125:
592:
102:
289:
were considered a sister group to all other archostematan beetles. However, other authors have considered their placement within beetles as a whole uncertain, due to their mix characteristics of typical
Archostemata, as well as
348:
542:
327:
2627:
2516:
311:
259:
218:
111:
2601:
2640:
2645:
629:
484:"Early evolution of Cupedidae revealed by a mid-Cretaceous reticulated beetle from Myanmar (Coleoptera: Archostemata)"
573:
482:
Li, Yan-Da; Liu, Zhen-Hua; Jarzembowski, Edmund A.; Yin, Zi-Wei; Huang, Di-Ying; Cai, Chen-Yang (October 2019).
441:
124:
2699:
622:
265:
2671:
2531:
397:
2694:
2666:
2658:
2632:
1768:
495:
456:
8:
2569:
1987:
615:
596:
483:
499:
460:
277:, and assigned to this family. Their placement is uncertain, but are usually considered
602:
565:
519:
235:
119:
2653:
2588:
2256:
1673:
561:
523:
511:
335:
569:
2353:
2132:
2127:
2050:
1808:
824:
557:
503:
464:
352:
442:"The most mysterious beetles: Jurassic Jurodidae (Insecta: Coleoptera) from China"
2593:
2554:
2042:
1995:
1840:
1824:
1500:
1260:
1229:
1144:
1062:
953:
921:
2445:
2396:
2251:
2058:
1726:
1710:
1649:
1544:
1515:
1443:
1407:
1391:
1307:
1252:
1157:
1054:
903:
865:
845:
741:
2688:
2427:
2231:
2222:
1963:
1934:
1926:
1918:
1892:
1867:
1763:
1750:
1689:
1607:
1510:
1505:
1386:
1320:
1291:
1275:
1239:
1209:
1162:
1116:
1083:
999:
991:
962:
800:
607:
515:
468:
406:
225:
59:
440:
Yan, Evgeny V.; Wang, Bo; Ponomarenko, Alexander G.; Zhang, Haichun (2014).
2404:
2328:
2074:
2066:
2021:
2005:
1955:
1876:
1832:
1784:
1702:
1615:
1484:
1471:
1451:
1412:
1312:
1299:
1270:
1204:
1192:
1088:
1045:
1028:
1020:
970:
808:
778:
709:
698:
682:
278:
270:
186:
2580:
263:, known from the Middle-Late Jurassic of Asia. In 1996, a living species,
2614:
2563:
2490:
2461:
2435:
2422:
2362:
2320:
2304:
2296:
2288:
2272:
2264:
2200:
2158:
2150:
2119:
2034:
2013:
1979:
1971:
1884:
1816:
1800:
1792:
1755:
1742:
1718:
1636:
1628:
1578:
1570:
1528:
1492:
1463:
1435:
1430:
1346:
1341:
1333:
1283:
1219:
1214:
1128:
1093:
1078:
1070:
986:
978:
795:
34:
2417:
2378:
2370:
2312:
2175:
2137:
2090:
2082:
1902:
1850:
1734:
1665:
1536:
1520:
1476:
1417:
1381:
1361:
1325:
1234:
1224:
1098:
832:
816:
507:
79:
44:
2619:
2482:
2474:
2466:
2453:
2409:
2208:
2183:
2111:
1939:
1910:
1594:
1425:
1399:
1351:
1265:
1167:
1136:
1108:
941:
916:
911:
892:
857:
746:
721:
670:
658:
373:
356:
291:
156:
136:
84:
2606:
2525:
2548:
2386:
2336:
2280:
2239:
2188:
1845:
1776:
1657:
1369:
1175:
1149:
1007:
840:
787:
766:
676:
591:
369:
331:
295:
74:
69:
54:
49:
39:
2103:
2095:
1947:
1681:
1602:
1586:
274:
89:
64:
1694:
1554:
1187:
664:
652:
639:
254:
176:
166:
146:
550:
Journal of
Zoological Systematics and Evolutionary Research
543:"Ocellar atavism in Coleoptera: plesiomorphy or apomorphy?"
