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Fezouata Formation

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1034: 1408: 1621: 645: 468: 3541: 2167: 278: 1336: 1589: 2457: 1470: 303: 294: 285: 1128: 2504: 104: 551:(IUGS) included the 'Ordovician Fezouata Shale Fossil Site at Jbeltizagzaouine' in its assemblage of 100 'geological heritage sites' around the world in a listing published in October 2022. The organisation defines an IUGS Geological Heritage Site as 'a key place with geological elements and/or processes of international scientific relevance, used as a reference, and/or with a substantial contribution to the development of geological sciences through history.' 3986: 7119: 380:, have been recovered from the formations in addition to a less abundant shelly fauna. The make-up of the community varies significantly through the stratigraphic sequence, with both abundances and faunal composition changing as time progresses. Major burrowing is not present, but there are small (1-3 mm wide) burrows in the sediment, which may indicate that there is not enough oxygen in the water or sediment. Particularly notable is the presence of 2072: 681: 2646: 260: 361: 479:(offshore to lower shoreface transition), at between 50 and 150 metres (160 and 490 ft) water depth. Organisms were likely buried in situ. Because of its placement above storm wave base, storms would have mobilized sediment that could be quickly deposited, trapping animals and leading to their preservation. Consequently, the assemblage is dominated by benthic organisms. 2757:
oxygenated waters. The rarity of carbonate platforms and hard substrates in the Gondwanan area delayed the diversification of crinoids and edrioasteroids in the region. This also provided more space for the establishment of a distinctive South Polar ecosystem dominated by eocrinoids, mitrates, solutans, and eventually diploporites.
495:. Non-mineralized appendages are often preserved. While the formation as a whole is over 1,000 metres (3,300 ft) thick, only two intervals, 25 metres (82 ft) and 15 metres (49 ft) thick, provide exceptional preservation. Both of these intervals are located near the top of the lower formation, corresponding to the 1057:
An uncommon medium-sized endemic asaphid. Some specimens have been fossilized with unusual flattened antennae, the only reliable attributions of these structures in asaphids. The antennae may have had a particular sensory function to assist in hunting or scavenging. Found in relatively shallow (lower
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This is unusual relative to other Ordovician sponge ecosystems, such as the Builth Inlier of Wales. Another difference is how Fezouata's sponge fauna consists mostly of protomonaxonids, with a few reticulosans occupying course-grained shallow seabeds. Conversely, in Wales there is a clear succession
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A large isoteline asaphid up to 28.5 centimetres (11.2 in) long, not counting a pronounced spine on the pygidium. Some specimens preserving gut traces and appendages showing new anatomical traits not seen in other trilobites. The cephalic appendages hosted long spines capable of combing through
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Nevertheless, there are quite a few exceptions which contradict these broad rules. Many sites record diverse deep-water ecosystems protected from both storms and insufficient oxygen. Numerous species of echinoderms and other invertebrates coexist at these localities, even if one echinoderm species
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A common nileid with gregarious habits, often associated with graptolite debris at multiple water depths. Some specimens have been fossilized in the process of a mass moulting event, including some of the few known fossils of post-moult trilobites. Known from multiple life stages, including mid to
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Overall diversity is rather low, and species which were common in temperate and tropical seas are apparently absent. The Fezouata Formation appears to be an exemplar of the 'subpolar domain', an assemblage of cold-water coastal conodonts native to the South Polar region of the Early Ordovician.
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Fezouata reconstructs the uneven nature of the Cambrian-Ordovician transition of echinoderm faunas. Cosmopolitan Late Cambrian hallmarks (such as cornute stylophorans) maintain their abundance in oxygen-poor areas, while newer groups (crinoids, diploporites, asterozoans) make inroads into more
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A common bivalve, tentatively identified as one of the oldest lucinids (hatchet shells). It was likely an infaunal filter-feeder capable of rapid vertical movement within sediment. Like modern lucinids, it may have been a specialist in oxygen-poor environments thanks to endosymbiotic
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A common and endemic eocrinoid which can be abundant in certain layers. It is a small but robust species often found in densely jumbled fossiliferous lenses. These lenses are interpreted as debris from shallow-water "meadows", uprooted and washed down into deeper areas by storms.
6602:, which has a broader and less dense distribution in the formation. Periodic unstable seafloor conditions (potentially related to seasonal disruptions) would have favored species-poor colonization events over short time periods, rather than a stable and diverse equilibrium. 8047:
Saleh, Farid; Vaucher, Romain; Vidal, Muriel; Hariri, Khadija El; Laibl, Lukáš; Daley, Allison C.; Gutiérrez-Marco, Juan Carlos; Candela, Yves; Harper, David A. T.; Ortega-Hernández, Javier; Ma, Xiaoya; Rida, Ariba; Vizcaïno, Daniel; Lefebvre, Bertrand (2022-12-13).
