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Commensalism

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22: 137: 557: 478: 2045: 1762: 208:, well before the cultivation of crops or the domestication of other animals. The dog is often hypothesised to be a classic example of a domestic animal that likely traveled a commensal pathway into domestication. Archaeological evidence, such as the Bonn-Oberkassel dog dating to ~14,000BP, supports the hypothesis that dog domestication preceded the emergence of agriculture and began close to the 233:
population that remained in a boreal coniferous forest. Although these two populations spend a period of the year in the same place, and though there was evidence of gene flow between them, the difference in prey–habitat specialization has been sufficient to maintain genetic and even coloration divergence.
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early hunters, assisted in the capture of prey, or provided defense from large competing predators at kills. However, the extent to which proto-domestic wolves could have become dependent on this way of life prior to domestication and without human provisioning is unclear and highly debated. In contrast,
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with unique mitochondrial signatures. The skull shape, tooth wear, and isotopic signatures suggested these remains were derived from a population of specialist megafauna hunters and scavengers that became extinct while less specialized wolf ecotypes survived. Analogous to the modern wolf ecotype that
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The commensal pathway was traveled by animals that fed on refuse around human habitats or by animals that preyed on other animals drawn to human camps. Those animals established a commensal relationship with humans in which the animals benefited but the humans received little benefit or harm. Those
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Historically, commensalism has been recognized as the usual type of association between brittle stars and octocorals. In this association, the ophiurans benefit directly by being elevated through facilitating their feeding by suspension, while the octocorals do not seem to benefit or be harmed by
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A mitochondrial, microsatellite, and Y-chromosome assessment of two wolf populations in North America combined with satellite telemetry data revealed significant genetic and morphological differences between one population that migrated with and preyed upon caribou and another territorial ecotype
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may have become fully dependent on a commensal lifestyle before being domesticated by preying on other commensal animals, such as rats and mice, without any human provisioning. Debate over the extent to which some wolves were commensal with humans prior to domestication stems from debate over the
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The wolves more likely drawn to human camps were the less-aggressive, subdominant pack members with lowered flight response, higher stress thresholds, and less wary around humans, and therefore better candidates for domestication. Proto-dogs might have taken advantage of carcasses left on site by
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are "nutritional pirates" that may intercept substantial amounts of nutrients that would otherwise go to the host plant. Large numbers of epiphytes can also cause tree limbs to break or shade the host plant and reduce its rate of photosynthesis. Similarly, phoretic mites may hinder their host by
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The earliest sign of domestication in dogs was the neotenization of skull morphology and the shortening of snout length that results in tooth crowding, reduction in tooth size, and a reduction in the number of teeth, which has been attributed to the strong selection for reduced aggression. This
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population to a domestic one could only have taken place after the animals had progressed from anthropophily to habituation, to commensalism and partnership, at which point the establishment of a reciprocal relationship between animal and human would have laid the foundation for domestication,
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Recent studies in the Gulf of Mexico have suggested that there are actually some benefits to the octocorals, such as receiving a cleaning action by the brittle star as it slowly moves around the coral. In some cases, a close relationship occurs between cohabiting species, with the interaction
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is capable of living under considerable environmental stress, and thus is capable of colonising the upper gastrointestinal tract where relatively few examples of the body's gut flora can survive due to highly acidic or alkaline conditions produced by gastric acid and digestive juices. While
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Commensalistic relationships between microorganisms include situations in which the waste product of one microorganism is a substrate for another species. One good example is nitrification-the oxidation of ammonium ion to nitrate. Nitrification occurs in two steps: first, bacteria such as
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has evolved to track and prey upon caribou, a Pleistocene wolf population could have begun following mobile hunter-gatherers, thus slowly acquiring genetic and phenotypic differences that would have allowed them to more successfully adapt to the human habitat.
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The commensal (the species that benefits from the association) may obtain nutrients, shelter, support, or locomotion from the host species, which is substantially unaffected. The commensal relation is often between a larger host and a smaller commensal; the
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formation provides another example. The colonization of a newly exposed surface by one type of microorganism (an initial colonizer) makes it possible for other microorganisms to attach to the microbially modified surface.
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stimulates the proliferation of more acid-tolerant microorganisms, which may be only a minor part of the microbial community at neutral pH. A good example is the succession of microorganisms during milk spoilage.
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Williams, E. H.; Mignucci-Giannoni, A. A.; Bunkley-Williams, L.; Bonde, R. K.; Self-Sullivan, C.; Preen, A.; Cockcroft, V. G. (2003). "Echeneid-sirenian associations, with information on sharksucker diet".