101:
1010:(furniture beetles, death watch beetles, spider beetles)
439:
481:
307:
257:
that was originally described for the extinct genus
214:
2686:
1887:(jewel beetles, or metallic wood-boring beetles)
1610:(true weevils, bark beetles, ambrosia beetles)
777:
637:
623:
540:
1853:(ironclad beetles, cylindrical bark beetles)
630:
616:
100:
2517:List of subgroups of the order Coleoptera
1597:(straight snout weevils, New York weevil)
696:
2024:(cicada beetle, cicada parasite beetles)
384:Yan, Wang, Ponomarenko & Zhang, 2014
367:Yan, Wang, Ponomarenko & Zhang, 2014
2687:
2114:(false fireflies, long-lipped beetles)
939:
890:
541:Leschen, R.A.B.; Beutel, R.G. (2004).
2530:
2529:
938:
889:
764:
763:
695:
611:
2140:(false metallic wood-boring beetles)
13:
534:
14:
2711:
584:
1751:Perimylopidae, or Promecheilidae
1336:(minute brown scavenger beetles)
590:
562:10.1046/j.0947-5745.2003.00241.x
123:
32:
1966:(variegated mud-loving beetles)
475:
433:
301:
1:
1795:(narrow-waisted bark beetles)
1487:(parasitic flat bark beetles)
1438:(short-winged flower beetles)
1294:(lady beetles, or God's cows)
1031:(tooth-necked fungus beetles)
426:
2307:(sand-loving scarab beetles)
1697:(minute tree-fungus beetles)
1523:(silvanid flat bark beetles)
7:
2456:(primitive carrion beetles)
2412:(minute mud-loving beetles)
2275:(bumble bee scarab beetles)
2045:(soft-bodied plant beetles)
2016:(soft bodied plant beetles)
819:(predaceous diving beetles)
10:
2716:
2291:(scavenger scarab beetles)
2283:(enigmatic scarab beetles)
1456:Lamingtonium binnaberrense
1170:(soft-wing flower beetles)
386:Dahougou, China, Callovian
372:, China, Middle Jurassic (
273:mountains in southeastern
2538:
2514:
2444:
2430:(water scavenger beetles)
2395:
2361:
2352:
2230:
2221:
2174:
2163:Rhinorhipus tamborinensis
2149:
2033:
2004:
1942:(long-toed water beetles)
1901:
1875:
1866:
1835:(polypore fungus beetles)
1737:(palm and flower beetles)
1721:(tumbling flower beetles)
1668:(ant-like flower beetles)
1648:
1627:
1553:
1446:(lined flat bark beetles)
1420:(pleasing fungus beetles)
1360:
1315:(handsome fungus beetles)
1251:
1107:
1053:
1044:
1019:
961:
952:
948:
934:
899:
885:
860:(burrowing water beetles)
773:
759:
705:
691:
647:
355:, Russia, Late Jurassic (
334:, Russia, Late Jurassic (
241:
234:
211:
206:
120:Scientific classification
118:
108:
99:
23:
2477:(feather-winged beetles)
1811:(false longhorn beetles)
1758:(jugular-horned beetles)
1705:(false darkling beetles)
1676:(cryptic fungus beetles)
1495:(shining flower beetles)
835:(crawling water beetles)
734:Sikhotealinia zhiltzovae
469:10.1016/j.gr.2013.04.002
416:Sikhotealinia zhiltzovae
266:Sikhotealinia zhiltzovae
1958:(forest stream beetles)
1819:(synchroa bark beetles)
1779:(dead log bark beetles)
1771:(fire-coloured beetles)
1660:(ant-like leaf beetles)
1394:(silken fungus beetles)
1328:(well polished beetles)
1302:(minute fungus beetles)
1121:Acanthocnemus nigricans
