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Pantherophis obsoletus

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649:, as researchers have observed that snakes have used this method of tail vibration as a defense mechanism against predation long before the emergence of rattlesnakes. They are also capable of producing a foul-smelling musk, which they will release onto predators if picked up. They spread the musk with their tails in hopes of deterring the threat. When cornered or provoked, black snakes are known to stand their ground and can become aggressive. Counterattacks on large birds of prey, often committed by large snakes in excess of 150 cm (59 in) in length, have resulted in violent prolonged struggles. Utilizing its infamous agility and the great strength of its muscular coils, the black rat snake is sometimes able to overwhelm and kill formidable avian predators such as 102: 77: 33: 452: 57: 527:
are diurnally-active and live in a variety of habitats; some overlap each other. They have adapted to a variety of habitats, including bayou, prairie, and rock outcrops, but they seem to have a particular preference for wooded areas, especially oak trees. Rat snakes are excellent climbers and spend
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such as corn snakes, kingsnakes, milksnakes, and hognose snakes. Opinions vary on the western rat snake's disposition, but captive-bred individuals are reported to be more docile than their wild counterparts. With appropriate care, this species may be expected to live 15 years in captivity, and
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Account of an Expedition from Pittsburgh to the Rocky Mountains, Performed in the Years 1819 and '20, by Order of the Hon. J.C. Calhoun, Sec'y of War: Under the Command of Major Stephen H. Long. From the Notes of Major Long, Mr. T. Say, and other Gentlemen of the Exploring Party. Vol.
762:, which are 36–60 mm (1.4–2.4 in) long by 20–26.5 mm (0.79–1.04 in) wide. The eggs hatch about 65 to 70 days later in late August to early October. The hatchlings are 28–41 cm (11–16 in) in total length, and they look like miniature fox snakes. 597:). Darkening occurs rapidly as they grow. Adults are glossy black above with white lips, chin, and throat. Sometimes traces of the "obsolete" juvenile pattern are still discernible in the skin between the scales, especially when stretched after a heavy meal. 680:
to the point of cardiovascular collapse due to obstructive shock, coiling around small animals and tightening its grip until the prey can no longer circulate blood and dies of profound hypotension, before being eaten. Though it will often consume
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suggested this species be divided into three species based on geographic patterns of mitochondrial DNA diversity. He assigned new common names and resurrected old scientific names, resulting in the following combinations: eastern ratsnake
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can range up to 0.5 to 2.2 kg (1.1 to 4.9 lb) in adults, although most adults are on the smaller end of this scale, per herpetology research sites, with weights most commonly between 0.77 and 1 kg (1.7 and 2.2 lb).
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colonies as a single large snake will readily consume a number of eggs, hatchlings, and adults each summer. Several rat snake repelling methods have been offered to those putting up martin houses, but most are mixed in success.
693:, the western rat snake is far from a specialist at this kind of prey and will readily consume any small vertebrate it can catch. Other prey opportunistically eaten by this species can include other 539:. When found in homes, often in groups. This association gave rise to one of its common names, pilot black snake, and the superstition that this nonvenomous species led the venomous ones to the den. 1618:
Gibbs, H. Lisle; Corey, Sarah J.; Blouin-Demers, Gabriel; Prior, Kent A.; Weatherhead, Patrick J. (2006). "Hybridization between mtDNA-defined phylogeographic lineages of black ratsnakes (
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include: gray rat snake, black chicken snake, black coluber, chicken snake, mountain black snake, mountain pilot snake, pilot, rat snake, rusty black snake, scaly black snake, cow snake,
