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Baldwin effect

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referred to behavioural adaptations to prevailing climates in different species of bees as "what had merely been an idea, therefore, and opposed to instinct, has thus by slow degrees become an instinctive habit". The Baldwin effect theory subsequently became more controversial, with scholars divided
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Selected offspring would tend to have an increased capacity for learning new skills rather than being confined to genetically coded, relatively fixed abilities. In effect, it places emphasis on the fact that the sustained behaviour of a species or group can shape the evolution of that species. The
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If animals entered a new environment—or their old environment rapidly changed—those that could flexibly respond by learning new behaviours or by ontogenetically adapting would be naturally preserved. This saved remnant would, over several generations, have the opportunity to exhibit spontaneously
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According to Dennett, also in 2003, recent work has rendered the Baldwin effect "no longer a controversial wrinkle in orthodox Darwinism". Potential genetic mechanisms underlying the Baldwin effect have been proposed for the evolution of natural (genetically determinant) antibodies. In 2009,
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Thanks to the Baldwin effect, species can be said to pretest the efficacy of particular different designs by phenotypic (individual) exploration of the space of nearby possibilities. If a particularly winning setting is thereby discovered, this discovery will
317:. The Baldwin effect includes genetic accommodation, of which one type is genetic assimilation. Science historian Laurent Loison has written that "the Baldwin effect and genetic assimilation, even if they are quite close, should not be conflated". 270:
who are searching for scenarios in which a population can get itself by behavioral trial and error onto a "hard to find" part of the fitness landscape in which human brain, language, and mind can rapidly coevolve. They are searching for what
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congenital variations similar to their acquired traits and have these variations naturally selected. It would look as though the acquired traits had sunk into the hereditary substance in a Lamarckian fashion, but the process would really be
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Still, David Depew observed in 2003, "it is striking that a rather diverse lot of contemporary evolutionary theorists, most of whom regard themselves as supporters of the Modern Synthesis, have of late become 'Baldwin boosters'". These
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Initially Baldwin's ideas were not incompatible with the prevailing, but uncertain, ideas about the mechanism of transmission of hereditary information and at least two other biologists put forward very similar ideas in 1896. In 1901,
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in the 1960s–1970s. This new version included an explicit role for the social in shaping subsequent natural change in humans (both evolutionary and developmental), with reference to alterations of selection pressures.
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their parents' acquired characteristics. The Baldwin effect only posits that learning ability, which is genetically based, is another variable in / contributor to environmental adaptation. First proposed during the
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and evolution in the effect. Baldwin's account of how neurophysiological and conscious mental factors may contribute to the effect brings into focus the question of the possible survival value of consciousness.
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approach and that "Mendelism and later genetic theory so conclusively ruled out the extreme neo-Lamarckian position that reconciliation came to seem unnecessary". In 1942, the evolutionary biologist
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Lindahl, B. I. B. (2001). "Consciousness, behavioural patterns and the direction of biological evolution: implications for the mind–brain problem". In Pylkkänen, Paavo; Vadén, Tere (eds.).
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Griffiths, Paul E. (2003). "Beyond the Baldwin effect: James Mark Baldwin's 'social heredity', epigenetic inheritance, and niche construction". In Weber, Bruce H.; Depew, David J. (eds.).
1235: 76:. It posits that subsequent selection might reinforce the originally learned behaviors, if adaptive, into more in-born, instinctive ones. Though this process appears similar to 220:
the argument is stated in terms of the individual genotype, whereas what is really exposed to the selection pressure is a phenotypically and genetically variable population;
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new behaviours (e.g. to acclimatise to a new stressor) will affect its reproductive success and will therefore have an effect on the genetic makeup of its species through
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2017, 6(4), 47; doi:10.3390/biology6040047, quoting Bateson, P. The adaptability driver: Links between behaviour and evolution. Biol. Theory 2006, 1, 342–345. See also
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a new selection pressure: organisms that are closer in the adaptive landscape to that discovery will have a clear advantage over those more distant.
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and Steven Nowlan demonstrated by computer simulation that learning can accelerate evolution, and they associated this with the Baldwin effect.
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in the late 19th century, this effect has been independently proposed several times, and today it is generally recognized as part of the
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promoted the Baldwin effect as part of the modern synthesis, saying the concept had been unduly neglected by evolutionists.
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Hall, Brian K. (2001). "Organic Selection: Proximate Environmental Effects on the Evolution of Morphology and Behaviour".
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Puentedura, Ruben R. (2003). "The Baldwin effect in the age of computation". In Weber, Bruce H.; Depew, David J. (eds.).
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The Baldwin effect has been confused with, and sometimes conflated with, a different evolutionary theory also based on
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it is not sufficiently emphasized that the degree of modification of the phenotype is in itself genetically controlled;
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Paul Griffiths suggests two reasons for the continuing interest in the Baldwin effect. The first is the role
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in 1953. Simpson "admitted that the idea was theoretically consistent, that is, not inconsistent with the
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The effect, then unnamed, was put forward in 1896 in a paper "A New Factor in Evolution" by the American
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Anderson, Russell (1996). "How the adaptive antibodies facilitate the evolution of natural antibodies".