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epochs. Two stratigraphic intervals of the formation are fossiliferous: the lower is Late Tremadocian and sits 260 to 330 metres (850 to 1,080 ft) above the base of the formation; the upper, at 570 to 620 metres (1,870 to 2,030 ft), is mid-Floian in age.
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Martin, Emmanuel L.O.; Pittet, Bernard; Gutiérrez-Marco, Juan-Carlos; Vannier, Jean; El Hariri, Khadija; Lerosey-Aubril, Rudy; Masrour, Moussa; Nowak, Hendrik; Servais, Thomas; Vandenbroucke, Thijs R.A.; Van Roy, Peter; Vaucher, Romain; Lefebvre, Bertrand (2016).
861:, up to 4.8 centimetres (1.9 in) long, which is common in relatively deep (upper offshore) sediments. Fossils of this species are known to preserve appendages similar in form to those of Cambrian trilobites. Known from multiple life stages, including late 7400:
Saleh, Farid; Antcliffe, Jonathan B.; Birolini, Enzo; Candela, Yves; Corthésy, Nora; Daley, Allison C.; Dupichaud, Christophe; Gibert, Corentin; Guenser, Pauline; Laibl, Lukáš; Lefebvre, Bertrand; Michel, Soline; Potin, Gaëtan J.-M. (6 September 2024).
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indicate a normal range of salinity, and the overall shelly assemblage is not significantly different from the normal shelly fauna expected in open Ordovician waters. The non-mineralized cohort contains a range of forms familiar from the Burgess Shale:
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Similar conodont faunas are known from Early Ordovician deposits in Central Europe, which was also located near the South Pole. The 'subpolar domain' survived into the Middle Ordovician and expanded into areas now found in the Middle East.
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worm. Though most palaeoscolecids are inferred to be burrowing carnivores, remains from the Fezouata Formation are more consistent with an opportunistic epibenthic lifestyle, patrolling for organic matter on the surface of the seabed.
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A large endemic nileid, up to 23 centimetres (9.1 in) long. Typically considered a single species. Known from multiple life stages, including early to late meraspids. Found in relatively shallow (lower nearshore) sediments.
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Saleh, Farid; Vidal, Muriel; Laibl, Lukáš; Sansjofre, Pierre; Gueriau, Pierre; Pérez-Peris, Francesc; Lustri, Lorenzo; Lucas, Victoire; Lefebvre, Bertrand; Pittet, Bernard; El Hariri, Khadija; Daley, Allison C. (2021).
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Lefebvre, Bertrand; Allaire, Ninon; Guensburg, Thomas E.; Hunter, Aaron W.; KouraĂŻss, Khaoula; Martin, Emmanuel L.O.; Nardin, Elise; Noailles, Fleur; Pittet, Bernard; Sumrall, Colin D.; Zamora, Samuel (2016).
4134:(hatchet shells). It was likely an infaunal filter-feeder capable of rapid vertical movement within sediment. Like modern lucinids, it may have been a specialist in oxygen-poor environments thanks to 7292: 7269: 7246: 825:
lobsters, either as biological aggregations (for spawning or moulting), or migrating groups due to environmental pressures such as storms. Other trilobites occasionally become entangled with the
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have been found growing around the edge of a cephalopod shell, one of the few examples of hard substrate to be found in the formation. This association is similar to the ecology reported for
4639:, an extinct group of gastropod-like molluscs with asymmetrically coiled shells. It was gregarious and primarily found in shallow oxygenated waters represented at the Oued Beni Zoli locality. 7736: 7709: 5429:
interval” (10). Not all of the index taxa which these zones are named for are known from the Fezouata Formation. The Tremadocian-Floian boundary is approximately at the level between the
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Lefebvre, Bertrand; Guensburg, Thomas E.; Martin, Emmanuel L.O.; Mooi, Rich; Nardin, Elise; Nohejlová, Martina; Saleh, Farid; Kouraïss, Khaoula; El Hariri, Khadija; David, Bruno (2019).
10423:"Palaeobiogeography, palaeoecology and evolution of Lower Ordovician conulariids and Sphenothallus (Medusozoa, Cnidaria), with emphasis on the Fezouata Shale of southeastern Morocco" 2485:
marrellomorph which has not been formally named or described in detail. It had three pairs of cephalic appendages, 13 pairs of trunk appendages, and a cephalic shield with six large
7223: 141:>1,000 m (3,300 ft), combined for upper and lower formations, the two exceptionally preserved intervals ~25 m (82 ft) and 15 m (49 ft) respectively 6610:
and thick-walled hexactinellids in shallow reefs and other energetic areas, with protomonaxonids at intermediate depths, and reticulosans in the deepest and calmest environments.
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Fossils of the Fezouata Formation, which are usually squashed flat (although some do retain some degree of their original three-dimensionality) are often coated with a dusting of
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Lefebvre, B.; Botting, J. (2007). "First report of the mitrate Peltocystis cornuta Thoral (Echinodermata, Stylophora) in the Lower Ordovician of central Anti-Atlas (Morocco)".