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can become quite large, with a reported typical total length (including tail) of 106.5–183 cm (3 ft 6 in â€“ 6 ft 0 in). It is the largest snake found in
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mating takes place in late May and early June. The male snake wraps its tail around the female with their vents nearly touching. The male then everts one of its sex organs, a
1398: 1313: 846:). However, these three species are not morphologically distinct and overlap in all examined morphological characters. More recent investigations have indicated 733:
eggs. Cavity-nesting bird species are seemingly especially prevalent in this snake's diet. The western rat snake has been noted as perhaps the top predator at
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When not fully grown, rat snakes are subject to predation by many animals, including other snakes. Once they attain maturity, they are readily preyed on by
922: 1866: 1918: 1379: 1990: 637:). When startled, they may freeze and wrinkle themselves into a series of kinks. If they feel further threatened, they may flee quickly or 1144:. (with 108 drawings by Edmond Malnate). New York and London: D. Appleton-Century. Frontispiece map + viii + 163 pp. + Plates A-C, 1-32. ( 2020: 510:, which is a brown-to-black variant, often with tinges of orange or red, that can be found in southern Oklahoma, Texas, and Louisiana. 1840: 1544:, Burbrink FT (2009). "Neogene diversification and taxonomic stability in the snake tribe Lampropeltini (Serpentes: Colubridae)". 1124: 1105: 1879: 1046: 934: 2025: 1884: 1086: 1727: 1265: 1038: 893:
The western rat snake is available captive-bred in the United States pet trade, and it has been bred for mutations such as
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Aside from the usual variety that is black or has patches of black on a lighter background, color variations include the
1175: 2035: 1246: 1021: 261: 2005: 1795: 1472: 1923: 64: 1666: 1541: 858: 368: 355: 1158: 555:. The record total length is 256.5 cm (8 ft 5 in), making it (officially) the longest snake in 1282: 1066: 959: 2015: 1070: 2010: 1419: 824: 101: 938: 971: 579:
Juveniles are strongly patterned with brown blotches on a gray background (like miniature fox snakes:
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Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America, Fourth Edition
1435: 1910: 1617: 885:, 2007. The justification for this nomenclatural change has been removed by more recent research. 758:. The mating lasts a few minutes to a few hours. After five weeks, the female lays about 12 to 20 1704: 1376: 1137: 795: 304: 229: 1423: 1142:
What Snake Is That? A Field Guide to the Snakes of the United States East of the Rocky Mountains
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a significant amount of their time in trees. The black rat snake is also a competent swimmer.
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A Field Guide to Reptiles and Amphibians of Eastern and Central North America, Second Edition
1093:, p. 388 + Figure 161 on p. 334 + Plate 36 on p. 335 + Map on p. 385 + Figure 180 on p. 386). 206: 1949: 661:, though in many cases the bird is able to kill the snake and both combatants may even die. 1897: 1804: 1745: 1500: 1283:"Behavioral Plasticity and the Origins of Novelty: The Evolution of the Rattlesnake Rattle" 1217: 882: 593: 433: 338: 571:), with a portion of its tail missing, measured 111 inches (2.8 m). The body mass of 8: 2000: 1005: 658: 581: 46: 1647: 1600: 1431: 1415: 1102: 536: 321: 238: 96: 641:; although this act of tail vibration is indeed a behavior that rat snakes share with 1944: 1827: 1639: 1635: 1561: 1520: 1242: 1082: 1074: 1017: 468: 351: 1957: 1832: 1651: 1631: 1592: 1553: 1512: 770:
This species has previously been placed (and is still placed by many) in the genus
654: 414: 278: 1931: 1476: 1402: 1383: 1317: 1109: 1016:. Ithaca and London: Comstock. 1,105 pp. (in 2 volumes) (Seventh printing, 1985) 650: 634: 507: 429: 1557: 1516: 1439: 638: 308: 265: 1979: 1892: 1780: 1503:(2007). "How and when did Old World ratsnakes disperse into the New World?". 1188: 759: 734: 706: 626: 556: 441: 421: 291: 86: 81: 1643: 1565: 1524: 783: 673: 630: 523: 183: 1936: 1669:(2008). "An alternative classification of the New World Rat Snakes (genus 1905: 1853: 1819: 1789: 642: 386: 1733: 1467: 1281:
Allf, Bradley C.; Durst, Paul A. P.; Pfennig, David W. (July 28, 2016).