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Burman, J. T. (2013). "Updating the Baldwin Effect: The biological levels behind Piaget's new theory".
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between "Baldwin boosters" and "Baldwin skeptics". The theory was first called the "Baldwin effect" by
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empirical evidence for the Baldwin effect was provided from the colonisation of North America by the
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into its developmental genetic or epigenetic repertoire. In the words of the philosopher of science
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it is assumed that phenotypic rigidity is selectively superior to phenotypic flexibility.
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Subsequent research shows that Baldwin was not the first to identify the process;
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Scheiner, Samuel M. (2014). "The Baldwin Effect: Neglected and Misunderstood".
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is understood to play in the effect. The second is the connection between
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Bateson, Patrick (2004). "The Active Role of Behaviour in Evolution".
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Philosophical Transactions of the Royal Society B: Biological Sciences
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Darwin and the Emergence of Evolutionary Theories of Mind and Behavior
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Adaptive Individuals In Evolving Populations: Models And Algorithms
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Depew, David J. (2003), "Baldwin Boosters, Baldwin Skeptics" in:
817:; Nowlan, Steven J. (1987). "How learning can guide evolution". 341: â€“ Thought or idea that can be shared, in analogy to a gene 216:
contended that the Baldwin effect theory was untenable because
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Philosophical Transactions of the Royal Society of London B
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Sznajder, B.; Sabelis, M. W.; Egas, M. (September 2012).
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Evolution and Learning: The Baldwin Effect Reconsidered
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Evolution and Learning: The Baldwin Effect Reconsidered
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Evolution and Learning: The Baldwin Effect Reconsidered
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Evolution and Learning: The Baldwin Effect Reconsidered
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describes an effect of learned behaviour on evolution.
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as a result of its interaction with its environment
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Archived from 1504: 1494: 1465: 1440:(121): 634–636. 1426:Baldwin, J. Mark 1421: 1404:(354): 441–451. 1390:Baldwin, J. Mark 1385: 1348: 1347: 1345: 1335: 1311: 1305: 1304: 1286: 1262: 1256: 1255: 1246:(5): 1199–1209. 1231: 1225: 1224: 1214: 1190: 1184: 1171: 1165: 1164: 1128: 1122: 1121: 1111: 1079: 1070: 1069: 1057: 1047: 1041: 1040: 1029:10.1037/h0063996 1009:Baldwin, J. Mark 1005: 999: 998: 959:Baldwin, J. Mark 955: 949: 948: 936: 926: 920: 919: 903: 893: 887: 886: 868: 866:10.1038/329761a0 833: 827: 826: 811: 805: 804: 792: 779: 773: 772: 752: 746: 745: 701: 695: 694: 673: 658: 657: 646: 640: 639: 614:(100): 786–789. 