320:
287:
381:
372:
363:
354:
345:
336:
327:
467:
Dinosaur remains are among the fossils that have been recovered from the formation. The formation is about 45 metres (148 ft) thick and consists primarily of clay with interbeds of sand, representing an estuarine environment.
1127:
L. A. Nessov. 1995. Dinozavri severnoi
Yevrazii: Novye dannye o sostave kompleksov, ekologii i paleobiogeografii . Institute for Scientific Research on the Earth's Crust, St. Petersburg State University, St. Petersburg
1140:
A. K. Rozhdestvensky. 1964. Novye dannye o mestonakhozhdeniyakh dinozavrov na territorii
Kazakhstana i Srednei Azii . Tashkentskii Gosudarstvennyi Universitet, Nauchnye Trudy: Geologiya 234:227-241
1124:
E. G. Kordikova, S. M. Kurzanov, and G. F. Gunnell. 1997. Unusual dinosaur claw phalanxes from the Upper
Cretaceous of the northeastern Aral Sea region, Kazakhstan. Journal of Morphology 232(3):278
1137:
L. A. Nessov. 1981. Amfibii i reptilii v ekosistemakh Mela sredney Azii . In I. V. Darevsky (ed.), The
Problems of Herpetology. Fifth Herpetological Conference. Abstracts 91–92
890:"Frontal bones of non-avian theropod dinosaurs from the Upper Cretaceous (Santonian-?Campanian) Bostobe Formation of the northeastern Aral Sea region, Kazakhstan"
775:
1134:
L. A. Nessov. 1984. Data on late
Mesozoic turtles from the USSR. Studia Geologica Salamanticensia, vol. especial 1 (Studia Palaeocheloniologica I) 1:215-223
710:
1203:
1121:
A. O. Averianov. 2007. Theropod dinosaurs from Late
Cretaceous deposits in the northeastern Aral Sea region, Kazakhstan. Cretaceous Research 28:532-544
134:
520:
307:
670:
513:
506:
677:
1153:
1158:
770:
1213:
319:
1131:
L. A. Nessov. 1988. Late
Mesozoic amphibians and lizards of Soviet Middle Asia. Acta Zoologica Cracoviensia 31(14):475-486
1208:
894:
1063:
1026:
Averianov, A.O (2004), "New Data on
Cretaceous Flying Reptiles (Pterosauria) of Russia, Kazakhstan, and Kyrgyzstan",
235:
196:
1163:
1080:"Small vertebrates from the late Cretaceous and early Tertiary of the northeastern Aral Sea region, Kazakhstan"
488:
recovered from the formation, was initially thought to belong to a hadrosaurid dinosaur and classified as cf.
1178:
1007:
938:
1198:
189:
968:"New mammal remains from the Late Cretaceous Bostobe Formation (northeast Aral Sea Region, Kazakhstan)"
967:
1218:
1193:
1188:
1183:
1173:
1168:
461:
1087:
1028:
1078:
Kordikova, E.G.; Polly, P.D.; Alifanov, V.A.; RoÄŤek, Z.; Gunnell, G.F.; Averianov, A.O. (2001),
903:
790:
144:
124:
8:
1041:
928:
Danilov, Igor G.; Vitek, Natasha S.; Averianov, Alexander O.; Glinskiy, Vadim N. (2015),
411:
1049:
907:
1002:
929:
780:
1079:
1104:
1059:
785:
1096:
984:
947:
911:
889:
615:
1053:
988:
604:
423:
403:
795:
730:
609:
593:
574:
533:
228:
1100:
1147:
1108:
628:
554:
250:
237:
211:
198:
74:
27:
1045:
915:
724:
663:
599:
561:
490:
482:
477:
952:
975:
547:
458:
49:
966:
Averianov, A.O.; Archibald, J.D.; Skutschas, P.P.; Dyke, G.J. (2014a),
656:
454:
415:
292:
270:
94:
59:
580:
442:
438:
431:
427:
419:
274:
177:
167:
99:
43:
35:
31:
757:
744:
635:
89:
84:
69:
64:
54:
696:
690:
684:
450:
446:
104:
79:
965:
1003:"A new eutherian mammal from the Late Cretaceous of Kazakhstan"
649:
485:
927:
1058:, Berkeley: University of California Press, pp. 1–880,
1077:
934:
from the Upper
Cretaceous Bostobe Formation of Kazakhstan"
1040:
1000:
776:
List of fossiliferous stratigraphic units in
Kazakhstan
807:
1001:
Averianov, A.; Archibald, J.D.; Dyke, G.J. (2014b),
930:"A new softshelled trionychid turtle of the genus
1145:
846:
837:
830:
828:
869:
816:
422:, Kazakhastan whose strata date back to the
825:
1204:Fossiliferous stratigraphic units of Asia
1025:
951:
887:
862:
860:
858:
380:
371:
362:
353:
344:
335:
326:
771:List of dinosaur-bearing rock formations
1146:
855:
813:Weishampel et al. , 2004, pp.593-600
445:of the formation were deposited in
13:
1115:
895:Canadian Journal of Earth Sciences
14:
1230:
1154:Geologic formations of Kazakhstan
471:
379:
370:
361:
352:
343:
334:
325:
318:
285:
47:
16:Stratigraphic Unit in Kazakhstan
1159:Upper Cretaceous Series of Asia
881:
434:stages, approximately 85 Ma).
388:Bostobe Formation (Kazakhstan)
1:
1214:Geography of Karaganda Region
1008:Acta Palaeontologica Polonica
939:Acta Palaeontologica Polonica
801:
154:About 45 m (148 ft)
989:10.1016/j.palwor.2014.10.009
562:Batyrosaurus rozhdestvenskyi
229:Approximate paleocoordinates
7:
1055:The Dinosauria, 2nd edition
764:
555:Arstanosaurus akkurganensis
10:
1235:
1209:Paleontology in Kazakhstan
1101:10.1017/S0022336000018187
664:Khunnuchelys lophorhothon
629:Aralazhdarcho bostobensis
478:Khunnuchelys lophorhothon
407:
313:
303:
298:
280:
266:
227:
188:
183:
173:
163:
158:
150:
140:
130:
120:
26:
21:
1088:Journal of Paleontology
1029:Paleontological Journal
852:Averianov et al., 2014a
843:Averianov et al., 2014b
616:Tyrannosauroidea indet.
548:Aralosaurus tuberiferus
916:10.1139/cjes-2015-0099
822:Kordikova et al., 2001
605:Therizinosauria indet.
1164:Cretaceous Kazakhstan
953:10.4202/app.2013.0045
888:Averianov, A (2016),
678:Shachemys baibolatica
1179:Sandstone formations
1042:Weishampel, David B.
834:Danilov et al., 2014
791:Karabastau Formation
514:Zhalmouzia bazhanovi
145:Zhirkindek Formation
125:Geological formation
1199:Lacustrine deposits
908:2016CaJES..53..168A
685:Trionyx kansaiensis
671:Lindholmemys gravis
600:Troodontidae indet.
247: /
208: /
28:Stratigraphic range
781:Bissekty Formation
657:Anatolemys maximus
410:) is a geological
1050:OsmĂłlska, Halszka
786:Madygen Formation
521:Parazhelestes sp.
507:Beleutinus orlovi
400:Bostobe Formation
396:
395:
22:Bostobe Formation
1226:
1219:Kyzylorda Region
1194:Fluvial deposits
1189:Deltaic deposits
1184:Shale formations
1111:
1084:
1074:
1073:
1072:
1037:
1022:
1021:
1020:
997:
996:
995:
983:(3–4): 314–320,
972:
962:
961:
960:
955:
924:
923:
922:
876:
873:
867:
864:
853:
850:
844:
841:
835:
832:
823:
820:
814:
811:
691:T. onomatoplokos
636:Samrukia nessovi
409:
383:
382:
374:
373:
365:
364:
356:
355:
347:
346:
338:
337:
329:
328:
322:
291:
289:
288:
262:
261:
259:
258:
257:
252:
248:
245:
244:
243:
240:
223:
222:
220:
219:
218:
213:
209:
206:
205:
204:
201:
135:Akzhar Formation
116:
46:
19:
18:
1234:
1233:
1229:
1228:
1227:
1225:
1224:
1223:
1174:Santonian Stage
1169:Campanian Stage
1144:
1143:
1118:
1116:Further reading
1082:
1070:
1068:
1066:
1018:
1016:
993:
991:
970:
958:
956:
920:
918:
884:
879:
875:Averianov, 2004
874:
870:
866:Averianov, 2016
865:
856:
851:
847:
842:
838:
833:
826:
821:
817:
812:
808:
804:
767:
762:
758:Ulmaceae indet.
