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Plant genetics

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micro-projectile bombardment, which wounds the tissue. Wounding of the target tissue causes the release of phenolic compounds by the plant, which induces invasion of the tissue by Agrobacterium. Because of this, microprojectile bombardment often increases the efficiency of infection with Agrobacterium. The marker is used to find the organism which has successfully taken up the desired gene. Tissues of the organism are then transferred to a medium containing an
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this is less of a problem. Polyploid individuals are created frequently by a variety of processes; however, once created, they usually cannot cross back to the parental type. Polyploid individuals that are capable of self-fertilizing can give rise to a new, genetically distinct lineage, which can be the start of a new species. This is often called "instant
498:, use this sequence of DNA to their advantage to better find and understand the role of different genes within a given genome. Through research and plant breeding, manipulation of different plant genes and loci encoded by the DNA sequence of the plant chromosomes by various methods can be done to produce different or desired 193:. Mendel studied "trait inheritance", patterns in the way traits are handed down from parents to offspring. He observed that organisms (most famously pea plants) inherit traits by way of discrete "units of inheritance". This term, still used today, is a somewhat ambiguous definition of what is referred to as a 600:, and some are nearly exclusively self-fertilizers. This means that a plant can be both mother and father to its offspring, a rare occurrence in animals. Scientists and hobbyists attempting to make crosses between different plants must take special measures to prevent the plants from self-fertilizing. In 967:
that currently available food derived from GM crops poses no greater risk to human health than conventional food, but that each GM food needs to be tested on a case-by-case basis before introduction. Nonetheless, members of the public are much less likely than scientists to perceive GM foods as safe.
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had invented a method that led to the first hybrid maize seed that were available commercially. The large demand for hybrid seed in the U.S. Corn Belt by the mid 1930s led to a rapid growth in the seed production industry and ultimately seed research. The strict requirements for producing hybrid seed
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approximately two micrometers in diameter. The particles are placed in a vacuum chamber and the plant tissue to be engineered is placed below the chamber. The particles are propelled at high velocity using a short pulse of high pressure helium gas, and hit a fine mesh baffle placed above the tissue
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on the other hand, while having only 7 distinct chromosomes, is considered a hexaploid and has 6 copies of each chromosome, for a total of 42. In animals, inheritable germline polyploidy is less common, and spontaneous chromosome increases may not even survive past fertilization. In plants however
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Genetically modifying plants is an important economic activity: in 2017, 89% of corn, 94% of soybeans, and 91% of cotton produced in the US were from genetically modified strains. Since the introduction of GM crops, yields have increased by 22%, and profits have increased to farmers, especially in
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present is also killed by the antibiotic. Only tissues expressing the marker will survive and possess the gene of interest. Thus, subsequent steps in the process will only use these surviving plants. In order to obtain whole plants from these tissues, they are grown under controlled environmental
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In the early 1900s, botanists and statisticians began to examine the segregation ratios put forth by Mendel. W.E. Castle discovered that while individual traits may segregate and change over time with selection, that when selection is stopped and environmental effects are taken into account, the
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found has been dated to 11,000 years before present in ancestral wheat. While initially selection may have happened unintentionally, it is very likely that by 5,000 years ago farmers had a basic understanding of heredity and inheritance. This selection over time gave rise to new crop species and
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that contains the genetic instructions used in the development and functioning of all known living organisms and some viruses. The main role of DNA molecules is the long-term storage of information. DNA is often compared to a set of blueprints or a recipe, or a code, since it contains the
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inducing (Ti) region is removed from the T-DNA (transfer DNA) and replaced with the desired gene and a marker, which is then inserted into the organism. This may involve direct inoculation of the tissue with a culture of transformed Agrobacterium, or inoculation following treatment with
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led to the development of careful population and inbred line maintenance, keeping plants isolated and unable to out-cross, which produced plants that better allowed researchers to tease out different genetic concepts. The structure of these populations allowed scientist such a
336:, recognized its importance and re-published an English translation. Mendel died in 1884. The significance of Mendel's work was not recognized until the turn of the 20th century. Its rediscovery prompted the foundation of modern genetics. His discoveries, deduction of 448:. Identifying traits and their underlying genetics allowed for transferring useful genes and the traits they controlled from either wild or mutant plants to crop plants. Understanding and manipulating of plant genetics was in its heyday during the 439:
and seek out wild species that had given rise to domesticated species after selection. Determining how crops changed over time with selection was initially based on morphological features. It developed over time to chromosomal analysis, then
332:, which were later named after him. His seminal work on genetics, “Versuche über Pflanzen-Hybriden” (Experiments on Plant Hybrids), was published in 1866, but went almost entirely unnoticed until 1900 when prominent botanists in the UK, like 624:. Polyploid organisms have more than two sets of homologous chromosomes. For example, humans have two sets of homologous chromosomes, meaning that a typical human will have 2 copies each of 23 different chromosomes, for a total of 46. 200:
Plants, like all known organisms, use DNA to pass on their traits. Animal genetics often focuses on parentage and lineage, but this can sometimes be difficult in plant genetics due to the fact that plants can, unlike most animals, be
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has increased nearly five-fold in the past century due in part to the discovery and proliferation of hybrid corn varieties. Plant genetics can be used to predict which combination of plants may produce a plant with
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to the understanding of plant genetics. It was the first plant to ever have its genome sequenced in the year 2000. It has a small genome, making the initial sequencing more attainable. It has a genome size of 125
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The legal and regulatory status of GM foods varies by country, with some nations banning or restricting them, and others permitting them with widely differing degrees of regulation. There are still ongoing
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Banjara, Manoj; Zhu, Longfu; Shen, Guoxin; Payton, Paxton; Zhang, Hong (2012-01-01). "Expression of an Arabidopsis sodium/proton antiporter gene (AtNHX1) in peanut to improve salt tolerance - Springer".
