115:
1998:
140:
1993:
741:. They were mid-sized predators who probably hunted in trees or ambushed prey from a branch. Calculations of the size following a method of statistical analysis for predicting body size, derived from 164 millimetres for the greatest length of the skull, indicate a weight of 22.6 or 24.0 kilograms. Another estimate using
762:
in the northwest of
Queensland, at sediments dated to the later Oligocene to early Miocene. The assumption of at least partially arboreal habits is by the forest types that existed during the Oligo-Miocene at Riversleigh, open woodlands with the later development of rainforest. The humerus morphology
796:
The first fossil, a right dentary fragment, was found in 1967 by a group of field workers at
Bullock Creek in the Northern Territory. A left dentary of the species in a good state of preservation was described in 1896. Since then more fossils have been recovered, including a well-preserved
696:
is a largely complete skull retaining some teeth and alveoli, with other materials such as the lower jaw and a humerus associated with the new species as paratypes. The discovery of more complete evidence of
Oligocene species of the marsupial lion lineages prompted the authors to emend the
523:
Fossil material of this species is fragmentary and rare, and it is only known amongst the
Alcoota local fauna. Further evidence of the animals cranial and dental features were examined in 2014, leading to a revision of
723:, was determined to have diverged from this genus during early period of the fossil record, and scant evidence of the species did not allow revising authors to assign a taxonomic placement with any confidence.
1041:"New craniodental remains of Wakaleo alcootaensis (Diprotodontia: Thylacoleonidae) a carnivorous marsupial from the late Miocene Alcoota Local Fauna of the Northern Territory, Australia"
870:
B. Rauscher. 1987. Priscileo pitikantensis, a new genus and species of thylacoleonid marsupial (Marsupialia: Thylacoleonidae) from the
Miocene Etaduma formation, South Australia.
1326:
922:
1286:"Functional morphology of Wakaleo postcrania from the middle to late Miocene of central Australia reveals new insights in the evolution of marsupial hypercarnivores"
962:"Functional morphology of Wakaleo postcrania from the middle to late Miocene of central Australia reveals new insights in the evolution of marsupial hypercarnivores"
637:, and was approximately the size of a cat. They were mid-sized predators who probably hunted in trees or ambushed prey from a branch. Like the later discovery,
441:
910:
763:
is proposed to support this conception as a scansorial and arboreal predator, being similar to the structures of the arm and shoulder in species such as
427:
681:
651:
2086:
1709:
467:
2165:
751:, a very large carnivore, resulted in a value around 5 kg that the describing authors considered too small. The dentition suggests that
1927:
790:
1679:
1641:
1422:
416:
238:
1253:
Gillespie, A. K. (2023). "Two new marsupial lion taxa (Marsupialia, Thylacoleonidae) from the early and Middle
Miocene of Australia".
2073:
1353:
650:
It is known only from a few post cranial bones and a poorly preserved maxillary fragment found at the Lake
Pitikanta site of the
643:, the species possesses three premolars and four molars which distinguishes them from others of the genus. A little smaller than
2160:
1150:
Wroe, Stephen; Myers, Troy; Seebacher, Frank; Kear, Ben; Gillespie, Anna; Crowther, Mathew; Salisbury, Steve (September 2003).
2155:
2145:
817:
lifestyle of earlier species), and to grappling prey with their forelimbs, likely correlated with increasing prey size.
786:
517:
444:
in southern
Australia in 1971. They found a nearly complete left dentary which included a few well-preserved teeth.
2170:
759:
1167:
114:
482:
is a genus of the thylacoleonid family of predatory mammals, which are known as marsupial lions. The size of
1098:
1192:
914:
685:
2150:
1346:
755:
may have been omnivorous, but reveals the transition to hyper-carnivory of the genus during the
Miocene.
1285:
961:
516:
of
Australia. The holotype was a single fossil maxilla fragment found in 1974 by the palaeontologist
17:
1997:
139:
498:
are estimated to have had bodymasses of 30 kilograms (66 lb) and 50 kilograms (110 lb).