1845: 1695: 1604: 902: 729:, and bird eggs. One snake was observed to consume an entire clutch of 561: 425: 417: 223: 173: 428:
that are recognized as being valid. Its color variations include the
1858: 1440:"Molecular Systematics and Phylogeny of Old and New World ratsnakes, 894: 751: 622: 484: 113: 1751: 1596: 972:
https://dx.doi.org/10.2305/IUCN.UK.2019-2.RLTS.T90069553A90069569.en
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Black snake profile at Smithsonian National Zoological Park website
726: 714: 710: 500: 492: 480: 476: 451: 153: 133: 432:. Along with other snakes of the eastern United States, like the 1739: 1579:
Burbrink FT (2001). "Systematics of the Eastern Ratsnake complex
1349:. Smithsonian Institution U.S. National Museum Bulletin, no. 170. 1336:. Smithsonian Institution U.S. National Museum Bulletin, no. 167. 730: 722: 646: 406: 143: 1414: 1081:. Boston and New York: Houghton Mifflin Harcourt. xiv + 494 pp. 898: 810:
has been further confirmed by additional phylogenetic studies.
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Found in Gaston county NC 8-21-19 species over 14’ in length.
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Auct., and related genera (Reptilia, Squamata, Colubridae)"
1241:. Boston: Houghton Mifflin. xviii + 429 pp. + Plates 1-48. 970:
Red List of Threatened Species 2019: e.T90069553A90069569.
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Fact Sheet at Smithsonian National Zoological Park Website
629:, which weighs no more than an adult rat snake) and large 565:) are known to exceed it, and one wild-caught pine snake ( 40:
Western rat snake swimming on water in northeastern Texas
690: 1228:, pp. 148-150, Figure 40 + Plate 16, center, on p. 336). 901:, and scaleless. However, it is not as popular as other 1480: 531:
During winter it hibernates in shared dens, often with
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Life Histories of North American Birds of Prey. Part 2
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Life Histories of North American Birds of Prey. Part 1
697:(including both those of its own and other species), 1491: 1489: 1222:
Field Book of Snakes of the United States and Canada
1014:Handbook of Snakes of the United States and Canada 1536: 1534: 1486: 1191:. Herpetological Education & Research Project 991:, The Reptile Database. www.reptile-database.org. 1977: 1280: 1224:. New York: G.P. Putnam's Sons. xiii + 365 pp. ( 1178:. Qrg.northwestern.edu. Retrieved on 2012-12-19. 1166:. Reptilia.forumpro.fr. Retrieved on 2012-12-19. 1312:Typically when found in homes, found in groups 1271:. Natural Heritage Endangered Program. mass.gov 1712:Philadelphia: H.C. Carey and I. Lea. 503 pp. ( 1664: 1531: 1495: 1405:. Nationalzoo.si.edu. Retrieved on 2012-12-19. 1320:. Nationalzoo.si.edu. Retrieved on 2012-12-19. 881:) in response to the findings of Burbrink and 1540: 1028:, pp. 230-234, Figure 72 + Map 24 on p. 235). 1386:. Purplemartin.org. Retrieved on 2012-12-19. 1578: 1127:. Dnr.state.mn.us. Retrieved on 2012-12-19. 