4263:
382:
2352:
4258:
136:
4286:
118:
2496:
607:
similar morphology to the median thoracic osteoderms of the holotype specimen of
Scolosaurus. The osteoderm has a length of 15 cm, a width of 12 cm and a height of 10 cm. The smallest ossicle has a diameter of 1 cm. The surface texture of the osteoderms are smooth to uniformly pitted with a sparse patterns of grooves as in
606:
osteoderm and numerous ossicles. The osteoderm and ossicles of the right scapula and another series of osteoderms preserved in situ display a rosette arrangement of small ossicles surrounding the osteoderms. The largest osteoderm pertains to the thoracic region behind the cervical region based on the
589:
fragments that probably represent parts of the cervicals and/or dorsals were recovered and the position of the vertebrae are based on their association with elements of the pectoral girdles and forelimb. A few vertebrae were dorsoventrally tall dorsal vertebrae, based on a few of the more complete
573:
is massive and has a length of 31 cm. The left humerus preserves a deltopectoral crest that is well-developed and measures 15 cm across the widest portion of the humerus. The lateral margin of the deltopectoral crest forms a 23° angle in posterior view with the long axis of the humerus. Bounded by
564:
shearing, the prominent overhang of the acromion on the left scapula is absent, although this feature is not a taphonomic artefact on the right scapulocoracoid. However, the right scapulocoracoid is cracked, its junction with the scapular blade is visible, well-preserved, and verifies its natural
397:
was discovered from the Hunter Wash Member in the lower part of the
Kirtland Formation, New Mexico at the Ah-shi-sle-pah Wilderness Study Area. The specimen was later collected over consecutive field seasons from 2005 to 2009 and was subsequently described in 2011 by Michael Burns and
2958:
2413:. Based on the abundance of angiosperms with leaves that have entire or nearly entire margins and drip points, the Kirtland Formation may have had a warm temperate to subtropical climate. The Kirtland Formation was better drained than the underlying
527:, by the superficial osteodermal surface texture, characterized by uniformly distributed pitted rugosity, and sparse distribution of reticular neurovascular grooves with neurovascular foramina extending perpendicularly to obliquely into the bone.
574:
the deltopectoral crest and humeral head is a broad, bicipital fossa that is present on the anterior face of the humerus. On the anterior face of the humerus is a hemispherical radial condyle. The proximal articular surface of the left
559:
is present along the dorsal margin of the scapula opposite the glenoid cavity. The acromion attains its front most extension towards the back and sides to the most anterior extent of the glenoid cavity. Due to
598:
Numerous osteoderms and associated osteoderm fragments were associated with the holotype specimen. Most osteoderms pertain to the thoracic region while others might pertain to the pelvic region and
555:, the scapula does not narrow towards the upper surface and sides relative to the coracoid and does not exhibit a scapular neck. A rugose, ridge-like structure that is identified as the
2894:"A New Ankylosaurid Dinosaur from the Upper Cretaceous (Kirtlandian) of New Mexico with Implications for Ankylosaurid Diversity in the Upper Cretaceous of Western North America"
2678:
Arbour, V.M.; Zanno, L.E.; Gates, T. (2016). "Ankylosaurian dinosaur palaeoenvironmental associations were influenced by extirpation, sea-level fluctuation, and geodispersal".
3033:"Re-evaluation of pachycephalosaurids from the Fruitland-Kirtland transition (Kirtlandian, late Campanian), San Juan Basin, New Mexico, with a description of a new species of
2959:"The Kirtlandian land-vertebrate "age"-faunal composition, temporal position and biostratigraphic correlation in the nonmarine Upper Cretaceous of western North America"
4433:
2892:
Arbour, Victoria M.; Burns, Michael E.; Sullivan, Robert M.; Lucas, Spencer G.; Cantrell, Amanda K.; Fry, Joshua; Suazo, Thomas L. (24 September 2014).
4379:
2556:"A new ankylosaurid from the Upper Cretaceous Kirtland Formation, San Juan Basin, with comments on the diversity of ankylosaurids in New Mexico"
551:. The glenoid cavity forms an arc that projects the scapular margin posteroventrally at an 80° angle relative to the coracoid margin. As in
569:
are prominent and creates an area or the attachment of muscles. The left forelimb is more than 50% complete. As in ankylosaurids, the left
4453:
590:
vertebrae appearing to have laterally compressed neural canals. All other vertebrae lack open sutural facets for the neural arches.
1636:
In contrast, an earlier analysis by Arbour & Currie (2015) favors an ankylosaurid position. Their results are reproduced below.
4366:
476:
of the vertebrae, and the re-modelled osteoderm cores. The holotype falls within the same size range as adult specimens of other
2715:"The most basal ankylosaurine dinosaur from the Albian–Cenomanian of China, with implications for the evolution of the tail club"
3170:
4443:
3217:
3032:
4262:
3175:, n. gen., n. sp., (Pterosauria, ?Azhdarchidae) from the Upper Cretaceous Kirtland Formation (upper Campanian) of New Mexico"
602:. All osteoderms are either keeled or circular with an off-center apex. The distal portion of the right scapula preserves an
4458:
535:
The right scapula preserves osteoderms on the lateral side at the most distal portion and is associated with fragments of
4448:
3238:
3126:
2773:
Rivera-Sylva, H.E.; Frey, E.; Stinnesbeck, W.; Carbot-Chanona, G.; Sanchez-Uribe, I.E.; Guzmán-Gutiérrez, J.R. (2018).
578:
is oval in proximal view and is concave. The articular surface is 8 cm long, proportionally twice the diameter of the
2635:
Arbour, V. M.; Currie, P. J. (2015). "Systematics, phylogeny and palaeobiogeography of the ankylosaurid dinosaurs".
3067:"Transitional evolutionary forms in chasmosaurine ceratopsid dinosaurs: evidence from the Campanian of New Mexico"
4279:
2509:
3117:
344:
2990:
4463:
1946:
1478:
1454:
2991:"New Late Cretaceous leaf locality from lower Kirtland Shale member, Bisti area, San Juan Basin, New Mexico"
513:
based on the dorsolateral overhang of the scapular acromion process to 25% of the dorsoventral width of the
4257:
3122:, gen. et sp. nov., a new tyrannosauroid from New Mexico and the origin of deep snouts in Tyrannosauroidea"
435:
135:
749:(2018) is reproduced below, favoring a nodosaurid position. Clade names have been added following Madzia
3210:
480:
taxa outside of North
America and represents an individual that was larger than juvenile specimens of
1495:
450:
3308:
565:
projection towards the lower edge and sides. Towards the sides of the acromion, the borders of the
439:
434:, and other unidentifiable postcranial fragments. The holotype specimen is currently housed at the
641:
at the base of
Ankylosauridae, and Wiersma and Irmis (2018) placed it in a polytomy consisting of
4344:
2394:
2369:
is known from the Hunter Wash Member of the
Kirtland Formation which has been dated to the upper
4410:
2775:"Paleodiversity of Late Cretaceous Ankylosauria from Mexico and their phylogenetic significance"
461:" (shield). The specific name, minor, is in reference to its small adult size relative to other
4306:
4384:
4405:
4397:
3335:
3203:
2830:
Madzia, D.; Arbour, V.M.; Boyd, C.A.; Farke, A.A.; CruzadoCaballero, P.; Evans, D.C. (2021).