9715:"Morphological disparity and systematic revision of the eocrinoid genus Rhopalocystis (Echinodermata, Blastozoa) from the Lower Ordovician of the central Anti-Atlas (Morocco)" 8344:
Vannier, Jean; Vidal, Muriel; Marchant, Robin; El Hariri, Khadija; Kouraiss, Khaoula; Pittet, Bernard; El Albani, Abderrazak; Mazurier, Arnaud; Martin, Emmanuel (2019-10-17).
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The Lagerstätten were first identified in the late 1990s when a local fossil collector, Ben Moula, showed some of the finds to a PhD student who was then working in the area.
9894:"The bivalve fauna from the Fezouata Formation (Lower Ordovician) of Morocco and its significance for palaeobiogeography, palaeoecology and early diversification of bivalves" 9247:
Laibl, Lukáš; Gueriau, Pierre; Saleh, Farid; Pérez-Peris, Francesc; Lustri, Lorenzo; Drage, Harriet B.; Bath Enright, Orla G.; Potin, Gaëtan J.-M.; Daley, Allison C. (2023).
3475:. Crushed but complete skeletons of this species are abundant at Oued Beni Zoli locality Z-F5, a site preserving a diverse assemblage of storm-influenced siltstone in the 10703: 5337:
Graptolites in the Fezouata Formation are important for biostratigraphic correlation to other regions. Most species found in the formation are assigned to planktic (
7937:"Radiodont frontal appendages from the Fezouata Biota (Morocco) reveal high diversity and ecological adaptations to suspension-feeding during the Early Ordovician" 7191:
Lefebvre, Bertrand; Gutiérrez-Marco, Juan C; Lehnert, Oliver; Martin, Emmanuel L. O; Nowak, Hendrik; Akodad, Mustapha; El Hariri, Khadija; Servais, Thomas (2017).
1033: 8416:"Systematics, morphology, and appendages of an Early Ordovician pilekiine trilobite Anacheirurus from Fezouata Shale and the early diversification of Cheiruridae" 7762: 6996: 2710:. Fezouata stylophoran fossils include soft tissue preserved among the skeletal elements, helping to unravel controversial details of their anatomy and ecology. 9039:"Sequence of post-moult exoskeleton hardening preserved in a trilobite mass moult assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco" 6520: 5021: 4727: 3306: 2713:
Specific echinoderm species may form dense fossil beds in some layers of the formation, a phenomenon which is particularly common in the mid-late Tremadocian (
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A filter-feeding aegirocassisine hurdiid radiodont. Originally considered enigmatic until 2023, where it was redescribed as a radiodont. Like the contemporary
9665:"Solutan echinoderms from the Fezouata Shale Lagerstätte (Lower Ordovician, Morocco): diversity, exceptional preservation, and palaeoecological implications" 8302:"Biostratigraphic and palaeoenvironmental controls on the trilobite associations from the Lower Ordovician Fezouata Shale of the central Anti-Atlas, Morocco" 5043: 10776: 7103:(armored annelid worm). This species is the only machaeridian known to preserve soft tissue and jaw material, helping to demonstrate their affinities with 6361: 4999: 4712: 1385: 726:
The largest trilobite individuals in the Fezouata Formation tend to inhabit deep oxygenated waters with minimal influence from storms or larger predators.
10320: 6589: 4744: 3676: 3551: 2691: 10175:"Conodonts from the Lower Ordovician of Morocco — Contributions to age and faunal diversity of the Fezouata Lagerstätte and peri-Gondwana biogeography" 6839: 6025: 4397: 3964: 3530: 3143:
A locally abundant cornute with a large sample size, allowing for a reconstruction of decay pathways in stylophoran soft tissue and skeletal material.
3110:. The dominant echinoderm at sites Z-F26 and Z-F23, which preserve diverse fossil assemblages buried in calm-water fine-grained siltstone of the ? 2683: 1928: 10868: 7104: 6346: 5975: 5723: 5197: 5093: 4477: 4432: 3469: 2137: 2012: 1713: 1367: 1346: 6598:
Though at least 27 sponge species have been recorded in the biota, nearly all occurrences are monospecific death assemblages, with the exception of
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Dupichaud, Christophe; Lefebvre, Bertrand; Milne, Claire H.; Mooi, Rich; Nohejlová, Martina; Roch, Renaud; Saleh, Farid; Zamora, Samuel (2023).
9249:"Early developmental stages of a Lower Ordovician marrellid from Morocco suggest simple ontogenetic niche differentiation in early euarthropods" 6867: 6570: 6562: 6448: 6419: 6332: 5917: 5648: 5618: 5564: 5493: 5294: 5216: 5081: 4874: 4832: 4509: 4075: 3664:. Paddles of this species are abundant at sites Z-F13c and Z-F24, a pair of diverse faunal assemblages emplaced in sandy storm deposits of the 3639: 3512: 3024: 2818: 2656: 2261: 2023: 1996: 1196: 9577:"Exceptionally preserved soft parts in fossils from the Lower Ordovician of Morocco clarify stylophoran affinities within basal deuterostomes" 8895:"Digestive and appendicular soft-parts, with behavioural implications, in a large Ordovician trilobite from the Fezouata Lagerstätte, Morocco" 6933:
An abundant small conulariid (less than 5 centimetres (2.0 in) tall). Frequently found attached to brachiopods or even other conulariids.
