVSc DipACVIM - Google Boeken
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For some individuals who experience difficulty in achieving full glanular engorgement of glans penis, they may be diagnosed with soft glans syndrome (
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1982:
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1151:, a small pouch at the distal end of the glans. Unlike the human glans, the glans of a horse's penis extends backwards on its shaft.
1023:
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678:, plexus. Smaller paired venules run into the frenulum and the glans from its ventral surface. The deep dorsal vein, one of the two
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fetus the absence of testosterone will stop the growth of the phallus causing the tubercle to shrink and form the clitoris. In the
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allowing the glans to be completely exposed when needed. By the age of eighteen most boys will have a fully retractable foreskin.
4783:
2705:
2634:
1519:
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HSU, G‐L., et al. "The distribution of elastic fibrous elements within the human penis." BJU International 73.5 (1994): 566-571.
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363:. In adults, the foreskin can generally be retracted over and past the glans manually or sometimes automatically during an
3942:
2063:
698:. In adults, glans exposure can be easily achieved by manual retraction of the foreskin or sometimes automatically during
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report that excessive washing with soap may dry the mucous membrane which covers the glans penis and cause non-specific
3309:
2953:
2442:
2231:
1677:
1602:
4044:
2356:
Hsu, Geng-Long; Hsieh, Cheng-Hsing; Wen, Hsien-Sheng; Chen, Yi-Chang; Chen, Shyh-Chyan; Mok, Martin S. (2003-11-12).
1312:
Yang, C. C.; W.E. Bradley (July 1998). "Neuroanatomy of the penile portion of the human dorsal nerve of the penis".
335:, which creates a smooth texture and glossy appearance. In humans, the glans is located over the distal ends of the
1278:
981:
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of the phenotypically indifferent organs. When exposed to testosterone, the genital tubercle elongates to form the
969:
habits or have not been circumcised. It has many causes, including irritation or infection with a wide variety of
4620:
4330:
4252:
4247:
3046:
Merz, Eberhard; Bahlmann, F. (2004). Ultrasound in
Obstetrics and Gynecology. Vol. 1. Thieme Medical Publishers.
2981:
Merz, Eberhard; Bahlmann, F. (2004). Ultrasound in
Obstetrics and Gynecology. Vol. 1. Thieme Medical Publishers.
3573:
507:
375:
3631:
McCurnin's
Clinical Textbook for Veterinary Technicians - Joanna M Bassert, Dennis M McCurnin - Google Boeken
218:
4778:
4496:
4469:
4425:
2397:
Velazquez, Elsa F.; Bock, Adelaida; Soskin, Ana; Codas, Ricardo; Arbo, Manuel; Cubilla, Antonio L. (2003).
2299:"Meatal stenosis after surgical correction of short frenulum: Is the "pull-and-burn" method the way to go?"
992:
3337:
2925:
Arulkumaran, Sabaratnam; Regan, Lesley; Papageorghiou, Aris; Monga, Ash; Farquharson, David (2011-06-23).
946:. The condition is described as an inflammation of the skin, often caused by an irritating substance or a
409:
The glans is more commonly known as the "head" or the "tip" of the penis, and colloquially referred to in
34:
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can most readily be distinguished from its close relatives by its partially corrugated glans penis. In
658:
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of typical skin. The papillary layer of the dermis blends into the dense connective tissue forming the
146:
3595:
995:). It is often undiagnosed in the general population due to the lack of a standardized nomenclature.
4711:
4688:
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At birth, the inner foreskin is usually fused to the glans penis and should not be forcibly retracted
950:. Sensitivity to chemicals in certain products can cause an allergic reaction, including irritation,
679:
591:
340:
170:
158:
30:
1954:
1293:...the most pleasurable of all body parts when stimulated sexually: the glans (or tip) of the penis.
1109:
is long and robust, averaging 7.3 mm (0.29 in) long and 4.6 mm (0.18 in) broad.
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exit the penis. The circumference of the base of the glans forms a rounded projecting border, the
4743:
4726:
4556:
4521:
4447:
4437:
4388:
4220:
4114:
3508:
Bradley, R.D.; Schmidley, D.J. (1987). "The glans penes and bacula in Latin
American taxa of the
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by curving the tip of the glans penis backward. In cats, the glans penis is covered with spines.
903:
683:
626:
491:
336:
2660:"CUA guideline on the care of the normal foreskin and neonatal circumcision in Canadian infants"
2589:"CUA guideline on the care of the normal foreskin and neonatal circumcision in Canadian infants"
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494:, extending farther on their upper than on their lower surfaces. It is the expanded cap of the
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1247:
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Comparative
Reproductive Biology - Heide Schatten, Gheorghe M. Constantinescu - Google Boeken
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2004:
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The glans is completely or partially covered by a double-layered fold of skin, known as the
4773:
4317:
3763:
Horse
Conformation: Structure, Soundness, and Performance - Equine Research - Google Boeken
1878:"Male circumcision: Global trends and determinants of prevalence, safety and acceptability"
1120:
977:, swabs and cultures, and biopsy are essential in order to determine the proper treatment.
890:, which secrete a large amount of hormones called androgens. These hormones will cause the
845:
674:
corona form a venous plexus at the neck of the penis, known as the retro-coronal, or retro-
541:
At the summit of the glans is the slit-like vertical external urethral orifice, called the
134:
8:
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3664:
Horseman's
Veterinary Encyclopedia, Revised and Updated - Equine Research - Google Boeken
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398:) initiates the tubercle's development into a penis making the glans penis anatomically
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399:
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McKinnon, Angus O; Squires, Edward L; Vaala, Wendy E; Varner, Dickson D (2011-07-05).
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where it may appear smooth, spiny, elongated or divided. It is externally lined with
2808:
Bleustein, Clifford B.; Eckholdt, Haftan; Arezzo, Joseph C.; Melman, Arnold (2003).
2438:
1673:
1634:"The neuroanatomical basis for the protopathic sensibility of the human glans penis"
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269:
112:
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4303:
4161:
4136:
3793:
Evans, Warren J; Borton, Anthony; Hintz, Harold; Dale Van Vleck, L (1990-02-15).
