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Human taxonomy

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2921: 4808:(e.g., Stringer, 1984; Andrews, 1984; Tattersall, 1986; Wood, 1984, 1991a, b; Schwartz and Tattersall, 2000) ... Unlike the European lineage, in my opinion, the taxonomic issues surrounding Asian vs. African H. erectus are more intractable. The issue was most pointedly addressed with the naming of H. ergaster on the basis of the type mandible KNM-ER 992, but also including the partial skeleton and isolated teeth of KNM-ER 803 among other Koobi Fora remains (Groves and Mazak, 1975). Recently, this specific name was applied to most early African and Georgian H. erectus in recognition of the less-derived nature of these remains vis Ă  vis conditions in Asian H. erectus (see Wood, 1991a, p. 268; Gabunia et al., 2000a). At least portions of the paratype of H. ergaster (e.g., KNM-ER 1805) are not included in most current conceptions of that taxon. The 138: 4526:
Gongdibé; Fronty, Pierre; Geraads, Denis; Lehmann, Thomas; Lihoreau, Fabrice; Louchart, Antoine; Mahamat, Adoum; Merceron, Gildas; Mouchelin, Guy; Otero, Olga; Campomanes, Pablo Pelaez; De Leon, Marcia Ponce; Rage, Jean-Claude; Sapanet, Michel; Schuster, Mathieu; Sudre, Jean; Tassy, Pascal; Valentin, Xavier; Vignaud, Patrick; Viriot, Laurent; Zazzo, Antoine; Zollikofer, Christoph (11 July 2002). "A new hominid from the Upper Miocene of Chad, Central Africa".
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Statement of the Principle of Coordination applied to species-group names. A name established for a taxon at either rank in the species group is deemed to have been simultaneously established by the same author for a taxon at the other rank in the group; both nominal taxa have the same name-bearing
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Brunet, Michel; Guy, Franck; Pilbeam, David; Mackaye, Hassane Taisso; Likius, Andossa; Ahounta, Djimdoumalbaye; Beauvilain, Alain; Blondel, Cécile; Bocherens, Hervé; Boisserie, Jean-Renaud; De Bonis, Louis; Coppens, Yves; Dejax, Jean; Denys, Christiane; Duringer, Philippe; Eisenmann, Véra; Fanone,
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may have appeared some 2 million years ago. Fossils dated to as much as 1.8 million years ago have been found both in Africa and in Southeast Asia, and the oldest fossils by a narrow margin (1.85 to 1.77 million years ago) were found in the Caucasus, so that it is unclear whether
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is rarely used before the 1940s. In 1946, John Wendell Bailey attributes the name to Linnaeus (1758) explicitly: "Linnaeus. Syst. Nat. ed. 10, Vol. 1. pp. 20, 21, 22, lists five races of man, viz: Homo sapiens sapiens (white — Caucasian) ", This is a misattribution, but
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question remains famously unresolved (e.g., Stringer, 1984; Tattersall, 1986; Wood, 1991a, 1994; Rightmire, 1998b; Gabunia et al., 2000a; Schwartz and Tattersall, 2000), in no small part because the original diagnosis provided no comparison with the Asian fossil
1779:" hoax of 1912 was the fraudulent presentation of such a transitional species. Since the mid-20th century, knowledge of the development of Hominini has become much more detailed, and taxonomical terminology has been altered a number of times to reflect this. 4848:
Lordkipanidze, David; Ponce de Leòn, Marcia S.; Margvelashvili, Ann; Rak, Yoel; Rightmire, G. Philip; Vekua, Abesalom; Zollikofer, Christoph P. E. (18 October 2013). "A Complete Skull from Dmanisi, Georgia, and the Evolutionary Biology of Early Homo".
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over an extended time of hybridization, spanning roughly 10 to 6 million years ago, with possible admixture as late as 4 million years ago. A subtribe of uncertain validity, grouping archaic "pre-human" or "para-human" species younger than the
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DiMaggio EN, Campisano CJ, Rowan J, Dupont-Nivet G, Deino AL, Bibi F, et al. (March 2015). "Paleoanthropology. Late Pliocene fossiliferous sedimentary record and the environmental context of early Homo from Afar, Ethiopia".
1630:, but contemporary anthropologists reject the concept of race as a useful tool to understanding humanity, and instead view humanity as a complex, interrelated genetic continuum. Taxonomy of the hominins continues to evolve. 2987:
A subspecies cannot be recognized independently: a species will either be recognized as having no subspecies at all or at least two (including any that are extinct). Therefore, the designation of an extant subspecies
5714:"A new human species once lived in this Philippine cave – Archaeologists in Luzon Island have turned up the bones of a distantly related species, Homo luzonensis, further expanding the human family tree" 4804:), once thought so primitive, was in fact more derived than its African counterparts. These morphological differences were interpreted by some as evidence that more than one species might be included in 5522:
Bischoff JL, Shamp DD, Aramburu A, et al. (March 2003). "The Sima de los Huesos Hominids Date to Beyond U/Th Equilibrium (>350 kyr) and Perhaps to 400–500 kyr: New Radiometric Dates".
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during the 1730s to 1750s. He was the first to develop the idea that, like other biological entities, groups of people could too share taxonomic classifications. He named the human species as
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has long been assumed to be close to 200,000 years, but since 2017 there have been a number of suggestions extending this time to as high as 300,000 years. In 2017, fossils found in
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Curnoe D (June 2010). "A review of early Homo in southern Africa focusing on cranial, mandibular and dental remains, with the description of a new species (Homo gautengensis sp. nov.)".
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has increased drastically, and a number of intermediate taxa have been proposed in the 20th and early 21st centuries. The most widely accepted taxonomy grouping takes the genus
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has a complicated history. The rank of subspecies in zoology is introduced for convenience, and not by objective criteria, based on pragmatic consideration of factors such as
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De Heinzelin, J; Clark, JD; White, T; Hart, W; Renne, P; Woldegabriel, G; Beyene, Y; Vrba, E (1999). "Environment and behavior of 2.5-million-year-old Bouri hominids".
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A species proposed in 2010 based on the fossil remains of three individuals dated between 1.9 and 0.6 million years ago. The same fossils were also classified as
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Gray, J. E. (1825). "An outline of an attempt at the disposition of Mammalia into Tribes and Families, with a list of genera apparently appertaining to each Tribe".
2920: 7587: 7582: 1763:, which even if "archaic" is recognizable as clearly human, late 19th to early 20th century anthropology for a time was occupied with finding the supposedly " 3569:
Hominins with "proto-Homo" traits may have lived as early as 2.8 million years ago, as suggested by a fossil jawbone classified as transitional between
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Designations alternative to Hominina have been proposed: Australopithecinae (Gregory & Hellman 1939) and Preanthropinae (Cela-Conde & Altaba 2002);
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Muttoni G, Scardia G, Kent DV, Swisher CC, Manzi G (2009). "Pleistocene magnetochronology of early hominin sites at Ceprano and Fontana Ranuccio, Italy".
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Intensive Course in Biological Anthrpology, 1st Summer School of the European Anthropological Association, 16–30 June, 2007, Prague, Czech Republic
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between 300 and 243 thousand years ago is conventional, and makes use of the fact that there is no known fossil in this period. Examples of
2907: 3849:). There is a fossil gap in Europe between 300 and 243 kya, and by convention, fossils younger than 243 kya are called "Neanderthal". 5769: 7714: 5775: 3364:, but even in this case there is a dissenting view arguing that "the skulls may not be distinctive enough to warrant a new subspecies name". 5559:
Dean D, Hublin JJ, Holloway R, Ziegler R (May 1998). "On the phylogenetic position of the pre-Neandertal specimen from Reilingen, Germany".
3209:(2014) frames this as a debate that is unresolvable in principle, "since there is no definition of species perfectly describing the case." 7094: 2347: 1960:
jawbone fossil discovered in 2013, dated to 2.8 Mya, has been argued as being transitional between the two. It is also disputed whether
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the only known genus within Hominina. Alternatively, following Cela-Conde and Ayala (2003), the "pre-human" or "proto-human" genera of
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may have speciated by as early as 315,000 years ago. Genetic evidence has been adduced for an age of roughly 270,000 years.
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exclusively, so that Hominini would have two subtribes, Australopithecina and Hominina, with the only known genus in Hominina being
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since its introduction in 1892 has been divided into numerous subspecies, many of them formerly considered individual species of
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The first humans with "proto-Neanderthal traits" lived in Eurasia as early as 0.6 to 0.35 million years ago (classified as
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1938) is a proposal by Gregory & Hellman (1939). More recently proposed additions to the Australopithecina subtribe include
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in the second half of the 20th century. Derived from the convention, widespread in the 1980s, of considering two subspecies,
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because it represents a chronological grouping rather than being based on clear morphological distinctions from either
3637:. Formerly dated to as late as 50,000 to 40,000 years ago, a 2011 study pushed back the date of its extinction of 3139: 3021: 2977: 7173: 6388: 6058: 6009: 5974: 5524: 4509: 1979: 5964: 5467:"Deeply divergent archaic mitochondrial genome provides lower time boundary for African gene flow into Neanderthals" 7521: 7365: 4944: 3792: 2112:) There appears to be a recent trend, with the availability of ever more difficult-to-classify fossils such as the 1122: 2013:
by definition, to the inclusion of Australopithecines and other possible pre-human or para-human species (such as
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emerged in Africa and migrated to Eurasia, or if, conversely, it evolved in Eurasia and migrated back to Africa.
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fossils are dated to between 2.1 and 1.5 million years ago. This date range overlaps with the emergence of
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A proposal by Wood and Richmond (2000) would introduce Hominina as a subtribe alongside Australopithecina, with
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continue to be considered separate species by some authorities, but the 2010s discovery of genetic evidence of
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The discovery of the first extinct archaic human species from the fossil record dates to the mid 19th century:
6285:"Neanderthal taxonomy reconsidered: implications of 3D primate models of intra- and interspecific differences" 5329:
Ni X, Ji Q, Wu W, Shao Q, Ji Y, Zhang C, Liang L, Ge J, Guo Z, Li J, Li Q, GrĂĽn R, Stringer C (25 June 2021).
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fossils. However, the discovery of more fossils of this type has opened up the debate on the delineation of
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became popular for "modern humans" in the context of Neanderthals being considered a subspecies of
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as originating between two and three million years ago, divided into at least two species, archaic
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is variously considered equivalent or subordinate to the rank of subspecies, and the division of
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Dirks PH, Roberts EM, Hilbert-Wolf H, Kramers JD, Hawks J, Dosseto A, et al. (May 2017).
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Posth C, WiĂźing C, Kitagawa K, Pagani L, van Holstein L, Racimo F, et al. (July 2017).
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Indriati E, Swisher CC, Lepre C, Quinn RL, Suriyanto RA, Hascaryo AT, et al. (2011).
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and Out of Africa: Recent Developments in Paleoanthropology and Prehistoric Archaeology"
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and Out of Africa: Recent Developments in Paleoanthropology and Prehistoric Archaeology"
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DĂ©troit F, Mijares AS, Corny J, Daver G, Zanolli C, Dizon E, et al. (April 2019).
5628: 5595: 5499: 5466: 5406: 5379: 5355: 5330: 5306: 5281: 5193: 5158: 5134: 5099: 5064: 5010: 4882: 4727: 4705: 4666: 4644: 4639: 4559: 4475: 4453: 4448: 4429: 4170: 4088: 4061: 3995: 3970: 3929: 3235: 3154: 2911: 2145: 2039:(2001) has been proposed as a possible ancestor of Hominina but not Australopithecina. 1801: 1668:
Human taxonomy on one hand involves the placement of humans within the taxonomy of the
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Chang CH, Kaifu Y, Takai M, Kono RT, GrĂĽn R, Matsu'ura S, et al. (January 2015).
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Perkins, Sid (17 October 2013). "Skull suggests three early human species were one".
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Ferring R, Oms O, AgustĂ­ J, Berna F, Nioradze M, Shelia T, et al. (June 2011).
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to 143,000 years ago at the latest, more likely before 550,000 years ago.
