Knowledge

Nedoceratops

Source đź“ť

849:, may be the result of pathology, injury, and/or deformation of the skull after burial. Two of the features that have been used to diagnose Nedoceratops - the position of the squamosal, and the upright brow horns- are seen on one side of the skull, but not the other. This, along with the fact that the entire skull can be seen to be twisted when seen head-on, have been used to argue that these features result from post-mortem distortion of the fossil, rather than reflecting the anatomy of the animal during life. Furthermore, the presence of numerous holes in the frill suggests pathology due to injury or illness, the supposedly unique 'parietal fenestrae' may therefore be the result of an injury. Tanke & Farke (2007) noted that the supposed parietal fenestra had an irregular shape with swollen margins and an irregularly vascularized texture. This is similar to a parietal hole, also interpreted as the result of an injury by Marshall & Barreto (2001), in a specimen of 450: 2167: 2173: 669: 2723: 136: 807: 2753: 2729: 621: 1455: 118: 825:
is the remarkably short, rounded nasal "horn". Scanella and Horner proposed that the nasal horn of the USNM 2412 skull could have been lost when the animal was alive or when it became fossilized. However, it has been noted the horns of ceratopsids show a great deal of variation between age groups and
651:
proper is that it has a solid, unperforated fill). However, at least some of these holes show evidence that they are the result of injury or disease. The nasal horn of this specimen is low and rounded, compared with the larger, pointed nose horns of typical
1216:"New insights into chasmosaurine (Dinosauria: Ceratopsidae) skulls from the Upper Cretaceous (Campanian) of Alberta, and an update on the distribution of accessory frill fenestrae in Chasmosaurinae" 803:
occupying variable positions with respect to the other two but generally outside the range of variation, concluding that "the size of USNM 2412 is a plausible intermediate, but the shape is not."
529:
paper had been written by Hatcher, and Lull had only contributed the name and published the paper after Hatcher's death, Lull was not quite as convinced of the distinctiveness of
473: 2879: 2833: 2820: 830:
have a similar nasal horn shape. In most features- the short, saddle-shape frill, the s-shaped snout- the animal closely resembles
2914: 2884: 871: 2919: 1509: 2899: 1488: 2166: 811: 459: 2172: 2894: 2746: 1214:
Campbell, James A.; Ryan, Michael J.; Schröder-Adams, Claudia J.; Evans, David C.; Holmes, Robert B. (2018-07-03).
783:. In 2012, Farke proposed a counterargument, and suggested that the bone surface texture and shape of the horns of 643:, but on closer examination, it differs: specifically, the brow horns stand almost vertically compared to typical 514:
to complete the monograph in 1905, publishing Hatcher's description of a skull separately and giving it the name
2909: 2904: 716:
Period, which ended roughly 66 million years ago. All ceratopsians became extinct at the end of this era.
2722: 135: 449: 779:). These authors suggested that all three "genera" actually represent different aged individuals of 1579: 1481: 2874: 1697: 2851: 1278:
Maiorino, Leonardo; Farke, Andrew A.; Kotsakis, Tassos; Piras, Paolo; Butler, Richard J (2013).
656:
specimens, though this feature appears to be within the known range of individual variation for
2811: 2773: 2838: 2846: 2756: 1606: 1002: 971: 853:. The frill is only preserved on one side, which makes it difficult to test this hypothesis. 787:
indicate an "old adult". A follow-up study by Leonardo Maiorino and colleagues in 2013 using
311: 2728: 2889: 1420: 1361: 1295: 1177: 1014: 924: 337: 288: 8: 1862: 1474: 1040: 1424: 1365: 1299: 1181: 1018: 928: 765:(parietal bone) that had "incipient" openings (contrasting with no openings in subadult 2582: 2532: 1964: 1384: 1345: 1318: 1279: 1250: 1215: 1193: 1089: 947: 908: 511: 345: 130: 2665: 2603: 2566: 2040: 1927: 1389: 1323: 1255: 1237: 952: 499: 1277: 1197: 1093: 579: 377: 2695: 2493: 2318: 2020: 1715: 1460: 1428: 1379: 1369: 1313: 1303: 1245: 1227: 1185: 1081: 1022: 983: 942: 932: 624: 605: 1432: 2589: 2541: 2500: 2436: 2339: 2290: 1992: 1916: 1871: 1745: 1731: 1690: 1564: 1548: 1374: 1308: 1189: 937: 469: 412: 409: 406: 30: 2596: 2555: 2486: 2473: 2444: 2429: 2399: 2360: 2346: 2263: 2224: 2103: 2013: 1978: 1971: 1851: 1801: 1724: 1704: 742: 668: 591: 435:
a valid, distinct taxon, while others consider it to be an unusual specimen of
428: 267: 254: 212: 1123: 1026: 987: 635:
The nearly complete skull known as USNM 2412 is the only fossil attributed to
612:, means "horned face". It was named in reference to its lack of a nasal horn. 574:(Förster, 1868), Andrey Sergeevich Ukrainsky gave the animal its current name 2868: 2796: 2674: 2653: 2623: 2614: 2514: 2408: 2353: 2332: 2304: 2297: 2283: 2276: 2082: 2071: 2064: 2006: 1999: 1985: 1943: 1878: 1844: 1815: 1768: 1284:? Geometric Morphometric Evidence of Late Maastrichtian Ceratopsid Dinosaurs" 1241: 788: 705: 697: 416: 68: 727:, or perhaps its nearest relative. An ongoing debate concerns the status of 2688: 2639: 2632: 2548: 2521: 2507: 2388: 2381: 2374: 2325: 2311: 2203: 2190: 2121: 2110: 2095: 2057: 2049: 2031: 1936: 1902: 1885: 1837: 1830: 1786: 1591: 1539: 1393: 1327: 1259: 956: 685: 647:
skulls, and there also are several holes in the frill (a unique feature of
628: 563:; largely attributing its differences to being that of an aged individual. 534: 492: 399: 241: 196: 806: 2805: 2681: 2575: 2458: 2422: 2245: 1957: 1822: 754: 543: 510:
in 1904 at the age of 42, leaving the paper still incomplete. It fell to
488: 484: 437: 363: 43: 1168:
Marsh, 1889 (Ceratopsidae: Chasmosaurinae): synonymy through ontogeny".
