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Aspergillus oryzae

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are pectinase and peptidase. Pectinase drives starch hydrolysis by breaking down the pectin in the cell walls of plant materials like soybeans, in the case of miso and soy sauce production, while peptidases like leucine aminopeptidase cleave amino acids from proteins and polypeptides like glutamic
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Merz, Michael; Eisele, Thomas; Berends, Pieter; Appel, Daniel; Rabe, Swen; Blank, Imre; Stressler, Timo; Fischer, Lutz (17 June 2015). "Flavourzyme, an Enzyme Preparation with Industrial Relevance: Automated Nine-Step Purification and Partial Characterization of Eight Enzymes".
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Lin, Weimin; Song, Jiajia; Hu, Wenfeng; Miao, Jianyin; Gao, Xiangyang (2016). "Relationship Between Extracellular Cellulase, Pectinase and Xylanase Activity of Isolated Aspergillus oryzae Strains Grown on Koji and the Umami-Tasting Amino Acid Content of Soy Sauce".
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are responsible for the function of proteins and cellular processes such as hydrolase, transporters, and metabolism. The extensive array of secretory hydrolase and transporters allows the mold to break down or secrete various compounds effectively. Typically, when
554:. Its strength is that it gives rise to a rich, fruity, refreshing taste, so despite the difficulties and great skill required, it is still used by some manufacturers. It is popular amongst young people who previously had no interest in typically strong potato 1020: 654:
a rich aroma with a slightly sweet, mellow taste. Its spores disperse easily, covering production facilities and workers' clothes in a layer of black. Such issues led to it falling out of favour, but due to the development of new
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Kiyota, Takuro; Hamada, Ryoko; Sakamoto, Kazutoshi; Iwashita, Kazuhiro; Yamada, Osamu; Mikami, Shigeaki (May 2011). "Aflatoxin non-productivity of Aspergillus oryzae caused by loss of function in the aflJ gene product".
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Wang, D.; Zheng, Z. Y.; Feng, J.; Zhan, X. B.; Zhang, L. M.; Wu, J. R.; Lin, C. C. (2013-07-04). "A high salt tolerant neutral protease from Aspergillus Oryzae: Purification, characterization and kinetic properties".
636:. In 1901, Tamaki Inui, lecturer at University of Tokyo succeeded in the first isolating and culturing. In 1910, Genichirō Kawachi succeeded for the first time in culturing var. kawachi, a variety of subtaxa of 1473:
Zhao, Guozhong; Yao, Yunping; Wang, Chunling; Hou, Lihua; Cao, Xiaohong (2013-06-17). "Comparative genomic analysis of Aspergillus oryzae strains 3.042 and RIB40 for soy sauce fermentation".
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exposed to high concentrations of foods like rice, soybean, wheat, etc. during fermentation, its growth may be negatively affected. However, over time this may potentially allow the
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secretes a number of salt-tolerant alkaline proteases which makes it particularly stable in the high-sodium conditions required for the production of miso and soy sauce. The strain
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have cell walls that are difficult to break down which makes gene insertion/editing complicated. However, scientists have recently started utilizing CRISPR/Cas9 in
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Atanasov, Atanas G.; Zotchev, Sergey B.; Dirsch, Verena M.; Supuran, Claudiu T.; et al. (The International Natural Product Sciences Taskforce) (2021-01-28).
853: 943:) in China in 300 BCE. Its development is a milestone in Chinese food technology, for it provides the conceptual framework for three major fermented soy foods: 2514: 1300: 772:
mold has not been found to produce those toxins. Furthermore, no carcinogenic substances have been discovered in the mold. A study has shown that even when
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Wang, H.; Liu, L.; Guo, Y. -X.; Dong, Y. -S.; Zhang, D. -J.; Xiu, Z. -L. (2007). "Biotransformation of piceid in Polygonum cuspidatum to resveratrol by
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Osamu Yamada; Masayuki Machida; Akira Hosoyama; Masatoshi Goto; Toru Takahashi; Taiki Futagami; Youhei Yamagata; et al. (December 2016).
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can easily sour during fermentation. This makes it difficult to use in warmer regions such as Kyūshū, and gradually black and white
2885: 2753: 2607: 303:. While the two fungi share the same cluster of genes that encode for aflatoxin synthesis, this gene cluster is non-functional in 1280: 640:. This improved the efficiency of shōchū production. It produces plenty of citric acid which helps to prevent the souring of the 2959: 1729: 2143: 1537: 671:
makers because of the depth and quality of the taste it produced. Several popular brands now explicitly state they use black
821:. This increased mutation rates in the genome which was not possible in the past since the mold only reproduced asexually. 1216: 804: 2475: 341:, and a range of other traditional Japanese foods. His proposal was approved at the society's annual meeting in 2006. 1355: 2810: 734: 2470: 2677: 2511: 1306: 1319: 439:
acid, an amino acid that contributes to the characteristic umami flavor of these fermented soybean products.
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Lafayette, California: Soyinfo Center. 660 pp. (1,560 references; 142 photos and illustrations, Free online)
2264:"Advances in Genetic Engineering Technology and Its Application in the Industrial Fungus Aspergillus oryzae" 2572: 1119: 606:
is easy to cultivate and its enzymes promote rapid saccharization; as a result, it is used to produce most
470: 1759: 56: 2044: 2969: 2926: 2877: 2745: 2687: 2599: 2331: 1094: 837: 2165: 319:) in the journal of the Brewing Society of Japan, because of its importance not only for making the 2079:
Barbesgaard, Peder; Heldt-Hansen, HansPeter; Diderichsen, Børge (February 1992). "On the safety of
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RIB40, for example, appears to have specific salt tolerance genes that regulate K transport.