1537:, with about a dozen further suggestions for species without universal recognition. 7203: 6956: 6773: 6735: 6582: 6316: 6306: 6263: 6245: 6188: 6171: 6089: 5675: 5623: 5615: 5568: 5533: 5494: 5486: 5440: 5435: 5401: 5391: 5350: 5342: 5301: 5293: 5262: 5188: 5178: 5129: 5119: 5074: 5037: 4994: 4948: 4866: 4830: 4773: 4661: 4653: 4610: 4600: 4563: 4543: 4470: 4462: 4421: 4416:
Kaplan, Matt (8 August 2012). "Fossils point to a big family for human ancestors".
4390: 4326: 4244: 4234: 4152: 4083: 4075: 3990: 3982: 3913: 3896: 2961: 2861: 2486: 2253: 2239: 2185: 1784: 1737: 1729: 1661: 984: 933: 395: 3947: 7609: 7535: 7010: 6983: 6881: 6839: 6451: 6345: 5183: 5100:"Earliest human occupations at Dmanisi (Georgian Caucasus) dated to 1.85-1.78 Ma" 4952: 4394: 4272: 3750: 3313:
into subspecies declined. An early authority explicitly avoiding the division of
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made in the 20th century. Many of the original proposals were not using explicit
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heissen" ("the Europid racial group, considered as a subspecies, would be named
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The Journal of the Royal Anthropological Institute of Great Britain and Ireland
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Proceedings of the National Academy of Sciences of the United States of America
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has been taken to originate some two million years ago, since the discovery of
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is a socially controversial subject. Biologists originally classified races as
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Delson, Eric; Tattersall, Ian; Couvering, John Van; Brooks, Alison S. (2004).
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Since the introduction of systematic names in the 18th century, knowledge of
1357: 1025: 345: 266: 84: 6311: 6250: 6093: 5984: 5380:"Homo naledi and associated sediments in the Rising Star Cave, South Africa" 5124: 4998: 4870: 4605: 4331: 4306: 4239: 4197: 7004: 6992: 6934: 6808: 6756: 6721: 6646: 6553: 6330: 6277: 6200: 5956: 5687: 5637: 5572: 5537: 5508: 5415: 5364: 5315: 5202: 5143: 5049: 5006: 4878: 4847: 4787: 4675: 4624: 4555: 4484: 4402: 4340: 4258: 4166: 4097: 4004: 3986: 3925: 3858:
younger than 450 kya, either between 190–130 or between 70–10 kya
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There were variations and additions to the categories of Linnaeus, such as
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Bulletin der Schweizerische Gesellschaft fĂĽr Anthropologie und Ethnologie
5709: 3702: 3696: 3690: 3452: 3440: 3428: 3366: 3250: 3145: 3091: 3041: 2929: 2839: 2673: 2597: 2573: 2118: 1994:) at about 2 million years ago, or alternatively as transitional between 1971: 1742: 1553: 1336: 1079: 703: 405: 59: 6259: 5490: 5396: 4943:. Vol. 1. In collaboration with Thorolf Hardt. Berlin, Heidelberg: 4547: 6873: 6794: 5803:
has since often been attributed to Linnaeus. In actual fact, Linnaeus,
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Philosophical Transactions of the Royal Society B: Biological Sciences
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Harrison, T. (2013). Kimbel, William H.; Martin, Lawrence B. (eds.).
5231:. Vol. 1. European Anthropological Association. pp. 35–41. 4796:
specimens, particularly in Africa, led to the realization that Asian
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Stringer, Chris (June 12, 2003). "Human evolution: Out of Ethiopia".
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In the 1970s a tendency developed to regard the Javanese variety of
6192: 5653:"A new species of Homo from the Late Pleistocene of the Philippines" 4157: 4132: 3917: 6801: 6766: 6743: 6464: 6143:
The tragedy of evolution: the human animal confronts modern society
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After World War II, the practice of dividing extant populations of
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Mammal Species of the World: A Taxonomic and Geographic Reference
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is the only extant human subspecies" appears in the early 1990s.
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only makes sense if at least one other subspecies is recognized.
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Rightmire, GP (June 3, 1983). "The Lake Ndutu cranium and early
5431:"Oldest Homo sapiens fossil claim rewrites our species' history" 6042: 5377: 4931:
Schrenk F, Kullmer O, Bromage T (2007). "The Earliest Putative
3779:, dated to ca. 0.6 million years ago. The transition from 2806: 2009:("hominins") be designated as comprising all species after the 1502:, differentiated, according to some, from the direct ancestor, 209: 179: 159: 7026: 6917: 6907: 6037:
among other names suggested for fossils later subsumed under
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type, whether that type was fixed originally or subsequently.
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as subspecies, following Linnaeus (1758), who had recognized
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M. R. Drennan, "An Australoid Skull from the Cape Flats",
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is widely recognized as the species directly ancestral to
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and current humans as belonging to the same subspecies).
1450: 5650: 4062:"Anthropologists' views on race, ancestry, and genetics" 3380:
has reopened the details of taxonomy of archaic humans.
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There are a number of proposals of extinct varieties of
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The Human Career: Human biological and cultural origins
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Reynolds, Sally C; Gallagher, Andrew (March 29, 2012).
4307:"Early Homo at 2.8 Ma from Ledi-Geraru, Afar, Ethiopia" 3360:, 2003) has gained wide recognition as a subspecies of 2230:
110–140 cm (3 ft 7 in – 4 ft 7 in)
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have been proposed, with varying degrees of consensus.
3300:(McCown & Keith, 1932). Rightmire (1983) proposed 2122:
fossils (2015) to subsume all archaic varieties under
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into an African and an Asian variety; the African is
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1978) is now seen as either a third early species of
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Other proposed genera, now mostly considered part of
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is attributed to "Linnaeus (1758)" by the taxonomic
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Wood and Richmond (2000) proposed that Gray's tribe
1603:. An even more extreme view rejects the division of 1557:(chimpanzees). The two genera are estimated to have 1498:. Current humans have been designated as subspecies 411: 401: 391: 381: 371: 361: 351: 341: 331: 321: 6282: 5593: 3682:is now grouped into various subspecies, including 1615:would imply the reclassification of chimpanzees as 516: 6930:(archaic homo sapiens, anatomically modern humans) 4726: 4501:African Genesis: Perspectives on Hominin Evolution 4289:Journal of the East Africa Natural History Society 3736:. The distinction from descendant species such as 3185:(1899) also had categories based on race, such as 6413:Beiträge zur Ur- und FrĂĽhgeschichte Mitteleuropas 4700:. Archived from the original on October 15, 2013. 4497: 4349:taxon as invalid, made up of fossil specimens of 3201:was proposed by King (1864) as an alternative to 1694:in 1758, as the only member species of the genus 1660:The taxonomic classification of humans following 7745: 6377:"Implications arising from Pithecanthropus VIII" 6369:, with the Chinese variety being referred to as 6283:Harvati, K.; Frost, S.R.; McNulty, K.P. (2004). 5752:as far as I know, there is no type material for 6342:Wiley-Blackwell Encyclopedia of Human Evolution 6290:Proceedings of the National Academy of Sciences 6229:Proceedings of the National Academy of Sciences 6001:Species, Species Concepts and Primate Evolution 5881:26-27 (1945), p. 359.; J. Desmond Clark (ed.), 4902:"Beautiful skull spurs debate on human history" 4576: 4218:Proceedings of the National Academy of Sciences 4210: 3040:, for the "wild" form which he identified with 2908:Interbreeding between archaic and modern humans 1672:(great apes), and on the other the division of 6381:Paleoanthropology: Morphology and Paleoecology 6046:Encyclopedia of Human Evolution and Prehistory 3393:Homo erectus § Descendants and subspecies 3242:by Wilson & Reeder (2005). These include: 1684:. Modern zoological taxonomy was developed by 1508:(with some other research instead classifying 7405: 6436: 5823:Volume 21 (1944), p. 18 (arguing not against 5776:International Code of Zoological Nomenclature 5587: 5280:Ji Q, Wu W, Ji Y, Li Q, Ni X (25 June 2021). 