1740:"Black Snakes": Identification and Ecology 75: 55: 31: 1481:The Center for North American Herpetology 613:, sleepy John, and white-throated racer. 450: 1261: 1259: 1047:Integrated Taxonomic Information System 786:, and placed this species in the genus 1978: 1756: 1755: 1546:Molecular Phylogenetics and Evolution 1505:Molecular Phylogenetics and Evolution 1209: 1207: 1205: 1120: 1118: 982: 980: 935:Ontario Ministry of Natural Resources 559:. Unofficially, indigo snakes (genus 1461: 1256: 1253:, pp. 193-194 + Plate 28 + Map 149). 1001: 999: 997: 778:. However, Utiger et al. found that 1991:IUCN Red List least concern species 1031: 931:Natural Heritage Information Centre 459: 420:. The species is native to central 13: 2021:Fauna of the Eastern United States 1742:– University of Florida fact sheet 1688: 1202: 1148:, pp. 56-58 + Plate 8, Figure 23). 1115: 977: 14: 2047: 1748:, Reptiles and Amphibians of Iowa 1721: 994: 754:, into the female sex organ, the 1636:10.1111/j.1365-294X.2006.03056.x 1160:Pantherophis obsoletus obsoletus 100: 1658: 1611: 1572: 1408: 1389: 1370: 1352: 1339: 1323: 1303: 1274: 1231: 1181: 1169: 1151: 888: 741: 600: 448:), it is called “black snake”. 1451:Russian Journal of Herpetology 1130: 1125:Species profile: Minnesota DNR 1096: 1060: 953: 915: 854:interbreed freely in Ontario. 542: 1: 2026:Reptiles of the United States 1734:Reptarium.cz Reptile Database 1675:Journal of Kansas Herpetology 974:. Downloaded on 05 July 2021. 909: 1716:, new species, p. 140). 471:, from eastern and southern 7: 1558:10.1016/j.ympev.2009.02.008 1517:10.1016/j.ympev.2006.09.009 1268:Pantherophis alleghaniensis 865:for Burbrink's three taxa ( 825:Pantherophis alleghaniensis 765: 616: 10: 2052: 2031:Reptiles described in 1823 1986:NatureServe secure species 838:), and western rat snake ( 676:, meaning it squeezes its 664: 513: 1764: 1585:Herpetological Monographs 244: 237: 212: 205: 97:Scientific classification 95: 73: 53: 44: 39: 30: 23: 2036:Taxa named by Thomas Say 1483:. Accessed 20 June 2008. 1251:Elaphe obsoleta obsoleta 1226:Elaphe obsoleta obsoleta 1146:Elaphe obsoleta obsoleta 1026:Elaphe obsoleta obsoleta 871:Scotophis alleghaniensis 782:is broadly construed as 2006:Snakes of North America 1736:. Accessed 2 June 2008. 790:. In addition, because 605:Other common names for 1810:Pantherophis_obsoletus 1796:Pantherophis obsoletus 1766:Pantherophis obsoletus 1729:Pantherophis obsoletus 1362:Pantherophis obsoletus 1091:Pantherophis obsoletus 988:Pantherophis obsoletus 964:Pantherophis obsoletus 861:resurrected the genus 844:Pantherophis obsoletus 835:Pantherophis spiloides 798:becomes the masculine 645:, it is not a form of 568:Pituophis melanoleucus 487:, westward to eastern 456: 382:Pantherophis obsoletus 363:Pantherophis obsoletus 316:Pantherophis obsoletus 216:Pantherophis obsoletus 25:Pantherophis obsoletus 857:In 2008, Collins and 828:), central ratsnake ( 820:Elaphe alleghaniensis 467:is found west of the 454: 1364:(Eastern Rat Snake)" 1176:Black Rat Snake Info 659:red-shouldered hawks 623:mammalian carnivores 491:, northward through 434:eastern indigo snake 371:& Burbrink, 2009 2016:Reptiles of Ontario 879:Scotophis obsoletus 875:Scotophis spiloides 537:timber rattlesnakes 446:Coluber constrictor 438:Drymarchon couperi) 405:, is a nonvenomous 346:Scotophis obsoletus 329:Pituophis obsoletus 273:Scotophis obsoletus 47:Conservation status 2011:Reptiles of Canada 1475:2009-04-14 at the 1401:2016-08-25 at the 1382:2012-11-22 at the 1316:2016-08-25 at the 1266:Eastern Ratsnake. 