3730:
Veterinary
Anatomy of Domestic Mammals: Textbook and Colour Atlas - Google Boeken
2041:
1805:
1351:
985:
984:(opening) of the urethra located at the tip of the glans might become subject to
707:
630:
605:
Halata & Spathe (1997) reported; "the glans penis contains a predominance of
570:
523:
487:
414:
410:
352:
316:
3261:"Meatal Stenosis: Symptoms, Diagnosis & Treatment - Urology Care Foundation"
2901:
2036:. Advances in Experimental Medicine and Biology. Vol. 424. pp. 265–6.
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2497:
Dobanovacki D, Lucić Prostran B, Sarac D, Antić J, Petković M, Lakić T (2012).
2358:"Penile Venous Anatomy: An Additional Description and Its Clinical Implication"
2031:
1914:
1148:
1106:
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1073:
911:
763:
735:
718:
703:
682:, serves as a common vessel receiving blood drained from the glans and the two
638:
599:
595:
566:
542:
344:
308:
1715:
1491:
Schünke, Michael; Schulte, Erik; Lamperti, Edward D.; Schumacher, Udo (2006).
1249:
Carnal Knowledge: A Navel Gazer's Dictionary of Anatomy, Etymology, and Trivia
1071:. In male dogs the glans penis is smooth and consists of two parts called the
4762:
4668:
4432:
4415:
4410:
4405:
4084:
2683:
2612:
2565:
2483:
2422:
2324:
2219:
2119:
2030:
Halata, Zdenek; A. Spaethe (1997). "Sensory Innervation of the Human Penis".
1723:
1657:
1086:
1068:
1064:
1060:
902:—elongated spindle-shaped structures that contribute to the formation of the
871:
860:
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for the protein TDF (testis determining factor) and triggers the creation of
813:
662:
634:
550:
391:
251:
211:
2398:
2268:
2165:
808:. Initially undifferentiated, the tubercle develops into a penis during the
4625:
4486:
4281:
4199:
4179:
4146:
4036:
3200:
3182:
2878:
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2794:
2756:
2701:
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2573:
2524:
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2383:
2342:
2195:
2127:
1978:
1926:
1741:
1700:"Physiology of Penile Erection and Pathophysiology of Erectile Dysfunction"
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129:
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4397:
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4225:
4141:
3570:"The Stallion: Breeding Soundness Examination & Reproductive Anatomy"
3137:"Balanitis: Types, Symptoms, Causes, Treatments, Prevention & Relief"
3097:. Archived from the original on 13 June 2013. Retrieved 27 October 2015.
2515:
2498:
895:
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503:
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304:
79:
2731:"Pathologic and physiologic phimosis: Approach to the phimotic foreskin"
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the foreskin is fused to the glans, it remains non retractable in early
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1094:
943:
931:
837:
312:
107:
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Dave, Sumit; Afshar, Kourosh; Braga, Luis H.; Anderson, Peter (2018).
2604:
2587:
Dave, Sumit; Afshar, Kourosh; Braga, Luis H.; Anderson, Peter (2018).
1562:
236:
4189:
4075:
2954:"Genetic Mechanisms of Sex Determination | Learn Science at Scitable"
1160:
962:
886:
fetus the presence of a Y chromosome leads to the development of the
874:
of the penis and this shared embryonic origin makes these two organs
805:
731:
359:, it is completely or partially covered by a fold of skin called the
3525:
3453:
3108:"Contact Dermatitis: Irritants, Allergies, Symptoms & Treatment"
522:
of the corpus spongiosum behind the glans. The external lining with
4464:
4337:
4093:
3597:
Mating Males: An Evolutionary Perspective on Mammalian Reproduction
2496:
2093:
1494:
Thieme Atlas of Anatomy: General Anatomy and Musculoskeletal System
1428:(Second, Revised ed.). Butterworth-Heinemann. pp. 57–79.
1383:
1171:
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887:
817:
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695:
534:
395:
364:
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62:
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4652:
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935:
919:
915:
739:
727:
499:
91:
3727:
König, Horst Erich; Hans-Georg, Hans-Georg; Bragulla, H (2007).
2892:
Warren, John (August 2010). "Physical Effects of Circumcision".
2772:
2729:
McGregor, Thomas B.; Pike, John G.; Leonard, Michael P. (2007).
2003:
Greenberg, Jerrold; Bruess, Clint; Conklin, Sarah (2010-03-10).
1545:
Lee, Shin-Hyo; Ha, Tae-Jun; Koh, Ki-Seok; Song, Wu-Chul (2019).
1490:
386:
as an outgrowth in the caudal region that later develops into a
4491:
2468:(3rd ed.). New York, NY: Informa Healthcare. p. 283.
879:
797:
642:
530:
515:
379:
332:
328:
141:
3792:
3694:
Weese, Scott; Graham Munroe, Dr; Munroe, Graham (2011-03-15).
1518:
Schatten, Heide; Constantinescu, Gheorghe M. (21 March 2008).
526:
is responsible for its typical smooth texture and appearance.
2807:
2255:
Quartey, J. K.M. (2006), Schreiter, F.; Jordan, G.H. (eds.),
2152:
Quartey, J. K.M. (2006), Schreiter, F.; Jordan, G.H. (eds.),
1913:
Weech, David; Ameer, Muhammad Atif; Ashurst, John V. (2022),
1595:"penis | Description, Anatomy, & Physiology | Britannica"
1090:
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546:
324:
293:
182:
165:
122:
55:
3920:
A monographic study of the American marsupial, Caēnolestes …
3507:
1163:
species. In most marsupials, the glans is divided, but male
955:
951:
883:
478:
470:
248:
153:
54:
This article is about the glans penis. For other uses, see
3858:
3826:
Schatten, Heide; Constantinescu, Gheorghe M (2008-03-21).
3089:
Sloane, Ethel (2002). Biology of Women. Cengage Learning.
2856:
1349:
1273:
Olausson, Håkan; Wessberg, Johan; Morrison, India (2016).
726:
Foreskin rectractability gradually increases with age. In
3891:
3404:
Evans, Howard E.; de Lahunta, Alexander (7 August 2013).
1511:
1033:
any relevant information into other sections or articles.