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Global Aspergillus oryzae Market Report 2020 - Market Size, Share, Price, Trend and Forecast
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Wilkinson, Heather; Tsitsigiannis, Dimitrios; Zhang, Yongqiang; Keller, Nancy (2004-08-30).
768:, which are known to secrete toxins called aflatoxins that cause severe food poisoning, the 2949: 1903: 1842: 1074: 1030:
colonies grown at 37 °C (99 °F) for three days on rich media. The bottom two are
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is hard to study due to difficulties in conventional genetic manipulation. This is because
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is put under conditions favorable to express and secrete aflatoxin, the aflatoxin genes in
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was released by a consortium of Japanese biotechnology companies in late 2005. The eight
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History of Koji - Grains and/or Soybeans Enrobed with a Mold Culture (300 BCE to 2012).
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42149; its morphology, growth, and enzyme production are typical of strains used for
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Genomic analysis has led some scholars to believe that the Japanese domesticated the
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However, in the production of fermented foods of soybeans such as soy sauce and
2872: 2740: 2594: 2413: 1118:(in Japanese). The Society for Biotechnology, Japan. p. 75. Archived from 118: 98: 2280: 2196: 1955: 1650:(5). Japan Society for Bioscience, Biotechnology, and Agrochemistry: 574–584. 1443: 1222: 1149:
Biotechnology in Agriculture and Food Processing: Opportunities and Challenges
2943: 2731: 2421: 2358: 2349: 2289: 2240: 2104: 2008: 1733: 1494: 1451: 1407: 1258: 940: 400: 348:(麹) is used in several meanings, and in some cases it specifically refers to 180: 1999: 1696: 2439: 2376: 2307: 2248: 2204: 2026: 1963: 1925: 1864: 1811: 1714: 1665: 1502: 1365: 1266: 427: 266: 192: 2112: 1656: 1639: 2766: 2620: 2566: 1516: 1197:(in Japanese). The Society of Yeast Scientists. p. 2. Archived from 881: 873: 709: 463: 423: 229: 207: 128: 2061: 1942:
Rokas, A. (2009). "The effect of domestication on the fungal proteome".
1916: 1879: 1855: 1826: 935:) which means mold used in fermented foods, was first mentioned in the 836:
factory because of its relatively few endogenous secondary metabolites.
2890: 2758: 2625: 2612: 2096: 1983:: Learning from the History of Koji Mold and Exploration of Its Future" 692: 408: 296: 108: 88: 2771: 2523: 2916: 2784: 2651: 1802: 1785: 944: 889: 696: 610:
today. It gives rise to a drink with a refreshing, mild, sweet taste.
338: 219: 184: 68: 2528: 1342:. Advances in Applied Microbiology. Vol. 51. pp. 129–153. 2911: 2844: 2779: 2725: 2646: 2551: 2262:
Jin, Feng-Jie; Hu, Shuang; Wang, Bao-Teng; Jin, Long (2021-02-23).
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by Genichirō Kawachi in 1918. This effect was researched and white
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strains are important in rice saccharification for sake brewing:
215: 2392:"Natural products in drug discovery: advances and opportunities" 2503: 961: 792: 688: 642: 285: 278: 196: 78: 2321:
Anyaogu, Diana Chinyere; Mortensen, Uffe Hasbro (2015-02-10).
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to gain new transporters due to the environment's conditions.
558:, playing a role in its recent revival. Thus, white and black 362:
and other molds, so care should be taken to avoid confusion.