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Pérez-Peris, Francesc; Laibl, Lukáš; Lustri, Lorenzo; Gueriau, Pierre; Antcliffe, Jonathan B; Bath Enright, Orla G; Daley, Allison C (2021).
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Martin, Emmanuel L.O.; Vidal, Muriel; VizcaĂŻno, Daniel; Vaucher, Romain; Sansjofre, Pierre; Lefebvre, Bertrand; Destombes, Jacques (2016).
6574: 6433: 5235: 5009: 4780: 4681: 4408: 3487: 2962: 2424: 1903: 1782: 1631: 1613: 1066: 855: 10646:
Nowak, Hendrik; Servais, Thomas; Pittet, Bernard; Vaucher, Romain; Akodad, Mustapha; Gaines, Robert R.; Vandenbroucke, Thijs R.A. (2016).
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up to 3.8 centimetres (1.5 in) long. Found at intermediate depths in the transitional zone between shoreface and offshore sediments.
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Lehnert, Oliver; Nowak, Hendrik; Sarmiento, Graciela N.; Gutiérrez-Marco, Juan Carlos; Akodad, Mustapha; Servais, Thomas (October 2016).
9952:"Palaeogeography and palaeoecology of early Floian (Early Ordovician) cephalopods from the Upper Fezouata Formation, Anti-Atlas, Morocco" 6943: 6491: 6404: 6317: 6261: 5374: 4649: 4541: 2082: 1511: 1388:
up to 33.7 centimetres (13.3 in) long. Often found in mass death assemblages, possibly for spawning akin to modern horseshoe crabs.
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A rare "azygograptid", a unique type of early graptoloid with a single uniserial stipe, convergently similar to the Silurian-Devonian
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In respect of the 'exceptional fossil preservation bridging the Cambrian Explosion and the Great Ordovician Biodiversification', the
10472:"Palaeoscolecid worms from the Lower Ordovician Fezouata Lagerstätte, Morocco: Palaeoecological and palaeogeographical implications" 7467:
Botting, J. (2007). "'Cambrian' demosponges in the Ordovician of Morocco: Insights into the early evolutionary history of sponges".
10689: 10511:"X-ray microtomography applied to fossils preserved in compression: Palaeoscolescid worms from the Lower Ordovician Fezouata Shale" 4783:. It was likely a shallowly infaunal filter-feeder capable of rapid reburial, with the majority of the shell embedded in sediment. 2717:
graptolite zone). Most echinoderm beds are dominated by only a few species, often representatives of stylophorans or the eocrinoid
1813: 1480: 8625:"Rusophycus carleyi (James, 1885), Trace Fossils from the Lower Ordovician of Southern Morocco, and the Trilobites that Made Them" 567:
Apart from the three named species of Fezouata radiodonts, three other unnamed species occur in the formation: a third species of
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etc.), and other indeterminate forms. Many of the sponges have affinities with Cambrian taxa common in Burgess Shale-type faunas.
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A common and endemic eocrinoid. It is a large but fragile species which is often found disarticulated in siltstone beds. Unlike
1407: 277: 7769:. The Economist, print edition: January 26th 2019, p. 72 (1/2-page based on article in Geology--in prior footnote--1/Feb/2019) 2729:
usurp this niche. Conversely, stylophorans are opportunistic colonizers of deeper low-oxygen seabeds in some intervals of the
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A small isoteline asaphid up to 5.6 centimetres (2.2 in) long. Common in relatively shallow (lower nearshore) sediments.
9451:"Revision of the mollisoniid chelicerate(?) Thelxiope, with a new species from the middle Cambrian Wheeler Formation of Utah" 9022: 958:. Associated body fossils and trace fossils help to identify this species as the tracemaker for a distinctive resting trace, 522:. Stratigraphically productive layers are found through a 1.1 kilometres (0.68 mi)-thick column of rock that spans the 10838: 9714: 9403: 9134:"A new aglaspidid euarthropod with a six-segmented trunk from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco" 8832: 8187: 2733:
zone. Echinoderms are uncommon at intermediate depths, which have a higher proportion of brachiopod and trilobite fossils.
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Kouraiss, Khaoula; El Hariri, Khadija; El Albani, Abderrazak; Azizi, Abdelfattah; Mazurier, Arnaud; Vannier, Jean (2018).
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Saleh, Farid; Lefebvre, Bertrand; Dupichaud, Christophe; Martin, Emmanuel L.O.; Nohejlová, Martina; Spaccesi, Léa (2023).