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Thalmann O, Shapiro B, Cui P, Schuenemann VJ, Sawyer SK, Greenfield DL, et al. (November 2013). "Complete mitochondrial genomes of ancient canids suggest a European origin of domestic dogs".
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Some biologists argue that any close interaction between two organisms is unlikely to be completely neutral for either party, and that relationships identified as commensal are likely
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Commensalistic associations also occur when one microbial group modifies the environment to make it better suited for another organism. The synthesis of acidic waste products during
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animals that were most capable of taking advantage of the resources associated with human camps would have been the 'tamer' individuals: less aggressive, with shorter
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Leonard JA, Vilà C, Fox-Dobbs K, Koch PL, Wayne RK, Van Valkenburgh B (July 2007). "Megafaunal extinctions and the disappearance of a specialized wolf ecomorph".
2089: 1905:"Preadaptive plateau in Rhabditida (Nematoda) allowed the repeated evolution of zooparasites, with an outlook on evolution of life cycles within Spiroascarida" 304:, a common bacterial species, is known best for its numerous pathogenic strains that can cause numerous illnesses and conditions. However, many strains of 91:
The word "commensalism" is derived from the word "commensal", meaning "eating at the same table" in human social interaction, which in turn comes through
1889: 3020: 2273: 1931:"Relationships between phoretic mites and their carrier, the banana pseudostem weevil Odoiporus longicollis Oliver (Coleoptera: Curculionidae)" 229:
process may have begun during the initial commensal stage of dog domestication, even before humans began to be active partners in the process.
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making flight more difficult, which may affect its aerial hunting ability or cause it to expend extra energy while carrying these passengers.
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Metabiosis is a more indirect dependency, in which one organism creates or prepares a suitable environment for a second. Examples include
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are specially adapted to attach themselves to larger fish (or other animals, in this case a sea turtle) that provide locomotion and food.
3234: 766:"Large-scale assessment of commensalistic-mutualistic associations between African birds and herbivorous mammals using internet photos" 634:
is one animal associating with another until the latter dies, then the former feeds on the corpse of the latter. Examples include some
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organism is unmodified, whereas the commensal species may show great structural adaptation consistent with its habits, as in the
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also benefits from the variable ambient conditions created by the body's mucous membranes, and as such can be found in the
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distances. Later, these animals developed closer social or economic bonds with humans and led to a domestic relationship.
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normally produces no symptoms, in individuals who are immunocompromised or suffering from existing conditions such as
2526: 1606: 1681:"Mutualistic symbiosis with ophiuroids limited the impact of the Deepwater Horizon oil spill on deep-sea octocorals" 3825: 2987: 3269: 2982: 2849: 2561: 3654: 3527: 668:– long-term interactions between different biological species, which can be mutualistic, commensal or parasitic 3700: 3299: 3254: 3006: 2259: 2176: 2049: 2786: 1506: 3132: 2156: 2151: 2116: 3489: 2380: 821: 180:
including captivity and then human-controlled breeding. From this perspective, animal domestication is a
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Like all ecological interactions, commensalisms vary in strength and duration from intimate, long-lived
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gain benefits while those of the other species neither benefit nor are harmed. This is in contrast with
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are metabiotic commensals, and are present on roughly 20 to 30% of the human population as part of the
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feed on the leftovers of their hosts' meals. Numerous birds perch on bodies of large mammal
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process in which a population responds to selective pressure while adapting to a novel
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was the first domesticated animal, and was domesticated and widely established across
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A different study has identified the remains of a population of extinct Pleistocene
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level of human intentionality in the domestication process, which remains untested.
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In deep-sea, benthic environments there is an associative relationship between
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live on and in the human body as part of its natural flora. The fungal genus
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oxidize ammonium to nitrite; and second, nitrite is oxidized to nitrate by
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that benefit at least one of the participants and cause harm to neither.
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is the use of a second organism for permanent housing. Examples are
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is one animal attached to another exclusively for transport, mainly
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and other fishes. Remoras feed on their hosts' fecal matter, while
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Text was copied from this source, which is available under a
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spp. because they use nitrite to obtain energy for growth.
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Whether the relationship between humans and some types of
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in a subtle way that has not been detected. For example,
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that includes another species with evolving behaviors.
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Creative Commons Attribution 4.0 International License
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or feed on the insects turned up by grazing mammals.
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Annual Review of Ecology, Evolution, and Systematics
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to brief, weak interactions through intermediaries.