2389: 2323:"Heterologous production of fungal secondary metabolites in 1640:"The white koji fungus Aspergillus luchuensis mut. kawachii" 1235: 1046: 967: 955: 733:. The sequenced strain isolated in 1950 is called RIB40 or 700: 530: 333: 224: 201: 416:: pleasant fragrance; accumulation of flavoring compounds 965:, not to mention grain-based wines (including Japanese 2314: 2225: 1824: 1681:"Genome sequence of Aspergillus luchuensis NBRC 4314" 473:(1883 -1948), who is said to be the father of modern 2383: 1877: 452: 1563:"Other terminology relating to Shochu and Awamori" 1104: 1102: 707:is thus one-third larger than that of two related 481:succeeded in the first isolation and culturing of 1977:Machida, M.; Yamada, O.; Gomi, K. (August 2008). 1976: 1428: 1221:(in Japanese). Discover Japan Inc. Archived from 461:Three varieties of kōji mold are used for making 2941: 2320: 1825:Machida, Masayuki; et al. (December 2005). 1298: 1292: 1108: 915: 2049:The Journal of General and Applied Microbiology 1099: 799:. "Flavourzyme", a protease blend derived from 2178: 1637: 1472: 1214: 1187: 1160: 791:can be efficiently cleaved from its glucoside 1727: 1384: 295:that had mutated and ceased to produce toxic 199:in the making of alcoholic beverages such as 2261: 1752: 1581: 1579: 602:was successfully grown independently. White 505:developed by Kawachi has also been used for 477:and Tamaki Inui (1873 -1946), a lecturer at 1644:Bioscience, Biotechnology, and Biochemistry 542:is extremely sensitive to temperature; its 497:, which led to great progress in producing 2229:Journal of Agricultural and Food Chemistry 1937: 1935: 1475:International Journal of Food Microbiology 1305:(in Japanese). p. 643. Archived from 422:: low production of deferriferrichrome (a 31: 2429: 2366: 2348: 2297: 2279: 2060: 2016: 1998: 1915: 1878:Galagan JE, et al. (December 2005). 1854: 1801: 1704: 1655: 1576: 1555: 434:Two of the key enzyme groups secreted by 2042: 1672: 1631: 1331: 1239:Journal of Bioscience and Bioengineering 1181: 824: 594:was discovered as a mutation from black 1932: 1783: 1691:(6). Oxford University Press: 507–515. 1208: 1007:with conidia of the microscopic fungus 795:through the process of fermentation by 783: 574:) is usually used in the production of 2942: 2185:Applied Microbiology and Biotechnology 2085:Applied Microbiology and Biotechnology 1509: 1154: 725:. Many of the extra genes present in 663:) in the mid-1980s, interest in black 467:, each with distinct characteristics. 2533: 2532: 2074: 2072: 2038: 2036: 1941: 1530: 1432:Applied Biochemistry and Microbiology 713:species, the genetics model organism 562:are mainly used in the production of 2927:5667faa9-b9ba-4479-bb3c-6032379a0f89 2688:247e8502-eac3-4577-a727-ee791320ef3e 2172: 1615:. Kawauchi-kin honpo. Archived from 1589:. Kawauchi-kin honpo. Archived from 1325: 981:(the Chinese forerunner of Japanese 550:became more common in production of 1827:"Genome sequencing and analysis of 805:enzyme-hydrolyzed vegetable protein 389:growth on and into the rice kernels 265:is also used for the production of 13: 2136:Secondary Metabolite Gene Clusters 2069: 2033: 1191:家畜化された微生物、麹菌 その分子細胞生物学的解析から見えてきたこと 14: 2986: 2471:Making Rice Koji from Koji Spores 2464: 2045:"Classification of the Koji Mold" 1487:10.1016/j.ijfoodmicro.2013.03.027 1279:Fujita, Chieko, Tokyo Foundation 744:The increased number of genes in 327:brewing, but also for making the 2476:Sake World's description of koji 1784:Goffeau, André (December 2005). 1019: 997: 729:are predicted to be involved in 277:(米麹) are made by fermenting the 55: 2446: 2255: 2219: 2138:. CRC Press. pp. 355–385. 2127: 1970: 1871: 1818: 1777: 1721: 1605: 1466: 1422: 1378: 1313: 1215:Kiyoko Hayashi (19 July 2021). 846:1,3,6,8-tetrahydroxynaphthalene 1884:and comparative analysis with 1728:Takeo Koizumi (5 April 2018). 1273: 1229: 1141: 1129: 1088: 719:and the potentially dangerous 493:, and a variety of subtaxa of 430:, and other colored substances 1: 2960:Molds used in food production 2397:Nature Reviews Drug Discovery 1400:10.1080/08905436.2016.1244768 1348:10.1016/S0065-2164(02)51004-2 1082: 695:together comprise 37 million 257:is sometimes used instead of 228:. It is one of the different 1332:Kitamoto, Katsuhiko (2002). 1251:10.1016/j.jbiosc.2010.12.022 766:A. flavus and A. parasiticus 374:The following properties of 315:fungus a "national fungus" ( 244:used for food fermentation. 