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Rail (bird)

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benefit the population as an entirety, rather than many young that would exhibit less fitness. Unfortunately, with the human occupation of most islands in the past 5,000 to 35,000 years, selection has undoubtedly reversed the tolerance into a wariness of humans and predators, causing species unequipped for the change to become susceptible to extinction.
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Hugueney, Marguerite; Berthet, Didier; Bodergat, Anne-Marie; Escuillié, François; Mourer-Chauviré, Cécile; Wattinne, Aurélia (2003). "La limite OligocÚne-MiocÚne en Limagne: changements fauniques chez les mammifÚres, oiseaux et ostracodes des différents niveaux de Billy-Créchy (Allier, France) ".
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demands, reduces the flightless rail's energy expenditures. For this reason, flightlessness makes it easier to survive and colonize an island where resources may be limited. This also allows for the evolution of multiple sizes of flightless rails on the same island as the birds diversify to fill
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exhibit lower aggression and reduced territorial defense behaviors than do their mainland European counterparts, but this tolerance may be limited to close relatives. The resulting kin-selecting altruistic phenomena reallocate resources to produce fewer young that are more competitive and would
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In addition to energy conservation, certain morphological traits also affect rail evolution. Rails have relatively small flight muscles and wings to begin with. In rails, the flight muscles make up only 12–17% of their overall body mass. This, in combination with their terrestrial habits and
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Another factor that contributes to the occurrence of the flightless state is a climate that does not necessitate seasonal long-distance migration; this is evidenced by the tendency to evolve flightlessness at a much greater occurrence in tropical islands than in temperate or polar islands.
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in dense vegetation. In general, they are shy, secretive, and difficult to observe. Most species walk and run vigorously on strong legs, and have long toes that are well adapted to soft, uneven surfaces. They tend to have short, rounded wings, and although they are generally weak
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The rails are a family of small to medium-sized, ground-living birds. They vary in length from 12 to 63 cm (5 to 25 in) and in weight from 20 to 3,000 g (0.7 oz to 6 lb 10 oz). Some species have long necks and in many cases are laterally compressed.
671:, at 13 cm (5.1 in) and 25 g. The larger species are also sometimes given other names. The black coots are more adapted to open water than their relatives, and some other large species are called gallinules and swamphens. The largest of this group is the 466:
in the animal kingdom. Of the roughly 150 historically known rail species, 31 extant or recently extinct species evolved flightlessness from volant (flying) ancestors. This process created the endemic populations of flightless rails seen on Pacific islands today.
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of Rallidae species vary and are often quite loud. Some are whistle-like or squeak-like, while others seem unbirdlike. Loud calls are useful in dense vegetation, or at night where seeing another member of the species is difficult. Some calls are
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to lose the power of flight and evolve the reduced, stubby wings only useful to keep balance when running quickly. Indeed, some argue that measuring the evolution of flightlessness in rails in generations rather than millennia might be possible.
540:, although the species is clearly polyphyletic (it has more than one ancestral species), it is not the ancestor of most of its flightless descendants, revealing that the flightless condition evolved in rails before speciation was complete. 715:
as small as one or as large as 15 eggs are known. Egg clutches may not always hatch at the same time. Chicks become mobile after a few days. They often depend on their parents until fledging, which happens around 1 month old.
536:, exhibit a persistently high ability to disperse long distances among tropic Pacific islands, though only the latter two gave rise to flightless endemic species throughout the Pacific Basin. In examining the phylogeny of 455:, a characteristic that has led them to colonize many isolated oceanic islands. Furthermore, these birds often prefer to run rather than fly, especially in dense habitat. Some are also flightless at some time during their 3396: 862:
below showing the phylogeny of the living and recently extinct Rallidae is based on a study by Juan Garcia-R and collaborators published in 2020. The genera and number of species are taken from the list maintained by
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The rails have suffered disproportionally from human changes to the environment, and an estimated several hundred species of island rails have become extinct because of this. Several island species of rails remain
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Granjon, L., and G. Cheylan (1989): The fate of black rats (rattus-rattus, l) introduced on an island, as revealed by radio-tracking. Comptes Rendus De L Académie des Sciences, Série III Sciences de la Vie
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A consequence of lowered energy expenditure in flightless island rails has also been associated with evolution of their "tolerance" and "approachability". For example, the (non-Rallidae) Corsican
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It is paradoxical, since rails appear loath to fly, that the evolution of flightless rails would necessitate high dispersal to isolated islands. Nonetheless, three species of small-massed rails,
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Livezey, B. (2003): Evolution of Flightlessness in Rails (Gruiformes: Rallidae): Phylogenetic, Ecomorphological, and Ontogenetic Perspectives. Ornithological Monographs No. 53. (Book)
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GĂĄl, Erika; HĂ­r, JĂĄnos; Kessler, EugĂ©n & KĂłkay, JĂłzsef (1998–99): KözĂ©psĂ”-miocĂ©n Ă”smaradvĂĄnyok, a MĂĄtraszĂ”lĂ”s, RĂĄkĂłczi-kĂĄpolna alatti ĂștbevĂĄgĂĄsbĂłl. I. A MĂĄtraszĂ”lĂ”s 1. lelĂ”hely .
4250:; Fleischer, R.C. (2002). "Rapid, independent evolution of flightlessness in four species of Pacific Island rails (Rallidae): an analysis based on mitochondrial sequence data". 772:
Due to their tendencies towards flightlessness, many island species have been unable to cope with introduced species. The most dramatic human-caused extinctions occurred in the
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GarcĂ­a-R, J.C.; Gibb, G.C.; Trewick, S.A. (2014). "Deep global evolutionary radiation in birds: Diversification and trait evolution in the cosmopolitan bird family Rallidae".
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from the Greater Antilles was formerly considered to be a rail, but based on DNA evidence is now known to be an independent lineage of gruiform more closely related to
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McNab, B.K.; Ellis, H.I. (2006). "Flightless rails endemic to islands have lower energy expenditures and clutch sizes than flighted rails on islands and continents".
839:, this may or may not be correct; it certainly seems more justified than most of the Sibley-Ahlquist proposals. However, such a group would probably also include the 4159: 4402:
Taylor, Barry., van Perlo, Ber. Rails: A Guide to Rails, Crakes, Gallinules and Coots of the World. United Kingdom: Bloomsbury Publishing, 2010. ISBN 9781408135372
804:, but some of the last remaining individuals were taken into captivity and are breeding well, though attempts at reintroduction have met with mixed results. 4776: 3465: 788:, during which an estimated 750–1800 species of birds became extinct, half of which were rails. Some species that came close to extinction, such as the 3828: 2928:"Phylogeny based on ultra-conserved elements clarifies the evolution of rails and allies (Ralloidea) and is the basis for a revised classification" 487:
behavioral flightlessness, is a significant contributor to the rail's remarkably fast loss of flight; as few as 125,000 years were needed for the
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Many island rails are flightless because small island habitats without mammalian predators eliminate the need to fly or move long distances.
2449:(recently extinct; flightless, single island, lost by 1760 to overhunting, destruction of habitat by tortoise hunters, and introduced cats) 4352:; Tennyson, A.J.D.; Jones, C.; McNamara, J.A.; Douglas, B.J. (2007). "Miocene waterfowl and other birds from central Otago, New Zealand". 2393: 3302: 835:, which has been widely accepted in America, raises the family to ordinal level as the Ralliformes. Given uncertainty about gruiform 3889: 4766: 4606: 4645: 4405:
Australia's Amazing Wildlife. 2009, (Original Publisher: Cornell University). United Kingdom: Bay Books, 1985. ISBN 9780858358300
2767: 287:, and flooded fields or open forest. They are especially fond of dense vegetation for nesting. The rail family is found in every 3927:
Dinkins, Walter (2014): The Rail Bird Hunter's Bible. A History of Rail Bird Hunting in the USA. Virtualbookworm.com Publishing.
3723:(1974). "A new species of Nesotrochis from Hispaniola, with notes on other fossil rails from the West Indies (Aves: Rallidae)". 3563: 2360: 3496: 2007: 872: 3435: 4650: 3598: 4190: 4354: 3006: 2727:
Rallidae gen. et sp. indet. UMMP V55013-55014; UMMP V55012/V45750/V45746 (Rexroad Late Pliocene of Saw Rock Canyon, USA)
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The undescribed Fernando de Noronha rail, genus and species undetermined, survived to historic times. The extinct genus
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and flight muscles taking up to 40% of a bird's weight. Reducing the flight muscles, with a corresponding lowering of
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Perret, P.; Blondel, J. (1993). "Experimental-evidence of the territorial defense hypothesis in insular blue tits".