1929:
1309:
272:
2713:
Wenjie Zheng; Xingsheng Jin; Yoichi Azuma; Qiongying Wang; Kazunori Miyata; Xing Xu (2018).
4438:
4353:
4289:
3135:
2911:
2786:
2726:
2687:
2644:
2424:
1292:
249:
8:
3695:
3592:
2414:
1664:
637:
469:
3139:
2915:
2790:
2730:
2691:
2648:
3431:
3151:
3093:
3066:
3013:
2934:
2893:
2871:
2858:
2831:
2812:
2747:
2714:
2660:
2609:
2583:
gen. et sp. nov., from the upper
Campanian Kaiparowits Formation of southern Utah, USA"
2578:
566:
423:
399:
348:
330:
130:
4392:
4188:
3452:
3098:
3017:
2939:
2875:
2863:
2816:
2804:
2752:
2664:
2614:
2555:
2117:
701:
628:
547:. The coracoid is roughly 16 cm long and the coracoid foramen is anterodorsal to the
472:
or adult individual based on the complete fusion of the scapulocoracoids, centra and
381:
3155:
686:
On the other hand, several other studies have favored a nodosaurid position. Arbour
543:
and has a blunt, rugose process along the anterodorsal margin for the attachment of
3653:
3600:
3438:
3143:
3088:
3078:
3005:
2929:
2919:
2853:
2843:
2794:
2742:
2734:
2712:
2695:
2652:
2604:
2594:
2501:
2464:
427:
2656:
2351:
4118:
4082:
4022:
3988:
3981:
3874:
3815:
3687:
3584:
3500:
3460:
3293:
2924:
2699:
2461:
2410:
1904:
1731:
1349:
1115:
872:
848:
714:
544:
415:
411:
322:
304:
30:
611:. One osteoderm has a histological condition typical of that of ankylosaurids.
4329:
4181:
4167:
4153:
4061:
4034:
3930:
3911:
3826:
3772:
3637:
3607:
3577:
3540:
3516:
2738:
2480:
2468:
2451:
2440:
2433:
2200:
2174:
1643:
1179:
1065:
815:
696:
667:
643:
624:
523:
477:
3147:
2799:
2774:
539:
that are mostly visible on the medial side. The scapula is fully fused to the
4427:
4144:
4054:
4047:
3999:
3890:
3856:
3703:
3678:
3660:
3563:
3509:
3482:
3474:
2808:
1862:
1370:
1217:
906:
892:
575:
548:
462:
454:
419:
68:
4174:
4160:
4089:
3973:
3897:
3842:
3793:
3765:
3750:
3737:
3710:
3629:
3614:
3570:
3550:
3417:
3399:
3377:
3347:
3320:
3268:
3226:
3182:
Fossil Record 3. New Mexico Museum of
Natural History and Science, Bulletin
3102:
3048:
Fossil Record 3. New Mexico Museum of
Natural History and Science, Bulletin
2943:
2867:
2756:
2618:
2560:
Fossil Record 3. New Mexico Museum of
Natural History and Science, Bulletin
2473:
2446:
2436:
2157:
2069:
1420:
1275:
1024:
959:
945:
798:
760:
679:
673:
552:
492:
based on humeral size; however, the scapulocoracoid is larger than that of
394:
369:
357:
311:
228:
196:
4338:
4209:
4126:
4075:
4015:
3966:
3808:
3786:
3718:
3667:
3467:
3445:
3277:
3009:
2772:
2224:
1886:
1829:
1772:
1241:
1139:
923:
781:
720:
655:
212:
43:
2848:
4371:
4202:
4195:
4133:
4008:
3959:
3951:
3904:
3849:
3835:
3801:
3779:
3645:
3489:
3424:
3083:
2599:
2428:
2398:
2027:
1812:
1755:
1714:
1437:
1193:
1041:
1000:
730:
649:
340:
318:
88:
53:
4216:
4096:
3883:
3863:
3725:
3622:
3524:
2476:
2378:
2374:
2370:
2100:
1986:
1387:
1258:
983:
579:
561:
431:
363:
326:
147:
93:
37:
4300:
3030:
4323:
4223:
3259:
3250:
2898:
2456:
2382:
2241:
691:
661:
599:
586:
556:
540:
473:
403:
390:
314:
180:
167:
117:
83:
78:
63:
58:
48:
3168:
2963:
Late
Cretaceous Vertebrates from the Western Interior: Bulletin 35
2553:
4358:
4111:
4103:
2406:
2010:
570:
514:
98:
73:
2576:
3244:
3195:
2373:
stage, 74.5 Ma. The Kirtland Formation consists of interbedded
157:
2989:
R. Robison, Coleman; Hunt, Adrian; L. Wolberg, Donald (1982).
4068:
2390:
329:
stage, 74.5 Ma) in what is now the Hunter Wash Member of the
307:
4230:
2907:
2402:
2393:. The Kirtland Formation lies on the western margin of the
2386:
2832:"The phylogenetic nomenclature of ornithischian dinosaurs"
2634:
536:
343:
of a small subadult or adult individual. It was named in
2988:
2891:
521:
differs from other ankylosaurids, with the exception of
2961:. In G. Lucas, Spencer; M. Sullivan, Sullivan (eds.).
3064:
2829:
453:, the locality from which the holotype came, and the
2491:
385:
Location and stratigraphy of the Kirtland Formation.
2956:
3169:Robert M. Sullivan & Denver W. Fowler (2011).
3109:
2768:
2766:
2554:Michael E. Burns & Robert M. Sullivan (2011).
745:A phylogenetic analysis conducted by Rivera-Sylva
488:but smaller than Talarurus and adult specimens of
2680:Palaeogeography, Palaeoclimatology, Palaeoecology
627:, although this was not based on the result of a
4425:
2677:
3116:Carr, Thomas D.; Williamson, Thomas E. (2010).
3031:Steven E. Jasinski; Robert M. Sullivan (2011).
3024:
2984:
2982:
2980:
2957:M. Sullivan, Robert; G. Lucas, Spencer (2006).
2763:
3292:
3162:
3115:
631:. Likewise, Arbour & Currie (2015) placed
619:Burns & Sullivan (2011) originally placed
3211:
2887:
2885:
430:fragments, complete and fragmentary thoracic
3065:Fowler, D.W.; Freedman Fowler, E.A. (2020).
2977:
2950:
2630:
2628:
2549:
2547:
2545:
2358:, a pachycephalosaurid contemporaneous with
303:(meaning "Ah-shi-sle-pah Wash shield") is a
3397:
2577:Jelle P. Wiersma; Randall B. Irmis (2018).