7: 1638:Taiki Futagami (May 2022). 1039: 684:Initially kept secret, the 10: 2991: 2452:Shurtleff, W.; Aoyagi, A. 2414:10.1038/s41573-020-00114-z 990: 699:and 12 thousand predicted 626:is mainly used to produce 2834: 2711: 2541: 2332:Frontiers in Microbiology 2281:10.3389/fmicb.2021.644404 2268:Frontiers in Microbiology 2197:10.1007/s00253-007-0874-3 2043:Murakami, Hideya (1971). 1956:10.1016/j.tig.2008.11.003 1444:10.1134/S0003683813040170 1335:Molecular Biology of the 941:Rites of the Zhou dynasty 679: 239: 157: 150: 52:Scientific classification 50: 39: 30: 23: 2350:10.3389/fmicb.2015.00077 2241:10.1021/acs.jafc.5b01665 1301: 1217: 1190: 1163: 1111: 403:and glucoamylase); some 366:Properties desirable in 43:growing on rice to make 1299:Kenichiro Matsushima. 1109:Kenichiro Matsushima. 832:is a good choice as a 534:, and at one time all 501:in Japan. Since then, 395:: strong secretion of 2000:10.1093/dnares/dsn020 1697:10.1093/dnares/dsw032 1587:"初代 河内源一郎(1883~1948)" 1302:醤油づくりと麹菌の利用ー今までとこれからー 854:3-methylorcinaldehyde 825:Secondary metabolites 803:, is used to produce 1882:Aspergillus nidulans 1281:Koji, an Aspergillus 1225:on 10 November 2022. 1204:on 13 November 2022. 1188:Katsuhiko Kitamoto. 1161:Katsuhiko Kitamoto. 1147:Parmjit S. Panesar, 1138:Kikkoman Corporation 1075:Rhizopus oligosporus 834:secondary metabolite 784:Use in biotechnology 780:were not expressed. 731:secondary metabolism 2713:Aspergillus flavus 2062:10.2323/jgam.17.281 1917:10.1038/nature04341 1908:2005Natur.438.1105G 1902:(7071): 1105–1115. 1856:10.1038/nature04300 1847:2005Natur.438.1157M 1841:(7071): 1157–1161. 1796:(7071): 1092–1093. 1730:"黒麹菌の役割 発酵中の雑菌繁殖防ぐ" 1657:10.1093/bbb/zbac033 1177:on 31 October 2022. 842:polyketide synthase 840:types can produce: 764:is closely related 528:is used to produce 479:University of Tokyo 453:Varieties used for 370:brewing and testing 2573:Aspergillus oryzae 2543:Aspergillus oryzae 2517:2021-11-08 at the 2482:Aspergillus oryzae 2181:Aspergillus oryzae 2097:10.1007/BF00183230 2081:Aspergillus oryzae 1981:Aspergillus oryzae 1944:Trends in Genetics 1829:Aspergillus oryzae 1388:Food Biotechnology 1286:2009-05-22 at the 904:, originally from 746:Aspergillus oryzae 566:, but only yellow 538:. However, yellow 359:Monascus purpureus 344:The Japanese word 307:Eiji Ichishima of 293:Aspergillus flavus 176:Aspergillus oryzae 161:Aspergillus oryzae 25:Aspergillus oryzae 16:Filamentous fungus 2937: 2936: 2819:Open Tree of Life 2696:Open Tree of Life 2535:Taxon identifiers 2235:(23): 5682–5693. 2145:978-0-8247-5655-0 1786:"Multiple moulds" 1538:"What is Shochu?" 1322:Marukome co.,ltd. 1112:しょうゆづくりの歩みと麹菌の関わり 1058:Aspergillus sojae 898:co-transformation 789:Trans-resveratrol 667:resurged amongst 471:Genichirō Kawachi 309:Tohoku University 299:, giving rise to 254:Aspergillus sojae 172: 171: 167:(Ahlburg) E. Cohn 2982: 2970:Japanese cuisine 2930: 2929: 2920: 2919: 2907: 2906: 2894: 2893: 2881: 2880: 2868: 2867: 2855: 2854: 2853: 2827: 2826: 2814: 2813: 2801: 2800: 2798:BMSSYS0000042725 2788: 2787: 2775: 2774: 2762: 2761: 2749: 2748: 2736: 2735: 2734: 2716: 2704: 2703: 2691: 2690: 2681: 2680: 2668: 2667: 2665:NHMSYS0020535053 2655: 2654: 2642: 2641: 2629: 2628: 2616: 2615: 2603: 2602: 2590: 2589: 2577: 2576: 2575: 2562: 2561: 2560: 2530: 2529: 2459: 2450: 2444: 2443: 2433: 2406:Nature Portfolio 2387: 2381: 2380: 2370: 2352: 2318: 2312: 2311: 2301: 2283: 2259: 2253: 2252: 2223: 2217: 2216: 2176: 2170: 2169: 2163: 2159: 2157: 2149: 2131: 2125: 2124: 2076: 2067: 2066: 2064: 2040: 2031: 2030: 2020: 2002: 1974: 1968: 1967: 1939: 1930: 1929: 1919: 1875: 1869: 1868: 1858: 1822: 1816: 1815: 1805: 1803:10.1038/4381092b 1781: 1775: 1774: 1772: 1771: 1762:. Archived from 1756: 1750: 1749: 1747: 1745: 1736:. Archived from 1725: 1719: 1718: 1708: 1676: 1670: 1669: 1659: 1635: 1629: 1628: 1626: 1624: 1613:"元祖 源一郎さんの生マッコリ" 1609: 1603: 1602: 1600: 1598: 1583: 1574: 1572: 1570: 1569: 1559: 1553: 1552: 1550: 1549: 1540:. Archived from 1534: 1528: 1526: 1524: 1523: 1513: 1507: 1506: 1470: 1464: 1463: 1426: 1420: 1419: 1382: 1376: 1375: 1373: 1372: 1329: 1323: 1317: 1311: 1310: 1309:on 21 June 2022. 