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Wilhelm Johannsen

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536:(The way that the phenotype manifests itself, whether it shows itself by traits that examined qualitatively or quantitatively, says absolutely nothing beforehand about the genes. Very conspicuous phenotypic differences could show themselves where no genotypic difference is present, and there are also cases where in the case of genotypic variety, the phenotypes are alike. Thus it is precisely of the greatest importance to separate clearly the concept of phenotype (a type of appearance) from the concept of genotype (one might say a type of plan). We will admittedly not be able to operate with this latter concept – a genotype just does not appear in pure form ; the derived concept of a genotypic difference will, however, often be of use to us.) 33: 614:(from γί-γ (ε)ν-ομαι, to become). Only the meaning of this latter comes into consideration here ; just the basic idea – that a trait in the developing organism can be determined or is influenced by "something" in the gametes – should find expression. No hypothesis about the nature of this "something" should be postulated or supported by it. For that reason it seems simplest to use in isolation the last syllable 622:. Thus we will say simply "gene" and "genes" for "pangene" and "pangenes". The word gene is completely free of any hypothesis ; it expresses only the established fact that in any case many traits of the organism are determined by specific, separable, and thus independent "conditions", "foundations", "plans" – in short, precisely what we want to call genes.) 505:
Johannsen distinguished between an organism's outward appearance (which he designated as its "phenotype") and its inherent genetic heritage (which he designated as its "genotype"). He stressed that an organism's appearance need not correspond exactly to its genetic heritage. So on p. 123 he defines
508:"Darum könnte man den statistisch hervortretenden Typus passend als Erscheinungstypus bezeichnen oder, kurz und klar, als "Phaenotypus". 1) … Ein gebener Phaenotypus mag Ausdruck einer biologischen Einheit sein; er braucht es aber durchaus nicht zu sein. 1) Von φαίν-ομαι, scheinen." 510:(Therefore one could designate the statistically prominent type appropriately as a type of appearance or, clearly and concisely, as a "phenotype". 1) … A given phenotype may be an expression of a biological unit; but it definitely need not be so. 1) From φαίν-ομαι, to appear.) 732: 602:. Many other terms have been suggested, mostly unfortunately in closer connection with certain hypothetical opinions. The word "pangene", which was introduced by Darwin, is perhaps used most frequently in place of 760:
Anker, Jean (1932) Wilhelm Johannsen, pp. 177–180 in: Meisen, V. Prominent Danish Scientists through the Ages. University Library of Copenhagen 450th Anniversary. Levin & Munksgaard, Copenhagen.
880: 448:. 4th edn by Warming and Johannsen 1900-01). German edn 1907-09: Lehrbuch der allgemeinen Botanik (from the 4th edn, by E. P. Meinecke). Berlin, Borntraeger. 667 pp. 763:
Roll-Hansen, Nils (1983) The Death of Spontaneous Generation and the Birth of the Gene: Two Case Studies of Relativism. Social Studies of Science 13 (4): 481–519.
845: 598:(This "something" in the gametes or in the zygote, which has crucial importance for the character of the organism, is usually called by the quite ambiguous term 885: 773:
Kim, Kyung-Man (1991) On the Reception of Johannsen's Pure Line Theory: Toward a Sociology of Scientific Validity. Social Studies of Science 21 (4): 649–679.
557: 870: 534:"Die Art, wie die Phaenotypen sich manifestieren, … der abgeleitete Begriff genotypischer Unteschied wird uns aber vielfach von Nutzen sein." 265: 213:
and Germany beginning in 1872 until passing his pharmacist's exam in 1879. In 1881, he became assistant in the chemistry department at the
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Roll-Hansen, Nils (1979). "The Genotype Theory of Wilhelm Johannsen and its Relation to Plant Breeding and the Study of Evolution".
444:& W. Johannsen (1895) Den almindelige Botanik (General Botany): En Lærebog, nærmest til Brug for Studerende og Lærere. 3rd edn, 840: 546: 875: 825: 800: 186: 855: 618:
from Darwin's well-known word, which alone is of interest to us, in order to replace, with it, the poor, ambiguous word
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Johannsen coins the term "genotype" on p. 130, where he stresses the distinction between phenotype and genotype:
835: 403: 335: 830: 422: 596:"Dieses "etwas" in den Gameten bezw. in der Zygote, … – kurz, was wir eben Gene nennen wollen – bedingt sind." 554: 860: 402:, becoming vice-chancellor in 1917. In December 1910, Johannsen was invited to give an address before the 865: 790: 583: 521: 494: 399: 142: 87: 606:. However, the word "pangene" was not well chosen, as it is a compound word containing the roots 295:. This was attributable to resource provision to the mother plant and to the position of seeds in 820: 815: 467: 214: 572:
Johannsen, W.L. (1905) Arvelighedslærens elementer (The Elements of Heredity). Copenhagen.
