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Heaviside's dolphin

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644:) are found throughout the range and occasionally both dolphin species are sighted in mixed groups. Where both species overlap in prey selection, Heaviside's take larger prey items, potentially because they are outcompeted by the larger dusky dolphins for their preferred, smaller sized prey. In central Namibia (especially Walvis Bay) Heaviside's dolphins overlap with a small populations of fewer than 100 common bottlenose dolphins REF. The bottlenose dolphin population use only water less than 15m depth while in this area Heaviside's dolphins are almost always encounteres in water deeper than 20 m (66 ft), suggesting some form of competitive exclusion. 31: 653:
outside of the detectable frequencies of killer whales. Although NBHF clicks are limited in acoustic range, they have a better resolution for small targets and are thought to provide a foraging advantage in the often cluttered, nearshore environment in which these species occur. Heaviside's also produce a second click type, of lower frequency and broader bandwidth, that is within the hearing range of killer whales. These calls are produced most frequently in groups engaging in social behaviour. It is likely that the dolphins use these calls when socialising away from predator threat and switch to high frequency clicks when foraging and travelling.
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coastal waves. Iconic vertical leaps clear the water before re-entering headfirst with almost no splash. Heaviside's dolphins use echolocation to find and capture prey. Mating typically occurs in social groups of 3–7 individuals which remain in a small area exhibiting extensive rolling, touching and position changes with frequent leaps by one of pairs of animals which potentially serve a competitive function.
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170 km into Angola but well south of the defined northern boundary of the Benguela Ecosystem. The northern boundary of the Benguela current shifts north and south seasonally and as Heaviside's dolphins appear closely linked to its cool waters, their northern range limit may shift along with water conditions. Systematic surveys have dedicated effort to describing the distribution in southern
487:, the real southern limit beyond which sightings are extremely rare is Hout Bay, some 40 km (25 mi) to the north (a considerable distance for a species which shows very high site fidelity to quite small spatial scales when nearshore). The species occurs more or less continuously for 2,500 km (1,600 mi) to the north of this along the South African coast, through 246: 728:
behaviour in Walvis Bay includes a reduction in resting behaviour and an increase in socialising behaviour in the presence of tour boats (indurkyhua). One Heaviside's dolphin was documented in 2010 with evidence of a propeller strike along its flank. As individuals have small home ranges, they may be vulnerable to localised threats.
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most bays throughout South Africa and Namibia, but they are rarely seen in the protected shallows of these bays. Outside of bays, they show relatively high densities along exposed sandy beaches, but these may be a secondary choice after a preference for areas where there is a high abundance of their main prey item; juvenile hake (
740:(IUCN) listed the Heaviside's as 'Data Deficient' however, as of 2017 the status was changed to 'Near Threatened', owing to improved knowledge on the species from multiple studies. Despite this, the overall population trend remains unknown, and there are many aspects of the species biology that remain to be studied. 440:
Information on reproduction is limited for Heaviside's dolphins, however they are thought to be comparable to Hector's and Commerson's dolphins. Females and males reach sexual maturity approximately between 5–9 years. Mating is thought to occur year-round, however individual females may only produce
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are 508 and 494 respectively. A visual and acoustic line-transect ship survey estimated an average of 1594 dolphins in the Namibian Islands' Marine Protected Area (NIMPA), which spans 400 km (250 mi) of coastline along southern Namibia (REF Martin et al. 2020). Quantification of abundance
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Heaviside's dolphins typically remain nearshore in the mornings where they typically socialise and rest. When nearshore their distribution patterns are remarkably predictable within and between years, with the animals showing highly consistent use of aggregation sites at the exposed western tips of
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Heaviside's dolphins are energetic and social animals, especially when nearshore in the mornings. Behaviour when offshore tends to be less playful and may include a rest phase when moving offshore to feed. They are attracted to boats and frequently bow-ride. Individuals can also be seen surfing in
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move offshore in the afternoon to feed on prey rising vertically to the surface at night. Movement inshore to rest and socialise occurs in the morning. However, the pattern is different in Luderitz and Walvis Bay, Namibia where the movement is less pronounced and dolphins appear to stay inshore
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Heaviside's are small and stocky with adults reaching a maximum length and weight of 1.7 m (5.6 ft) and 75 kg (165 lb) respectively. The dolphin has a distinct black, grey and white body pattern, and is not easily confused with any other species in its range. The head is cone
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As is the case with all species in the genus, Heaviside's dolphins produce narrow-band high-frequency (NBHF) echolocation clicks (centred around 125–130 kHz), and do not whistle. This adaptation is theorised to allow acoustic crypsis from eavesdropping predators, as the sounds produced are
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Heaviside's dolphins may be exposed to increase in the marine eco-tourism business in Namibia, which has grown without regulation in Walvis Bay (Leeney, 2014). Negative effects have been demonstrated for other coastal cetacean species, for example the impact of tour boats on bottlenose dolphin
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warm-water currents at both its northern and southern limits. Consequently, small increases in water temperature could result in all shelf waters in this area becoming too warm for both species, and populations are expected to decline rather than shift poleward due to a lack of shelf habitat.
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Heaviside's dolphins are listed amongst the cetacean species most vulnerable to climate change as they are limited to a distribution range that includes both suitable shelf habitat and cool water temperatures (Best, 2007). The Benguela Current is the only eastern-boundary current bordered by
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where the northern boundary for the species remains poorly defined. Several dolphins have been sighted or accidentally caught by fishing vessels north of the Angola–Namibia border, but no sightings were reported during a series of coastal scientific surveys at Tombua which is approximately
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There has been limited research into Heaviside's diving behaviour, however a study of two dolphins fitted with satellite tags was undertaken in South Africa in 1997. The maximum dive depth recorded was 147 meters; however, the majority of dives were less than 50 meters. Dive duration were
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Recent genetic research has demonstrated evidence of population structure across the range, indicating two metapopulations (north and south) with limited genetic exchange. This pattern of fragmentation is a common feature amongst the other three species in the genus
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Heaviside's dolphins are exposed to several poorly quantified and rapidly changing human threats including fisheries bycatch related mortality and illegal directed catch (Elwen and Gopal, 2018; Alfaro-Shigueto et al., 2019). Recently developed mid water trawls for
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Kyhn, L. A.; Jensen, F. H.; Beedholm, K.; Tougaard, J.; Hansen, M.; Madsen, P. T. (2010). "Echolocation in sympatric Peale's dolphins (Lagenorhynchus australis) and Commerson's dolphins (Cephalorhynchus commersonii) producing narrow-band high-frequency clicks".
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Heaviside's dolphins are exposed to a variety of threats given their limited range in coastal shallow waters which are subject to a range of anthropogenic activities. Directed catch has occurred historically, with meat being used for human consumption.
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Heaviside's have small home ranges of 50–80 km (31–50 mi) as measured using satellite telemetry over 2–3 months and photographic resightings over up to 3 years. Some individuals have been resighted at the same location for up to 10 years.
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Typically occurs in small groups of 2–3, but numbers of 1–10 are frequent and large aggregations of ~100 individuals or more are known to form in high density areas. Nursery groups (exclusively females and calves) are not formed in this species.
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Hamner, Rebecca M.; Pichler, Franz B.; Heimeier, Dorothea; Constantine, Rochelle; Baker, C. Scott (2012). "Genetic differentiation and limited gene flow among fragmented populations of New Zealand endemic Hector's and Maui's dolphins".
323:, who was known for his own biological collections at the time. "Heaviside's dolphin" is the recognised common name, though amongst others, "Haviside's dolphin" and "Benguela dolphin" are also used, the latter especially in Namibia. 1490:
Elwen, Simon H.; Best, Peter B.; Reeb, Desray; Thornton, Meredith (2009). "Diurnal Movements and Behaviour of Heaviside's Dolphins,Cephalorhynchus heavisidii, with some Comparative Data for Dusky Dolphins,Lagenothynchus obscutus".
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Elwen, S.H.; Thornton, M.; Reeb, D.; Best, P.B. (2010). "Near-shore distribution of Heaviside's (Cephalorhynchus heavisidii) and dusky dolphins (Lagenorhynchus obscurus) at the southern limit of their range in South Africa".
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Davis, RW; David, JHM; Meÿer, MA; Sekiguchi, K; Best, PB; Dassis, M; Rodríguez, DH (2014). "Home range and diving behaviour of Heaviside's dolphins monitored by satellite off the west coast of South Africa".