1296: 1290: 1277: 1271: 1270: 1233: 1227: 1226: 1212: 1206: 1205: 1203: 1196: 1185: 1179: 1178: 1176: 1169: 1158: 1152: 1151:CRC Press (2014) 1145: 1139: 1133: 1127: 1126: 1125:on 26 July 2020. 1124: 1117: 1106: 1097: 1092: 1023: 1013:light microscope 1001: 920:in a broad sense 703:. The genome of 675:on their labels. 485:species such as 405:carboxypeptidase 243: 241: 163: 60: 59: 35: 21: 20: 2990: 2989: 2985: 2984: 2983: 2981: 2980: 2979: 2940: 2939: 2938: 2933: 2925: 2923: 2915: 2910: 2902: 2897: 2889: 2884: 2876: 2871: 2863: 2858: 2849: 2848: 2843: 2836:Eurotium oryzae 2830: 2822: 2817: 2809: 2804: 2796: 2791: 2783: 2778: 2770: 2765: 2757: 2752: 2744: 2739: 2730: 2729: 2724: 2714: 2707: 2699: 2694: 2686: 2684: 2676: 2671: 2663: 2658: 2650: 2645: 2637: 2632: 2624: 2619: 2611: 2606: 2598: 2593: 2585: 2580: 2571: 2570: 2565: 2556: 2555: 2550: 2537: 2519:Wayback Machine 2467: 2462: 2451: 2447: 2388: 2384: 2319: 2315: 2260: 2256: 2224: 2220: 2177: 2173: 2161: 2160: 2151: 2150: 2146: 2132: 2128: 2077: 2070: 2041: 2034: 1975: 1971: 1940: 1933: 1880:"Sequencing of 1876: 1872: 1823: 1819: 1782: 1778: 1769: 1767: 1760:"Shochu Circle" 1758: 1757: 1753: 1743: 1741: 1740:on 4 April 2023 1726: 1722: 1677: 1673: 1636: 1632: 1622: 1620: 1611: 1610: 1606: 1596: 1594: 1585: 1584: 1577: 1567: 1565: 1561: 1560: 1556: 1547: 1545: 1536: 1535: 1531: 1521: 1519: 1515: 1514: 1510: 1471: 1467: 1427: 1423: 1383: 1379: 1370: 1368: 1358: 1330: 1326: 1318: 1314: 1303: 1297: 1293: 1288:Wayback Machine 1278: 1274: 1234: 1230: 1219: 1213: 1209: 1201: 1194: 1192: 1186: 1182: 1174: 1167: 1165: 1159: 1155: 1146: 1142: 1134: 1130: 1122: 1115: 1113: 1107: 1100: 1093: 1089: 1085: 1080: 1069:Medicinal molds 1042: 1035: 1024: 1015: 1002: 993: 922: 910:Gateway cloning 827: 786: 682: 646:. Of all three 459: 372: 240:nihon kōji kabi 233: 232:ニホンコウジカビ (日本麹黴) 168: 165: 159: 146: 54: 17: 12: 11: 5: 2988: 2978: 2977: 2975:Fungus species 2972: 2967: 2965:Fungi of Japan 2962: 2957: 2952: 2935: 2934: 2932: 2931: 2921: 2908: 2895: 2882: 2869: 2856: 2840: 2838: 2832: 2831: 2829: 2828: 2815: 2802: 2789: 2776: 2763: 2750: 2737: 2721: 2719: 2709: 2708: 2706: 2705: 2692: 2682: 2669: 2656: 2643: 2630: 2617: 2604: 2591: 2578: 2563: 2547: 2545: 2539: 2538: 2527: 2526: 2521: 2509: 2478: 2473: 2466: 2465:External links 2463: 2461: 2460: 2445: 2382: 2313: 2254: 2218: 2191:(4): 763–768. 2171: 2144: 2126: 2091:(5): 569–572. 2068: 2055:(4): 281–309. 2032: 1993:(4): 173–183. 1969: 1931: 1870: 1817: 1776: 1751: 1720: 1671: 1630: 1604: 1575: 1554: 1529: 1508: 1481:(2): 148–154. 1465: 1438:(4): 378–385. 1421: 1394:(4): 278–291. 1377: 1356: 1324: 1312: 1291: 1272: 1245:(5): 512–517. 1228: 1207: 1180: 1153: 1140: 1128: 1098: 1095:Index Fungorum 1086: 1084: 1081: 1079: 1078: 1071: 1066: 1061: 1054: 1049: 1043: 1041: 1038: 1037: 1036: 1025: 1018: 1016: 1003: 996: 992: 989: 921: 914: 862:terrequinone A 826: 823: 785: 782: 681: 678: 677: 676: 669:honkaku shōchū 611: 579: 536:honkaku shōchū 458: 451: 432: 431: 417: 411: 390: 371: 364: 211:, and also to 170: 169: 166: 155: 154: 148: 147: 143:A. oryzae 140: 138: 134: 133: 126: 122: 121: 119:Aspergillaceae 116: 112: 111: 106: 102: 101: 99:Eurotiomycetes 96: 92: 91: 86: 82: 81: 76: 72: 71: 66: 62: 61: 48: 47: 37: 36: 28: 27: 15: 9: 6: 4: 3: 2: 2987: 2976: 2973: 2971: 2968: 2966: 2963: 2961: 2958: 2956: 2953: 2951: 2948: 2947: 2945: 2928: 2922: 2918: 2913: 2909: 2905: 2900: 2896: 2892: 2887: 2883: 2879: 2874: 2870: 2866: 2861: 2857: 2852: 2846: 2842: 2841: 2839: 2837: 2833: 2825: 2820: 2816: 2812: 2807: 2803: 2799: 2794: 2790: 2786: 2781: 2777: 2773: 2768: 2764: 2760: 2755: 2751: 2747: 2742: 2738: 2733: 2727: 2723: 2722: 2720: 2718: 2710: 2702: 2697: 2693: 2689: 2683: 2679: 2674: 2670: 2666: 2661: 2657: 2653: 2648: 2644: 2640: 2635: 2631: 2627: 2622: 2618: 2614: 2609: 2605: 2601: 2596: 