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as proof of gradual genetic variation on which selection could act. Only with the
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Johannsen, W. (1903) Om arvelighed i samfund og i rene linier. Oversigt over det
365:(Elements of the exact theory of heredity). This book was based in large part on 702: 384: 319: 218: 32: 809: 254: 718: 675: 323: 276:. He taught plant physiology. His best-known research concerned so-called 281: 230: 801:
6 min silent movie showing Johannsen in action as teacher and scientist
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for all traits, i.e. without genetic variation, seed size followed a
178: 111: 649: 357: 307: 246: 226: 190: 182: 162: 128: 107: 53: 210: 158: 57: 410:. In 1911, he was invited to give a series of four lectures at 296: 269: 98: 592:] (in German). Jena, Germany: Gustav Fischer. p. 124. 530:] (in German). Jena, Germany: Gustav Fischer. p. 130. 503:] (in German). Jena, Germany: Gustav Fischer. p. 123. 369:("On heredity in society and in pure lines") and in his book 238: 375: 234: 174: 103: 391:. The book became one of the founding texts of genetics. 373:. It was in this book Johannsen also introduced the term 299:
and of pods on the plant. This led him to coin the terms
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Anon (1911). "Professor Johannsen's Columbia Lectures".
379:. This term was coined in opposition to the then common 791:
Wilhelm Johannsen Centre for Functional Genome Research
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Johannsen's findings led him to oppose contemporary
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Members of the Royal Society of Sciences in Uppsala
394:Also in 1905, Johannsen was appointed professor of 338:, was it established that variation needed to be 807: 551:Erblichkeit in Populationen und in reinen Linien 846:Academic staff of the University of Copenhagen 886:Members of the American Philosophical Society 205:. While very young, he was apprenticed to a 417:Johannsen was a corresponding member of the 361:, first using them in his book in German as 266:Royal Veterinary and Agricultural University 549:Forhandlinger, vol. 3: 247-270. German ed. 457: 419:Academy of Natural Sciences of Philadelphia 157:(3 February 1857 – 11 November 1927) was a 31: 665: 631: 581: 519: 492: 173:. He is best known for coining the terms 610:(the neuter form of Πας all, every) and 590:Elements of the exact theory of heredity 547:Kongelige Danske Videnskabernes Selskabs 528:Elements of the exact theory of heredity 501:Elements of the exact theory of heredity 367:Om arvelighed i samfund og i rene linier 871:Rectors of the University of Copenhagen 808: 585:Elemente der exakten Erblichkeitslehre 523:Elemente der exakten Erblichkeitslehre 496:Elemente der exakten Erblichkeitslehre 421:(elected 1915). He was elected to the 363:Elemente der exakten Erblichkeitslehre 264:In 1892, he was appointed lecturer at 634:"The Genotype Conception of Heredity" 688: 284:. He was able to show that even in 13: 480:10.1111/j.1600-0498.1979.tb00589.x 14: 897: 784: 841:University of Copenhagen alumni 725: 406:. This talk was printed in the 404:American Society of Naturalists 342:to act as the raw material for 682: 625: 575: 566: 539: 513: 486: 451: 435: 423:American Philosophical Society 268:and later became professor of 16:Danish botanist and geneticist 1: 876:19th-century Danish botanists 826:20th-century Danish botanists 553:(1903) Gustav Fischer, Jena. 428: 326:, who held the occurrence of 97:proving the constancy of the 196: 7: 371:Arvelighedslærens Elementer 249:could be broken by various 10: 902: 856:Carlsberg Laboratory staff 754: 703:10.1126/science.34.876.484 148: 138: 124: 117: 93: 83: 64: 39: 30: 23: 400:University of Copenhagen 221:. Johannsen studied the 143:University of Copenhagen 88:University of Copenhagen 25:Wilhelm Ludvig Johannsen 638:The American Naturalist 187:"pure line" experiments 836:Danish science writers 632:Johannsen, W. (1911). 