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begins. This process entails selection of the seeds with the desired traits and then retesting and growing to make sure that the entire process has been completed successfully with the desired results.
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National Academies Of Sciences, Engineering; Division on Earth Life Studies; Board on Agriculture Natural Resources; Committee on Genetically Engineered Crops: Past Experience Future Prospects (2016).
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Bidney, D; Scelonge, C; Martich, J; Burrus, M; Sims, L; Huffman, G (January 1992). "Microprojectile bombardment of plant tissues increases transformation frequency by Agrobacterium tumefaciens".
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Emmenecker, CĂ´me; MĂ©zard, Christine; Kumar, Rajeev (March 2023). "Repair of DNA double-strand breaks in plant meiosis: role of eukaryotic RecA recombinases and their modulators".
741:(TAIR) has been established as a repository for multiple data sets and information on the species. Information housed in TAIR include the complete genome sequence along with 225:, possess their own DNA. Chloroplasts thus provide an additional reservoir for genes and genetic diversity, and an extra layer of genetic complexity not found in animals. 2897: 456:. During this time, the molecule of heredity, DNA, was also discovered, which allowed scientists to actually examine and manipulate genetic information directly. 2052: 2851: 1965: 383:, recognized that by crossing plants and forming hybrids, they were not only able to combine traits from two desirable parents, but the crop also experienced 328:
traits of pea plants, such as their height, flower color, and seed characteristics. Mendel showed that the inheritance of these traits follows two particular
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with ease, since flowers develop at the ends of branches composed of somatic cells. People have known of this for centuries, and mutant branches are called "
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Panchin, Alexander Y.; Tuzhikov, Alexander I. (January 14, 2016). "Published GMO studies find no evidence of harm when corrected for multiple comparisons".
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Alonso-Blanco, Carlos; Koornneef, Maarten (2000-01-01). "Naturally occurring variation in Arabidopsis: an underexploited resource for plant genetics".
1731: 3086: 2993: 2280: 1955:, Prepared by the UK GM Science Review panel (July 2003). Chairman Professor Sir David King, Chief Scientific Advisor to the UK Government, P 9 893:. Genetic engineering techniques allow for the introduction of new traits as well as greater control over traits than previous methods such as 3024: 811:
is a popular model organism for both plant-pathogen and transgenic studies. Because its broad leaves are easily transiently transformed with
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and rice. This approach literally shoots genes into plant cells and plant cell chloroplasts. DNA is coated onto small particles of gold or
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species, brachypodium is diploid with a relatively small genome (~355 Mbp) with a short life-cycle, making genomic studies on it simpler.
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molecules. The DNA segments that carry this genetic information are called genes, and their location within the genome are referred to as
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is an experimental model grass that has many attributes that make it an excellent model for temperate cereals. Unlike wheat, a tetra or
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related to food safety, regulation, labeling, environmental impact, research methods, and the fact that some GM seeds are subject to
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to increase yields, confer pest and disease resistance, provide resistance to herbicides, or to increase their nutritional value.
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over time and the statistics underlying plant genetics. Their work laid the foundations for future genetic discoveries such as
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Dudley, J. W. (1993-07-08). "Molecular Markers in Plant Improvement: Manipulation of Genes Affecting Quantitative Traits".
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Castle, W. E. (1903). "The Laws of Heredity of Galton and Mendel, and Some Laws Governing Race Improvement by Selection".
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Arabidopsis thaliana, growing from between a crack in a sidewalk; it is considered a key model organism in plant genetics.
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that encodes about 25,000 genes. Because an incredible amount of research has been done on the plant, a database called
633:". Polyploids generally have larger fruit, an economically desirable trait, and many human food crops, including wheat, 4051: 3845: 3689: 2074:"Genetically modified Golden Rice falls short on lifesaving promises | The Source | Washington University in St. Louis" 1948: 1816: 488:, but other DNA sequences have structural purposes, or are involved in regulating the use of this genetic information. 445: 83: 3699: 3017: 2459: 1517: 123: 344:
have not only been used in research to gain a better understanding of plant genetics, but also play a large role in
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species of modern crop plants. Botanists between the 1920s and 1960s often would travel to regions of high plant
364:. This was independently discovered by G. H. Hardy and W. Weinberg, which ultimately gave rise to the concept of 57: 90: 3753: 3422: 1162: 365: 61: 17: 984:
Genetic modification has been the cause for much research into modern plant genetics, and has also led to the
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Haslberger, Alexander G. (2003). "Codex guidelines for GM foods include the analysis of unintended effects".
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Griffiths, Anthony J. F.; Miller, Jeffrey H.; Suzuki, David T.; Lewontin, Richard C.; Gelbart, eds. (2000).
765:") are available and have been useful in genetic research. This natural variation has been used to identify 4137: 3225: 3132: 3010: 1755: 72: 3733: 3359: 3344: 3288: 3279: 3230: 3155: 3096: 3081: 1533:
Lewontin, R. C.; Kojima, Ken-ichi (December 1960). "The Evolutionary Dynamics of Complex Polymorphisms".