2543:
2541:
2539:
2537:
2535:
2533:
2531:
2529:
2527:
2525:
4434:Late Cretaceous dinosaurs of North America
3218:
3204:
3058:
2882:
2570:
734:and more deeply nested taxa. Rivera-Sylva
116:
3092:
3082:
2933:
2923:
2857:
2847:
2823:
2798:
2746:
2671:
2625:
2608:
2598:
2579:"A new southern Laramidian ankylosaurid,
2522:
2350:
504:
380:
376:
4426:
509:Burns & Sullivan (2011) diagnosed
4305:
4304:
3237:
3199:
738:(2018) placed it as sister taxon to
623:as an ankylosaurid within the clade
351:. Based on the size of the humerus,
2706:
2637:Journal of Systematic Palaeontology
2346:
700:and a juvenile nodosaurid from the
468:The holotype specimen represents a
339:, is known only from an incomplete
13:
3127:Journal of Vertebrate Paleontology
14:
4475:
4454:Campanian genus first appearances
614:
4285:
4284:
4261:
4256:
3225:
2494:
2417:due to the lack of coal swamps.
134:
41:
2510:Timeline of ankylosaur research
389:In 2005, a partial postcranial
2779:Swiss Journal of Palaeontology
499:
1:
4444:Fossil taxa described in 2011
2657:10.1080/14772019.2015.1059985
2515:
1947:Pinacosaurus mephistocephalus
1479:Struthiosaurus transylvanicus
1455:Struthiosaurus languedocensis
593:
530:
486:Pinacosaurus mephistocephalus
358:Pinacosaurus mephistocephalus
333:. The type and only species,
2925:10.1371/journal.pone.0108804
2700:10.1016/j.palaeo.2016.02.033
436:State Museum of Pennsylvania
418:, a proximal portion of the
125:Restoration as a nodosaurid
7:
4459:Campanian genus extinctions
2487:
10:
4480:
4449:Paleontology in New Mexico
2739:10.1038/s41598-018-21924-7
2725:(1): Article number 3711.
291:Burns & Sullivan, 2011
16:Extinct genus of dinosaurs
4313:
4274:
4254:
4143:
4042:
4033:
3998:
3938:
3929:
3873:
3825:
3745:
3736:
3677:
3558:
3549:
3538:
3499:
3412:
3408:
3393:
3374:
3346:
3328:
3319:
3301:
3288:
3233:
3148:10.1080/02724630903413032
2800:10.1007/s13358-018-0153-1
2238:
2221:
2214:
2197:
2171:
2154:
2147:
2140:
2114:
2097:
2090:
2083:
2066:
2059:
2024:
2007:
2000:
1983:
1976:
1969:
1943:
1926:
1919:
1900:
1883:
1876:
1859:
1852:
1826:
1809:
1802:
1795:
1769:
1752:
1745:
1728:
1711:
1704:
1678:
1661:
1654:
1647:
1496:Struthiosaurus austriacus
1492:
1475:
1468:
1451:
1434:
1417:
1410:
1384:
1367:
1360:
1353:
1306:
1289:
1272:
1255:
1238:
1231:
1214:
1207:
1190:
1183:
1173:
1160:
1153:
1136:
1129:
1112:
1105:
1079:
1062:
1055:
1038:
1021:
1014:
997:
980:
973:
956:
949:
920:
903:
896:
886:
869:
862:
845:
838:
812:
795:
778:
771:
764:
712:in a polytomy containing
690:(2016) recovered it in a
278:
271:
131:Scientific classification
129:
124:
115:
23:
410:, consists of a partial
2581:Akainacephalus johnsoni
2401:that was abundant with
2395:Western Interior Seaway
3120:Bistahieversor sealeyi
3039:Texascephale langstoni
3037:and a reassessment of
2363:
386:
321:that lived during the
248:Burns & Sullivan,
4406:Paleobiology Database
3173:Navajodactylus boerei
2354:
1930:Pinacosaurus grangeri
1310:Edmontonia rugosidens
629:phylogenetic analysis
505:Distinguishing traits
482:Pinacosaurus grangeri
384:
370:Pinacosaurus grangeri
347:by Michael Burns and
4464:Ornithischian genera
3010:10.58799/NMG-v4n3.42
1293:Edmontonia longiceps
377:Discovery and naming
341:postcranial skeleton
3140:2010JVPal..30....1C
2916:2014PLoSO...9j8804A
2849:10.7717/peerj.12362
2791:2018SwJP..137...83R
2731:2018NatSR...8.3711Z
2692:2016PPP...449..289A
2649:2016JSPal..14..385A
2423:coexisted with the
2415:Fruitland Formation
635:as sister taxon to
451:Ah-shi-sle-pah Wash
336:Ahshislepelta minor
285:Ahshislepelta minor
3084:10.7717/peerj.9251
2998:New Mexico Geology
2719:Scientific Reports
2600:10.7717/peerj.5016
2425:pachycephalosaurid
2364:
708:(2018) positioned
567:infraspinous fossa
449:, is derived from
445:The generic name,
400:Robert M. Sullivan
387:
349:Robert M. Sullivan
331:Kirtland Formation
4421:
4420:
4393:Open Tree of Life
4307:Taxon identifiers
4298:
4297:
4270:
4269:
4252:
4251:
4248:
4247:
4244:
4243:
4240:
4239:
4189:Nodocephalosaurus
3925:
3924:
3921:
3920:
3534:
3533:
3453:Serendipaceratops
3389:
3388:
3385:
3384:
3370:
3369:
3366:
3365:
2397:and represents a
2343:
2342:
2334:
2333:
2325:
2324:
2316:
2315:
2307:
2306:
2298:
2297:
2289:
2288:
2280:
2279:
2271:
2270:
2262:
2261:
2253:
2252:
2186:
2185:
2129:
2128:
2118:Nodocephalosaurus
2048:
2047:
2039:
2038:
1958:
1957:
1841:
1840:
1784:
1783:
1693:
1692:
1633:
1632:
1624:
1623:
1615:
1614:
1606:
1605:
1597:
1596:
1588:
1587:
1579:
1578:
1570:
1569:
1561:
1560:
1552:
1551:
1543:
1542:
1534:
1533:
1525:
1524:
1516:
1515:
1507:
1506:
1399:
1398:
1339:
1338:
1330:
1329:
1321:
1320:
1094:
1093:
935:
934:
827:
826:
702:Paw Paw Formation
361:but smaller than
296:
295:
252:
4471:
4414:
4413:
4401:
4400:
4388:
4387:
4375:
4374:
4362:
4361:
4349:
4348:
4347:
4334:
4333:
4332:
4302:
4301:
4288:
4287:
4265:
4260:
4040:
4039:
3936:
3935:
3743:
3742:
3654:Propanoplosaurus
3601:Glyptodontopelta
3556:
3555:
3547:
3546:
3439:Priodontognathus
3410:
3409:
3406:
3405:
3395:
3394:
3326:
3325:
3299:
3298:
3290:
3289:
3283:
3282:
3235:
3234:
3220:
3213:
3206:
3197:
3196:
3190:
3189:
3179:
3166:
3160:
3159:
3113:
3107:
3106:
3096:
3086:
3062:
3056:
3055:
3045:
3028:
3022:
3021:
2995:
2986:
2975:
2974:
2972:
2970:
2954:
2948:
2947:
2937:
2927:
2889:
2880:
2879:
2861:
2851:
2827:
2821:
2820:
2802:
2770:
2761:
2760:
2750:
2710:
2704:
2703:
2675:
2669:
2668:
2632:
2623:
2622:
2612:
2602:
2574:
2568:
2567:
2551:
2504:
2502:Dinosaurs portal
2499:
2498:
2497:
2411:flowering plants
2347:Paleoenvironment
2217:
2216:
2150:
2149:
2143:
2142:
2093:
2092:
2086:
2085:
2062:
2061:
2003:
2002:
1979:
1978:
1972:
1971:
1922:
1921:
1879:
1878:
1855:
1854:
1805:
1804:
1798:
1797:
1748:
1747:
1707:
1706:
1657:
1656:
1650:
1649:
1640:
1639:
1471:
1470:
1413:
1412:
1363:
1362:
1356:
1355:
1234:
1233:
1210:
1209:
1186:
1185:
1176:
1175:
1156:
1155:
1132:
1131:
1108:
1107:
1058:
1057:
1017:
1016:
976:
975:
952:
951:
899:
898:
889:
888:
865:
864:
841:
840:
774:
773:
767:
766:
757:
756:
428:dorsal vertebrae
287:
283:
247:
240:
227:
211:
195:
139:
138:
120:
110:
40:
29:Temporal range:
21:
20:
4479:
4478:
4474:
4473:
4472:
4470:
4469:
4468:
4424:
4423:
4422:
4417:
4409:
4404:
4396:
4391:
4383:
4378:
4370:
4365:
4357:
4352:
4343:
4342:
4337:
4328:
4327:
4322:
4309:
4299:
4294:
4266:
4236:
4139:
4119:Tianzhenosaurus
4083:Minotaurasaurus
4029:
4023:Zhongyuansaurus
3994:
3989:Zhejiangosaurus
3982:Liaoningosaurus
3917:
3875:Struthiosaurini
3869:
3821:
3816:Tatankacephalus
3732:
3688:Gargoyleosaurus
3673:
3585:Gargoyleosaurus
3543:
3530:
3501:Parankylosauria
3495:
3461:Sinankylosaurus
3402:
3381:
3362:
3342:
3315:
3309:Avemetatarsalia
3294:Avemetatarsalia
3284:
3240:
3239:
3229:
3224:
3194:
3193:
3177:
3167:
3163:
3114:
3110:
3063:
3059:
3043:
3029:
3025:
2993:
2987:
2978:
2968:
2966:
2965:. pp. 7–30
2955:
2951:
2890:
2883:
2828:
2824:
2771:
2764:
2711:
2707:
2676:
2672:
2633:
2626:
2575:
2571:
2552:
2523:
2518:
2500:
2495:
2493:
2490:
2349:
2344:
2335:
2326:
2317:
2308:
2299:
2290:
2281:
2272:
2263:
2254:
2187:
2130:
2049:
2040:
1959:
1905:Zhejiangosaurus
1842:
1785:
1732:Liaoningosaurus
1694:
1634:
1625:
1616:
1607:
1598:
1589:
1580:
1571:
1562:
1553:
1544:
1535:
1526:
1517:
1508:
1400:
1350:Struthiosaurini
1340:
1331:
1322:
1116:Tatankacephalus
1095:
936:
873:Gargoyleosaurus
828:
715:Tatankacephalus
617:
596:
533:
507:
502:
465:ankylosaurids.
416:scapulocoracoid
379:
355:is larger than
323:Late Cretaceous
292:
289:
281:
280:
267:
246:
238:
225:
209:
193:
133:
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
31:Late Cretaceous
27:
17:
12:
11:
5:
4477:
4467:
4466:
4461:
4456:
4451:
4446:
4441:
4436:
4419:
4418:
4416:
4415:
4402:
4389:
4376:
4363:
4350:
4335:
4319:
4317:
4311:
4310:
4296:
4295:
4293:
4292:
4282:
4275:
4272:
4271:
4268:
4267:
4255:
4253:
4250:
4249:
4246:
4245:
4242:
4241:
4238:
4237:
4235:
4234:
4227:
4220:
4213:
4206:
4199:
4192:
4185:
4182:Euoplocephalus
4178:
4171:
4168:Anodontosaurus
4164:
4157:
4154:Akainacephalus
4149:
4147:
4141:
4140:
4138:
4137:
4130:
4123:
4115:
4108:
4100:
4093:
4086:
4079:
4072:
4065:
4062:Crichtonsaurus
4058:
4051:
4043:
4037:
4035:Ankylosaurinae
4031:
4030:
4028:
4027:
4019:
4012:
4004:
4002:
3996:
3995:
3993:
3992:
3985:
3978:
3970:
3963:
3956:
3948:
3939:
3933:
3931:Ankylosauridae
3927:
3926:
3923:
3922:
3919:
3918:
3916:
3915:
3912:Struthiosaurus
3908:
3901:
3894:
3887:
3879:
3877:
3871:
3870:
3868:
3867:
3860:
3853:
3846:
3839:
3831:
3829:
3827:Panoplosaurini
3823:
3822:
3820:
3819:
3812:
3805:
3798:
3790:
3783:
3776:
3773:Niobrarasaurus
3769:
3762:
3754:
3746:
3740:
3734:
3733:
3731:
3730:
3722:
3715:
3707:
3700:
3692:
3683:
3681:
3675:
3674:
3672:
3671:
3664:
3657:
3650:
3642:
3638:Polacanthoides
3634:
3626:
3619:
3611:
3608:Horshamosaurus
3604:
3597:
3589:
3581:
3578:Dongyangopelta
3574:
3567:
3559:
3553:
3544:
3541:Euankylosauria
3539:
3536:
3535:
3532:
3531:
3529:
3528:
3521:
3517:Kunbarrasaurus
3513:
3505:
3503:
3497:
3496:
3494:
3493:
3486:
3479:
3471:
3464:
3457:
3449:
3442:
3435:
3428:
3421:
3413:
3403:
3398:
3391:
3390:
3387:
3386:
3383:
3382:
3375:
3372:
3371:
3368:
3367:
3364:
3363:
3361:
3360:
3359:
3358:
3352:
3350:
3344:
3343:
3341:
3340:
3339:
3338:
3329:
3323:
3317:
3316:
3314:
3313:
3312:
3311:
3302:
3296:
3286:
3285:
3281:
3280:
3271:
3262:
3253:
3247:
3231:
3230:
3223:
3222:
3215:
3208:
3200:
3192:
3191:
3161:
3108:
3057:
3023:
2976:
2949:
2881:
2822:
2762:
2705:
2670:
2624:
2569:
2520:
2519:
2517:
2514:
2513:
2512:
2506:
2505:
2489:
2486:
2481:Navajodactylus
2469:Bistahieversor
2465:tyrannosauroid
2441:Navajoceratops
2348:
2345:
2341:
2340:
2337:
2336:
2332:
2331:
2328:
2327:
2323:
2322:
2319:
2318:
2314:
2313:
2310:
2309:
2305:
2304:
2301:
2300:
2296:
2295:
2292:
2291:
2287:
2286:
2283:
2282:
2278:
2277:
2274:
2273:
2269:
2268:
2265:
2264:
2260:
2259:
2256:
2255:
2251:
2250:
2247:
2246:
2237:
2234:
2233:
2230:
2229:
2220:
2215:
2213:
2210:
2209:
2206:
2205:
2201:Euoplocephalus
2196:
2193:
2192:
2189:
2188:
2184:
2183:
2180:
2179:
2175:Anodontosaurus
2170:
2167:
2166:
2163:
2162:
2153:
2148:
2146:
2141:
2139:
2136:
2135:
2132:
2131:
2127:
2126:
2123:
2122:
2113:
2110:
2109:
2106:
2105:
2096:
2091:
2089:
2084:
2082:
2079:
2078:
2075:
2074:
2065:
2060:
2058:
2055:
2054:
2051:
2050:
2046:
2045:
2042:
2041:
2037:
2036:
2033:
2032:
2023:
2020:
2019:
2016:
2015:
2006:
2001:
1999:
1996:
1995:
1992:
1991:
1982:
1977:
1975:
1970:
1968:
1965:
1964:
1961:
1960:
1956:
1955:
1952:
1951:
1942:
1939:
1938:
1935:
1934:
1925:
1920:
1918:
1915:
1914:
1911:
1910:
1899:
1896:
1895:
1892:
1891:
1882:
1877:
1875:
1872:
1871:
1868:
1867:
1858:
1853:
1851:
1848:
1847:
1844:
1843:
1839:
1838:
1835:
1834:
1825:
1822:
1821:
1818:
1817:
1808:
1803:
1801:
1796:
1794:
1791:
1790:
1787:
1786:
1782:
1781:
1778:
1777:
1768:
1765:
1764:
1761:
1760:
1751:
1746:
1744:
1741:
1740:
1737:
1736:
1727:
1724:
1723:
1720:
1719:
1710:
1705:
1703:
1700:
1699:
1696:
1695:
1691:
1690:
1687:
1686:
1677:
1674:
1673:
1670:
1669:
1660:
1655:
1653:
1648:
1646:
1644:Ankylosauridae
1638:
1631:
1630:
1627:
1626:
1622:
1621:
1618:
1617:
1613:
1612:
1609:
1608:
1604:
1603:
1600:
1599:
1595:
1594:
1591:
1590:
1586:
1585:
1582:
1581:
1577:
1576:
1573:
1572:
1568:
1567:
1564:
1563:
1559:
1558:
1555:
1554:
1550:
1549:
1546:
1545:
1541:
1540:
1537:
1536:
1532:
1531:
1528:
1527:
1523:
1522:
1519:
1518:
1514:
1513:
1510:
1509:
1505:
1504:
1501:
1500:
1491:
1488:
1487:
1484:
1483:
1474:
1469:
1467:
1464:
1463:
1460:
1459:
1450:
1447:
1446:
1443:
1442:
1433:
1430:
1429:
1426:
1425:
1416:
1411:
1409:
1406:
1405:
1402:
1401:
1397:
1396:
1393:
1392:
1383:
1380:
1379:
1376:
1375:
1366:
1361:
1359:
1354:
1352:
1346:
1345:
1342:
1341:
1337:
1336:
1333:
1332:
1328:
1327:
1324:
1323:
1319:
1318:
1315:
1314:
1305:
1302:
1301:
1298:
1297:
1288:
1285:
1284:
1281:
1280:
1271:
1268:
1267:
1264:
1263:
1254:
1251:
1250:
1247:
1246:
1237:
1232:
1230:
1227:
1226:
1223:
1222:
1213:
1208:
1206:
1203:
1202:
1199:
1198:
1189:
1184:
1182:
1180:Panoplosaurini
1174:
1172:
1169:
1168:
1165:
1164:
1159:
1154:
1152:
1149:
1148:
1145:
1144:
1135:
1130:
1128:
1125:
1124:
1121:
1120:
1111:
1106:
1104:
1101:
1100:
1097:
1096:
1092:
1091:
1088:
1087:
1078:
1075:
1074:
1071:
1070:
1066:Niobrarasaurus
1061:
1056:
1054:
1051:
1050:
1047:
1046:
1037:
1034:
1033:
1030:
1029:
1020:
1015:
1013:
1010:
1009:
1006:
1005:
996:
993:
992:
989:
988:
979:
974:
972:
969:
968:
965:
964:
955:
950:
948:
942:
941:
938:
937:
933:
932:
929:
928:
919:
916:
915:
912:
911:
902:
897:
895:
887:
885:
882:
881:
878:
877:
868:
863:
861:
858:
857:
854:
853:
844:
839:
837:
834:
833:
830:
829:
825:
824:
821:
820:
816:Dongyangopelta
811:
808:
807:
804:
803:
794:
791:
790:
787:
786:
777:
772:
770:
765:
763:
755:
704:, while Zheng
697:Niobrarasaurus
668:Anodontosaurus
644:Euoplocephalus
625:Ankylosaurinae
616:
615:Classification
613:
609:Euoplocephalus
595:
592:
549:glenoid cavity
532:
529:
524:Euoplocephalus
506:
503:
501:
498:
463:North American
378:
375:
294:
293:
290:
276:
275:
269:
268:
260:
258:
254:
253:
236:
232:
231:
223:
216:
215:
207:
200:
199:
191:
184:
183:
178:
171:
170:
165:
161:
160:
155:
151:
150:
145:
141:
140:
127:
126:
122:
121:
113:
112:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
28:
15:
9:
6:
4:
3:
2:
4476:
4465:
4462:
4460:
4457:
4455:
4452:
4450:
4447:
4445:
4442:
4440:
4437:
4435:
4432:
4431:
4429:
4412:
4407:
4403:
4399:
4394:
4390:
4386:
4381:
4377:
4373:
4368:
4364:
4360:
4355:
4351:
4346:
4345:Ahshislepelta
4340:
4336:
4331:
4325:
4321:
4320:
4318:
4316:
4315:Ahshislepelta
4312:
4308:
4303:
4291:
4283:
4281:
4277:
4276:
4273:
4264:
4259:
4233:
4232:
4228:
4226:
4225:
4221:
4219:
4218:
4214:
4212:
4211:
4207:
4205:
4204:
4200:
4198:
4197:
4193:
4191:
4190:
4186:
4184:
4183:
4179:
4177:
4176:
4172:
4170:
4169:
4165:
4163:
4162:
4158:
4156:
4155:
4151:
4150:
4148:
4146:
4145:Ankylosaurini
4142:
4136:
4135:
4131:
4129:
4128:
4124:
4121:
4120:
4116:
4114:
4113:
4109:
4106:
4105:
4101:
4099:
4098:
4094:
4092:
4091:
4087:
4085:
4084:
4080:
4078:
4077:
4073:
4071:
4070:
4066:
4064:
4063:
4059:
4057:
4056:
4055:Crichtonpelta
4052:
4050:
4049:
4048:Bissektipelta
4045:
4044:
4041:
4038:
4036:
4032:
4025:
4024:
4020:
4018:
4017:
4013:
4011:
4010:
4006:
4005:
4003:
4001:
4000:Shamosaurinae
3997:
3991:
3990:
3986:
3984:
3983:
3979:
3976:
3975:
3971:
3969:
3968:
3964:
3962:
3961:
3957:
3954:
3953:
3949:
3946:
3945:
3944:Ahshislepelta
3941:
3940:
3937:
3934:
3932:
3928:
3914:
3913:
3909:
3907:
3906:
3902:
3900:
3899:
3895:
3893:
3892:
3891:Hungarosaurus
3888:
3886:
3885:
3881:
3880:
3878:
3876:
3872:
3866:
3865:
3861:
3859:
3858:
3857:Panoplosaurus
3854:
3852:
3851:
3847:
3845:
3844:
3840:
3838:
3837:
3833:
3832:
3830:
3828:
3824:
3818:
3817:
3813:
3811:
3810:
3806:
3804:
3803:
3799:
3796:
3795:
3791:
3789:
3788:
3784:
3782:
3781:
3777:
3775:
3774:
3770:
3768:
3767:
3763:
3760:
3759:
3758:Ahshislepelta
3755:
3753:
3752:
3748:
3747:
3744:
3741:
3739:
3735:
3728:
3727:
3723:
3721:
3720:
3716:
3713:
3712:
3708:
3706:
3705:
3704:Hoplitosaurus
3701:
3698:
3697:
3693:
3690:
3689:
3685:
3684:
3682:
3680:
3679:Polacanthinae
3676:
3670:
3669:
3665:
3663:
3662:
3661:Rhadinosaurus
3658:
3656:
3655:
3651:
3648:
3647:
3643:
3640:
3639:
3635:
3632:
3631:
3627:
3625:
3624:
3620:
3617:
3616:
3612:
3610:
3609:
3605:
3603:
3602:
3598:
3595:
3594:
3590:
3587:
3586:
3582:
3580:
3579:
3575:
3573:
3572:
3568:
3566:
3565:
3564:Acanthopholis
3561:
3560:
3557:
3554:
3552:
3548:
3545:
3542:
3537:
3527:
3526:
3522:
3519:
3518:
3514:
3512:
3511:
3510:Antarctopelta
3507:
3506:
3504:
3502:
3498:
3492:
3491:
3487:
3485:
3484:
3483:Tianchisaurus
3480:
3477:
3476:
3475:Stegosaurides
3472:
3470:
3469:
3465:
3463:
3462:
3458:
3455:
3454:
3450:
3448:
3447:
3443:
3441:
3440:
3436:
3434:
3433:
3429:
3427:
3426:
3422:
3420:
3419:
3415:
3414:
3411:
3407:
3404:
3401:
3396:
3392:
3380:
3379:
3373:
3356:
3355:
3354:
3353:
3351:
3349:
3345:
3337:
3333:
3332:
3331:
3330:
3327:
3324:
3322:
3318:
3310:
3306:
3305:
3304:
3303:
3300:
3297:
3295:
3291:
3287:
3279:
3275:
3272:
3270:
3266:
3263:
3261:
3257:
3254:
3252:
3248:
3246:
3242:
3241:
3236:
3232:
3228:
3221:
3216:
3214:
3209:
3207:
3202:
3201:
3198:
3187:
3183:
3176:
3174:
3165:
3157:
3153:
3149:
3145:
3141:
3137:
3133:
3129:
3128:
3123:
3121:
3112:
3104:
3100:
3095:
3090:
3085:
3080:
3076:
3072:
3068:
3061:
3053:
3049:
3042:
3040:
3036:
3027:
3019:
3015:
3011:
3007:
3003:
2999:
2992:
2985:
2983:
2981:
2964:
2960:
2953:
2945:
2941:
2936:
2931:
2926:
2921:
2917:
2913:
2909:
2905:
2901:
2900:
2895:
2888:
2886:
2877:
2873:
2869:
2865:
2860:
2855:
2850:
2845:
2841:
2837:
2833:
2826:
2818:
2814:
2810:
2806:
2801:
2796:
2792:
2788:
2784:
2780:
2776:
2769:
2767:
2758:
2754:
2749:
2744:
2740:
2736:
2732:
2728:
2724:
2720:
2716:
2709:
2701:
2697:
2693:
2689:
2685:
2681:
2674:
2666:
2662:
2658:
2654:
2650:
2646:
2642:
2638:
2631:
2629:
2620:
2616:
2611:
2606:
2601:
2596:
2592:
2588:
2584:
2582:
2573:
2565:
2561:
2557:
2550:
2548:
2546:
2544:
2542:
2540:
2538:
2536:
2534:
2532:
2530:
2528:
2526:
2521:
2511:
2508:
2507:
2503:
2492:
2485:
2483:
2482:
2478:
2475:
2471:
2470:
2466:
2463:
2462:eutyrannosaur
2459:
2458:
2454:ankylosaurid
2453:
2452:ankylosaurine
2449:
2448:
2443:
2442:
2438:
2435:
2434:chasmosaurine
2431:
2430:
2426:
2422:
2421:Ahshislepelta
2418:
2416:
2412:
2408:
2404:
2400:
2396:
2392:
2388:
2384:
2380:
2376:
2372:
2368:
2367:Ahshislepelta
2361:
2360:Ahshislepelta
2357:
2353:
2339:
2338:
2330:
2329:
2321:
2320:
2312:
2311:
2303:
2302:
2294:
2293:
2285:
2284:
2276:
2275:
2267:
2266:
2258:
2257:
2249:
2248:
2245:
2244:
2243:
2236:
2235:
2232:
2231:
2228:
2227:
2226:
2219:
2218:
2212:
2211:
2208:
2207:
2204:
2203:
2202:
2195:
2194:
2191:
2190:
2182:
2181:
2178:
2177:
2176:
2169:
2168:
2165:
2164:
2161:
2160:
2159:
2152:
2151:
2145:
2144:
2138:
2137:
2134:
2133:
2125:
2124:
2121:
2120:
2119:
2112:
2111:
2108:
2107:
2104:
2103:
2102:
2095:
2094:
2088:
2087:
2081:
2080:
2077:
2076:
2073:
2072:
2071:
2064:
2063:
2057:
2056:
2053:
2052:
2044:
2043:
2035:
2034:
2031:
2030:
2029:
2022:
2021:
2018:
2017:
2014:
2013:
2012:
2005:
2004:
1998:
1997:
1994:
1993:
1990:
1989:
1988:
1981:
1980:
1974:
1973:
1967:
1966:
1963:
1962:
1954:
1953:
1950:
1949:
1948:
1941:
1940:
1937:
1936:
1933:
1932:
1931:
1924:
1923:
1917:
1916:
1913:
1912:
1909:
1908:
1906:
1898:
1897:
1894:
1893:
1890:
1889:
1888:
1881:
1880:
1874:
1873:
1870:
1869:
1866:
1865:
1864:
1863:Crichtonpelta
1857:
1856:
1850:
1849:
1846:
1845:
1837:
1836:
1833:
1832:
1831:
1824:
1823:
1820:
1819:
1816:
1815:
1814:
1807:
1806:
1800:
1799:
1793:
1792:
1789:
1788:
1780:
1779:
1776:
1775:
1774:
1767:
1766:
1763:
1762:
1759:
1758:
1757:
1750:
1749:
1743:
1742:
1739:
1738:
1735:
1734:
1733:
1726:
1725:
1722:
1721:
1718:
1717:
1716:
1709:
1708:
1702:
1701:
1698:
1697:
1689:
1688:
1685:
1684:
1683:
1682:Ahshislepelta
1676:
1675:
1672:
1671:
1668:
1667:
1666:
1659:
1658:
1652:
1651:
1645:
1642:
1641:
1637:
1629:
1628:
1620:
1619:
1611:
1610:
1602:
1601:
1593:
1592:
1584:
1583:
1575:
1574:
1566:
1565:
1557:
1556:
1548:
1547:
1539:
1538:
1530:
1529:
1521:
1520:
1512:
1511:
1503:
1502:
1499:
1498:
1497:
1490:
1489:
1486:
1485:
1482:
1481:
1480:
1473:
1472:
1466:
1465:
1462:
1461:
1458:
1457:
1456:
1449:
1448:
1445:
1444:
1441:
1440:
1439:
1432:
1431:
1428:
1427:
1424:
1423:
1422:
1415:
1414:
1408:
1407:
1404:
1403:
1395:
1394:
1391:
1390:
1389:
1382:
1381:
1378:
1377:
1374:
1373:
1372:
1371:Hungarosaurus
1365:
1364:
1358:
1357:
1351:
1348:
1347:
1344:
1343:
1335:
1334:
1326:
1325:
1317:
1316:
1313:
1312:
1311:
1304:
1303:
1300:
1299:
1296:
1295:
1294:
1287:
1286:
1283:
1282:
1279:
1278:
1277:
1270:
1269:
1266:
1265:
1262:
1261:
1260:
1253:
1252:
1249:
1248:
1245:
1244:
1243:
1236:
1235:
1229:
1228:
1225:
1224:
1221:
1220:
1219:
1218:Panoplosaurus
1212:
1211:
1205:
1204:
1201:
1200:
1197:
1196:
1195:
1188:
1187:
1181:
1178:
1177:
1171:
1170:
1167:
1166:
1163:
1158:
1157:
1151:
1150:
1147:
1146:
1143:
1142:
1141:
1134:
1133:
1127:
1126:
1123:
1122:
1119:
1118:
1117:
1110:
1109:
1103:
1102:
1099:
1098:
1090:
1089:
1086:
1085:
1084:
1083:Ahshislepelta
1077:
1076:
1073:
1072:
1069:
1068:
1067:
1060:
1059:
1053:
1052:
1049:
1048:
1045:
1044:
1043:
1036:
1035:
1032:
1031:
1028:
1027:
1026:
1019:
1018:
1012:
1011:
1008:
1007:
1004:
1003:
1002:
995:
994:
991:
990:
987:
986:
985:
978:
977:
971:
970:
967:
966:
963:
962:
961:
954:
953:
947:
944:
943:
940:
939:
931:
930:
927:
926:
925:
918:
917:
914:
913:
910:
909:
908:
907:Hoplitosaurus
901:
900:
894:
893:Polacanthinae
891:
890:
884:
883:
880:
879:
876:
875:
874:
867:
866:
860:
859:
856:
855:
852:
851:
850:
843:
842:
836:
835:
832:
831:
823:
822:
819:
818:
817:
810:
809:
806:
805:
802:
801:
800:
793:
792:
789:
788:
785:
784:
783:
776:
775:
769:
768:
762:
759:
758:
754:
752:
748:
743:
741:
740:Niobrarasurus
737:
733:
732:
727:
726:Niobrarasurus
723:
722:
717:
716:
711:
710:Ahshislepelta
707:
703:
699:
698:
693:
689:
684:
682:
681:
676:
675:
670:
669:
664:
663:
658:
657:
652:
651:
646:
645:
640:
639:
634:
633:Ahshislepelta
630:
626:
622:
621:Ahshislepelta
612:
610:
605:
601:
591:
588:
583:
581:
577:
572:
568:
563:
558:
554:
550:
546:
542:
538:
528:
526:
525:
520:
519:Ahshislepelta
516:
512:
511:Ahshislepelta
497:
495:
491:
487:
483:
479:
475:
474:neural arches
471:
466:
464:
460:
456:
455:Ancient Greek
452:
448:
447:Ahshislepelta
443:
441:
437:
433:
429:
425:
421:
417:
413:
409:
405:
401:
396:
392:
383:
374:
372:
371:
366:
365:
360:
359:
354:
353:Ahshislepelta
350:
346:
342:
338:
337:
332:
328:
324:
320:
316:
313:
309:
306:
302:
301:
300:Ahshislepelta
288:
286:
277:
274:
273:Binomial name
270:
266:
265:
264:A. minor
259:
256:
255:
251:
245:
244:
243:Ahshislepelta
237:
234:
233:
230:
224:
221:
218:
217:
214:
208:
205:
202:
201:
198:
192:
189:
186:
185:
182:
179:
176:
173:
172:
169:
166:
163:
162:
159:
156:
153:
152:
149:
146:
143:
142:
137:
132:
128:
123:
119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
26:
25:Ahshislepelta
22:
19:
4314:
4229:
4222:
4215:
4208:
4201:
4194:
4187:
4180:
4175:Dyoplosaurus
4173:
4166:
4161:Ankylosaurus
4159:
4152:
4132:
4125:
4117:
4110:
4102:
4095:
4090:Pinacosaurus
4088:
4081:
4074:
4067:
4060:
4053:
4046:
4021:
4014:
4007:
3987:
3980:
3974:Hylaeosaurus
3972:
3965:
3958:
3950:
3943:
3942:
3910:
3903:
3898:Pawpawsaurus
3896:
3889:
3882:
3862:
3855:
3848:
3843:Denversaurus
3841:
3834:
3814:
3807:
3800:
3794:Peloroplites
3792:
3785:
3778:
3771:
3766:Borealopelta
3764:
3757:
3756:
3751:Acantholipan
3749:
3738:Nodosaurinae
3724:
3717:
3711:Peloroplites
3709:
3702:
3694:
3686:
3666:
3659:
3652:
3644:
3636:
3630:Mymoorapelta
3628:
3621:
3615:Hylaeosaurus
3613:
3606:
3599:
3591:
3583:
3576:
3571:Anoplosaurus
3569:
3562:
3551:Nodosauridae
3523:
3515:
3508:
3488:
3481:
3473:
3466:
3459:
3451:
3444:
3437:
3430:
3423:
3418:Cryptosaurus
3416:
3400:Ankylosauria
3378:Ankylosauria
3376:
3348:Ankylosauria
3336:Ornithischia
3321:Ornithischia
3273:
3269:Ornithischia
3264:
3255:
3227:Ankylosauria
3185:
3181:
3172:
3164:
3131:
3125:
3119:
3111:
3074:
3070:
3060:
3051:
3047:
3038:
3034:
3026:
3004:(3): 42–45.