841:
801:
641:
of the glans, as well as scattered throughout the deeper
482:
The glans penis as the expansion of the corpus spongiosum
383:
278:
272:
3917:
Osgood, Wilfred Hudson; Herrick, Charles Judson (1921).
3693:
3310:"Soft Glans / Cold Glans / Glans Insufficiency Syndrome"
2396:
1821:
Sexual Selection and the Origins of Human Mating Systems
1272:
502:
within the penis maintaining it as a viable channel for
3726:
2818:. Vol. 2, no. 1. Chicago, IL. pp. 8–12.
2814:. American Urological Association 98th Annual Meeting.
2657:
2586:
1275:
Affective Touch and the Neurophysiology of CT Afferents
61:"Bellend" redirects here. For the English village, see
3825:
3164:
2499:"Prepuce in boys and adolescents: What when, and how?"
2002:
1517:
3947:. Urologic & Cutaneous Press. 1920. pp. 677–
1846:"Penis Frenulum: Location, Function & Conditions"
284:
281:
3628:
Bassert, Joanna M; McCurnin, Dennis M (2013-04-01).
2086:"3 - Anatomy and physiology of genital organs – men"
1808:." Evolution and Human Behavior 24.4 (2003): 277-289
1350:
Renfree, Marilyn; Hugh Tyndale-Biscoe (1987-01-30).
840:. The Y chromosome contains a sex-determining gene (
784:
Development of external genitalia in the male embryo
738:. The skin begins to loosen up significantly during
370:
The glans penis develops as the terminal end of the
275:
3005:
Introduction to Clinical Reproductive Endocrinology
2761:
most foreskins will become retractile by adulthood.
2728:
2257:"Anatomy and Blood Supply of the Urethra and Penis"
2154:"Anatomy and Blood Supply of the Urethra and Penis"
1953:Yang, Claire C.; Bradley, William E. (1999-01-01).
878:(different versions of the same structure). In the
2811:Effects of circumcision on male penile sensitivity
1419:
1170:The glans penis is also divided into two parts in
1154:
1085:'s penis extends about halfway down the shaft and
866:Both the penis and clitoris develop from the same
474:Diagram of the arteries of the penis and its glans
355:, runs on its ventral surface. In men who are not
3910:
3572:. University of Wisconsin-Madison. Archived from
3555:Skurski, D., J. Waterman. 2005. "Xerus inauris",
2928:Oxford Desk Reference: Obstetrics and Gynaecology
1915:"Anatomy, Abdomen and Pelvis, Penis Dorsal Nerve"
1912:
914:and the labioscrotal swellings unite to form the
686:through the circumflex veins that surround them.
319:pleasure. The glans penis is present in the male
299:meaning "acorn") is the bulbous structure at the
4760:
3430:
3397:
2463:
2029:
1415:
1413:
1411:
1409:
1311:
1219:Greenburg, Jerrold S.; Bruess, Clint E. (2016).
906:on the belly aspect of the genital tubercle—the
569:and a triangle or a V-shaped area under it. The
3627:
3403:
2355:
2083:
1921:, Treasure Island (FL): StatPearls Publishing,
1218:
669:, the terminal branches of the dorsal arteries
498:, a sponge-like region that surrounds the male
490:that is moulded on the rounded ends of the two
2009:. Jones & Bartlett Learning. p. 132.
1799:The human penis as a semen displacement device
1252:. St. Martin's Publishing Group. p. 179.
1225:. Jones & Bartlett Learning. p. 267.
1159:The shape of the glans varies among different
4052:
3993:Hyman, Libbie Henrietta (15 September 1992).
3959:
3916:
1761:Encyclopedia of Reproduction (Second Edition)
1547:"Ligamentous structures in human glans penis"
1406:
1345:
1343:
4066:
3600:. Cambridge University Press. 30 June 2012.
3165:Sorokan SK, Finlay JC, Jefferies AL (2015).
2998:
2951:
2263:, Berlin, Heidelberg: Springer, p. 16,
2196:"The blood supply to preputial island flaps"
2160:, Berlin, Heidelberg: Springer, p. 14,
2084:Clement, Pierre; Giuliano, Francois (2015).
1763:, Oxford: Academic Press, pp. 367–375,
1755:Hsu, Geng-Long; Lu, Hsiu-Chen (2018-01-01),
1697:
1631:
1544:
1467:: CS1 maint: multiple names: authors list (
961:Inflammation of the glans penis is known as
852:for the embryo's development into a male.
788:The glans develops as the terminal end of a
661:, which also supplies the foreskin, and the
3892:Australian Mammal Society (December 1978).
3885:
3549:
3501:
3424:
3342:. Cornell University Press. pp. 116–.
3331:
3329:
2490:
2296:
2006:Exploring the Dimensions of Human Sexuality
1952:
1420:W.George, D.Wilson, Fredrick, Jean (1984).
1222:Exploring the Dimensions of Human Sexuality
1067:also occur on the glans of male and female
15:
4059:
4045:
3363:
2537:
2403:The American Journal of Surgical Pathology
1698:Dean, Robert C.; Lue, Tom F. (2005–2011).
1340:
351:. An elastic band of tissue, known as the
16:
3986:
3965:
3898:. Australian Mammal Society. pp. 73–
3190:
3007:. Butterworth-Heinemann. pp. 36–41.
2746:
2691:
2620:
2555:
2514:
2373:
2332:
2314:
1731:
1570:
3660:
3326:
1632:Halata, Z.; Munger, B. L. (1986-04-23).
1307:
1305:
1303:
1301:
925:
779:
712:
653:The glans penis receives blood from the
477:
469:
419:
343:of the penis. At the summit appears the
307:that is the human male's most sensitive
24:This is an accepted version of this page
2664:Canadian Urological Association Journal
2593:Canadian Urological Association Journal
2254:
2151:
2023:
1245:
1185:have a narrow, egg-shaped glans penis.
424:Internal anatomy of human glans penis:
376:embryonic development of the male fetus
14:
4761:
3996:Hyman's Comparative Vertebrate Anatomy
3369:
3238:from the original on 12 September 2017
3225:
3062:
2891:
2838:from the original on February 11, 2023
2193:
1955:"Innervation of the human glans penis"
1817:
1089:except at the tip. In comparison, the
810:development of the reproductive system
776:Development of the reproductive system
4040:
3992:
3966:Griffiths, Mervyn (2 December 2012).