1108:2017-05-06 at the 794:is masculine, the 611:schwartze Schlange 518:Rat snakes of the 475:southward through 457: 257:Elaphis holbrookii 1973: 1972: 1945:Open Tree of Life 1758:Taxon identifiers 1714:Coluber obsoletus 1630:(12): 3755–3767. 1624:Molecular Ecology 1581:(Elaphe obsoleta) 1237:Conant R (1975). 1164: 1087:978-0-544-12997-9 941:on March 22, 2012 848:P. alleghaniensis 655:great horned owls 469:Mississippi River 399:pilot black snake 391:western rat snake 378: 377: 286:Elaphis obsoletus 248:Coluber obsoletus 198:P. obsoletus 90: 68: 2043: 1966: 1965: 1953: 1952: 1940: 1939: 1927: 1926: 1914: 1913: 1901: 1900: 1888: 1887: 1875: 1874: 1862: 1861: 1849: 1848: 1836: 1835: 1823: 1822: 1813: 1812: 1800: 1799: 1798: 1785: 1784: 1783: 1753: 1752: 1683: 1682: 1662: 1656: 1655: 1615: 1609: 1608: 1576: 1570: 1569: 1538: 1529: 1528: 1493: 1484: 1465: 1459: 1458: 1448: 1412: 1406: 1393: 1387: 1374: 1368: 1367: 1356: 1350: 1345:Bent AC (1938). 1343: 1337: 1327: 1321: 1307: 1301: 1300: 1298: 1296: 1287: 1278: 1272: 1263: 1254: 1235: 1229: 1211: 1200: 1199: 1197: 1196: 1185: 1179: 1173: 1167: 1162: 1155: 1149: 1134: 1128: 1122: 1113: 1112:, Herps of Texas 1103:Eastern Ratsnake 1100: 1094: 1064: 1058: 1057: 1055: 1053: 1041:Elaphe obsoleta 1035: 1029: 1003: 992: 984: 975: 957: 951: 950: 948: 946: 937:. Archived from 925:Elaphe obsoleta 919: 830:Elaphe spiloides 796:specific epithet 651:red-tailed hawks 635:red-tailed hawks 499:to southeastern 460:Geographic range 372: 359: 342: 325: 312: 295: 282: 269: 253: 218: 105: 104: 84: 79: 78: 62: 59: 58: 35: 21: 20: 16:Species of snake 2051: 2050: 2046: 2045: 2044: 2042: 2041: 2040: 1976: 1975: 1974: 1969: 1961: 1956: 1948: 1943: 1935: 1932:Observation.org 1930: 1922: 1917: 1909: 1904: 1896: 1891: 1883: 1878: 1870: 1865: 1857: 1852: 1844: 1839: 1831: 1826: 1818: 1816: 1808: 1803: 1794: 1793: 1788: 1779: 1778: 1773: 1760: 1746:Black Rat Snake 1724: 1719: 1691: 1689:Further reading 1686: 1663: 1659: 1616: 1612: 1597:10.2307/1467037 1577: 1573: 1539: 1532: 1494: 1487: 1477:Wayback Machine 1469:Elaphe obsoleta 1466: 1462: 1446: 1413: 1409: 1403:Wayback Machine 1394: 1390: 1384:Wayback Machine 1375: 1371: 1358: 1357: 1353: 1344: 1340: 1328: 1324: 1318:Wayback Machine 1308: 1304: 1294: 1292: 1285: 1279: 1275: 1264: 1257: 1236: 1232: 1212: 1203: 1194: 1192: 1187: 1186: 1182: 1174: 1170: 1156: 1152: 1135: 1131: 1123: 1116: 1110:Wayback Machine 1101: 1097: 1065: 1061: 1051: 1049: 1037: 1036: 1032: 1004: 995: 985: 978: 958: 954: 944: 942: 921: 920: 916: 912: 906:possibly more. 891: 840:Elaphe obsoleta 802:. The split of 776:Elaphe obsoleta 768: 744: 667: 625:(including the 619: 603: 545: 516: 508:Texas