5070:Evolution and Prehistory: The Human Challenge 4491: 3633:, is considered the latest known survival of 2146: 1964:was the first hominin to use stone tools, as 1944:) by definition, its type specimen being the 545: 6415:35). Beier & Beran, LangenweiĂźbach 2002. 5702: 5209: 4637: 4449:"Human evolution: taxonomy and paleobiology" 4446: 4440: 4364: 3234:, even though they are still cited as valid 2932:costume. Linnaeus is sometimes named as the 6915: 5644: 5371: 1720:) means "human being". The systematic name 7412: 7398: 6443: 6429: 5939:Neanderthal Man: In Search of Lost Genomes 5328: 4447:Wood, Bernard; Richmond, Brian G. (2000). 4133:"Long shadow of Linnaeus's human taxonomy" 3971:"The origin and evolution of Homo sapiens" 3383: 3378:archaic human admixture with modern humans 3020:as grouping the native populations of the 2898: 2660: 2153: 2139: 552: 538: 6320: 6310: 6267: 6249: 6081:American Journal of Physical Anthropology 6073: 5627: 5498: 5405: 5395: 5354: 5305: 5192: 5182: 5133: 5123: 4777: 4765:American Journal of Physical Anthropology 4724: 4665: 4614: 4604: 4474: 4361:, Westview Press, New York, 2001, p. 111. 4330: 4304: 4248: 4238: 4156: 4087: 4067:American Journal of Physical Anthropology 4059: 3994: 7335:Human evolutionary developmental biology 6161: 6147:We are the only surviving subspecies of 6140: 5997: 5428: 5279: 5215: 4688: 4211:Cela-Conde, C. J.; Ayala, F. J. (2003). 3968: 3893: 3063:for the native population of Australia. 2919: 1655: 7677: 6374: 6114:English translation (1972–1975): 4824: 4357:. Tattersall, I. & Schwartz, J.H., 4345:. Some paleoanthropologists regard the 3036:, respectively. Linnaeus also included 1611:as separate genera, which based on the 14: 7746: 6357: 6355: 6221: 5955: 5708: 5027: 4899: 4577:Cela-Conde, C.J.; Ayala, F.J. (2003). 4415: 4034:"Chimps are human, gene study implies" 1446: 7393: 7120:Evolutionary models of human drug use 6424: 6028:Vol. 59 (Jul. - Dec., 1929), 417-427. 5933: 5739: 4792:By the 1980s, the growing numbers of 4757: 4130: 4031: 2948:The recognition or nonrecognition of 2196: 2191: 2184: 2181: 2178: 2166: 2011:chimpanzee–human last common ancestor 530: 423:other species or subspecies suggested 7360: 6405:Der fossile Mensch von Bilzingsleben 5919:Evolutionary History of the Primates 5819:. This is explicitly pointed out in 5596:"The first archaic Homo from Taiwan" 4187: 4117:American Anthropological Association 3948:"Herto skulls (Homo sapiens idaltu)" 3075:" categories to as many as fifteen: 2663: 2652:150–190 cm (4 ft 11 in – 6 ft 3 in) 2360: 2088:in particular. One proposal divides 1728:of the great apes was introduced by 1389: 1367: 1345: 1321: 1301: 1281: 1257: 1237: 1214: 1191: 1171: 1151: 1131: 1111: 519: 7419: 6450: 6352: 5247:Earth and Planetary Science Letters 4689:Hazarika, Manji (16–30 June 2007). 4467:10.1046/j.1469-7580.2000.19710019.x 3327:(1974). The trinomial nomenclature 2879: 2816: 2802: 2699: 2611:150 cm (4 ft 11 in) 2602: 2584: 2569: 2513: 2498: 2473: 2398: 27:Classification of the human species 24: 6383:. Walter de Gruyter. p. 328. 6167:"Human evolution: Out of Ethiopia" 6116:Grzimek's Animal Life Encyclopedia 5917:Frederick S. Szalay, Eric Delson, 5847:, Oxford University Press (1974), 2619: 2238: 1796:, in the tribe Hominini is due to 1740:including both chimpanzees (genus 25: 7765: 7715:Commonly fossilised invertebrates 6131:, Oxford University Press (1974). 5525:Journal of Archaeological Science 4900:Switek, Brian (17 October 2013). 4760:"Natural history of Homo erectus" 3503:) (de Lumley and de Lumley, 1971) 2964:. The informal taxonomic rank of 2849: 2777: 2740: 2655:50–100 kg (110–220 lb) 2544: 2433: 2321: 2281: 7371: 7359: 7348: 7347: 5740:Ralph, Bob (February 19, 1987). 4658:10.1111/j.1469-7580.2008.00871.x 1974:(discovered in 1972, designated 136: 57: 6397: 6379:. In Tuttle, Russell H. (ed.). 6215: 6155: 6134: 6121: 6108: 6067: 6031: 6018: 5991: 5949: 5927: 5911: 5896: 5883:The Cambridge History of Africa 5867: 5854: 5761: 5733: 5552: 5515: 5458: 5422: 5322: 5273: 5238: 5150: 5091: 5056: 5021: 4969: 4924: 4893: 4841: 4818: 4751: 4718: 4682: 4638:Wood, B.; Lonergan, N. (2008). 4631: 4570: 4518: 4409: 4298: 4265: 3861: 3852: 3827: 3802: 3769: 3673: 3644: 3618: 3601: 3580: 3563: 3546: 3516:Homo erectus bilzingslebenensis 3254:(Klaatsch & Hauser, 1910), 2692:55–70 kg (121–154 lb) 2233:33–55 kg (73–121 lb) 2082:as a subspecies. This concerns 1699: 5873:See e.g. John Wendell Bailey, 5862:Systema naturæ. Regnum animale 5429:Callaway, Ewan (7 June 2017). 4504:. Cambridge University Press. 4204: 4181: 4124: 4104: 4053: 4011: 3962: 3940: 3887: 1490:, is designed to include both 1110: 13: 1: 5750:. No. 1548. p. 59. 5073:(8th ed.). Belmont, CA: 4941:Handbook of Paleoanthropology 4579:"Genera of the human lineage" 4213:"Genera of the human lineage" 3881: 3797:Homo erectus bilzingslebensis 3350:Since the 2000s, the extinct 3198:Homo sapiens neanderthalensis 2893: 2732:100 cm (3 ft 3 in) 2425:175 cm (5 ft 9 in) 2315:100 cm (3 ft 3 in) 1899:(Broom & Anderson 1949), 1622:Categorizing humans based on 1464:is the classification of the 5184:10.1371/journal.pone.0021562 4953:10.1007/978-3-540-33761-4_52 4395:10.1126/science.284.5414.625 4131:Marks, Jonathan (May 2007). 4060:K. Wagner, Jennifer (2016). 4032:Hecht, Jeff (May 19, 2003). 3715:Homo erectus palaeojavanicus 3540: 3472:Homo erectus palaeojavanicus 3281:Homo sapiens cro-magnonensis 3071:(1825) extended Linnaeus's " 2640:(anatomically modern humans) 2056:At least a dozen species of 1788:as a third genus, alongside 1640:History of hominoid taxonomy 7: 7378:Evolutionary biology Portal 6375:Sartono, S. (12 May 2011). 6344:. Chichester, West Sussex: 6224:"The origin of Neandertals" 4735:University of Chicago Press 3535:Timeline of human evolution 3530:Names for the human species 3523: 3343:, the explicit claim that " 3266:(Giuffrida-Ruggeri, 1915), 2358:850 (early) – 1,100 (late) 2098:Homo erectus sensu stricto. 1976:Pithecanthropus rudolfensis 1732:(1825). Gray also supplied 1447: 669: 10: 7770: 5887:Cambridge University Press 5561:Journal of Human Evolution 5347:10.1016/j.xinn.2021.100130 5298:10.1016/j.xinn.2021.100132 5267:10.1016/j.epsl.2009.06.032 5042:10.1016/j.jchb.2010.04.002 4287:by Weinert (1938). Leaky, 3873:Homo sapiens ssp. Denisova 3727:Homo erectus tautavelensis 3496:Homo erectus tautavelensis 3390: 3285:Homo sapiens grimaldiensis 3251:Homo aurignacensis hauseri 2970:anatomically modern humans 2905: 2492:a single fossil, possibly 2049: 2045: 1637: 1633: 1492:anatomically modern humans 7727: 7668: 7560: 7453: 7435: 7428: 7343: 7323:Evolutionary anthropology 7300: 7274: 7227: 7165: 7084: 7043: 7036: 6982: 6906: 6829: 6754: 6708: 6701: 6684: 6644: 6580: 6551: 6515: 6506: 6473: 6458: 6348:. 2013. pp. 328–331. 6141:Kitahara, Michio (1991). 6004:. Springer. p. 361. 5833:Homo sapiens eurpoaeus L. 5680:10.1038/s41586-019-1067-9 5445:10.1038/nature.2017.22114 4835:10.1038/nature.2013.13972 4710:: CS1 maint: unfit URL ( 4426:10.1038/nature.2012.11144 3697:Homo erectus lantianensis 3691:Homo erectus yuanmouensis 3639:H. e. soloensis 3598:by other anthropologists. 3436:Homo erectus lantianensis 3424:Homo erectus yuanmouensis 3303:Homo sapiens rhodesiensis 3256:Notanthropus eurafricanus 3183:Georges Vacher de Lapouge 2998:Principle of Coordination 2468:90 kg (200 lb) 2428:90 kg (200 lb) 2389:East and Southern Africa 2355:60 kg (130 lb) 1804:as a subtribe containing 1674:archaic and modern humans 1494:and extinct varieties of 1480:taxonomy. The systematic 507:Broom & Anderson 1949 436: 429: 302: 297: 284: 277: 133:Scientific classification 131: 34: 5963:; Reeder, D. M. (eds.). 5860:LinnĂ©, Carl von (1758). 5063:Haviland WA, Walrath D, 4802:H. erectus sensu stricto 3847:H. neanderthalensis 3785:H. neanderthalensis 3703:Homo erectus nankinensis 3448:Homo erectus nankinensis 3298:Paleanthropus palestinus 2760:classification uncertain 2735:25 kg (55 lb) 2720:classification uncertain 2674:H. neanderthalensis 2614:45 kg (99 lb) 2454:H. neanderthalensis 2110:Homo erectus sensu lato. 1830:(2001). The position of 562: 7229:Origin of modern humans 6371:Homo erectus pekinensis 6312:10.1073/pnas.0308085100 6251:10.1073/pnas.0904119106 6094:10.1002/ajpa.1330610214 5875:The Mammals of Virginia 5259:2009E&PSL.286..255M 5229:EAA Summer School eBook 5125:10.1073/pnas.1106638108 4999:10.1126/science.aaa1415 4871:10.1126/science.1238484 4693:Homo erectus / ergaster 4606:10.1073/pnas.0832372100 4332:10.1126/science.aaa1343 4240:10.1073/pnas.0832372100 4192:. New Series: 337–344. 