596: 590: 589: 546: 540: 525: 519: 518: 498: 492: 491: 467: 456: 450: 444: 438: 427: 426: 410: 397: 388: 382: 376: 370: 351:Weismann barrier 311:C. H. Waddington 199:modern synthesis 177:Douglas Spalding 91:modern synthesis 70:ability to learn 4384: 4383: 4379: 4378: 4377: 4375: 4374: 4373: 4349: 4348: 4347: 4342: 4284: 4270:Neoevolutionism 4177: 4161:Species complex 4126:Group selection 4064:Research topics 4059: 4035:Neuropsychology 3937: 3923:Substance abuse 3845:Sex differences 3839: 3753:Coolidge effect 3714: 3626:Neuroergonomics 3591: 3582: 3506: 3408: 3342:Folk psychology 3223: 3207: 3077: 3070: 3013: 3008: 2978: 2973: 2945: 2872:Group selection 2845: 2770: 2674: 2601: 2563:Tempo and modes 2557: 2412: 2316: 2133: 2092: 1968: 1961: 1938:Species complex 1751: 1742:History of life 1718: 1713: 1671: 1624: 1610:10.2307/2405746 1548: 1541: 1502: 1468: 1359: 1356: 1351: 1312: 1308: 1263: 1259: 1232: 1228: 1191: 1187: 1172: 1168: 1129: 1125: 1080: 1073: 1066: 1048: 1044: 1006: 1002: 956: 952: 945: 927: 923: 916: 894: 890: 834: 830: 819:Complex Systems 812: 808: 801: 780: 776: 769: 753: 749: 702: 698: 691: 674: 661: 647: 643: 597: 593: 564:(99): 733–740. 547: 543: 526: 522: 499: 495: 488: 470:Dennett, Daniel 468: 459: 451: 447: 439: 430: 423: 398: 391: 383: 379: 371: 364: 360: 333:Maternal effect 323: 303: 297: 232:Geoffrey Hinton 185: 110:Robert Richards 99: 28: 21:Baldwin's rules 17: 12: 11: 5: 4382: 4372: 4371: 4366: 4361: 4344: 4343: 4341: 4340: 4327: 4314: 4301: 4293: 4290: 4289: 4286: 4285: 4283: 4282: 4277: 4272: 4267: 4262: 4257: 4252: 4247: 4246: 4245: 4240: 4235: 4230: 4225: 4220: 4215: 4210: 4205: 4191: 4185: 4183: 4179: 4178: 4176: 4175: 4174: 4173: 4168: 4163: 4158: 4153: 4148: 4143: 4138: 4133: 4128: 4123: 4118: 4113: 4108: 4098: 4093: 4088: 4083: 4078: 4073: 4067: 4065: 4061: 4060: 4058: 4057: 4052: 4047: 4042: 4037: 4032: 4027: 4022: 4017: 4012: 4007: 4002: 3997: 3992: 3975: 3966: 3956: 3954: 3947: 3943: 3942: 3939: 3938: 3936: 3935: 3930: 3925: 3920: 3915: 3910: 3905: 3900: 3895: 3890: 3885: 3880: 3875: 3870: 3865: 3860: 3855: 3849: 3847: 3841: 3840: 3838: 3837: 3832: 3827: 3814: 3805: 3800: 3795: 3790: 3785: 3780: 3775: 3770: 3765: 3760: 3755: 3750: 3745: 3740: 3735: 3730: 3724: 3722: 3716: 3715: 3713: 3712: 3711: 3710: 3705: 3700: 3695: 3685: 3680: 3675: 3670: 3665: 3660: 3655: 3653:Mind-blindness 3650: 3645: 3640: 3635: 3630: 3629: 3628: 3623: 3618: 3613: 3608: 3597: 3595: 3584: 3583: 3581: 3580: 3575: 3570: 3565: 3560: 3555: 3550: 3545: 3540: 3527: 3522: 3516: 3514: 3508: 3507: 3505: 3504: 3499: 3498: 3497: 3487: 3486: 3485: 3475: 3474: 3473: 3468: 3463: 3453: 3448: 3447: 3446: 3436: 3435: 3434: 3429: 3418: 3416: 3410: 3409: 3407: 3406: 3405: 3404: 3399: 3394: 3384: 3379: 3374: 3365: 3364: 3363: 3358: 3348: 3346:theory of mind 3339: 3330: 3329: 3328: 3323: 3318: 3308: 3303: 3298: 3293: 3292: 3291: 3286: 3281: 3276: 3271: 3257: 3252: 3251: 3250: 3245: 3240: 3229: 3227: 3213: 3209: 3208: 3206: 3205: 3200: 3199: 3198: 3193: 3188: 3179: 3169: 3168: 3167: 3157: 3152: 3147: 3142: 3141: 3140: 3130: 3125: 3120: 3115: 3113:Baldwin effect 3110: 3109: 3108: 3103: 3098: 3088: 3082: 3080: 3072: 3071: 3069: 3068: 3063: 3062: 3061: 3056: 3051: 3046: 3041: 3031: 3030: 3029: 3018: 3015: 3014: 3007: 3006: 2999: 2992: 2984: 2975: 2974: 2972: 2971: 2961: 2950: 2947: 2946: 2944: 2943: 2938: 2933: 2928: 2923: 2922: 2921: 2911: 2906: 2901: 2896: 2891: 2890: 2889: 2884: 2879: 2869: 2864: 2859: 2853: 2851: 2847: 2846: 2844: 2843: 2838: 2837: 2836: 2831: 2826: 2825: 2824: 2814: 2809: 2804: 2799: 2794: 2784: 2778: 2776: 2772: 2771: 2769: 2768: 2763: 2758: 2753: 2748: 2743: 2738: 2733: 2728: 2723: 2722: 2721: 2712:Charles Darwin 