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the developing world, by 68%. An important side effect of GM crops has been decreased land requirements,
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of single-site insertions of the foreign DNA, which allows for easier monitoring. In this method, the
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and scientist born on 20 July 1822 in Austria-Hungary. He worked at the Abbey of St. Thomas in
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Ministry for the Environment NZ – Report of the Royal Commission on Genetic Modification, 2001
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can be easier in many plants due to unique genetic abilities, such as being well adapted to
179:. Plant genetics is similar in many ways to animal genetics but differs in a few key areas. 97: 4072: 3890: 3679: 3454: 3220: 2381: 2251: 1835: 1589: 1439: 1363: 1095: 964: 720: 413: 53: 2776:"Evidence for Absolute Moral Opposition to Genetically Modified Food in the United States" 2372:
Yang, Y.T.; Chen, B. (2016). "Governing GMOs in the USA: science, law and public health".
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Wright, Sewall (May 1940). "Breeding Structure of Populations in Relation to Speciation".
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and for better nutrient profiles. Other such crops include the economically important GM
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between plant species for economic and aesthetic reasons. For example, the yield of
213:. Plants are unique in that they are able to produce energy-dense carbohydrates via 4157: 4132: 3557: 2954: 2790: 2723: 2715: 2665: 2645: 2571: 2570:. The National Academies of Sciences, Engineering, and Medicine (US). p. 149. 2468: 2389: 2341: 2295: 2259: 2205: 2197: 2134: 2114: 1912: 1843: 1789: 1711: 1703: 1669: 1634: 1611: 1597: 1542: 1472: 1387: 1371: 1327: 1322: 1314: 1247: 1177: 1172: 1036: 925: 578: 565:. Plants however are unique from other living organisms in the fact that they have 495: 449: 2810: 2719: 2345: 2236: 1281: 728:, a species of fruit fly, was to the understanding of early genetics, so has been 4017: 3606: 3490: 3449: 3408: 3318: 2997: 2263: 2118: 2100:"An overview of the last 10 years of genetically engineered crop safety research" 2098:
Nicolia, Alessandro; Manzo, Alberto; Veronesi, Fabio; Rosellini, Daniele (2013).
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instructions needed to construct other components of cells, such as proteins and
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The study of plant genetics has major economic impacts: many staple crops are
4211: 3807: 3684: 3622: 3439: 3328: 3215: 2794: 2299: 2237:"A literature review on the safety assessment of genetically modified plants" 1924: 1857: 1646: 1554: 1484: 1415: 1383: 1259: 1089: 1051: 1023: 998: 822: 770: 409: 405: 298: 222: 183: 2185: 2020:. The International Service for the Acquisition of Agri-biotech Applications 1945: 1102:. Once the plants are grown and produce seed, the process of evaluating the 1022:(within a living organism) transformation and has been especially useful in 4097: 3977: 3596: 3495: 3474: 3323: 3145: 2802: 2737: 2657: 2593: 2401: 2353: 2271: 2219: 2126: 1932: 1865: 1707: 1401: 1267: 1238:
Allard, Robert W. (December 1999). "History of Plant Population Genetics".
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delayed-ripening tomato. Most food modifications have primarily focused on
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Plants are generally more capable of surviving, and indeed flourishing, as
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began. Maize that has been self-pollinated experiences a phenomenon called
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genetic ratio stops changing and reach a sort of stasis, the foundation of
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varieties that are the basis of the crops we grow, eat and research today.
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Plants, like all other known living organisms, pass on their traits using
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While breeding experiments were taking place, other scientists such as
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method is considered preferable to the gene gun, because of a greater
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on selection provided the basis for much of genetics as a discipline.