2592: 2588: 2583: 2579: 2574: 2568: 2564: 2559: 2553: 2549: 2548: 2546: 2544: 2540: 2536: 2531: 2525: 2522: 2520: 2516: 2513: 2510: 2508: 2506: 2501: 2497: 2493: 2489: 2485: 2483: 2479: 2477: 2474: 2472: 2469: 2468: 2457: 2456: 2449: 2441: 2437: 2432: 2427: 2423: 2419: 2415: 2411: 2407: 2403: 2399: 2398: 2393: 2386: 2378: 2374: 2369: 2364: 2360: 2356: 2351: 2346: 2342: 2338: 2334: 2333: 2328: 2326: 2317: 2309: 2305: 2300: 2295: 2291: 2287: 2282: 2277: 2273: 2269: 2265: 2258: 2250: 2246: 2242: 2238: 2234: 2230: 2222: 2214: 2210: 2206: 2202: 2198: 2194: 2190: 2186: 2182: 2175: 2167: 2155: 2147: 2141: 2137: 2130: 2122: 2118: 2114: 2110: 2106: 2102: 2098: 2094: 2090: 2086: 2083:: a review". 2082: 2075: 2073: 2063: 2058: 2054: 2050: 2046: 2039: 2037: 2028: 2024: 2019: 2014: 2010: 2006: 2001: 1996: 1992: 1988: 1984: 1982: 1979:"Genomics of 1973: 1965: 1961: 1957: 1953: 1949: 1945: 1938: 1936: 1927: 1923: 1918: 1913: 1909: 1905: 1901: 1897: 1893: 1891: 1887: 1883: 1874: 1866: 1862: 1857: 1852: 1848: 1844: 1840: 1836: 1832: 1830: 1821: 1813: 1809: 1804: 1799: 1795: 1791: 1787: 1780: 1766:on 2007-12-10 1765: 1761: 1755: 1739: 1735: 1734:Okinawa Times 1731: 1724: 1716: 1712: 1707: 1702: 1698: 1694: 1690: 1686: 1682: 1675: 1667: 1663: 1658: 1653: 1649: 1645: 1641: 1634: 1619:on 7 May 2020 1618: 1614: 1608: 1593:on 7 May 2020 1592: 1588: 1582: 1580: 1564: 1558: 1544:on 2007-09-28 1543: 1539: 1533: 1518: 1512: 1504: 1500: 1496: 1492: 1488: 1484: 1480: 1476: 1469: 1461: 1457: 1453: 1449: 1445: 1441: 1437: 1433: 1425: 1417: 1413: 1409: 1405: 1401: 1397: 1393: 1389: 1381: 1367: 1363: 1359: 1357:9780120026531 1353: 1349: 1345: 1341: 1340: 1336: 1328: 1321: 1316: 1308: 1304: 1295: 1289: 1285: 1282: 1276: 1268: 1264: 1260: 1256: 1252: 1248: 1244: 1240: 1232: 1224: 1220: 1211: 1200: 1193: 1184: 1173: 1166: 1157: 1150: 1144: 1137: 1132: 1121: 1114: 1105: 1103: 1096: 1091: 1087: 1077: 1076: 1072: 1070: 1067: 1065: 1062: 1060: 1059: 1055: 1053: 1050: 1048: 1045: 1044: 1033: 1029: 1022: 1017: 1014: 1010: 1006: 1005:Conidiophores 1000: 995: 994: 988: 986: 985: 980: 976: 975: 970: 969: 964: 963: 958: 957: 952: 951: 946: 942: 938: 934: 930: 926: 919: 913: 911: 907: 903: 899: 895: 891: 887: 883: 879: 875: 871: 867: 863: 859: 855: 851: 850:alternapyrone 847: 843: 839: 835: 831: 822: 820: 816: 812: 808: 806: 802: 798: 794: 790: 781: 779: 775: 771: 767: 763: 758: 756: 752: 747: 742: 740: 736: 732: 728: 724: 723: 718: 717: 712: 711: 706: 702: 698: 694: 690: 687: 674: 670: 666: 662: 658: 653: 649: 645: 644: 639: 635: 634: 629: 625: 623: 622: 621:A. luchuensis 617: 612: 609: 605: 601: 597: 593: 591: 590: 585: 580: 577: 573: 569: 565: 561: 557: 553: 549: 545: 541: 537: 533: 532: 527: 525: 521: 516: 515: 514: 512: 508: 504: 500: 496: 492: 488: 484: 480: 476: 472: 468: 466: 465: 456: 450: 448: 444: 440: 437: 429: 425: 421: 418: 415: 412: 410: 406: 402: 398: 394: 391: 388: 384: 381: 380: 379: 377: 369: 363: 361: 360: 355: 351: 347: 342: 340: 336: 335: 330: 326: 322: 318: 314: 310: 306: 302: 298: 294: 289: 287: 284: 280: 276: 273:(麦麹) or rice 272: 268: 267:rice vinegars 264: 260: 256: 255: 250: 245: 237: 231: 227: 226: 221: 217: 214: 210: 209: 204: 203: 198: 194: 190: 186: 182: 178: 177: 164: 162: 156: 153: 152:Binomial name 149: 145: 144: 139: 136: 135: 132: 131: 127: 124: 123: 120: 117: 114: 113: 110: 107: 104: 103: 100: 97: 94: 93: 90: 87: 84: 83: 80: 77: 74: 73: 70: 67: 64: 63: 58: 53: 49: 46: 42: 38: 34: 29: 26: 22: 19: 2835: 2712: 2542: 2504: 2499: 2495: 2491: 2487: 2481: 2453: 2448: 2401: 2395: 2385: 2336: 2330: 2324: 2316: 2271: 2267: 2257: 2232: 2228: 2221: 2188: 2184: 2180: 2174: 2135: 2129: 2088: 2084: 2080: 2052: 2048: 1990: 1987:DNA Research 1986: 1980: 1972: 1950:(2): 60–63. 