2709: 2708: 2707: 2695: 2690: 2684: 2682: 2676: 2675: 2673: 2672: 2667: 2662: 2657: 2652: 2650:Non-ecological 2647: 2642: 2637: 2632: 2627: 2622: 2617: 2611: 2609: 2603: 2602: 2600: 2599: 2590: 2581: 2567: 2565: 2559: 2558: 2556: 2555: 2550: 2549: 2548: 2543: 2538: 2533: 2528: 2518: 2513: 2508: 2503: 2498: 2493: 2488: 2483: 2478: 2473: 2468: 2467: 2466: 2456: 2451: 2446: 2441: 2440: 2439: 2434: 2423: 2421: 2414: 2413: 2411: 2410: 2409: 2408: 2403: 2401:nervous system 2398: 2393: 2388: 2380: 2379: 2378: 2373: 2368: 2363: 2358: 2353: 2343: 2338: 2333: 2327: 2325: 2318: 2317: 2315: 2314: 2309: 2304: 2299: 2294: 2293: 2292: 2282: 2281: 2280: 2275: 2274: 2273: 2268: 2258: 2253: 2248: 2243: 2238: 2237: 2236: 2231: 2221: 2211: 2206: 2205: 2204: 2194: 2189: 2184: 2179: 2178: 2177: 2167: 2162: 2161: 2160: 2150: 2144: 2142: 2135: 2134: 2132: 2131: 2126: 2121: 2116: 2111: 2106: 2100: 2098: 2094: 2093: 2091: 2090: 2085: 2080: 2075: 2074: 2073: 2068: 2063: 2053: 2048: 2043: 2038: 2033: 2032: 2031: 2026: 2016: 2011: 2006: 2005: 2004: 1994: 1989: 1984: 1979: 1973: 1971: 1963: 1962: 1960: 1959: 1958: 1957: 1947: 1942: 1941: 1940: 1935: 1925: 1924: 1923: 1913: 1908: 1903: 1901:Origin of life 1898: 1893: 1888: 1886:Microevolution 1883: 1881:Macroevolution 1878: 1873: 1868: 1867: 1866: 1856: 1851: 1846: 1841: 1836: 1831: 1826: 1821: 1819:Common descent 1816: 1815: 1814: 1804: 1799: 1797:Baldwin effect 1794: 1793: 1792: 1787: 1777: 1772: 1767: 1761: 1759: 1753: 1752: 1750: 1749: 1744: 1739: 1734: 1729: 1723: 1720: 1719: 1712: 1711: 1704: 1697: 1689: 1683: 1682: 1677: 1670: 1669:External links 1667: 1666: 1665: 1637:(3): 301–310. 1622: 1604:(2): 110–117. 1590: 1546: 1544:on 2008-12-17. 1495: 1477:(2): 215–237. 1466: 1422: 1410:10.1086/276408 1386: 1368:(2): 283–298. 1355: 1352: 1350: 1349: 1306: 1257: 1252:10.1086/714999 1226: 1185: 1166: 1139:(2): 286–291. 1123: 1071: 1064: 1042: 1023:(3): 300–309. 1000: 950: 943: 921: 914: 888: 828: 806: 799: 774: 767: 747: 718:10.1086/677944 696: 689: 659: 641: 600:Osborne, H. F. 591: 541: 520: 509:(3): 363–373. 493: 486: 457: 445: 428: 421: 389: 377: 361: 359: 356: 355: 354: 348: 342: 336: 330: 322: 319: 296: 293: 273:Daniel Dennett 266:are typically 228: 227: 224: 221: 203:neo-Lamarckian 184: 181: 144:Daniel Dennett 98: 95: 62:Baldwin effect 15: 9: 6: 4: 3: 2: 4381: 4370: 4367: 4365: 4362: 4360: 4357: 4356: 4354: 4339: 4338: 4332: 4328: 4326: 4325: 4319: 4315: 4313: 4312: 4302: 4300: 4299: 4295: 4294: 4291: 4281: 4278: 4276: 4273: 4271: 4268: 4266: 4265:Neo-Darwinism 4263: 4261: 4258: 4256: 4253: 4251: 4250:Functionalism 4248: 4244: 4241: 4239: 4236: 4234: 4231: 4229: 4226: 4224: 4221: 4219: 4216: 4214: 4211: 4209: 4208:Connectionism 4206: 4204: 4201: 4200: 4199: 4198:indeterminism 4195: 4192: 4190: 4187: 4186: 4184: 4180: 4172: 4169: 4167: 4164: 4162: 4159: 4157: 4154: 4152: 4149: 4147: 4144: 4142: 4139: 4137: 4134: 4132: 4129: 4127: 4124: 4122: 4119: 4117: 4114: 4112: 4109: 4107: 4104: 4103: 4102: 4099: 4097: 4094: 4092: 4089: 4087: 4084: 4082: 4079: 4077: 4074: 4072: 4069: 4068: 4066: 4062: 4056: 4053: 4051: 4048: 4046: 4043: 4041: 4038: 4036: 4033: 4031: 4028: 4026: 4023: 4021: 4018: 4016: 4013: 4011: 4008: 4006: 4003: 4001: 3998: 3996: 3993: 3991: 3987: 3983: 3979: 3976: 3974: 3970: 3967: 3965: 3961: 3958: 3957: 3955: 3951: 3948: 3944: 3934: 3931: 3929: 3926: 3924: 3921: 3919: 3918:Schizophrenia 3916: 3914: 3911: 3909: 3906: 3904: 3903:Mental health 3901: 3899: 3896: 3894: 3891: 3889: 3886: 3884: 3881: 3879: 3876: 3874: 3871: 3869: 3866: 3864: 3861: 3859: 3856: 3854: 3851: 3850: 3848: 3846: 3842: 3836: 3833: 3831: 