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species dominate "meadow"-like ecosystems in shallow waters impacted by storms. In the youngest layers of the formations,
1774:, a trace fossil common throughout Ordovician Gondwana. Known from multiple life stages, including mid to late meraspids. 10648:"Palynomorphs of the Fezouata Shale (Lower Ordovician, Morocco): Age and environmental constraints of the Fezouata Biota" 7789: 2489:-bearing spines. Both adult and juvenile specimens are known of this species, including individuals caught in the act of 1029:
A large endemic asaphid up to 38 centimetres (15 in) long. Found in relatively shallow (lower nearshore) sediments.
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species, one which reaches 7.5 centimetres (3.0 in) in length, and another reaching 4.2 centimetres (1.7 in).
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which often occurs in distinctive linear clusters. These clusters are interpreted as collective behavior akin to modern
9538:"Palaeoecological aspects of the diversification of echinoderms in the Lower Ordovician of central Anti-Atlas, Morocco" 8997:
Gutiérrez-Marco, Juan Carlos; Rábano, Isabel; García-Bellido, Diego C. (2019), Owen, Alan W.; Bruton, David L. (eds.),
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clusters. This species is widely distributed in time and is most common in relatively deep (upper offshore) sediments.
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Van Iten, Heyo; Muir, Lucy; Simões, Marcello G.; Leme, Juliana M.; Marques, Antonio C.; Yoder, Naomi (October 2016).
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Allaire, Ninon; Lefebvre, Bertrand; Nardin, Elise; Martin, Emmanuel L.O.; Vaucher, Romain; Escarguel, Gilles (2017).
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from upper Tremadocian strata of the Moroccan Anti-Atlas: taxonomic reappraisal and palaeoenvironmental implications"
8965:"Parvilichas marochii: New genus and species of Lichidae from the Zagora region (Morocco); Early Ordovician (Floian)" 8483:
Laibl, Lukáš; Drage, Harriet B.; Pérez-Peris, Francesc; Schöder, Sebastian; Saleh, Farid; Daley, Allison C. (2023).
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A small and common asaphid up to 6.2 centimetres (2.4 in) long, without clear preferences for specific depths.
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The most well-preserved fossils in the Fezouata Formation generally come from strata of the late Tremadocian (late
7661:"The Lower Ordovician Fezouata Konservat-Lagerstätte from Morocco: Age, environment and evolutionary perspectives" 173: 10712: 10261:"Rhabdopleurid epibionts from the Ordovician Fezouata Shale biota and the longevity of cross-phylum interactions" 10219:"Biostratigraphy and palaeoecology of Lower Ordovician graptolites from the Fezouata Shale (Moroccan Anti-Atlas)" 348:, and the particular strata within the formation which exhibit exceptional preservation are generally termed the 2499:
for this species, first introduced in a PhD thesis. Multiple marrellid species may be present in the formation.
9619:"Skeletal elements controlled soft-tissue preservation in echinoderms from the Early Ordovician Fezouata Biota" 2682:
Many echinoderm species of the Fezouata Biota remain unnamed and undescribed. These include representatives of
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Fezouata conodonts are difficult to correlate to biostratigraphic systems in nearby temperate regions such as
8485:"Babies from the Fezouata Biota: Early developmental trilobite stages and their adaptation to high latitudes" 1584:, which is often considered a group of trilobites. Very small, at up to 0.9 centimetres (0.35 in) long. 8871:
Fortey, Richard A. (2012). "The first known complete Lichakephalid trilobite, lower Ordovician of Morocco".
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The graptolites of the Fezouata Formation are distributed over 10 biozones. In order, these zones are: the
166: 10087:"Gastropoda, Tergomya and Paragastropoda (Mollusca) from the Lower Ordovician Fezouata Formation, Morocco" 7708:
Saleh, Farid; Candela, Yves; Harper, David A. T.; Polechová, Marika; Lefebvre, Bertrand; Pittet, Bernard.
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which is common at intermediate depths in the transitional zone between shoreface and offshore sediments.
10833: 10321:"A new species of the problematic organism Webbyites from the Early Ordovician Fezouata Biota of Morocco" 9820:"A new somasteroid from the Fezouata Lagerstätte in Morocco and the Early Ordovician origin of Asterozoa" 7716:. Society for Sedimentary Geology PALAIOS, print edition: 21 December 2018, pp. 535-541 (Vol. 33, No. 12) 644: 514:
The fossils occur within an area of 500 square kilometres (190 sq mi), in southeast Morocco's
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Lefebvre, Bertrand; El Hariri, Khadija; Lerosey-Aubril, Rudy; Servais, Thomas; Van Roy, Peter (2016).
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from the Cambian, and thus serves as a transitional form among Ordovician euchelicerate evolution.