582:Johannsen, W. (1909). 520:Johannsen, W. (1909). 493:Johannsen, W. (1909). 201:Johannsen was born in 851:People from Helsingør 831:Agriculture educators 349:He created the terms 737:search.amphilsoc.org 733:"APS Member History" 280:of the self-fertile 215:Carlsberg Laboratory 861:Plant physiologists 560:30 May 2009 at the 472:1979Cent...22..201R 412:Columbia University 408:American Naturalist 330:trait variation in 293:normal distribution 253:compounds, such as 185:, and for his 1903 866:Danish geneticists 555:Scanned full text. 383:that stemmed from 328:normal distributed 217:under the chemist 167:plant physiologist 245:. He showed that 155:Wilhelm Johannsen 152: 151: 119:Scientific career 893: 748: 747: 745: 743: 729: 723: 722: 686: 680: 679: 669: 629: 623: 593: 579: 573: 570: 564: 543: 537: 531: 517: 511: 504: 490: 484: 483: 455: 449: 439: 396:plant physiology 336:modern synthesis 274:plant physiology 133:Plant physiology 71: 68:11 November 1927 49: 47: 35: 21: 20: 901: 900: 896: 895: 894: 892: 891: 890: 806: 805: 787: 757: 752: 751: 741: 739: 731: 730: 726: 687: 683: 644:(531): 129–59. 630: 626: 580: 576: 571: 567: 562:Wayback Machine 544: 540: 518: 514: 491: 487: 456: 452: 440: 436: 431: 318:, most notably 199: 131: 101: 84:Alma mater 79: 73: 69: 60: 51: 50:3 February 1857 45: 43: 26: 17: 12: 11: 5: 899: 889: 888: 883: 878: 873: 868: 863: 858: 853: 848: 843: 838: 833: 828: 823: 818: 804: 803: 798: 793: 786: 785:External links 783: 782: 781: 771: 761: 756: 753: 750: 749: 724: 681: 650:10.1086/279202 624: 594:From p. 124: 574: 565: 538: 512: 506:"phenotype": 485: 466:(3): 201–235. 450: 433: 432: 430: 427: 320:Francis Galton 219:Johan Kjeldahl 209:and worked in 198: 195: 150: 149: 146: 145: 140: 136: 135: 126: 122: 121: 115: 114: 95: 94:Known for 91: 90: 85: 81: 80: 74: 72:(aged 70) 66: 62: 61: 52: 41: 37: 36: 28: 27: 24: 15: 9: 6: 4: 3: 2: 898: 887: 884: 882: 879: 877: 874: 872: 869: 867: 864: 862: 859: 857: 854: 852: 849: 847: 844: 842: 839: 837: 834: 832: 829: 827: 824: 822: 819: 817: 814: 813: 811: 802: 799: 797: 796:Image gallery 794: 792: 789: 788: 780: 776: 772: 770: 766: 762: 759: 758: 738: 734: 728: 720: 716: 712: 708: 704: 700: 696: 692: 685: 677: 673: 668: 663: 659: 655: 651: 647: 643: 639: 635: 628: 621: 617: 613: 609: 605: 601: 597: 591: 587: 586: 578: 569: 563: 559: 556: 552: 548: 542: 535: 529: 525: 524: 516: 509: 502: 498: 497: 489: 481: 477: 473: 469: 465: 461: 454: 447: 443: 442:Warming, Eug. 438: 434: 426: 424: 420: 415: 413: 409: 405: 401: 397: 392: 390: 386: 382: 378: 377: 372: 368: 364: 360: 359: 354: 353: 347: 345: 341: 337: 333: 329: 325: 321: 317: 312: 310: 309: 304: 303: 298: 294: 290: 287: 283: 279: 275: 271: 267: 262: 260: 256: 255:diethyl ether 252: 248: 244: 240: 236: 232: 228: 224: 220: 216: 212: 208: 204: 194: 192: 188: 184: 180: 176: 172: 168: 164: 160: 156: 147: 144: 141: 137: 134: 130: 127: 123: 120: 116: 113: 109: 105: 100: 96: 92: 89: 86: 82: 77: 67: 63: 59: 55: 42: 38: 34: 29: 22: 19: 740:. 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Index


Elsinore
Denmark
Copenhagen
University of Copenhagen
genome
gene
genotype
phenotype
Genetics
Plant physiology
University of Copenhagen
Danish
botanist
plant physiologist
geneticist
gene
phenotype
genotype
"pure line" experiments
genetics
Copenhagen
pharmacist
Denmark
Carlsberg Laboratory
Johan Kjeldahl
metabolism
dormancy
germination
seeds

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