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Guide, Joe. Rail Bird Hunter's Bible. N.p.: Virtualbookworm.com Publishing, Incorporated, 2014. ISBN 9781621374527
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annually. The weakness of their flight, however, means they are easily blown off course, thus making them common
3457: 3056:"Phylogenomic reconstruction sheds light on new relationships and timescale of rails (Aves: Rallidae) evolution" 875:(IOC). The names of the subfamilies and tribes are those proposed by Jeremy Kirchman and collaborators in 2021. 447:
of those Rallidae able to fly, while not powerful, can be sustained for long periods of time, and many species
263:. The family exhibits considerable diversity in its forms, and includes such ubiquitous species as the crakes, 4658: 3486: 2439:(recently extinct; flightless, single island, lost by 1700 to overhunting and introduced pigs, cats and rats) 827:, as well as several smaller families of usually "primitive" midsized amphibious birds, to make up the order 816: 219: 748:
was hunted to extinction by the starving Japanese garrison after the island was cut off from supply during
404:, which is a fleshy, rearward extension of the upper bill. The most complex frontal shield is found in the 4528: 4515: 4168: 4164: 695:
The breeding behaviors of many Rallidae species are poorly understood or unknown. Most are thought to be
4533: 3547: 2302: 864: 4725: 4022:"Speciation of flightless rails on islands: A DNA-based phylogeny of the typical rails of the Pacific" 384:
is the most variable feature within the family. In some species, it is longer than the head (like the
4738: 4686: 4421: 757: 253: 152: 792:, and the takahē, have made modest recoveries due to the efforts of conservation organisations. The 4252: 832: 17: 3786: 2567:(Wasatch Early Eocene of Steamboat Springs, USA; Late Eocene – ?Oligocene of Isfara, Tadzhikistan) 3897: 4712: 4468: 3885: 2851:(Montmartre Late Eocene of France) is sometimes considered a rail and then placed in the genus 2370: 2178: 4291:"Flightlessness and phylogeny amongst endemic rails (Aves:Rallidae) of the New Zealand region" 3938: 2409:(recently extinct; flightless, single island, lost by early 1800s to introduced cats and rats) 279:(such as the coot), but many more are wading birds or shorebirds. The ideal rail habitats are 4707: 4637: 3670: 2481:(recently extinct; flightless, single island, lost by no later than 1400s to introduced rats) 4699: 3827:. Probably from a rail, but it is too damaged to determine its affiliations more precisely: 4554: 4362: 4302: 4109: 3953: 3760: 3244: 2011: 848: 704: 3294: 819:
in 1815. The family has traditionally been grouped with two families of larger birds, the
612:, they are, nevertheless, capable of covering long distances. Island species often become 8: 4068: 3646: 2548: 2463: 700: 696: 511: 4366: 4306: 4113: 4066:
McNab, B.K. (1994). "Energy conservation and the evolution of flightlessness in birds".
3957: 3764: 3248: 2419:(recently extinct; flightless, single island, lost by 1500s to introduced cats and rats) 667:
species are normally called rails irrespective of bill length. The smallest of these is
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Dozens of mostly broken isolated skull and limb bones of a rail or crake the size of a
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Comparative Biochemistry and Physiology Part A: Molecular & Integrative Physiology
4047: 3878: 2429:(recently extinct; flightless, two islands, lost between 1500 and 1700 to overhunting) 2356: 4761: 4694: 4541: 4445: 4336: 4279: 4179: 4148: 4122: 4098:"Minimizing energy expenditure facilitates vertebrate persistence on oceanic islands" 4097: 3969: 3920: 3816: 3555: 3331: 3272: 2978: 2426: 2416: 789: 684: 668: 412: 4392: 4221: 4088: 4055: 3902:) from the Oligo-Miocene of Riversleigh, Northwestern Queensland, Australia. (2005)" 3490: 4378: 4370: 4349: 4326: 4310: 4260: 4209: 4140: 4117: 4076: 4033: 4008: 3961: 3916: 3874: 3768: 3732: 3678: 3650: 3551: 3526: 3425: 3262: 3252: 3067: 2939: 2508: 2081: 868: 797: 745: 520: 256: 4012: 2944: 2927: 2730:
Rallidae gen. et sp. indet. UMMP V29080 (Rexroad Late Pliocene of Fox Canyon, USA)
2507:(recently extinct; flight ability uncertain, single island, lost by no later than 4546: 4247: 4228: 4194: 3863:
Ballmann, Peter (1969). "Les Oiseaux miocĂšnes de la Grive-Saint-Alban (IsĂšre) ".
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In general, members of the Rallidae are omnivorous generalists. Many species eat
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Rallidae gen. et sp. indet. (Bathans Early/Middle Miocene of Otago, New Zealand)
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Rallidae gen. et sp. indet. (Bathans Early/Middle Miocene of Otago, New Zealand)
271:; other rail species are extremely rare or endangered. Many are associated with 4771: 4593: 4580: 4271: 4144: 3708: 3521:. Bulletin of the American Museum of Natural History. Vol. 222. New York: 3383: 3323: 2895: 2871: 2718:
Rallidae gen. et sp. indet. (Sajóvölgyi Middle Miocene of MåtraszÔlÔs, Hungary)
2504: 2489: 2446: 2355:). These have not been listed here; see the genus accounts and the articles on 2203: 2133: 840: 753: 663:, long-billed species tend to be called rails and short-billed species crakes. 597: 532: 448: 401: 369: 303: 4374: 3772: 2998: 4755: 3430: 3290: 2754: 2745: 2648: 2556: 2522: 2320: 2293: 2153: 2072: 2047: 2028: 1483: 773: 712: 708: 664: 503: 299: 136: 86: 3257: 687:
organisations and governments continue to work to prevent their extinction.
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Additionally, many prehistoric rails of extant genera are known only from
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Rallidae gen. et sp. indet. (Middle Miocene of Grive-Saint-Alban, France)
2544: 2517: 2250: 2229:– swamphens and purple gallinules (10 living species, 2 recently extinct) 2139: 2036: 1836: 1689: 1407: 1283: 1258: 935: 656: 609: 499: 488: 475: 471: 444: 405: 61: 35: 3233:"Magnitude and variation of prehistoric bird extinctions in the Pacific" 2708:(Middle Miocene of Sansan, France -? Late Miocene of Rudabánya, Hungary) 2010:(IOC) contains 152 species divided into 43 genera. For more detail, see 1601:– swamphens, gallinules and takahes (12 species of which 2 are extinct) 4213: 2667: 2659:(Late Oligocene/?Early Miocene -? Middle Miocene of France) – includes 2478: 2366: 2348: 2335: 2258: 2242: 2098: 2087: 1913: 1707: 1655: 1637: 1233: 1122: 828: 785: 617: 577: 479: 327: 311: 284: 204: 106: 71: 4383: 4295:
Philosophical Transactions of the Royal Society B: Biological Sciences
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Analyse de la nature ou, Tableau de l'univers et des corps organisés
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Rallidae gen. et sp. indet. (Late Oligocene of Billy-Créchy, France)
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Neornithes incerta sedis (Late Oligocene of Riversleigh, Australia)
2759: 2678:(Late Oligocene? – Late Miocene of C Europe) – possibly belongs in 2526: 2493: 2436: 2374: 2266: 2192: 2053: 1818: 1758: 1458: 987: 813: 732:
is an example of an island species that has been badly affected by
652: 637: 593: 544: 184: 128: 101: 96: 81: 76: 66: 2559:, as well from the less comprehensively studied strata elsewhere: 3999: 3865: 3295:"Memorializing the Wake Island Rail: An Extinction Caused by War" 2724:
Rallidae gen. et sp. indet. (Late Miocene of Lemoyne Quarry, USA)
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Worthy, T.H. (1997). "A mid-Pleistocene rail from New Zealand".
3397:"Guam: Brown Tree Snake Responsible for Extinction of 5 Species" 3231:
Duncan, Richard P.; Boyer, Alison G.; Blackburn, Tim M. (2013).
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Garcia-R, J.C.; Lemmon, E.M.; Lemmon, A.R.; French, N. (2020).