3828: 3826: 3822: 3818: 3815: 3813: 3809: 3806: 3804: 3801: 3799: 3796: 3794: 3791: 3789: 3786: 3784: 3781: 3779: 3778:Mate guarding 3776: 3774: 3771: 3769: 3766: 3764: 3761: 3759: 3756: 3754: 3751: 3749: 3746: 3744: 3741: 3739: 3738:Age disparity 3736: 3734: 3731: 3729: 3726: 3725: 3723: 3721: 3717: 3709: 3706: 3704: 3701: 3699: 3696: 3694: 3691: 3690: 3689: 3686: 3684: 3681: 3679: 3676: 3674: 3671: 3669: 3668:Schizophrenia 3666: 3664: 3661: 3659: 3656: 3654: 3651: 3649: 3646: 3644: 3641: 3639: 3636: 3634: 3631: 3627: 3624: 3622: 3619: 3617: 3614: 3612: 3609: 3607: 3604: 3603: 3602: 3599: 3598: 3596: 3594: 3593:Mental health 3589: 3588:Human factors 3585: 3579: 3578:Socialization 3576: 3574: 3571: 3569: 3566: 3564: 3561: 3559: 3556: 3554: 3551: 3549: 3546: 3544: 3541: 3539: 3538:paternal bond 3535: 3531: 3528: 3526: 3523: 3521: 3518: 3517: 3515: 3513: 3509: 3503: 3500: 3496: 3493: 3492: 3491: 3488: 3484: 3481: 3480: 3479: 3476: 3472: 3469: 3467: 3464: 3462: 3459: 3458: 3457: 3454: 3452: 3449: 3445: 3442: 3441: 3440: 3437: 3433: 3430: 3428: 3425: 3424: 3423: 3420: 3419: 3417: 3415: 3411: 3403: 3402:NaĂŻve physics 3400: 3398: 3395: 3393: 3390: 3389: 3388: 3385: 3383: 3380: 3378: 3375: 3373: 3369: 3368:Motor control 3366: 3362: 3359: 3357: 3354: 3353: 3352: 3349: 3347: 3343: 3340: 3338: 3334: 3331: 3327: 3326:Ophidiophobia 3324: 3322: 3319: 3317: 3316:Arachnophobia 3314: 3313: 3312: 3309: 3307: 3304: 3302: 3299: 3297: 3294: 3290: 3287: 3285: 3282: 3280: 3277: 3275: 3272: 3270: 3267: 3266: 3265: 3261: 3258: 3256: 3253: 3249: 3246: 3244: 3243:Display rules 3241: 3239: 3236: 3235: 3234: 3231: 3230: 3228: 3226: 3221: 3217: 3214: 3210: 3204: 3201: 3197: 3194: 3192: 3189: 3187: 3183: 3180: 3178: 3175: 3174: 3173: 3170: 3166: 3163: 3162: 3161: 3158: 3156: 3153: 3151: 3148: 3146: 3145:Kin selection 3143: 3139: 3136: 3135: 3134: 3131: 3129: 3126: 3124: 3121: 3119: 3116: 3114: 3111: 3107: 3104: 3102: 3099: 3097: 3094: 3093: 3092: 3089: 3087: 3084: 3083: 3081: 3079: 3073: 3067: 3064: 3060: 3057: 3055: 3052: 3050: 3047: 3045: 3042: 3040: 3039:Adaptationism 3037: 3036: 3035: 3032: 3028: 3025: 3024: 3023: 3020: 3019: 3016: 3012: 3005: 3000: 2998: 2993: 2991: 2986: 2985: 2982: 2970: 2966: 2962: 2960: 2952: 2951: 2948: 2942: 2939: 2937: 2934: 2932: 2929: 2927: 2924: 2920: 2917: 2916: 2915: 2914:Phylogenetics 2912: 2910: 2907: 2905: 2902: 2900: 2897: 2895: 2892: 2888: 2885: 2883: 2880: 2878: 2875: 2874: 2873: 2870: 2868: 2865: 2863: 2860: 2858: 2855: 2854: 2852: 2848: 2842: 2839: 2835: 2832: 2830: 2827: 2823: 2820: 2819: 2818: 2817:Structuralism 2815: 2813: 2810: 2808: 2805: 2803: 2800: 2798: 2795: 2793: 2792:Catastrophism 2790: 2789: 2788: 2785: 2783: 2780: 2779: 2777: 2773: 2767: 2764: 2762: 2759: 2757: 2754: 2752: 2751:Neo-Darwinism 2749: 2747: 2744: 2742: 2739: 2737: 2734: 2732: 2729: 2727: 2724: 2720: 2719: 2715: 2714: 2713: 2710: 2706: 2705: 2701: 2700: 2699: 2696: 2694: 2691: 2689: 2686: 2685: 2683: 2681: 2677: 2671: 2668: 2666: 2665:Reinforcement 2663: 2661: 2658: 2656: 2653: 2651: 2648: 2646: 2643: 2641: 2638: 2636: 2633: 2631: 2628: 2626: 2623: 2621: 2618: 2616: 2613: 2612: 2610: 2608: 2604: 2598: 2597:Catastrophism 2594: 2591: 2589: 2588:Macromutation 2585: 2584:Micromutation 2582: 2580: 2576: 2572: 2569: 2568: 2566: 2564: 2560: 2554: 2551: 2547: 2544: 2542: 2539: 2537: 2534: 2532: 2529: 2527: 2524: 2523: 2522: 2519: 2517: 2514: 2512: 2509: 2507: 2504: 2502: 2499: 2497: 2494: 2492: 2491:Immune system 2489: 2487: 2484: 2482: 2479: 2477: 2474: 2472: 2469: 2465: 2462: 2461: 2460: 2457: 2455: 2452: 2450: 2447: 2445: 2442: 2438: 2435: 2433: 2430: 2429: 