rat snake 462: 455:Black rat snake 430:Texas rat snake 424:. There are no 395:black rat snake 366: 365: 349: 348: 332: 331: 319: 318: 302: 301: 299:Elaphe obsoleta 289: 288: 276: 275: 260: 259: 251: 250: 233: 220: 214: 201: 99: 91: 80: 76: 69: 60: 56: 49: 17: 12: 11: 5: 2049: 2039: 2038: 2033: 2028: 2023: 2018: 2013: 2008: 2003: 1998: 1993: 1988: 1971: 1970: 1968: 1967: 1954: 1941: 1928: 1915: 1902: 1889: 1876: 1863: 1850: 1837: 1824: 1814: 1801: 1786: 1770: 1768: 1762: 1761: 1750: 1749: 1743: 1737: 1723: 1722:External links 1720: 1718: 1717: 1692: 1690: 1687: 1685: 1684: 1657: 1610: 1571: 1552:(2): 524–529. 1530: 1511:(1): 173–189. 1485: 1460: 1420:Helfenberger N 1407: 1388: 1369: 1351: 1338: 1322: 1302: 1290:D. Pfennig Lab 1273: 1255: 1249:(paperback). ( 1230: 1201: 1180: 1168: 1150: 1129: 1114: 1095: 1059: 1030: 993: 976: 952: 913: 911: 908: 890: 887: 767: 764: 743: 740: 666: 663: 618: 615: 602: 599: 544: 541: 515: 512: 461: 458: 376: 375: 374: 373: 360: 343: 326: 313: 296: 283: 270: 262:A.M.C. Dumeril 254: 242: 241: 235: 234: 221: 210: 209: 203: 202: 195: 193: 189: 188: 181: 177: 176: 171: 167: 166: 161: 157: 156: 151: 147: 146: 141: 137: 136: 131: 127: 126: 121: 117: 116: 111: 107: 106: 93: 92: 74: 71: 70: 54: 51: 50: 45: 42: 41: 37: 36: 28: 27: 15: 9: 6: 4: 3: 2: 2048: 2037: 2034: 2032: 2029: 2027: 2024: 2022: 2019: 2017: 2014: 2012: 2009: 2007: 2004: 2002: 1999: 1997: 1994: 1992: 1989: 1987: 1984: 1983: 1981: 1964: 1959: 1955: 1951: 1946: 1942: 1938: 1933: 1929: 1925: 1920: 1916: 1912: 1907: 1903: 1899: 1894: 1890: 1886: 1881: 1877: 1873: 1868: 1864: 1860: 1855: 1851: 1847: 1842: 1838: 1834: 1829: 1825: 1821: 1815: 1811: 1806: 1802: 1797: 1791: 1787: 1782: 1776: 1772: 1771: 1769: 1767: 1763: 1759: 1754: 1747: 1744: 1741: 1738: 1735: 1731: 1730: 1726: 1725: 1715: 1711: 1706: 1703: 1702: 1697: 1694: 1693: 1680: 1676: 1672: 1668: 1661: 1653: 1649: 1645: 1641: 1637: 1633: 1629: 1625: 1621: 1614: 1606: 1602: 1598: 1594: 1590: 1586: 1582: 1575: 1567: 1563: 1559: 1555: 1551: 1547: 1543: 1537: 1535: 1526: 1522: 1518: 1514: 1510: 1506: 1502: 1498: 1492: 1490: 1482: 1478: 1474: 1471: 1470: 1464: 1457:(2): 105–124. 1456: 1452: 1445: 1443: 1437: 1433: 1429: 1425: 1421: 1417: 1411: 1404: 1400: 1397: 1392: 1385: 1381: 1378: 1373: 1365: 1363: 1355: 1348: 1342: 1335: 1331: 1326: 1319: 1315: 1311: 1306: 1291: 1284: 1277: 1270: 1269: 1262: 1260: 1252: 1248: 1247:0-395-19977-8 1244: 1240: 1234: 1227: 1223: 1219: 1215: 1210: 1208: 1206: 1190: 1184: 1177: 1172: 1165: 1161: 1154: 1147: 1143: 1139: 1133: 1126: 1121: 1119: 1111: 1107: 1104: 1099: 1092: 1088: 1084: 1080: 1076: 1072: 1068: 1063: 1048: 1044: 1042: 1034: 1027: 1023: 1022:0-8014-0463-0 1019: 1015: 1011: 1007: 1002: 1000: 998: 990: 989: 983: 981: 973: 969: 965: 961: 956: 940: 936: 932: 928: 926: 918: 914: 907: 904: 900: 896: 886: 884: 880: 876: 872: 868: 864: 860: 855: 853: 849: 845: 841: 837: 836: 831: 827: 826: 821: 816: 811: 809: 805: 801: 797: 793: 789: 785: 781: 777: 773: 763: 761: 757: 753: 749: 739: 736: 735:purple martin 732: 728: 724: 720: 716: 712: 708: 704: 700: 696: 692: 688: 684: 