2825: 2451: 2367: 2217: 1950: 1738: 1672: 1659: 1618: 1497: 1343: 1232: 775: 762: 733: 713: 609: 228: 88: 53: 1085: 758: 571: 495: 147: 93: 37: 2767: 2790: 2252: 2238: 2231: 1794: 1530: 1521: 608:
prefix meaning "insufficient". The suffix "ceratops", common among
402: 180: 167: 83: 78: 63: 58: 48: 1411:, a giant horned dinosaur from the Late Campanian of New Mexico". 837:
It has been noted that many of the features that seem to separate
620: 2646: 1808: 1752: 972:"Two new Ceratopsia from the Laramie of Converse County, Wyoming" 424: 98: 73: 578:
in 2007. Unaware that Ukrainsky had already renamed the animal,
1515: 1213: 1159: 745:(2010 and 2011) and claimed that the USNM 2412 skull (i.e., of 701: 507: 157: 117: 1064:
Mateus, Octávio (2008). "Two ornithischian dinosaurs renamed:
909:"Anatomy and Taxonomic Status of the Chasmosaurine Ceratopsid 1466: 1344:
Scannella, John B.; Horner, John R.; Claessens, Leon (2011).
420: 395: 913:
from the Upper Cretaceous Lance Formation of Wyoming, U.S.A"
826:
individuals, and some specimens more solidly attributed to
709: 700:
for "horned faces"), a group of herbivorous dinosaurs with
753:. Evidence for this hypothesis included the shapes of the 498:. Marsh died in 1899 before the work was completed, and 394:(meaning "insufficient horned face") is a controversial 1273: 1271: 1269: 541:
being a distinct genus and he put it in a subgenus of
1339: 1337: 1114:
Mateus, 2008 (Reptilia: Ornithischia: Ceratopidae)."
741:. In a series of publications, John B. Scannella and 1450: 1406: 1132:
Mateus, 2008 (Reptilia: Ornithischia: Ceratopidae)."
604:
means "insufficient horned face". The "nedo" is the
427:. Its status is the subject of ongoing debate among 1266: 1045:
Lull, 1905 (Reptilia: Ornithischia: Ceratopsidae)."
1334: 696:belonged to the Ceratopsia (the name is Latinised 1122:(1): 108. Translated as: Ukrainsky, A.S. (2009). 2866: 1160:John B. Scannella; John R. Horner (July 2010). " 1155: 1153: 1151: 1149: 1147: 1003:"Restoration of the horned dinosaur Diceratops" 1563: 1209: 1207: 1482: 1144: 907:Farke, A. A. (2011). Claessens, Leon (ed.). 2201: 1204: 468:, was found in eastern Wyoming in 1891, in 2880:Late Cretaceous dinosaurs of North America 1489: 1475: 1350:: An Example of a Transitional Morphology" 1106:Ukrainsky, A.S. (2009). "Sinonimiya rodov 1100: 1033: 537:. By 1933, Lull had second thoughts about 116: 1383: 1373: 1317: 1307: 1249: 1231: 946: 936: 1400: 805: 667: 619: 582:coined another new name for it in 2008, 448: 969: 902: 900: 898: 896: 894: 892: 444: 2867: 1063: 639:. Superficially, it resembles that of 2772: 2771: 2188: 1657: 1508: 1470: 906: 358:Lull vide Hatcher, 1905 (preoccupied) 1000: 889: 872:"Nedoceratops: To Be, or Not to Be?" 817:Another supposed difference between 719:Several authors have suggested that 1057: 13: 1170:Journal of Vertebrate Paleontology 14: 2931: 1446: 869: 799:being distinct, valid taxa, with 663: 419:. It is known only from a single 2752: 2751: 2727: 2721: 2189: 2171: 2165: 1453: 134: 41: 1658: 1496: 1407:Nicholas R. Longrich (2011). " 994: 963: 863: 749:) belonged to a "young adult" 615: 570:name was already in use for a 1: 2915:Ceratopsians of North America 2885:Fossil taxa described in 2007 1433:10.1016/j.cretres.2010.12.007 856: 723:may be directly ancestral to 704:-like beaks which thrived in 506:section. However, he died of 2920:Late Cretaceous ceratopsians 1375:10.1371/journal.pone.0028705 1309:10.1371/journal.pone.0081608 1190:10.1080/02724634.2010.483632 1041:"A new replacement name for 1001:Lull, Richard Swann (1905). 