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is minimal, however variation in the shape of the white patch covering the genital slit is distinct between genders. In males, the patch ends in a point, but in females widens out to cover the mammary slits.
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Martin, Morgan J.; Elwen, Simon H.; Kassanjee, Reshma; Gridley, Tess (2019). "To buzz or burst-pulse? The functional role of Heaviside's dolphin, Cephalorhynchus heavisidii, rapidly pulsed signals".
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Morisaka, Tadamichi; Karczmarski, Leszek; Akamatsu, Tomonari; Sakai, Mai; Dawson, Steve; Thornton, Meredith (2011). "Echolocation signals of Heaviside's dolphins (Cephalorhynchus heavisidii)".
912:"A conservation assessment of Cephalorhynchus heavisidii. In Child MF, Roxburgh L, Do Linh San E, Raimondo D, Davies-Mostert HT. The Red List of Mammals of South Africa, Swaziland and Lesotho" 1689:
Heinrich, Sonja; Elwen, Simon; Bräger, Stefan (2010). "Patterns of sympatry in Lagenorhynchus and Cephalorhynchus: dolphins in different habitats". In Würsig, Bernd; Wursig, Melany (eds.).
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Gopal, K; Karczmarski, L; Tolley, K.A (2019). "Patterns of geographic variation between mitochondrial and nuclear markers in Heaviside's (Benguela) dolphins (Cephalorhynchus heavisidii)".
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Elwen, Simon H.; Reeb, Desray; Thornton, Meredith; Best, Peter B. (2009). "A population estimate of Heaviside's dolphins,Cephalorhynchus heavisidii, at the southern end of their range".
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Leeney, R. H; Carslake, D; Elwen, S. H (2011). "Using static acoustic monitoring to describe echolocation behaviour of Heaviside's dolphins (Cephalorhynchus heavisidii) in Namibia".
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Morias, Miguel (2012). "Marine mammal sightings off the Angolan coast recorded from the R/V Dr. Fridtjof Nansenin August 2004 and July 2005". In Van Waerebeek, Koen (ed.).
545:) in adjacent offshore waters. Most commonly sighted within sea surface temperatures of 9 to 15 °C (48 to 59 °F) and depths less than 100 metres (330 ft). 999:
Pichler, F; D. Robineau, R; Goodall, M; Meyer, M; Olivarria, C; Baker, C (2001). "Origin and radiation of Southern Hemisphere coastal dolphins (genus Cephalorhynchus)".
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The genus name "Cephalorhynchus" comes from the Greek kephale for 'head' and rhynchos for 'beak'. For the species name "heavisidii" see the above description.
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Best, Peter B.; Abernethy, R. Blake (1994). "Heaviside's dolphin, Cephalorhynchus heavisidii (Gray, 1828)". In Ridgway, Sam H.; Harrison, Richard (eds.).
520:, South Africa. Sightings are common from land and there are several dolphin watching tour companies by which Heaviside's dolphins can be seen by boat. 4120: 4345: 737: 4198: 4159: 911: 589:) are also consumed especially in Namibia. Foraging occurs mostly at the seabed, in shallow depths. Feeding nearshore is rarely observed. 483:
The species is strongly associated with the cool waters of the Benguela Ecosystem. Although the southern limit of the range is defined as
751:. The Memorandum of Understanding was established in 2008 and aims to protect these species at a national, regional and global level. 1433:"Using static acoustic monitoring to describe echolocation behaviour of heaviside's dolphins (Cephalorhynchus heavisidii) in Namibia" 4375: 4355: 4317: 4094: 423:. The underbelly is white, with bands that extend onto the lower rear of the body. Small white patches are located just behind the 2005:"Global climate change, range changes and potential implications for the conservation of marine cetaceans: a review and synthesis" 1779:"Predation by killer whales (Orcinus orca) and the evolution of whistle loss and narrow-band high frequency clicks in odontocetes" 4385: 4133: 1432: 1576:"Range and Movements of Female Heaviside's Dolphins (Cephalorhynchus heavisidii), as Determined by Satellite-Linked Telemetry" 4211: 4177: 1700: 1180: 950: 885: 4138: 749:
Memorandum of Understanding Concerning the Conservation of the Manatee and Small Cetaceans of Western Africa and Macaronesia
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Weir, C. R (2019). "The Cetaceans (Whales and Dolphins) of Angola". In Huntley, B; Russo, V; Lages, F; Ferrand, N (eds.).
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shaped with a blunt beak. The dorsal fin is triangular in shape and centred in the middle of the back. The head and
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Elwen, Simon; Meÿer, Michael A.; Best, Peter B.; Kotze, P. G H.; Thornton, Meredith; Swanson, Stephan (2006).
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Best, P. B (1988). "The external appearance of Heaviside's dolphin, Cephalorhynchus heavisidii (Gray, 1828)".
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Dawson, S (26 February 2009). "Cephalorhynchus dolphins". In Würsig, B; Thewissen, J.G.M; Kovacs, K.M (eds.).
4185: 3251: 2976: 2967: 374: 673:, South Africa represents the southernmost populations in the species range. Local population estimates for 419:
and dorsal cape are a dark grey to almost black with a band that extends forward from the dorsal fin to the
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time is unknown. Maximum known lifespan is based on the oldest recorded individual at 26 years old.
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predominantly less than 2 minutes with most dives between 0 and 1 minutes (Davis et al. 2014).
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Early in the 19th century, a specimen was caught off the Cape of Good Hope and brought to the
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Best, P; Ros, G.J.B (1977). Exploitation of small cetaceans of the coast of Southern Africa.
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Payne, A. I. L; Brink, K. H; Mann, K. H; Hillborn, R (eds.) Benguela Trophic Functioning.
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South African National Biodiversity Institute and Endangered Wildlife Trust, South Africa
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estimate currently exists, however a population estimate of 6,345 for the region between
662: 529: 359: 351: 62: 3870: 3335: 3326: 3093: 2902: 2893: 2878: 2869: 2729: 2663: 2654: 2598: 2589: 2292: 1972: 1914: 1879: 1816: 1759: 1671: 1516: 1363: 1319: 1186: 1152:(CMS Technical Series No. 26 ed.). Bonn, Germany: CMS Secretariat. pp. 26–30. 1098: 1024: 706: 420: 109: 4224: 4055: 3736: 3418: 3409: 2821: 2738: 2449: 2440: 2026: 1964: 1919: 1901: 1860: 1852: 1808: 1800: 1795: 1778: 1751: 1743: 1696: 1663: 1597: 1508: 1472: 1413: 1399: 1355: 1311: 1230: 1190: 1176: 1123: 1102: 1090: 1016: 1012: 976: 946: 881: 428: 2056: 1976: 1820: 1675: 1520: 1367: 1323: 1028: 777: 4308: 4060: 3671: 3593: 3584: 3574: 3293: 3234: 3119: 2950: 2346: 2328: 2319: 2074: 2016: 1954: 1946: 1909: 1891: 1844: 1790: 1763: 1733: 1725: 1653: 1645: 1587: 1551: 1543: 1500: 1462: 1403: 1395: 1347: 1301: 1293: 1266: 1220: 1168: 1080: 1076: 1072: 1008: 795: 316: 235: 1950: 3054: 3036: 3027: 2917: 2532: 2514: 2496: 2364: 1690: 1649: 1172: 1117: 970: 940: 875: 370: 335: 274: 196: 186: 77: 4268: 4190: 4164: 2935: 2702: 2574: 2565: 2412: 2403: 2135: 1263:
Spatial and temporal patterns of habitat use of Heaviside's dolphins in Namibia
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during the night, which is likely associated with foraging on different prey.
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Martin, Morgan J.; Gridley, Tess; Elwen, Simon H.; Jensen, Frants H. (2018).