705:
490:
is estimated to have weighed approximately 23 kilograms (51 lb), comparable to a dog, while
709:, as a sister species to this taxon and contradicting a 2016 study that supported separation of
2117:
2013:
1339:
765:
618:. Fossil material examined by Rauscher was obtained at Lake Pitikanta, situated to the east of
2091:
923:"A new Oligo–Miocene marsupial lion from Australia and revision of the family Thylacoleonidae"
2112:
2104:
771:
1992:
1152:"An alternative method for predicting body mass: the case of the Pleistocene marsupial lion"
2060:
973:
719:
8:
839:
Clemens, W.A.; Plane, M. (1974). "Mid-Tertiary Thylacoleonidae (Marsupialia, Mammalia)".
814:
747:
742:
576:
570:
977:
1266:
1171:
1075:
1040:
997:
942:
692:
working on the PANGEA research project based at the University of New South Wales. The
565:. It had three unfused molar teeth instead of two fused molars as is the case with the
513:
431:
350:
134:
2099:
2051:
1305:
1270:
1080:
1062:
1015:
1001:
989:
848:
639:
602:
The description by Rauscher was published in 1987, naming a new species and genus as
1321:
1175:
1151:
946:
2140:
1784:
1739:
1297:
1258:
1236:
The Beagle: Occasional Papers of the Northern Territory Museum of Arts and Sciences
1163:
1132:
1070:
1052:
981:
934:
1301:
1262:
1227:
985:
938:
1829:
1820:
1809:
1749:
1573:
1461:
1413:
667:
562:
550:
393:
378:
214:
680:
The description of the species was published in 2017, the collaborating authors
1793:
1668:
1556:
727:
2134:
2036:
1948:
1892:
1530:
1392:
1362:
1309:
1066:
993:
852:
714:
412:
234:
201:
72:
588:
teeth, the latter were also used in slicing and stabbing during eating. The
508:
The first description was given in an examination of material discovered at
1955:
1802:
1634:
1618:
1473:
1084:
689:
300:
247:
123:
670:
and presumed to have diverged during the early to middle Oligocene epoch.
470:
in the Northern Territory. Many more specimens have been found since then.
2045:
1969:
1962:
1920:
1906:
1878:
1864:
1719:
1699:
1652:
1540:
566:
47:
730:, was honored by the authors in their proposed epithet for the species.
2078:
1934:
1857:
1843:
1604:
1513:
1443:
1057:
918:
813:
show increased adaptation to terrestrial locomotion (as opposed to the
92:
57:
1836:
1588:
1523:
1503:
1493:
1429:
630:
619:
608:
460:
Gillespie, Archer & Hand, 2017, a mid-sized species of the genus.
191:
151:
97:
2007:
1136:
2030:
1941:
1913:
1899:
1885:
1871:
1850:
1766:
1756:
1729:
1627:
1436:
1380:
693:
589:
585:
454:
in 1987 and placed with this genus in a revision published in 2017.
171:
87:
82:
67:
62:
52:
629:, it lived in Australia about 25 million years ago, from the late
726:
A painter celebrated for his reconstructions of ancient animals,
634:
581:
558:
546:
509:
486:
species increases over the course of the evolution of the genus.
397:
102:
77:
2065:
1331:
1204:
1690:
1597:
1483:
1386:
1374:
181:
161:
647:, this species also differs in the morphology of the humerus.
612:
was later recognised as a species of a revised description of
528:
that provided further support to the separation from earlier
389:
1284:
Warburton, Natalie M.; Yates, Adam. M. (10 December 2020).
1168:
10.1666/0094-8373(2003)029<0403:AAMFPB>2.0.CO;2
960:
Warburton, Natalie M.; Yates, Adam. M. (10 December 2020).
434:
in 1974. It was slightly larger than the other two species.
440:
was found by a group of scientists working in the Miocene
1123:
Myers, T.J. (2001). "Prediction of marsupial body mass".