938:10.1371/journal.pone.0016196 769:and large openings in adult 7: 2900:Controversial dinosaur taxa 1116:Paleontologicheskii zhurnal 1054:(2), 20 December 2007: 292. 1007:American Journal of Science 976:American Journal of Science 970:Hatcher, John Bell (1904). 502:endeavored to complete the 464:, the holotype specimen of 125:Skull from multiple angles 10: 2936: 457:The nearly complete skull 336:, preoccupied by Förster, 16:Extinct genus of dinosaurs 2780: 2741: 2719: 2664: 2622: 2613: 2565: 2531: 2481: 2472: 2398: 2271: 2262: 2212: 2197: 2184: 2162: 2120: 2090: 2081: 2039: 2030: 1926: 1911: 1900: 1861: 1781: 1777: 1766: 1714: 1685: 1681: 1670: 1666: 1653: 1617: 1599: 1590: 1572: 1559: 1504: 1027:10.2475/ajs.s4-20.120.420 988:10.2475/ajs.s4-20.120.413 522:means "two horned face." 479:The paper that described 453:1905 diagram of the skull 351: 344: 330:(Lull vide Hatcher, 1905) 317: 310: 131:Scientific classification 129: 124: 115: 23: 1124:"Synonymy of the genera 1039:Ukrainsky, A.S. (2007). 821:and fossils referred to 533:, thinking it primarily 431:: some authors consider 2895:Paleontology in Wyoming 1698:Micropachycephalosaurus 1135:Paleontological Journal 1074:Journal of Paleontology 810:Skull exhibited at the 483:was originally part of 1409:Titanoceratops ouranos 1282:Torosaurus Triceratops 1048:Zoosystematica Rossica 814: 681: 632: 454: 2847:Paleobiology Database 911:Nedoceratops hatcheri 809: 773:formerly assigned to 690:Nedoceratops hatcheri 672:Skull comparisons of 671: 637:Nedoceratops hatcheri 627:showing the possibly 623: 466:Nedoceratops hatcheri 452: 324:Nedoceratops hatcheri 2910:Ornithischian genera 2905:Maastrichtian genera 1128:Ukrainsky, 2007 and 876:Smithsonian Magazine 847:Triceratops horridus 832:Triceratops horridus 823:Triceratops horridus 445:History of discovery 1863:Archaeoceratopsidae 1425:2011CrRes..32..264L 1413:Cretaceous Research 1366:2011PLoSO...628705S 1300:2013PLoSO...881608M 1182:2010JVPal..30.1157S 1019:1905AmJS...20..420L 929:2011PLoSO...616196F 845:, and specifically 631:holes in the frills 516:Diceratops hatcheri 2583:Crittendenceratops 2533:Albertaceratopsini 1233:10.7717/peerj.5194 1110:Ukrainsky, 2007 i 815: 791:found support for 682: 633: 512:Richard Swann Lull 455: 2862: 2861: 2774:Taxon identifiers 2765: 2764: 2737: 2736: 2717: 2716: 2713: 2712: 2709: 2708: 2705: 2704: 2666:Pachyrhinosaurini 2604:Yehuecauhceratops 2567:Nasutoceratopsini 2468: 2467: 2180: 2179: 2160: 2159: 2156: 2155: 2152: 2151: 2148: 2147: 2144: 2143: 2140: 2139: 2041:Protoceratopsidae 1928:Leptoceratopsidae 1896: 1895: 1762: 1761: 1649: 1648: 1645: 1644: 1641: 1640: 1637: 1636: 500:John Bell Hatcher 387: 386: 381: 370: 359: 291: 2927: 2855: 2854: 2842: 2841: 2829: 2828: 2816: 2815: 2814: 2801: 2800: 2799: 2769: 2768: 2755: 2754: 2732: 2731: 2725: 2696:Pachyrhinosaurus 2620: 2619: 2494:Machairoceratops 2479: 2478: 2319:Coahuilaceratops 2269: 2268: 2210: 2209: 2199: 2198: 2186: 2185: 2175: 2170: 2169: 2088: 2087: 2037: 2036: 2021:Zhuchengceratops 1909: 1908: 1779: 1778: 1775: 1774: 1716:Chaoyangsauridae 1683: 1682: 1679: 1678: 1668: 1667: 1655: 1654: 1597: 1596: 1570: 1569: 1561: 1560: 1554: 1553: 1506: 1505: 1491: 1484: 1477: 1468: 1467: 1463: 1461:Dinosaurs portal 1458: 1457: 1456: 1437: 1436: 1404: 1398: 1397: 1387: 1377: 1341: 1332: 1331: 1321: 1311: 1275: 1264: 1263: 1253: 1235: 1211: 1202: 1201: 1176:(4): 1157–1168. 