2552: 2537: 2344: 2061: 1031: 633: 295: 174: 4450: 3677:(MNZ S.42658, S.42785) of a rail or crake the size of a large 2974:
Lapwings, Loons and Lousy Jacks: The How and Why of Bird Names
2737:(Blanco Late Pliocene/Early Pleistocene of Crosby County, USA) 4720: 4559: 3674: 2845: 2671:(Late Oligocene – Middle Miocene of NW Queensland, Australia) 852: 836: 812:
The family Rallidae was introduced (as Rallia) by the French
724: 589: 585: 560:, as well as fruit or seedlings. A few species are primarily 456: 342: 280: 264: 259:
of small- to medium-sized terrestrial and/or semi-amphibious
3458:"Is this Guam bird coming back from extinction in the wild?" 2171:– Austropacific rails (8 living species, 4 recently extinct) 675:, at 65 cm (26 in) and 2.7 kg (6.0 lb). 4348: 2816:(Bridger middle Eocene of Forbidden City, USA) – phasianid? 2217: 2144: 1526: 1147: 801: 581: 393: 381: 260: 194: 4235:
Farner, D.S.; King, J.R. & Parkes, Kenneth C. (eds.):
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Horsfall, Joseph A. & Robinson, Robert (2003): Rails.
3556:"Flufftails, finfoots, rails, trumpeters, cranes, limpkin" 3053: 2701:(Anversian Black Sand Middle Miocene of Antwerp, Belgium) 645: 641: 629: 625: 356: 2588:(Bridger Middle Eocene of Uinta County, USA) – includes 1305:– rails and woodhen (12 species of which 4 are extinct) 3140: 3138: 3136: 2643:(Late Oligocene/Early Miocene – Late Miocene of France) 1762:– rails and crakes (15 species of which 5 are extinct) 462:
Flightlessness in rails is one of the best examples of
310:). Members of Rallidae occur on every continent except 2652:(Late Oligocene/Early Miocene of Mainz Basin, Germany) 4276:
Extinction and Biogeography of Tropical Pacific Birds
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Extinction and Biogeography of Tropical Pacific Birds
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rails are richly documented from the well-researched
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Alcheringa: An Australasian Journal of Palaeontology
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History and Nomenclature of Avian Family-Group Names
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Kirchman, J.J.; Rotzel McInerney, N.; Giarla, T.C.;
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Rallidae gen. et sp. indet. (Bermuda, West Atlantic)
2633:(Corbula Middle/Late Oligocene of KolzsvĂĄr, Romania) 707:
have been reported. Most often, they lay five to 10
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Proceedings of the Biological Society of Washington
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Smithsonian Libraries - Smithsonian Research Online
2957: 2955: 368:) from behind, showing the short, soft, and fluffy 3495:(in French). Palermo: Self-published. p. 70. 3361: 3359: 3344: 2163:– (1 possibly extinct species, 1 recently extinct) 855:, and usually united with the rails in the Ralli. 443:The wings of all rails are short and rounded. The 392:); in others, it may be short and wide (as in the 322:"Rail" is the anglicized respelling of the French 3037: 3035: 3025: 3023: 2807:(Irdin Manha Late Eocene of Chimney Butte, China) 2221:– coots (10 living species, one recently extinct) 2213:– moorhens (5 living species, 2 recently extinct) 4753: 3353:Horsfall & Robinson (2003): pp. 209–210 3190: 3188: 2961:Horsfall & Robinson (2003): pp. 206–207 2952: 596:are a particularly favoured habitat. Those that 3356: 655:species are secretive (apart from loud calls), 3080: 3032: 3020: 2917: 2915: 1512:– moorhens (7 species of which 2 are extinct) 659:, and have laterally flattened bodies. In the 235: 4199: 3898:"A New Flightless Gallinule (Aves: Rallidae: 3185: 3178: 3176: 3174: 3172: 3117: 3115: 576:The most typical family members occupy dense 30:"Crake" redirects here. For the surname, see 3330:. University of Chicago Press. p. 296. 3215: 3049: 3047: 2598:(Bridger Middle Eocene of Henry's Fork, USA) 807: 438: 314:. Numerous unique island species are known. 4777:Taxa named by Constantine Samuel Rafinesque 4175:Cenozoic Birds of the World, Part 1: Europe 3890:Wake Island Rail BirdLife Species Factsheet 3793:. Museum of New Zealand Te Papa Tongarewa. 3694:Several limb bones of a smallish rail: GĂĄl 3221:Horsfall & Robinson (2003): p. 207 2912: 2779:These taxa may or may not have been rails: 1530:– coots (11 species of which 1 is extinct) 767: 4130: 3932:Folia Historico Naturalia Musei Matraensis 3485: 3169: 3112: 2823:(Early Oligocene of Trigonias Quarry, USA) 2388: 1126:– rails (2 species of which 1 is extinct) 620:following the introduction of terrestrial 127: 4382: 4330: 4121: 4037: 3266: 3256: 3071: 3044: 2943: 2774: 2715:(Shanwang Middle Miocene of Linqu, China) 2619:(Late Eocene -? Late Oligocene of France) 2605:(Middle Eocene of Lee-on-Solent, England) 2270:– (10 living species, 5 recently extinct) 843:(finfoots and sungrebes), an exclusively 796:came perilously close to extinction when 4019: 3862: 3322: 2392: 2365: 2091:– (1 living species, 1 recently extinct) 723: 551: 498: 355: 4288: 4231:(1985): Section X.D.2.b. Scolopacidae. 3562:. International Ornithologists' Union. 2970: 2768:Museum of New Zealand Te Papa Tongarewa 744:and their eggs collected for food. The 275:habitats, some being semi-aquatic like 14: 4754: 3750: 3542: 3540: 3365:Horsfall & Robinson (2003): p. 211 3086:Horsfall & Robinson (2003): p. 209 3041:Horsfall & Robinson (2003): p. 210 3029:Horsfall & Robinson (2003): p. 208 2977:. Exeter, UK: Pelagic Publishing Ltd. 2926:; Slikas, E.; Fleischer, R.C. (2021). 