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permanent freshwater ponds on the Cat Ba Island, the langurs take their moisture from rainwater caught in rock pockets and contained in vegetation. Temporary surface streams form briefly during rainy season, rapidly receding into caves and subterranean passages. The soil is derived from the erosion of the native limestone bedrock, and organic detritus from the vegetation. The Cat Ba langurs live in groups, usually one male with several females and their offspring. Each group has its own territory, defended by the adult male who also initiates the location of the group. The females usually give birth to a single baby every 2–3 years, which becomes mature at 4–6 years old. Langurs have an average life expectancy of 25 years. The environment provides an arborous and terrestrial habitat for langurs as well as meeting the needs of their folivorous diet. Food mainly consists of leaves, but also fresh shoots, flowers, bark, and some fruits. The leaves make up over 70% of the langur's diet.
356:. According to the Cat Ba Langur Conservation Project, the Cat Ba langur's skin is black and the pelage color is dark brown; head and shoulder are bright golden to yellowish-white. The tail is very long (ca. 85 cm (33 in)) compared with the body size (ca. 50 cm (20 in)). Babies are colored golden-orange; the pelage starts to change its color from about the fourth month on. Males and females look alike. Two adult females captured during translocation in 2012 weighted slightly more than 9 kilograms (20 lb) each.
84:
59:
31:
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weeks. A hunter can earn up to $ 50 from a single langur, which is a lot when the average annual per capita income is less than $ 350. Since it is so hard to find the langurs, the poachers do not go out with the intent on capturing one, but often capture them by chance when hunting other animals in the area. Between 1970 and 1986, an estimated 500 to 800 langurs were killed.
242:
461:
Hunting of the Cat Ba langur used to be common. The primary reason for hunting was to supply the traditional medicine industry. Cat Ba langurs were used to make a "monkey balm" believed to help with erectile dysfunction and other health issues. Since the langurs are so few, hunting takes two to four
408:
group found elsewhere in
Vietnam. During the summer the weather is warm, and rainy with temperatures averaging 25 °C (77 °F). In the winter, it is usually cold with little rain and high temperatures averaging less than 20 °C (68 °F). Because there are no rivers and streams and no
441:
There is a strictly protected sanctuary, a peninsula on the eastern coast of Cat Ba Island, in the
National Park and supports about 40% of the population. Fixed boundaries were set with blocking buoys and prohibition signs. Another step taken was to increase the number of rangers in the area. Local
364:
species. Cat Ba langurs are diurnal creatures and travel in groups of about four to eighteen animals. They prefer the steep limestone cliffs that make up most of Cat Ba Island. Most of the places that they are found are not accessible to humans by foot. Cat Ba
Langurs spend around 66% of their time
642:
Stenke, R., Phan Duy Thuc and Nadler, T. 2007. Golden-headed Langur or Cat Ba Langur. In: Primates in Peril: The World's 25 Most
Endangered Primates 2006–2008, R. A. Mittermeier et al. (compilers), pp.14-15. Unpublished report, IUCN/SSC Primate Specialist Group (PSG), International Primatological
359:
The Cat Ba langur, which lives on Cat Ba Island in
Vietnam, is one of the 25 most endangered primates. Conservation efforts are helping to prevent this, however, and have greatly increased their population since 2003, when there were only 40. Until recently, the Cat Ba langur was not considered a
469:
Cat Ba Island is in the process of creating a booming tourist industry. They are in the process of building luxury hotels around the island, with one hotel being 17 stories tall. To accommodate all the new tourists, the town of Cat Ba is building a new road that will connect the town to a small
465:
The hunters of the langurs have been known to attack people who get in their way. As part of a conservation effort, "body guards" for the langurs have been put into place. These guards are unarmed, and on several occasions have been severely hurt by the hunters. Teeth have been knocked out and
437:
and ZSCSP, the
Zoological Society for the Conservation of Species and Populations, started the Cat Ba Langur Conservation Project. The Cat Ba langur's distribution range was declared a National Park in 1986, however that did not stop the poaching and decline of the population. About 30% of the
438:
population is located outside of the
National Park until 2006. In 2006 Cat Ba National Park was expanded to include the entire Cat Ba langur population and Special Protection Zones were established to provide the most stringent protection available under Vietnamese law.