1856: 1804: 1747: 1667: 1601: 1547: 1512: 1476: 1417: 1359: 1234: 1094: 786: 637: 598: 517: 470:) are known predators and there is evidence of shark attack from body scars. 424: 382: 82: 48: 1297: 3611: 3174: 3165: 3155: 2283: 2163: 2147: 1923: 1896: 1864: 1812: 1755: 1315: 1020: 822: 670: 602: 497: 463: 416: 4250: 4107: 4001: 3372: 2613: 2212: 2141: 2066: 1504: 378: 166: 1738: 1617:
Heavy metal analysis in Heaviside's dolphins (Cephalorhyncus heavisidii)
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Spicilegia Zoologica: Original Figures and Short Systematic Descriptions
678: 505: 385:, is thought to have led to the subsequent speciation within the genus. 39: 4322: 4099: 2388: 2379: 2337: 2178: 1848: 674: 570: 501: 484: 412: 1959: 1729: 1658: 1556: 1408: 1306: 1270: 1207:
Findlay, K. P.; Best, P. B.; Ross, G. J. B.; Cockcroft, V. G. (1992).
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Convention on the Conservation of Migratory Species of Wild Animals
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and a single white patch extends between these fins on the chest.
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are coloured light grey with darker patches around the eye. The
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Conserving cetaceans and manatees in the western African region
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The three other species in the genus Cephalorhynchus are the
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and current research efforts focus on local populations in
1833: 1122:. Vol. 5 The First Book of Dolphins. Academic Press. 2479: 358:). All are located in cool temperate shelf waters in the 1877: 1634: 1265:(M.Sc). Pretoria, South Africa: University of Pretoria. 1206: 2161: 1884:
Proceedings of the Royal Society B: Biological Sciences
1709: 1619:(Thesis). Pokfulam, Hong Kong: University of Hong Kong. 1692:
The Dusky Dolphin: Master Acrobat Off Different Shores
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Sekiguchi, K.; Klages, N. T. W.; Best, P. B. (1992).
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Whales and Dolphins of the Southern African Subregion
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dolphins originated from a single common ancestor in
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Heaviside's dolphin is listed on Appendix II of the
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Cambridge University Press. p. 388. 636:Whilst typically found further from shore, 2096: 2089: 2075: 1493:South African Journal of Wildlife Research 942:Handbook of Whales, Dolphins and Porpoises 938: 869: 867: 865: 863: 462:Levels of predation are unknown; however, 330:Closely related species and genetic origin 244: 71: 47: 29: 3582: 3536: 2020: 1983: 1958: 1913: 1895: 1794: 1737: 1657: 1608: 1591: 1555: 1466: 1407: 1305: 1224: 1156: 1084: 799: 297: 1136: 1035: 975:(3rd ed.). London: Academic Press. 682:throughout the range is still required. 397: 3361: 2002: 1455:South African Journal of Marine Science 1260: 1249:South African Journal of Marine Science 1213:South African Journal of Marine Science 860: 290:ecosystem along the southwest coast of 4338: 3499: 2438: 1430: 1142: 968: 845: 843: 731: 694: 622: 573:species, predominantly juvenile hake ( 535: 4346:IUCN Red List near threatened species 3968: 3967: 2478: 2160: 2070: 1614: 710:) are considered an emerging threat. 3398: 1777:Morisaka, T.; Connor, R. C. (2007). 1695:. Academic Press. pp. 313–332. 1167:. Cham: Springer. pp. 445–470. 1162: 1041: 910:Gopal, K; Elwen, S; Plön, S (2016). 873: 849: 722: 656: 592: 557: 3444: 840: 787:IUCN Red List of Threatened Species 478: 13: 56:Size compared to an average human 14: 4397: 3111:Indo-Pacific bottlenose dolphin ( 2050: 1638:African Journal of Marine Science 713: 365:Genetic studies suggest that the 1796:10.1111/j.1420-9101.2007.01336.x 1400:10.1111/j.1748-7692.2008.00246.x 1013:10.1046/j.0962-1083.2001.01360.x 647: 113: 4376:Marine fauna of Southern Africa 4356:Cetaceans of the Atlantic Ocean 3252:Indo-Pacific finless porpoise ( 3129:Tamanend's bottlenose dolphin ( 2977:Indian Ocean humpback dolphin ( 2968:Indo-Pacific humpback dolphin ( 1837:Journal of Experimental Biology 1783:Journal of Evolutionary Biology 1424: 1241: 601:movement pattern is present in 581:species such as juvenile goby ( 473: 435: 302: 4386:Taxa named by John Edward Gray 2798:Southern right whale dolphin ( 2789:Northern right whale dolphin ( 2721:Atlantic white-sided dolphin ( 1077:10.1080/15627020.2010.11657256 972:Encyclopedia of Marine Mammals 815: 632:Sympatry with other delphinids 388: 1: 3835:Deraniyagala's beaked whale ( 3808:Ginkgo-toothed beaked whale ( 3019:Pantropical spotted dolphin ( 2986:Australian humpback dolphin ( 2748:Pacific white-sided dolphin ( 1951:10.1016/j.anbehav.2019.01.007 776:Elwen, S.; Gopal, K. (2018). 754: 562:Prey items consist of mostly 448: 393: 375:Antarctic Circumpolar Current 311:by a Captain Haviside of the 2831:Australian snubfin dolphin ( 2239:North Atlantic right whale ( 1650:10.2989/1814232X.2014.973903 1173:10.1007/978-3-030-03083-4_16 759: 614:Home range and site fidelity 587:Trachurus trachurus capensis 548: 457: 7: 3781:Blainville's beaked whale ( 3722:Tropical bottlenose whale ( 3698:Southern bottlenose whale ( 3689:Northern bottlenose whale ( 3138:Common bottlenose dolphin ( 2995:Atlantic humpback dolphin ( 2248:North Pacific right whale ( 2009:Endangered Species Research 1046:(Special issue 9): 279–299. 10: 4402: 4074:cephalorhynchus-heavisidii 4035:cephalorhynchus-heavisidii 4022:Cephalorhynchus_heavisidii 4008:Cephalorhynchus heavisidii 3978:Cephalorhynchus heavisidii 3880:Stejneger's beaked whale ( 3261:Yangtze finless porpoise ( 3046:Atlantic spotted dolphin ( 2631:Short-finned pilot whale ( 1119:Handbook of Marine Mammals 850:Gray, John Edward (1828). 