466:
was first found by field workers in 1967 in the Miocene
737:
and resembling a contemporary animal of the Oligocene,
450:
described by Rauscher as the type species of the genus
1149:
809:
are thought to have been carnivores. Later species of
785:
The type species fossils were originally found in the
592:
also had a crescent-shaped circumference for slicing.
1781:
1762:
1745:
1735:
1725:
1715:
1705:
1675:
1664:
1615:
1585:
1569:
1536:
1519:
1509:
1499:
1489:
1479:
1469:
1410:
1255:
Alcheringa: An Australasian Journal of Palaeontology
1099:"Thylacoleonid Genera and Species - Tertiary Genera"
333:
320:
307:
290:
277:
254:
909:
1008:
606:.The designation as the type species of the genus
396:that lived in Australia in the Late Oligocene and
789:area. It has been found in limestone deposits in
745:previously used in calculations of body size for
2132:
905:
903:
797:near-complete cranium with the mandible intact.
652:Lake Ngapakaldi to Lake Palankarinna Fossil Area
901:
899:
897:
895:
893:
891:
889:
887:
885:
883:
584:(upper) teeth to hold its food and sharpen the
1283:
1187:
1185:
959:
703:to include the type species of another genus,
1347:
1246:
834:
832:
830:
758:The distribution of the species included the
1219:
880:
838:
580:, this species is presumed to have used its
1182:
1034:
1032:
1354:
1340:
872:Possums and Opossums: Studies in Evolution
827:
113:
1252:
1074:
1056:
654:in northeastern South Australia. To date
1225:
1116:
1029:
769:, the related koala, and spotted cuscus
666:are the most ancient phylogenies of the
595:
1143:
501:
14:
2133:
778:
713:from the wakaleo clade. Another early
2012:
2011:
1335:
1122:
1038:
2166:Taxa named by William A. Clemens Jr.
866:
864:
862:
673:
535:
1039:Yates, Adam M. (12 November 2015).
927:Journal of Systematic Palaeontology
24:
1322:Tertiary Genera of Thylacoleonidae
1290:Journal of Vertebrate Paleontology
966:Journal of Vertebrate Paleontology
787:Bullock Creek (Northern Territory)
329:Gillespie, Archer & Hand, 2017
25:
2182:
1361:
859:
1996:
1991:
1226:Megirian, D. (28 January 1986).
138:
45:
1277:
1232:(Thylacoleonidae: Marsupialia)"
1201:Riversleigh faunal encyclopedia
760:Riversleigh World Heritage Area
384:, "little", "small"; and Latin
1091:
953:
625:The smallest known species of
474:
13:
1:
2161:Fossil taxa described in 1974
1302:10.1080/02724634.2021.1878203
1263:10.1080/03115518.2022.2152096
1203:. wakaleo.net. Archived from
1125:Australian Journal of Zoology
1016:"Australian Museum - Wakaleo"
986:10.1080/02724634.2021.1878203
939:10.1080/14772019.2017.1391885
820:
2156:Prehistoric marsupial genera
2146:Miocene mammals of Australia
7:
403:
299:Clemens & Plane, 1974 (
122:Life reconstruction of the
43:Late Oligocene–Late Miocene
27:Extinct genus of marsupials
10:
2187:
1197:Vanderleuer's Little Lion"
800:
419:. Five species are known:
2020:
1989:
1819:
1792:
1780:
1689:
1660:
1651:
1614:
1584:
1565:
1554:
1460:
1409:
1405:
1401:
1369:
1327:Mikko's Phylogeny Archive
426:was found in the Miocene
356:
349:
342:Clemens & Plane, 1974
274:
269:
264:Clemens & Plane, 1974
253:
246:
135:Scientific classification
133:
121:
112:
34:
388:, "lion") is an extinct
841:Journal of Paleontology
805:The various species of
706:Priscileo pitikantensis
604:Priscileo pitikantensis
411:was erected in 1974 by
360:Priscileo pitikantensis
286:Archer & Rich, 1984
2171:Carnivorous marsupials
766:Phascolarctos cinereus
733:A species assigned to
720:"Priscileo" roskellyae
664:"Priscileo' roskellyae
2113:Paleobiology Database
1103:www.naturalworlds.org
772:Spilocuscus maculatus
739:Wakaleo pitikantensis
597:Wakaleo pitikantensis
448:Wakaleo pitikantensis
1230:Wakaleo vanderleueri
1195:Wakaleo vanderleueri
743:regression equations
526:Wakaleo alcootaensis
503:Wakaleo alcootaensis
424:Wakaleo alcootaensis
978:2020JVPal..40E8203W
921:(6 December 2017).