1164:Marsh, 1891, is 1157: 1142: 1104: 1098: 1097: 1086:10.1666/07-069.1 1068:Bohlin 1953 and 1061: 1055: 1037: 1031: 1030: 1013:(120): 420–422. 998: 992: 991: 982:(120): 413–419. 967: 961: 960: 950: 940: 904: 887: 886: 884: 882: 867: 625:Life restoration 376: 368: 357: 326: 322: 303:N. hatcheri 286: 279: 266: 253: 240: 227: 211: 195: 139: 138: 120: 110: 40: 29:Temporal range: 21: 20: 2935: 2934: 2930: 2929: 2928: 2926: 2925: 2924: 2865: 2864: 2863: 2858: 2850: 2845: 2837: 2832: 2824: 2819: 2810: 2809: 2804: 2795: 2794: 2789: 2776: 2766: 2761: 2733: 2726: 2701: 2660: 2609: 2590:Furcatoceratops 2561: 2542:Albertaceratops 2527: 2501:Menefeeceratops 2464: 2437:Tatankaceratops 2394: 2340:Mercuriceratops 2291:Arrhinoceratops 2258: 2206: 2193: 2176: 2164: 2136: 2116: 2077: 2026: 1993:Montanoceratops 1922: 1917:Graciliceratops 1905: 1892: 1872:Archaeoceratops 1857: 1771: 1758: 1746:Xuanhuaceratops 1732:Hualianceratops 1710: 1691:Albalophosaurus 1675: 1662: 1633: 1613: 1586: 1580:Avemetatarsalia 1565:Avemetatarsalia 1555: 1549:Marginocephalia 1511: 1510: 1500: 1495: 1459: 1454: 1452: 1449: 1443: 1441: 1440: 1405: 1401: 1342: 1335: 1276: 1267: 1212: 1205: 1158: 1145: 1105: 1101: 1062: 1058: 1038: 1034: 999: 995: 968: 964: 905: 890: 880: 878: 868: 864: 859: 666: 618: 474:Lightning Creek 470:Niobrara County 447: 429:paleontologists 413:Lance Formation 407:Late Cretaceous 340: 331: 328: 320: 319: 306: 285: 277: 264: 251: 238: 225: 209: 193: 133: 111: 109: 108: 107: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 51: 46: 35: 34: 31:Late Cretaceous 27: 17: 12: 11: 5: 2933: 2923: 2922: 2917: 2912: 2907: 2902: 2897: 2892: 2887: 2882: 2877: 2875:Chasmosaurines 2860: 2859: 2857: 2856: 2843: 2830: 2817: 2802: 2786: 2784: 2778: 2777: 2763: 2762: 2760: 2759: 2749: 2742: 2739: 2738: 2735: 2734: 2720: 2718: 2715: 2714: 2711: 2710: 2707: 2706: 2703: 2702: 2700: 2699: 2692: 2685: 2678: 2670: 2668: 2662: 2661: 2659: 2658: 2650: 2643: 2636: 2628: 2626: 2617: 2611: 2610: 2608: 2607: 2600: 2597:Nasutoceratops 2593: 2586: 2579: 2571: 2569: 2563: 2562: 2560: 2559: 2556:Medusaceratops 2552: 2545: 2537: 2535: 2529: 2528: 2526: 2525: 2518: 2511: 2504: 2497: 2490: 2487:Diabloceratops 2482: 2476: 2474:Centrosaurinae 2470: 2469: 2466: 2465: 2463: 2462: 2455: 2448: 2445:Titanoceratops 2441: 2433: 2430:Regaliceratops 2426: 2419: 2412: 2404: 2402: 2400:Triceratopsini 2396: 2395: 2393: 2392: 2385: 2378: 2371: 2364: 2361:Sierraceratops 2357: 2350: 2347:Navajoceratops 2343: 2336: 2329: 2322: 2315: 2308: 2301: 2294: 2287: 2280: 2272: 2266: 2264:Chasmosaurinae 2260: 2259: 2257: 2256: 2249: 2242: 2235: 2228: 2225:Brachyceratops 2221: 2213: 2207: 2202: 2195: 2194: 2182: 2181: 2178: 2177: 2163: 2161: 2158: 2157: 2154: 2153: 2150: 2149: 2146: 2145: 2142: 2141: 2138: 2137: 2135: 2134: 2133: 2132: 2126: 2124: 2118: 2117: 2115: 2114: 2107: 2104:Turanoceratops 2100: 2091: 2085: 2079: 2078: 2076: 2075: 2068: 2061: 2054: 2045: 2043: 2034: 2028: 2027: 2025: 2024: 2017: 2014:Unescoceratops 