2743:Rallidae gen. et sp. indet. (formerly 2612:(Hordwell Late Eocene of Hordwell, UK) 2574:(Wasatch Early Eocene of Wyoming, USA) 2008:International Ornithological Committee 873:International Ornithological Committee 719: 333:. It is named from its harsh cry, in 4467: 4466: 4095: 4065: 3946:Molecular Phylogenetics and Evolution 3895: 3593: 3289: 2006:The list maintained on behalf of the 1917:– waterhen and bush-hens (5 species) 740:Some larger, more abundant rails are 400:). A few coots and gallinules have a 4726:90521459-8557-FFAA-FE8C-2D741630F9E1 4687:817745b3-ec2c-4890-974d-57b73dab7122 4599:99ffda06-d1f0-4dda-bebb-00a3f6b9e5c8 4242:: 174–175. Academic Press, New York. 3797:from the original on 24 October 2012 3516: 3096: 3094: 3092: 3599:"A Synopsis of the Fossil Rallidae" 3537: 3455: 2758:(mid-Pleistocene New Zealand). The 871:and David Donsker on behalf of the 776:as people colonised the islands of 24: 3856: 3823:piece of a bird about the size of 3787:"Pleistorallus flemingi; holotype" 3632:A small species of rail: Hugueney 3566:from the original on 14 March 2023 3523:American Museum of Natural History 2581:(Early – Middle Eocene of England) 2347:or subfossil remains, such as the 25: 4788: 4409: 4265:10.1034/j.1600-048X.2002.330103.x 3426:"Guam Rail (Gallirallus owstoni)" 3089: 2361:Late Quaternary prehistoric birds 4453:The New Student's Reference Work 4429: 4415: 4123:10.1046/j.1461-0248.2002.00365.x 3921:10.3853/j.0067-1975.57.2005.1441 3909:Records of the Australian Museum 3560:IOC World Bird List Version 12.2 2901:List of Gruiformes by population 2889: 2877: 2865: 2749:) (Late Pleistocene of Barbados) 2532: 2001: 1711:– crakes and rails (13 species) 1693:– rails and a crake (3 species) 474:makes intense demands, with the 419:or size. Two exceptions are the 151: 59: 4767:Extant Eocene first appearances 4278:. University of Chicago Press. 3841: 3809: 3779: 3744: 3714: 3701: 3688: 3660: 3639: 3626: 3615:from the original on 2018-07-29 3587: 3578: 3510: 3499:from the original on 2018-08-19 3479: 3468:from the original on 2018-07-29 3449: 3438:from the original on 2018-07-29 3418: 3407:from the original on 2018-07-29 3389: 3377: 3368: 3316: 3305:from the original on 2019-06-02 3283: 3224: 3206: 3197: 3156: 3147: 3124: 3103: 3009:from the original on 2021-06-10 2397:1888 color lithograph of a rail 690: 3487:Rafinesque, Constantine Samuel 2991: 2964: 2688:(Early Miocene of New Zealand) 2626:(Early Oligocene of WC Europe) 1437:– crakes and sora (3 species) 13: 1: 4013:10.1016/j.geobios.2003.01.002 3990:Firefly Encyclopedia of Birds 3879:10.1016/S0016-6995(69)80005-7 2906: 1822:– New Guinea flightless rail 817:Constantine Samuel Rafinesque 510:) skeleton on display at the 351: 34:. For the English river, see 3988:Perrins, Christopher (ed.): 3711:-sized rail: Ballmann (1969) 2830:(Early Oligocene of Germany) 2766:is in the collection of the 2065:– typical rails (14 species) 752:. At least two species, the 7: 3966:10.1016/j.ympev.2014.09.008 2945:10.1093/ornithology/ukab042 2858: 616:, and many of them are now 372:typical of flightless rails 10: 4793: 4145:10.1016/j.cbpa.2006.02.025 3462:BirdLife International.org 2840:(Pleistocene of Argentina) 2303:New Guinea flightless rail 580:in damp environments near 411:Rails exhibit very little 294:with the exception of dry 29: 4475: 4375:10.1017/S1477201906001957 3773:10.1080/03115519708619186 2987:– via Google Books. 2789:(Late Eocene) – includes 1910: 1892: 1885: 1867: 1860: 1833: 1815: 1808: 1801: 1773: 1755: 1748: 1740: 1704: 1686: 1679: 1652: 1634: 1627: 1620: 1612: 1594: 1586: 1523: 1505: 1498: 1480: 1473: 1462:– nativehens (2 species) 1455: 1448: 1430: 1423: 1404: 1396: 1389: 1298: 1280: 1273: 1255: 1248: 1230: 1223: 1205: 1198: 1162: 1144: 1137: 1119: 1112: 1105: 1078: 1060: 1053: 1046: 1028: 1021: 984: 966: 959: 932: 914: 907: 900: 892: 884: 808:Systematics and evolution 758:American purple gallinule 439:Flight and flightlessness 396:), or massive (as in the 302:or freezing regions, and 232: 227: 148:Scientific classification 146: 135: 126: 45: 4020:Kirchman, J. J. (2012). 3896:Boles, Walter E (2005). 3850:(1985), MlĂ­kovskĂœ (2002) 3517:Bock, Walter J. (1994). 3212:Granjon and Cheylan 1989 3109:McNab & Ellis (2006) 2844:The presumed scolopacid 2196:– nativehens (2 species) 2057:– wood rails (8 species) 833:Sibley-Ahlquist taxonomy 768:Threats and conservation 521:Gallirallus philippensis 234:Some 40 living, and see 4289:Trewick, S. A. (1997). 4272:Steadman, David William 4177:. Ninox Press, Prague. 3892:. Retrieved 2007-07-04. 3258:10.1073/pnas.1216511110 3203:Perret and Blondel 1993 3003:Encyclopedia Britannica 2795:"Palaeortyx" blanchardi 2466:(sometimes included in 2389:Recently extinct genera 2373:'s 1617 drawing of the 2339:– bush-hens (5 species) 847:group that is somewhat 760:, have been considered 317: 4315:10.1098/rstb.1997.0031 4048:10.1525/auk.2011.11096 4039:10.1525/auk.2011.11096 3886:BirdLife International 3554:, eds. (August 2022). 2775:Doubtfully placed here 2764:Pleistorallus flemingi 2470:; extinct around 1900) 2398: 2385: 2371:Pieter van den Broecke 2179:spot-flanked gallinule 737: 515: 373: 366:Porphyrio hochstetteri 4708:Paleobiology Database 3525:. pp. 136, 252. 2971:Reedman, Ray (2016). 2396: 2369: 1840:– white-browed crake 1780:– crakes (4 species) 1659:– crakes (2 species) 1035:– rails (14 species) 921:– crakes (2 species) 727: 552:Behaviour and ecology 502: 359: 4594:Fauna Europaea (new) 4426:at Wikimedia Commons 4355:J. Syst. Palaeontol. 4096:McNab, B.K. (2002). 3144:McNab and Ellis 2006 2383:Aphanapteryx bonasia 2012:List of rail species 1169:– rails (4 species) 991:– rails (8 species) 939:– rails (3 species) 4367:2007JSPal...5....1W 4307:1997RSPTB.352..429T 4114:2002EcolL...5..693M 3958:2014MolPE..81...96G 3765:1997Alch...21...71W 3249:2013PNAS..110.6436D 2464:New Caledonian rail 2363:for these species. 800:were introduced to 720:Rallidae and humans 527:Porphyrio porphyrio 512:Museum of Osteology 141:Gallinula tenebrosa 4214:10.1007/bf01928800 4193:2011-05-20 at the 3791:Collections Online 3707:Partial hand of a 3550:; Donsker, David; 3324:Steadman, David W. 2896:Biology portal 2872:Animals portal 2791:"Tringa" hoffmanni 2399: 2386: 2353:Rallus eivissensis 2312:white-browed crake 2023:grey-throated rail 1641:– ocellated crake 831:. The alternative 738: 734:introduced species 516: 464:parallel evolution 374: 4749: 4748: 4695:Open Tree of Life 4469:Taxon identifiers 4420:Media related to 4350:Worthy, Trevor H. 4301:(1352): 429–446. 