3923:. University of Chicago. pp. 64–
3759:
3498:Hooper and Musser, 1964, table 1
3255:
3253:
2250:
2248:
1948:
1946:
1908:
1906:
1904:
1872:
1870:
1754:
1704:The Urologic Clinics of North America
1627:
1625:
1623:
1621:
1619:
1386:Maternal Child Nursing Care in Canada
1353:Reproductive Physiology of Marsupials
1298:
762:physical changes do occur, including
4019:Bates et al., 2006, pp. 306–307
3483:
3335:
3290:"Glans Insufficiency Syndrome | CSM"
3207:from the original on 30 October 2019
3001:"Chapter 4 - Sexual differentiation"
2557:10.1046/j.1464-410x.1999.0830s1034.x
2445:from the original on 8 November 2022
1486:
1484:
1482:
1480:
1478:
1007:
734:and it continues to be tight during
557:, overhanging a deep retroglandular
486:The glans penis is a body of spongy
3862:Equine Reproduction - Google Boeken
3489:Hooper and Musser, 1964, p. 13
3431:Köhncke, M.; Leonhardt, K. (1986).
3158:
2952:Hake, Laura & O'Connor, Clare.
2464:Kirby R, Carson C, Kirby M (2009).
1883:. World Health Organization. 2007.
48:
3250:
3013:10.1016/B978-0-7506-1171-8.50008-8
2375:10.1002/j.1939-4640.2003.tb03145.x
2245:
2102:10.1016/B978-0-444-63247-0.00003-1
1943:
1901:
1867:
1616:
1434:10.1016/B978-0-407-00366-8.50008-3
1093:is short and spiny, while that of
855:Although the sex of the infant is
49:
4795:
4032:
3944:The Urologic and Cutaneous Review
3471:from the original on 21 June 2010
2538:Cold, C.J.; Taylor, J.R. (1999).
1475:
1130:
1024:list of miscellaneous information
378:. The tubercle is present in the
2871:10.1111/j.1743-6109.2007.00471.x
2415:10.1097/00000478-200307000-00015
1759:, in Skinner, Michael K. (ed.),
1521:Comparative Reproductive Biology
1326:10.1046/j.1464-410x.1998.00669.x
1167:have an undivided glans penis.
1012:
998:
268:
90:
84:Human penile glans (dorsal view)
78:
4769:Mammal male reproductive system
4453:Trabeculae of corpus spongiosum
4443:Trabeculae of corpora cavernosa
4013:
3999:. University of Chicago Press.
3852:
3819:
3786:
3753:
3720:
3687:
3661:Research, Equine (2005-07-01).
3654:
3621:
3588:
3562:
3540:
3492:
3302:
3282:
3271:from the original on 2022-09-25
3226:Koenig JF (22 September 2016).
3219:
3147:from the original on 2022-09-25
3129:
3118:from the original on 2022-09-25
3100:
3083:
3056:
3040:
3029:from the original on 2022-09-22
2992:
2975:
2964:from the original on 2017-08-19
2945:
2918:
2885:
2850:
2801:
2766:
2722:
2708:from the original on 2022-10-28
2651:
2637:from the original on 2022-10-28
2580:
2531:
2457:
2390:
2349:
2297:Gyftopoulos, Kostis I. (2018).
2290:
2261:Urethral Reconstructive Surgery
2234:from the original on 2022-11-09
2187:
2158:Urethral Reconstructive Surgery
2145:
2134:from the original on 2020-10-17
2077:
2066:from the original on 2006-06-20
1996:
1985:from the original on 2021-11-21
1933:from the original on 2023-04-05
1890:from the original on 2015-07-15
1856:from the original on 2022-09-24
1838:
1811:
1791:
1782:
1748:
1691:
1680:from the original on 2022-09-25
1605:from the original on 2022-09-24
1587:
1538:
1450:from the original on 2022-09-22
1279:Springer Science+Business Media
1155:Marsupials, monotremes and bats
934:of the glans penis consists of
648:
586:The glans and the frenulum are
449:Navicular fossa of male urethra
4784:Human male reproductive system
3171:Paediatrics & Child Health
2999:C.L.Lachelin, Gillian (1991).
2090:Handbook of Clinical Neurology
2033:The Fate of the Male Germ Cell
1377:
1356:. Cambridge University Press.
1266:
1239:
1212:
910:closes completely to form the
769:
581:
508:stratified squamous epithelium
262:, commonly referred to as the
13:
1:
3969:The Biology of the Monotremes
2824:10.1016/s1540-0352(11)70011-7
2787:10.1016/j.urology.2004.11.007
2212:10.1016/s0022-5347(17)38584-1
1971:10.1016/S0022-5347(01)62075-5
1426:Fetal Physiology and Medicine
1205:
1050:
812:depending on the exposure to
706:, while also maintaining the
339:and is a continuation of the
3667:. Rowman & Littlefield.
3634:. Elsevier Health Sciences.
3410:. Elsevier Health Sciences.
2042:10.1007/978-1-4615-5913-9_48
1650:10.1016/0006-8993(86)90357-4
1422:"2 - Sexual Differentiation"
993:glans insufficiency syndrome
506:. The glans is covered by a
465:
7:
4584:Internal urethral sphincter
3407:Miller's Anatomy of the Dog
3167:"Newborn male circumcision"
3069:. Delmar Thomson Learning.
2902:10.1007/978-90-481-9446-9_7
1797:Gallup, Gordon G., et al. "
1188:
1135:When erect, the glans of a
689:
10:
4800:
4717:Posterior scrotal arteries
2859:Journal of Sexual Medicine
1824:. OUP Oxford. p. 68.
1757:"Penis Structure—Erection"
1314:British Journal of Urology
1123:is large with a prominent
1100:
1003:
870:that become the glans and
804:during the early weeks of
773:
659:dorsal artery of the penis
347:and at the base forms the
147:Dorsal artery of the penis
60:
53:
4712:Anterior scrotal arteries
4689:Internal spermatic fascia
4674:External spermatic fascia
4651:
4633:External urethral orifice
4614:External sphincter muscle
4589:Internal urethral orifice
4396:
4387:
4316:
4208:
4160:
4092:
4083:
4074:
3895:Australian Mammal Society
3865:. John Wiley & Sons.