373: 9951: 9290:"Novel marrellomorph moulting behaviour preserved in the Lower Ordovician Fezouata Shale, Morocco" 9038: 8624: 8415: 962:. Found at intermediate depths in the transitional zone between shoreface and offshore sediments. 364:
Reconstruction of the Fezouata Biota, featuring roughly 50 different species. The largest animal,
10647: 10591:"Jaw elements in Plumulites bengtsoni confirm that machaeridians are extinct armoured scaleworms" 10510: 10471: 10378: 10218: 10174: 10130: 10086: 9618: 9576: 9537: 8670:
Fortey, Richard A. (2009). "A New Giant Asaphid Trilobite from the Lower Ordovician of Morocco".
8484: 8301: 7871:"The Fezouata fossils of Morocco; an extraordinary record of marine life in the Early Ordovician" 6075: 5368: 10259:
Nanglu, Karma; Waskom, Madeleine E.; Richards, Jared C.; Ortega-Hernández, Javier (2023-10-11).
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Van Roy, P.; Orr, P. J.; Botting, J. P.; Muir, L. A.; Vinther, J.; Lefebvre, B.; Hariri, K. E.;
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A basal graptoloid with a mesh-like conical colony form transitional between dendroids (such as
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fossils, filling an important preservational window beyond the earlier and more common Cambrian
10131:"An unusual onychochilid mollusc from the Ordovician (Tremadocian) Fezouata Formation, Morocco" 8893:
Gutiérrez-Marco, Juan C.; García-Bellido, Diego C.; Rábano, Isabel; Sá, Artur A. (2017-01-10).
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Conodonts from the Fezouata Formation are mostly coniform elements of Early Ordovician species.
821: 8120:"A spinose appendage fragment of a problematic arthropod from the Early Ordovician of Morocco" 2669:
An aglaspidid, one of the first expectionally-preserved invertebrates found in the formation.
1687:, the only diagnostic member of the family to be found in Africa. Previously misidentified as 10812: 8833:"First occurrence of well-preserved Ordovician trilobites of the family Olenidae from Africa" 5521: 448: 4704:
protobranch bivalve known from a single specimen. It was likely an infaunal deposit feeder.
10848: 10659: 10522: 10483: 10434: 10390: 10230: 10186: 10142: 10098: 10012: 10001:"Ancestral morphology of crown-group molluscs revealed by a new Ordovician stem aculiferan" 9966: 9905: 9782: 9726: 9630: 9588: 9549: 9415: 9347: 9205: 9145: 9091: 8906: 8797: 8705: 8636: 8597: 8496: 8357: 8313: 8199: 8061: 7994: 7882: 7672: 7613: 7569: 7518: 7476: 7358: 6375: 121: 9775:"Origin of the subphylum Asterozoa and redescription of a Moroccan Ordovician somasteroid" 7870: 8: 8831:
Hopkins, Melanie J.; Gutiérrez-Marco, Juan Carlos; Di Silvestro, Gianpaolo (2024-01-15).
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A common basal graptoloid with a mesh-like conical colony form. Potentially a species of
3661: 3533: 676:, this genus was most likely a filter feeder, but it could have preyed upon larger food. 655: 10663: 10526: 10487: 10438: 10394: 10293: 10260: 10234: 10190: 10146: 10102: 10016: 9970: 9909: 9786: 9730: 9634: 9592: 9553: 9419: 9376: 9351: 9335: 9209: 9149: 9095: 9037:
Drage, Harriet B.; Vandenbroucke, Thijs R.A.; Van Roy, Peter; Daley, Allison C. (2019).
8910: 8801: 8709: 8640: 8601: 8500: 8361: 8317: 8203: 8065: 7998: 7983:"Anomalocaridid trunk limb homology revealed by a giant filter-feeder with paired flaps" 7886: 7676: 7617: 7573: 7522: 7480: 7437: 7362: 3540: 2166: 10731: 10623: 10590: 10054: 9852: 9485: 9450: 9169: 8935: 8894: 8386: 8345: 8090: 8049: 7906: 7542: 7382: 5581:
A multiramous anisograptid. Very rare, only a single fossil may indicate its presence.
5553: 368:(just over 2 metres in length), is depicted in a pair swimming just above the seafloor. 4533:
A rare afghanodesmatid protobranch bivalve. It was likely an infaunal deposit feeder.