4186: 3552:Rasmussen, Pamela 3403:. February 1996. 3243:(16): 6436–6441. 3073:10.3390/d12020070 2417:Saint Helena rail 1998: 1997: 1989: 1988: 1980: 1979: 1971: 1970: 1962: 1961: 1953: 1952: 1944: 1943: 1935: 1934: 1926: 1925: 1849: 1848: 1789: 1788: 1729: 1728: 1720: 1719: 1668: 1667: 1575: 1574: 1566: 1565: 1557: 1556: 1548: 1547: 1539: 1538: 1487:– lesser moorhen 1387:Himantornithinae 1377: 1376: 1368: 1367: 1359: 1358: 1350: 1349: 1341: 1340: 1332: 1331: 1323: 1322: 1314: 1313: 1237:– invisible rail 1187: 1186: 1178: 1177: 1094: 1093: 1009: 1008: 1000: 999: 948: 947: 798:brown tree snakes 790:Lord Howe woodhen 533:Porzana tabuensis 425:Gallicrex cinerea 413:sexual dimorphism 398:purple gallinules 306:areas (above the 283:areas, including 243: 242: 223: 57: 16:(Redirected from 4784: 4742: 4741: 4729: 4728: 4716: 4715: 4703: 4702: 4690: 4689: 4680: 4679: 4667: 4666: 4664:NBNSYS0000160870 4654: 4653: 4641: 4640: 4628: 4627: 4615: 4614: 4602: 4601: 4589: 4588: 4576: 4575: 4563: 4562: 4550: 4549: 4537: 4536: 4524: 4523: 4511: 4510: 4509: 4496: 4495: 4494: 4464: 4463: 4459: 4457: 4448: 4434:Data related to 4433: 4419: 4396: 4386: 4344: 4334: 4268: 4248:Olson, Storrs L. 4229:Olson, Storrs L. 4225: 4178: 4172: 4156: 4127: 4125: 4092: 4059: 4041: 4016: 3992:. Firefly Books. 3977: 3924: 3906: 3882: 3851: 3845: 3839: 3837: 3813: 3807: 3806: 3804: 3802: 3783: 3777: 3776: 3748: 3742: 3740: 3721:Olson, Storrs L. 3718: 3712: 3705: 3699: 3692: 3686: 3679:buff-banded rail 3664: 3658: 3651:buff-banded rail 3643: 3637: 3630: 3624: 3623: 3621: 3620: 3614: 3603: 3595:Olson, Storrs L. 3591: 3585: 3584:MlĂ­kovskĂœ (2002) 3582: 3576: 3575: 3573: 3571: 3544: 3535: 3534: 3514: 3508: 3507: 3505: 3504: 3483: 3477: 3476: 3474: 3473: 3456:Hurrell, Shaun. 3453: 3447: 3446: 3444: 3443: 3422: 3416: 3415: 3413: 3412: 3393: 3387: 3381: 3375: 3372: 3366: 3363: 3354: 3351: 3342: 3341: 3320: 3314: 3313: 3311: 3310: 3299:TheRevelator.org 3287: 3281: 3280: 3270: 3260: 3228: 3222: 3219: 3213: 3210: 3204: 3201: 3195: 3192: 3183: 3180: 3167: 3160: 3154: 3151: 3145: 3142: 3131: 3128: 3122: 3119: 3110: 3107: 3101: 3098: 3087: 3084: 3078: 3077: 3075: 3051: 3042: 3039: 3030: 3027: 3018: 3017: 3015: 3014: 2995: 2989: 2988: 2968: 2962: 2959: 2950: 2949: 2947: 2919: 2894: 2893: 2884:Birds portal 2882: 2881: 2880: 2870: 2869: 2746:Fulica podagrica 2540:species of long- 2509:Late Pleistocene 1888: 1887: 1874:– striped crake 1863: 1862: 1811: 1810: 1804: 1803: 1751: 1750: 1743: 1742: 1682: 1681: 1630: 1629: 1623: 1622: 1615: 1614: 1589: 1588: 1501: 1500: 1476: 1475: 1451: 1450: 1426: 1425: 1411:– Nkulengu rail 1402:Himantornithini 1399: 1398: 1392: 1391: 1287:– chestnut rail 1276: 1275: 1251: 1250: 1226: 1225: 1201: 1200: 1140: 1139: 1115: 1114: 1108: 1107: 1085:– Rouget's rail 1067:– African crake 1056: 1055: 1049: 1048: 1024: 1023: 973:– uniform crake 962: 961: 910: 909: 903: 902: 895: 894: 887: 886: 879: 878: 869:Pamela Rasmussen 746:Wake Island rail 600:do so at night. 508:Fulica americana 218: 156: 155: 131: 121: 58: 52: 49:Temporal range: 43: 42: 21: 4792: 4791: 4787: 4786: 4785: 4783: 4782: 4781: 4752: 4751: 4750: 4745: 4737: 4732: 4724: 4719: 4711: 4706: 4698: 4693: 4685: 4683: 4675: 4670: 4662: 4657: 4649: 4644: 4636: 4631: 4623: 4618: 4610: 4605: 4597: 4592: 4584: 4579: 4571: 4566: 4558: 4553: 4545: 4540: 4532: 4527: 4519: 4514: 4505: 4504: 4499: 4490: 4489: 4484: 4471: 4443: 4412: 4195:Wayback Machine 4162: 4160:MlĂ­kovskĂœ, JirĂ­ 4102:Ecology Letters 3904: 3859: 3857:Further reading 3854: 3846: 3842: 3831: 3821:carpometacarpus 3814: 3810: 3800: 3798: 3785: 3784: 3780: 3749: 3745: 3736: 3731:(38): 439–450. 3719: 3715: 3706: 3702: 3693: 3689: 3673:S.40957) and 2 3665: 3661: 3644: 3640: 3631: 3627: 3618: 3616: 3612: 3601: 3592: 3588: 3583: 3579: 3569: 3567: 3545: 3538: 3515: 3511: 3502: 3500: 3484: 3480: 3471: 3469: 3454: 3450: 3441: 3439: 3424: 3423: 3419: 3410: 3408: 3395: 3394: 3390: 3382: 3378: 3373: 3369: 3364: 3357: 3352: 3345: 3338: 3321: 3317: 3308: 3306: 3288: 3284: 3229: 3225: 3220: 3216: 3211: 3207: 3202: 3198: 3193: 3186: 3181: 3170: 3161: 3157: 3152: 3148: 3143: 3134: 3129: 3125: 3120: 3113: 3108: 3104: 3100:Kirchman (2012) 3099: 3090: 3085: 3081: 3052: 3045: 3040: 3033: 3028: 3021: 3012: 3010: 2997: 2996: 2992: 2985: 2969: 2965: 2960: 2953: 2920: 2913: 2909: 2888: 2878: 2876: 2864: 2861: 2849:Limosa gypsorum 2777: 2657:Paraortygometra 2535: 2407:Ascension crake 2391: 2317:Aenigmatolimnas 2237:ocellated crake 2004: 1999: 1990: 1981: 1972: 1963: 1954: 1945: 1936: 1927: 1871:Aenigmatolimnas 1850: 1799:Amaurornithini 1790: 1730: 1721: 1669: 1576: 1567: 1558: 1549: 1540: 1378: 1369: 1360: 1351: 1342: 1333: 1324: 1315: 1212:– Calayan rail 1188: 1179: 1095: 1010: 1001: 949: 882:Rallidae  810: 770: 722: 693: 554: 538:G. philippensis 441: 354: 348:("to scrape"). 320: 252:) are a large, 217: 150: 122: 120: 119: 114: 109: 104: 99: 94: 89: 84: 79: 74: 69: 64: 51: 50: 47: 39: 32:Crake (surname) 28: 27:Family of birds 23: 22: 15: 12: 11: 5: 4790: 4780: 4779: 4774: 4769: 4764: 4747: 4746: 4744: 4743: 4730: 4717: 4704: 4691: 4681: 4668: 4655: 4642: 4629: 4616: 4603: 4590: 4581:Fauna Europaea 4577: 4564: 4551: 4538: 4525: 4512: 4497: 4481: 4479: 4473: 4472: 4461: 4460: 4441: 4440:at Wikispecies 4427: 4411: 4410:External links 4408: 4407: 4406: 4403: 4400: 4397: 4346: 4286: 4269: 4253:J. Avian Biol. 4243: 4226: 4197: 4157: 4139:(3): 295–311. 4128: 4108:(5): 693–704. 4093: 4081:10.1086/285697 4075:(4): 628–642. 4063: 4060: 4017: 4007:(6): 719–731. 3993: 3982: 3978: 3941: 3928: 3925: 3915:(2): 179–190. 