3832:. John Wiley & Sons.
3796:The Horse - Google Boeken
2735:Canadian Family Physician
1716:10.1016/j.ucl.2005.08.007
1524:. John Wiley & Sons.
1246:Hodgson, Charles (2015).
757:reasons some men undergo
710:in a moist environment.
680:dorsal veins of the penis
633:typically found in thick
592:dorsal nerve of the penis
452:Tunica albuginea of penis
229:
217:
205:
193:
181:
176:
171:Dorsal nerve of the penis
164:
159:Dorsal veins of the penis
152:
140:
128:
118:
106:
101:
89:
77:
72:
4732:Posterior scrotal nerves
4537:Internal pudendal artery
4068:Male reproductive system
3376:. Rastogi Publications.
2816:Clinical Prostate Cancer
1818:Dixson, Alan F. (2009).
1390:Elsevier Health Sciences
1021:This section contains a
657:through its branch, the
655:internal pudental artery
598:, both divisions of the
31:latest accepted revision
4744:Posterior scrotal veins
4727:Anterior scrotal nerves
4557:Internal pudendal veins
4221:Tubuli seminiferi recti
2269:10.1007/3-540-29385-x_3
2166:10.1007/3-540-29385-x_3
1183:Racey's pipistrelle bat
1105:The glans penis of the
828:that carries either an
545:, through which urine,
492:corpora cavernosa penis
3760:Hedge, Juliet (2004).
3546:Voss, 2003, p. 11
3370:Mathur, Reena (2010).
3063:Sloane, Ethel (2002).
2200:The Journal of Urology
822:sexual differentiation
792:structure, called the
785:
723:
483:
475:
462:
231:Anatomical terminology
3733:. Schattauer Verlag.
1200:Pearly penile papules
1195:Corona of glans penis
1097:is smooth and long.
926:Clinical significance
824:is determined by the
796:, which forms in the
783:
716:
627:Meissner's corpuscles
551:pre-ejaculatory fluid
510:and a dense layer of
481:
473:
446:Meatus of the urethra
423:
4375:Bulbourethral glands
4216:Seminiferous tubules
4152:Blood–testis barrier
3972:. Elsevier Science.
3514:Journal of Mammalogy
3373:Animal Behaviour 3/e
3336:Ewer, R. F. (1973).
3183:10.1093/pch/20.6.311
2516:10.2298/MPNS1208295D
2362:Journal of Andrology
1121:cape ground squirrel
846:transcription factor
623:Merkel nerve endings
96:Glans (ventral view)
4779:Human penis anatomy
4638:Lacunae of Morgagni
4426:Suspensory ligament
4299:Testicular arteries
4175:Spermatocytogenesis
2316:10.4103/UA.UA_25_18
2194:Hinman, F. (1991).
898:. By fusion of the
859:from the moment of
745:In some cases, for
609:, numerous genital
321:reproductive organs
21:Page version status
4684:Cremasteric fascia
4421:Fundiform ligament
4348:Seminal colliculus
4294:Rectovesical pouch
4275:Pampiniform plexus
4253:Superior ligaments
4248:Inferior ligaments
3435:Cryptoprocta ferox
2896:. pp. 75–79.
2503:Medicinski Pregled
2096:. pp. 19–37.
1959:Journal of Urology
1804:2018-02-13 at the
1599:www.britannica.com
1551:Journal of Anatomy
1139:increases by 3 to
1111:Winkelmann's mouse
1079:pars longa glandis
786:
724:
607:free nerve endings
514:equivalent to the
484:
476:
463:
417:as the "bellend".
388:primordial phallus
27:
4756:
4755:
4752:
4751:
4448:Corpus spongiosum
4438:Corpora cavernosa
4383:
4382:
4353:Ejaculatory ducts
4312:
4311:
4185:Spermatidogenesis
4006:978-0-226-87013-7
3979:978-0-323-15331-7
3872:978-0-470-96187-2
3839:978-0-470-39025-2
3806:978-0-7167-1811-6
3773:978-1-59228-487-0
3740:978-3-7945-2485-3
3707:978-1-84076-608-0
3674:978-0-7627-9451-5
3641:978-1-4557-2884-8
3607:978-1-107-00001-8
3557:Mammalian Species
3510:Peromyscus boylii
3445:Mammalian Species
3417:978-0-323-26623-9
3383:978-81-7133-747-7
3349:978-0-8014-8493-3
3265:urologyhealth.org
3228:"Meatal stenosis"
3095:978-0-7668-1142-3
3076:978-0-7668-1142-3
3052:978-1-58890-147-7
3022:978-0-7506-1171-8
2987:978-1-58890-147-7
2938:978-0-19-162087-4
2911:978-90-481-9445-2
2676:10.5489/cuaj.5033
2605:10.5489/cuaj.5033
2544:BJU International
2475:978-1-4398-0807-8
2278:978-3-540-29385-9
2175:978-3-540-29385-9
2111:978-0-444-63247-0
2092:. Vol. 130.
2051:978-0-306-45696-1
2016:978-0-7637-7660-2
1831:978-0-19-156973-9
1770:978-0-12-815145-7
1563:10.1111/joa.12896
1531:978-0-470-39025-2
1504:978-1-58890-387-7
1443:978-0-407-00366-8
1399:978-0-323-75920-5
1363:978-0-521-33792-2
1288:978-1-4939-6418-5
1259:978-1-46689-043-5
1232:978-1-28408-154-1
1147:opens within the
1116:Thomasomys ucucha
1081:. The glans of a
1061:urinate backwards
1048:
1047:
844:) that encodes a
721:of a glans penis
684:corpora cavernosa
590:by the bilateral
512:connective tissue
496:corpus spongiosum
455:Corpus spongiosum
431:Corpus cavernosum
341:corpus spongiosum
337:corpora cavernosa
245:
244:
240:
135:Urogenital system
18:
4791:
4522:Arteries of bulb
4509:Tunica albuginea
4394:
4393:
4326:Seminal vesicles
4318:Accessory glands
4304:Testicular veins
4115:Tunica vasculosa
4110:Tunica albuginea
4105:Tunica vaginalis
4090:
4089:
4081:
4080:
4061:
4054:
4047:
4038:
4037:
4020:
4017:
4011:
4010:
3990:
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3141:Cleveland Clinic
3133:
3127:
3126:
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3112:Cleveland Clinic
3104:
3098:
3087:
3081:
3080:
3066:Biology of Women
3060:
3054:
3044:
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3037:
3035:
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2979:
2973:
2972:
2970:
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2916:
2915:
2894:Genital Autonomy
2889:
2883:
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2584:
2578:
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2529:
2528:
2518:
2509:(7–8): 295–300.