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formation that has been invaded by the ocean. The topography is of limestone that has been worn away through erosion, which later formed ridges, towers, fissures, sinkholes and other types of landforms. The Cat Ba Langur are diurnal animals, adapted to living in
400:
habitat. The sleeping caves, ledges, and overhangs used by the langurs are thought to be used as protection from predators and extreme weather. Accessible caves were used as a hunting ground for humans to capture or kill langurs as they slept.
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citizens, especially fishermen, were informed and rangers were given permission to remove people and take away any poaching equipment they found. Any existing and potential caves and hunter trails are registered, regulated, and controlled.
599:; Wallis, J.; Rylands, A. B.; Ganzhorn, J. U.; Oates, J. F.; Williamson, E. A.; Palacios, E.; Heymann, E.W.; Kierulff, M. C. M.; Long Yongcheng; Supriatna, J.; Roos, C.; Walker, S.; Cortés-Ortiz, L.; Schwitzer, C., eds. (November 2009).
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village on the northern edge of the island where a ferry will be to take tourists to another popular destination: Ha Long Bay. The road runs just along a border of the park which may attract more hunters to the area.
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The greatest populations of the Cat Ba Langur are expected to recover with the appropriate conservation of the limestone habitats. There are currently 3 Cat Ba
Langurs held at the Endangered Primate Center in
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445:
At the end of 2001, there was no more hunting of the Cat Ba Langur. Since the beginning of the conservation efforts, nine langurs have been born and only three have died of natural causes.
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resting and the rest moving, foraging, and socializing, with the distribution changing between summer and winter. They eat less and rest more in the winter and the opposite in the summer.
608:. Illustrated by S.D. Nash. Arlington, VA.: IUCN/SSC Primate Specialist Group (PSG), International Primatological Society (IPS), and Conservation International (CI). pp. 1–92.
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Cat Ba island is the largest island within the Cat Ba archipelago, with 366 smaller islands and tidally exposed rocks surrounding it. As all members of the
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and is assumed to have declined by 80% over the last three generations. It is estimated that there are less than 70 of these langurs left in the world.
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Despite its very small distribution and population, research indicates that this species persisted on Cat Ba Island since at least the latest
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The Cat Ba langur lives in a moist tropical rain forest on limestone hills, a type of habitat used by 6–7 other genera of the
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in the world, and possibly the rarest primate in Asia, with population size estimated at less than 70 individuals.
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several have been stabbed. Because of the fear of being attacked, the guards do not want to do their job anymore.
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959:
https://web.archive.org/web/20110728055728/http://www.stiftung-artenschutz.org/eng/set.html?projects%2Fgold.html
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777:"Behavioral Observations and Notes on the Vertical Ranging Pattern of the Critically Endangered Cat Ba Langur (
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63:
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Bradon-Jones, D. 1995. A revision of the Asian pied leaf monkeys (Mammalia: Cercopithecidae: Superspecies
416:, about 12,000 years ago, and never reached a population over 4,000 - 5,000 individuals even at its peak.
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Schneider, Isabell; Tielen, Inge H.M.; Rode, Johanna; Levelink, Pieter; Schrudde, Daniela (2010-12-01).
231:
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Ben, S (5 March 2003). "Experts Say Viet
Tourism Overtaking Conservation, as Monkey Nears Extinction".
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1989:
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391:, specifically in Cat Ba Island, hence its common name "Cat Ba Langur". This landscape is known as a
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835:"Landscapes and Ecosystems of Tropical Limestone: Case Study of the Cat Ba Islands, Vietnam"
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954:
http://floydssecrets.blogspot.com/2007/10/golden-headed-langurvietnam.html
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Species endangered by human consumption for medicinal or magical purposes
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503:(3rd ed.). Baltimore: Johns Hopkins University Press. p. 177.
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are overall blackish, but the crown, cheeks and neck are yellowish in
1681:
1014:
397:
95:
2402:
602:
Primates in Peril: The World's 25 Most
Endangered Primates 2008–2010
2425:
1033:
751:"The Cat Ba Langur: a primate walks the razor's edge of extinction"
670:
281:
115:
934:
671:"Primates-SG - Vietnamese Journal of Primatology Volume 1 Issue 1"
643:
Society (IPS), and Conservation International (CI), Arlington, VA.