3893: 3888:(BLI) (2007): 3883: 3858: 3855: 3853: 3852: 3840: 3808: 3778: 3743: 3713: 3709:common moorhen 3700: 3687: 3659: 3647:slaty-breasted 3638: 3625: 3586: 3577: 3536: 3509: 3478: 3448: 3417: 3388: 3376: 3367: 3355: 3343: 3336: 3315: 3291:Platt, John R. 3282: 3223: 3214: 3205: 3196: 3184: 3168: 3155: 3146: 3132: 3130:Trewick (1997) 3123: 3111: 3102: 3088: 3079: 3043: 3031: 3019: 2990: 2983: 2963: 2951: 2938:(4): ukab042. 2910: 2908: 2905: 2904: 2903: 2898: 2886: 2874: 2860: 2857: 2842: 2841: 2834: 2831: 2824: 2817: 2808: 2801: 2799:"P." hoffmanni 2776: 2773: 2772: 2771: 2750: 2741: 2738: 2731: 2728: 2725: 2722: 2719: 2716: 2709: 2702: 2695: 2692: 2689: 2682: 2680:Palaeoaramides 2672: 2663: 2653: 2644: 2641:Palaeoaramides 2637: 2634: 2627: 2620: 2613: 2606: 2599: 2596:Fulicaletornis 2592: 2582: 2575: 2568: 2534: 2531: 2513: 2512: 2505:Hova gallinule 2497: 2490:Viti Levu rail 2482: 2471: 2450: 2447:Rodrigues rail 2440: 2430: 2427:Hawkins's rail 2423:Diaphorapteryx 2420: 2410: 2390: 2387: 2341: 2340: 2332: 2323: 2314: 2305: 2296: 2287: 2279: 2271: 2263: 2262:– (13 species) 2255: 2247: 2239: 2230: 2222: 2214: 2206: 2204:lesser moorhen 2197: 2189: 2181: 2172: 2164: 2156: 2147: 2136: 2134:invisible rail 2127: 2118: 2110: 2101: 2092: 2084: 2075: 2066: 2058: 2050: 2041: 2033: 2025: 2003: 2000: 1996: 1995: 1992: 1991: 1987: 1986: 1983: 1982: 1978: 1977: 1974: 1973: 1969: 1968: 1965: 1964: 1960: 1959: 1956: 1955: 1951: 1950: 1947: 1946: 1942: 1941: 1938: 1937: 1933: 1932: 1929: 1928: 1924: 1923: 1920: 1919: 1909: 1906: 1905: 1902: 1901: 1891: 1886: 1884: 1881: 1880: 1877: 1876: 1866: 1861: 1859: 1856: 1855: 1852: 1851: 1847: 1846: 1843: 1842: 1832: 1829: 1828: 1825: 1824: 1814: 1809: 1807: 1802: 1800: 1796: 1795: 1792: 1791: 1787: 1786: 1783: 1782: 1772: 1769: 1768: 1765: 1764: 1754: 1749: 1747: 1741: 1739: 1736: 1735: 1732: 1731: 1727: 1726: 1723: 1722: 1718: 1717: 1714: 1713: 1703: 1700: 1699: 1696: 1695: 1685: 1680: 1678: 1675: 1674: 1671: 1670: 1666: 1665: 1662: 1661: 1651: 1648: 1647: 1644: 1643: 1633: 1628: 1626: 1621: 1619: 1613: 1611: 1608: 1607: 1604: 1603: 1593: 1592:Porphyrionini 1587: 1585: 1582: 1581: 1578: 1577: 1573: 1572: 1569: 1568: 1564: 1563: 1560: 1559: 1555: 1554: 1551: 1550: 1546: 1545: 1542: 1541: 1537: 1536: 1533: 1532: 1522: 1519: 1518: 1515: 1514: 1504: 1499: 1497: 1494: 1493: 1490: 1489: 1479: 1474: 1472: 1469: 1468: 1465: 1464: 1454: 1449: 1447: 1444: 1443: 1440: 1439: 1429: 1424: 1422: 1418: 1417: 1414: 1413: 1403: 1397: 1395: 1390: 1388: 1384: 1383: 1380: 1379: 1375: 1374: 1371: 1370: 1366: 1365: 1362: 1361: 1357: 1356: 1353: 1352: 1348: 1347: 1344: 1343: 1339: 1338: 1335: 1334: 1330: 1329: 1326: 1325: 1321: 1320: 1317: 1316: 1312: 1311: 1308: 1307: 1297: 1294: 1293: 1290: 1289: 1279: 1274: 1272: 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157: 144: 143: 133: 132: 124: 123: 115: 110: 105: 100: 95: 90: 85: 80: 75: 70: 65: 60: 48: 26: 9: 6: 4: 3: 2: 4789: 4778: 4775: 4773: 4770: 4768: 4765: 4763: 4760: 4759: 4757: 4740: 4735: 4731: 4727: 4722: 4718: 4714: 4709: 4705: 4701: 4696: 4692: 4688: 4682: 4678: 4673: 4669: 4665: 4660: 4656: 4652: 4647: 4643: 4639: 4634: 4630: 4626: 4621: 4617: 4613: 4608: 4604: 4600: 4595: 4591: 4587: 4582: 4578: 4574: 4569: 4565: 4561: 4556: 4552: 4548: 4543: 4539: 4535: 4530: 4526: 4522: 4517: 4513: 4508: 4502: 4498: 4493: 4487: 4483: 4482: 4480: 4478: 4474: 4470: 4465: 4456: 4454: 4447: 4442: 4439: 4438: 4432: 4428: 4425: 4424: 4418: 4414: 4413: 4404: 4401: 4398: 4394: 4390: 4385: 4380: 4376: 4372: 4368: 4364: 4360: 4357: 4356: 4351: 4347: 4342: 4338: 4333: 4328: 4324: 4320: 4316: 4312: 4308: 4304: 4300: 4296: 4292: 4287: 4285: 4284:0-226-77142-3 4281: 4277: 4273: 4270: 4266: 4262: 4258: 4255: 4254: 4249: 4244: 4241: 4238: 4237:Avian Biology 4234: 4230: 4227: 4223: 4219: 4215: 4211: 4207: 4203: 4198: 4196: 4192: 4189: 4185: 4184:80-901105-3-3 4181: 4176: 4170: 4166: 4161: 4158: 4154: 4150: 4146: 4142: 4138: 4134: 4129: 4124: 4119: 4115: 4111: 4107: 4103: 4099: 4094: 4090: 4086: 4082: 4078: 4074: 4071: 4070: 4064: 4061: 4057: 4053: 4049: 4045: 4040: 4035: 4031: 4027: 4023: 4018: 4014: 4010: 4006: 4002: 4001: 3994: 3991: 3987: 3983: 3979: 3975: 3971: 3967: 3963: 3959: 3955: 3951: 3947: 3942: 3940: 3936: 3933: 3929: 3926: 3922: 3918: 3914: 3910: 3903: 3901: 3894: 3891: 3887: 3884: 3880: 3876: 3872: 3868: 3867: 3861: 3860: 3849: 3844: 3835: 3830: 3826: 3822: 3818: 3812: 3796: 3792: 3788: 3782: 3774: 3770: 3766: 3762: 3758: 3754: 3747: 3739: 3734: 3730: 3726: 3722: 3717: 3710: 3704: 3697: 3691: 3684: 3680: 3676: 3672: 3668: 3663: 3656: 3652: 3648: 3642: 3635: 3629: 3611: 3607: 3600: 3596: 3590: 3581: 3565: 3561: 3557: 3553: 3549: 3543: 3541: 3532: 3528: 3524: 3520: 3513: 3498: 3494: 3493: 3488: 3482: 3467: 3463: 3459: 3452: 3437: 3433: 3432: 3431:San Diego Zoo 3427: 3421: 3406: 3402: 3398: 3392: 3385: 3380: 3371: 3362: 3360: 3350: 3348: 3339: 3337:9780226771427 3333: 3329: 3325: 3319: 3304: 3300: 3296: 3292: 3286: 3278: 3274: 3269: 3264: 3259: 3254: 3250: 3246: 3242: 3238: 3234: 3227: 3218: 3209: 3200: 3194:Kirchman 2012 3191: 3189: 3179: 3177: 3175: 3173: 3165: 3159: 3150: 3141: 3139: 3137: 3127: 3118: 3116: 3106: 3097: 3095: 3093: 3083: 3074: 3069: 3065: 3061: 3057: 3050: 3048: 3038: 3036: 3026: 3024: 3008: 3004: 3000: 2999:"Rail (bird)" 2994: 2986: 2984:9781784270933 2980: 2976: 2975: 2967: 2958: 2956: 2946: 2941: 2937: 2933: 2929: 2925: 2918: 2916: 2911: 2902: 2899: 2897: 2892: 2887: 2885: 2875: 2873: 2868: 2863: 2862: 2856: 2854: 2850: 2847: 2839: 2835: 2832: 2829: 2825: 2822: 2818: 2815: 2814: 2809: 2806: 2802: 2800: 2796: 2792: 2788: 2787: 2782: 2781: 2780: 2769: 2765: 2761: 2757: 2756: 2755:Pleistorallus 2751: 2748: 2747: 2742: 2739: 2736: 2732: 2729: 2726: 2723: 2720: 2717: 2714: 2710: 2707: 2703: 2700: 2696: 2693: 2690: 2687: 2683: 2681: 2677: 2673: 2670: 2669: 2664: 2662: 2658: 2654: 2651: 2650: 2649:Rhenanorallus 2645: 2642: 2638: 2635: 2632: 2628: 2625: 2621: 2618: 2614: 2611: 2607: 2604: 2600: 2597: 2593: 2591: 2587: 2583: 2580: 2576: 2573: 2569: 2566: 2562: 2561: 2560: 2558: 2557:North