1947: 1943: 1899: 1895: 1889: 1886:A. fumigatus 1885: 1881: 1873: 1838: 1834: 1828: 1820: 1793: 1789: 1779: 1768:. Retrieved 1764:the original 1754: 1742:. Retrieved 1738:the original 1723: 1688: 1685:DNA Research 1684: 1674: 1647: 1643: 1633: 1621:. Retrieved 1617:the original 1607: 1595:. Retrieved 1591:the original 1566:. Retrieved 1557: 1546:. Retrieved 1542:the original 1532: 1520:. Retrieved 1511: 1478: 1474: 1468: 1435: 1431: 1424: 1391: 1387: 1380: 1369:. Retrieved 1338: 1334: 1327: 1315: 1307:the original 1294: 1275: 1242: 1238: 1231: 1223:the original 1210: 1199:the original 1183: 1172:the original 1156: 1148: 1143: 1131: 1120:the original 1090: 1073: 1056: 1031: 1027: 1008: 982: 978: 972: 971:and Chinese 966: 960: 954: 948: 936: 932: 928: 924: 923: 917: 905: 870:pyripyropene 829: 828: 818: 814: 810: 809: 800: 796: 787: 777: 773: 769: 765: 761: 759: 754: 750: 745: 743: 738: 726: 722:A. fumigatus 720: 714: 708: 704: 685: 683: 672: 668: 664: 660: 656: 651: 647: 641: 637: 631: 627: 619: 615: 613: 607: 603: 599: 595: 587: 583: 581: 575: 571: 567: 563: 559: 555: 551: 547: 543: 539: 535: 529: 523: 519: 517: 510: 506: 502: 498: 494: 490: 486: 482: 474: 469: 462: 460: 454: 446: 442: 441: 435: 433: 419: 413: 392: 382: 375: 373: 367: 357: 353: 349: 345: 343: 332: 328: 324: 320: 316: 312: 304: 300: 292: 290: 282: 274: 270: 262: 258: 252: 248: 246: 223: 206: 200: 193:sweet potato 175: 174: 173: 160: 158: 142: 141: 129: 44: 40: 24: 18: 2950:Aspergillus 2767:iNaturalist 2621:iNaturalist 2567:Wikispecies 2502:nalysed at 2408:: 200–216. 2162:|work= 1028:Aspergillus 931:, Japanese 916:History of 906:A. nidulans 888:insertion; 882:andrastin A 874:aphidicolin 838:Transformed 716:A. nidulans 710:Aspergillus 693:chromosomes 589:A. kawachii 503:aspergillus 487:A. kawachii 483:aspergillus 424:siderophore 311:called the 218:for making 130:Aspergillus 2944:Categories 2325:Aspergilli 2274:: 644404. 1770:2007-12-11 1573:(Japanese) 1568:2007-01-27 1548:2007-01-24 1527:(Japanese) 1522:2007-01-24 1517:"In-depth" 1371:2008-01-03 1218:日本の発酵技術と歴史 1083:References 1052:Akira Endo 902:aspyridone 878:terretonin 697:base pairs 638:A. awamori 513:in Korea. 491:A. awamori 414:Aesthetics 409:tyrosinase 305:A. oryzae. 297:aflatoxins 230:koji molds 189:saccharify 109:Eurotiales 89:Ascomycota 85:Division: 2955:Rice wine 2851:Q80832328 2558:Q22662068 2486:from the 2422:1474-1776 2359:1664-302X 2341:Frontiers 2290:1664-302X 2164:ignored ( 2154:cite book 2105:0175-7598 2009:1340-2838 1890:A. oryzae 1495:0168-1605 1460:254189777 1452:1608-3024 1408:0890-5436 1259:1389-1723 1032:A. oryzae 1009:A. oryzae 945:soy sauce 927:(Chinese 890:paxilline 844:-derived 830:A. oryzae 819:A. oryzae 815:A. oryzae 811:A. oryzae 801:A. oryzae 797:A. oryzae 778:A. oryzae 774:A. oryzae 762:A. oryzae 760:Although 751:A. oryzae 741:brewing. 727:A. oryzae 705:A. oryzae 686:A. oryzae 657:kuro-kōji 572:A. oryzae 524:A. oryzae 511:makgeolli 495:A. oryzae 447:A. oryzae 443:A. oryzae 436:A. oryzae 401:α-amylase 376:A. oryzae 350:A. oryzae 339:soy sauce 301:A. oryzae 283:A. oryzae 269:. Barley 263:A. oryzae 259:A. oryzae 220:soy sauce 185:East Asia 137:Species: 75:Kingdom: 69:Eukaryota 41:A. oryzae 2912:MycoBank 2904:10644360 2873:Fungorum 2845:Wikidata 2780:MycoBank 2741:Fungorum 2726:Wikidata 2647:MycoBank 2639:10388277 2595:Fungorum 2552:Wikidata 2515:Archived 2490:atabase 2440:33510482 2377:25713568 2308:33708187 2249:25996918 2213:13139293 2205:17333175 2027:18820080 1964:19081651 1926:16372000 1865:16372010 1812:16371993 1744:10 April 1715:27651094 1666:35238900 1503:23673060 1416:88669475 1366:12236056 1284:Archived 1267:21342785 1040:See also 1034:strains. 974:huangjiu 894:aflatrem 866:tennelin 858:citrinin 397:amylases 387:mycelial 385:: rapid 354:A. sojae 236:Japanese 216:soybeans 183:used in 115:Family: 65:Domain: 2891:2597225 2824:5484773 2759:5259826 2732:Q131911 2626:1148191 2613:5259827 2524:(DOGAN) 2498:enomes 2431:7841765 2368:4322707 2299:7940364 2121:7348198 2113:1368061 2018:2575883 1904:Bibcode 1843:Bibcode 1706:5144674 1623:9 April 1597:9 April 1136:麹菌ゲノム解読 1064:Lactase 991:Gallery 984:amazake 886:plasmid 633:awamori 518:Yellow 428:flavins 393:Enzymes 213:ferment 125:Genus: 105:Order: 95:Class: 2924:NZOR: 2917:225012 2878:225012 2811:748371 2785:130238 2772:488517 2746:130238 2717:oryzae 2701:188996 2685:NZOR: 2652:184394 2600:184394 2484:genome 2438:  2428:  2420:  2375:  2365:  2357:  2343:: 77. 