2428: 2425: 2424: 2422: 2420: 2415: 2407: 2404: 2402: 2399: 2397: 2394: 2392: 2389: 2387: 2384: 2383: 2381: 2377: 2374: 2372: 2369: 2367: 2364: 2362: 2359: 2357: 2354: 2352: 2351:symbiogenesis 2349: 2348: 2347: 2344: 2342: 2339: 2337: 2334: 2332: 2329: 2328: 2326: 2324: 2319: 2313: 2310: 2308: 2305: 2303: 2300: 2298: 2295: 2291: 2288: 2287: 2286: 2283: 2279: 2276: 2272: 2269: 2267: 2264: 2263: 2262: 2259: 2257: 2254: 2252: 2249: 2247: 2244: 2242: 2239: 2235: 2232: 2230: 2227: 2226: 2225: 2222: 2220: 2217: 2216: 2215: 2212: 2210: 2207: 2203: 2200: 2199: 2198: 2195: 2193: 2190: 2188: 2185: 2183: 2180: 2176: 2173: 2172: 2171: 2168: 2166: 2163: 2159: 2156: 2155: 2154: 2151: 2149: 2146: 2145: 2143: 2141: 2136: 2130: 2127: 2125: 2122: 2120: 2117: 2115: 2112: 2110: 2107: 2105: 2102: 2101: 2099: 2095: 2089: 2086: 2084: 2081: 2079: 2076: 2072: 2069: 2067: 2064: 2062: 2059: 2058: 2057: 2054: 2052: 2049: 2047: 2044: 2042: 2039: 2037: 2034: 2030: 2027: 2025: 2022: 2021: 2020: 2019:Kin selection 2017: 2015: 2014:Genetic drift 2012: 2010: 2007: 2003: 2000: 1999: 1998: 1995: 1993: 1990: 1988: 1985: 1983: 1980: 1978: 1975: 1974: 1972: 1970: 1964: 1956: 1953: 1952: 1951: 1948: 1946: 1943: 1939: 1936: 1934: 1931: 1930: 1929: 1926: 1922: 1919: 1918: 1917: 1914: 1912: 1909: 1907: 1904: 1902: 1899: 1897: 1894: 1892: 1889: 1887: 1884: 1882: 1879: 1877: 1874: 1872: 1869: 1865: 1862: 1861: 1860: 1857: 1855: 1852: 1850: 1847: 1845: 1842: 1840: 1837: 1835: 1832: 1830: 1827: 1825: 1822: 1820: 1817: 1813: 1810: 1809: 1808: 1805: 1803: 1800: 1798: 1795: 1791: 1788: 1786: 1783: 1782: 1781: 1778: 1776: 1773: 1771: 1768: 1766: 1763: 1762: 1760: 1758: 1754: 1748: 1745: 1743: 1740: 1738: 1735: 1733: 1730: 1728: 1725: 1724: 1721: 1717: 1710: 1705: 1703: 1698: 1696: 1691: 1690: 1687: 1681: 1678: 1676: 1673: 1672: 1662: 1658: 1653: 1648: 1644: 1640: 1636: 1632: 1628: 1623: 1619: 1615: 1611: 1607: 1603: 1599: 1595: 1591: 1587: 1583: 1579: 1575: 1571: 1567: 1563: 1559: 1555: 1551: 1547: 1540: 1536: 1532: 1528: 1524: 1520: 1516: 1513:(2): 97–104. 1512: 1508: 1501: 1496: 1492: 1488: 1484: 1480: 1476: 1472: 1467: 1463: 1459: 1455: 1451: 1447: 1443: 1439: 1435: 1431: 1427: 1423: 1419: 1415: 1411: 1407: 1403: 1399: 1395: 1391: 1387: 1383: 1379: 1375: 1371: 1367: 1363: 1358: 1357: 1344: 1339: 1334: 1329: 1325: 1321: 1317: 1310: 1302: 1298: 1294: 1290: 1285: 1280: 1276: 1272: 1268: 1261: 1253: 1249: 1245: 1241: 1237: 1230: 1222: 1218: 1213: 1208: 1204: 1200: 1196: 1189: 1182: 1178: 1177: 1170: 1162: 1158: 1154: 1150: 1146: 1142: 1138: 1134: 1127: 1119: 1115: 1110: 1105: 1101: 1097: 1093: 1089: 1085: 1078: 1076: 1067: 1061: 1056: 1055: 1046: 1038: 1034: 1030: 1026: 1022: 1018: 1014: 1010: 1004: 996: 992: 988: 984: 980: 976: 972: 968: 964: 960: 954: 946: 944:9780262232296 940: 935: 934: 925: 917: 915:9780262232296 911: 907: 902: 901: 892: 884: 880: 876: 872: 867: 862: 858: 854: 850: 846: 842: 838: 832: 824: 820: 816: 810: 802: 796: 791: 790: 784: 778: 770: 764: 760: 759: 751: 743: 739: 735: 731: 727: 723: 719: 715: 711: 707: 700: 692: 686: 682: 681: 672: 670: 668: 666: 664: 655: 651: 645: 637: 633: 629: 625: 621: 617: 613: 609: 605: 601: 595: 587: 583: 579: 575: 571: 567: 563: 559: 555: 551: 550:Morgan, C. L. 545: 538: 537:Stigler's law 534: 530: 524: 516: 512: 508: 504: 497: 489: 483: 479: 478: 471: 466: 464: 462: 454: 449: 442: 437: 435: 433: 424: 418: 414: 409: 408: 402: 396: 394: 386: 381: 374: 373:Baldwin 1896a 369: 367: 362: 352: 349: 346: 343: 340: 337: 334: 331: 328: 325: 324: 318: 316: 312: 308: 302: 292: 290: 285: 283: 276: 274: 269: 263: 256: 251: 247: 244: 240: 235: 233: 