4112:"AAA Statement on Race" 3835:H. heidelbergensis 3789:H. heidelbergensis 3781:H. heidelbergensis 3709:Homo erectus pekinensis 3668:Pithecanthropus erectus 3660:Sinanthropus pekinensis 3460:Homo erectus pekinensis 3292:(Kleinschmidt, 1931), 2982:human genetic variation 2494:H. heidelbergensis 2447:H. heidelbergensis 1434:P a r a n t h r o p u s 1293:Dispersal beyond Africa 6222:Hublin, J. J. (2009). 5817:Homo sapiens Europaeus 5573:10.1006/jhev.1998.0214 5538:10.1006/jasc.2002.0834 4947:. pp. 1611–1631. 4935:Fossils". In Henke W, 4305:Villmoare, B. (2015). 3987:10.1098/rstb.2015.0237 3766:is not entirely clear. 3733:Homo erectus georgicus 3721:Homo erectus soloensis 3626:Homo erectus soloensis 3508:Homo erectus georgicus 3484:Homo erectus soloensis 3367:H. s. neanderthalensis 3337:H. s. neanderthalensis 3232:trinomial nomenclature 2978:major racial groupings 2945: 2928:wearing a traditional 2790:H. tsaichangensis 1967:Australopithecus garhi 1665: 565:−10 — 7237:Recent African origin 6475:Last common ancestors 6338:Homo neanderthalensis 5600:Nature Communications 5471:Nature Communications 5220:Homo erectus/ergaster 4806:H. erectus sensu lato 4275:(discovered in 1932, 3969:Stringer, C. (2016). 3791:are fossils found at 3650:Now also included in 3443:) (Woo Ju-Kang, 1964) 3391:Further information: 3269:Telanthropus capensis 3216:Homo sapiens fossilis 3203:Homo neanderthalensis 2923: 2906:Further information: 2686:Europe, Western Asia 2130:Comparative table of 1856:)as separate genera. 1756:Homo neanderthalensis 1702:corresponding to the 1659: 1638:Further information: 1613:Principle of Priority 1476:: "wise man") within 655:−1 — 645:−2 — 635:−3 — 625:−4 — 615:−5 — 605:−6 — 595:−7 — 585:−8 — 575:−9 — 416:Homo neanderthalensis 7252:Behavioral modernity 7242:Multiregional origin 7022:archaic Homo sapiens 7017:Homo heidelbergensis 6962:Red Deer Cave people 6409:Bilzingsleben. Bd. 6 6367:Homo erectus erectus 6179:(6941): 692–3, 695. 5903:Annals of Philosophy 5879:Journal of Mammalogy 5813:Homo sapiens sapiens 5796:Homo sapiens sapiens 5742:"Conforming to type" 5067:, McBride B (2007). 4758:AntĂłn, S.C. (2003). 4190:Annals of Philosophy 4023:The Third Chimpanzee 3795:(also classified as 3757:Homo heidelbergensis 3685:Homo erectus erectus 3491:) (Oppenoorth, 1932) 3412:Homo erectus erectus 3329:Homo sapiens sapiens 3317:into subspecies was 2990:Homo sapiens sapiens 2958:geographic isolation 2778:several individuals 2715:H. floresiensis 2558:H. rhodesiensis 2465:180 cm (5 ft 11 in) 2352:180 cm (5 ft 11 in) 2295:H. gautengensis 1782:The introduction of 1746:) and humans (genus 1680:and, if applicable, 1576:(proposed in 1939). 1500:Homo sapiens sapiens 1249:Earliest stone tools 376:Homo heidelbergensis 314:Homo sapiens sapiens 18:Homo sapiens sapiens 6889:H. neanderthalensis 6809:H. e. tautavelensis 6303:2004PNAS..101.1147H 6242:2009PNAS..10616022H 6185:2003Natur.423..692S 6118:, Volume 11, p. 55. 6049:(Second ed.). 5905:11, London (1826), 5864:(10 ed.). pp. 18ff. 5837:H. s. europeaeus L. 5672:2019Natur.568..181D 5612:2015NatCo...6.6037C 5491:10.1038/ncomms16046 5483:2017NatCo...816046P 5397:10.7554/eLife.24231 5216:Hazarika M (2007). 5175:2011PLoSO...621562I 5116:2011PNAS..10810432F 4991:2015Sci...347.1355D 4912:on October 17, 2013 4906:National Geographic 4863:2013Sci...342..326L 4597:2003PNAS..100.7684C 4548:10.1038/nature00879 4540:2002Natur.418..145B 4387:1999Sci...284..625D 4323:2015Sci...347.1352V 4317:(6228): 1352–1355. 4277:Homo florisbadensis 4271:Introduced for the 4231:2003PNAS..100.7684C 4149:2007Natur.447Q..28M 3910:2003Natur.423..692S 3775:The type fossil is 3577:discovered in 2015. 3431:) (Li et al., 1977) 3353:Homo sapiens idaltu 3065:Bory de St. Vincent 2689:170 cm (5 ft 7 in) 2585:Single or very few 2487:H. cepranensis 2300:also classified as 2254:H. rudolfensis 2160: 1905:(Arambourg, 1954), 1761:H. neanderthalensis 1736:as the name of the 1505:Homo sapiens idaltu 7076:Self-domestication 6867:H. heidelbergensis 6816:H. e. yuanmouensis 6781:H. e. lantianensis 6508:Australopithecines 5893:(with references). 5877:(1946), p. 356.; 5719:The New York Times 5620:10.1038/ncomms7037 4779:10.1002/ajpa.10399 4725:Klein, R. (1999). 4645:Journal of Anatomy 4454:Journal of Anatomy 4080:10.1002/ajpa.23120 3981:(1698): 20150237. 3867:provisional names 3373:H. s. rhodesiensis 3320:Grzimeks Tierleben 3296:(Joleaud, 1931) = 3221:Cro-Magnon fossils 3034:Sub-Saharan Africa 2946: 2912:Early modern human 2830:H. luzonensis 2412:H. antecessor 2129: 1956:. Especially, the 1700:several subspecies 1666: 1050:H. heidelbergensis 7741: 7740: 7723: 7722: 7705:Cultivated plants 7700: 7699: 7664: 7663: 7387: 7386: 7328:Paleoanthropology 7270: 7269: 7247:Archaic admixture 7125:Stoned ape theory 7061:Endurance running 6978: 6977: 6974: 6973: 6970: 6969: 6825: 6824: 6788:H. e. nankinensis 6744:H. tsaichangensis 6680: 6679: 6365:as a subspecies, 6165:(June 12, 2003). 5712:(10 April 2019). 5666:(7751): 181–186. 5084:978-0-495-38190-7 5075:Thomson Wadsworth 4962:978-3-540-32474-4 4857:(6156): 326–331. 4591:(13): 7684–7689. 4534:(6894): 145–151. 4225:(13): 7684–7689. 3950:. talkorigins org 3904:(6941): 693–695. 3745:Homo floresiensis 3288:(Gregory, 1921), 3278:(Dubois, 1921), 3264:proto-aethiopicus 3248:(Lapouge, 1906), 3089:(with subraces), 3069:Essai sur l'Homme 3061:H. s. tasmanianus 3054:H. s. troglodytes 2924:1737 painting of 2916:Race and genetics 2891: 2890: 2800: 2761: 2721: 2567: 2496: 2456: 2383: 2304: 2264: 2219: 2096:and the Asian is 1911:(Coppens, 1965). 1875:(Haeckel, 1895), 1802:Australopithecina 1573:Australopithecina 1544:is placed in the 1468:(systematic name 1459: 1458: 1451:million years ago 1410: 1409: 1388: 1387: 1366: 1365: 1358:Earliest rock art 1344: 1343: 1320: 1319: 1313:Earliest language 1300: 1299: 1280: 1279: 1256: 1255: 1236: 1235: 1226:Earliest sign of 1213: 1212: 1203:Earliest sign of 1190: 1189: 1170: 1169: 1150: 1149: 1130: 1129: 773:Ou. macedoniensis 515: 514: 508: 500: 492: 484: 476: 468: 460: 452: 442: 356:Homo floresiensis 273: 16:(Redirected from 7761: 7675: 7674: 7451: 7450: 7433: 7432: 7414: 7407: 7400: 7391: 7390: 7375: 7363: 7362: 7351: 7350: 7287:Human prehistory 7262:Recent evolution 7257:Early migrations 7199:Thermoregulation 7100:Expensive tissue 7071:Sexual selection 7041: 7040: 6913: 6912: 6795:H. e. pekinensis 6706: 6705: 6699: 6698: 6614:A. bahrelghazali 6583:Australopithecus 6513: 6512: 6483:Chimpanzee–human 6471: 6470: 6445: 6438: 6431: 6422: 6421: 6416: 6401: 6395: 6394: 6359: 6350: 6349: 6334: 6324: 6314: 6281: 6271: 6253: 6219: 6213: 6212: 6159: 6153: 6152: 6138: 6132: 6125: 6119: 6112: 6106: 6105: 6071: 6065: 6064: 6039:neanderthalensis 6035: 6029: 6022: 6016: 6015: 5995: 5989: 5988: 5953: 5947: 5946: 5931: 5925: 5915: 5909: 5900: 5894: 5871: 5865: 5858: 5852: 5794: 5788: 5787: 5778:. Archived from 5774:(4th ed.). 5768:"article 46.1". 5765: 5759: 5758: 5737: 5731: 5730: 5728: 5726: 5706: 5700: 5699: 5657: 5648: 5642: 5641: 5631: 5591: 5585: 5584: 5556: 5550: 5549: 5519: 5513: 5512: 5502: 5462: 5456: 5455: 5453: 5451: 5426: 5420: 5419: 5409: 5399: 5375: 5369: 5368: 5358: 5326: 5320: 5319: 5309: 5277: 5271: 5270: 5253:(1–2): 255–268. 5242: 5236: 5235: 5226: 5213: 5207: 5206: 5196: 5186: 5154: 5148: 5147: 5137: 5127: 5095: 5089: 5088: 5060: 5054: 5053: 5025: 5019: 5018: 4985:(6228): 1355–9. 4973: 4967: 4966: 4928: 4922: 4921: 4919: 4917: 4908:. Archived from 4897: 4891: 4890: 4845: 4839: 4838: 4822: 4816: 4815: 4781: 4755: 4749: 4748: 4732: 4722: 4716: 4715: 4709: 4701: 4699: 4686: 4680: 4679: 4669: 4635: 4629: 4628: 4618: 4608: 4574: 4568: 4567: 4522: 4516: 4515: 4495: 4489: 4488: 4478: 4444: 4438: 4437: 4413: 4407: 4406: 4368: 4362: 4351:Australopithecus 4344: 4334: 4302: 4296: 4269: 4263: 4262: 4252: 4242: 4208: 4202: 4201: 4185: 4179: 4178: 4160: 4128: 4122: 4121: 4108: 4102: 4101: 4091: 4057: 4051: 4050: 4048: 4046: 4015: 4009: 4008: 3998: 3966: 3960: 3959: 3957: 3955: 3944: 3938: 3937: 3891: 3876: 3865: 3859: 3856: 3850: 3837:, also called a 3831: 3825: 3806: 3800: 3773: 3767: 3677: 3671: 3648: 3642: 3622: 3616: 3605: 3599: 3596:Australopithecus 3592:H. ergaster 3584: 3578: 3571:Australopithecus 3567: 3561: 3550: 3275:Homo wadjakensis 3258:(Sergi, 1911), 3213:(1869) proposed 3050:H. s. monstrosus 3006:H. s. americanus 2962:sexual selection 2862:Denisova hominin 2794: 2759: 2719: 2694:(heavily built) 2641: 2562: 2535:Northeast China 2491: 2451: 2378: 2374:H. ergaster 2299: 2258: 2213: 2186:Cranial capacity 2161: 2155: 2148: 2141: 2128: 1996:Australopithecus 1954:Australopithecus 1930:, in the 1960s. 1893:(Dreyer, 1935), 1887:(Pycraft, 1928) 1869:(Dubois, 1894), 1838:Australopithecus 1806:Australopithecus 1785:Australopithecus 1730:John Edward Gray 1662:John Edward Gray 1585:Australopithecus 1439: 1437: 1436: 1422: 1420: 1404: 1395: 1390: 1382: 1380:Earliest clothes 1373: 1368: 1360: 1351: 1346: 1327: 1322: 1307: 1302: 1287: 1282: 1269:Earliest sign of 1263: 1258: 1243: 1238: 1228:Australopithecus 1220: 1215: 1197: 1192: 1183:Earliest bipedal 1177: 1172: 1163:Chimpanzee split 1157: 1152: 1137: 1132: 1117: 1112: 1098: 1097: 1083: 1082: 1066: 1052: 1038: 1010: 997: 977: 964: 936: 934:Australopithecus 923: 908: 891: 878: 854: 841: 828: 815: 802: 789: 777: 758: 745: 732: 720: 706: 693: 680: 678: 666: 661: 656: 651: 646: 641: 636: 631: 626: 621: 616: 611: 606: 601: 596: 591: 586: 581: 576: 571: 566: 554: 547: 540: 534: 524: 523:Hominin timeline 517: 506: 498: 490: 482: 474: 466: 458: 450: 440: 413: 403: 396:Homo rudolfensis 393: 383: 373: 363: 353: 343: 333: 323: 265: 141: 140: 126: 56: 41:Temporal range: 32: 31: 21: 7769: 7768: 7764: 7763: 7762: 7760: 7759: 7758: 7754:Human evolution 7744: 7743: 7742: 7737: 7719: 7696: 7660: 7556: 7523:B. integrifolia 7449: 7424: 7418: 7388: 7383: 7339: 7296: 7282:Human evolution 7266: 7223: 7167: 7161: 7140:Cooperative eye 7085:Specific models 7080: 7032: 7011:Homo antecessor 6966: 6902: 6896:H. rhodesiensis 6860:H. floresiensis 6821: 6802:H. e. soloensis 6774:H. e. georgicus 6750: 6714:H. gautengensis 6689: 6687: 6676: 6640: 6576: 6547: 6502: 6493:Orangutan–human 6462: 6454: 6452:Human evolution 6449: 6419: 6403:Emanuel VlÄŤek: 6402: 6398: 6391: 6360: 6353: 6346:Wiley-Blackwell 6335: 6236:(38): 16022–7. 