America 2554: 2550: 2546: 2543: 2539: 2533:Fossil record 2530: 2528: 2524: 2523:Sarothruridae 2520: 2519: 2510: 2506: 2502: 2498: 2495: 2491: 2487: 2483: 2480: 2476: 2475:Capellirallus 2472: 2469: 2465: 2461: 2457: 2456: 2451: 2448: 2444: 2443:Erythromachus 2441: 2438: 2434: 2431: 2428: 2424: 2421: 2418: 2414: 2411: 2408: 2404: 2401: 2400: 2395: 2384: 2380: 2376: 2372: 2368: 2364: 2362: 2358: 2354: 2350: 2346: 2338: 2337: 2333: 2331: 2327: 2324: 2322: 2321:striped crake 2318: 2315: 2313: 2309: 2306: 2304: 2300: 2297: 2295: 2294:Nkulengu rail 2291: 2288: 2286:– (3 species) 2285: 2284: 2280: 2278:– (4 species) 2277: 2276: 2272: 2269: 2268: 2264: 2261: 2260: 2256: 2254:– (3 species) 2253: 2252: 2248: 2246:– (2 species) 2245: 2244: 2240: 2238: 2234: 2231: 2228: 2227: 2223: 2220: 2219: 2215: 2212: 2211: 2207: 2205: 2201: 2200:Paragallinula 2198: 2195: 2194: 2190: 2188:– (3 species) 2187: 2186: 2182: 2180: 2176: 2173: 2170: 2169: 2165: 2162: 2161: 2157: 2155: 2154:chestnut rail 2151: 2148: 2146: 2142: 2141: 2137: 2135: 2131: 2128: 2126: 2122: 2119: 2117:– (4 species) 2116: 2115: 2111: 2109: 2105: 2102: 2100: 2096: 2093: 2090: 2089: 2085: 2083: 2082:Rouget's rail 2079: 2076: 2074: 2073:African crake 2070: 2067: 2064: 2063: 2059: 2056: 2055: 2051: 2049: 2048:uniform crake 2045: 2042: 2039: 2038: 2034: 2032:– (4 species) 2031: 2030: 2029:Mustelirallus 2026: 2024: 2020: 2017: 2016: 2015: 2013: 2009: 2002:Extant genera 1994: 1993: 1985: 1984: 1976: 1975: 1967: 1966: 1958: 1957: 1949: 1948: 1940: 1939: 1931: 1930: 1922: 1921: 1918: 1916: 1915: 1908: 1907: 1904: 1903: 1900: 1898: 1897: 1890: 1889: 1883: 1882: 1879: 1878: 1875: 1873: 1872: 1865: 1864: 1858: 1857: 1854: 1853: 1845: 1844: 1841: 1839: 1838: 1831: 1830: 1827: 1826: 1823: 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Retrieved 3790: 3781: 3759:(1): 71–78. 3756: 3752: 3746: 3728: 3724: 3716: 3703: 3695: 3690: 3682: 3662: 3654: 3641: 3633: 3628: 3617:. Retrieved 3605: 3589: 3580: 3568:. Retrieved 3559: 3518: 3512: 3501:. Retrieved 3491: 3481: 3470:. Retrieved 3461: 3451: 3440:. Retrieved 3429: 3420: 3409:. Retrieved 3400: 3391: 3379: 3370: 3327: 3318: 3307:. Retrieved 3298: 3293:(May 2018). 3285: 3240: 3236: 3226: 3217: 3208: 3199: 3163: 3158: 3153:Livezey 2003 3149: 3126: 3121:McNab (1994) 3105: 3082: 3063: 3059: 3011:. Retrieved 3002: 2993: 2973: 2966: 2935: 2931: 2852: 2848: 2843: 2837: 2827: 2820: 2811: 2804: 2798: 2794: 2790: 2784: 2778: 2763: 2753: 2744: 2734: 2712: 2705: 2698: 2685: 2679: 2675: 2666: 2660: 2656: 2647: 2640: 2630: 2623: 2617:Quercyrallus 2616: 2609: 2602: 2595: 2589: 2585: 2578: 2572:Palaeorallus 2571: 2564: 2536: 2516: 2514: 2500: 2485: 2474: 2467: 2460:Chatham rail 2453: 2442: 2433:Aphanapteryx 2432: 2422: 2412: 2402: 2382: 2352: 2342: 2334: 2325: 2316: 2307: 2298: 2289: 2281: 2273: 2265: 2257: 2249: 2241: 2232: 2224: 2216: 2208: 2199: 2191: 2183: 2174: 2168:Hypotaenidia 2166: 2158: 2149: 2138: 2129: 2125:Calayan rail 2121:Aptenorallus 2120: 2112: 2108:snoring rail 2103: 2094: 2086: 2077: 2068: 2060: 2052: 2044:Amaurolimnas 2043: 2035: 2027: 2018: 2005: 1912: 1911: 1899:– watercock 1894: 1893: 1869: 1868: 1835: 1834: 1817: 1816: 1775: 1774: 1757: 1756: 1706: 1705: 1688: 1687: 1654: 1653: 1636: 1635: 1618:Laterallini 1596: 1595: 1525: 1524: 1507: 1506: 1482: 1481: 1457: 1456: 1432: 1431: 1406: 1405: 1302:Hypotaenidia 1300: 1299: 1282: 1281: 1257: 1256: 1232: 1231: 1209:Aptenorallus 1207: 1206: 1164: 1163: 1146: 1145: 1121: 1120: 1080: 1079: 1062: 1061: 1030: 1029: 986: 985: 970:Amaurolimnas 968: 967: 934: 933: 916: 915: 857: 811: 771: 750:World War II 739: 694: 691:Reproduction 685:conservation 677: 650: 602: 575: 555: 542: 537: 531: 525: 519: 517: 507: 494: 485: 469: 461: 442: 432: 429:little crake 424: 410: 390:the Americas 386:clapper rail 379: 375: 365: 345: 338: 335:Vulgar Latin 330: 323: 321: 285:rice paddies 254:cosmopolitan 249: 245: 244: 233: 214: 140: 54:Early Eocene 40: 4620:iNaturalist 4501:Wikispecies 4361:(1): 1–39. 4345:(Full text) 4259:(1): 5–14. 4202:Experientia 4163: [ 3873:: 157–204. 3832: [ 3570:14 November 3548:Gill, Frank 2932:Ornithology 2924:Olson, S.L. 2853:Montirallus 2828:Rupelrallus 2661:Microrallus 2624:Belgirallus 2579:Parvirallus 2545:prehistoric 2518:Nesotrochis 2468:Gallirallus 2308:Poliolimnas 2290:Himantornis 2251:Coturnicops 2175:Porphyriops 2150:Eulabeornis 2140:Gallirallus 2104:Aramidopsis 2040:(3 species) 2037:Pardirallus 1837:Poliolimnas 1690:Coturnicops 1408:Himantornis 1284:Eulabeornis 1259:Gallirallus 936:Pardirallus 699:, although 657:crepuscular 571:territorial 562:herbivorous 489:Laysan rail 406:horned coot 289:terrestrial 36:River Crake 4756:Categories 4384:2440/43360 3952:: 96–108. 3937:: 33–78. 3738:10088/8374 3619:2018-07-29 3503:2018-02-21 3472:2018-07-29 3442:2018-07-29 3411:2018-07-29 3401:AP Archive 3374:BLI (2007) 3309:2018-07-29 3182:McNab 2002 3013:2021-06-03 2907:References 2838:Euryonotus 2821:Palaeocrex 2735:Creccoides 2713:Youngornis 2686:Litorallus 2676:Pararallus 2668:Australlus 2549:formations 2486:Vitirallus 2479:Snipe-rail 2413:Aphanocrex 2349:Ibiza rail 2336:Amaurornis 2259:Laterallus 2243:Rufirallus 2233:Micropygia 2130:Habroptila 2099:corn crake 2088:Dryolimnas 2019:Canirallus 1914:Amaurornis 1746:Zapornini 1708:Laterallus 1656:Rufirallus 1638:Micropygia 1234:Habroptila 1123:Dryolimnas 865:Frank Gill 849:convergent 829:Gruiformes 786:Micronesia 697:monogamous 681:endangered 614:flightless 578:vegetation 427:) and the 415:in either 352:Morphology 328:Old French 312:Antarctica 220:Rafinesque 205:Gruiformes 4208:: 94–98. 3900:Gallinula 3819:. A left 3817:QM F40203 3815:Specimen 3698:(1998–99) 3681:: Worthy 3653:: Worthy 3649:or small 3066:(2): 70. 