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of many plant genomes. Today there are two predominant procedures of
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Commercial sale of genetically modified foods began in 1994, when
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generally. Central elements of eukaryotic meiosis are pairing of
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Around this same time, genetic and plant breeding experiments in
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European Commission. Directorate-General for Research (2010).
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of pathogen genes introduced into a plant or test new genetic
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Final Report of the PABE research project (December 2001).
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Mirzaghaderi, Ghader; Hörandl, Elvira (14 September 2016).
979: 609: 534: 194: 148: 2921:"Agricultural Biotechnology: Background and Recent Issues" 2615:"Frequently asked questions on genetically modified foods" 1756:"Why is the Wheat Genome So Complicated? | Colorado Wheat" 301:, who is often called the "father of genetics". He was an 3916: 3911: 2607: 2165:. Food and Agriculture Organization of the United Nations 1018:
using biological components). This technique is used for
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Proceedings of the American Academy of Arts and Sciences
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while the DNA coating continues into any target cell or
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Genetically Engineered Crops: Experiences and Prospects
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Proceedings of the Royal Society B: Biological Sciences
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An image of multiple chromosomes, taken from many cells
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research community. Many natural inbred accessions of
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regularly, but these mutations can contribute to the
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List of varieties of genetically modified maize/corn
2875:"Over Half of E.U. Countries Are Opting Out of GMOs" 2281:"An Illusory Consensus behind GMO Health Assessment" 1834:(6814). The Arabidopsis Genome Initiative: 796–815. 1778: 2774:Scott, Sydney E.; Inbar, Yoel; Rozin, Paul (2016). 2695: 2672: 2535: 1815:The Arabidopsis Genome Initiative (December 2000). 1692:"The evolution of meiotic sex and its alternatives" 1689: 1685: 1683: 1282:"1. Gregor Mendel: Versuche ĂĽber Pflanzen-Hybriden" 297:The field of plant genetics began with the work of 1190: 881:Genetically modified (GM) foods are produced from 4209: 2825:"Restrictions on Genetically Modified Organisms" 2704:"Public views on GMOs: deconstructing the myths" 2527:: CS1 maint: bot: original URL status unknown ( 2235:Domingo, JosĂ© L.; Bordonaba, Jordi GinĂ© (2011). 2234: 1680: 537:that are homologous to recombinases employed by 2892:Lynch, Diahanna; Vogel, David (April 5, 2001). 2773: 2331: 518:, double-strand breaks are introduced into the 221:. Chloroplasts, like the superficially similar 2912: 2896:. Council on Foreign Relations. Archived from 2461:A decade of EU-funded GMO research (2001–2010) 2374:Journal of the Science of Food and Agriculture 1532: 1219: 1005: 952:which are resistant to the highly destructive 3839: 3018: 1352:"Mendelian Proportions in a Mixed Population" 870: 2679:Funk, Cary; Rainie, Lee (January 29, 2015). 2177: 1578:"Studies on the Origin of Cultivated Plants" 1014:method is also referred to as "biolistics" ( 885:that have had changes introduced into their 171:, but intersects frequently with many other 3660:Detection of genetically modified organisms 715:History of research on Arabidopsis thaliana 468:The structure of part of a DNA double helix 3846: 3832: 3025: 3011: 2891: 2850:. American Bar Association. Archived from 2678: 2635: 2433: 2006: 1505: 1087:, depending on which marker was used. The 780: 753:, publications, and information about the 556: 2872: 2866: 2727: 2209: 1966:"USDA ERS - Recent Trends in GE Adoption" 1847: 1715: 1639:10.2135/cropsci1993.0011183X003300040003x 1601: 1391: 1326: 1205:(7th ed.). New York: W. H. Freeman. 1199:"Genetics and the Organism: Introduction" 348:. Mendel's works along with the works of 175:and is strongly linked with the study of 124:Learn how and when to remove this message 2873:Sifferlin, Alexandra (October 3, 2015). 