780:Cephalorhynchus heavisidii 685: 528:and most prevalent in the 313:British East India Company 265:Cephalorhynchus heavisidii 253:Heaviside's dolphin range 229:Cephalorhynchus heavisidii 4381:Mammals described in 1828 4292: 3976: 3925: 3913:Shepherd's beaked whale ( 3901: 3734: 3710: 3669: 3627: 3610: 3573: 3527: 3490: 3456:Araguaian river dolphin ( 3435: 3389: 3352: 3324: 3273: 3232: 3215: 3187: 3163: 3154: 3091: 3067: 3007: 2948: 2915: 2891: 2867: 2843: 2810: 2769: 2700: 2676: 2652: 2640:Long-finned pilot whale ( 2611: 2587: 2563: 2512: 2495: 2491: 2474: 2429: 2401: 2377: 2281: 2264: 2210: 2186: 2177: 2173: 2156: 2106: 1593:10.1644/05-MAMM-A-307R2.1 1468:10.2989/02577619209504746 1352:10.1007/s10592-012-0347-9 1226:10.2989/02577619209504706 939:Carwardine, Mark (2020). 252: 243: 225: 218: 110:Scientific classification 108: 91: 69: 60: 55: 46: 37: 28: 23: 3799:Gervais's beaked whale ( 3754:Sowerby's beaked whale ( 3465:Bolivian river dolphin ( 2057:Namibian Dolphin Project 1548:10.1578/AM.37.2.2011.151 441:calves every 2–4 years. 38:Heaviside's dolphin off 4371:Mammals of South Africa 3937:Cuvier's beaked whale ( 3862:Perrin's beaked whale ( 3826:Hector's beaked whale ( 3790:Ramari's beaked whale ( 3763:Andrews' beaked whale ( 3639:Arnoux's beaked whale ( 3312:Burmeister's porpoise ( 3079:Rough-toothed dolphin ( 2503:(Oceanic dolphins) 2302:Antarctic minke whale ( 1298:10.1111/1749-4877.12380 874:Best, Peter B. (2007). 747:and is included in the 642:Lagenorhynchus obscurus 583:Sufflogobius bibarbatus 577:) and octopus, however 3648:Baird's beaked whale ( 3548:Ganges river dolphin ( 3474:Amazon river dolphin ( 2855:Orca or killer whale ( 2779:(Right whale dolphins) 2712:White-beaked dolphin ( 2230:Southern right whale ( 1897:10.1098/rspb.2018.1178 1165:Biodiversity of Angola 585:) and horse mackerel ( 403: 298:Taxonomy and evolution 4238:Paleobiology Database 3889:Spade-toothed whale ( 3853:True's beaked whale ( 3844:Strap-toothed whale ( 3817:Gray's beaked whale ( 3772:Hubbs' beaked whale ( 3657:Sato's beaked whale ( 3557:Indus river dolphin ( 3285:Spectacled porpoise ( 3101:(Bottlenose dolphins) 2542:Heaviside's dolphin ( 2524:Commerson's dolphin ( 2329:Pygmy Bryde's whale ( 1431:Leeney, Ruth (2011). 1388:Marine Mammal Science 1340:Conservation Genetics 401: 4294:Delphinus heavisidii 3871:Pygmy beaked whale ( 3744:(Mesoplodont whales) 2903:False killer whale ( 2879:Melon-headed whale ( 2599:Pygmy killer whale ( 2293:Common minke whale ( 2003:MacLeod, CD (2009). 1615:Serot, J. L (2013). 1580:Journal of Mammalogy 1505:10.3957/056.039.0204 823:"Appendices | CITES" 794:: e.T4161A50352086. 24:Heaviside's dolphin 3679:(Bottlenose whales) 3419:Dwarf sperm whale ( 3410:Pygmy sperm whale ( 3242:(Finless porpoises) 2958:(Humpback dolphins) 2822:Irrawaddy dolphin ( 2739:Hourglass dolphin ( 2450:Pygmy right whale ( 1286:Integrative Zoology 1261:Golaski, S (2015). 736:Prior to 2018, the 732:Conservation status 695:Bycatch and hunting 623:Dive time and depth 575:Merluccius capensis 543:Merluccius capensis 536:Habitat preferences 360:Southern Hemisphere 348:C. commersonii 344:Commerson's dolphin 260:Heaviside's dolphin 63:Conservation status 4366:Mammals of Namibia 3594:La Plata dolphin ( 3294:Harbour porpoise ( 3263:N. asiaeorientalis 3120:Burrunan dolphin ( 2688:Fraser's dolphin ( 2551:Hector's dolphin ( 1890:(1883): 20181178. 1849:10.1242/jeb.042440 707:Trachurus capensis 491:and into southern 404: 211:C. heavisidii 4361:Mammals of Angola 4333: 4332: 4225:Open Tree of Life 3970:Taxon identifiers 3961: 3960: 3957: 3956: 3953: 3952: 3949: 3948: 3747: 3682: 3621: 3606: 3605: 3569: 3568: 3523: 3522: 3486: 3485: 3431: 3430: 3385: 3384: 3348: 3347: 3336:Dall's porpoise ( 3245: 3226: 3211: 3210: 3150: 3149: 3104: 3055:Spinner dolphin ( 3037:Striped dolphin ( 3028:Clymene dolphin ( 2961: 2782: 2730:Peale's dolphin ( 2664:Risso's dolphin ( 2624: 2533:Chilean dolphin ( 2506: 2485: 2470: 2469: 2466: 2465: 2462: 2461: 2425: 2424: 2275: 2260: 2259: 2223: 2167: 1843:(11): 1940–1949. 1730:10.1121/1.3519401 1702:978-0-08-092035-1 1182:978-3-030-03082-7 1001:Molecular Ecology 952:978-1-4729-7715-1 887:978-0-521-89710-5 723:Boat interactions 657:Population status 593:Movement patterns 558:Diet and Foraging 429:Sexual dimorphism 268:) is one of four 257: 256: 103: 86: 16:Species of mammal 4393: 4326: 4325: 4313: 4312: 4311: 4285: 4284: 4272: 4271: 4259: 4258: 4246: 4245: 4233: 4232: 4220: 4219: 4207: 4206: 4194: 4193: 4191:NHMSYS0000376516 4181: 4180: 4168: 4167: 4155: 4154: 4142: 4141: 4129: 4128: 4116: 4115: 4103: 4102: 4090: 4089: 4077: 4076: 4064: 4063: 4051: 4050: 4038: 4037: 4025: 4024: 4012: 4011: 4010: 3997: 3996: 3995: 3965: 3964: 3746: 3745: 3741: 3681: 3680: 3676: 3625: 3624: 3620: 3619: 3615: 3580: 3579: 3534: 3533: 3497: 3496: 3458:I. araguaiaensis 3442: 3441: 3396: 3395: 3375:P. macrocephalus 3359: 3358: 3244: 3243: 3239: 3230: 3229: 3225: 3224: 3220: 3161: 3160: 3103: 3102: 3098: 2960: 2959: 2955: 2936:Guiana dolphin ( 2781: 2780: 2776: 2633:G. macrorhynchus 2623: 2622: 2618: 2575:Common dolphin ( 2510: 2509: 2505: 2504: 2500: 2493: 2492: 2484:(Toothed whales) 2483: 2476: 2475: 2436: 2435: 2413:Humpback whale ( 2295:B. acutorostrata 2279: 2278: 2274: 2273: 2269: 2222: 2221: 2217: 2184: 2183: 2175: 2174: 2165: 2158: 2157: 2091: 2084: 2077: 2068: 2067: 2044: 2041: 2035: 2034: 2024: 2022:10.3354/esr00197 2000: 1994: 1987: 1981: 1980: 1962: 1939:Animal Behaviour 1934: 1928: 1927: 1917: 1899: 1875: 1869: 1868: 1831: 1825: 1824: 1798: 1789:(4): 1439–1458. 