30:Middle Jurassic-Present
2469:(round fungus beetles)
1803:(false flower beetles)
1787:(wedge-shaped beetles)
1745:(false blister beetle)
1729:(hairy fungus beetles)
1684:(conifer bark beetles)
1581:(leaf-rolling weevils)
1242:(bark-gnawing beetles)
1002:(wounded-tree beetles)
868:(false ground beetles)
790:(trout-stream beetles)
269:was discovered in the
2667:Paleobiology Database
2381:(false clown beetles)
2203:(plate-thigh beetles)
2085:(false click beetles)
1990:(water-penny beetles)
1639:(ship-timber beetles)
1618:(pine flower weevils)
1466:(root-eating beetles)
1286:(minute bark beetles)
1197:Phloiophilus edwardsi
724:(reticulated beetles)
488:Systematic Entomology
400:, Mongolia, Tithonian
391:Jurodes shartegiensis
364:Jurodes daohugouensis
2259:(false stag beetles)
2244:Belohina inexpectata
2106:(net-winged beetles)
2069:(rare click beetles)
1982:(travertine beetles)
1974:(minute mud beetles)
1539:(dry-fungus beetles)
1374:Agapytho foveicollis
1131:(false skin beetles)
994:(Jacobson's beetles)
714:Crowsoniella relicta
2257:Diphyllostomatidae
1589:(primitive weevils)
1402:(flat bark beetles)
1152:(checkered beetles)
1139:(fruitworm beetles)
827:(whirligig beetles)
500:2019SysEn..44..777L
461:2014GondR..25..214Y
349:Glushkovo Formation
2122:(glowworm beetles)
1827:(darkling beetles)
1674:Archeocrypticidae
1531:(palmetto beetles)
1278:(dry bark beetles)
1065:(longhorn beetles)
508:10.1111/syen.12355
2682:
2681:
2654:Open Tree of Life
2532:Taxon identifiers
2523:
2522:
2510:
2509:
2506:
2505:
2502:
2501:
2494:
2486:
2485:(carrion beetles)
2478:
2470:
2457:
2431:
2413:
2382:
2374:
2348:
2347:
2340:
2332:
2324:
2316:
2308:
2300:
2292:
2284:
2276:
2268:
2260:
2247:
2217:
2216:
2204:
2196:
2166:
2141:
2123:
2115:
2107:
2099:
2086:
2078:
2070:
2062:
2061:(soldier beetles)
2054:
2051:Brachypsectridae
2046:
2025:
2017:
1991:
1983:
1975:
1967:
1959:
1951:
1943:
1930:
1922:
1914:
1888:
1862:
1861:
1854:
1836:
1828:
1820:
1812:
1809:Stenotrachelidae
1804:
1796:
1788:
1780:
1772:
1759:
1746:
1738:
1730:
1722:
1714:
1713:(blister beetles)
1706:
1698:
1685:
1677:
1669:
1661:
1640:
1619:
1611:
1598:
1590:
1582:
1574:
1564:
1540:
1532:
1524:
1496:
1488:
1480:
1467:
1459:
1447:
1439:
1421:
1403:
1395:
1377:
1337:
1329:
1316:
1303:
1295:
1287:
1279:
1261:Akalyptoischiidae
1243:
1200:
1183:
1171:
1153:
1140:
1132:
1124:
1074:
1066:
1040:
1039:
1032:
1011:
1003:
995:
982:
974:
930:
929:
907:
881:
880:
877:
876:
869:
861:
853:
836:
828:
820:
812:
804:
791:
755:
754:
737:
725:
717:
449:Gondwana Research
420:
395:
347:
346:Ponomarenko, 1990
328:Ichetuy Formation
326:
325:Ponomarenko, 1985
322:Jurodes ignoramus
317:
248:
247:
243:Sikhotealiniidae
202:
201:Ponomarenko, 1985
16:Family of beetles
2707:
2675:
2674:
2662:
2661:
2649:
2648:
2636:
2635:
2623:
2622:
2610:
2609:
2597:
2596:
2584:
2583:
2574:
2573:
2572:
2559:
2558:
2557:
2527:
2526:
2492:
2484:
2476:
2468:
2455:
2429:
2411:
2380:
2372:
2359:
2358:
2354:Staphyliniformia
2338:
2330:
2322:
2314:
2306:
2298:
2290:
2282:
2274:
2266:
2258:
2241:
2228:
2227:
2202:
2193:Declinia relicta
2190:
2160:
2139:
2133:Sinopyrophoridae
2128:Rhagophthalmidae
2121:
2113:
2105:
2097:
2084:
2076:
2068:
2060:
2052:
2044:
2043:Artematopodidae
2023:
2015:
1989:
1981:
1973:
1965:
1957:
1950:(riffle beetles)
1949:
1941:
1929:(turtle beetles)
1928:
1920:
1912:
1886:
1873:
1872:
1852:
1834:
1826:
1818:
1810:
1802:
1794:
1786:
1778:
1770:
1757:
1744:
1736:
1728:
1720:
1712:
1704:
1696:
1683:
1675:
1667:
1659:
1638:
1617:
1609:
1596:
1588:
1580:
1573:(fungus weevils)
1572:
1563:
1562:
1558:
1545:Tasmosalpingidae
1538:
1530:
1522:
1494:
1486:
1478:
1465:
1453:
1445:
1437:
1419:
1401:
1393:
1371:
1335:
1327:
1314:
1301:
1293:
1285:
1277:
1241:
1194:
1177:
1169:
1151:
1138:
1130:
1118:
1117:Acanthocnemidae
1072:
1064:
1051:
1050:
1030:
1009:
1001:
993:
980:
972:
959:
958:
950:
949:
936:
935:
905:
887:
886:
867:
859:
847:
834:
826:
818:
810:
803:(ground beetles)
802:
789:
775:
774:
761:
760:
731:
723:
711:
710:Crowsoniellidae
693:
692:
632:
625:
618:
609:
608:
595:Data related to
594:
580:
578:
572:. Archived from
547:
528:
527:
479:
473:
472:
446:
437:
418:
412:
393:
385:
381:Jurodes pygmaeus
368:
345:
324:
316:Ponomarenko 1985
315:
309:
281:. In one study,
216:
200:
128:
127:
104:
94:
31:
27:Temporal range:
21:
20:
2715:
2714:
2710:
2709:
2708:
2706:
2705:
2704:
2700:Beetle families
2685:
2684:
2683:
2678:
2670:
2665:
2657:
2652:
2644:
2639:
2631:
2626:
2618:
2613:
2605:
2600:
2592:
2587:
2579:
2577:
2568:
2567:
2562:
2553:
2552:
2547:
2534:
2524:
2519:
2498:
2440:
2391:
2373:(clown beetles)
2344:
2315:(betsy beetles)
2223:Scarabaeiformia
2213:
2170:
2145:
2077:(click beetles)
2053:(Texas beetles)
2029:
2000:
1996:Ptilodactylidae
1921:(cedar beetles)
1897:
1858:
1841:Trictenotomidae
1727:Mycetophagidae
1644:
1623:
1560:
1559:
1557:
1549:
1501:Phloeostichidae
1452:Lamingtoniidae
1444:Laemophloeidae
1392:Cryptophagidae
1356:
1247:
1230:Thanerocleridae
1193:Phloiophilidae
1180:Metaxina ornata
1145:Chaetosomatidae
1103:
1036:
1015:
973:(auger beetles)
954:Bostrichiformia
944:
926:
922:Torridincolidae
906:(skiff beetles)
904:Hydroscaphidae
895:
873:
866:Trachypachidae
811:(tiger beetles)
769:
751:
701:
687:
643:
636:
587:
576:
545:
537:
535:Further reading
532:
531:
480:
476:
444:
438:
434:
429:
410:
383:
366:
304:
253:is a family of
199:
122:
95:
93:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
37:
29:
28:
25:
17:
12:
11:
5:
2713:
2703:
2702:
2697:
2680:
2679:
2677:
2676:
2663:
2650:
2637:
2624:
2611:
2598:
2585:
2575:
2560:
2544:
2542:
2536:
2535:
2521:
2520:
2515:
2512:
2511:
2508:
2507:
2504:
2503:
2500:
2499:
2497:
2496:
2493:(rove beetles)
2491:Staphylinidae
2488:
2480:
2472:
2464:
2459:
2450:
2448:
2446:Staphylinoidea
2442:
2441:
2439:
2438:
2433:
2428:Hydrophilidae
2425:
2420:
2415:
2407:
2401:
2399:
2397:Hydrophiloidea
2393:
2392:
2390:
2389:
2384:
2376:
2367:
2365:
2356:
2350:
2349:
2346:
2345:
2343:
2342:
2339:(hide beetles)
2334:
2326:
2323:(rain beetles)
2318:
2310:
2302:
2299:(stag beetles)
2294:
2286:
2278:
2270:
2262:
2254:
2252:Bolboceratidae
2249:
2236:
2234:
2225:
2219:
2218:
2215:
2214:
2212:
2211:
2206:
2198:
2186:
2180:
2178:
2172:
2171:
2169:
2168:
2159:Rhinorhipidae
2155:
2153:
2151:Rhinorhipoidea
2147:
2146:
2144:
2143:
2135:
2130:
2125:
2117:
2109:
2101:
2093:
2088:
2080:
2072:
2064:
2056:
2048:
2039:
2037:
2031:
2030:
2028:
2027:
2019:
2010:
2008:
2002:
2001:
1999:
1998:
1993:
1985:
1977:
1969:
1964:Heteroceridae
1961:
1953:
1945:
1937:
1932:
1927:Chelonariidae
1924:
1919:Callirhipidae
1916:
1913:(pill beetles)
1907:
1905:
1899:
1898:
1896:
1895:
1890:
1881:
1879:
1870:
1864:
1863:
1860:
1859:
1857:
1856:
1848:
1843:
1838:
1830:
1825:Tenebrionidae
1822:
1814:
1806:
1798:
1790:
1782:
1774:
1766:
1761:
1753:
1748:
1740:
1732:
1724:
1716:
1708:
1700:
1692:
1687:
1679:
1671:
1663:
1654:
1652:
1650:Tenebrionoidea
1646:
1645:
1643:
1642:
1633:
1631:
1625:
1624:
1622:
1621:
1613:
1608:Curculionidae
1605:
1600:
1592:
1584:
1576:
1567:
1565:
1555:Curculionoidea
1551:
1550:
1548:
1547:
1542:
1534:
1526:
1518:
1516:Protocucujidae
1513:
1508:
1503:
1498:
1490:
1482:
1474:
1469:
1461:
1449:
1441:
1433:
1428:
1423:
1415:
1410:
1408:Cybocephalidae
1405:
1397:
1389:
1384:
1379:
1366:
1364:
1358:
1357:
1355:
1354:
1349:
1344:
1339:
1331:
1323:
1318:
1310:
1308:Discolomatidae
1305:
1297:
1292:Coccinellidae
1289:
1281:
1276:Bothrideridae
1273:
1268:
1263:
1257:
1255:
1253:Coccinelloidea
1249:
1248:
1246:
1245:
1240:Trogossitidae
1237:
1232:
1227:
1222:
1217:
1212:
1207:
1202:
1190:
1185:
1173:
1165:
1160:
1158:Lophocateridae
1155:
1147:
1142:
1134:
1126:
1113:
1111:
1105:
1104:
1102:
1101:
1096:
1091:
1086:
1081:
1076:
1073:(leaf beetles)
1071:Chrysomelidae
1068:
1059:
1057:
1055:Chrysomeloidea
1048:
1042:
1041:
1038:
1037:
1035:
1034:
1025:
1023:
1017:
1016:
1014:
1013:
1005:
1000:Nosodendridae
997:
992:Jacobsoniidae
989:
984:
981:(skin beetles)
976:
967:
965:
956:
946:
945:
932:
931:
928:
927:
925:
924:
919:
914:
909:
900:
897:
896:
883:
882:
879:
878:
875:
874:
872:
871:
863:
855:
850:Meru phyllisae
843:
838:
830:
822:
814:
806:
798:
793:
784:
782:
771:
770:
757:
756:
753:
752:
750:
749:
744:
742:Micromalthidae
739:
727:
719:
706:
703:
702:
689:
688:
686:
685:
679:
673:
667:
661:
655:
648:
645:
644:
635:
634:
627:
620:
612:
606:
605:
600:
599:at Wikispecies
586:
585:External links
583:
582:
581:
579:on 2019-03-02.
536:
533:
530:
529:
494:(4): 777β786.
474:
455:(1): 214β225.
431:
430:
428:
425:
424:
423:
422:
421:
403:
402:
401:
387:
377:
360:
339:
303:
300:
279:archostematans
246:
245:
239:
238:
232:
231:
230:
229:
222:
209:
208:
204:
203:
194:
190:
189:
184:
180:
179:
174:
170:
169:
164:
160:
159:
154:
150:
149:
144:
140:
139:
134:
130:
129:
116:
115:
106:
105:
97:
96:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
33:
26:
15:
9:
6:
4:
3:
2:
2712:
2701:
2698:
2696:
2693:
2692:
2690:
2673:
2668:
2664:
2660:
2655:
2651:
2647:
2642:
2638:
2634:
2629:
2625:
2621:
2616:
2612:
2608:
2603:
2599:
2595:
2590:
2586:
2582:
2576:
2571:
2565:
2561:
2556:
2550:
2546:
2545:
2543:
2541:
2537:
2533:
2528:
2518:
2513:
2495:
2489:
2487:
2481:
2479:
2473:
2471:
2465:
2463:
2460:
2458:
2452:
2451:
2449:
2447:
2443:
2437:
2434:
2432:
2426:
2424:
2421:
2419:
2416:
2414:
2408:
2406:
2403:
2402:
2400:
2398:
2394:
2388:
2385:
2383:
2379:Sphaeritidae
2377:
2375:
2369:
2368:
2366:
2364:
2360:
2357:
2355:
2351:
2341:
2335:
2333:
2329:Scarabaeidae
2327:
2325:
2319:
2317:
2311:
2309:
2303:
2301:
2295:
2293:
2287:
2285:
2279:
2277:
2271:
2269:
2267:(dor beetles)
2263:
2261:
2255:
2253:
2250:
2248:
2245:
2238:
2237:
2235:
2233:
2232:Scarabaeoidea
2229:
2226:
2224:
2220:
2210:
2207:
2205:
2199:
2197:
2194:
2187:
2185:
2182:
2181:
2179:
2177:
2173:
2167:
2164:
2157:
2156:
2154:
2152:
2148:
2142:
2136:
2134:
2131:
2129:
2126:
2124:
2118:
2116:
2110:
2108:
2102:
2100:
2094:
2092:
2089:
2087:
2081:
2079:
2073:
2071:
2067:Cerophytidae
2065:
2063:
2057:
2055:
2049:
2047:
2041:
2040:
2038:
2036:
2032:
2026:
2022:Rhipiceridae
2020:
2018:
2012:
2011:
2009:
2007:
2003:
1997:
1994:
1992:
1986:
1984:
1978:
1976:
1970:
1968:
1962:
1960:
1956:Eulichadidae
1954:
1952:
1946:
1944:
1938:
1936:
1935:Cneoglossidae
1933:
1931:
1925:
1923:
1917:
1915:
1909:
1908:
1906:
1904:
1900:
1894:
1893:Schizopodidae
1891:
1889:
1883:
1882:
1880:
1878:
1874:
1871:
1869:
1868:Elateriformia
1865:
1855:
1849:
1847:
1844:
1842:
1839:
1837:
1833:Tetratomidae
1831:
1829:
1823:
1821:
1815:
1813:
1807:
1805:
1799:
1797:
1791:
1789:
1785:Ripiphoridae
1783:
1781:
1775:
1773:
1769:Pyrochroidae
1767:
1765:
1764:Pterogeniidae
1762:
1760:
1754:
1752:
1749:
1747:
1741:
1739:
1733:
1731:
1725:
1723:
1717:
1715:
1709:
1707:
1703:Melandryidae
1701:
1699:
1693:
1691:
1690:Chalcodryidae
1688:
1686:
1680:
1678:
1672:
1670:
1664:
1662:
1656:
1655:
1653:
1651:
1647:
1641:
1635:
1634:
1632:
1630:
1626:
1620:
1616:Nemonychidae
1614:
1612:
1606:
1604:
1601:
1599:
1593:
1591:
1585:
1583:
1577:
1575:
1569:
1568:
1566:
1556:
1552:
1546:
1543:
1541:
1535:
1533:
1527:
1525:
1519:
1517:
1514:
1512:
1511:Propalticidae
1509:
1507:
1506:Priasilphidae
1504:
1502:
1499:
1497:
1491:
1489:
1485:Passandridae
1483:
1481:
1479:(sap beetles)
1475:
1473:
1470:
1468:
1462:
1460:
1457:
1450:
1448:
1442:
1440:
1434:
1432:
1429:
1427:
1424:
1422:
1416:
1414:
1411:
1409:
1406:
1404:
1398:
1396:
1390:
1388:
1387:Cavognathidae
1385:
1383:
1380:
1378:
1375:
1368:
1367:
1365:
1363:
1359:
1353:
1350:
1348:
1345:
1343:
1340:
1338:
1332:
1330:
1324:
1322:
1321:Eupsilobiidae
1319:
1317:
1313:Endomychidae
1311:
1309:
1306:
1304:
1300:Corylophidae
1298:
1296:
1290:
1288:
1282:
1280:
1274:
1272:
1269:
1267:
1264:
1262:
1259:
1258:
1256:
1254:
1250:
1244:
1238:
1236:
1233:
1231:
1228:
1226:
1223:
1221:
1218:
1216:
1215:Protopeltidae
1213:
1211:
1210:Prionoceridae
1208:
1206:
1203:
1201:
1198:
1191:
1189:
1186:
1184:
1181:
1174:
1172:
1166:
1164:
1163:Mauroniscidae
1161:
1159:
1156:
1154:
1148:
1146:
1143:
1141:
1135:
1133:
1127:
1125:
1122:
1115:
1114:
1112:
1110:
1106:
1100:
1097:
1095:
1092:
1090:
1087:
1085:
1084:Megalopodidae
1082:
1080:
1077:
1075:
1069:
1067:
1063:Cerambycidae
1061:
1060:
1058:
1056:
1052:
1049:
1047:
1043:
1033:
1029:Derodontidae
1027:
1026:
1024:
1022:
1021:Derodontoidea
1018:
1012:
1006:
1004:
998:
996:
990:
988:
987:Endecatomidae
985:
983:
977:
975:
971:Bostrichidae
969:
968:
966:
964:
963:Bostrichoidea
960:
957:
955:
951:
947:
943:
937:
933:
923:
920:
918:
917:Sphaeriusidae
915:
913:
910:
908:
902:
901:
898:
894:
888:
884:
870:
864:
862:
856:
854:
851:
844:
842:
839:
837:
831:
829:
823:
821:
815:
813:
809:Cicindelidae
807:
805:
799:
797:
794:
792:
786:
785:
783:
780:
776:
772:
768:
762:
758:
748:
745:
743:
740:
738:
735:
728:
726:
720:
718:
715:
708:
707:
704:
700:
694:
690:
684:
680:
678:
674:
672:
668:
666:
662:
660:
656:
654:
650:
649:
646:
641:
633:
628:
626:
621:
619:
614:
613:
610:
604:
601:
598:
593:
589:
588:
575:
571:
567:
563:
559:
555:
551:
544:
539:
538:
525:
521:
517:
513:
509:
505:
501:
497:
493:
489:
485:
478:
470:
466:
462:
458:
454:
450:
443:
436:
432:
417:
414:
413:
409:
408:
407:Sikhotealinia
404:
399:
392:
388:
382:
378:
375:
371:
365:
361:
358:
354:
350:
344:
343:Jurodes minor
340:
337:
333:
329:
323:
319:
318:
314:
313:
306:
305:
299:
297:
293:
288:
284:
283:Sikhotealinia
280:
276:
272:
268:
267:
262:
261:
256:
252:
244:
240:
237:
233:
228:
227:
226:Sikhotealinia
223:
221:
220:
213:
212:
210:
205:
198:
195:
192:
191:
188:
185:
182:
181:
178:
175:
172:
171:
168:
165:
162:
161:
158:
155:
152:
151:
148:
145:
142:
141:
138:
135:
132:
131:
126:
121:
117:
114:
113:
107:
103:
98:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
36:
22:
19:
2695:Archostemata
2539:
2418:Helophoridae
2410:Georissidae
2405:Epimetopidae
2321:Pleocomidae
2305:Ochodaeidae
2289:Hybosoridae
2273:Glaphyridae
2265:Geotrupidae
2243:
2240:Belohinidae
2201:Eucinetidae
2192:
2189:Decliniidae
2162:
2120:Phengodidae
2059:Cantharidae
2014:Dascillidae
2006:Dascilloidea
1988:Psephenidae
1980:Lutrochidae
1972:Limnichidae
1885:Buprestidae
1877:Buprestoidea
1817:Synchroidae
1801:Scraptiidae
1793:Salpingidae
1756:Prostomidae
1743:Oedemeridae
1719:Mordellidae
1637:Lymexylidae
1629:Lymexyloidea
1579:Attelabidae
1571:Anthribidae
1529:Smicripidae
1493:Phalacridae
1477:Nitidulidae
1472:Myraboliidae
1464:Monotomidae
1455:
1436:Kateretidae
1413:Cyclaxyridae
1373:
1370:Agapythidae
1334:Latridiidae
1284:Cerylonidae
1271:Anamorphidae
1205:Phycosecidae
1196:
1179:
1176:Metaxinidae
1129:Biphyllidae
1120:
1089:Orsodacnidae
1046:Cucujiformia
979:Dermestidae
849:
788:Amphizoidae
733:
729:
713:
699:Archostemata
683:Holometabola
681:Superorder:
675:Infraclass:
603:Illustration
574:the original
556:(1): 63β69.