2010: 2003: 1996: 1989: 1982: 1979:Ischioceratops 1975: 1972:Gryphoceratops 1968: 1961: 1954: 1947: 1940: 1932: 1930: 1924: 1923: 1921: 1920: 1912: 1906: 1901: 1898: 1897: 1894: 1893: 1891: 1890: 1882: 1875: 1867: 1865: 1859: 1858: 1856: 1855: 1852:Sasayamagnomus 1848: 1841: 1834: 1827: 1819: 1812: 1805: 1802:Auroraceratops 1798: 1791: 1782: 1772: 1767: 1764: 1763: 1760: 1759: 1757: 1756: 1749: 1742: 1735: 1728: 1725:Chaoyangsaurus 1720: 1718: 1712: 1711: 1709: 1708: 1705:Psittacosaurus 1701: 1694: 1686: 1676: 1671: 1664: 1663: 1651: 1650: 1647: 1646: 1643: 1642: 1639: 1638: 1635: 1634: 1632: 1631: 1630: 1629: 1623: 1621: 1615: 1614: 1612: 1611: 1610: 1609: 1600: 1594: 1588: 1587: 1585: 1584: 1583: 1582: 1573: 1567: 1557: 1556: 1552: 1551: 1542: 1533: 1524: 1518: 1502: 1501: 1494: 1493: 1486: 1479: 1471: 1465: 1464: 1448: 1447:External links 1445: 1439: 1438: 1419:(3): 264–276. 1399: 1360:(12): e28705. 1333: 1294:(11): e81608. 1265: 1203: 1143: 1099: 1056: 1032: 993: 962: 888: 870:Black, Riley. 861: 860: 858: 855: 743:John R. Horner 676:(A. Left) and 665: 664:Classification 662: 617: 614: 592:junior synonym 580:Octávio Mateus 446: 443: 423:discovered in 385: 384: 383: 382: 371: 360: 349: 348: 342: 341: 329: 315: 314: 308: 307: 299: 297: 293: 292: 275: 271: 270: 268:Triceratopsini 262: 258: 257: 255:Chasmosaurinae 249: 245: 244: 236: 232: 231: 223: 216: 215: 213:Neornithischia 207: 200: 199: 191: 184: 183: 178: 171: 170: 165: 161: 160: 155: 151: 150: 145: 141: 140: 127: 126: 122: 121: 113: 112: 104: 103: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 52: 47: 42: 28: 15: 9: 6: 4: 3: 2: 2932: 2921: 2918: 2916: 2913: 2911: 2908: 2906: 2903: 2901: 2898: 2896: 2893: 2891: 2888: 2886: 2883: 2881: 2878: 2876: 2873: 2872: 2870: 2853: 2848: 2844: 2840: 2835: 2831: 2827: 2822: 2818: 2813: 2807: 2803: 2798: 2792: 2788: 2787: 2785: 2783: 2779: 2775: 2770: 2758: 2750: 2748: 2744: 2743: 2740: 2730: 2724: 2698: 2697: 2693: 2691: 2690: 2686: 2684: 2683: 2679: 2677: 2676: 2675:Achelousaurus 2672: 2671: 2669: 2667: 2663: 2656: 2655: 2654:Styracosaurus 2651: 2649: 2648: 2644: 2642: 2641: 2637: 2635: 2634: 2630: 2629: 2627: 2625: 2624:Centrosaurini 2621: 2618: 2616: 2615:Eucentrosaura 2612: 2606: 2605: 2601: 2599: 2598: 2594: 2592: 2591: 2587: 2585: 2584: 2580: 2578: 2577: 2573: 2572: 2570: 2568: 2564: 2558: 2557: 2553: 2551: 2550: 2546: 2544: 2543: 2539: 2538: 2536: 2534: 2530: 2524: 2523: 2519: 2517: 2516: 2515:Wendiceratops 2512: 2510: 2509: 2505: 2503: 2502: 2498: 2496: 2495: 2491: 2489: 2488: 2484: 2483: 2480: 2477: 2475: 2471: 2461: 2460: 2456: 2454: 2453: 2449: 2447: 2446: 2442: 2439: 2438: 2434: 2432: 2431: 2427: 2425: 2424: 2420: 2418: 2417: 2413: 2411: 2410: 2409:Eotriceratops 2406: 2405: 2403: 2401: 2397: 2391: 2390: 2386: 2384: 2383: 2379: 2377: 2376: 2372: 2370: 2369: 2365: 2363: 2362: 2358: 2356: 2355: 2354:Pentaceratops 2351: 2349: 2348: 2344: 2342: 2341: 2337: 2335: 2334: 2333:Kosmoceratops 2330: 2328: 2327: 2323: 2321: 2320: 2316: 2314: 2313: 2309: 2307: 2306: 2305:Bravoceratops 2302: 2300: 2299: 2298:Bisticeratops 2295: 2293: 2292: 2288: 2286: 2285: 2284:Anchiceratops 2281: 2279: 2278: 2277:Agujaceratops 2274: 2273: 2270: 2267: 2265: 2261: 2255: 2254: 2250: 2248: 2247: 2243: 2241: 2240: 2236: 2234: 2233: 2229: 2227: 2226: 2222: 2220: 2219: 2215: 2214: 