780:Wakaleo vanderleuri
748:Thylacoleo carnifex
697:circumscription of
577:Thylacoleo carnifex
571:Thylacoleo carnifex
464:Wakaleo vanderleuri
2151:Miocene marsupials
1058:10.7717/peerj.1408
514:Northern Territory
442:Wipijiri Formation
432:Northern Territory
2128:
2127:
2100:Open Tree of Life
2014:Taxon identifiers
2005:
2004:
1987:
1986:
1983:
1982:
1979:
1978:
1776:
1775:
1550:
1549:
675:Wakaleo schouteni
660:Wakaleo schouteni
640:Wakaleo schouteni
543:Wakaleo oldfieldi
537:Wakaleo oldfieldi
458:Wakaleo schouteni
438:Wakaleo oldfieldi
370:
369:
364:
343:
330:
317:
304:
287:
259:Wakaleo oldfieldi
242:
128:Wakaleo oldfieldi
16:(Redirected from
2178:
2121:
2120:
2108:
2107:
2095:
2094:
2082:
2081:
2069:
2068:
2056:
2055:
2054:
2041:
2040:
2039:
2009:
2008:
2000:
1995:
1790:
1789:
1785:Diprotodontoidea
1783:
1764:
1747:
1740:Rhizophascolonus
1737:
1727:
1717:
1707:
1677:
1666:
1658:
1657:
1617:
1587:
1571:
1563:
1562:
1538:
1521:
1511:
1501:
1491:
1481:
1471:
1412:
1407:
1406:
1403:
1402:
1356:
1349:
1342:
1333:
1332:
1314:
1313:
1281:
1275:
1274:
1250:
1244:
1243:
1228:"The dentary of
1223:
1217:
1216:
1214:
1212:
1207:on 10 March 2015
1189:
1180:
1179:
1147:
1141:
1140:
1120:
1114:
1113:
1111:
1109:
1095:
1089:
1088:
1078:
1060:
1036:
1027:
1026:
1024:
1022:
1012:
1006:
1005:
957:
951:
950:
907:
878:
868:
857:
856:
836:
711:P. pitikantensis
656:W. pitikantensis
392:of medium-sized
362:
341:
335:
328:
322:
315:
312:W. pitikantensis
309:
298:
292:
285:
279:
256:
233:
226:
213:
143:
142:
117:
107:
44:
40:Temporal range:
32:
31:
21:
2186:
2185:
2181:
2180:
2179:
2177:
2176:
2175:
2131:
2130:
2129:
2124:
2116:
2111:
2103:
2098:
2090:
2085:
2077:
2072:
2064:
2059:
2050:
2049:
2044:
2035:
2034:
2029:
2016:
2006:
2001:
1975:
1830:Alkwertatherium
1821:Diprotodontidae
1815:
1810:Propalorchestes
1772:
1750:Sedophascolomys
1685:
1647:
1610:
1580:
1574:Thylacoleonidae
1559:
1546:
1462:Phascolarctidae
1456:
1414:Thylacoleonidae
1397:
1365:
1360:
1318:
1317:
1296:(6): e1878203.
1282:
1278:
1251:
1247:
1224:
1220:
1210:
1208:
1191:
1190:
1183:
1148:
1144:
1137:10.1071/ZO01009
1121:
1117:
1107:
1105:
1097:
1096:
1092:
1037:
1030:
1020:
1018:
1014:
1013:
1009:
972:(6): e1878203.