10741: 10628: 10610: 10571: 10342: 10298: 10280: 10058: 10046: 10038: 9893: 9857: 9839: 9819: 9800: 9752: 9686: 9490: 9472: 9431: 9381: 9363: 9336:"Lower Ordovician synziphosurine reveals early euchelicerate diversity and evolution" 9311: 9270: 9229: 9221: 9193: 9173: 9161: 9133: 9107: 9018: 8998: 8964: 8940: 8922: 8850: 8813: 8785: 8759: 8721: 8693: 8652: 8391: 8373: 8215: 8095: 8077: 8018: 8010: 7982: 7958: 7910: 7898: 7690: 7637: 7629: 7546: 7534: 7424: 7402: 7374: 4861: 4492: 794: 9194:"An acercostracan marrellomorph (Euarthropoda) from the Lower Ordovician of Morocco" 7710:"Storm-Induced Community Dynamics in the Fezouata Biota (Lower Ordovician, Morocco)" 7346: 3974:-like echinoderm with a broad pentagonal body. Originally described in a new genus, 2608:. One of the few liwiid fossils with soft tissue of the digestive system preserved. 2308:
Many arthropods of the Fezouata Biota remain unnamed and undescribed. These include
376:
specimens, representing 50 distinct taxa that have a composition similar to earlier
247: 10807: 10751: 10667: 10618: 10602: 10563: 10530: 10491: 10450: 10442: 10398: 10332: 10288: 10272: 10238: 10194: 10150: 10106: 10028: 10020: 9974: 9913: 9847: 9831: 9790: 9742: 9734: 9676: 9638: 9596: 9557: 9480: 9462: 9423: 9371: 9355: 9301: 9260: 9213: 9153: 9099: 9053: 9010: 8930: 8914: 8840: 8805: 8713: 8644: 8605: 8504: 8430: 8381: 8365: 8321: 8207: 8085: 8069: 8002: 7948: 7890: 7813: 7791:
Exceptionally Preserved Faunas from the Lower Ordovician of the Anti-Atlas, Morocco
7680: 7621: 7577: 7526: 7484: 7432: 7414: 7386: 7366: 7204: 4269: 1895:
A very large ogygiocaridinine asaphid, up to 49.1 centimetres (19.3 in) long.
627: 10681: 10379:"Diversity and ecology of sponges in the Early Ordovician Fezouata Biota, Morocco" 10154: 9999:
Vinther, Jakob; Parry, Luke; Briggs, Derek E. G.; Van Roy, Peter (February 2017).
9795: 9774: 9642: 9600: 9079: 8508: 7488: 7403:"Highly resolved taphonomic variations within the Early Ordovician Fezouata Biota" 571:, an aegirocassisine, and a predatory hurdiid formerly mistaken for a eurypterid. 10853: 10756: 10671: 10534: 10495: 10446: 10402: 10242: 10198: 10110: 9917: 9561: 9103: 9080:"Ordovician edrioasteroids from Morocco: faunal exchanges across the Rheic Ocean" 8648: 8609: 8583:"The Fezouata Shale (Lower Ordovician, Anti-Atlas, Morocco): A historical review" 8325: 7581: 7299: 7276: 7253: 7230: 6555: 5666: 4346: 4312: 4212: 4208: 2321: 2159: 1335: 905: 519: 10589:
Parry, Luke A.; Edgecombe, Gregory D.; Sykes, Dan; Vinther, Jakob (2019-07-24).
8346:"Collective behaviour in 480-million-year-old trilobite arthropods from Morocco" 2721:, with few other animal fossils. Through nearly the entire the formation, small 1588: 1220:
An endemic bavarillid common in relatively shallow (lower nearshore) sediments.
10276: 9449:
Lerosey-Aubril, Rudy; Skabelund, Jacob; Ortega-Hernández, Javier (2020-04-09).
9359: 8786:"Trilobites of the genus Dikelokephalina from Ordovician Gondwana and Avalonia" 8369: 8073: 7419: 7041: 6803: 6786:
Many animals from the Fezouata biota remain unnamed and undescribed, including
6226: 6138: 5999: 5774: 5517: 5379: 4977: 4633: 4602: 4168: 4088: 3985: 3946: 2536: 2456: 2445: 2309: 1959: 1877:
A very large ogygiocaridinine asaphid, up to 45 centimetres (18 in) long.
1167: 816: 205: 9681: 9664: 9427: 9306: 9289: 9265: 9248: 9217: 9157: 9014: 8414:
Pérez-Peris, Francesc; Laibl, Lukáš; Vidal, Muriel; Daley, Allison C. (2021).
8211: 7953: 7936: 7601: 1469: 337: 10827: 10771: 10761: 10736: 10726: 10614: 10575: 10346: 10284: 10042: 10000: 9843: 9804: 9756: 9690: 9476: 9435: 9367: 9315: 9274: 9225: 9165: 9111: 8926: 8854: 8817: 8725: 8656: 8377: 8219: 8081: 8014: 7962: 7902: 7735:
Saleh, Farid; Pittet, Bernard; Perrillat, Jean-Philippe; Lefebvre, Bertrand.
7694: 7685: 7660: 7633: 7428: 7131: 6585: 6422: 6393: 6335: 6158: 5834: 5637: 5362: 5186: 4919:. Most species appear to correspond to the time interval stretching from the 4806: 4701: 4668: 4078: 3967: 2951: 2707: 2540: 2467: 2341: 2337: 2333: 2038: 1649: 637: 491:, and tin; this aspect of the fossil preservation is very similar to that at 428: 424: 377: 227: 214: 188: 175: 109: 62: 27: 4349:
known from tiny juveniles which died in relatively deep, oxygen-poor waters.
10766: 10746: 10632: 10606: 10470:
Martin, Emmanuel L.O.; Lerosey-Aubril, Rudy; Vannier, Jean (October 2016).