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(30 December 2010). 1847:European Journal of Soil Biology 1760: 1414:10.1111/j.1365-294x.2007.03458.x 743:10.1046/j.1095-8649.2003.00236.x 604:shells to protect their bodies. 566:growing on an oak tree in Mexico 212:when hunter-gatherers preyed on 3270:Relative abundance distribution 2983:Plant defense against herbivory 2850:Competitive exclusion principle 2562:Mesopredator release hypothesis 1977: 1922: 1896: 1873: 1772: 1721: 1672: 1623: 1574: 697:Sociobiology: The New Synthesis 674:– species commensal with humans 607: 2855:Consumer–resource interactions 1688:Marine Ecology Progress Series 1636:Marine Ecology Progress Series 1505:Wolpert S (14 November 2013). 854: 839:Animal parasites and messmates 829: 808: 713: 551: 1: 3701:Biological data visualization 3528:Environmental niche modelling 3255:Population viability analysis 1955:10.1080/01647954.2019.1656286 1867:10.1016/S1164-5563(01)01108-6 1538:Clinical Microbiology Reviews 1145:10.1126/science.276.5319.1687 678: 587: 3186:Density-dependent inhibition 1309:10.1016/0305-4403(92)90049-9 951:10.3998/jar.0521004.0068.201 626: 440: 409:Octocorals and Brittle Stars 86: 7: 3655:Liebig's law of the minimum 3490:Resource selection function 2381:Metabolic theory of ecology 1734:Frontiers in Marine Science 822:Online Etymology Dictionary 641: 427:Metallogorgia melanotrichos 382:spp. and similar bacteria. 10: 3847: 3555:Niche apportionment models 3275:Relative species abundance 2479:Primary nutritional groups 2376:List of feeding behaviours 2167:Behavior-altering parasite 1880:Hogan, C. Michael (2011). 1587:Advances in Marine Biology 1255:10.1016/j.tree.2016.05.001 500: 254: 108:, meaning "together", and 43:) in which members of one 3804: 3736:Ecosystem based fisheries 3678: 3578: 3503: 3376: 3348:Interspecific competition 3313: 3240:Minimum viable population 3173: 3098:Maximum sustainable yield 3083:Intraspecific competition 3078:Effective population size 3041: 2958:Anti-predator adaptations 2943: 2822: 2749: 2706: 2628: 2595: 2492: 2469:Photosynthetic efficiency 2404: 2298: 2177:Host–parasite coevolution 2112: 1990:Journal of Insect Science 1781:Biology of the Bromeliads 1747:10.3389/fmars.2021.735039 1466:10.1016/j.cub.2007.05.072 1108:10.1016/j.jas.2018.01.004 867:Evolutionary Applications 487:Pseudolynchia canariensis 161:cosmopolitan distribution 126:Pierre-Joseph van Beneden 3726:Ecological stoichiometry 3691:Alternative stable state 1522:Statement by Wayne, R.K. 848:10.5962/bhl.title.132633 693:"Ch.17-Social Symbiosis" 512:, examples of which are 472: 324:, as well as inside the 3826:Biological interactions 3570:Ontogenetic niche shift 3433:Ideal free distribution 3343:Ecological facilitation 3093:Malthusian growth model 3063:Consumer-resource model 2920:Paradox of the plankton 2885:Energy systems language 2605:Chemoorganoheterotrophy 2572:Optimal foraging theory 2547:Heterotrophic nutrition 2102:biological interactions 2061:Encyclopædia Britannica 1362:Fieldiana: Anthropology 1191:10.1126/science.1243650 1047:10.1073/pnas.1203005109 723:Journal of Fish