679: 675: 671: 662: 660: 656: 652: 648: 644: 640: 636: 632: 631:birds of prey 628: 627:American mink 624: 614: 612: 608: 598: 596: 595: 590: 589: 584: 583: 577: 574: 570: 569: 564: 563: 558: 557:North America 554: 550: 540: 538: 534: 529: 526: 525: 521: 511: 509: 504: 502: 498: 494: 490: 486: 482: 478: 474: 470: 466: 453: 449: 447: 443: 442:eastern racer 439: 435: 431: 427: 423: 422:North America 419: 416: 412: 408: 404: 400: 396: 392: 388: 385:, also known 384: 383: 370: 364: 361: 357: 353: 347: 344: 340: 336: 330: 327: 323: 317: 314: 310: 306: 300: 297: 293: 287: 284: 280: 274: 271: 267: 263: 258: 255: 249: 246: 245: 243: 240: 236: 231: 228: 225: 219: 217: 211: 208: 207:Binomial name 204: 200: 199: 194: 191: 190: 187: 186: 182: 179: 178: 175: 172: 169: 168: 165: 162: 159: 158: 155: 152: 149: 148: 145: 142: 139: 138: 135: 132: 129: 128: 125: 122: 119: 118: 115: 112: 109: 108: 103: 98: 94: 88: 83: 82:Least Concern 72: 66: 52: 48: 43: 38: 34: 29: 26: 22: 19: 1996:Pantherophis 1765: 1728: 1713: 1708: 1700: 1699: 1678: 1674: 1671:Pantherophis 1670: 1665:Collins JT, 1660: 1627: 1623: 1620:Pantherophis 1619: 1613: 1588: 1584: 1580: 1574: 1549: 1545: 1508: 1504: 1468: 1463: 1454: 1450: 1441: 1410: 1391: 1372: 1361: 1354: 1346: 1341: 1333: 1325: 1305: 1293:. Retrieved 1289: 1276: 1267: 1250: 1238: 1233: 1225: 1221: 1193:. Retrieved 1183: 1171: 1159: 1153: 1145: 1141: 1132: 1098: 1090: 1078: 1062: 1050:. Retrieved 1040: 1033: 1025: 1013: 987: 963: 960:Hammerson GA 955: 943:. Retrieved 939:the original 930: 924: 917: 892: 889:In captivity 878: 874: 870: 866: 862: 856: 852:P. spiloides 851: 847: 843: 839: 833: 829: 823: 819: 812: 807: 804:Pantherophis 803: 799: 792:Pantherophis 791: 788:Pantherophis 787: 784:paraphyletic 779: 775: 771: 769: 748:P. obsoletus 747: 745: 742:Reproduction 670:P. obsoletus 669: 668: 643:rattlesnakes 639:tail vibrate 633:(especially 620: 610: 607:P. obsoletus 606: 604: 601:Common names 592: 588:P. ramspotti 586: 580: 578: 573:P. obsoletus 572: 566: 560: 549:P. obsoletus 548: 546: 530: 524:Pantherophis 522: 517: 505: 495:and eastern 465:P. obsoletus 464: 463: 445: 437: 402: 401:, or simply 398: 394: 390: 381: 380: 379: 362: 345: 328: 324:et al., 2002 315: 298: 285: 272: 256: 247: 226: 215: 213: 197: 196: 185:Pantherophis 184: 24: 18: 1906:NatureServe 1854:iNaturalist 1790:Wikispecies 1497:Burbrink FT 1163:(Say, 1823) 1052:29 November 945:October 19, 721:, juvenile 717:, juvenile 674:constrictor 594:P. vulpinus 543:Description 533:copperheads 483:to western 403:black snake 65:NatureServe 2001:Rat snakes 1980:Categories 1667:Taggart TW 1436:Ziswiler V 1377:Rat Snakes 1295:January 2, 1214:Schmidt KP 1195:2015-05-21 1136:Conant R, 1075:Collins JT 910:References 562:Drymarchon 547:Adults of 426:subspecies 418:Colubridae 174:Colubridae 160:Suborder: 1963:obsoletus 1428:Schmidt C 1424:Schätti B 1138:Bridges W 1010:Wright AA 1006:Wright AH 962:(2019). " 903:colubrids 895:leucistic 863:Scotophis 813:In 2001, 800:obsoletus 752:hemipenis 727:songbirds 715:squirrels 711:chipmunks 582:P. gloydi 485:Louisiana 305:Stejneger 279:Kennicott 252:Say, 1823 192:Species: 164:Serpentes 120:Kingdom: 114:Eukaryota 1911:2.840332 1872:10643814 1817:BioLib: 1775:Wikidata 1707:(1823). 