30:
1045:
500:
Mammal Species of the World: A Taxonomic and Geographic Reference
293:
289:
135:
2509:
595:
2468:
1039:
1027:
543:
Rawson, B.M.; Leonard, N.; Covert, H. & Nadler, T. (2020).
278:
155:
125:
105:
2647:
392:
321:
1886:
774:
845:(1). Translated by Van; Duc, Thanh; Van, Huy Dinh: 23–36.
931:
Cat Ba Langur Conservation Project. Accessed 2008-07-15.
964:
http://www.ape-rescue.co.uk/golden-headed-langur.html
360:
species but rather a subspecies of one of two other
875:
304:The Cat Ba langur was considered a subspecies of
2677:
568:10.2305/IUCN.UK.2020-2.RLTS.T39871A17959804.en
424:The Cat Ba langur is considered to be one of "
2083:
991:
656:), with the description of a new subspecies.
591:
589:
587:
585:
882:. Charlestown, Rhode Island: Pogonia Press.
646:
2686:IUCN Red List critically endangered species
832:
2090:
2076:
998:
984:
878:The Pictorial Guide to the Living Primates
779:Trachypithecus Poliocephalus Poliocephalus
582:
240:
57:
29:
858:
796:
566:
873:
324:was formerly considered a subspecies of
2099:The World's 25 Most Endangered Primates
426:The World's 25 Most Endangered Primates
249:Cat Ba langur range (includes range of
2726:Taxa named by Édouard Louis Trouessart
2678:
538:
536:
534:
532:
530:
528:
489:
485:
483:
456:
383:monkey is found in limestone forests.
2407:
2406:
2071:
979:
948:http://www.catbalangur.org/Langur.htm
905:
903:
901:
899:
828:
826:
824:
839:Journal of Ecology and Field Biology
722:
720:
718:
2711:Critically endangered fauna of Asia
1963:Black-and-white snub-nosed monkey (
909:
554:IUCN Red List of Threatened Species
525:
480:
13:
896:
821:
14:
2737:
922:
715:
82:
1699:Black-crested Sumatran langur (
935:http://ippl.org/2001-langur.php
867:
768:
419:
743:
687:
663:
636:
331:
1:
1753:East Sumatran banded langur (
1393:Northern plains gray langur (
473:
2474:trachypithecus-poliocephalus
2461:Trachypithecus_poliocephalus
2447:Trachypithecus poliocephalus
2417:Trachypithecus poliocephalus
2270:Trachypithecus poliocephalus
1164:Miss Waldron's red colobus (
729:Trachypithecus poliocephalus
697:Trachypithecus poliocephalus
547:Trachypithecus poliocephalus
379:species group, this social,
266:Trachypithecus poliocephalus
224:Trachypithecus poliocephalus
7:
2215:Nigeria-Cameroon chimpanzee
2114:Madame Berthe's mouse lemur
1990:Myanmar snub-nosed monkey (
755:Mongabay Environmental News
740:ARKive. Accessed 2008-07-15
387:, this langur resides near
299:
10:
2742:
2696:Primates of Southeast Asia
2381:Ateles fusciceps fusciceps
2377:Brown-headed spider monkey
1981:Tonkin snub-nosed monkey (
1954:Golden snub-nosed monkey (
1744:Robinson's banded langur (
1667:Indochinese black langur (
1514:Selangor silvered langur (
1402:Black-footed gray langur (
1299:Ulindi River red colobus (
860:10.5141/JEFB.2010.33.1.023
833:Quan Nguyen (March 2010).
368:
348:, while they are white in
2716:Mammals described in 1911
2415:
2319:
2228:
2157:
2128:Lepilemur septentrionalis
2106:
2054:
2026:
2002:
1934:
1884:
1877:
1679:
1541:Indochinese grey langur (
1441:
1346:
1339:
1311:
1245:Niger Delta red colobus (
1135:
1067:
1060:
1022:
1006:Extant species of family
292:. It is among the rarest
248:
239:
220:
213:
79:Scientific classification
77:
55:
46:
37:
28:
23:
2706:Endemic fauna of Vietnam
2387:Geoffroy's spider monkey
2296:Skywalker hoolock gibbon
1972:Gray snub-nosed monkey (
1735:Raffles' banded langur (
1708:Black-and-white langur (
1281:Oustalet's red colobus (
1236:Tana River red colobus (
1077:(Black and white colobi)
787:(Submitted manuscript).