2845: 2839: 2371: 980:Modern ways to genetically modify plants 799: 655: 463: 288: 217:, a process which is achieved by use of 163:. It is generally considered a field of 135: 52:Relevant discussion may be found on the 4179:List of genetics research organizations 3032: 2817: 2557: 2434:Pinholster, Ginger (October 25, 2012). 2288:Science, Technology, & Human Values 2278: 1575: 1042: 940:have been engineered for resistance to 706: 293:Gregor Mendel, the "Father of genetics" 14: 4210: 2701: 2183: 1624: 1462: 1304: 1237: 960:(it is however still in development). 525:and these breaks can be repaired by a 3827: 3006: 2918: 2783:Perspectives on Psychological Science 2012: 1349: 828: 27:Study of genes and heredity in plants 2755:. Commission of European Communities 2032: 1509:Genetics of the Evolutionary Process 1233: 1231: 739:The Arabidopsis Information Resource 670: 502:that result in different or desired 245: 186:, a late 19th-century scientist and 62:adding citations to reliable sources 29: 2827:. Library of Congress. June 9, 2015 2545:. Library of Congress. June 9, 2015 1203:An Introduction to Genetic Analysis 1054:has been successfully practiced in 24: 3690:Genetic use restriction technology 2846:Bashshur, Ramona (February 2013). 1732:"Plant and Soil Sciences eLibrary" 1547:10.1111/j.1558-5646.1960.tb03113.x 1420:USDA Agricultural Research Service 920:in high demand by farmers such as 664: 596:Some plant species are capable of 577:. Like animals, plants experience 25: 4229: 2984: 2334:Critical Reviews in Biotechnology 2107:Critical Reviews in Biotechnology 1228: 1058:, i.e. broadleaf plants, such as 956:, and the nutritionally improved 4192: 4191: 912:first marketed its unsuccessful 674: 529:process employing gene products 249: 34: 3708:Cartagena Protocol on Biosafety 2938: 2767: 2496: 2451: 2409: 2226: 2155: 2091: 2066: 2045: 1982: 1958: 1939: 1896: 1872: 1808: 1772: 1748: 1724: 1653: 1618: 1569: 1526: 1506:Dobzhansky, Theodosius (1970). 1499: 817:, it is used to study both the 281:The earliest evidence of plant 182:The discoverer of genetics was 45:needs additional citations for 2919:Cowan, Tadlock (18 Jun 2011). 2184:Ronald, Pamela (May 5, 2011). 1946:GM Science Review First Report 1456: 1432: 1408: 1343: 1298: 1274: 1163:Genetically modified organisms 976:rights owned by corporations. 833:Other models include the alga 573:, chloroplasts have their own 13: 1: 3470:Somatic cell nuclear transfer 2346:10.3109/07388551.2015.1130684 1917:10.1016/S1360-1385(99)01510-1 1512:. Columbia University Press. 1184: 324:. Mendel's work tracked many 4138:Missing heritability problem 3853: 2848:"FDA and Regulation of GMOs" 2264:10.1016/j.envint.2011.01.003 2119:10.3109/07388551.2013.823595 1252:10.1146/annurev.genet.33.1.1 745:, gene product information, 416:concepts as well as explore 7: 2720:10.1093/embo-reports/kve142 2617:. World Health Organization 2202:10.1534/genetics.111.128553 1782:Plant Biotechnology Reports 1350:Hardy, G. H. (1908-07-10). 1110: 857:(snapdragon), the C4 grass 10: 4234: 1674:10.1007/s00497-022-00443-6 938:Genetically modified crops 874: 871:Genetically modified crops 712: 509: 366:Hardy–Weinberg equilibrium 239: 235: 4187: 4166: 4065: 4016: 3960: 3899: 3861: 3793: 3762: 3716: 3698: 3645: 3615: 3583: 3528:Genetically modified food 3520: 3513: 3483: 3430: 3421: 3384: 3337: 3287: 3278: 3208: 3190: 3172: 3154: 3131: 3113: 3095: 3062: 3053: 3040: 2279:Krimsky, Sheldon (2015). 2244:Environment International 1951:October 16, 2013, at the 1794:10.1007/s11816-011-0200-5 1376:10.1126/science.28.706.49 1240:Annual Review of Genetics 992:genes in organisms: the " 877:Genetically modified food 836:Chlamydomonas reinhardtii 814:Agrobacterium tumefaciens 2795:10.1177/1745691615621275 2300:10.1177/0162243915598381 431:were interested in wild 2702:Marris, Claire (2001). 1905:Trends in Plant Science 1576:Vavilov, N. I. (1926). 1465:The American Naturalist 1328:2027/hvd.32044106445109 1225:Hartl D, Jones E (2005) 789:Brachypodium distachyon 782:Brachypodium distachyon 761:(often referred to as " 557:Plant Specific Genetics 444:analysis, and eventual 2038:Weasel, Lisa H. 2009. 1708:10.1098/rspb.2016.1221 1138:Experimental evolution 1123:Biological engineering 661: 543:homologous chromosomes 469: 459: 422:linkage disequilibrium 395:. By the early 1920s, 294: 141: 4174:List of genetic codes 3141:Roundup ready soybean 2900:on September 29, 2016 2683:. Pew Research Center 2055:. Fda.gov. 2009-03-06 2013:James, Clive (1996). 