2211: 2208: 2205: 2200: 2196: 2192: 2187: 2183: 2174: 2168: 2130: 2129: 2128: 2127: 2125: 2123: 2119: 2113: 2112: 2108: 2106: 2105: 2101: 2098: 2097: 2093: 2092: 2089: 2086: 2084: 2083:Ceratopsoidea 2080: 2074: 2073: 2072:Protoceratops 2069: 2067: 2066: 2065:Breviceratops 2062: 2060: 2059: 2055: 2052: 2051: 2047: 2046: 2044: 2042: 2038: 2035: 2033: 2029: 2023: 2022: 2018: 2016: 2015: 2011: 2009: 2008: 2007:Udanoceratops 2004: 2002: 2001: 2000:Prenoceratops 1997: 1995: 1994: 1990: 1988: 1987: 1986:Leptoceratops 1983: 1981: 1980: 1976: 1974: 1973: 1969: 1967: 1966: 1962: 1960: 1959: 1955: 1953: 1952: 1948: 1946: 1945: 1944:Bainoceratops 1941: 1939: 1938: 1934: 1933: 1931: 1929: 1925: 1919: 1918: 1914: 1913: 1910: 1907: 1904: 1899: 1888: 1887: 1883: 1881: 1880: 1879:Helioceratops 1876: 1874: 1873: 1869: 1868: 1866: 1864: 1860: 1854: 1853: 1849: 1847: 1846: 1845:Mosaiceratops 1842: 1840: 1839: 1835: 1833: 1832: 1828: 1825: 1824: 1820: 1818: 1817: 1816:Koreaceratops 1813: 1811: 1810: 1806: 1804: 1803: 1799: 1797: 1796: 1792: 1789: 1788: 1784: 1783: 1780: 1776: 1773: 1770: 1769:Neoceratopsia 1765: 1755: 1754: 1750: 1748: 1747: 1743: 1741: 1740: 1736: 1734: 1733: 1729: 1727: 1726: 1722: 1721: 1719: 1717: 1713: 1707: 1706: 1702: 1700: 1699: 1695: 1693: 1692: 1688: 1687: 1684: 1680: 1677: 1674: 1669: 1665: 1661: 1656: 1652: 1627: 1626: 1625: 1624: 1622: 1620: 1616: 1608: 1604: 1603: 1602: 1601: 1598: 1595: 1593: 1589: 1581: 1577: 1576: 1575: 1574: 1571: 1568: 1566: 1562: 1558: 1550: 1546: 1543: 1541: 1537: 1534: 1532: 1528: 1525: 1523: 1519: 1517: 1513: 1512: 1507: 1503: 1499: 1492: 1487: 1485: 1480: 1478: 1473: 1472: 1469: 1462: 1451: 1444: 1434: 1430: 1426: 1422: 1418: 1414: 1410: 1403: 1395: 1391: 1386: 1381: 1376: 1371: 1367: 1363: 1359: 1355: 1351: 1349: 1340: 1338: 1329: 1325: 1320: 1315: 1310: 1305: 1301: 1297: 1293: 1289: 1285: 1283: 1274: 1272: 1270: 1261: 1257: 1252: 1247: 1243: 1239: 1234: 1229: 1225: 1221: 1217: 1210: 1208: 1199: 1195: 1191: 1187: 1183: 1179: 1175: 1171: 1167: 1163: 1156: 1154: 1152: 1150: 1148: 1140: 1136: 1133: 1131: 1127: 1121: 1117: 1113: 1109: 1103: 1095: 1091: 1087: 1083: 1079: 1075: 1071: 1067: 1066:Microceratops 1060: 1053: 1049: 1046: 1044: 1036: 1028: 1024: 1020: 1016: 1012: 1008: 1004: 997: 989: 985: 981: 977: 973: 966: 958: 954: 949: 944: 939: 934: 930: 926: 923:(1): e16196. 922: 918: 914: 912: 903: 901: 899: 897: 895: 893: 877: 873: 866: 862: 854: 852: 848: 844: 840: 835: 833: 829: 824: 820: 813: 808: 804: 802: 798: 794: 790: 789:morphometrics 786: 782: 778: 777: 772: 768: 764: 761:bones, and a 760: 756: 752: 748: 744: 740: 736: 735: 730: 726: 722: 717: 715: 711: 707: 706:North America 703: 699: 695: 691: 687: 679: 675: 670: 661: 659: 655: 650: 646: 642: 638: 630: 626: 622: 613: 611: 607: 603: 599: 597: 593: 589: 585: 581: 577: 573: 569: 564: 562: 558: 554: 550: 546: 545: 540: 536: 532: 528: 523: 521: 517: 513: 509: 505: 501: 497: 494: 490: 486: 482: 477: 475: 471: 467: 463: 461: 451: 442: 440: 439: 434: 430: 426: 422: 418: 417:North America 414: 411: 408: 404: 401: 397: 393: 392: 379: 375: 372: 367: 365: 361: 356: 353: 352: 350: 347: 343: 339: 335: 327: 325: 316: 313: 312:Binomial name 309: 305: 304: 298: 295: 294: 290: 284: 283: 276: 273: 272: 269: 263: 260: 259: 256: 250: 247: 246: 243: 237: 234: 233: 230: 224: 221: 218: 217: 