2306:  2296:  2288:  2247:  2211:  2203:  2142:  2119:  2111:  2103:  2025:  2015:  2007:  1962:  1924:  1896:Nature 1863:  1835:Nature 1810:  1790:Nature 1713:  1703:  1664:  1501:  1493:  1458:  1450:  1414:  1406:  1364:  1354:  1265:  1257:  1011:under 977:) and 962:douchi 959:, and 937:Zhouli 900:; and 880:, and 852:, and 793:piceid 689:genome 680:Genome 652:shōchū 643:moromi 628:shōchū 614:Black 608:shōchū 582:White 564:shōchū 556:shōchū 552:shōchū 544:moromi 499:shōchū 475:shōchū 464:shōchū 457:making 455:shōchū 407:; low 383:Growth 317:kokkin 286:hyphae 279:grains 208:shōchū 197:barley 195:, and 191:rice, 2899:IRMNG 2865:3D2L9 2634:IRMNG 2404:(3). 2209:S2CID 2117:S2CID 1456:S2CID 1412:S2CID 1339:Molds 1202:(PDF) 1195:(PDF) 1175:(PDF) 1168:(PDF) 1123:(PDF) 1116:(PDF) 1026:Four 950:jiang 908:, by 701:genes 420:Color 281:with 179:is a 79:Fungi 2886:GBIF 2806:NCBI 2754:GBIF 2715:var. 2678:5062 2673:NCBI 2608:GBIF 2587:HC8N 2436:PMID 2418:ISSN 2373:PMID 2355:ISSN 2304:PMID 2286:ISSN 2245:PMID 2201:PMID 2166:help 2140:ISBN 2109:PMID 2101:ISSN 2023:PMID 2005:ISSN 1960:PMID 1922:PMID 1888:and 1861:PMID 1808:PMID 1746:2023 1711:PMID 1662:PMID 1625:2023 1599:2023 1499:PMID 1491:ISSN 1448:ISSN 1404:ISSN 1362:PMID 1352:ISBN 1337:Koji 1320:麹のこと 1263:PMID 1255:ISSN 1164:麹菌物語 968:sake 956:miso 933:kōji 892:and 770:kōji 755:kōji 739:sake 735:ATCC 673:kōji 665:kōji 661:kōji 659:(NK- 648:kōji 630:and 616:kōji 604:kōji 600:kōji 596:kōji 584:kōji 576:sake 568:kōji 560:kōji 548:kōji 540:kōji 531:sake 520:kōji 509:and 507:soju 368:sake 352:and 346:kōji 334:miso 331:for 329:kōji 325:sake 323:for 321:kōji 313:kōji 275:kōji 271:kōji 249:miso 225:miso 222:and 205:and 202:sake 181:mold 45:koji 2860:CoL 2793:NBN 2660:NBN 2582:CoL 2507:ITE 2426:PMC 2410:doi 2363:PMC 2345:doi 2294:PMC 2276:doi 2237:doi 2193:doi 2183:". 2093:doi 2057:doi 2013:PMC 1995:doi 1952:doi 1912:doi 1900:438 1851:doi 1839:438 1798:doi 1794:438 1701:PMC 1693:doi 1652:doi 1483:doi 1479:164 1440:doi 1396:doi 1344:doi 1247:doi 1243:111 987:). 896:by 884:by 426:), 261:. 187:to 2946:: 2914:: 2901:: 2888:: 2875:: 2862:: 2847:: 2821:: 2808:: 2795:: 2782:: 2769:: 2756:: 2743:: 2728:: 2698:: 2675:: 2662:: 2649:: 2636:: 2623:: 2610:: 2597:: 2584:: 2569:: 2554:: 2494:f 2434:. 2424:. 2416:. 2402:20 2400:. 2394:. 2371:. 2361:. 2353:. 2339:. 2335:. 2329:. 2302:. 2292:. 2284:. 2272:12 2270:. 2266:. 2243:. 2233:63 2231:. 2207:. 2199:. 2189:75 2187:. 2158:: 2156:}} 2152:{{ 2115:. 2107:. 2099:. 2089:36 2087:. 2071:^ 2053:17 2051:. 2047:. 2035:^ 2021:. 2011:. 2003:. 1991:15 1989:. 1985:. 1958:. 1948:25 1946:. 1934:^ 1920:. 1910:. 1898:. 1894:. 1859:. 1849:. 1837:. 1833:. 1806:. 1792:. 1788:. 1732:. 1709:. 1699:. 1689:23 1687:. 1683:. 1660:. 1648:86 1646:. 1642:. 1578:^ 1497:. 1489:. 1477:. 1454:. 1446:. 1436:49 1434:. 1410:. 1402:. 1392:30 1390:. 1360:. 1350:. 1261:. 1253:. 1241:. 1101:^ 1047:Qū 979:li 947:, 929:qū 912:. 876:, 872:, 868:, 864:; 860:; 856:; 848:, 807:. 624:) 489:, 337:, 288:. 251:, 238:: 2505:N 2500:A 2496:G 2492:o 2488:D 2442:. 2412:: 2379:. 2347:: 2337:6 2327:" 2310:. 2278:: 2251:. 2239:: 2215:. 2195:: 2168:) 2148:. 2123:. 2095:: 2065:. 2059:: 2029:. 1997:: 1966:. 1954:: 1928:. 1914:: 1906:: 1892:" 1867:. 1853:: 1845:: 1831:" 1814:. 1800:: 1773:. 1748:. 1717:. 1695:: 1668:. 1654:: 1627:. 1601:. 1571:. 1551:. 1525:. 1505:. 1485:: 1462:. 1442:: 1418:. 1398:: 1374:. 1346:: 1269:. 1249:: 953:/ 939:( 925:麹 918:麹 618:( 592:) 586:( 578:. 570:( 526:) 522:( 399:( 242:) 234:(

Index


Scientific classification
Edit this classification
Eukaryota
Fungi
Ascomycota
Eurotiomycetes
Eurotiales
Aspergillaceae
Aspergillus
Binomial name
mold
East Asia
saccharify
sweet potato
barley
sake
shōchū
ferment
soybeans
soy sauce
miso
koji molds
Japanese
Aspergillus sojae
rice vinegars
grains
hyphae
aflatoxins
Tohoku University

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