497:; Reeder, D. M. (eds.).
451:Cuc Phuong National Park
376:Trachypithecus francoisi
192:Trachypithecus francoisi
2219:Pan troglodytes ellioti
2205:Niger Delta red colobus
2189:Erythrocebus baumstarki
2179:Cercocebus chrysogaster
2175:Golden-bellied mangabey
2124:Sahafary sportive lemur
1825:Sabah grizzled langur (
1717:Black Sumatran langur (
1254:Bouvier's red colobus (
1191:Thollon's red colobus (
944:Retrieved June 30, 2008
703:www.mammaldiversity.org
2290:Rhinopithecus brelichi
2286:Gray snub-nosed monkey
2276:Raffles' banded langur
1861:Natuna Island surili (
1780:White-fronted surili (
1771:White-thighed surili (
1550:Phayre's leaf monkey (
1227:Zanzibar red colobus (
1218:Udzungwa red colobus (
1182:Preuss's red colobus (
352:, as suggested by its
2635:Paleobiology Database
2361:Plecturocebus grovesi
2327:Buffy-headed marmoset
2260:Semnopithecus vetulus
2199:Cercopithecus roloway
2185:Southern patas monkey
2169:Paragalogo rondoensis
2148:Propithecus coquereli
2134:Blue-eyed black lemur
1640:White-headed langur (
1595:Gee's golden langur (
1586:Shortridge's langur (
1420:Purple-faced langur (
1375:Kashmir gray langur (
1263:Semliki red colobus (
1209:Ugandan red colobus (
1155:Western red colobus (
912:San Jose Mercury News
727:White-headed langur (
654:Semnopithecus auratus
561:: e.T39871A17959804.
385:Critically endangered
275:critically endangered
269:), also known as the
206:T. poliocephalus
64:Critically Endangered
2331:Callithrix flaviceps
2240:Nycticebus javanicus
1944:(Snub-nosed monkeys)
1913:Black-shanked douc (
1834:Maroon leaf monkey (
1411:Tufted gray langur (
1290:Lomami red colobus (
1272:Lang's red colobus (
950:Accessed 2011-07-23.
798:10.1896/052.025.0104
785:Primate Conservation
271:golden-headed langur
40:Cát Bà National Park
2367:Brown howler monkey
2357:Groves' titi monkey
2351:Cebus aequatorialis
2347:Ecuadorian capuchin
2310:Pongo tapanuliensis
2280:Presbytis femoralis
2256:Purple-faced langur
2250:Tarsius sangirensis
2209:Piliocolobus epieni
2038:Pig-tailed langur (
1922:Gray-shanked douc (
1658:Delacour's langur (
1532:Dusky leaf monkey (
1505:Tenasserim lutung (
1469:West Javan langur (
1460:East Javan langur (
1384:Tarai gray langur (
1366:Nepal gray langur (
1200:Foa's red colobus (
1173:Pennant's colobus (
1010:(Old World monkeys)
851:2010JEcEn..33...23V
699:(Trouessart, 1911)"
658:Raffles Bull. Zool.
457:Threats to survival
314:white-headed langur
251:white-headed langur
49:Conservation status
2701:Mammals of Vietnam
2306:Tapanuli orangutan
2165:Rondo dwarf galago
2138:Eulemur flavifrons
2118:Microcebus berthae
2014:Proboscis monkey (
1904:Red-shanked douc (
1613:François' langur (
1487:Germain's langur (
969:2017-11-29 at the
940:2008-10-06 at the
929:The Cat Ba Langur.
736:2008-07-24 at the
675:www.primate-sg.org
597:Mittermeier, R. A.
433:In November 2000,
312:) until 1995. The
38:Cat Ba langurs at
2673:
2672:
2622:Open Tree of Life
2409:Taxon identifiers
2400:
2399:
2144:Coquerel's sifaka
2065:
2064:
2050:
2049:
1947:
1897:
1873:
1872:
1843:Mentawai langur (
1816:Miller's langur (
1798:Thomas's langur (
1692:
1496:Annamese langur (
1359:
1335:
1334:
1148:
1123:Mantled guereza (
1080:
1011:
889:978-0-9648825-0-8
874:Rowe, N. (1996).