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2206:(6): 1232–1235.
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1850:Cleveland Clinic
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1016:
1015:
1008:
908:urogenital sinus
900:urogenital folds
794:genital tubercle
631:mechanoreceptors
529:The increase of
520:tunica albuginea
372:genital tubercle
291:
290:
287:
286:
283:
280:
277:
274:
237:edit on Wikidata
234:
113:Genital tubercle
94:
82:
70:
69:
51:End of the penis
39:6 September 2024
4799:
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4794:
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4792:
4790:
4789:
4788:
4759:
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4757:
4748:
4647:
4643:Urethral glands
4621:Navicular fossa
4572:Pudendal nerves
4527:Dorsal arteries
4379:
4308:
4282:Vasa deferentia
4270:Spermatic cords
4204:
4162:Spermatogenesis
4156:
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3766:. Lyons Press.
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3745:
3743:
3741:
3725:
3721:
3712:
3710:
3708:
3692:
3688:
3679:
3677:
3675:
3659:
3655:
3646:
3644:
3642:
3626:
3622:
3612:
3610:
3608:
3594:
3593:
3589:
3579:
3577:
3568:
3567:
3563:
3554:
3550:
3545:
3541:
3526:10.2307/1381595
3506:
3502:
3497:
3493:
3488:
3484:
3474:
3472:
3468:
3454:10.2307/3503919
3439:
3429:
3425:
3418:
3402:
3398:
3388:
3386:
3384:
3368:
3364:
3354:
3352:
3350:
3334:
3327:
3318:
3316:
3308:
3307:
3303:
3294:
3292:
3288:
3287:
3283:
3274:
3272:
3259:
3258:
3251:
3241:
3239:
3224:
3220:
3210:
3208:
3163:
3159:
3150:
3148:
3135:
3134:
3130:
3121:
3119:
3106:
3105:
3101:
3088:
3084:
3077:
3061:
3057:
3045:
3041:
3032:
3030:
3023:
2997:
2993:
2980:
2976:
2967:
2965:
2950:
2946:
2939:
2923:
2919:
2912:
2890:
2886:
2855:
2851:
2841:
2839:
2806:
2802:
2771:
2767:
2727:
2723:
2711:
2709:
2656:
2652:
2640:
2638:
2585:
2581:
2536:
2532:
2495:
2491:
2476:
2462:
2458:
2448:
2446:
2395:
2391:
2354:
2350:
2295:
2291:
2283:
2281:
2279:
2253:
2246:
2237:
2235:
2192:
2188:
2180:
2178:
2176:
2150:
2146:
2137:
2135:
2112:
2082:
2078:
2069:
2067:
2052:
2028:
2024:
2017:
2001:
1997:
1988:
1986:
1951:
1944:
1936:
1934:
1911:
1902:
1893:
1891:
1887:
1880:
1876:
1875:
1868:
1859:
1857:
1844:
1843:
1839:
1832:
1816:
1812:
1806:Wayback Machine
1796:
1792:
1787:
1783:
1775:
1773:
1771:
1753:
1749:
1696:
1692:
1683:
1681:
1630:
1617:
1608:
1606:
1593:
1592:
1588:
1543:
1539:
1532:
1516:
1512:
1505:
1489:
1476:
1460:
1459:
1453:
1451:
1444:
1418:
1407:
1400:
1392:. p. 501.
1382:
1378:
1368:
1366:
1364:
1348:
1341:
1310:
1299:
1289:
1281:. p. 305.
1271:
1267:
1260:
1244:
1240:
1233:
1217:
1213:
1208:
1191:
1157:
1140:
1133:
1103:
1091:glans of felids
1053:
1044:
1038:
1035:
1028:
1017:
1013:
1006:
1001:
986:meatal stenosis
948:contact allergy
938:tissue. Birley
928:
904:urethral groove
892:masculinization
778:
772:
722:
692:
651:
584:
488:erectile tissue
468:
461:
437:Corona of glans
411:British English
327:and most other
271:
267:
241:
97:
85:
66:
59:
52:
47:
46:
45:
44:
43:
42:
26:
12:
11:
5:
4797:
4787:
4786:
4781:
4776:
4771:
4754:
4753:
4750:
4749:
4747:
4746:
4741:
4740:
4739:
4737:Perineal nerve
4734:
4729:
4721:
4720:
4719:
4714:
4706:
4701:
4698:
4693:
4692:
4691:
4686:
4681:
4676:
4671:
4666:
4657:
4655:
4649:
4648:
4646:
4645:
4640:
4635:
4630:
4629:
4628:
4618:
4617:
4616:
4611:
4606:
4601:
4596:
4591:
4586:
4576:
4575:
4574:
4569:
4561:
4560:
4559:
4554:
4549:
4541:
4540:
4539:
4534:
4529:
4524:
4516:
4511:
4506:
4505:
4504:
4499:
4489:
4484:
4483:
4482:
4480:Septum glandis
4477:
4472:
4467:
4457:
4456:
4455:
4450:
4445:
4440:
4430:
4429:
4428:
4423:
4418:
4413:
4402:
4400:
4391:
4385:
4384:
4381:
4380:
4378:
4377:
4372:
4371:
4370:
4365:
4360:
4355:
4350:
4345:
4343:Urethral crest
4335:
4334:
4333:
4331:Excretory duct
4322:
4320:
4314:
4313:
4310:
4309:
4307:
4306:
4301:
4296:
4291:
4290:
4289:
4279:
4278:
4277:
4267:
4262:
4261:
4260:
4255:
4250:
4245:
4235:
4234:
4233:
4231:Efferent ducts
4228:
4223:
4212:
4210:
4206:
4205:
4203:
4202:
4197:
4195:Spermiogenesis
4192:
4187:
4182:
4177:
4172:
4170:Spermatogonium
4166:
4164:
4158:
4157:
4155:
4154:
4149:
4144:
4139:
4134:
4129:
4124:
4119:
4118:
4117:
4112:
4107:
4098:
4096:
4087:
4078:
4072:
4071:
4064:
4063:
4056:
4049:
4041:
4034:
4033:External links
4031:
4022:
4021:
4012:
4005:
3985:
3978:
3958:
3934:
3909:
3884:
3871:
3851:
3838:
3818:
3805:
3785:
3772:
3752:
3739:
3719:
3706:
3686:
3673:
3653:
3640:
3620:
3606:
3587:
3561:
3548:
3539:
3520:(3): 595–615.