Biology 191: 149:Columba livia domestica 3716:Ecological forecasting 3660:Marginal value theorem 3458:Landscape epidemiology 3393:Cross-boundary subsidy 3328:Biological interaction 2678:Microbial intelligence 2366:Green world hypothesis 567: 490: 204:before the end of the 164: 71:that ride attached to 37:biological interaction 29: 3721:Ecological humanities 3620:Ecological energetics 3565:Niche differentiation 3428:Habitat fragmentation 3196:Ecological extinction 3143:Small population size 2895:Feed conversion ratio 2875:Ecological succession 2807:San Francisco Estuary 2721:Ecological efficiency 2663:Microbial cooperation 2002:10.1673/031.012.12801 1886:Encyclopedia of Earth 559: 480: 453:is still unanswered. 301:Staphylococcus aureus 296:grow out of control. 288:, a condition called 139: 24: 3746:Evolutionary ecology 3711:Ecological footprint 3706:Ecological economics 3630:Ecological threshold 3625:Ecological indicator 3495:Source–sink dynamics 3448:Land change modeling 3443:Insular biogeography 3295:Species distribution 3034:Modelling ecosystems 2693:Microbial metabolism 2532:Intraguild predation 2321:Biogeochemical cycle 2287:Modelling ecosystems 2192:Parasitic castration 2132:Deception in animals 2052:at Wikimedia Commons 1845:) in birdsˈ nests". 1550:10.1128/CMR.10.3.505 621:species interactions 210:Last Glacial Maximum 3796:Theoretical ecology 3771:Natural environment 3635:Ecosystem diversity 3605:Ecological collapse 3595:Bateman's principle 3550:Limiting similarity 3463:Landscape limnology 3285:Species homogeneity 3123:Population modeling 3118:Population dynamics 2935:Trophic state index 1947:2019IJAca..45..361B 1859:2001EJSB...37..321T 1812:1991Biotr..23..204D 1779:Benzing DH (1980). 1700:2016MEPS..549...89G 1648:1990MEPS...64..113F 1458:2007CBio...17.1146L 1405:2007MolEc..16.4149M 1339:1999AmSci..87.....T 1301:1992JArSc..19..181M 1246:2016TEcoE..31..633H 1183:2013Sci...342..871T 1139:(5319): 1687–1689. 1100:2018JArSc..92..126J 1038:2012PNAS..109.8878L 879:2018EvApp..11.1598T 735:2003JFBio..63.1176W 563:Tillandsia bourgaei 433:this relationship. 265:Numerous genera of 3807:Outline of ecology 3756:Industrial ecology 3751:Functional ecology 3615:Ecological deficit 3560:Niche construction 3523:Ecosystem engineer 3300:Species–area curve 3221:Introduced species 3036:: Other components 2968:Deimatic behaviour 2870:Ecological network 2802:North Pacific Gyre 2787:hydrothermal vents 2726:Ecological pyramid 2673:Microbial food web 2484:Primary production 2429:Foundation species 2231:Cleaning symbiosis 1787:: Mad River Press. 1785:Eureka, California 1657:10.3354/meps064113 1347:10.1511/1999.2.160 1327:American Scientist 783:10.7717/peerj.4520 691:Wilson EO (1975). 568: 491: 423:Ophiocreas oedipus 332:species including 165: 30: 3813: 3812: 3696:Balance of nature 3453:Landscape ecology 3338:Community ecology 3280:Species diversity 3216:Indicator species 3211:Gradient analysis 3088:Logistic function 2996: 2995: 2953:Animal coloration 2930:Trophic mutualism 2668:Microbial ecology 2459:Photoheterotrophs 2444:Myco-heterotrophy 2356:Ecosystem ecology 2341:Carrying capacity 2306:Abiotic component 2249: 2248: 2048:Media related to 1888:. Washington DC: 1708:10.3354/meps11697 1393:Molecular Ecology 887:10.1111/eva.12650 815:Harper, Douglas. 