3060:Diversity 2786:Ludiortyx 2706:Miorallus 2699:Miofulica 2631:Rallicrex 2610:Ibidopsis 2590:Protogrus 2586:Aletornis 2527:adzebills 2379:Mauritius 2330:watercock 2326:Gallicrex 2283:Gymnocrex 2226:Porphyrio 2210:Gallinula 2078:Rougetius 2069:Crecopsis 1896:Gallicrex 1598:Porphyrio 1509:Gallinula 1421:Fulicini 1082:Rougetius 1064:Crecopsis 890:Rallinae 860:cladogram 837:monophyly 794:Guam rail 782:Polynesia 778:Melanesia 730:Guam rail 705:polyandry 661:Old World 638:mongooses 622:predators 594:Reed beds 545:blue tits 480:metabolic 459:periods. 421:watercock 308:snow line 277:waterfowl 269:gallinule 171:Kingdom: 165:Eukaryota 4762:Rallidae 4521:Rallidae 4507:Rallidae 4486:Wikidata 4477:Rallidae 4437:Rallidae 4423:Rallidae 4393:85230857 4274:(2006): 4222:23665106 4191:Archived 4173:(2002): 4153:16632395 4089:86511951 4069:Am. Nat. 4056:85940913 3974:25255711 3795:Archived 3667:Quadrate 3610:Archived 3597:(1977). 3564:Archived 3531:2246/830 3497:Archived 3489:(1815). 3466:Archived 3436:Archived 3405:Archived 3384:Steadman 3326:(2006). 3303:Archived 3277:23530197 3007:Archived 2859:See also 2813:Amitabha 2805:Telecrex 2760:holotype 2603:Latipons 2501:Hovacrex 2494:Holocene 2437:Red rail 2375:red rail 2299:Megacrex 2267:Zapornia 2193:Tribonyx 2054:Aramides 1819:Megacrex 1759:Zapornia 1459:Tribonyx 1019:Rallini 988:Aramides 845:tropical 825:bustards 814:polymath 756:and the 713:Clutches 701:polygyny 624:such as 483:niches. 453:vagrants 250:Rallidae 215:Rallidae 211:Family: 185:Chordata 181:Phylum: 175:Animalia 161:Domain: 56:– Recent 18:Rallidae 4458:. 1914. 4363:Bibcode 4341:9163823 4332:1691940 4303:Bibcode 4110:Bibcode 4026:The Auk 4000:Geobios 3954:Bibcode 3866:Geobios 3801:18 July 3761:Bibcode 3268:3631643 3245:Bibcode 3162:Slikas 2542:extinct 2455:Cabalus 2275:Rallina 2185:Porzana 2160:Cabalus 2114:Lewinia 1777:Rallina 1434:Porzana 1262:– weka 1166:Lewinia 918:Neocrex 653:reedbed 634:weasels 618:extinct 598:migrate 449:migrate 417:plumage 370:remiges 341:, from 339:rascula 326:, from 292:habitat 273:wetland 228:Genera 201:Order: 191:Class: 4739:159003 4700:414340 4684:NZOR: 4651:176205 4638:105005 4573:1RALLF 4492:Q26623 4455:  4391:  4339:  4329:  4321:  4282:  4220:  4182:  4151:  4087:  4054:  4046:  3972:  3838:(2005) 3696:et al. 3685:(2007) 3683:et al. 3675:femora 3657:(2007) 3655:et al. 3636:(2003) 3634:et al. 3386:(2006) 3334:  3275:  3265:  3166:(2002) 3164:et al. 2981:  2836:Genus 2826:Genus 2819:Genus 2810:Genus 2803:Genus 2783:Genus 2752:Genus 2733:Genus 2711:Genus 2704:Genus 2697:Genus 2684:Genus 2674:Genus 2665:Genus 2655:Genus 2646:Genus 2639:Genus 2629:Genus 2622:Genus 2615:Genus 2608:Genus 2601:Genus 2594:Genus 2584:Genus 2577:Genus 2570:Genus 2565:Eocrex 2563:Genus 2553:Europe 2538:Fossil 2499:Genus 2484:Genus 2473:Genus 2452:Genus 2403:Mundia 2357:fossil 2345:fossil 2218:Fulica 2062:Rallus 1527:Fulica 1032:Rallus 853:grebes 821:cranes 784:, and 742:hunted 683:, and 673:takahē 644:, and 610:fliers 590:rivers 586:swamps 564:. The 530:, and 472:Flight 445:flight 362:takahē 346:rādere 304:alpine 296:desert 267:, and 257:family 222:, 1815 4772:Rails 4734:WoRMS 4721:Plazi 4713:39513 4633:IRMNG 4586:10767 4389:S2CID 4323:56680 4319:JSTOR 4218:S2CID 4171:] 4085:S2CID 4052:S2CID 4044:JSTOR 3905:(PDF) 3848:Olson 3836:] 3829:Boles 3613:(PDF) 3602:(PDF) 2846:wader 851:with 762:pests 651:Many 630:foxes 603:Most 588:, or 582:lakes 566:calls 457:moult 394:coots 343:Latin 331:rasle 300:polar 281:marsh 265:coots 261:birds 246:Rails 236:below 46:Rails 4677:9119 4672:NCBI 4646:ITIS 4612:9342 4607:GBIF 4568:EPPO 4560:7581 4534:1450 4529:BOLD 4337:PMID 4280:ISBN 4180:ISBN 4149:PMID 3970:PMID 3803:2010 3572:2022 3332:ISBN 3273:PMID 3237:PNAS 2979:ISBN 2555:and 2525:and 2462:and 2359:and 2145:weka 2095:Crex 1148:Crex 858:The 823:and 802:Guam 728:The 709:eggs 703:and 646:pigs 642:rats 626:cats 605:nest 476:keel 382:bill 380:The 324:rĂąle 318:Name 195:Aves 62:PreꞒ 4659:NBN 4625:154 4555:EoL 4547:FJ9 4542:CoL 4516:AFD 4379:hdl 4371:doi 4327:PMC 4311:doi 4299:352 4261:doi 4233:In: 4210:doi 4141:doi 4137:145 4118:doi 4077:doi 4073:144 4034:doi 4030:129 4009:doi 3986:In: 3962:doi 3917:doi 3875:doi 3769:doi 3733:hdl 3671:MNZ 3527:hdl 3263:PMC 3253:doi 3241:110 3068:doi 2940:doi 2936:138 2762:of 2551:of 2377:of 435:). 388:of 4758:: 4736:: 4723:: 4710:: 4697:: 4674:: 4661:: 4648:: 4635:: 4622:: 4609:: 4596:: 4583:: 4570:: 4557:: 4544:: 4531:: 4518:: 4503:: 4488:: 4449:. 4387:. 4377:. 4369:. 4335:. 4325:. 4317:. 4309:. 4297:. 4293:. 4257:33 4216:. 4206:49 4204:. 4169:fr 4167:; 4165:de 4147:. 4135:. 4116:. 4104:. 4100:. 4083:. 4050:. 4042:. 4028:. 4024:. 4005:36 4003:. 3968:. 3960:. 3950:81 3948:. 3935:23 3913:57 3911:. 3907:. 3869:. 3834:de 3789:. 3767:. 3757:21 3755:. 3729:87 3727:. 3608:. 3604:. 3558:. 3539:^ 3464:. 3460:. 3434:. 3428:. 3399:. 3358:^ 3346:^ 3301:. 3297:. 3271:. 3261:. 3251:. 3239:. 3235:. 3187:^ 3171:^ 3135:^ 3114:^ 3091:^ 3064:12 3062:. 3058:. 3046:^ 3034:^ 3022:^ 3005:. 3001:. 2954:^ 2934:. 2930:. 2914:^ 2855:. 2797:, 2793:, 2529:. 2503:– 2488:– 2477:– 2458:– 2445:– 2435:– 2425:– 2415:– 2405:– 2381:, 2328:– 2319:– 2310:– 2301:– 2292:– 2235:– 2202:– 2177:– 2152:– 2143:– 2132:– 2123:– 2106:– 2097:– 2080:– 2071:– 2046:– 2021:– 2014:. 867:, 780:, 764:. 711:. 648:. 640:, 636:, 632:, 628:, 592:. 584:, 573:. 524:, 408:. 298:, 238:. 139:, 112:Pg 4395:. 4381:: 4373:: 4365:: 4359:5 4343:. 4313:: 4305:: 4267:. 4263:: 4240:8 4224:. 4212:: 4155:. 4143:: 4126:. 4120:: 4112:: 4106:5 4091:. 4079:: 4058:. 4036:: 4015:. 4011:: 3976:. 3964:: 3956:: 3923:. 3919:: 3881:. 3877:: 3871:2 3805:. 3775:. 3771:: 3763:: 3741:. 3735:: 3669:( 3622:. 3574:. 3533:. 3529:: 3506:. 3475:. 3445:. 3414:. 3340:. 3312:. 3279:. 3255:: 3247:: 3076:. 3070:: 3016:. 2948:. 2942:: 2770:. 2511:) 2496:) 2351:( 736:. 514:. 506:( 431:( 423:( 364:( 337:* 117:N 107:K 102:J 97:T 92:P 87:C 82:D 77:S 72:O 67:Ꞓ 38:. 20:)

Index

Rallidae
Crake (surname)
River Crake
Early Eocene
PreꞒ
Ꞓ
O
S
D
C
P
T
J
K
Pg
N

Dusky moorhen
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Aves
Gruiformes
Rallidae
Rafinesque
below
cosmopolitan
family

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