3001:
2997:
2967:. Retrieved
2962:
2952:
2903:
2897:
2839:
2835:
2825:
2785:(1): 83–93.
2782:
2778:
2722:
2718:
2708:
2683:
2679:
2673:
2640:
2636:
2590:
2586:
2580:
2572:
2563:
2559:
2479:
2467:
2455:
2447:Terminocavus
2445:
2439:
2427:
2420:
2419:
2366:
2365:
2359:
2355:
2240:
2239:
2223:
2222:
2199:
2198:
2173:
2172:
2158:Ankylosaurus
2156:
2155:
2116:
2115:
2099:
2098:
2070:Dyoplosaurus
2068:
2067:
2026:
2025:
2009:
2008:
1985:
1984:
1945:
1944:
1928:
1927:
1902:
1901:
1885:
1884:
1861:
1860:
1828:
1827:
1811:
1810:
1771:
1770:
1754:
1753:
1730:
1729:
1713:
1712:
1681:
1680:
1679:
1663:
1662:
1635:
1494:
1493:
1477:
1476:
1453:
1452:
1436:
1435:
1421:Pawpawsaurus
1419:
1418:
1386:
1385:
1369:
1368:
1308:
1307:
1291:
1290:
1276:Denversaurus
1274:
1273:
1257:
1256:
1240:
1239:
1216:
1215:
1192:
1191:
1161:
1138:
1137:
1114:
1113:
1082:
1081:
1080:
1064:
1063:
1040:
1039:
1025:Acantholipan
1023:
1022:
999:
998:
982:
981:
960:Peloroplites
958:
957:
946:Nodosaurinae
922:
921:
905:
904:
871:
870:
847:
846:
814:
813:
799:Mymoorapelta
797:
796:
780:
779:
761:Nodosauridae
750:
746:
744:
739:
735:
729:
725:
719:
713:
709:
705:
695:
687:
685:
680:Dyoplosaurus
678:
674:Ankylosaurus
672:
666:
660:
654:
648:
642:
636:
632:
620:
618:
608:
603:
597:
584:
534:
522:
518:
510:
508:
493:
489:
485:
481:
478:ankylosaurid
467:
458:
446:
444:
407:
388:
368:
362:
356:
352:
335:
334:
305:monospecific
299:
298:
297:
284:
279:
263:
262:
242:
241:
229:Ankylosauria
219:
203:
197:Ornithischia
187:
174:
24:
18:
4439:Ankylosaurs
4339:Wikispecies
4210:Scolosaurus
4127:Tsagantegia
4076:Jinyunpelta
4016:Shamosaurus
3967:Chuanqilong
3809:Silvisaurus
3787:Patagopelta
3719:Polacanthus
3668:Sauroplites
3468:Spicomellus
3446:Sarcolestes
3278:Thyreophora
3134:(1): 1–16.
2910:: e108804.
2686:: 289–299.
2643:(5): 1–60.
2437:ceratopsids
2225:Scolosaurus
1887:Tsagantegia
1830:Shamosaurus
1773:Chuanqilong
1242:Patagopelta
1140:Silvisaurus
924:Polacanthus
782:Sauroplites
721:Silvisaurus
656:Scolosaurus
553:nodosaurids
500:Description
490:P. grangeri
422:, numerous
408:SMP VP-1930
213:Thyreophora
4428:Categories
4278:See also:
4203:Platypelta
4196:Oohkotokia
4134:Zaraapelta
4009:Gobisaurus
3960:Cedarpelta
3952:Aletopelta
3905:Stegopelta
3850:Edmontonia
3836:Animantarx
3802:Sauropelta
3780:Nodosaurus
3646:Priconodon
3490:Vectipelta
3425:Dracopelta
3357:see below↓
3260:Dinosauria
3188:: 393–404.
3054:: 202–215.
3035:Stegoceras
2969:19 January
2842:: e12362.
2566:: 169–178.
2516:References
2474:azhdarchid
2472:, and the
2429:Stegoceras
2399:floodplain
2356:Stegoceras
2028:Zaraapelta
1813:Gobisaurus
1756:Cedarpelta
1715:Aletopelta
1438:Stegopelta
1194:Animantarx
1042:Nodosaurus
1001:Sauropelta
731:Nodosaurus
650:Oohkotokia
594:Osteoderms
562:taphonomic
531:Postcrania
440:Harrisburg
432:osteoderms
414:, partial
406:specimen,
395:ankylosaur
319:New Mexico
312:ankylosaur
181:Dinosauria
36:74.5
4217:Talarurus
4097:Saichania
3884:Europelta
3864:Texasetes
3726:Taohelong
3623:Invictarx
3525:Stegouros
3243:Kingdom:
3077:: e9251.
3018:259446701
2876:245111393
2817:134924657
2809:1664-2376
2665:214625754
2593:: e5016.
2477:pterosaur
2379:siltstone
2375:sandstone
2371:Campanian
2101:Talarurus
1987:Saichania
1388:Europelta
1259:Texasetes
984:Taohelong
753:(2021).
600:forelimbs
587:vertebrae
585:Numerous
580:diaphysis
494:Talarurus
364:Talarurus
327:Campanian
257:Species:
154:Kingdom:
148:Eukaryota
4359:42418600
4330:Q1705974
4324:Wikidata
4290:Category
4280:Timeline
4224:Ziapelta
3696:Gastonia
3593:Gastonia
3251:Chordata
3249:Phylum:
3245:Animalia
3156:54029279
3103:32547873
2944:25250819
2899:PLOS ONE
2868:34966571
2757:29487376
2619:30065856
2488:See also
2457:Ziapelta
2407:conifers
2383:mudstone
2242:Ziapelta
1665:Gastonia
1162:CPC 273
849:Gastonia
692:polytomy
662:Ziapelta
638:Gastonia
557:acromion
541:coracoid
470:subadult
424:cervical
404:holotype
391:skeleton
315:dinosaur
168:Chordata
164:Phylum:
158:Animalia
144:Domain:
4398:6150239
4385:1479641
4372:8698972
4112:Tarchia
4104:Shanxia
3136:Bibcode
3094:7278894
2935:4177562
2912:Bibcode
2859:8667728
2787:Bibcode
2748:5829254
2727:Bibcode
2688:Bibcode
2645:Bibcode
2610:6063217
2011:Tarchia
604:in situ
571:humerus
545:muscles
515:scapula
426:and/or
282:†
261:†
235:Genus:
105:↓
4411:295547
3154:
3101:
3091:
3016:
2942:
2932:
2874:
2866:
2856:
2815:
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