6220: 6216: 6193:10.1038/423692a 6163:Stringer, Chris 6160: 6156: 6139: 6135: 6127:John R. Baker, 6126: 6122: 6113: 6109: 6072: 6068: 6061: 6036: 6032: 6023: 6019: 6012: 5996: 5992: 5977: 5954: 5950: 5932: 5928: 5916: 5912: 5901: 5897: 5872: 5868: 5859: 5855: 5807:ed. 10 Vol. 1. 5785: 5783: 5767: 5766: 5762: 5738: 5734: 5724: 5722: 5707: 5703: 5655: 5649: 5645: 5592: 5588: 5557: 5553: 5520: 5516: 5463: 5459: 5449: 5447: 5427: 5423: 5376: 5372: 5327: 5323: 5278: 5274: 5243: 5239: 5224: 5214: 5210: 5155: 5151: 5110:(26): 10432–6. 5096: 5092: 5085: 5077:. p. 162. 5061: 5057: 5026: 5022: 4974: 4970: 4963: 4929: 4925: 4915: 4913: 4898: 4894: 4846: 4842: 4823: 4819: 4756: 4752: 4745: 4733:. Chicago, IL: 4723: 4719: 4703: 4702: 4697: 4687: 4683: 4636: 4632: 4575: 4571: 4523: 4519: 4512: 4496: 4492: 4461:(Pt 1): 19–60. 4445: 4441: 4414: 4410: 4381:(5414): 625–9. 4369: 4365: 4303: 4299: 4273:Florisbad Skull 4270: 4266: 4209: 4205: 4186: 4182: 4158:10.1038/447028a 4129: 4125: 4110: 4109: 4105: 4058: 4054: 4044: 4042: 4016: 4012: 3967: 3963: 3953: 3951: 3946: 3945: 3941: 3918:10.1038/423692a 3892: 3888: 3884: 3879: 3866: 3862: 3857: 3853: 3843:H. erectus 3832: 3828: 3820:) suggest that 3810:H. sapiens 3807: 3803: 3774: 3770: 3751:Homo antecessor 3680:H. erectus 3678: 3674: 3652:H. erectus 3649: 3645: 3635:H. erectus 3623: 3619: 3613:H. erectus 3608:H. erectus 3606: 3602: 3588:H. habilis 3585: 3581: 3568: 3564: 3554:H. habilis 3551: 3547: 3543: 3526: 3479:) (Tyler, 2001) 3395: 3389: 3294:Homo galilensis 3272:(Broom, 1917), 3014:H. s. asiaticus 3010:H. s. europaeus 2918: 2904: 2896: 2884: 2833: 2797:H. erectus 2793: 2758: 2718: 2693: 2677: 2664:—— 2644:c. 300–present 2639: 2638: 2634:H. sapiens 2620:15 individuals 2565:H. sapiens 2561: 2490: 2462:Europe, Africa 2450: 2381:H. erectus 2377: 2335:H. erectus 2302:H. habilis 2298: 2257: 2212: 2209:H. habilis 2201: 2188: 2168: 2159: 2054: 2048: 1881:(Black, 1927), 1866:Pithecanthropus 1698:, divided into 1654: 1652:Pithecanthropus 1636: 1595:, and possibly 1517:human evolution 1455: 1454: 1442: 1441: 1440: 1432: 1431: 1429: 1425: 1424: 1423: 1418:H o m i n i d s 1416: 1414: 1406: 1405: 1400: 1393: 1384: 1383: 1378: 1371: 1362: 1361: 1356: 1349: 1340: 1339: 1325: 1316: 1315: 1305: 1296: 1295: 1285: 1276: 1275: 1270: 1261: 1252: 1251: 1241: 1232: 1231: 1218: 1209: 1208: 1195: 1186: 1185: 1175: 1166: 1165: 1155: 1146: 1145: 1135: 1126: 1125: 1115: 1108: 1107: 1106: 1101: 1100: 1099: 1093: 1092: 1088: 1086: 1085: 1084: 1078: 1077: 1073: 1071: 1070: 1069: 1062: 1056: 1055: 1054: 1048: 1043: 1041: 1040: 1039: 1030: 1022: 1014: 1013: 1006: 1000: 999: 998: 989: 981: 980: 973: 967: 966: 965: 956: 948: 940: 939: 932: 926: 925: 924: 915: 911: 910: 909: 900: 896: 895: 894: 887: 881: 880: 879: 870: 862: 858: 857: 856: 850: 845: 844: 843: 837: 832: 831: 830: 824: 819: 818: 817: 811: 806: 805: 804: 798: 793: 792: 791: 787:Chororapithecus 785: 780: 779: 778: 769: 761: 760: 754: 749: 748: 747: 743:Samburupithecus 741: 736: 735: 734: 728: 723: 722: 721: 714: 710: 709: 708: 702: 697: 696: 695: 689: 684: 683: 682: 676: 674: 667: 664: 662: 659: 657: 654: 652: 649: 647: 644: 642: 639: 637: 634: 632: 629: 627: 624: 622: 619: 617: 614: 612: 609: 607: 604: 602: 599: 597: 594: 592: 589: 587: 584: 582: 579: 577: 574: 572: 569: 567: 564: 558: 532: 526: 522: 511: 472:Pithecanthropus 459:Arambourg, 1954 444: 443: 386:Homo luzonensis 326:Homo antecessor 293: 290: 264: 135: 127: 125: 124: 123: 122: 117: 112: 107: 102: 97: 92: 87: 82: 77: 72: 67: 62: 51: 50: 39: 28: 23: 22: 15: 12: 11: 5: 7767: 7757: 7756: 7739: 7738: 7736: 7731: 7729: 7725: 7724: 7721: 7720: 7718: 7717: 7712: 7707: 7701: 7698: 7697: 7695: 7694: 7689: 7681: 7679: 7672: 7666: 7665: 7662: 7661: 7659: 7658: 7657: 7656: 7649: 7639: 7634: 7629: 7622: 7617: 7612: 7607: 7602: 7597: 7592: 7591: 7590: 7585: 7580: 7572: 7564: 7562: 7558: 7557: 7555: 7554: 7547: 7540: 7533: 7528: 7527: 7526: 7512: 7505: 7498: 7493: 7486: 7485: 7484: 7472: 7465: 7457: 7455: 7448: 7447: 7441: 7439: 7430: 7426: 7425: 7417: 7416: 7409: 7402: 7394: 7385: 7384: 7382: 7381: 7369: 7357: 7344: 7341: 7340: 7338: 7337: 7332: 7331: 7330: 7320: 7315: 7310: 7304: 7302: 7298: 7297: 7295: 7294: 7292:Human timeline 7289: 7284: 7278: 7276: 7272: 7271: 7268: 7267: 7265: 7264: 7259: 7254: 7249: 7244: 7239: 7233: 7231: 7225: 7224: 7222: 7221: 7216: 7211: 7206: 7201: 7196: 7191: 7186: 7181: 7176: 7170: 7168: 7163: 7162: 7160: 7159: 7158: 7157: 7152: 7144: 7143: 7142: 7137: 7129: 7128: 7127: 7122: 7117: 7115:Drunken monkey 7109: 7108: 7107: 7102: 7097: 7088: 7086: 7082: 7081: 7079: 7078: 7073: 7068: 7063: 7058: 7053: 7047: 7045: 7044:General models 7038: 7034: 7033: 7031: 7030: 6988: 6986: 6980: 6979: 6976: 6975: 6972: 6971: 6968: 6967: 6965: 6964: 6959: 6954: 6949: 6944: 6937: 6932: 6923: 6921: 6910: 6904: 6903: 6901: 6900: 6892: 6885: 6878: 6870: 6863: 6856: 6848: 6843: 6835: 6833: 6831:Archaic humans 6827: 6826: 6823: 6822: 6820: 6819: 6812: 6805: 6798: 6791: 6784: 6777: 6770: 6762: 6760: 6752: 6751: 6749: 6748: 6740: 6736:H. rudolfensis 6732: 6725: 6718: 6709: 6703: 6696: 6682: 6681: 6678: 6677: 6675: 6674: 6667: 6660: 6657:P. aethiopicus 6652: 6650: 6642: 6641: 6639: 6638: 6631: 6624: 6617: 6610: 6603: 6596: 6588: 6586: 6578: 6577: 6575: 6574: 6567: 6559: 6557: 6549: 6548: 6546: 6545: 6538: 6535:Sahelanthropus 6531: 6524: 6521:Nakalipithecus 6516: 6510: 6504: 6503: 6501: 6500: 6495: 6490: 6485: 6479: 6477: 6468: 6456: 6455: 6448: 6447: 6440: 6433: 6425: 6418: 6417: 6396: 6389: 6351: 6297:(5): 1147–52. 6214: 6154: 6145:. p. xi. 6133: 6120: 6107: 6066: 6059: 6030: 6017: 6010: 5990: 5975: 5948: 5945:. p. 237. 5926: 5910: 5895: 5866: 5853: 5839:"). See also: 5829:H. s. albus L. 5811:does not have 5760: 5732: 5701: 5643: 5586: 5567:(5): 485–508. 5551: 5532:(3): 275–280. 5514: 5457: 5421: 5370: 5335:The Innovation 5321: 5286:The Innovation 5272: 5237: 5208: 5149: 5090: 5083: 5055: 5020: 4968: 4961: 4923: 4892: 4840: 4817: 4750: 4743: 4717: 4681: 4652:(4): 354–376. 4630: 4569: 4517: 4510: 4490: 4439: 4408: 4363: 4359:Extinct Humans 4297: 4264: 4203: 4180: 4123: 4103: 4074:(2): 318–327. 4052: 4028:Morris Goodman 4010: 3961: 3939: 3885: 3883: 3880: 3878: 3877: 3869:Homo sp. Altai 3860: 3851: 3826: 3801: 3768: 3672: 3643: 3617: 3600: 3579: 3562: 3544: 3542: 3539: 3538: 3537: 3532: 3525: 3522: 3521: 3520: 3512: 3504: 3492: 3480: 3468: 3456: 3444: 3432: 3420: 3388: 3382: 3260:Homo fossilis 3245:Homo grimaldii 3042:feral children 2903: 2897: 2895: 2892: 2889: 2888: 2885: 2881: 2878: 2876: 2874: 2872: 2867: 2864: 2858: 2857: 2854: 2851: 2850:3 individuals 2848: 2846: 2844: 2842: 2837: 2834: 2825: 2824: 2821: 2818: 2815: 2813: 2811: 2809: 2804: 2801: 2785: 2784: 2782: 2779: 2776: 2774: 2772: 2770: 2765: 2762: 2749: 2748: 2745: 2742: 2741:7 individuals 2739: 2736: 2733: 2730: 2725: 2722: 2710: 2709: 2706: 2703: 2698: 2695: 2690: 2687: 2684: 2678: 2669: 2668: 2665: 2662: 2659: 2656: 2653: 2650: 2645: 2642: 2628: 2627: 2624: 2621: 2618: 2615: 2612: 2609: 2604: 2601: 2598:H. naledi 2593: 2592: 2589: 2586: 2583: 2580: 2578: 2576: 2571: 2568: 2553: 2552: 2549: 2546: 2543: 2540: 2538: 2536: 2533: 2530: 2522: 2521: 2518: 2515: 2512: 2509: 2507: 2505: 2500: 2497: 2482: 2481: 2478: 2475: 2472: 2469: 2466: 2463: 2460: 2457: 2442: 2441: 2438: 2435: 2432: 2429: 2426: 2423: 2421:Western Europe 2418: 2415: 2407: 2406: 2403: 2400: 2397: 2394: 2392: 2390: 2387: 2384: 2369: 2368: 2365: 2362: 2359: 2356: 2353: 2350: 2341: 2338: 2330: 2329: 2326: 2323: 2322:3 individuals 2320: 2318: 2316: 2313: 2308: 2305: 2290: 2289: 2286: 2283: 2280: 2277: 2275: 2273: 2268: 2265: 2259:membership in 2249: 2248: 2245: 2242: 2237: 2234: 2231: 2228: 2223: 2220: 2214:membership in 2204: 2203: 2198: 2195: 2190: 2183: 2180: 2177: 2174: 2167:Temporal range 2165: 2158: 2157: 2150: 2143: 2135: 2114:Dmanisi skulls 2100:(Inclusion of 2094:Homo ergaster, 2050:Main article: 2047: 2044: 2022:Sahelanthropus 1908:Tchadanthropus 1890:Africanthropus 1854:incertae sedis 1850:Sahelanthropus 1833:Sahelanthropus 1648:Anthropomorpha 1635: 1632: 1619:(or similar). 