974:intellectual property 954:Papaya ringspot virus 889:using the methods of 842:Physcomitrella patens 808:Nicotiana benthamiana 801:Nicotiana benthamiana 659: 473:Deoxyribonucleic acid 467: 381:Nils Heribert-Nilsson 377:inbreeding depression 292: 139: 4073:Behavioural genetics 3680:Reverse transfection 3455:Genetic transduction 2638:Nature Biotechnology 1994:Alliance for Science 1819:Arabidopsis thaliana 965:scientific consensus 721:Arabidopsis thaliana 708:Arabidopsis thaliana 414:evolutionary biology 379:. Researchers, like 299:Gregor Johann Mendel 230:genetically modified 58:improve this article 4153:Population genomics 4143:Molecular evolution 4103:Genetic engineering 3670:Genetics in fiction 3602:Genetic enhancement 3404:Hepatitis B vaccine 3034:Genetic engineering 2650:10.1038/nbt0703-739 2386:2016JSFA...96.1851Y 2256:2011EnInt..37..734D 1884:www.arabidopsis.org 1840:2000Natur.408..796T 1594:1926Natur.118..392T 1368:1908Sci....28...49H 1050:Transformation via 891:genetic engineering 848:Medicago truncatula 397:Donald Forsha Jones 368:published in 1908. 362:Population Genetics 242:History of genetics 177:information systems 4113:Genetic monitoring 3780:Stem cell research 3399:Ice-minus bacteria 2996:2010-06-02 at the 2959:10.1007/bf00034957 2053:"Consumer Q&A" 1880:"TAIR - Home Page" 1702:(1838): 20161221. 1662:Plant Reproduction 1168:Ice-minus bacteria 895:selective breeding 829:Other model plants 769:important in both 686:. You can help by 662: 598:self-fertilization 470: 338:segregation ratios 295: 261:. You can help by 142: 4205: 4204: 4128:He Jiankui affair 4118:Genetic genealogy 4108:Genetic diversity 4037:the British Isles 3942:Genetic variation 3821: 3820: 3785:Synthetic biology 3675:Human enhancement 3665:Genetic pollution 3641: 3640: 3509: 3508: 3417: 3416: 3380: 3379: 3274: 3273: 2585:978-0-309-43738-7 2482:978-92-79-16344-9 2394:10.1002/jsfa.7523 2042:Amacom Publishing 1760:coloradowheat.org 1588:(2967): 392–393. 1444:www.cabdirect.org 1212:978-0-7167-3520-5 1158:Genetic pollution 1006:"Gene gun" method 996:" method and the 899:mutation breeding 854:Antirrhinum majus 704: 703: 593:may be obtained. 579:somatic mutations 496:plant geneticists 452:brought about by 340:, and subsequent 334:Sir Gavin de Beer 279: 278: 153:genetic variation 134: 133: 126: 108: 16:(Redirected from 4225: 4195: 4194: 4158:Reverse genetics 4133:Medical genetics 3848: 3841: 3834: 3825: 3824: 3558:Dow AgroSciences 3518: 3517: 3428: 3427: 3285: 3284: 3060: 3059: 3051: 3050: 3027: 3020: 3013: 3004: 3003: 2979: 2978: 2942: 2936: 2935: 2933: 2931: 2925: 2916: 2910: 2909: 2907: 2905: 2889: 2883: 2882: 2870: 2864: 2863: 2861: 2859: 2854:on June 21, 2018 2843: 2837: 2836: 2834: 2832: 2821: 2815: 2814: 2780: 2771: 2765: 2764: 2762: 2760: 2748: 2742: 2741: 2731: 2699: 2693: 2692: 2690: 2688: 2676: 2670: 2669: 2633: 2627: 2626: 2624: 2622: 2611: 2605: 2604: 2602: 2600: 2561: 2555: 2554: 2552: 2550: 2539: 2533: 2532: 2526: 2518: 2516: 2514: 2508: 2500: 2494: 2493: 2491: 2489: 2466: 2455: 2449: 2447: 2445: 2443: 2431: 2429: 2427: 2421: 2413: 2407: 2405: 2380:(6): 1851–1855. 2365: 2325: 2323: 2322: 2316: 2310:. Archived from 2285: 2275: 2241: 2230: 2224: 2223: 2213: 2181: 2175: 2174: 2172: 2170: 2159: 2153: 2152: 2150: 2149: 2143: 2137:. Archived from 2104: 2095: 2089: 2088: 2086: 2085: 2070: 2064: 2063: 2061: 2060: 2049: 2043: 2036: 2030: 2029: 2027: 2025: 2019: 2010: 2004: 2003: 2001: 2000: 1986: 1980: 1979: 1977: 1976: 1970:www.ers.usda.gov 1962: 1956: 1943: 1937: 1936: 1900: 1894: 1893: 1891: 1890: 1876: 1870: 1869: 1851: 1849:10.1038/35048692 1825: 1812: 1806: 1805: 1776: 1770: 1769: 1767: 1766: 1752: 1746: 1745: 1743: 1742: 1728: 1722: 1721: 1719: 1687: 1678: 1677: 1657: 1651: 1650: 1622: 1616: 1615: 1605: 1603:10.1038/118392a0 1573: 1567: 1566: 1530: 1524: 1523: 1503: 1497: 1496: 1471:(752): 232–248. 1460: 1454: 1453: 1451: 1450: 1436: 1430: 1429: 1427: 1426: 1416:"Improving Corn" 1412: 1406: 1405: 1395: 1347: 1341: 1340: 1330: 1319:10.2307/20021870 1302: 1296: 1295: 1293: 1292: 1278: 1272: 1271: 1235: 1226: 1223: 1217: 1216: 1194: 1178:Paratransgenesis 1173:Monsanto Company 699: 696: 678: 671: 604:, people create 450:Green Revolution 446:genomic analysis 274: 271: 253: 246: 159:specifically in 147:is the study of 129: 122: 118: 115: 109: 107: 73:"Plant genetics" 66: 38: 30: 21: 4233: 4232: 4228: 4227: 4226: 4224: 4223: 4222: 4208: 4207: 4206: 4201: 4183: 4162: 4061: 4052:the Middle East 4018:Archaeogenetics 4012: 3956: 3895: 3857: 3852: 3822: 3817: 3789: 3758: 3712: 3694: 3647: 3637: 3611: 3607:Genetic testing 3589: 3579: 3505: 3491:Recombinant DNA 3479: 