1774: 1768: 1767: 1741: 1713: 1707: 1706: 1686: 1680: 1679: 1661: 1632: 1621: 1620: 1612: 1606: 1605: 1595: 1571: 1562: 1561: 1559: 1531: 1525: 1524: 1487: 1481: 1480: 1470: 1446: 1440: 1439: 1437: 1428: 1422: 1421: 1411: 1383: 1372: 1371: 1334: 1328: 1327: 1309: 1281: 1275: 1274: 1258: 1252: 1245: 1239: 1238: 1228: 1204: 1195: 1194: 1160: 1154: 1153: 1151: 1140: 1134: 1133: 1113: 1107: 1106: 1088: 1059: 1048: 1047: 1039: 1033: 1032: 1007:(9): 2215–2223. 996: 987: 986: 966: 957: 956: 936: 923: 922: 916: 907: 892: 891: 871: 858: 857: 847: 838: 837: 835: 833: 819: 813: 812: 810: 808: 803: 773: 530:Hector's dolphin 479:Geographic range 352:Hector's dolphin 340:C. eutropia 317:John Edward Gray 248: 231: 118: 117: 97: 80: 75: 74: 51: 33: 21: 20: 4401: 4400: 4396: 4395: 4394: 4392: 4391: 4390: 4351:Cephalorhynchus 4336: 4335: 4334: 4329: 4321: 4316: 4307: 4306: 4301: 4288: 4280: 4275: 4267: 4262: 4254: 4249: 4241: 4236: 4228: 4223: 4215: 4210: 4202: 4197: 4189: 4184: 4176: 4171: 4163: 4158: 4150: 4145: 4137: 4132: 4124: 4119: 4111: 4106: 4098: 4093: 4085: 4080: 4072: 4067: 4059: 4054: 4046: 4041: 4033: 4028: 4020: 4015: 4006: 4005: 4000: 3991: 3990: 3985: 3972: 3962: 3945: 3921: 3897: 3783:M. densirostris 3743: 3742: 3740: 3730: 3706: 3678: 3677: 3675: 3665: 3618:(Beaked whales) 3617: 3616: 3614: 3602: 3565: 3519: 3482: 3427: 3381: 3344: 3320: 3269: 3254:N. phocaenoides 3241: 3240: 3238: 3222: 3221: 3219: 3207: 3183: 3146: 3100: 3099: 3097: 3087: 3063: 3057:S. longirostris 3039:S. coeruleoalba 3003: 2957: 2956: 2954: 2944: 2911: 2887: 2863: 2839: 2824:O. brevirostris 2806: 2778: 2777: 2775: 2765: 2757:Dusky dolphin ( 2696: 2672: 2648: 2620: 2619: 2617: 2607: 2583: 2559: 2515:Cephalorhynchus 2502: 2501: 2499: 2487: 2458: 2421: 2415:M. novaeangliae 2397: 2373: 2347:Omura's whale ( 2320:Bryde's whale ( 2271: 2270: 2268: 2266:Balaenopteridae 2256: 2219: 2218: 2216: 2206: 2198:Bowhead whale ( 2169: 2166:(Baleen whales) 2152: 2102: 2095: 2053: 2048: 2047: 2042: 2038: 2001: 1997: 1988: 1984: 1935: 1931: 1876: 1872: 1832: 1828: 1775: 1771: 1714: 1710: 1703: 1687: 1683: 1633: 1624: 1613: 1609: 1572: 1565: 1536:Aquatic Mammals 1532: 1528: 1488: 1484: 1447: 1443: 1435: 1429: 1425: 1384: 1375: 1346:(4): 987–1002. 1335: 1331: 1282: 1278: 1259: 1255: 1246: 1242: 1205: 1198: 1183: 1161: 1157: 1149: 1141: 1137: 1130: 1114: 1110: 1065:African Zoology 1060: 1051: 1040: 1036: 997: 990: 983: 967: 960: 953: 937: 926: 914: 908: 895: 888: 872: 861: 848: 841: 831: 829: 821: 820: 816: 806: 804: 774: 767: 762: 757: 734: 725: 716: 697: 688: 659: 650: 634: 625: 616: 595: 560: 551: 538: 526:Cephalorhynchus 481: 476: 460: 451: 438: 396: 391: 371:Southern Africa 367:Cephalorhynchus 356:C. hectori 336:Chilean dolphin 332: 305: 300: 275:Cephalorhynchus 239: 233: 227: 214: 198:Cephalorhynchus 112: 104: 87: 78:Near Threatened 76: 72: 65: 17: 12: 11: 5: 4399: 4389: 4388: 4383: 4378: 4373: 4368: 4363: 4358: 4353: 4348: 4331: 4330: 4328: 4327: 4314: 4298: 4296: 4290: 4289: 4287: 4286: 4273: 4260: 4247: 4234: 4221: 4208: 4195: 4182: 4169: 4156: 4143: 4130: 4117: 4104: 4091: 4078: 4065: 4052: 4039: 4026: 4013: 3998: 3982: 3980: 3974: 3973: 3959: 3958: 3955: 3954: 3951: 3950: 3947: 3946: 3944: 3943: 3939:Z. cavirostris 3933: 3931: 3923: 3922: 3920: 3919: 3909: 3907: 3899: 3898: 3896: 3895: 3886: 3877: 3868: 3859: 3850: 3841: 3832: 3823: 3814: 3805: 3796: 3787: 3778: 3769: 3760: 3750: 3748: 3732: 3731: 3729: 3728: 3718: 3716: 3708: 3707: 3705: 3704: 3695: 3685: 3683: 3667: 3666: 3664: 3663: 3654: 3645: 3635: 3633: 3622: 3608: 3607: 3604: 3603: 3601: 3600: 3596:P. blainvillei 3590: 3588: 3577: 3571: 3570: 3567: 3566: 3564: 3563: 3554: 3544: 3542: 3531: 3525: 3524: 3521: 3520: 3518: 3517: 3507: 3505: 3494: 3488: 3487: 3484: 3483: 3481: 3480: 3476:I. geoffrensis 3471: 3467:I. boliviensis 3462: 3452: 3450: 3439: 3433: 3432: 3429: 3428: 3426: 3425: 3416: 3406: 3404: 3393: 3387: 3386: 3383: 3382: 3380: 3379: 3369: 3367: 3356: 3350: 3349: 3346: 3345: 3343: 3342: 3332: 3330: 3322: 3321: 3319: 3318: 3314:P. spinipinnis 3309: 3300: 3291: 3281: 3279: 3271: 3270: 3268: 3267: 3258: 3248: 3246: 3227: 3213: 3212: 3209: 3208: 3206: 3205: 3195: 3193: 3185: 3184: 3182: 3181: 3175:Beluga whale ( 3171: 3169: 3166:Delphinapterus 3158: 3152: 3151: 3148: 3147: 3145: 3144: 3135: 3126: 3117: 3107: 3105: 3089: 3088: 3086: 3085: 3081:S. bredanensis 3075: 3073: 3065: 3064: 3062: 3061: 3052: 3043: 3034: 3025: 3015: 3013: 3005: 3004: 3002: 3001: 2992: 2983: 2974: 2964: 2962: 2946: 2945: 2943: 2942: 2933: 2929:S. fluviatilis 2923: 2921: 2913: 2912: 2910: 2909: 2899: 2897: 2889: 2888: 2886: 2885: 2875: 2873: 2865: 2864: 2862: 2861: 2851: 2849: 2841: 2840: 2838: 2837: 2828: 2818: 2816: 2808: 2807: 2805: 2804: 2795: 2785: 2783: 2767: 2766: 2764: 2763: 2754: 2750:L. obliquidens 2745: 2736: 2727: 2718: 2714:L. albirostris 2708: 2706: 2703:Lagenorhynchus 2698: 2697: 2695: 2694: 2684: 2682: 2674: 2673: 2671: 2670: 2660: 2658: 2650: 2649: 2647: 2646: 2637: 2627: 2625: 2621:(Pilot whales) 2609: 2608: 2606: 2605: 2595: 2593: 2585: 2584: 2582: 2581: 2571: 2569: 2561: 2560: 2558: 2557: 2548: 2539: 2530: 2526:C. commersonii 2520: 2518: 2507: 2489: 2488: 2472: 2471: 2468: 2467: 2464: 2463: 2460: 2459: 2457: 2456: 2446: 2444: 2433: 2427: 2426: 2423: 2422: 2420: 2419: 2409: 2407: 2399: 2398: 2396: 2395: 2385: 2383: 2375: 2374: 2372: 2371: 2365:Rice's whale ( 2362: 2353: 2344: 2335: 2326: 2317: 2308: 2304:B. bonaerensis 2299: 2289: 2287: 2276: 2262: 2261: 2258: 2257: 2255: 2254: 2245: 2236: 2226: 2224: 2220:(Right whales) 2208: 2207: 2205: 2204: 2194: 2192: 2181: 2171: 2170: 2154: 2153: 2151: 2150: 2144: 2138: 2136:Laurasiatheria 2132: 2126: 2120: 2114: 2107: 2104: 2103: 2094: 2093: 2086: 2079: 2071: 2065: 2064: 2059: 2052: 2051:External links 2049: 2046: 2045: 2036: 1995: 1982: 1929: 1870: 1826: 1769: 1724:(1): 449–457. 