706:978-0-674-00089-6 372:spp. and certain 3838: 3513:Ecological niche 3485:selection theory 3305:Umbrella species 3290:Species richness 3226:Invasive species 3206:Flagship species 3113:Population cycle 3108:Overexploitation 3073:Ecological yield 3023: 3016: 3009: 3000: 2999: 2905:Mesotrophic soil 2845:Climax community 2777:Marine food webs 2716:Biomagnification 2517:Chemoorganotroph 2371:Keystone species 2331:Biotic component 2276: 2269: 2262: 2253: 2252: 2240: 2233: 2219: 2212: 2187:Kleptoparasitism 2172:Brood parasitism 2092: 2085: 2078: 2069: 2068: 2064: 2063:. 29 April 2023. 2047: 2032: 2031: 2021: 1981: 1975: 1974: 1941:(6–7): 361–365. 1926: 1920: 1919: 1909: 1900: 1894: 1893: 1877: 1871: 1870: 1838: 1832: 1831: 1795: 1789: 1788: 1776: 1770: 1764: 1759: 1749: 1725: 1719: 1718: 1716: 1714: 1685: 1676: 1670: 1669: 1659: 1642:(1/2): 113–122. 1627: 1621: 1620: 1578: 1572: 1571: 1561: 1529: 1523: 1521: 1519: 1517: 1502: 1496: 1495: 1477: 1441: 1435: 1434: 1416: 1384: 1378: 1377: 1357: 1351: 1350: 1322: 1313: 1312: 1284: 1278: 1277: 1267: 1257: 1225: 1219: 1218: 1166: 1157: 1156: 1128: 1122: 1121: 1119: 1085: 1076: 1070: 1069: 1059: 1049: 1017: 1011: 1010: 982: 963: 962: 934: 917: 916: 906: 873:(9): 1598–1608. 858: 852: 851: 833: 827: 826: 812: 806: 805: 795: 785: 761: 755: 754: 729:(5): 1176–1183. 717: 711: 710: 688: 638:and some mites. 540:or beetles, and 449:is commensal or 238:Beringian wolves 175:The leap from a 3846: 3845: 3841: 3840: 3839: 3837: 3836: 3835: 3816: 3815: 3814: 3809: 3800: 3786:Systems ecology 3674: 3645:Extinction debt 3610:Ecological debt 3600:Bioluminescence 3581: 3574: 3543:marine habitats 3518:Ecological trap 3499: 3379: 3372: 3315: 3309: 3265:Rapoport's rule 3260:Priority effect 3201:Endemic species 3169: 3128:Population size 3044: 3037: 3027: 2997: 2992: 2945: 2939: 2925:Trophic cascade 2835:Bioaccumulation 2818: 2745: 2702: 2624: 2591: 2488: 2400: 2361:Ecosystem model 2294: 2280: 2250: 2245: 2236: 2229: 2215: 2208: 2108: 2096: 2055: 2040: 2035: 1982: 1978: 1927: 1923: 1912:Palaeodiversity 1907: 1901: 1897: 1878: 1874: 1839: 1835: 1820:10.2307/2388309 1796: 1792: 1777: 1773: 1726: 1722: 1712: 1710: 1683: 1677: 1673: 1628: 1624: 1609: 1579: 1575: 1530: 1526: 1515: 1513: 1503: 1499: 1452:(13): 1146–50. 1446:Current Biology 1442: 1438: 1399:(19): 4149–70. 1385: 1381: 1358: 1354: 1323: 1316: 1285: 1281: 1226: 1222: 1177:(6160): 871–4. 1167: 1160: 1129: 1125: 1117:1854/LU-8550758 1083: 1077: 1073: 1032:(23): 8878–83. 