225: 222: 219: 218: 217: 215: 210: 208: 207:Julian Huxley 204: 200: 196: 191: 180: 178: 173: 170: 166: 162: 156: 154: 147: 145: 141: 137: 133: 129: 122: 120: 119:neo-Darwinian 113: 111: 107: 104: 94: 92: 88: 83: 79: 75: 71: 67: 63: 59: 51: 47: 43: 39: 34: 30: 26: 22: 4335: 4322: 4309: 4296: 4055:Sociobiology 3913:Neuroscience 3893:Intelligence 3439:Anthropology 3392:Color vision 3377:Multitasking 3356:Flynn effect 3351:Intelligence 3333:Folk biology 3112: 3076:Evolutionary 2926:Polymorphism 2909:Astrobiology 2857:Biogeography 2812:Saltationism 2802:Orthogenesis 2787:Alternatives 2716: 2702: 2635:Cospeciation 2630:Cladogenesis 2579:Saltationism 2536:Mating types 2459:Color vision 2444:Avian flight 2366:mitochondria 2104:Canalisation 1982:Biodiversity 1796: 1727:Introduction 1680:Bibliography 1634: 1630: 1601: 1597: 1561: 1557: 1539:the original 1510: 1506: 1474: 1470: 1437: 1433: 1401: 1397: 1365: 1361: 1323: 1319: 1309: 1274: 1270: 1260: 1243: 1239: 1229: 1202: 1198: 1188: 1180: 1175: 1169: 1136: 1132: 1126: 1091: 1087: 1053: 1045: 1020: 1016: 1003: 970: 966: 953: 932: 924: 899: 891: 848: 844: 831: 822: 818: 809: 788: 777: 757: 750: 709: 705: 699: 679: 653: 644: 611: 607: 594: 561: 557: 544: 532: 523: 506: 502: 496: 476: 448: 441:Simpson 1953 406: 385:Baldwin 1897 380: 327:Evolvability 304: 286: 278: 265: 260: 242: 238: 236: 229: 211: 186: 174: 158: 152: 149: 124: 115: 103:psychologist 100: 81: 69: 61: 55: 29: 4194:Determinism 4106:Coevolution 4050:Primatology 3888:Gender role 3793:Orientation 3673:Screen time 3530:Affectional 3512:Development 3191:Mate choice 3118:By-products 3086:Adaptations 3049:Cognitivism 2936:Systematics 2807:Mutationism 2625:Catagenesis 2553:Snake venom 2486:Eusociality 2464:in primates 2454:Cooperation 2382:In animals 2202:butterflies 2175:Cephalopods 2165:Brachiopods 2097:Development 2071:Mate choice 1824:Convergence 1807:Coevolution 1765:Abiogenesis 783:Mayr, Ernst 527:Noble, R.; 453:Newman 2002 282:house finch 255:house finch 243:development 161:Jean Piaget 4353:Categories 4141:Population 4136:Lamarckism 3982:behavioral 3960:Behavioral 3908:Narcissism 3853:Aggression 3643:Hypophobia 3633:Depression 3520:Attachment 3502:Universals 3466:Psychology 3444:Biological 3432:Musicology 3422:Aesthetics 3321:Basophobia 3128:Exaptation 3106:Reciprocal 2797:Lamarckism 2775:Philosophy 2698:David Hume 2660:Peripatric 2655:Parapatric 2640:Ecological 2620:Anagenesis 2615:Allopatric 2607:Speciation 2571:Gradualism 2496:Metabolism 2356:chromosome 2346:Eukaryotes 2124:Modularity 2041:Population 1967:Population 1928:Speciation 1906:Panspermia 1859:Extinction 1854:Exaptation 1829:Divergence 1802:Cladistics 1790:Reciprocal 1770:Adaptation 1354:References 825:: 495–502. 345:Pangenesis 214:Ernst Mayr 165:Paul Weiss 112:explains: 78:Lamarckism 46:Waddington 4369:Selection 3986:cognitive 3978:Affective 3863:Cognition 3817:Sexuality 3803:Pair bond 3563:Education 3220:Cognition 3138:Inclusive 3078:processes 3066:Criticism 2931:Protocell 2782:Darwinism 2670:Sympatric 2419:processes 2307:Tetrapods 2256:Kangaroos 2182:Dinosaurs 2119:Inversion 2088:Variation 2009:Gene flow 2002:Inclusive 1812:Mutualism 1757:Evolution 1598:Evolution 1392:(1896a). 1271:Evolution 1011:(1896c). 961:(1896b). 726:0003-0147 529:Noble, D. 82:inherited 4255:Memetics 4015:Ethology 3973:genetics 3808:Physical 3773:Jealousy 3728:Activity 3534:maternal 3490:Religion 3478:Morality 3456:Language 3337:taxonomy 3150:Mismatch 3096:Cheating 3091:Altruism 2959:Category 2834:Vitalism 2829:Theistic 2822:Spandrel 2506:Morality 2501:Monogamy 2376:plastids 2341:Flagella 2297:Reptiles 2278:sea cows 2261:primates 2170:Molluscs 2148:Bacteria 2036:Mutation 1969:genetics 1945:Taxonomy 1891:Mismatch 1871:Homology 1785:Cheating 1780:Altruism 1661:22923852 1586:17734840 1552:(1896). 