1708: 1701: 1681: 1644:(4): 455–466. 1622: 1607: 1586:(5): 866–877. 1563: 1542:(2): 151–160. 1526: 1499:(2): 143–154. 1482: 1461:(1): 843–861. 1441: 1423: 1394:(1): 107–124. 1373: 1329: 1292:(5): 506–526. 1276: 1253: 1240: 1219:(1): 237–270. 1196: 1181: 1155: 1135: 1129:978-0125885058 1128: 1108: 1049: 1034: 988: 981: 958: 951: 924: 893: 886: 859: 839: 814: 764: 763: 761: 758: 756: 753: 733: 730: 724: 721: 715: 714:Climate change 712: 704:(specifically 702:horse mackerel 696: 693: 687: 684: 658: 655: 649: 646: 638:dusky dolphins 633: 630: 624: 621: 615: 612: 605:, whereby the 594: 591: 559: 556: 550: 547: 537: 534: 480: 477: 475: 472: 459: 456: 450: 447: 437: 434: 395: 392: 390: 387: 331: 328: 321:John Heaviside 309:United Kingdom 304: 301: 299: 296: 255: 254: 250: 249: 241: 240: 234: 223: 222: 216: 215: 208: 206: 202: 201: 194: 190: 189: 184: 180: 179: 174: 170: 169: 164: 160: 159: 154: 150: 149: 144: 140: 139: 134: 130: 129: 124: 120: 119: 106: 105: 92: 89: 88: 70: 67: 66: 61: 58: 57: 53: 52: 44: 43: 35: 34: 26: 25: 15: 9: 6: 4: 3: 2: 4398: 4387: 4384: 4382: 4379: 4377: 4374: 4372: 4369: 4367: 4364: 4362: 4359: 4357: 4354: 4352: 4349: 4347: 4344: 4343: 4341: 4324: 4319: 4315: 4310: 4304: 4300: 4299: 4297: 4295: 4291: 4283: 4278: 4274: 4270: 4265: 4261: 4257: 4252: 4248: 4244: 4239: 4235: 4231: 4226: 4222: 4218: 4213: 4209: 4205: 4200: 4196: 4192: 4187: 4183: 4179: 4174: 4170: 4166: 4161: 4157: 4153: 4148: 4144: 4140: 4135: 4131: 4127: 4122: 4118: 4114: 4109: 4105: 4101: 4096: 4092: 4088: 4083: 4079: 4075: 4070: 4066: 4062: 4057: 4053: 4049: 4044: 4040: 4036: 4031: 4027: 4023: 4018: 4014: 4009: 4003: 3999: 3994: 3988: 3984: 3983: 3981: 3979: 3975: 3971: 3966: 3942: 3940: 3935: 3934: 3932: 3930: 3929: 3924: 3918: 3916: 3911: 3910: 3908: 3906: 3905: 3900: 3894: 3892: 3887: 3885: 3883: 3882:M. stejnegeri 3878: 3876: 3874: 3873:M. peruvianus 3869: 3867: 3865: 3860: 3858: 3856: 3851: 3849: 3847: 3842: 3840: 3838: 3833: 3831: 3829: 3824: 3822: 3820: 3815: 3813: 3811: 3810:M. ginkgodens 3806: 3804: 3802: 3797: 3795: 3793: 3788: 3786: 3784: 3779: 3777: 3775: 3774:M. carlhubbsi 3770: 3768: 3766: 3761: 3759: 3757: 3752: 3751: 3749: 3739: 3738: 3733: 3727: 3725: 3720: 3719: 3717: 3715: 3714: 3709: 3703: 3701: 3700:H. planifrons 3696: 3694: 3692: 3691:H. ampullatus 3687: 3686: 3684: 3674: 3673: 3668: 3662: 3660: 3655: 3653: 3651: 3646: 3644: 3642: 3637: 3636: 3634: 3632: 3631: 3626: 3623: 3613: 3609: 3599: 3597: 3592: 3591: 3589: 3587: 3586: 3581: 3578: 3576: 3575:Pontoporiidae 3572: 3562: 3560: 3555: 3553: 3551: 3546: 3545: 3543: 3541: 3540: 3535: 3532: 3530: 3529:Platanistidae 3526: 3516: 3514: 3513:L. vexillifer 3509: 3508: 3506: 3504: 3503: 3498: 3495: 3493: 3489: 3479: 3477: 3472: 3470: 3468: 3463: 3461: 3459: 3454: 3453: 3451: 3449: 3448: 3443: 3440: 3438: 3434: 3424: 3422: 3417: 3415: 3413: 3408: 3407: 3405: 3403: 3402: 3397: 3394: 3392: 3388: 3378: 3376: 3373:Sperm whale ( 3371: 3370: 3368: 3366: 3365: 3360: 3357: 3355: 3351: 3341: 3339: 3334: 3333: 3331: 3329: 3328: 3323: 3317: 3315: 3310: 3308: 3306: 3301: 3299: 3297: 3292: 3290: 3288: 3283: 3282: 3280: 3278: 3277: 3272: 3266: 3264: 3259: 3257: 3255: 3250: 3249: 3247: 3237: 3236: 3231: 3228: 3218: 3214: 3204: 3202: 3197: 3196: 3194: 3192: 3191: 3186: 3180: 3178: 3173: 3172: 3170: 3168: 3167: 3162: 3159: 3157: 3153: 3143: 3141: 3136: 3134: 3132: 3127: 3125: 3123: 3118: 3116: 3114: 3109: 3108: 3106: 3096: 3095: 3090: 3084: 3082: 3077: 3076: 3074: 3072: 3071: 3066: 3060: 3058: 3053: 3051: 3049: 3044: 3042: 3040: 3035: 3033: 3031: 3026: 3024: 3022: 3017: 3016: 3014: 3012: 3011: 3006: 3000: 2998: 2993: 2991: 2989: 2988:S. sahulensis 2984: 2982: 2980: 2975: 2973: 2971: 2966: 2965: 2963: 2953: 2952: 2947: 2941: 2939: 2938:S. guianensis 2934: 2932: 2930: 2925: 2924: 2922: 2920: 2919: 2914: 2908: 2906: 2905:P. crassidens 2901: 2900: 2898: 2896: 2895: 2890: 2884: 2882: 2877: 2876: 2874: 2872: 2871: 2870:Peponocephala 2866: 2860: 2858: 2853: 2852: 2850: 2848: 2847: 2842: 2836: 2834: 2829: 2827: 2825: 2820: 2819: 2817: 2815: 2814: 2809: 2803: 2801: 2796: 2794: 2792: 2787: 2786: 2784: 2774: 2773: 2768: 2762: 2760: 2755: 2753: 2751: 2746: 2744: 2742: 2737: 2735: 2733: 2728: 2726: 2724: 2719: 2717: 2715: 2710: 2709: 2707: 2705: 2704: 2699: 2693: 2691: 2686: 2685: 2683: 2681: 2680: 2679:Lagenodelphis 2675: 2669: 2667: 2662: 2661: 2659: 2657: 2656: 2651: 2645: 2643: 2638: 2636: 2634: 2629: 2628: 2626: 2616: 2615: 2610: 2604: 2602: 2597: 2596: 2594: 2592: 2591: 2586: 2580: 2578: 2573: 2572: 2570: 2568: 2567: 2562: 2556: 2554: 2549: 2547: 2545: 2544:C. heavisidii 2540: 2538: 2536: 2531: 2529: 2527: 2522: 2521: 2519: 2517: 2516: 2511: 2508: 2498: 2494: 2490: 2486: 2477: 2473: 2455: 2453: 2448: 2447: 2445: 2443: 2442: 2437: 2434: 2432: 2431:Cetotheriidae 2428: 2418: 2416: 2411: 2410: 2408: 2406: 2405: 2400: 2394: 2392: 2387: 2386: 2384: 2382: 2381: 2376: 2370: 2368: 2363: 2361: 2359: 2354: 2352: 2350: 2345: 2343: 2341: 2336: 2334: 2332: 2327: 2325: 2323: 2318: 2316: 2314: 2309: 2307: 2305: 2300: 2298: 2296: 2291: 2290: 2288: 2286: 2285: 2280: 2277: 2267: 2263: 2253: 2251: 2246: 2244: 2242: 2237: 2235: 2233: 2228: 2227: 2225: 2215: 2214: 2209: 2203: 2201: 2200:B. mysticetus 2196: 2195: 2193: 2191: 2190: 2185: 2182: 2180: 2176: 2172: 2168: 2159: 2155: 2149: 2145: 2143: 2139: 2137: 2133: 2131: 2127: 2125: 2121: 2119: 2115: 2113: 2109: 2108: 2105: 2100: 2092: 2087: 2085: 2080: 2078: 2073: 2072: 2069: 2063: 2062:uk.whales.org 2060: 2058: 2055: 2054: 2040: 2032: 2028: 2023: 2018: 2014: 2010: 2006: 1999: 1992: 1986: 1978: 1974: 1970: 1966: 1961: 1956: 1952: 1948: 1944: 1940: 1933: 1925: 1921: 1916: 1911: 1907: 1903: 1898: 1893: 1889: 1885: 1881: 1874: 1866: 1862: 1858: 1854: 1850: 1846: 1842: 1838: 1830: 