615:978-1-934151-34-1
257:
256:
72:
16:Species of monkey
2733:
2666:
2665:
2656:
2655:
2643:
2642:
2630:
2629:
2617:
2616:
2604:
2603:
2591:
2590:
2578:
2577:
2565:
2564:
2552:
2551:
2539:
2538:
2526:
2525:
2513:
2512:
2500:
2499:
2487:
2486:
2477:
2476:
2464:
2463:
2451:
2450:
2449:
2436:
2435:
2434:
2404:
2403:
2391:Ateles geoffroyi
2371:Alouatta guariba
2337:Ka'apor capuchin
2300:Hoolock tianxing
2236:Javan slow loris
2092:
2085:
2078:
2069:
2068:
1946:
1945:
1941:
1896:
1895:
1891:
1882:
1881:
1852:Siberut langur (
1762:Sarawak surili (
1726:Mitered langur (
1691:
1690:
1686:
1649:Laotian langur (
1642:T. leucocephalus
1633:T. poliocephalus
1516:T. selangorensis
1478:Silvery lutung (
1429:Nilgiri langur (
1358:
1357:
1353:
1344:
1343:
1147:
1146:
1142:
1114:Ursine colobus (
1096:Angola colobus (
1079:
1078:
1074:
1065:
1064:
1009:
1008:Cercopithecidae
1000:
993:
986:
977:
976:
916:
915:
907:
894:
893:
881:
871:
865:
864:
862:
830:
819:
818:
800:
772:
766:
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763:
762:
747:
741:
724:
713:
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709:
691:
685:
684:
682:
681:
667:
661:
650:
644:
640:
634:
633:
631:
630:
624:
618:. Archived from
607:
593:
580:
579:
577:
575:
570:
540:
523:
522:
487:
350:T. leucocephalus
346:T. poliocephalus
342:T. leucocephalus
338:T. poliocephalus
326:T. poliocephalus
318:T. leucocephalus
306:François' langur
244:
226:
87:
86:
66:
61:
60:
33:
21:
20:
2741:
2740:
2736:
2735:
2734:
2732:
2731:
2730:
2676:
2675:
2674:
2669:
2661:
2659:
2651:
2646:
2638:
2633:
2625:
2620:
2612:
2609:Observation.org
2607:
2599:
2594:
2586:
2581:
2573:
2568:
2560:
2555:
2547:
2542:
2534:
2529:
2521:
2516:
2508:
2503:
2495:
2490:
2482:
2480:
2472:
2467:
2459:
2454:
2445:
2444:
2439:
2430:
2429:
2424:
2411:
2401:
2396:
2315:
2246:Sangihe tarsier
2224:
2153:
2102:
2096:
2066:
2061:
2046:
2022:
1998:
1943:
1942:
1940:
1930:
1893:
1892:
1890:
1878:Odd-nosed group
1869:
1807:Hose's langur (
1688:
1687:
1685:
1675:
1631:Cat Ba langur (
1622:Hatinh langur (
1577:Capped langur (
1437:
1355:
1354:
1352:
1331:
1323:Olive colobus (
1307:
1265:P. semlikiensis
1238:P. rufomitratus
1211:P. tephrosceles
1144:
1143:
1141:
1131:
1087:Black colobus (
1076:
1075:
1073:
1056:
1018:
1004:
971:Wayback Machine
942:Wayback Machine
925:
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166:Cercopithecidae
81:
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62:
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17:
12:
11:
5:
2739:
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2698:
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2691:Trachypithecus
2688:
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2222:
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2195:Roloway monkey
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2159:
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1789:Javan surili (
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1764:P. chrysomelas
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1624:T. hatinhensis
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1592:
1588:T. shortridgei
1583:
1566:
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1556:
1547:
1543:T. crepusculus
1538:
1521:
1520:
1511:
1502:
1493:
1484:
1475:
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1449:
1447:
1444:Trachypithecus
1439:
1438:
1436:
1435:
1426:
1417:
1408:
1399:
1390:
1381:
1372:
1368:S. schistaceus
1362:
1360:
1356:(Gray langurs)
1341:
1337:
1336:
1333:
1332:
1330:
1329:
1319:
1317:
1309:
1308:
1306:
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1292:P. parmentieri
1287:
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923:External links
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791:(1): 111–117.