214: 208: 205: 202: 201: 198: 192: 189: 186: 185: 182: 179: 176: 173: 172: 169: 166: 163: 162: 159: 156: 153: 152: 149: 146: 143: 142: 137: 132: 128: 123: 119: 114: 100: 95: 90: 85: 80: 75: 70: 65: 60: 55: 50: 45: 39: 32: 26: 22: 19: 2812:Nedoceratops 2782:Nedoceratops 2781: 2694: 2689:Stellasaurus 2687: 2680: 2673: 2652: 2645: 2640:Coronosaurus 2638: 2633:Centrosaurus 2631: 2602: 2595: 2588: 2581: 2574: 2554: 2549:Lokiceratops 2547: 2540: 2522:Xenoceratops 2520: 2513: 2508:Sinoceratops 2506: 2499: 2492: 2485: 2457: 2450: 2443: 2435: 2428: 2421: 2416:Nedoceratops 2415: 2414: 2407: 2389:Vagaceratops 2387: 2382:Utahceratops 2380: 2375:Terminocavus 2373: 2366: 2359: 2352: 2345: 2338: 2331: 2326:Judiceratops 2324: 2317: 2312:Chasmosaurus 2310: 2303: 2296: 2289: 2282: 2275: 2251: 2244: 2237: 2230: 2223: 2216: 2204:Ceratopsidae 2191:Ceratopsidae 2122:Ceratopsidae 2111:Zuniceratops 2109: 2102: 2096:Ajkaceratops 2094: 2070: 2063: 2058:Bagaceratops 2056: 2050:Ajkaceratops 2048: 2032:Coronosauria 2019: 2012: 2005: 1998: 1991: 1984: 1977: 1970: 1963: 1956: 1949: 1942: 1937:Asiaceratops 1935: 1915: 1903:Euceratopsia 1886:Yamaceratops 1884: 1877: 1870: 1850: 1843: 1838:Microceratus 1836: 1831:Liaoceratops 1829: 1821: 1814: 1807: 1800: 1793: 1787:Ajkaceratops 1785: 1751: 1744: 1737: 1730: 1723: 1703: 1696: 1689: 1607:Ornithischia 1592:Ornithischia 1544: 1540:Ornithischia 1535: 1526: 1442: 1416: 1412: 1408: 1402: 1357: 1353: 1348:Nedoceratops 1347: 1291: 1287: 1281: 1223: 1219: 1173: 1169: 1165: 1161: 1138: 1134: 1129: 1126:Nedoceratops 1125: 1119: 1115: 1111: 1108:Nedoceratops 1107: 1102: 1077: 1073: 1072:Lull 1905". 1069: 1065: 1059: 1051: 1047: 1042: 1035: 1010: 1009:. Series 4. 1006: 996: 979: 978:. Series 4. 975: 965: 920: 916: 910: 879:. Retrieved 875: 865: 850: 846: 842: 839:Nedoceratops 838: 836: 831: 827: 822: 819:Nedoceratops 818: 816: 801:Nedoceratops 800: 796: 792: 785:Nedoceratops 784: 780: 774: 770: 766: 750: 747:Nedoceratops 746: 739:Nedoceratops 738: 732: 728: 724: 721:Nedoceratops 720: 718: 694:Nedoceratops 693: 689: 686:type species 683: 678:Nedoceratops 677: 673: 657: 653: 648: 644: 640: 636: 634: 629:pathological 610:ceratopsians 602:Nedoceratops 601: 600: 596:Nedoceratops 595: 587: 583: 576:Nedoceratops 575: 572:hymenopteran 567: 566:Because the 565: 560: 559:, including 556: 552: 548: 542: 538: 535:pathological 530: 526: 524: 519: 515: 503: 493:Ceratopsidae 481:Nedoceratops 480: 478: 465: 458: 456: 436: 433:Nedoceratops 432: 391:Nedoceratops 390: 389: 388: 373: 366:(Diceratops) 362: 354: 333: 323: 318: 302: 301: 282:Nedoceratops 281: 280: 242:Ceratopsidae 219: 203: 197:Ornithischia 187: 174: 25:Nedoceratops 24: 18: 2890:Lance fauna 2806:Wikispecies 2682:Einiosaurus 2576:Avaceratops 2459:Triceratops 2423:Ojoceratops 2246:Monoclonius 1958:Ferrisaurus 1823:Kulceratops 1166:Triceratops 843:Triceratops 828:T. horridus 812:Smithsonian 793:Triceratops 781:Triceratops 771:Triceratops 767:Triceratops 755:epoccipital 751:Triceratops 729:Triceratops 725:Triceratops 712:during the 674:Triceratops 658:Triceratops 654:Triceratops 649:Triceratops 645:Triceratops 641:Triceratops 616:Description 549:Triceratops 544:Triceratops 504:Triceratops 489:magnum opus 485:O. C. Marsh 438:Triceratops 364:Triceratops 332:Originally 287:Ukrainsky, 248:Subfamily: 36:67–66  2869:Categories 2745:See also: 2452:Torosaurus 2368:Spiclypeus 2218:Agathaumas 2131:see below↓ 1951:Cerasinops 1739:Stenopelix 1673:Ceratopsia 1660:Ceratopsia 1628:see below↓ 1619:Ceratopsia 1531:Dinosauria 1498:Ceratopsia 1162:Torosaurus 1141:43(1):116. 