1018: 1014: 983: 966: 935: 920: 859: 855: 834: 830: 813: 809: 762: 758: 718: 714: 707: 689: 685: 681: 644: 629: 610: 590: 554: 534:pseudoscorpions 503: 475: 443: 411: 363: 263: 253: 194: 170:fight-or-flight 134: 89: 35:is a long-term 17: 12: 11: 5: 3844: 3834: 3833: 3828: 3811: 3810: 3805: 3802: 3801: 3799: 3798: 3793: 3788: 3783: 3778: 3773: 3768: 3766:Microecosystem 3763: 3758: 3753: 3748: 3743: 3738: 3733: 3728: 3723: 3718: 3713: 3708: 3703: 3698: 3693: 3688: 3682: 3680: 3676: 3675: 3673: 3672: 3667: 3665:Thorson's rule 3662: 3657: 3652: 3647: 3642: 3637: 3632: 3627: 3622: 3617: 3612: 3607: 3602: 3597: 3592: 3590:Assembly rules 3586: 3584: 3576: 3575: 3573: 3572: 3567: 3562: 3557: 3552: 3547: 3546: 3545: 3535: 3530: 3525: 3520: 3515: 3509: 3507: 3501: 3500: 3498: 3497: 3492: 3487: 3475: 3473:Patch dynamics 3470: 3468:Metapopulation 3465: 3460: 3455: 3450: 3445: 3440: 3435: 3430: 3425: 3420: 3415: 3410: 3405: 3400: 3395: 3390: 3384: 3382: 3374: 3373: 3371: 3370: 3365: 3363:Storage effect 3360: 3355: 3350: 3345: 3340: 3335: 3330: 3325: 3319: 3317: 3311: 3310: 3308: 3307: 3302: 3297: 3292: 3287: 3282: 3277: 3272: 3267: 3262: 3257: 3252: 3247: 3245:Neutral theory 3242: 3237: 3232: 3230:Native species 3223: 3218: 3213: 3208: 3203: 3198: 3193: 3188: 3183: 3177: 3175: 3171: 3170: 3168: 3167: 3162: 3161: 3160: 3155: 3145: 3140: 3135: 3130: 3125: 3120: 3115: 3110: 3105: 3103:Overpopulation 3100: 3095: 3090: 3085: 3080: 3075: 3070: 3065: 3060: 3055: 3049: 3047: 3039: 3038: 3026: 3025: 3018: 3011: 3003: 2994: 2993: 2991: 2990: 2985: 2980: 2975: 2970: 2965: 2960: 2955: 2949: 2947: 2941: 2940: 2938: 2937: 2932: 2927: 2922: 2917: 2912: 2910:Nutrient cycle 2907: 2902: 2900:Feeding frenzy 2897: 2892: 2887: 2882: 2880:Energy quality 2877: 2872: 2867: 2862: 2857: 2852: 2847: 2842: 2840:Cascade effect 2837: 2832: 2826: 2824: 2820: 2819: 2817: 2816: 2815: 2814: 2809: 2804: 2799: 2794: 2789: 2784: 2774: 2769: 2764: 2759: 2753: 2751: 2747: 2746: 2744: 2743: 2738: 2733: 2728: 2723: 2718: 2712: 2710: 2704: 2703: 2701: 2700: 2695: 2690: 2685: 2683:Microbial loop 2680: 2675: 2670: 2665: 2660: 2655: 2650: 2648:Lithoautotroph 2645: 2640: 2634: 2632: 2630:Microorganisms 2626: 2625: 2623: 2622: 2617: 2612: 2607: 2601: 2599: 2593: 2592: 2590: 2589: 2587:Prey switching 2584: 2579: 2574: 2569: 2564: 2559: 2554: 2549: 2544: 2539: 2534: 2529: 2524: 2519: 2514: 2509: 2504: 2498: 2496: 2490: 2489: 2487: 2486: 2481: 2476: 2471: 2466: 2464:Photosynthesis 2461: 2456: 2451: 2446: 2441: 2436: 2431: 2426: 2421: 2419:Chemosynthesis 2416: 2410: 2408: 2402: 2401: 2399: 2398: 2393: 2388: 2383: 2378: 2373: 2368: 2363: 2358: 2353: 2348: 2343: 2338: 2333: 2328: 2323: 2318: 2313: 2311:Abiotic stress 2308: 2302: 2300: 2296: 2295: 2279: 2278: 2271: 2264: 2256: 2247: 2246: 2244: 2243: 2242: 2241: 2234: 2222: 2221: 2220: 2213: 2201: 2200: 2199: 2194: 2189: 2184: 2179: 2174: 2169: 2159: 2154: 2149: 2144: 2139: 2134: 2129: 2124: 2119: 2113: 2110: 2109: 2095: 2094: 2087: 2080: 2072: 2066: 2065: 2057:"Commensalism" 2053: 2039: 2038:External links 2036: 2034: 2033: 1976: 1921: 1895: 1882:"Commensalism" 1872: 1833: 1790: 1771: 1720: 1671: 1622: 1607: 1573: 1524: 1511:UCLA News Room 1497: 1436: 1379: 1352: 1314: 1295:(2): 181–204. 1279: 1240:(8): 633–645. 1220: 1158: 1123: 1071: 1012: 964: 945:(2): 161–190. 