958:
954:
911:Gillespie, A.K.
908:
881:
869:
860:
837:
828:
823:
803:
783:
678:
668:Thylacoleonidae
600:
563:South Australia
551:marsupial lions
540:
506:
496:W. alcootaensis
492:W. vanderleueri
477:
428:Waite Formation
406:
282:W. alcootaensis
265:
262:
232:
224:
215:Thylacoleonidae
211:
137:
108:
106:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
42:
41:
38:
28:
23:
22:
15:
12:
11:
5:
2184:
2174:
2173:
2168:
2163:
2158:
2153:
2148:
2143:
2126:
2125:
2123:
2122:
2109:
2096:
2083:
2070:
2057:
2042:
2026:
2024:
2018:
2017:
2003:
2002:
1990:
1988:
1985:
1984:
1981:
1980:
1977:
1976:
1974:
1973:
1966:
1959:
1952:
1945:
1938:
1931:
1924:
1917:
1910:
1903:
1896:
1889:
1882:
1875:
1868:
1861:
1854:
1847:
1840:
1833:
1825:
1823:
1817:
1816:
1814:
1813:
1806:
1798:
1796:
1794:Palorchestidae
1787:
1778:
1777:
1774:
1773:
1771:
1770:
1760:
1753:
1743:
1733:
1723:
1713:
1703:
1695:
1693:
1687:
1686:
1684:
1683:
1673:
1661:
1655:
1649:
1648:
1646:
1645:
1638:
1631:
1623:
1621:
1612:
1611:
1609:
1608:
1601:
1593:
1591:
1582:
1581:
1579:
1578:
1566:
1560:
1557:Vombatomorphia
1555:
1552:
1551:
1548:
1547:
1545:
1544:
1534:
1527:
1517:
1507:
1497:
1487:
1477:
1466:
1464:
1458:
1457:
1455:
1454:
1447:
1440:
1433:
1426:
1418:
1416:
1399:
1398:
1396:
1395:
1389:
1383:
1377:
1370:
1367:
1366:
1359:
1358:
1351:
1344:
1336:
1330:
1329:
1324:
1316:
1315:
1276:
1245:
1218:
1181:
1162:(3): 403–411.
1142:
1115:
1090:
1028:
1007:
952:
879:
858:
847:(4): 653–660.
825:
824:
822:
819:
802:
799:
782:
777:
728:Peter Schouten
682:Anna Gillespie
677:
672:
599:
594:
539:
534:
518:Michael Archer
505:
500:
476:
473:
472:
471:
461:
455:
445:
435:
405:
402:
394:thylacoleonids
368:
367:
366:
365:
363:Rauscher, 1987
354:
353:
347:
346:
345:
344:
338:W. vanderleuri
331:
318:
316:Rauscher, 1987
305:
288:
272:
271:
267:
266:
263:
251:
250:
244:
243:
222:
218:
217:
209:
205:
204:
199:
195:
194:
189:
185:
184:
179:
175:
174:
169:
165:
164:
159:
155:
154:
149:
145:
144:
131:
130:
119:
118:
110:
109:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
39:
26:
9:
6:
4:
3:
2:
2183:
2172:
2169:
2167:
2164:
2162:
2159:
2157:
2154:
2152:
2149:
2147:
2144:
2142:
2139:
2138:
2136:
2119:
2114:
2110:
2106:
2101:
2097:
2093:
2088:
2084:
2080:
2075:
2071:
2067:
2062:
2058:
2053:
2047:
2043:
2038:
2032:
2028:
2027:
2025:
2023:
2019:
2015:
2010:
1999:
1994:
1972:
1971:
1967:
1965:
1964:
1960:
1958:
1957:
1953:
1951:
1950:
1949:Raemeotherium
1946:
1944:
1943:
1939:
1937:
1936:
1932:
1930:
1929:
1925:
1923:
1922:
1918:
1916:
1915:
1911:
1909:
1908:
1904:
1902:
1901:
1897:
1895:
1894:
1893:Meniscolophus
1890:
1888:
1887:
1883:
1881:
1880:
1876:
1874:
1873:
1869:
1867:
1866:
1862:
1860:
1859:
1855:
1853:
1852:
1848:
1846:
1845:
1841:
1839:
1838:
1834:
1832:
1831:
1827:
1826:
1824:
1822:
1818:
1812:
1811:
1807:
1805:
1804:
1800:
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324:
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295:W. oldfieldi
294:
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281:
280:
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257:
248:Type species
228:
227:
188:Infraclass:
127:
124:type species
35:
29:
2046:Wikispecies
1970:Zygomaturus
1963:Sthenomerus
1921:Nototherium
1907:Ngapakaldia
1879:Kolopsoides
1865:Hulitherium
1720:Phascolonus
1700:Lasiorhinus
1653:Vombatoidea
1541:Priscakoala
791:Riversleigh
686:Mike Archer
567:Pleistocene
557:, found in
549:species of
475:Description
192:Marsupialia
2135:Categories
1935:Plaisiodon
1928:Pitikantia
1858:Euryzygoma
1844:Diprotodon
1691:Vombatidae
1605:Kuterintja
1514:Nimiokoala
1444:Thylacoleo
1021:17 January
919:Hand, S.J.
915:Archer, M.
821:References
633:to middle
586:mandibular
574:. As with
1837:Ambulator
1680:Mukupirna
1642:Wynyardia
1589:Ilariidae
1524:Perikoala
1504:Madakoala
1494:Litokoala
1430:Lekaneleo
1423:Enigmaleo
1373:Kingdom:
1310:0272-4634
1271:256157821
1131:(2): 99.
1108:1 October
1067:2167-8359
1051:: e1408.
1002:233793061
994:0272-4634
853:0022-3360
717:species,
631:Oligocene
620:Lake Eyre
609:Priscileo
590:premolars
582:maxillary
532:species.
452:Priscileo
158:Kingdom:
152:Eukaryota
18:Priscileo
2031:Wikidata
1942:Pyramios
1914:Nimbadon
1900:Neohelos
1886:Maokopia
1872:Kolopsis
1851:Euowenia
1767:Warendja
1757:Vombatus
1730:Ramsayia
1628:Muramura
1437:Microleo
1387:Mammalia
1381:Chordata
1379:Phylum:
1375:Animalia
1257:: 1–16.
1242:: 71–79.
1176:85776991
1085:26587359
947:90758394
877::423-432
815:arboreal
694:holotype
417:M. Plane
404:Taxonomy
400:Epochs.
351:Synonyms
270:Species
208:Family:
182:Mammalia
172:Chordata
168:Phylum:
162:Animalia
148:Domain:
2141:Wakaleo
2105:3607108
2092:1030255
2079:4826094
2066:4529428
2052:Wakaleo
2037:Q140445
2022:Wakaleo
1451:Wakaleo
1391:Order:
1385:Class:
1076:4647553
974:Bibcode
811:Wakaleo
807:Wakaleo
801:Ecology
735:Wakaleo
700:Wakaleo
635:Miocene
627:Wakaleo
615:Wakaleo
559:Miocene
555:Wakaleo
547:extinct
530:Wakaleo
512:in the
510:Alcoota
484:Wakaleo
480:Wakaleo
430:in the
409:Wakaleo
398:Miocene
374:Wakaleo
235:Clemens
229:Wakaleo
221:Genus:
198:Order:
178:Class:
36:Wakaleo
1669:Marada
1598:Ilaria
1484:Koobor
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1211:28 May
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241:, 1974
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1267:S2CID
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1045:PeerJ
998:S2CID
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390:genus
239:Plane
2074:GBIF
1306:ISSN
1213:2019
1110:2022
1081:PMID
1063:ISSN
1023:2011
990:ISSN
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688:and
662:and
494:and
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301:type
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