3450:Electroporation 3413: 3409:Oncolytic virus 3388: 3376: 3333: 3319:Herman the Bull 3270: 3204: 3186: 3168: 3150: 3127: 3109: 3091: 3045: 3043: 3036: 3031: 2998:Wayback Machine 2987: 2982: 2947:Plant Mol. Biol 2943: 2939: 2929: 2927: 2923: 2917: 2913: 2903: 2901: 2890: 2886: 2871: 2867: 2857: 2855: 2844: 2840: 2830: 2828: 2823: 2822: 2818: 2778: 2772: 2768: 2758: 2756: 2749: 2745: 2700: 2696: 2686: 2684: 2677: 2673: 2634: 2630: 2620: 2618: 2613: 2612: 2608: 2598: 2596: 2586: 2562: 2558: 2548: 2546: 2541: 2540: 2536: 2520: 2519: 2512: 2510: 2506: 2502: 2501: 2497: 2487: 2485: 2483: 2464: 2456: 2452: 2441: 2439: 2425: 2423: 2419: 2415: 2414: 2410: 2320: 2318: 2314: 2283: 2239: 2231: 2227: 2182: 2178: 2168: 2166: 2161: 2160: 2156: 2147: 2145: 2141: 2102: 2096: 2092: 2083: 2081: 2072: 2071: 2067: 2058: 2056: 2051: 2050: 2046: 2037: 2033: 2023: 2021: 2017: 2011: 2007: 1998: 1996: 1988: 1987: 1983: 1974: 1972: 1964: 1963: 1959: 1953:Wayback Machine 1944: 1940: 1901: 1897: 1888: 1886: 1878: 1877: 1873: 1823: 1813: 1809: 1777: 1773: 1764: 1762: 1754: 1753: 1749: 1740: 1738: 1730: 1729: 1725: 1688: 1681: 1658: 1654: 1623: 1619: 1574: 1570: 1531: 1527: 1520: 1504: 1500: 1461: 1457: 1448: 1446: 1438: 1437: 1433: 1424: 1422: 1414: 1413: 1409: 1348: 1344: 1303: 1299: 1290: 1288: 1286:www.bshs.org.uk 1280: 1279: 1275: 1236: 1229: 1224: 1220: 1213: 1195: 1191: 1187: 1182: 1153:Genetic erosion 1113: 1048: 1008: 982: 970:public concerns 879: 873: 860:Setaria viridis 831: 804: 785: 747:gene expression 717: 711: 700: 694: 691: 684:needs expansion 669: 666:Model organisms 559: 512: 462: 429:Nikolai Vavilov 275: 269: 266: 259:needs expansion 244: 238: 130: 119: 113: 110: 67: 65: 51: 39: 28: 23: 22: 15: 12: 11: 5: 4231: 4221: 4220: 4218:Plant genetics 4203: 4202: 4200: 4199: 4188: 4185: 4184: 4182: 4181: 4176: 4170: 4168: 4164: 4163: 4161: 4160: 4155: 4150: 4148:Plant genetics 4145: 4140: 4135: 4130: 4125: 4120: 4115: 4110: 4105: 4100: 4095: 4090: 4088:Genome editing 4085: 4080: 4075: 4069: 4067: 4066:Related topics 4063: 4062: 4060: 4059: 4054: 4049: 4044: 4039: 4034: 4029: 4023: 4021: 4014: 4013: 4011: 4010: 4005: 4000: 3995: 3990: 3988:Immunogenetics 3985: 3980: 3975: 3970: 3964: 3962: 3958: 3957: 3955: 3954: 3949: 3944: 3939: 3934: 3929: 3924: 3919: 3914: 3909: 3903: 3901: 3900:Key components 3897: 3896: 3894: 3893: 3888: 3883: 3878: 3873: 3868: 3862: 3859: 3858: 3851: 3850: 3843: 3836: 3828: 3819: 3818: 3816: 3815: 3810: 3805: 3800: 3794: 3791: 3790: 3788: 3787: 3782: 3777: 3772: 3766: 3764: 3763:Similar fields 3760: 3759: 3757: 3756: 3751: 3746: 3736: 3731: 3726: 3720: 3718: 3714: 3713: 3711: 3710: 3704: 3702: 3696: 3695: 3693: 3692: 3687: 3682: 3677: 3672: 3667: 3662: 3657: 3651: 3649: 3643: 3642: 3639: 3638: 3636: 3635: 3633:Gene targeting 3630: 3628:Gene knockdown 3625: 3619: 3617: 3613: 3612: 3610: 3609: 3604: 3599: 3593: 3591: 3581: 3580: 3578: 3577: 3576: 3575: 3570: 3565: 3563:DuPont Pioneer 3560: 3555: 3550: 3542: 3537: 3536: 3535: 3524: 3522: 3521:In agriculture 3515: 3511: 3510: 3507: 3506: 3504: 3503: 3498: 3493: 3487: 3485: 3481: 3480: 3478: 3477: 3472: 3467: 3465:Microinjection 3462: 3457: 3452: 3447: 3442: 3436: 3434: 3425: 3419: 3418: 3415: 3414: 3412: 3411: 3406: 3401: 3395: 3393: 3382: 3381: 3378: 3377: 3375: 3374: 3369: 3364: 3363: 3362: 3357: 3347: 3341: 3339: 3335: 3334: 3332: 3331: 3326: 3321: 3316: 3311: 3310: 3309: 3304: 3302:Knockout mouse 3293: 3291: 3282: 3276: 3275: 3272: 3271: 3269: 3268: 3263: 3258: 3253: 3248: 3243: 3238: 3233: 3228: 3223: 3218: 3212: 3210: 3206: 3205: 3203: 3202: 3196: 3194: 3188: 3187: 3185: 3184: 3178: 3176: 3170: 3169: 3167: 3166: 3160: 3158: 3152: 3151: 3149: 3148: 3143: 3137: 3135: 3129: 3128: 3126: 3125: 3119: 3117: 3111: 3110: 3108: 3107: 3101: 3099: 3093: 3092: 3090: 3089: 3084: 3079: 3074: 3068: 3066: 3057: 3048: 3038: 3037: 3030: 3029: 3022: 3015: 3007: 3001: 3000: 2986: 2985:External links 2983: 2981: 2980: 2937: 2911: 2884: 2865: 2838: 2816: 2789:(3): 315–324. 2766: 2743: 2714:(7): 545–548. 2694: 2671: 2644:(7): 739–741. 2628: 2606: 2584: 2576:10.17226/23395 2556: 2534: 2495: 2481: 2450: 2408: 2340:(2): 213–217. 2294:(6): 883–914. 2250:(4): 734–742. 2225: 2176: 2154: 2090: 2065: 2044: 2031: 2005: 1981: 1957: 1938: 1895: 1871: 1807: 1771: 1747: 1736:passel.unl.edu 1723: 1679: 1652: 1633:(4): 660–668. 1617: 1568: 1541:(4): 458–472. 1525: 1518: 1498: 1477:10.1086/280891 1455: 1431: 1407: 1362:(706): 49–50. 1342: 1313:(8): 223–242. 