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2341:Cebus kaapori
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2019:
2017:
2012:
2011:
2009:
2007:
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1995:
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1988:
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1984:
1979:
1977:
1975:
1970:
1968:
1966:
1961:
1959:
1957:
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1937:Rhinopithecus
1933:
1927:
1925:
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1918:
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1911:
1909:
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1889:
1888:
1883:
1880:
1876:
1866:
1864:
1859:
1857:
1855:
1850:
1848:
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1845:P. potenziani
1841:
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1701:P. melalophos
1697:
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1559:Popa langur (
1557:
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1529:
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1523:
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1440:
1434:
1432:
1427:
1425:
1423:
1418:
1416:
1414:
1409:
1407:
1405:
1404:S. hypoleucos
1400:
1398:
1396:
1391:
1389:
1387:
1382:
1380:
1378:
1373:
1371:
1369:
1364:
1363:
1361:
1351:
1350:
1349:Semnopithecus
1345:
1342:
1338:
1328:
1326:
1321:
1320:
1318:
1316:
1315:
1310:
1304:
1302:
1301:P. lulindicus
1297:
1295:
1293:
1288:
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1279:
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1220:P. gordonorum
1216:
1214:
1212:
1207:
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1203:
1198:
1196:
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1189:
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1153:
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1140:
1139:
1134:
1128:
1126:
1121:
1119:
1117:
1116:C. vellerosus
1112:
1110:
1108:
1103:
1101:
1099:
1098:C. angolensis
1094:
1092:
1090:
1085:
1084:
1082:
1072:
1071:
1066:
1063:
1061:African group
1059:
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1041:
1037:
1035:
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982:
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836:
829:
827:
825:
816:
812:
808:
804:
799:
794:
790:
786:
782:
781:) in Vietnam"
780:
771:
756:
752:
746:
739:
735:
732:
730:
723:
721:
719:
704:
700:
698:
690:
676:
672:
666:
659:
655:
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625:on 2011-07-23
621:
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531:
529:
520:
516:
512:
510:0-801-88221-4
506:
502:
501:
496:
495:Wilson, D. E.
492:
491:Groves, C. P.
486:
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479:
471:
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454:
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378:
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366:
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355:
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347:
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329:
328:until 2007.
327:
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315:
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307:
297:
295:
291:
287:
286:Cát Bà Island
283:
280:
276:
272:
268:
267:
262:
261:Cat Ba langur
252:
247:
243:
238:
233:
227:
225:
219:
216:
215:Binomial name
212:
208:
207:
202:
199:
198:
195:
193:
189:
186:
185:Species group
183:
182:
179:
178:
174:
171:
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167:
164:
161:
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157:
154:
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150:
147:
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137:
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127:
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117:
114:
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110:
107:
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100:
97:
94:
91:
90:
85:
80:
76:
70:
65:
54:
50:
45:
41:
36:
32:
27:
22:
19:
2416:
2390:
2380:
2370:
2360:
2350:
2340:
2330:
2309:
2299:
2289:
2279:
2269:
2265:
2259:
2249:
2239:
2218:
2208:
2198:
2188:
2178:
2168:
2147:
2137:
2127:
2117:
2056:
2039:
2027:
2015:
2003:
1991:
1983:R. avunculus
1982:
1973:
1964:
1956:R. roxellana
1955:
1935:
1923:
1914:
1905:
1885:
1862:
1853:
1844:
1836:P. rubicunda
1835:
1826:
1817:
1808:
1799:
1790:
1781:
1773:P. siamensis
1772:
1763:
1754:
1746:P. robinsoni
1745:
1737:P. femoralis
1736:
1727:
1719:P. sumatrana
1718:
1709:
1700:
1680:
1668:
1660:T. delacouri
1659:
1650:
1641:
1632:
1630:
1623:
1615:T. francoisi
1614:
1607:T. francoisi
1606:
1605:
1596:
1587:
1578:
1570:
1569:
1560:
1551:
1542:
1533:
1525:
1524:
1515:
1506:
1498:T. margarita
1497:
1488:
1480:T. cristatus
1479:
1471:T. mauritius
1470:
1461:
1454:T. cristatus
1453:
1452:
1442:
1430:
1421:
1412:
1403:
1394:
1385:
1376:
1367:
1347:
1340:Langur group
1324:
1312:
1300:
1291:
1283:P. oustaleti
1282:
1273:
1264:
1255:
1246:
1237:
1228:
1219:
1210:
1201:
1192:
1183:
1175:P. pennantii
1174:
1166:P. waldronae
1165:
1156:
1145:(Red colobi)
1138:Piliocolobus
1136:
1124:
1115:
1107:C. polykomos
1106:
1097:
1088:
1068:
911:
877:
869:
842:
838:
788:
784:
778:
770:
759:. Retrieved
757:. 2015-08-31
754:
745:
728:
706:. Retrieved
702:
696:
689:
678:. Retrieved
674:
665:
657:
653:
648:
638:
627:. Retrieved
620:the original
601:
572:. Retrieved
558:
552:
546:
499:
468:
464:
460:
447:
444:
440:
432:
428:
423:
420:Conservation
411:
406:T. francoisi
405:
403:
374:
372:
361:
358:
349:
345:
341:
337:
335:
325:
317:
310:T. francoisi
309:
303:
270:
265:
264:
260:
258:
223:
221:
205:
204:
191:
184:
176:
152:Infraorder:
18:
2531:iNaturalist
2441:Wikispecies
2101:(2022–2023)
2040:S. concolor
2016:N. larvatus
1992:R. strykeri
1974:R. brelichi
1915:P. nigripes
1818:P. canicrus
1782:P. frontata
1579:T. pileatus
1571:T. pileatus
1534:T. obscurus
1526:T. obscurus
1489:T. germaini
1395:S. entellus
1256:P. bouvieri
1193:P. tholloni
1013:(subfamily
453:, Vietnam.
435:Münster Zoo
414:Pleistocene
389:Ha Long Bay
332:Description
277:species of
156:Simiiformes
2680:Categories
2107:Madagascar
1924:P. cinerea
1906:P. nemaeus
1863:P. natunae
1800:P. thomasi
1755:P. percura
1728:P. mitrata
1710:P. bicolor
1552:T. phayrei
1462:T. auratus
1422:S. vetulus
1314:Procolobus
1184:P. preussi
1125:C. guereza
1089:C. satanas
1052:Haplorhini
1050:Suborder:
761:2015-10-29
708:2021-11-18
680:2021-11-18
629:2010-08-06
474:References
232:Trouessart
146:Haplorhini
142:Suborder:
42:, Vietnam
1887:Pygathrix
1854:P. siberu
1827:P. sabana
1791:P. comata
1689:(Surilis)
1682:Presbytis
1669:T. ebenus
1651:T. laotum
1507:T. barbei
1431:S. johnii
1386:S. hector
1247:P. epieni
1229:P. kirkii
1157:P. badius
1026:Kingdom:
1015:Colobinae
807:0898-6207
398:limestone
200:Species:
102:Kingdom:
96:Eukaryota
2648:Species+
2588:12100742
2481:BioLib:
2426:Wikidata
2058:Category
1965:R. bieti
1809:P. hosei
1413:S. priam
1325:P. verus
1274:P. langi
1046:Primates
1040:Mammalia
1034:Chordata
1032:Phylum:
1028:Animalia
967:Archived
938:Archived
815:55362144
734:Archived
660:43: 3-43
519:62265494
493:(2005).
300:Taxonomy
294:primates
162:Family:
136:Primates
126:Mammalia
116:Chordata
112:Phylum:
106:Animalia
92:Domain:
69:IUCN 3.1
2663:7139845
2575:1000713
2523:7262019
2510:4453635
2432:Q929055
2005:Nasalis
1894:(Doucs)
1597:T. geei
1561:T. popa
1377:S. ajax
1202:P. foai
1070:Colobus
1044:Order:
1038:Class:
847:Bibcode
574:19 June
381:diurnal
369:Habitat
290:Vietnam
282:endemic
273:, is a
234:, 1911)
172:Genus:
132:Order:
122:Class:
67: (
2660:uBio:
2640:238690
2627:250658
2614:201366
2601:465719
2549:944702
2536:879166
2484:302712
2469:ARKive
2158:Africa
2029:Simias
886:
813:
805:
612:
517:
507:
279:langur
2562:22045
2497:57SDP
1609:group
1573:group
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