1527:11937679 1491:80821399 1462:17781159 1428:(1897). 1382:85267141 1326:(1826). 1293:17714500 1221:29763656 1205:: 4–13. 1161:20023879 1118:19324617 1037:17835006 995:17780356 839:(1987). 785:(1963). 734:25354416 652:(1901). 636:17734840 602:(1896). 586:17735249 552:(1896). 403:(1987). 321:See also 230:In 1987 136:organism 4156:Species 3928:Suicide 3763:Fantasy 3743:Arousal 3525:Bonding 3414:Culture 3238:Display 3225:Emotion 3133:Fitness 3022:History 2850:Related 2680:History 2541:Meiosis 2476:Empathy 2471:Emotion 2371:nucleus 2312:Viruses 2302:Spiders 2214:Mammals 2197:Insects 1997:Fitness 1933:Species 1732:Outline 1652:3423563 1618:2405746 1566:Bibcode 1558:Science 1535:1162454 1442:Bibcode 1434:Science 1418:7059820 1343:8059632 1301:9292273 1153:8799730 1109:2666683 975:Bibcode 967:Science 883:5476916 875:3670381 853:Bibcode 742:9214778 616:Bibcode 608:Science 566:Bibcode 558:Science 533:Biology 4334:  4321:  4308:  3898:Memory 3858:Autism 3825:female 3758:Desire 3495:Origin 3471:Speech 3461:Origin 3233:Affect 2969:Portal 2645:Hybrid 2481:Ethics 2323:organs 2285:Plants 2271:lemurs 2266:humans 2251:horses 2241:hyenas 2229:wolves 2224:canids 2158:origin 1659:  1649:  1616:  1584:  1533:  1525:  1489:  1460:  1416:  1380:  1340:  1299:  1291:  1219:  1159:  1151:  1116:  1106:  1062:  1035:  993:  941:  912:  908:–234. 881:  873:  845:Nature 797:  765:  740:  732:  724:  687:  634:  584:  484:  419:  167:, and 153:create 60:, the 44:, and 3868:Crime 3451:Crime 3382:Sleep 3372:skill 3212:Areas 2432:Death 2427:Aging 2406:brain 2192:Fungi 2153:Birds 2066:Fungi 1864:Event 1747:Index 1614:JSTOR 1542:(PDF) 1531:S2CID 1503:(PDF) 1487:S2CID 1414:S2CID 1378:S2CID 1297:S2CID 1157:S2CID 879:S2CID 738:S2CID 358:Notes 132:trait 3821:male 3182:Male 2919:Tree 2391:hair 2331:Cell 2234:dogs 2219:cats 2209:Life 2187:Fish 2140:taxa 1657:PMID 1582:PMID 1523:PMID 1458:PMID 1289:PMID 1217:PMID 1149:PMID 1114:PMID 1060:ISBN 1033:PMID 991:PMID 939:ISBN 910:ISBN 871:PMID 795:ISBN 763:ISBN 730:PMID 722:ISSN 685:ISBN 632:PMID 582:PMID 482:ISBN 417:ISBN 339:Meme 253:The 239:mind 3720:Sex 3397:Eye 2417:Of 2386:eye 2336:DNA 2321:Of 2138:Of 1647:PMC 1639:doi 1606:doi 1574:doi 1515:doi 1479:doi 1450:doi 1406:doi 1370:doi 1338:PMC 1328:doi 1324:376 1279:doi 1248:doi 1207:doi 1141:doi 1104:PMC 1096:doi 1092:364 1025:doi 983:doi 906:219 861:doi 849:329 714:doi 710:184 624:doi 574:doi 511:doi 413:399 313:'s 56:In 48:'s 4355:: 1655:. 1645:. 1635:39 1633:. 1629:. 1612:. 1600:. 1580:. 1572:. 1560:. 1556:. 1529:. 1521:. 1511:27 1509:. 1505:. 1485:. 1475:16 1473:. 1456:. 1448:. 1436:. 1432:. 1412:. 1402:30 1400:. 1396:. 1376:. 1366:19 1364:. 1336:. 1322:. 1318:. 1295:. 1287:. 1275:61 1273:. 1269:. 1244:88 1242:. 1238:. 1215:. 1203:88 1201:. 1197:. 1155:. 1147:. 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Index

Baldwin's rules
Baldwin effect (astronomy)

Lamarck's theory of evolution
Darwinian evolution
Waddington
genetic assimilation
evolutionary biology
James Mark Baldwin
natural selection
Lamarckism
Eclipse of Darwinism
modern synthesis
psychologist
James Mark Baldwin
Robert Richards
neo-Darwinian
evolutionary developmental biology
trait
organism
becomes gradually assimilated
Daniel Dennett
Jean Piaget
Paul Weiss
Conrad Waddington
Douglas Spalding
Maurice Maeterlinck
George Gaylord Simpson
modern synthesis
neo-Lamarckian

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