1698:(1823). 1681:: 16–18. 1644:17032272 1622:spp.)". 1591:: 1–53. 1566:19236930 1542:Pyron RA 1525:17113316 1501:Lawson R 1473:Archived 1438:(2002). 1416:Utiger U 1399:Archived 1380:Archived 1332:(1937). 1314:Archived 1220:(1941). 1218:Davis DD 1189:"Snakes" 1140:(1939). 1106:Archived 1077:(2016). 1071:Conant R 1067:Powell R 1012:(1957). 815:Burbrink 766:Taxonomy 723:opossums 617:Behavior 501:Nebraska 493:Oklahoma 481:Arkansas 477:Missouri 440:and the 387:commonly 335:Burbrink 239:Synonyms 170:Family: 154:Squamata 144:Reptilia 134:Chordata 130:Phylum: 124:Animalia 110:Domain: 87:IUCN 3.1 1885:1081826 1846:2455676 1781:Q998324 1732:at the 1705:James E 1652:3727973 1605:1467037 1330:Bent AC 966:". The 859:Taggart 731:mallard 719:rabbits 703:lizards 665:Feeding 647:mimicry 514:Habitat 413:in the 407:species 389:as the 356:Taggart 352:Collins 309:Barbour 232:, 1823) 180:Genus: 150:Order: 140:Class: 85: ( 63: ( 61:Secure 1950:530777 1937:101976 1650:  1642:  1603:  1564:  1523:  1442:Elaphe 1245:  1085:  1020:  899:albino 883:Lawson 877:, and 842:, now 832:, now 822:, now 808:Elaphe 780:Elaphe 772:Elaphe 756:cloaca 695:snakes 689:, and 591:, and 553:Canada 497:Kansas 415:family 358:, 2008 354:& 341:, 2007 339:Lawson 337:& 322:Utiger 311:, 1917 307:& 294:, 1883 292:Garman 281:, 1860 268:, 1854 266:Bibron 264:& 1924:39099 1898:63864 1867:IRMNG 1859:73888 1833:4CGYF 1820:58574 1696:Say T 1648:S2CID 1601:JSTOR 1447:(PDF) 1432:Ruf M 1286:(PDF) 806:from 774:, as 707:moles 699:frogs 687:voles 672:is a 520:genus 489:Texas 411:snake 369:Pyron 230:James 1919:NCBI 1893:IUCN 1880:ITIS 1841:GBIF 1673:)". 1640:PMID 1562:PMID 1521:PMID 1297:2024 1243:ISBN 1083:ISBN 1054:2006 1018:ISBN 968:IUCN 947:2010 867:i.e. 850:and 760:eggs 691:rats 683:mice 678:prey 657:and 535:and 479:and 473:Iowa 1828:CoL 1805:ADW 1701:In: 1632:doi 1593:doi 1583:". 1554:doi 1513:doi 1479:at 1089:. ( 1024:. ( 746:In 409:of 224:Say 1982:: 1960:: 1958:RD 1947:: 1934:: 1921:: 1908:: 1895:: 1882:: 1869:: 1856:: 1843:: 1830:: 1807:: 1792:: 1777:: 1710:I. 1679:26 1677:. 1646:. 1638:. 1628:15 1626:. 1599:. 1589:15 1587:. 1560:. 1550:52 1548:. 1533:^ 1519:. 1509:43 1507:. 1499:, 1488:^ 1453:. 1449:. 1434:, 1430:, 1426:, 1422:, 1418:, 1288:. 1258:^ 1216:, 1204:^ 1117:^ 1073:, 1069:, 1045:. 1008:, 996:^ 979:^ 933:. 929:. 897:, 873:, 869:, 725:, 713:, 709:, 705:, 701:, 685:, 653:, 585:, 503:. 397:, 393:, 367:— 350:— 333:— 320:— 303:— 290:— 277:— 227:in 1654:. 1634:: 1607:. 1595:: 1568:. 1556:: 1527:. 1515:: 1455:9 1366:. 1360:" 1299:. 1198:. 1056:. 1043:" 1039:" 949:. 927:" 923:" 818:( 444:( 436:( 222:( 89:) 67:)

Index


Conservation status
NatureServe
Least Concern
IUCN 3.1
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Reptilia
Squamata
Serpentes
Colubridae
Pantherophis
Binomial name
Say
James
Synonyms
A.M.C. Dumeril
Bibron
Kennicott
Garman
Stejneger
Barbour
Utiger
Burbrink
Lawson
Collins
Taggart

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