1597:Sahelanthropus 1496:archaic humans 1462:Human taxonomy 1457: 1456: 1448: 1444: 1443: 1428: 1427: 1426: 1413: 1412: 1411: 1408: 1407: 1399: 1398: 1396: 1386: 1385: 1377: 1376: 1374: 1364: 1363: 1355: 1354: 1352: 1342: 1341: 1331: 1330: 1328: 1318: 1317: 1311: 1310: 1308: 1298: 1297: 1291: 1290: 1288: 1278: 1277: 1267: 1266: 1264: 1254: 1253: 1247: 1246: 1244: 1234: 1233: 1224: 1223: 1221: 1211: 1210: 1201: 1200: 1198: 1188: 1187: 1181: 1180: 1178: 1168: 1167: 1161: 1160: 1158: 1148: 1147: 1141: 1140: 1138: 1128: 1127: 1121: 1120: 1118: 1109: 1104: 1103: 1102: 1091: 1090: 1089: 1087: 1076: 1075: 1074: 1072: 1059: 1058: 1057: 1046: 1045: 1044: 1042: 1003: 1002: 1001: 985:H. rudolfensis 970: 969: 968: 929: 928: 927: 914: 913: 912: 899: 898: 897: 884: 883: 882: 861: 860: 859: 848: 847: 846: 839:Graecopithecus 835: 834: 833: 826:Sahelanthropus 822: 821: 820: 809: 808: 807: 796: 795: 794: 783: 782: 781: 756:Ouranopithecus 752: 751: 750: 739: 738: 737: 730:Nakalipithecus 726: 725: 724: 713: 712: 711: 700: 699: 698: 687: 686: 685: 672: 671: 670: 668: 665:0 — 663: 658: 653: 648: 643: 638: 633: 628: 623: 618: 613: 608: 603: 598: 593: 588: 583: 578: 573: 568: 563: 560: 559: 557: 556: 549: 542: 531: 528: 527: 520: 513: 512: 510: 509: 501: 496:Tchadanthropus 493: 485: 477: 469: 461: 453: 448:Africanthropus 439: 438: 437: 434: 433: 427: 426: 420: 419: 409: 399: 389: 379: 369: 359: 349: 339: 329: 319: 318: 317: 300: 299: 295: 294: 292:Linnaeus, 1758 291: 282: 281: 275: 274: 257: 253: 252: 247: 243: 242: 237: 233: 232: 227: 223: 222: 217: 213: 212: 207: 203: 202: 197: 193: 192: 187: 183: 182: 177: 173: 172: 167: 163: 162: 157: 153: 152: 147: 143: 142: 129: 128: 120: 119: 118: 113: 108: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 40: 26: 9: 6: 4: 3: 2: 7766: 7755: 7752: 7751: 7749: 7735: 7732: 7730: 7726: 7716: 7713: 7711: 7710:Invertebrates 7708: 7706: 7703: 7702: 7693: 7690: 7688: 7687: 7683: 7682: 7680: 7676: 7673: 7671: 7667: 7655: 7654: 7650: 7648: 7645: 7644: 7643: 7640: 7638: 7635: 7633: 7630: 7628: 7627: 7623: 7621: 7618: 7616: 7613: 7611: 7608: 7606: 7603: 7601: 7598: 7596: 7593: 7589: 7586: 7584: 7581: 7579: 7576: 7575: 7573: 7571: 7570: 7566: 7565: 7563: 7559: 7553: 7552: 7548: 7546: 7545: 7541: 7539: 7538: 7534: 7532: 7529: 7525: 7524: 7520: 7519: 7518: 7517: 7513: 7511: 7510: 7506: 7504: 7503: 7499: 7497: 7494: 7492: 7491: 7487: 7483: 7482: 7478: 7477: 7476: 7473: 7471: 7470: 7466: 7464: 7463: 7459: 7458: 7456: 7452: 7446: 7443: 7442: 7440: 7438: 7434: 7431: 7427: 7422: 7415: 7410: 7408: 7403: 7401: 7396: 7395: 7392: 7380: 7379: 7374: 7370: 7368: 7367: 7358: 7356: 7355: 7346: 7345: 7342: 7336: 7333: 7329: 7326: 7325: 7324: 7321: 7319: 7316: 7314: 7311: 7309: 7306: 7305: 7303: 7299: 7293: 7290: 7288: 7285: 7283: 7280: 7279: 7277: 7273: 7263: 7260: 7258: 7255: 7253: 7250: 7248: 7245: 7243: 7240: 7238: 7235: 7234: 7232: 7230: 7226: 7220: 7217: 7215: 7212: 7210: 7207: 7205: 7202: 7200: 7197: 7195: 7192: 7190: 7187: 7185: 7182: 7180: 7177: 7175: 7172: 7171: 7169: 7164: 7156: 7153: 7151: 7148: 7147: 7146:Life history 7145: 7141: 7138: 7136: 7133: 7132: 7130: 7126: 7123: 7121: 7118: 7116: 7113: 7112: 7110: 7106: 7103: 7101: 7098: 7096: 7093: 7092: 7090: 7089: 7087: 7083: 7077: 7074: 7072: 7069: 7067: 7064: 7062: 7059: 7057: 7054: 7052: 7049: 7048: 7046: 7042: 7039: 7035: 7029: 7028: 7023: 7019: 7018: 7013: 7012: 7007: 7006: 7001: 7000: 6999:Homo ergaster 6995: 6994: 6990: 6989: 6987: 6985: 6981: 6963: 6960: 6958: 6955: 6953: 6950: 6948: 6945: 6943: 6942: 6938: 6936: 6933: 6931: 6929: 6928:H. s. sapiens 6925: 6924: 6922: 6920: 6919: 6914: 6911: 6909: 6908:Modern humans 6905: 6898: 6897: 6893: 6891: 6890: 6886: 6884: 6883: 6882:H. luzonensis 6879: 6876: 6875: 6871: 6869: 6868: 6864: 6862: 6861: 6857: 6854: 6853: 6849: 6847: 6844: 6842: 6841: 6840:H. antecessor 6837: 6836: 6834: 6832: 6828: 6818: 6817: 6813: 6811: 6810: 6806: 6804: 6803: 6799: 6797: 6796: 6792: 6790: 6789: 6785: 6783: 6782: 6778: 6776: 6775: 6771: 6769: 6768: 6767:H. e. erectus 6764: 6763: 6761: 6759: 6758: 6753: 6746: 6745: 6741: 6738: 6737: 6733: 6731: 6730: 6726: 6724: 6723: 6719: 6716: 6715: 6711: 6710: 6707: 6704: 6700: 6697: 6695: 6693: 6683: 6673: 6672: 6668: 6666: 6665: 6661: 6659: 6658: 6654: 6653: 6651: 6649: 6648: 6643: 6637: 6636: 6632: 6630: 6629: 6625: 6623: 6622: 6621:A. deyiremeda 6618: 6616: 6615: 6611: 6609: 6608: 6604: 6602: 6601: 6597: 6595: 6594: 6590: 6589: 6587: 6585: 6584: 6579: 6573: 6572: 6568: 6566: 6565: 6561: 6560: 6558: 6556: 6555: 6550: 6544: 6543: 6542:Kenyanthropus 6539: 6537: 6536: 6532: 6530: 6529: 6525: 6523: 6522: 6518: 6517: 6514: 6511: 6509: 6505: 6499: 6496: 6494: 6491: 6489: 6488:Gorilla–human 6486: 6484: 6481: 6480: 6478: 6476: 6472: 6469: 6466: 6461: 6457: 6453: 6446: 6441: 6439: 6434: 6432: 6427: 6426: 6423: 6414: 6410: 6406: 6400: 6392: 6390:9783110810691 6386: 6382: 6378: 6372: 6368: 6364: 6358: 6356: 6347: 6343: 6340:King, 1864". 6339: 6332: 6328: 6323: 6318: 6313: 6308: 6304: 6300: 6296: 6292: 6291: 6286: 6279: 6275: 6270: 6265: 6261: 6257: 6252: 6247: 6243: 6239: 6235: 6231: 6230: 6225: 6218: 6210: 6206: 6202: 6198: 6194: 6190: 6186: 6182: 6178: 6174: 6173: 6168: 6164: 6158: 6151: 6150: 6144: 6137: 6130: 6124: 6117: 6111: 6103: 6099: 6095: 6091: 6088:(2): 245–54. 6087: 6083: 6082: 6077: 6070: 6062: 6060:9781135582272 6056: 6052: 6048: 6047: 6040: 6034: 6027: 6021: 6013: 6011:9781489937452 6007: 6003: 6002: 5994: 5986: 5982: 5978: 5976:0-801-88221-4 5972: 5968: 5967: 5962: 5961:Wilson, D. E. 5958: 5957:Groves, C. P. 5952: 5944: 5940: 5936: 5935:Pääbo, Svante 5930: 5924: 5920: 5914: 5908: 5904: 5899: 5892: 5888: 5884: 5880: 5876: 5870: 5863: 5857: 5850: 5846: 5842: 5841:John R. Baker 5838: 5834: 5830: 5827:but against " 5826: 5825:H. s. sapiens 5822: 5818: 5814: 5810: 5806: 5802: 5801:H. s. sapiens 5797: 5793: 5782:on 2016-03-03 5781: 5777: 5773: 5772: 5771:ICZN glossary 5764: 5757: 5755: 5749: 5748: 5747:New Scientist 5743: 5736: 5721: 5720: 5715: 5711: 5705: 5697: 5693: 5689: 5685: 5681: 5677: 5673: 5669: 5665: 5661: 5654: 5647: 5639: 5635: 5630: 5625: 5621: 5617: 5613: 5609: 5605: 5601: 5597: 5590: 5582: 5578: 5574: 5570: 5566: 5562: 5555: 5547: 5543: 5539: 5535: 5531: 5527: 5526: 5518: 5510: 5506: 5501: 5496: 5492: 5488: 5484: 5480: 5476: 5472: 5468: 5461: 5446: 5442: 5438: 5437: 5432: 5425: 5417: 5413: 5408: 5403: 5398: 5393: 5389: 5385: 5381: 5374: 5366: 5362: 5357: 5352: 5348: 5344: 5341:(3): 100130. 5340: 5336: 5332: 5325: 5317: 5313: 5308: 5303: 5299: 5295: 5292:(3): 100132. 5291: 5287: 5283: 5276: 5268: 5264: 5260: 5256: 5252: 5248: 5241: 5234: 5230: 5223: 5221: 5212: 5204: 5200: 5195: 5190: 5185: 5180: 5176: 5172: 5169:(6): e21562. 5168: 5164: 5160: 5153: 5145: 5141: 5136: 5131: 5126: 5121: 5117: 5113: 5109: 5105: 5101: 5094: 5086: 5080: 5076: 5072: 5071: 5066: 5059: 5051: 5047: 5043: 5039: 5036:(3): 151–77. 5035: 5031: 5024: 5016: 5012: 5008: 5004: 5000: 4996: 4992: 4988: 4984: 4980: 4972: 4964: 4958: 4954: 4950: 4946: 4942: 4938: 4934: 4927: 4911: 4907: 4903: 4896: 4888: 4884: 4880: 4876: 4872: 4868: 4864: 4860: 4856: 4852: 4844: 4836: 4832: 4828: 4821: 4814: 4811: 4807: 4803: 4799: 4795: 4789: 4785: 4780: 4775: 4771: 4767: 4766: 4761: 4754: 4746: 4740: 4736: 4731: 4730: 4721: 4713: 4707: 4696: 4694: 4685: 4677: 4673: 4668: 4663: 4659: 4655: 4651: 4647: 4646: 4641: 4634: 4626: 4622: 4617: 4612: 4607: 4602: 4598: 4594: 4590: 4586: 4585: 4580: 4573: 4565: 4561: 4557: 4553: 4549: 4545: 4541: 4537: 4533: 4529: 4521: 4513: 4511:9781107019959 4507: 4503: 4502: 4494: 4486: 4482: 4477: 4472: 4468: 4464: 4460: 4456: 4455: 4450: 4443: 4435: 4431: 4427: 4423: 4419: 4412: 4404: 4400: 4396: 4392: 4388: 4384: 4380: 4376: 4375: 4367: 4360: 4356: 4352: 4348: 4342: 4338: 4333: 4328: 4324: 4320: 4316: 4312: 4308: 4301: 4294: 4290: 4286: 4282: 4278: 4274: 4268: 4260: 4256: 4251: 4246: 4241: 4236: 4232: 4228: 4224: 4220: 4219: 4214: 4207: 4199: 4195: 4191: 4184: 4176: 4172: 4168: 4164: 4159: 4154: 4150: 4146: 4142: 4138: 4134: 4127: 4119: 4118: 4113: 4107: 4099: 4095: 4090: 4085: 4081: 4077: 4073: 4069: 4068: 4063: 4056: 4041: 4040: 4039:New Scientist 4035: 4029: 4025: 4024: 4019: 4018:Jared Diamond 4014: 4006: 4002: 3997: 3992: 3988: 3984: 3980: 3976: 3972: 3965: 3949: 3943: 3935: 3931: 3927: 3923: 3919: 3915: 3911: 3907: 3903: 3899: 3898: 3890: 3886: 3874: 3870: 3864: 3855: 3848: 3844: 3840: 3839:chronospecies 3836: 3830: 3823: 3819: 3815: 3811: 3805: 3798: 3794: 3793:Bilzingsleben 3790: 3786: 3782: 3778: 