3500:
3491:
3482:
3423:
3416:
3396:
3382:
3362:
3348:
3339:The Carnivores
3325:
3301:
3281:
3249:
3218:
3177:(6): 311–320.
3157:
3128:
3099:
3082:
3075:
3055:
3039:
3021:
2991:
2974:
2944:
2937:
2931:. OUP Oxford.
2917:
2910:
2884:
2865:(3): 667–674.
2849:
2800:
2781:(4): 773–777.
2765:
2741:(3): 445–448.
2721:
2670:(2): E76–E99.
2650:
2599:(2): E76–E99.
2579:
2530:
2489:
2474:
2456:
2409:(7): 994–998.
2389:
2368:(6): 921–927.
2348:
2309:(4): 354–357.
2303:Urology Annals
2289:
2277:
2244:
2186:
2174:
2144:
2110:
2076:
2050:
2022:
2015:
1995:
1942:
1900:
1866:
1837:
1830:
1810:
1790:
1781:
1769:
1747:
1690:
1644:(2): 205–230.
1638:Brain Research
1615:
1586:
1537:
1530:
1510:
1503:
1474:
1442:
1405:
1398:
1376:
1362:
1339:
1297:
1287:
1265:
1258:
1238:
1231:
1210:
1209:
1207:
1204:
1203:
1202:
1197:
1190:
1187:
1156:
1153:
1149:urethral fossa
1132:
1131:Perissodactyls
1129:
1107:marsh rice rat
1102:
1099:
1074:bulbus glandis
1069:spotted hyenas
1052:
1049:
1046:
1045:
1020:
1018:
1011:
1005:
1002:
1000:
997:
927:
924:
912:spongy urethra
820:. In mammals,
800:regardless of
774:Main article:
771:
768:
766:of the glans.
764:keratinization
736:preadolescence
719:Keratinization
717:
704:urinary meatus
691:
688:
650:
647:
639:dermal papilla
600:pudendal nerve
596:perineal nerve
583:
580:
573:is the highly
567:septum glandis
563:coronal sulcus
555:corona glandis
543:urinary meatus
524:mucosal tissue
467:
464:
460:
459:
456:
453:
450:
447:
444:
441:
438:
435:
434:Coronal sulcus
432:
429:
425:
404:clitoral glans
390:. Exposure to
349:corona glandis
345:urinary meatus
333:mucosal tissue
309:erogenous zone
243:
242:
233:
227:
226:
221:
215:
214:
209:
203:
202:
197:
191:
190:
185:
179:
178:
174:
173:
168:
162:
161:
156:
150:
149:
144:
138:
137:
132:
126:
125:
120:
116:
115:
110:
104:
103:
99:
98:
95:
87:
86:
83:
75:
74:
50:
28:
22:
19:
17:
9:
6:
4:
3:
2:
4796:
4785:
4782:
4780:
4777:
4775:
4772:
4770:
4767:
4766:
4764:
4745:
4742:
4738:
4735:
4733:
4730:
4728:
4725:
4724:
4722:
4718:
4715:
4713:
4710:
4709:
4707:
4705:
4702:
4699:
4697:
4694:
4690:
4687:
4685:
4682:
4680:
4677:
4675:
4672:
4670:
4669:Dartos fascia
4667:
4665:
4662:
4661:
4659:
4658:
4656:
4654:
4650:
4644:
4641:
4639:
4636:
4634:
4631:
4627:
4624:
4623:
4622:
4619:
4615:
4612:
4610:
4607:
4605:
4602:
4600:
4597:
4595:
4594:Pre-prostatic
4592:
4590:
4587:
4585:
4582:
4581:
4580:
4577:
4573:
4570:
4568:
4567:Dorsal nerves
4565:
4564:
4562:
4558:
4555:
4553:
4550:
4548:
4545:
4544:
4542:
4538:
4535:
4533:
4532:Deep arteries
4530:
4528:
4525:
4523:
4520:
4519:
4517:
4515:
4512:
4510:
4507:
4503:
4500:
4498:
4495:
4494:
4493:
4490:
4488:
4485:
4481:
4478:
4476:
4473:
4471:
4468:
4466:
4463:
4462:
4461:
4458:
4454:
4451:
4449:
4446:
4444:
4441:
4439:
4436:
4435:
4434:
4431:
4427:
4424:
4422:
4419:
4417:
4414:
4412:
4409:
4408:
4407:
4404:
4403:
4401:
4399:
4395:
4392:
4390:
4386:
4376:
4373:
4369:
4366:
4364:
4361:
4359:
4356:
4354:
4351:
4349:
4346:
4344:
4341:
4340:
4339:
4336:
4332:
4329:
4328:
4327:
4324:
4323:
4321:
4319:
4315:
4305:
4302:
4300:
4297:
4295:
4292:
4288:
4285:
4284:
4283:
4280:
4276:
4273:
4272:
4271:
4268:
4266:
4265:Paradidymides
4263:
4259:
4256:
4254:
4251:
4249:
4246:
4244:
4241:
4240:
4239:
4236:
4232:
4229:
4227:
4224:
4222:
4219:
4218:
4217:
4214:
4213:
4211:
4207:
4201:
4198:
4196:
4193:
4191:
4188:
4186:
4183:
4181:
4178:
4176:
4173:
4171:
4168:
4167:
4165:
4163:
4159:
4153:
4150:
4148:
4147:Sertoli cells
4145:
4143:
4140:
4138:
4135:
4133:
4130:
4128:
4125:
4123:
4120:
4116:
4113:
4111:
4108:
4106:
4103:
4102:
4100:
4099:
4097:
4095:
4091:
4088:
4086:
4085:Seminal tract
4082:
4079:
4077:
4073:
4069:
4062:
4057:
4055:
4050:
4048:
4043:
4042:
4039:
4030:
4027:
4016:
4008:
4002:
3998:
3997:
3989:
3981:
3975:
3971:
3970:
3962:
3946:
3945:
3938:
3922:
3921:
3913:
3897:
3896:
3888:
3874:
3868:
3864:
3863:
3855:
3841:
3835:
3831:
3830:
3822:
3808:
3802:
3799:. Macmillan.