918: 853: 828: 817:"commensalism" 807: 756: 712: 705: 682: 680: 677: 676: 675: 669: 663: 657: 651: 643: 640: 628: 625: 609: 606: 589: 586: 580:(such as many 553: 550: 502: 499: 474: 471: 442: 439: 410: 407: 374:crenarchaeotes 362: 352: 347:S. epidermidis 341:S. lugdunensis 330:Staphylococcus 322:nasal cavities 261:Staphylococcus 252: 250:Staphylococcus 243: 193: 190: 182:coevolutionary 133: 130: 97:Medieval Latin 88: 85: 15: 9: 6: 4: 3: 2: 3843: 3832: 3829: 3827: 3824: 3823: 3821: 3808: 3803: 3797: 3794: 3792: 3791:Urban ecology 3789: 3787: 3784: 3782: 3779: 3777: 3774: 3772: 3769: 3767: 3764: 3762: 3759: 3757: 3754: 3752: 3749: 3747: 3744: 3742: 3739: 3737: 3734: 3732: 3729: 3727: 3724: 3722: 3719: 3717: 3714: 3712: 3709: 3707: 3704: 3702: 3699: 3697: 3694: 3692: 3689: 3687: 3684: 3683: 3681: 3677: 3671: 3668: 3666: 3663: 3661: 3658: 3656: 3653: 3651: 3650:Kleiber's law 3648: 3646: 3643: 3641: 3638: 3636: 3633: 3631: 3628: 3626: 3623: 3621: 3618: 3616: 3613: 3611: 3608: 3606: 3603: 3601: 3598: 3596: 3593: 3591: 3588: 3587: 3585: 3583: 3577: 3571: 3568: 3566: 3563: 3561: 3558: 3556: 3553: 3551: 3548: 3544: 3541: 3540: 3539: 3536: 3534: 3531: 3529: 3526: 3524: 3521: 3519: 3516: 3514: 3511: 3510: 3508: 3506: 3502: 3496: 3493: 3491: 3488: 3486: 3484: 3480: 3476: 3474: 3471: 3469: 3466: 3464: 3461: 3459: 3456: 3454: 3451: 3449: 3446: 3444: 3441: 3439: 3436: 3434: 3431: 3429: 3426: 3424: 3423:Foster's rule 3421: 3419: 3416: 3414: 3411: 3409: 3406: 3404: 3401: 3399: 3396: 3394: 3391: 3389: 3386: 3385: 3383: 3381: 3375: 3369: 3366: 3364: 3361: 3359: 3356: 3354: 3351: 3349: 3346: 3344: 3341: 3339: 3336: 3334: 3331: 3329: 3326: 3324: 3321: 3320: 3318: 3312: 3306: 3303: 3301: 3298: 3296: 3293: 3291: 3288: 3286: 3283: 3281: 3278: 3276: 3273: 3271: 3268: 3266: 3263: 3261: 3258: 3256: 3253: 3251: 3248: 3246: 3243: 3241: 3238: 3236: 3233: 3231: 3227: 3224: 3222: 3219: 3217: 3214: 3212: 3209: 3207: 3204: 3202: 3199: 3197: 3194: 3192: 3189: 3187: 3184: 3182: 3179: 3178: 3176: 3172: 3166: 3163: 3159: 3156: 3154: 3151: 3150: 3149: 3146: 3144: 3141: 3139: 3136: 3134: 3131: 3129: 3126: 3124: 3121: 3119: 3116: 3114: 3111: 3109: 3106: 3104: 3101: 3099: 3096: 3094: 3091: 3089: 3086: 3084: 3081: 3079: 3076: 3074: 3071: 3069: 3066: 3064: 3061: 3059: 3056: 3054: 3051: 3050: 3048: 3046: 3040: 3035: 3031: 3024: 3019: 3017: 3012: 3010: 3005: 3004: 3001: 2989: 2986: 2984: 2981: 2979: 2976: 2974: 2971: 2969: 2966: 2964: 2961: 2959: 2956: 2954: 2951: 2950: 2948: 2942: 2936: 2933: 2931: 2928: 2926: 2923: 2921: 2918: 2916: 2913: 2911: 2908: 2906: 2903: 2901: 2898: 2896: 2893: 2891: 2888: 2886: 2883: 2881: 2878: 2876: 2873: 2871: 2868: 2866: 2863: 2861: 2858: 2856: 2853: 2851: 2848: 2846: 2843: 2841: 2838: 2836: 2833: 2831: 2828: 2827: 2825: 2821: 2813: 2810: 2808: 2805: 2803: 2800: 2798: 2795: 2793: 2790: 2788: 2785: 2783: 2780: 2779: 2778: 2775: 2773: 2770: 2768: 2765: 2763: 2760: 2758: 2755: 2754: 2752: 2748: 2742: 2741:Trophic level 2739: 2737: 2734: 2732: 2729: 2727: 2724: 2722: 2719: 2717: 2714: 2713: 2711: 2709: 2705: 2699: 2698:Phage ecology 2696: 2694: 2691: 2689: 2688:Microbial mat 2686: 2684: 2681: 2679: 2676: 2674: 2671: 2669: 2666: 2664: 2661: 2659: 2656: 2654: 2651: 2649: 2646: 2644: 2643:Bacteriophage 2641: 2639: 2636: 2635: 2633: 2631: 2627: 2621: 2618: 2616: 2613: 2611: 2610:Decomposition 2608: 2606: 2603: 2602: 2600: 2598: 2594: 2588: 2585: 2583: 