1297: 1273: 1227: 1218: 1211: 1188: 1186: 1183: 1181: 1180: 1175: 1170: 1165: 1160: 1155: 1150: 1145: 1140: 1135: 1130: 1125: 1120: 1114: 1112: 1109: 1096:tissue culture 1094:conditions in 1047: 1041: 1007: 1004: 981: 978: 875:Main article: 872: 869: 830: 827: 803: 798: 784: 779: 775:abiotic stress 743:gene structure 713:Main article: 710: 705: 702: 701: 681: 679: 668: 663: 602:plant breeding 558: 555: 511: 508: 461: 458: 454:Norman Borlaug 442:genetic marker 354:Alfred Wallace 350:Charles Darwin 346:plant breeding 277: 276: 256: 254: 237: 234: 215:photosynthesis 145:Plant genetics 132: 131: 56:. Please help 42: 40: 33: 26: 18:Plant Genetics 9: 6: 4: 3: 2: 4230: 4219: 4216: 4215: 4213: 4198: 4190: 4189: 4186: 4180: 4177: 4175: 4172: 4171: 4169: 4165: 4159: 4156: 4154: 4151: 4149: 4146: 4144: 4141: 4139: 4136: 4134: 4131: 4129: 4126: 4124: 4121: 4119: 4116: 4114: 4111: 4109: 4106: 4104: 4101: 4099: 4096: 4094: 4091: 4089: 4086: 4084: 4081: 4079: 4076: 4074: 4071: 4070: 4068: 4064: 4058: 4055: 4053: 4050: 4048: 4045: 4043: 4040: 4038: 4035: 4033: 4030: 4028: 4025: 4024: 4022: 4019: 4015: 4009: 4006: 4004: 4001: 3999: 3996: 3994: 3991: 3989: 3986: 3984: 3981: 3979: 3976: 3974: 3971: 3969: 3966: 3965: 3963: 3959: 3953: 3950: 3948: 3945: 3943: 3940: 3938: 3935: 3933: 3930: 3928: 3925: 3923: 3920: 3918: 3915: 3913: 3910: 3908: 3905: 3904: 3902: 3898: 3892: 3889: 3887: 3884: 3882: 3879: 3877: 3874: 3872: 3869: 3867: 3864: 3863: 3860: 3856: 3849: 3844: 3842: 3837: 3835: 3830: 3829: 3826: 3814: 3811: 3809: 3808:Biotechnology 3806: 3804: 3801: 3799: 3796: 3795: 3792: 3786: 3783: 3781: 3778: 3776: 3773: 3771: 3768: 3767: 3765: 3761: 3755: 3752: 3750: 3749:South America 3747: 3744: 3740: 3739:North America 3737: 3735: 3732: 3730: 3727: 3725: 3722: 3721: 3719: 3715: 3709: 3706: 3705: 3703: 3701: 3697: 3691: 3688: 3686: 3685:Transhumanism 3683: 3681: 3678: 3676: 3673: 3671: 3668: 3666: 3663: 3661: 3658: 3656: 3653: 3652: 3650: 3644: 3634: 3631: 3629: 3626: 3624: 3623:Gene knockout 3621: 3620: 3618: 3614: 3608: 3605: 3603: 3600: 3598: 3595: 3594: 3592: 3587: 3582: 3574: 3571: 3569: 3566: 3564: 3561: 3559: 3556: 3554: 3551: 3549: 3546: 3545: 3543: 3541: 3538: 3534: 3533:Controversies 3531: 3530: 3529: 3526: 3525: 3523: 3519: 3516: 3512: 3502: 3499: 3497: 3494: 3492: 3489: 3488: 3486: 3482: 3476: 3473: 3471: 3468: 3466: 3463: 3461: 3458: 3456: 3453: 3451: 3448: 3446: 3443: 3441: 3438: 3437: 3435: 3433: 3432:Inserting DNA 3429: 3426: 3424: 3420: 3410: 3407: 3405: 3402: 3400: 3397: 3396: 3394: 3392: 3387: 3383: 3373: 3370: 3368: 3365: 3361: 3358: 3356: 3353: 3352: 3351: 3348: 3346: 3343: 3342: 3340: 3338:Other animals 3336: 3330: 3327: 3325: 3322: 3320: 3317: 3315: 3312: 3308: 3305: 3303: 3300: 3299: 3298: 3295: 3294: 3292: 3290: 3286: 3283: 3281: 3277: 3267: 3264: 3262: 3259: 3257: 3254: 3252: 3249: 3247: 3244: 3242: 3239: 3237: 3234: 3232: 3229: 3227: 3224: 3222: 3219: 3217: 3214: 3213: 3211: 3207: 3201: 3198: 3197: 3195: 3193: 3189: 3183: 3180: 3179: 3177: 3175: 3171: 3165: 3162: 3161: 3159: 3157: 3153: 3147: 3144: 3142: 3139: 3138: 3136: 3134: 3130: 3124: 3121: 3120: 3118: 3116: 3112: 3106: 3103: 3102: 3100: 3098: 3094: 3088: 3085: 3083: 3080: 3078: 3075: 3073: 3070: 3069: 3067: 3065: 3061: 3058: 3056: 3052: 3049: 3047: 3039: 3035: 3028: 3023: 3021: 3016: 3014: 3009: 3008: 3005: 2999: 2995: 2992: 2989: 2988: 2976: 2972: 2968: 2964: 2960: 2956: 2953:(2): 301–13. 2952: 2948: 2941: 2922: 2915: 2899: 2895: 2888: 2880: 2876: 2869: 2853: 2849: 2842: 2826: 2820: 2812: 2808: 2804: 2800: 2796: 2792: 2788: 2784: 2777: 2770: 2754: 2747: 2739: 2735: 2730: 2725: 2721: 2717: 2713: 2709: 2705: 2698: 2682: 2675: 2667: 2663: 2659: 2655: 2651: 2647: 2643: 2639: 2632: 2616: 2610: 2595: 2591: 2587: 2581: 2577: 2573: 2569: 2568: 2560: 2544: 2538: 2530: 2524: 2505: 2499: 2484: 2478: 2474: 2473:10.2777/97784 2470: 2463: 2462: 2454: 2448: 2437: 2418: 2412: 2406: 2403: 2399: 2395: 2391: 2387: 2383: 2379: 2375: 2369: 2366: 2363: 2359: 2355: 2351: 2347: 2343: 2339: 2335: 2329: 2328:And contrast: 2326: 2317:on 2016-02-07 2313: 2309: 2305: 2301: 2297: 2293: 2289: 2282: 2276: 2273: 2269: 2265: 2261: 2257: 2253: 2249: 2245: 2238: 2232:But see also: 2229: 2221: 2217: 2212: 2207: 2203: 2199: 2195: 2191: 2187: 2180: 2164: 2158: 2144:on 2016-09-17 2140: 2136: 2132: 2128: 2124: 2120: 2116: 2112: 2108: 2101: 2094: 2079: 2075: 2069: 2054: 2048: 2041: 2035: 2016: 2009: 1995: 1991: 1985: 1971: 1967: 1961: 1954: 1950: 1947: 1942: 1934: 1930: 1926: 1922: 1918: 1914: 1910: 1906: 1899: 1885: 1881: 1875: 1867: 1863: 1859: 1855: 1850: 1845: 1841: 1837: 1833: 1829: 1822: 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