3798:
3797:
3789:
3775:
3769:
3765:
3764:
3756:
3742:
3736:
3732:
3731:
3723:
3709:
3703:
3700:. CRC Press.
3699:
3698:
3690:
3676:
3670:
3666:
3665:
3657:
3643:
3637:
3633:
3632:
3624:
3609:
3603:
3599:
3598:
3591:
3576:on 2007-07-16
3575:
3571:
3565:
3558:
3552:
3543:
3535:
3531:
3527:
3523:
3519:
3515:
3511:
3504:
3495:
3486:
3467:
3463:
3459:
3455:
3451:
3447:
3446:
3438:
3436:
3427:
3419:
3413:
3409:
3408:
3400:
3385:
3379:
3375:
3374:
3366:
3351:
3345:
3341:
3340:
3332:
3330:
3315:
3314:www.sdsm.info
3311:
3305:
3291:
3285:
3270:
3266:
3262:
3256:
3254:
3237:
3233:
3229:
3222:
3206:
3202:
3198:
3193:
3188:
3184:
3180:
3176:
3172:
3168:
3161:
3146:
3142:
3138:
3132:
3117:
3113:
3109:
3103:
3096:
3092:
3086:
3078:
3072:
3068:
3067:
3059:
3053:
3049:
3043:
3028:
3024:
3018:
3014:
3010:
3006:
3002:
2995:
2988:
2984:
2978:
2963:
2959:
2955:
2948:
2940:
2934:
2930:
2929:
2921:
2913:
2907:
2903:
2899:
2895:
2888:
2880:
2876:
2872:
2868:
2864:
2860:
2853:
2837:
2833:
2829:
2825:
2821:
2817:
2813:
2812:
2804:
2796:
2792:
2788:
2784:
2780:
2776:
2769:
2762:
2758:
2754:
2749:
2744:
2740:
2736:
2732:
2725:
2718:
2707:
2703:
2699:
2694:
2689:
2685:
2681:
2677:
2673:
2669:
2665:
2661:
2654:
2647:
2636:
2632:
2628:
2623:
2618:
2614:
2610:
2606:
2602:
2598:
2594:
2590:
2583:
2575:
2571:
2567:
2563:
2558:
2553:
2550:(S1): 34–44.
2549:
2545:
2541:
2540:"The prepuce"
2534:
2526:
2522:
2517:
2512:
2508:
2504:
2500:
2493:
2485:
2481:
2477:
2471:
2467:
2460:
2444:
2440:
2436:
2432:
2428:
2424:
2420:
2416:
2412:
2408:
2404:
2400:
2393:
2385:
2381:
2376:
2371:
2367:
2363:
2359:
2352:
2344:
2340:
2335:
2330:
2326:
2322:
2317:
2312:
2308:
2304:
2300:
2293:
2280:
2274:
2270:
2266:
2262:
2258:
2251:
2249:
2233:
2229:
2225:
2221:
2217:
2213:
2209:
2205:
2201:
2197:
2190:
2177:
2171:
2167:
2163:
2159:
2155:
2148:
2133:
2129:
2125:
2121:
2117:
2113:
2107:
2103:
2099:
2095:
2091:
2087:
2080:
2065:
2061:
2057:
2053:
2047:
2043:
2039:
2035:
2034:
2026:
2018:
2012:
2008:
2007:
1999:
1984:
1980:
1976:
1972:
1968:
1965:(1): 97–102.
1964:
1960:
1956:
1949:
1947:
1932:
1928:
1924:
1920:
1916:
1909:
1907:
1905:
1886:
1879:
1873:
1871:
1855:
1851:
1847:
1841:
1833:
1827:
1823:
1822:
1814:
1807:
1803:
1800:
1794:
1785:
1772:
1766:
1762:
1758:
1751:
1743:
1739:
1734:
1729:
1725:
1721:
1717:
1713:
1709:
1705:
1701:
1694:
1679:
1675:
1671:
1667:
1663:
1659:
1655:
1651:
1647:
1643:
1639:
1635:
1628:
1626:
1624:
1622:
1620:
1604:
1600:
1596:
1590:
1582:
1578:
1573:
1568:
1564:
1560:
1556:
1552:
1548:
1541:
1533:
1527:
1523:
1522:
1514:
1506:
1500:
1496:
1495:
1487:
1485:
1483:
1481:
1479:
1470:
1464:
1449:
1445:
1439:
1435:
1431:
1427:
1423:
1416:
1414:
1412:
1410:
1401:
1395:
1391:
1387:
1380:
1365:
1359:
1355:
1354:
1346:
1344:
1335:
1331:
1327:
1323:
1320:(1): 109–13.
1319:
1315:
1308:
1306:
1304:
1302:
1294:
1290:
1284:
1280:
1276:
1269:
1261:
1255:
1251:
1250:
1242:
1234:
1228:
1224:
1223:
1215:
1211:
1201:
1198:
1196:
1193:
1192:
1186:
1184:
1179:
1177:
1173:
1168:
1166:
1162:
1152:
1150:
1146:
1138:
1137:horse's penis
1128:
1126:
1122:
1118:
1117:
1112:
1108:
1098:
1096:
1092:
1088:
1084:
1080:
1076:
1075:
1070:
1066:
1065:Penile spines
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