2580: 2578: 2575: 2573: 2570: 2568: 2565: 2563: 2560: 2558: 2557:Mesopredators 2555: 2553: 2550: 2548: 2545: 2543: 2540: 2538: 2535: 2533: 2530: 2528: 2525: 2523: 2520: 2518: 2515: 2513: 2510: 2508: 2505: 2503: 2502:Apex predator 2500: 2499: 2497: 2495: 2491: 2485: 2482: 2480: 2477: 2475: 2472: 2470: 2467: 2465: 2462: 2460: 2457: 2455: 2452: 2450: 2447: 2445: 2442: 2440: 2437: 2435: 2432: 2430: 2427: 2425: 2422: 2420: 2417: 2415: 2412: 2411: 2409: 2407: 2403: 2397: 2394: 2392: 2389: 2387: 2384: 2382: 2379: 2377: 2374: 2372: 2369: 2367: 2364: 2362: 2359: 2357: 2354: 2352: 2349: 2347: 2344: 2342: 2339: 2337: 2336:Biotic stress 2334: 2332: 2329: 2327: 2324: 2322: 2319: 2317: 2314: 2312: 2309: 2307: 2304: 2303: 2301: 2297: 2292: 2288: 2284: 2277: 2272: 2270: 2265: 2263: 2258: 2257: 2254: 2239: 2235: 2232: 2228: 2227: 2226: 2223: 2218: 2214: 2211: 2207: 2206: 2205: 2202: 2198: 2195: 2193: 2190: 2188: 2185: 2183: 2182:Hyperparasite 2180: 2178: 2175: 2173: 2170: 2168: 2165: 2164: 2163: 2160: 2158: 2155: 2153: 2150: 2148: 2145: 2143: 2140: 2138: 2135: 2133: 2130: 2128: 2125: 2123: 2120: 2118: 2115: 2114: 2111: 2107: 2103: 2100: 2099:Inter-species 2093: 2088: 2086: 2081: 2079: 2074: 2073: 2070: 2062: 2058: 2054: 2051: 2046: 2042: 2041: 2029: 2025: 2020: 2015: 2011: 2007: 2003: 1999: 1995: 1991: 1987: 1980: 1972: 1968: 1964: 1960: 1956: 1952: 1948: 1944: 1940: 1936: 1932: 1925: 1917: 1913: 1906: 1899: 1891: 1887: 1883: 1876: 1868: 1864: 1860: 1856: 1852: 1848: 1844: 1837: 1829: 1825: 1821: 1817: 1813: 1809: 1805: 1801: 1794: 1786: 1782: 1775: 1768: 1763: 1757: 1753: 1748: 1743: 1739: 1735: 1731: 1724: 1709: 1705: 1701: 1697: 1693: 1689: 1682: 1675: 1667: 1663: 1658: 1653: 1649: 1645: 1641: 1637: 1633: 1626: 1618: 1614: 1610: 1608:9780123855299 1604: 1600: 1596: 1592: 1588: 1584: 1577: 1569: 1565: 1560: 1555: 1551: 1547: 1544:(3): 505–20. 1543: 1539: 1535: 1528: 1512: 1508: 1501: 1493: 1489: 1485: 1481: 1476: 1471: 1467: 1463: 1459: 1455: 1451: 1447: 1440: 1432: 1428: 1424: 1420: 1415: 1410: 1406: 1402: 1398: 1394: 1390: 1383: 1375: 1371: 1368:(3): 81–146. 1367: 1363: 1356: 1348: 1344: 1340: 1336: 1332: 1328: 1321: 1319: 1310: 1306: 1302: 1298: 1294: 1290: 1283: 1275: 1271: 1266: 1261: 1256: 1251: 1247: 1243: 1239: 1235: 1231: 1224: 1216: 1212: 1208: 1204: 1200: 1196: 1192: 1188: 1184: 1180: 1176: 1172: 1165: 1163: 1154: 1150: 1146: 1142: 1138: 1134: 1127: 1118: 1113: 1109: 1105: 1101: 1097: 1093: 1089: 1082: 1075: 1067: 1063: 1058: 1053: 1048: 1043: 1039: 1035: 1031: 1027: 1023: 1016: 1008: 1004: 1000: 996: 992: 988: 981: 979: 977: 975: 973: 971: 969: 960: 956: 952: 948: 944: 940: 933: 931: 929: 927: 925: 923: 914: 910: 905: 900: 896: 892: 888: 884: 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Index


Remora
biological interaction
symbiosis
species
mutualism
amensalism
parasitism
host
remoras
sharks
pilot fish
herbivores
French
Medieval Latin
com-
mensa
Oxford
Cambridge
Pierre-Joseph van Beneden

Domestic
feral pigeons
rock dove
cosmopolitan distribution
fight-or-flight
synanthropic
coevolutionary
niche
dog

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