1822: 1818: 1814: 1810: 1806: 1802: 1797: 1792: 1788: 1784: 1780: 1773: 1765: 1761: 1757: 1753: 1749: 1745: 1740: 1735: 1731: 1727: 1723: 1719: 1712: 1704: 1698: 1694: 1693: 1685: 1677: 1673: 1669: 1665: 1660: 1655: 1651: 1647: 1643: 1639: 1631: 1629: 1627: 1618: 1611: 1603: 1599: 1594: 1589: 1585: 1581: 1577: 1570: 1568: 1558: 1553: 1549: 1545: 1541: 1537: 1530: 1522: 1518: 1514: 1510: 1506: 1502: 1498: 1494: 1486: 1478: 1474: 1469: 1464: 1460: 1456: 1452: 1445: 1434: 1427: 1419: 1415: 1410: 1405: 1401: 1397: 1393: 1389: 1382: 1380: 1378: 1369: 1365: 1361: 1357: 1353: 1349: 1345: 1341: 1333: 1325: 1321: 1317: 1313: 1308: 1303: 1299: 1295: 1291: 1287: 1280: 1272: 1268: 1264: 1257: 1250: 1244: 1236: 1232: 1227: 1222: 1218: 1214: 1210: 1203: 1201: 1192: 1188: 1184: 1178: 1174: 1170: 1166: 1159: 1148: 1147: 1139: 1131: 1125: 1121: 1120: 1112: 1104: 1100: 1096: 1092: 1087: 1082: 1078: 1074: 1070: 1066: 1058: 1056: 1054: 1045: 1038: 1030: 1026: 1022: 1018: 1014: 1010: 1006: 1002: 995: 993: 984: 982:9780080919935 978: 974: 973: 965: 963: 954: 948: 944: 943: 935: 933: 931: 929: 920: 913: 906: 904: 902: 900: 898: 889: 883: 879: 878: 870: 868: 866: 864: 855: 854: 846: 844: 828: 824: 818: 802: 797: 793: 789: 788: 783: 781: 772: 770: 765: 752: 750: 746: 741: 739: 729: 720: 711: 709: 708: 703: 692: 683: 680: 676: 672: 668: 664: 654: 648:Vocalisations 645: 643: 639: 629: 620: 611: 608: 604: 600: 590: 588: 584: 580: 576: 572: 568: 565: 555: 546: 544: 533: 531: 527: 521: 519: 518:Britannia Bay 515: 511: 507: 503: 499: 494: 490: 486: 471: 469: 465: 464:killer whales 455: 446: 444: 433: 430: 426: 425:pectoral fins 422: 418: 414: 410: 400: 386: 384: 383:South America 380: 376: 372: 368: 363: 361: 357: 353: 349: 345: 341: 337: 327: 324: 322: 318: 314: 310: 295: 293: 289: 285: 281: 277: 276: 272:in the genus 271: 267: 266: 261: 251: 247: 242: 237: 232: 230: 224: 221: 220:Binomial name 217: 213: 212: 207: 204: 203: 200: 199: 195: 192: 191: 188: 185: 182: 181: 178: 175: 172: 171: 168: 165: 162: 161: 158: 155: 152: 151: 148: 145: 142: 141: 138: 135: 132: 131: 128: 125: 122: 121: 116: 111: 107: 101: 95: 90: 84: 79: 68: 64: 59: 54: 50: 45: 41: 36: 32: 27: 22: 19: 4293: 3977: 3938: 3926: 3915:T. shepherdi 3914: 3902: 3891:M. traversii 3890: 3881: 3872: 3863: 3854: 3845: 3836: 3827: 3818: 3809: 3801:M. europaeus 3800: 3791: 3782: 3773: 3764: 3755: 3735: 3724:I. pacificus 3723: 3711: 3699: 3690: 3670: 3658: 3649: 3640: 3628: 3595: 3583: 3558: 3550:P. gangetica 3549: 3537: 3512: 3500: 3475: 3466: 3457: 3445: 3420: 3412:K. breviceps 3411: 3399: 3374: 3362: 3354:Physeteridae 3337: 3327:Phocoenoides 3325: 3313: 3304: 3295: 3287:P. dioptrica 3286: 3274: 3262: 3253: 3233: 3201:M. monoceros 3200: 3188: 3176: 3164: 3156:Monodontidae 3140:T. truncatus 3139: 3131:T. erebennus 3130: 3122:T. australis 3121: 3112: 3092: 3080: 3068: 3056: 3048:S. frontalis 3047: 3038: 3029: 3021:S. attenuata 3020: 3008: 2996: 2987: 2978: 2970:S. chinensis 2969: 2949: 2937: 2928: 2916: 2904: 2892: 2880: 2868: 2856: 2844: 2833:O. heinsohni 2832: 2823: 2811: 2799: 2790: 2772:Lissodelphis 2770: 2758: 2749: 2740: 2732:L. australis 2731: 2722: 2713: 2701: 2689: 2677: 2665: 2653: 2641: 2632: 2614:Globicephala 2612: 2601:F. attenuata 2600: 2588: 2576: 2564: 2552: 2543: 2541: 2534: 2525: 2513: 2452:C. marginata 2451: 2439: 2414: 2402: 2390: 2389:Gray whale ( 2380:Eschrichtius 2378: 2366: 2357: 2348: 2339: 2338:Blue whale ( 2330: 2321: 2312: 2303: 2294: 2284:Balaenoptera 2282: 2249: 2241:E. glacialis 2240: 2232:E. australis 2231: 2211: 2199: 2187: 2148:Whippomorpha 2142:Artiodactyla 2039: 2012: 2008: 1998: 1990: 1985: 1942: 1938: 1932: 1887: 1883: 1873: 1840: 1836: 1829: 1786: 1782: 1772: 1739:10722/140937 1721: 1717: 1711: 1691: 1684: 1641: 1637: 1616: 1610: 1583: 1579: 1539: 1535: 1529: 1496: 1492: 1485: 1458: 1454: 1444: 1426: 1391: 1387: 1343: 1339: 1332: 1289: 1285: 1279: 1262: 1256: 1248: 1243: 1216: 1212: 1164: 1158: 1145: 1138: 1118: 1111: 1071:(1): 78–91. 1068: 1064: 1043: 1037: 1004: 1000: 971: 941: 918: 876: 852: 830:. Retrieved 826: 817: 805:. Retrieved 791: 785: 779: 742: 735: 726: 717: 705: 698: 689: 671:Lamberts Bay 660: 651: 641: 635: 626: 617: 603:South Africa 596: 586: 582: 574: 561: 552: 542: 539: 525: 522: 498:South Africa 482: 474:Distribution 468:Orcinus orca 467: 461: 452: 439: 436:Life history 405: 381:and then to 366: 364: 355: 347: 339: 333: 325: 315:. Zoologist 306: 303:Nomenclature 278:. The small 273: 264: 263: 259: 258: 228: 226: 210: 209: 197: 173:Infraorder: 167:Artiodactyla 18: 4251:SeaLifeBase 4108:iNaturalist 4002:Wikispecies 3846:M. layardii 3765:M. bowdoini 3713:Indopacetus 3296:P. phocoena 3235:Neophocoena 3223:(Porpoises) 3217:Phocoenidae 2791:L. borealis 2759:L. obscurus 2741:L. cruciger 2535:C. eutropia 2497:Delphinidae 2482:Odontoceti 2391:E. robustus 2358:B. physalus 2356:Fin whale ( 2340:B. musculus 2313:B. borealis 2311:Sei whale ( 2250:E. japonica 2134:Superorder 2128:Infraclass 2015:: 125–136. 1945:: 273–284. 1251:12:237–270. 