3772: 3765: 3764: 3759: 3758: 3753: 3752: 3747: 3746: 3741: 3740: 3739:Homo ergaster 3735: 3734: 3729: 3728: 3723: 3722: 3717: 3716: 3711: 3710: 3705: 3704: 3699: 3698: 3693: 3692: 3687: 3686: 3681: 3676: 3669: 3665: 3661: 3657: 3653: 3647: 3640: 3636: 3632: 3628: 3627: 3621: 3614: 3609: 3604: 3597: 3593: 3589: 3583: 3576: 3572: 3566: 3559: 3555: 3549: 3545: 3536: 3533: 3531: 3528: 3527: 3519:(VlÄŤek, 2002) 3518: 3517: 3513: 3510: 3509: 3505: 3502: 3498: 3497: 3493: 3490: 3486: 3485: 3481: 3478: 3474: 3473: 3469: 3466: 3462: 3461: 3457: 3454: 3450: 3449: 3445: 3442: 3438: 3437: 3433: 3430: 3426: 3425: 3421: 3418: 3414: 3413: 3409: 3408: 3407: 3405: 3401: 3400: 3394: 3386: 3381: 3379: 3375: 3374: 3369: 3368: 3363: 3359: 3355: 3354: 3348: 3346: 3345:H. s. sapiens 3342: 3341:H. s. sapiens 3338: 3334: 3330: 3326: 3322: 3321: 3316: 3312: 3307: 3305: 3304: 3299: 3295: 3291: 3290:Homo drennani 3287: 3286: 3282: 3277: 3276: 3271: 3270: 3265: 3261: 3257: 3253: 3252: 3247: 3246: 3241: 3237: 3233: 3229: 3224: 3222: 3219:based on the 3218: 3217: 3212: 3208: 3204: 3200: 3199: 3194: 3192: 3188: 3184: 3181:. Similarly, 3180: 3179: 3174: 3173: 3168: 3167: 3162: 3161: 3156: 3152: 3148: 3147: 3142: 3141: 3136: 3135: 3130: 3129: 3128:australasicus 3124: 3123: 3118: 3117: 3112: 3111: 3106: 3105: 3100: 3099: 3094: 3093: 3088: 3087: 3082: 3081:smooth-haired 3078: 3074: 3070: 3066: 3062: 3057: 3055: 3051: 3047: 3046:cryptozoology 3043: 3039: 3035: 3031: 3027: 3023: 3019: 3015: 3011: 3007: 3003: 2999: 2995: 2994:H. s. sapiens 2991: 2985: 2983: 2979: 2975: 2971: 2967: 2963: 2959: 2955: 2951: 2943: 2942:H. s. sapiens 2939: 2935: 2931: 2927: 2926:Carl Linnaeus 2922: 2917: 2913: 2909: 2901: 2886: 2882: 2877: 2875: 2873: 2871: 2868: 2865: 2863: 2860: 2859: 2855: 2852: 2847: 2845: 2843: 2841: 2838: 2835: 2832: 2831: 2827: 2826: 2822: 2819: 2817:1 individual 2814: 2812: 2810: 2808: 2805: 2798: 2792: 2791: 2787: 2786: 2783: 2780: 2775: 2773: 2771: 2769: 2766: 2763: 2757: 2756: 2753:Nesher Ramla 2751: 2750: 2746: 2743: 2737: 2734: 2731: 2729: 2726: 2723: 2717: 2716: 2712: 2711: 2707: 2704: 2702: 2696: 2691: 2688: 2685: 2683: 2679: 2676: 2675: 2671: 2670: 2666: 2657: 2654: 2651: 2649: 2646: 2643: 2637: 2636: 2635: 2630: 2629: 2625: 2622: 2616: 2613: 2610: 2608: 2605: 2600: 2599: 2595: 2594: 2590: 2587: 2581: 2579: 2577: 2575: 2572: 2566: 2560: 2559: 2555: 2554: 2550: 2547: 2545:1 individual 2541: 2539: 2537: 2534: 2531: 2529: 2528: 2527:H. longi 2524: 2523: 2519: 2516: 2510: 2508: 2506: 2504: 2501: 2495: 2489: 2488: 2484: 2483: 2479: 2476: 2470: 2467: 2464: 2461: 2458: 2455: 2449: 2448: 2444: 2443: 2439: 2436: 2430: 2427: 2424: 2422: 2419: 2416: 2414: 2413: 2409: 2408: 2404: 2401: 2395: 2393: 2391: 2388: 2385: 2382: 2376: 2375: 2371: 2370: 2366: 2363: 2357: 2354: 2351: 2349: 2345: 2342: 2339: 2337: 2336: 2332: 2331: 2327: 2324: 2319: 2317: 2314: 2312: 2309: 2306: 2303: 2297: 2296: 2292: 2291: 2287: 2284: 2278: 2276: 2274: 2272: 2269: 2266: 2262: 2256: 2255: 2251: 2250: 2246: 2243: 2241: 2235: 2232: 2229: 2227: 2224: 2221: 2217: 2211: 2210: 2206: 2205: 2199: 2194: 2193:Fossil record 2187: 2179:Adult height 2175: 2172: 2163: 2162: 2156: 2151: 2149: 2144: 2142: 2137: 2136: 2133: 2127: 2125: 2121: 2120: 2115: 2111: 2107: 2103: 2102:Homo ergaster 2099: 2095: 2091: 2087: 2086: 2085:Homo ergaster 2081: 2077: 2073: 2069: 2068: 2063: 2059: 2053: 2043: 2040: 2038: 2037: 2032: 2028: 2024: 2023: 2018: 2017: 2012: 2008: 2003: 2001: 1997: 1993: 1989: 1985: 1981: 1977: 1973: 1969: 1968: 1963: 1959: 1955: 1951: 1947: 1943: 1939: 1935: 1934: 1929: 1925: 1924:Olduvai Gorge 1921: 1917: 1912: 1910: 1909: 1904: 1903: 1898: 1897: 1892: 1891: 1886: 1885: 1884:Cyphanthropus 1880: 1879: 1874: 1873: 1872:Protanthropus 1868: 1867: 1862: 1857: 1855: 1851: 1847: 1846:Praeanthropus 1843: 1839: 1835: 1834: 1829: 1828: 1827:Kenyanthropus 1823: 1822: 1817: 1813: 1812: 1807: 1803: 1799: 1795: 1791: 1787: 1786: 1780: 1778: 1774: 1770: 1766: 1762: 1758: 1757: 1751: 1749: 1745: 1744: 1739: 1735: 1731: 1727: 1723: 1719: 1715: 1714: 1709: 1705: 1701: 1697: 1693: 1692: 1687: 1686:Carl Linnaeus 1683: 1679: 1675: 1671: 1663: 1658: 1653: 1649: 1645: 1641: 1631: 1629: 1625: 1620: 1618: 1617:Homo paniscus 1614: 1610: 1606: 1602: 1598: 1594: 1593:Praeanthropus 1590: 1586: 1582: 1577: 1575: 1574: 1569: 1565: 1560: 1556: 1555: 1550: 1547: 1543: 1538: 1536: 1535: 1530: 1529: 1524: 1523: 1518: 1513: 1511: 1507: 1506: 1501: 1497: 1493: 1489: 1488: 1483: 1479: 1475: 1471: 1467: 1466:human species 1463: 1452: 1445: 1438: 1435: 1421: 1419: 1403: 1402:Modern humans 1397: 1392: 1391: 1381: 1375: 1370: 1369: 1359: 1353: 1348: 1347: 1338: 1334: 1333:Earliest fire 1329: 1324: 1323: 1314: 1309: 1304: 1303: 1294: 1289: 1284: 1283: 1274: 1273: 1265: 1260: 1259: 1250: 1245: 1240: 1239: 1230: 1229: 1222: 1217: 1216: 1207: 1206: 1199: 1194: 1193: 1184: 1179: 1174: 1173: 1164: 1159: 1154: 1153: 1144: 1143:Gorilla split 1139: 1134: 1133: 1124: 1119: 1114: 1113: 1096: 1081: 1068: 1067: 1065: 1053: 1051: 1037: 1035: 1029: 1027: 1021: 1019: 1018:H. antecessor 1012: 1011: 1009: 996: 994: 988: 986: 979: 978: 976: 963: 961: 960:Au. anamensis 955: 953: 952:Au. afarensis 947: 945: 944:Au. africanus 938: 937: 935: 922: 920: 907: 905: 893: 892: 890: 877: 875: 874:O. tugenensis 869: 867: 855: 853: 842: 840: 829: 827: 816: 814: 803: 801: 790: 788: 776: 774: 768: 766: 759: 757: 746: 744: 733: 731: 719: 718: 707: 705: 694: 692: 681: 679: 561: 555: 550: 548: 543: 541: 536: 535: 529: 525: 518: 505: 502: 499:Coppens, 1965 497: 494: 489: 486: 483:Haeckel, 1895 481: 480:Protanthropus 478: 473: 470: 467:Pycraft, 1928 465: 464:Cyphanthropus 462: 457: 454: 449: 446: 445: 435: 432: 428: 425: 424: 418: 417: 410: 408: 407: 400: 398: 397: 390: 388: 387: 380: 378: 377: 370: 368: 367: 360: 358: 357: 350: 348: 347: 346:Homo ergaster 340: 338: 337: 330: 328: 327: 320: 316: 315: 311: 310: 309: 308: 304: 303: 301: 296: 289: 288: 283: 280: 276: 272: 268: 263: 262: 258: 255: 254: 251: 248: 245: 244: 241: 238: 235: 234: 231: 228: 225: 224: 221: 218: 215: 214: 211: 208: 205: 204: 201: 198: 195: 194: 191: 188: 185: 184: 181: 178: 175: 174: 171: 168: 165: 164: 161: 158: 155: 154: 151: 148: 145: 144: 139: 134: 130: 116: 111: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 55: 52:2.865–0  48: 44: 37: 33: 30: 19: 7684: 7670:Polyphyletic 7653:Homo sapiens 7652: 7651: 7625: 7567: 7550: 7543: 7536: 7522: 7515: 7508: 7501: 7489: 7480: 7468: 7460: 7437:Monophyletic 7376: 7364: 7352: 7219:Gender roles 7214:Intelligence 7027:Homo sapiens 7025: 7021: 7015: 7009: 7005:Homo erectus 7003: 6997: 6993:Homo habilis 6991: 6952:Manot people 6941:H. s. idaltu 6939: 6935:Jebel Irhoud 6927: 6918:Homo sapiens 6916: 6894: 6887: 6880: 6872: 6865: 6858: 6850: 6838: 6814: 6807: 6800: 6793: 6786: 6779: 6772: 6765: 6757:Homo erectus 6755: 6742: 6734: 6727: 6720: 6712: 6702:Proto-humans 6691: 6688:proto-humans 6669: 6662: 6655: 6647:Paranthropus 6645: 6633: 6626: 6619: 6612: 6607:A. anamensis 6605: 6600:A. africanus 6598: 6593:A. afarensis 6591: 6581: 6569: 6562: 6554:Ardipithecus 6552: 6540: 6533: 6526: 6519: 6498:Gibbon–human 6459: 6412: 6408: 6404: 6399: 6380: 6370: 6366: 6362: 6341: 6337: 6294: 6288: 6233: 6227: 6217: 6176: 6170: 6157: 6149:Homo sapiens 6148: 6146: 6142: 6136: 6128: 6123: 6115: 6110: 6085: 6079: 6078:in Africa". 6076:Homo sapiens 6075: 6069: 6045: 6038: 6033: 6025: 6020: 6000: 5993: 5965: 5951: 5941:. New York: 5938: 5929: 5918: 5913: 5902: 5898: 5882: 5878: 5874: 5869: 5856: 5844: 5836: 5832: 5828: 5824: 5820: 5816: 5812: 5804: 5800: 5795: 5790: 5784:. Retrieved 5780:the original 5770: 5763: 5754:Homo sapiens 5753: 5751: 5745: 5735: 5723:. Retrieved 5717: 5704: 5663: 5659: 5646: 5603: 5599: 5589: 5564: 5560: 5554: 5529: 5523: 5517: 5474: 5470: 5460: 5448:. Retrieved 5434: 5424: 5387: 5383: 5373: 5338: 5334: 5324: 5289: 5285: 5275: 5250: 5246: 5240: 5232: 5228: 5219: 5211: 5166: 5162: 5152: 5107: 5103: 5093: 5069: 5058: 5033: 5029: 5023: 4982: 4978: 4971: 4940: 4937:Tattersall I 4932: 4926: 4916:22 September 4914:. Retrieved 4910:the original 4895: 4854: 4850: 4843: 4826: 4820: 4809: 4805: 4801: 4797: 4793: 4791: 4769: 4763: 4753: 4728: 4720: 4692: 4684: 4649: 4643: 4633: 4588: 4582: 4572: 4531: 4527: 4520: 4500: 4493: 4458: 4452: 4442: 4417: 4411: 4378: 4372: 4366: 4358: 4354: 4350: 4346: 4314: 4310: 4300: 4288: 4280: 4276: 4267: 4222: 4216: 4206: 4189: 4183: 4143:(7140): 28. 