553:
549:
491:
487:
477:
452:
448:
435:
415:
405:
390:
380:
363:
342:
321:
310:
286:
282:
271:Sikhote-Alin
264:
258:
250:
249:
242:
224:
217:
196:
187:Archostemata
110:
18:
2615:iNaturalist
2564:Wikispecies
2462:Hydraenidae
2436:Spercheidae
2423:Hydrochidae
2387:Synteliidae
2371:Histeridae
2363:Histeroidea
2313:Passalidae
2281:Glaresidae
2138:Throscidae
2098:(fireflies)
2096:Lampyridae
2083:Eucnemidae
2075:Elateridae
2035:Elateroidea
1851:Zopheridae
1735:Mycteridae
1666:Anthicidae
1537:Sphindidae
1521:Silvanidae
1431:Hobartiidae
1418:Erotylidae
1347:Mycetaeidae
1342:Murmidiidae
1326:Euxestidae
1220:Rentoniidae
1094:Oxypeltidae
1079:Disteniidae
912:Lepiceridae
841:Hygrobiidae
833:Haliplidae
817:Dytiscidae
796:Aspidytidae
419:Lafer, 1996
411:Lafer, 1996
353:Zabaykalsky
302:Subdivision
2689:Categories
2483:Silphidae
2475:Ptiliidae
2467:Leiodidae
2454:Agyrtidae
2297:Lucanidae
2176:Scirtoidea
2112:Omethidae
2091:Jurasaidae
1940:Dryopidae
1911:Byrrhidae
1903:Byrrhoidea
1595:Brentidae
1400:Cucujidae
1382:Boganiidae
1362:Cucujoidea
1235:Thymalidae
1225:Rhadalidae
1168:Melyridae
1137:Byturidae
1099:Vesperidae
1008:Ptiniidae
858:Noteridae
825:Gyrinidae
801:Carabidae
730:Jurodidae
722:Cupedidae
669:Subclass:
659:Arthropoda
640:Coleoptera
427:References
183:Suborder:
177:Coleoptera
157:Arthropoda
109:Fossil of
2570:Jurodidae
2555:Q20735711
2540:Jurodidae
2337:Trogidae
2331:(scarabs)
2209:Scirtidae
2184:Clambidae
1777:Pythidae
1711:Meloidae
1658:Aderidae
1561:(weevils)
1426:Helotidae
1352:Teredidae
1266:Alexiidae
1150:Cleridae
1109:Cleroidea
942:Polyphaga
940:Suborder
893:Myxophaga
891:Suborder
846:Meruidae
765:Suborder
747:Ommatidae
697:Suborder
671:Pterygota
651:Kingdom:
597:Jurodidae
524:202852115
516:0307-6970
398:Shar Teeg
374:Callovian
357:Tithonian
336:Oxfordian
292:Polyphaga
251:Jurodidae
197:Jurodidae
143:Kingdom:
137:Eukaryota
24:Jurodidae
2578:BioLib:
2549:Wikidata
2104:Lycidae
1948:Elmidae
1846:Ulodidae
1682:Boridae
1587:Belidae
1188:Peltidae
781:families
767:Adephaga
677:Neoptera
657:Phylum:
653:Animalia
642:families
570:56327166
394:Yan 2014
370:Dahougou
332:Buryatia
296:Adephaga
236:Synonyms
193:Family:
153:Phylum:
147:Animalia
133:Domain:
2659:4662599
1695:Ciidae
1603:Caridae
665:Insecta
663:Class:
638:Extant
496:Bibcode
457:Bibcode
312:Jurodes
287:Jurodes
275:Siberia
260:Jurodes
255:beetles
219:Jurodes
207:Genera
173:Order:
167:Insecta
163:Class:
112:Jurodes
2672:212812
2646:678384
2633:102985
2620:575127
2581:238033
779:Extant
568:
522:
514:
2628:IRMNG
577:(PDF)
566:S2CID
546:(PDF)
520:S2CID
445:(PDF)
2641:ITIS
2607:7831
2602:GBIF
512:ISSN
294:and
285:and
35:Preκ
2594:BJQ
2589:CoL
558:doi
504:doi
465:doi
2691::
2669::
2656::
2643::
2630::
2617::
2604::
2591::
2566::
2551::
564:.
554:42
552:.
548:.
518:.
510:.
502:.
492:44
490:.
486:.
463:.
453:25
451:.
447:.
396:,
351:,
330:,
298:.
85:Pg
2246:)
2242:(
2195:)
2191:(
2165:)
2161:(
1458:)
1454:(
1376:)
1372:(
1199:)
1195:(
1182:)
1178:(
1123:)
1119:(
852:)
848:(
736:)
732:(
716:)
712:(
631:e
624:t
617:v
560::
526:.
506::
498::
471:.
467::
459::
389:β
379:β
376:)
362:β
359:)
341:β
338:)
320:β
308:β
215:β
90:N
80:K
75:J
70:T
65:P
60:C
55:D
50:S
45:O
40:κ
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