807:19 November 389:Description 379:New Zealand 187:Delphinidae 96:Appendix II 4340:Categories 4309:Q125572400 3904:Tasmacetus 3864:M. perrini 3837:M. hotaula 3828:M. hectori 3737:Mesoplodon 3672:Hyperoodon 3659:B. minimus 3650:B. bairdii 3641:B. arnuxii 3585:Pontoporia 3539:Platanista 3113:T. aduncus 3030:S. clymene 2997:S. teuszii 2979:S. plumbea 2881:P. electra 2800:L. peronii 2666:G. griseus 2577:D. delphis 2553:C. hectori 2480:Parvorder 2272:(Rorquals) 2179:Balaenidae 2164:Mysticeti 2162:Parvorder 1993:27:494-497 1960:2263/71067 1659:2263/43900 1557:2263/17051 1409:2263/10207 1307:2263/74315 1271:2263/57243 1086:2263/14283 832:14 January 755:References 675:Walvis bay 571:cephalopod 502:Walvis Bay 485:Cape Point 449:Group size 413:dorsal fin 394:Morphology 350:) and the 42:, Namibia 3756:M. bidens 3630:Berardius 3612:Ziphiidae 3492:Lipotidae 3303:Vaquita ( 3199:Narwhal ( 3177:D. leucas 2894:Pseudorca 2723:L. acutus 2566:Delphinus 2404:Megaptera 2349:B. omurai 2322:B. brydei 2213:Eubalaena 2146:Suborder 2031:1863-5407 1969:0003-3472 1906:0962-8452 1857:0022-0949 1805:1010-061X 1748:0001-4966 1668:1814-232X 1602:0022-2372 1513:0379-4369 1477:0257-7615 1418:0824-0469 1360:1566-0621 1235:0257-7615 1191:134352239 1103:219289679 1095:1562-7020 827:cites.org 760:Citations 667:Table Bay 663:abundance 661:No total 549:Behaviour 514:Cape Town 510:Table Bay 458:Predation 443:Gestation 205:Species: 133:Kingdom: 127:Eukaryota 4303:Wikidata 4264:Species+ 4178:14300039 4126:11241419 4087:46559320 3987:Wikidata 3928:Ziphius 3855:M. mirus 3819:M. grayi 3559:P. minor 3421:K. simus 3391:Kogiidae 3364:Physeter 3338:P. dalli 3305:P. sinus 3276:Phocoena 3094:Tursiops 3010:Stenella 2927:Tucuxi ( 2813:Orcaella 2690:L. hosei 2642:G. melas 2367:B. ricei 2331:B. edeni 2130:Eutheria 2124:Mammalia 2118:Chordata 2112:Animalia 2110:Kingdom 1977:72334526 1924:30051842 1865:20472781 1821:23902777 1813:17584238 1756:21303024 1676:56296143 1521:85572457 1368:17218356 1324:59306988 1316:30688009 1029:24368161 1021:11555263 679:Lüderitz 607:dolphins 564:demersal 506:Lüderitz 421:blowhole 288:Benguela 280:cetacean 270:dolphins 183:Family: 157:Mammalia 147:Chordata 143:Phylum: 137:Animalia 123:Domain: 83:IUCN 3.1 40:Lüderitz 4323:2440521 4165:1006409 4100:5220041 3993:Q301048 3792:M. eueu 3511:Baiji ( 3502:Lipotes 3437:Iniidae 3190:Monodon 2918:Sotalia 2857:O. orca 2846:Orcinus 2655:Grampus 2441:Caperea 2189:Balaena 2116:Phylum 2101:species 2099:Cetacea 2097:Extant 1915:6083265 1764:8219776 686:Threats 599:diurnal 579:pelagic 489:Namibia 342:), the 286:to the 284:endemic 193:Genus: 177:Cetacea 163:Order: 153:Class: 98: ( 81: ( 4282:254978 4230:860906 4217:254978 4204:103583 4139:180451 4048:462567 4030:ARKive 2590:Feresa 2140:Order 2122:Class 2029:  1975:  1967:  1922:  1912:  1904:  1863:  1855:  1819:  1811:  1803:  1762:  1754:  1746:  1699:  1674:  1666:  1600:  1519:  1511:  1475:  1416:  1366:  1358:  1322:  1314:  1233:  1189:  1179:  1126:  1101:  1093:  1027:  1019:  979:  949:  884:  516:) and 493:Angola 409:thorax 292:Africa 238:, 1828 4277:WoRMS 4256:69391 4243:82045 4121:IRMNG 4113:41517 4061:69J6N 3401:Kogia 3070:Steno 2951:Sousa 1973:S2CID 1817:S2CID 1760:S2CID 1672:S2CID 1517:S2CID 1436:(PDF) 1364:S2CID 1320:S2CID 1187:S2CID 1150:(PDF) 1099:S2CID 1025:S2CID 915:(PDF) 417:fluke 100:CITES 94:CITES 4318:GBIF 4269:4438 4212:OBIS 4199:NCBI 4152:4161 4147:IUCN 4134:ITIS 4095:GBIF 4043:BOLD 3447:Inia 2027:ISSN 1965:ISSN 1920:PMID 1902:ISSN 1861:PMID 1853:ISSN 1809:PMID 1801:ISSN 1752:PMID 1744:ISSN 1697:ISBN 1664:ISSN 1598:ISSN 1509:ISSN 1473:ISSN 1414:ISSN 1356:ISSN 1312:PMID 1231:ISSN 1177:ISBN 1124:ISBN 1091:ISSN 1017:PMID 977:ISBN 947:ISBN 882:ISBN 834:2022 809:2021 792:2018 677:and 669:and 569:and 567:fish 504:and 236:Gray 4186:NBN 4173:MSW 4160:MDD 4082:EoL 4069:CMS 4056:CoL 4017:ADW 2017:doi 1955:hdl 1947:doi 1943:150 1910:PMC 1892:doi 1888:285 1845:doi 1841:213 1791:doi 1734:hdl 1726:doi 1722:129 1654:hdl 1646:doi 1588:doi 1552:hdl 1544:doi 1501:doi 1463:doi 1404:hdl 1396:doi 1348:doi 1302:hdl 1294:doi 1267:hdl 1221:doi 1169:doi 1081:hdl 1073:doi 1009:doi 796:doi 282:is 4342:: 4320:: 4305:: 4279:: 4266:: 4253:: 4240:: 4227:: 4214:: 4201:: 4188:: 4175:: 4162:: 4149:: 4136:: 4123:: 4110:: 4097:: 4084:: 4071:: 4058:: 4045:: 4032:: 4019:: 4004:: 3989:: 2025:. 2011:. 2007:. 1971:. 1963:. 1953:. 1941:. 1918:. 1908:. 1900:. 1886:. 1882:. 1859:. 1851:. 1839:. 1815:. 1807:. 1799:. 1787:20 1785:. 1781:. 1758:. 1750:. 1742:. 1732:. 1720:. 1670:. 1662:. 1652:. 1642:36 1640:. 1625:^ 1596:. 1584:87 1582:. 1578:. 1566:^ 1550:. 1540:37 1538:. 1515:. 1507:. 1497:39 1495:. 1471:. 1459:12 1457:. 1453:. 1412:. 1402:. 1392:25 1390:. 1376:^ 1362:. 1354:. 1344:13 1342:. 1318:. 1310:. 1300:. 1290:14 1288:. 1229:. 1217:12 1215:. 1211:. 1199:^ 1185:. 1175:. 1097:. 1089:. 1079:. 1069:45 1067:. 1052:^ 1023:. 1015:. 1005:10 1003:. 991:^ 961:^ 927:^ 917:. 896:^ 862:^ 842:^ 825:. 790:. 784:. 768:^ 597:A 415:, 362:. 294:. 3941:) 3917:) 3893:) 3884:) 3875:) 3866:) 3857:) 3848:) 3839:) 3830:) 3821:) 3812:) 3803:) 3794:) 3785:) 3776:) 3767:) 3758:) 3726:) 3702:) 3693:) 3661:) 3652:) 3643:) 3598:) 3561:) 3552:) 3515:) 3478:) 3469:) 3460:) 3423:) 3414:) 3377:) 3340:) 3316:) 3307:) 3298:) 3289:) 3265:) 3256:) 3203:) 3179:) 3142:) 3133:) 3124:) 3115:) 3083:) 3059:) 3050:) 3041:) 3032:) 3023:) 2999:) 2990:) 2981:) 2972:) 2940:) 2931:) 2907:) 2883:) 2859:) 2835:) 2826:) 2802:) 2793:) 2761:) 2752:) 2743:) 2734:) 2725:) 2716:) 2692:) 2668:) 2644:) 2635:) 2603:) 2579:) 2555:) 2546:) 2537:) 2528:) 2454:) 2417:) 2393:) 2369:) 2360:) 2351:) 2342:) 2333:) 2324:) 2315:) 2306:) 2297:) 2252:) 2243:) 2234:) 2202:) 2090:e 2083:t 2076:v 2033:. 2019:: 2013:7 1979:. 1957:: 1949:: 1926:. 1894:: 1867:. 1847:: 1823:. 1793:: 1766:. 1736:: 1728:: 1705:. 1678:. 1656:: 1648:: 1604:. 1590:: 1560:. 1554:: 1546:: 1523:. 1503:: 1479:. 1465:: 1438:. 1420:. 1406:: 1398:: 1370:. 1350:: 1326:. 1304:: 1296:: 1273:. 1269:: 1237:. 1223:: 1193:. 1171:: 1132:. 1105:. 1083:: 1075:: 1031:. 1011:: 985:. 955:. 921:. 890:. 836:. 811:. 798:: 782:" 778:" 640:( 512:( 466:( 354:( 346:( 338:( 262:( 102:) 85:)

Index


Lüderitz

Conservation status
Near Threatened
IUCN 3.1
CITES
CITES
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Mammalia
Artiodactyla
Cetacea
Delphinidae
Cephalorhynchus
Binomial name
Gray

dolphins
Cephalorhynchus
cetacean
endemic
Benguela
Africa
United Kingdom
British East India Company
John Edward Gray

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