650:Adocus foveatus
474:
432:early Campanian
424:Late Cretaceous
408:Boztóbe svıtasy
392:
391:
390:
389:
386:
385:
384:
376:
375:
367:
366:
358:
357:
349:
348:
340:
339:
331:
330:
286:
284:
255:
253:
249:
246:
241:
238:
236:
234:
233:
216:
214:
210:
207:
202:
199:
197:
195:
194:
115:
114:
113:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
41:
40:
38:
36:early Campanian
17:
12:
11:
5:
1232:
1222:
1221:
1216:
1211:
1206:
1201:
1196:
1191:
1186:
1181:
1176:
1171:
1166:
1161:
1156:
1142:
1141:
1138:
1135:
1132:
1129:
1125:
1122:
1117:
1114:
1113:
1112:
1095:(2): 390–400,
1075:
1064:
1038:
1023:
998:
963:
925:
902:(2): 168–175,
883:
880:
878:
877:
868:
854:
845:
836:
824:
815:
805:
803:
800:
799:
798:
796:Kugitang Svita
793:
788:
783:
778:
773:
766:
763:
761:
760:
754:
753:
749:
748:
740:
739:
735:
734:
731:Eoscapherpeton
727:
721:
720:
716:
715:
706:
705:
704:Other reptiles
701:
700:
681:
674:
667:
660:
653:
645:
644:
640:
639:
632:
624:
623:
619:
618:
613:
610:Ornithomimidae
607:
602:
597:
594:Caenagnathidae
590:
589:
585:
584:
578:
575:Titanosauridae
571:
570:
566:
565:
558:
551:
543:
542:
538:
537:
534:Ankylosauridae
530:
529:
525:
524:
517:
510:
502:
501:
497:
473:
472:Fossil content
470:
394:
393:
387:
378:
377:
369:
368:
360:
359:
351:
350:
342:
341:
333:
332:
324:
323:
317:
316:
315:
314:
311:
310:
305:
301:
300:
296:
295:
282:
278:
277:
268:
264:
263:
231:
225:
224:
192:
186:
185:
181:
180:
175:
171:
170:
165:
161:
160:
156:
155:
152:
148:
147:
142:
138:
137:
132:
128:
127:
122:
118:
117:
110:
109:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
24:
23:
15:
9:
6:
4:
3:
2:
1231:
1220:
1217:
1215:
1212:
1210:
1207:
1205:
1202:
1200:
1197:
1195:
1192:
1190:
1187:
1185:
1182:
1180:
1177:
1175:
1172:
1170:
1167:
1165:
1162:
1160:
1157:
1155:
1152:
1151:
1149:
1139:
1136:
1133:
1130:
1126:
1123:
1120:
1119:
1110:
1106:
1102:
1098:
1094:
1090:
1089:
1081:
1076:
1067:
1065:0-520-24209-2
1061:
1057:
1056:
1051:
1047:
1046:Dodson, Peter
1043:
1039:
1035:
1031:
1030:
1024:
1014:
1010:
1009:
1004:
999:
990:
986:
982:
978:
977:
969:
964:
954:
949:
945:
941:
940:
935:
933:
926:
917:
913:
909:
905:
901:
897:
896:
891:
886:
885:
872:
863:
861:
859:
849:
840:
831:
829:
819:
810:
806:
797:
794:
792:
789:
787:
784:
782:
779:
777:
774:
772:
769:
768:
759:
756:
755:
751:
750:
747:
746:
742:
741:
737:
736:
733:
732:
728:
726:
723:
722:
718:
717:
714:
712:
708:
707:
703:
702:
699:
698:
693:
692:
687:
686:
682:
680:
679:
675:
673:
672:
668:
666:
665:
661:
659:
658:
654:
652:
651:
647:
646:
642:
641:
638:
637:
633:
631:
630:
626:
625:
621:
620:
617:
614:
611:
608:
606:
603:
601:
598:
595:
592:
591:
587:
586:
582:
579:
576:
573:
572:
568:
567:
564:
563:
559:
557:
556:
552:
550:
549:
545:
544:
540:
539:
535:
532:
531:
527:
526:
523:
522:
518:
516:
515:
511:
509:
508:
504:
503:
499:
498:
496:
494:
492:
487:
484:
480:
479:
469:
465:
463:
460:
456:
452:
448:
444:
440:
435:
433:
429:
425:
421:
417:
413:
405:
401:
321:
312:
309:
308:Bostobe River
306:
302:
297:
294:
283:
279:
276:
272:
269:
265:
260:
251:38.7°N 58.0°E
232:
230:
226:
221:
212:45.9°N 64.7°E
193:
191:
187:
182:
179:
176:
172:
169:
166:
162:
157:
153:
149:
146:
143:
139:
136:
133:
129:
126:
123:
119:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
45:
37:
33:
29:
25:
20:
1092:
1086:
1069:, retrieved
1054:
1033:
1027:
1017:, retrieved
1012:
1006:
992:, retrieved
980:
974:
957:, retrieved
943:
937:
932:Khunnuchelys
931:
919:, retrieved
899:
893:
882:Bibliography
871:
848:
839:
818:
809:
745:Gobiates sp.
743:
729:
725:Anura indet.
709:
697:T. riabinini
695:
689:
683:
676:
669:
662:
655:
648:
634:
627:
560:
553:
546:
519:
512:
505:
491:Lophorhothon
489:
476:
475:
466:
462:environments
457:and fluvial-
436:
399:
397:
299:Type section
976:Palaeoworld
946:: 155–161,
541:Ornithopods
528:Ankylosaurs
254: /
215: /
190:Coordinates
1148:Categories
1071:2019-02-21
1019:2020-03-23
994:2020-03-23
959:2020-03-23
921:2020-03-23
802:References
719:Amphibians
622:Pterosaurs
483:trionychid
455:lacustrine
443:claystones
439:sandstones
416:Qaraghandy
293:Kazakhstan
271:Qaraghandy
256:38.7; 58.0
217:45.9; 64.7
1109:0022-3360
1036:: 426–436
1015:: 537–542
588:Theropods
581:Sauropoda
569:Sauropods
447:estuarine
428:Santonian
420:Qyzylorda
412:formation
304:Named for
275:Qyzylorda
178:Claystone
168:Sandstone
159:Lithology
151:Thickness
131:Underlies
32:Santonian
1052:(2004),
765:See also
184:Location
141:Overlies
42:85
904:Bibcode
711:Bishara
643:Turtles
500:Mammals
459:deltaic
451:fluvial
281:Country
242:58°00′E
239:38°42′N
203:64°42′E
200:45°54′N
164:Primary
111:↓
1107:
1062:
612:indet.
596:indet.
583:indet.
577:indet.
536:indet.
486:turtle
418:&
404:Kazakh
290:
273:&
267:Region
1128:1–156
1083:(PDF)
971:(PDF)
752:Flora
713:backa
174:Other
1105:ISSN
1060:ISBN
738:Fish
481:, a
441:and
437:The
398:The
121:Type
50:Preęž’
1097:doi
985:doi
948:doi
912:doi
430:to
414:in
1150::
1103:,
1093:75
1091:,
1085:,
1048:;
1044:;
1034:38
1032:,
1013:59
1011:,
1005:,
981:23
979:,
973:,
944:60
942:,
936:,
910:,
900:53
898:,
892:,
857:^
827:^
694:,
688:,
495:.
493:sp
464:.
449:,
406::
100:Pg
44:Ma
30::
1099::
987::
950::
914::
906::
453:-
426:(
402:(
105:N
95:K
90:J
85:T
80:P
75:C
70:D
65:S
60:O
55:ęž’
39:~
34:-
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