1080:(2): 423. 1070:Diceratops 1043:Diceratops 857:References 851:Torosaurus 797:Torosaurus 776:Torosaurus 763:neck frill 734:Torosaurus 714:Cretaceous 680:(B. Right) 590:is thus a 568:Diceratops 561:T. obtusus 553:Diceratops 539:Diceratops 531:Diceratops 527:Diceratops 525:Since the 520:Diceratops 400:ceratopsid 369:Lull, 1933 355:Diceratops 334:Diceratops 229:Ceratopsia 181:Dinosauria 1514:Kingdom: 1242:2167-8359 1226:: e5194. 1130:Diceratus 1112:Diceratus 759:squamosal 588:Diceratus 584:Diceratus 496:monograph 405:from the 374:Diceratus 296:Species: 154:Kingdom: 148:Eukaryota 2791:Wikidata 2757:Category 2747:Timeline 2253:Polyonax 2239:Dysganus 2232:Ceratops 1795:Aquilops 1522:Chordata 1520:Phylum: 1516:Animalia 1394:22194891 1354:PLOS ONE 1328:24303058 1288:PLOS ONE 1260:30002987 1198:86767957 1094:86021954 957:21283763 917:PLOS ONE 881:11 March 557:hatcheri 403:dinosaur 346:Synonyms 235:Family: 168:Chordata 164:Phylum: 158:Animalia 144:Domain: 2839:1475528 2826:4965810 2797:Q132602 2647:Spinops 1965:Gremlin 1753:Yinlong 1421:Bibcode 1385:3241274 1362:Bibcode 1319:3841114 1296:Bibcode 1251:6034596 1178:Bibcode 1015:Bibcode 948:3024410 925:Bibcode 606:Russian 425:Wyoming 321:† 300:† 274:Genus: 261:Tribe: 105:↓ 2852:139255 1392:  1382:  1326:  1316:  1258:  1248:  1240:  1196:  1092:  955:  945:  737:, and 702:parrot 508:typhus 491:, his 410:period 380:, 2008 378:Mateus 2834:IRMNG 1545:Clade 1536:Clade 1527:Clade 1220:PeerJ 1194:S2CID 1090:S2CID 841:from 698:Greek 472:near 421:skull 396:genus 220:Clade 204:Clade 188:Clade 175:Clade 2821:GBIF 1605:see 1578:see 1390:PMID 1324:PMID 1280:"Is 1256:PMID 1238:ISSN 1139:2009 1120:2009 953:PMID 883:2011 795:and 757:and 710:Asia 708:and 684:The 462:2412 460:USNM 338:1868 289:2007 44:Preęž’ 1809:Beg 1429:doi 1380:PMC 1370:doi 1314:PMC 1304:doi 1246:PMC 1228:doi 1186:doi 1082:doi 1023:doi 984:doi 943:PMC 933:doi 688:is 594:of 487:'s 415:of 398:of 2871:: 2849:: 2836:: 2823:: 2808:: 2793:: 1547:: 1538:: 1529:: 1427:. 1417:32 1415:. 1388:. 1378:. 1368:. 1356:. 1352:. 1336:^ 1322:. 1312:. 1302:. 1290:. 1286:. 1268:^ 1254:. 1244:. 1236:. 1222:. 1218:. 1206:^ 1192:. 1184:. 1174:30 1172:. 1146:^ 1137:, 1118:, 1088:. 1078:82 1076:. 1052:16 1050:, 1021:. 1005:. 974:. 951:. 941:. 931:. 919:. 915:. 891:^ 874:. 834:. 731:, 692:. 660:. 598:. 586:. 555:) 547:: 518:; 476:. 441:. 222:: 206:: 190:: 177:: 94:Pg 38:Ma 33:, 2657:? 2440:? 2099:? 2053:? 1889:? 1826:? 1790:? 1490:e 1483:t 1476:v 1435:. 1431:: 1423:: 1396:. 1372:: 1364:: 1358:6 1346:" 1330:. 1306:: 1298:: 1292:8 1262:. 1230:: 1224:6 1200:. 1188:: 1180:: 1096:. 1084:: 1029:. 1025:: 1017:: 1011:4 990:. 986:: 980:4 959:. 935:: 927:: 921:6 885:. 551:( 278:† 265:† 252:† 239:† 226:† 210:† 194:† 99:N 89:K 84:J 79:T 74:P 69:C 64:D 59:S 54:O 49:ęž’

Index

Late Cretaceous
Ma
Preęž’
ęž’
O
S
D
C
P
T
J
K
Pg
N

Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Dinosauria
Ornithischia
Neornithischia
Ceratopsia
Ceratopsidae
Chasmosaurinae
Triceratopsini
Nedoceratops
2007
Binomial name

Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.

↑