4140: 4136: 4126: 4115: 4106: 4071: 4065: 4055: 4043:. Retrieved 4037: 4026:(1991), and 4021: 4013: 3978: 3974: 3964: 3952:. Retrieved 3942: 3901: 3895: 3889: 3872: 3868: 3863: 3854: 3846: 3842: 3834: 3829: 3822:Homo sapiens 3821: 3814:Jebel Irhoud 3809: 3804: 3796: 3788: 3784: 3780: 3771: 3763:Homo sapiens 3761: 3760:and indeed 3755: 3749: 3743: 3737: 3731: 3725: 3719: 3713: 3707: 3701: 3695: 3689: 3683: 3679: 3675: 3667: 3659: 3651: 3646: 3638: 3634: 3624: 3620: 3612: 3607: 3603: 3595: 3591: 3587: 3582: 3574: 3570: 3565: 3558:Homo erectus 3557: 3553: 3548: 3514: 3506: 3501:Tautavel Man 3494: 3482: 3477:Meganthropus 3470: 3458: 3446: 3434: 3422: 3410: 3403: 3399:Homo erectus 3397: 3396: 3385:Homo erectus 3384: 3371: 3365: 3362:Homo sapiens 3361: 3357: 3351: 3349: 3344: 3340: 3336: 3332: 3328: 3318: 3314: 3311:Homo sapiens 3310: 3308: 3301: 3297: 3293: 3289: 3279: 3273: 3267: 3263: 3259: 3255: 3249: 3243: 3239: 3228:Homo sapiens 3227: 3225: 3214: 3211:Louis Lartet 3202: 3196: 3195: 3190: 3186: 3176: 3170: 3164: 3158: 3155:crisp-haired 3150: 3144: 3138: 3132: 3126: 3120: 3114: 3108: 3102: 3096: 3090: 3084: 3076: 3068: 3060: 3058: 3053: 3049: 3037: 3026:West Eurasia 3017: 3013: 3009: 3005: 3001: 2993: 2989: 2986: 2973: 2954:Homo sapiens 2953: 2947: 2941: 2937: 2900:Homo sapiens 2899: 2828: 2803:c. 100 2796: 2788: 2754: 2713: 2697:1,200–1,900 2672: 2632: 2631: 2607:South Africa 2603:c. 300 2596: 2570:c. 300 2564: 2556: 2525: 2514:1 skull cap 2499:c. 450 2493: 2485: 2471:1,100–1,400 2453: 2445: 2410: 2386:1,800–1,300 2380: 2372: 2333: 2311:South Africa 2301: 2293: 2260: 2252: 2222:2,100–1,500 2215: 2207: 2131: 2124:Homo erectus 2123: 2117: 2109: 2106:Homo erectus 2105: 2101: 2097: 2093: 2090:Homo erectus 2089: 2083: 2080:Homo sapiens 2079: 2076:Homo erectus 2075: 2072:Homo sapiens 2071: 2067:Homo erectus 2065: 2062:Homo sapiens 2061: 2057: 2055: 2041: 2034: 2030: 2026: 2020: 2016:Ardipithecus 2014: 2004: 1999: 1995: 1991: 1987: 1983: 1975: 1965: 1961: 1953: 1949: 1941: 1937: 1933:Homo habilis 1931: 1915: 1913: 1906: 1902:Atlanthropus 1900: 1896:Telanthropus 1894: 1888: 1882: 1878:Sinanthropus 1876: 1870: 1864: 1860: 1858: 1853: 1852:(the latter 1849: 1845: 1842:Ardipithecus 1841: 1837: 1831: 1825: 1821:Ardipithecus 1819: 1811:Paranthropus 1809: 1805: 1798:Raymond Dart 1793: 1789: 1783: 1781: 1777:Piltdown Man 1772: 1768: 1765:missing link 1760: 1754: 1752: 1747: 1741: 1717: 1711: 1695: 1691:Homo sapiens 1689: 1667: 1621: 1616: 1608: 1604: 1600: 1596: 1592: 1589:Ardipithecus 1588: 1584: 1580: 1578: 1571: 1567: 1563: 1552: 1541: 1539: 1534:Homo sapiens 1532: 1528:Homo erectus 1526: 1520: 1514: 1509: 1503: 1499: 1485: 1470:Homo sapiens 1469: 1461: 1460: 1430: 1415: 1271: 1227: 1205:Ardipithecus 1204: 1123:Earlier apes 1080:Neanderthals 1064:Homo sapiens 1061: 1060: 1047: 1031: 1023: 1015: 1005: 1004: 990: 982: 972: 971: 957: 949: 941: 931: 930: 916: 901: 889:Ardipithecus 886: 885: 871: 863: 849: 836: 823: 813:Sivapithecus 810: 800:Oreopithecus 797: 784: 770: 762: 753: 740: 727: 715: 701: 688: 673: 504:Telanthropus 503: 495: 488:Sinanthropus 487: 479: 475:Dubois, 1894 471: 463: 456:Atlanthropus 455: 451:Dreyer, 1935 447: 422: 421: 414: 404: 394: 384: 374: 366:Homo habilis 364: 354: 344: 336:Homo erectus 334: 324: 312: 307:Homo sapiens 305: 287:Homo sapiens 285: 279:Type species 260: 206:Infraorder: 35: 29: 7642:Vertebrates 7632:Lepidoptera 7605:Brachiopoda 7574:Gastropoda 7551:Pachypodium 7496:Orchidaceae 7150:Grandmother 7105:Shore-based 7066:Aquatic ape 6957:Tam Pa Ling 6852:H. ergaster 6671:P. robustus 5943:Basic Books 4810:H. ergaster 4772:: 126–170. 4281:Homo helmei 4198:10026538759 4045:December 8, 3808:The age of 3629:, found in 3453:Nanjing Man 3441:Lantian Man 3429:Yuanmou Man 3172:hottentotus 3160:aethiopicus 3146:patagonicus 3122:neptunianus 3116:hyperboreus 3038:H. s. ferus 2840:Philippines 2799:or Denisova 2200:Publication 2182:Adult mass 2119:Homo naledi 2104:with Asian 2060:other than 1986:(alongside 1972:KNM-ER 1470 1920:stone tools 1863:, include: 1824:(1995) and 1808:as well as 1704:great races 1531:and modern 1026:H. ergaster 919:Ar. ramidus 904:Ar. kadabba 866:O. praegens 704:Pleistocene 533:This box: 491:Black, 1927 406:Homo naledi 226:Subfamily: 210:Simiiformes 7620:Dermaptera 7615:Scorpiones 7509:Adenanthos 7189:Skin color 7174:Bipedalism 7135:Killer ape 6947:Cro-Magnon 6846:Denisovans 6722:H. habilis 6686:Humans and 6571:A. ramidus 6564:A. kadabba 6363:H. erectus 5805:Syst. Nat. 5786:2018-06-04 5390:: e24231. 4798:H. erectus 4794:H. erectus 4744:0226439631 4347:H. habilis 4285:Lake Eyasi 3882:References 3666:(formerly 3658:(formerly 3656:Peking Man 3552:Confirmed 3465:Peking Man 3387:subspecies 3333:H. sapiens 3325:John Baker 3315:H. sapiens 3240:H. sapiens 3151:Oulotrichi 3140:americanus 3134:columbicus 3077:Leiotrichi 3018:H. s. afer 3002:H. sapiens 2974:H. sapiens 2950:subspecies 2938:H. sapiens 2902:subspecies 2894:Subspecies 2658:950–1,800 2417:1,200–800 2340:1,900–140 2307:1,900–600 2197:Discovery 2116:(2013) or 1992:H. erectus 1988:H. habilis 1962:H. habilis 1950:H. habilis 1914:The genus 1767:" between 1716:(genitive 1682:subspecies 1628:subspecies 1624:phenotypes 1570:split, is 1551:alongside 1540:The genus 1478:zoological 1095:Denisovans 1034:Au. sediba 1008:H. erectus 975:H. habilis 765:Ou. turkae 246:Subtribe: 200:Haplorhini 196:Suborder: 43:Piacenzian 38:("humans") 7647:Lemuridae 7637:Lucanidae 7626:Anopheles 7531:Cactaceae 7490:Narcissus 7475:Liliaceae 7308:Theorists 7275:Timelines 7155:Patriarch 7131:Behavior 7056:Gathering 6984:Ancestors 6729:H. naledi 6664:P. boisei 6635:A. sediba 6051:Routledge 5696:106411053 5546:0305-4403 5477:: 16046. 4706:cite news 3541:Footnotes 3467:) (1970s) 3419:) (1970s) 3178:melaninus 3086:japeticus 3030:East Asia 2980:based on 2934:lectotype 2795:possibly 2728:Indonesia 2661:(extant) 2648:Worldwide 2263:uncertain 2218:uncertain 2164:Lineages 2134:lineages 1980:Alekseyev 1722:Hominidae 993:Au. garhi 230:Homininae 220:Hominidae 156:Kingdom: 150:Eukaryota 7748:Category 7692:Diatomea 7686:Protista 7678:Protists 7600:Bivalvia 7595:Conoidea 7569:Animalia 7502:Triticum 7445:Bacteria 7421:Taxonomy 7354:Category 7209:Language 7179:Skeleton 6874:H. longi 6628:A. garhi 6465:Hominins 6460:Taxonomy 6331:14745010 6278:19805257 6260:40485013 6209:26693109 6201:12802315 5985:62265494 5959:(2005). 5937:(2014). 5921:(2013), 5889:(1982), 5725:10 April 5710:Zimmer C 5688:30971845 5638:25625212 5606:: 6037. 5509:28675384 5416:28483040 5365:34557770 5316:34557772 5203:21738710 5163:PLOS ONE 5144:21646521 5065:Prins HE 5050:20466364 5015:43455561 5007:25739409 4945:Springer 4939:(eds.). 4887:20435482 4879:24136960 4788:14666536 4676:18380861 4625:12794185 4556:12110880 4485:10999270 4434:87482930 4403:10213682 4341:25739410 4291:(1942), 4259:12794185 4175:33066198 4167:17476243 4098:27874171 4005:27298468 3934:26693109 3926:12802315 3664:Java Man 3524:See also 3489:Solo Man 3455:) (1993) 3417:Java Man 3262:infrasp. 3236:synonyms 3193:(etc.). 3191:spelaeus 3098:iranicus 3092:arabicus 3022:Americas 2936:of both 2880:2 sites 2820:2008(?) 2764:140–120 2532:309–138 2459:600–300 2434:2 sites 2396:700–850 2379:African 2282:2 sites 2236:510–660 2226:Tanzania 2202:of name 2176:Habitat 2007:Hominini 1958:LD 350-1 1936:(Leakey 1928:Tanzania 1800:(1925). 1734:Hominini 1724:for the 1670:hominids 1644:Hominini 1559:diverged 1549:Hominini 717:Hominini 691:Pliocene 441:Synonyms 431:Synonyms 298:Species 267:Linnaeus 250:Hominina 240:Hominini 216:Family: 190:Primates 180:Mammalia 170:Chordata 166:Phylum: 160:Animalia 146:Domain: 7734:Viruses 7610:Araneae 7561:Animals 7544:Drosera 7516:Banksia 7462:Plantae 7366:Commons 7318:Fossils 7184:Muscles 7095:Cooking 7051:Hunting 6528:Orrorin 6373:. 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The " 1718:hominis 1678:species 1634:History 1394:← 1372:← 1350:← 1337:cooking 1326:← 1306:← 1286:← 1262:← 1242:← 1219:← 1196:← 1176:← 1156:← 1136:← 1116:← 852:Orrorin 677:Miocene 660:– 650:– 640:– 630:– 620:– 610:– 600:– 590:– 580:– 570:– 256:Genus: 236:Tribe: 186:Order: 176:Class: 121:↓ 47:Present 7537:Citrus 7481:Tulipa 7469:Allium 7454:Plants 7423:of ... 7301:Others 7204:Speech 7166:Topics 7111:Drugs 7037:Models 6387:  6329:  6322:337021 6319:  6276:  6266:  6258:  6207:  6199:  6172:Nature 6100:  6057:  6008:  5983:  5973:  5891:p. 141 5694:  5686:  5660:Nature 5636:  5626:  5579:  5544:  5507:  5497:  5436:Nature 5414:  5404:  5363:  5353:  5314:  5304:  5201:  5191:  5142:  5132:  5081:  5048:  5013:  5005:  4959:  4885:  4877:  4827:Nature 4786:  4741:  4674:  4664:  4623:  4616:164648 4613:  4562:  4554:  4528:Nature 4508:  4483:  4473:  4432:  4418:Nature 4401:  4339:  4257:  4250:164648 4247:  4196:  4173:  4165:  4137:Nature 4096:  4086:  4003:  3993:  3932:  3924:  3897:Nature 3662:) and 3511:(1991) 3358:et al. 3073:racial 2914:, and 2836:c. 67 2807:Taiwan 2768:Israel 2582:1,300 2574:Zambia 2563:early 2542:1,420 2511:1,000 2452:early 2431:1,000 2344:Africa 2267:1,900 1938:et al. 1848:, and 1726:family 1706:. 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Index

Homo sapiens sapiens
Piacenzian
Present
Ma
Preęž’
ęž’
O
S
D
C
P
T
J
K
Pg
N
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Mammalia
Primates
Haplorhini
Simiiformes
Hominidae
Homininae
Hominini
Hominina
Homo

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