10302: 10050: 9979: 9861: 9835: 9494: 9385: 9233: 9132:
Ortega-Hernández, Javier; Van Roy, Peter; Lerosey-Aubril, Rudy (May 2016).
9057: 8944: 8434: 8395: 8099: 8022: 7641: 7538: 7506: 7378: 7342: 7100: 7022: 6205: 6153: 6056: 5748: 5524: 5349: 5155: 4135: 2638: 2442: 2355: 2325: 2292: 2148: 1255: 1228: 1152: 1127: 837: 603: 452: 415: 10567: 10337: 7817: 5967:
A "psigraptid"-type anisograptid, with a small number of reclined stipes.
10792: 9404:"A new nektaspid euarthropod from the Lower Ordovician strata of Morocco" 7869:
Van Roy, Peter; Briggs, Derek E. G.; Gaines, Robert R. (September 2015).
7787: 7303: 7280: 7257: 7234: 7160: 6854: 6275:
An enigmatic feather-shaped organism identified as a sessile graptolite.
4499:. The least rare nautiloid found in the formation, though still uncommon. 4496: 4193: 4180: 4153: 3013: 2789: 2726: 2699: 2545: 2532: 2503: 2495: 2482: 2177: 1768:
sediment for food. This species' feeding behavior may be responsible for
1724: 1553: 1418: 1280: 1244: 858: 523: 515: 394: 103: 37: 10455: 10024: 8845: 8717: 8623:
Gibb, Stacey; Chatterton, Brian D. E.; Gingras, Murray K. (2010-11-30).
8006: 7894: 7625: 7530: 7370: 10797: 9747: 9738: 9467: 9448: 7193:"Age calibration of the Lower Ordovician Fezouata Lagerstätte, Morocco" 7118: 7107: 7082: 6799: 6607: 6582: 6578: 6558: 6390: 6291: 5466: 5354: 5338: 5174: 4731: 4261: 4257: 4252:
A mollusc in the group Sachitida (the proposed clade containing living
4176: 2763: 2409: 2317: 1437: 1011:
An asaphid. Known from multiple life stages, including late meraspids.
934: 577: 503: 492: 432: 420: 410: 389: 385: 333: 82: 47: 8918: 8580: 7209: 7192: 2071: 1331:
A rare calymenid found in relatively deep (upper offshore) sediments.
1123:
A very large asaphine asaphid up to 51 centimetres (20 in) long.
680: 9131: 8892: 7763:"Shifts in Earth's Orbit Increase the Chances of Spectacular Fossils" 7164: 7111: 6807: 6687: 6666: 6163: 4987: 4253: 4172: 4131: 3472: 3350: 3107: 2645: 2616: 2594: 2573: 2551: 2368: 2313: 1581: 1577: 1462: 1403:
A large dikelokephalinid up to 29.5 centimetres (11.6 in) long.
1178: 955: 732: 633: 476: 456: 87: 10258: 10217:
Gutiérrez-Marco, Juan Carlos; Martin, Emmanuel L.O. (October 2016).
10172: 8809: 7797:. First international Conference and Exhibition, Marrakech, Morocco. 7737:"Orbital Control on Exceptional Fossil Preservation (abstract only)" 7190: 6720:
An indeterminate protomonaxonid. Previously considered a species of
1920:
Known from multiple life stages, including early to late meraspids.
8996: 7071: 7011: 6985: 6959: 6908: 6882: 6791: 5140: 4939: 4817: 4791: 4343: 4309: 4273: 4142: 4085: 4081: 4029: 3971: 3331: 2598: 2490: 2449: 2358: 2329: 1864: 1831: 1770: 1684: 1599: 1522: 952: 862: 402: 77: 72: 57: 52: 42: 9334:
Lustri, Lorenzo; Gueriau, Pierre; Daley, Allison C. (2024-05-07).
8830: 7657: 5552:
A "psigraptid"-type anisograptid, with a small number of reclined
2197:
A nileid found in relatively shallow (lower nearshore) sediments.
10553:"Pauxillites thaddei a new lower Ordovician hyolith from Morocco" 8050:"New fossil assemblages from the Early Ordovician Fezouata Biota" 6795: 6787: 6735: 6538: 6509: 6480: 6451: 6364: 4916: 4400:
protobranch bivalve. It may have been an infaunal filter-feeder.
4138: 3752: 2890: 2836: 2807: 2687: 2561: 1856:
A large asaphine asaphid up to 33 centimetres (13 in) long.
1144:
A large asaphine asaphid up to 36 centimetres (14 in) long.
1109:
A large asaphine asaphid up to 33 centimetres (13 in) long.
630: 444: 406: 398: 381: 360: 336:. It was deposited in a marine environment, and is known for its 329: 265: 92: 67: 8188:"Large trilobites in a stress-free Early Ordovician environment" 7935:
Potin, Gaëtan J.-M.; Gueriau, Pierre; Daley, Allison C. (2023).
6192:
A common multiramous dichograptid, also sometimes classified as
1652:
known from multiple life stages, including mid-stage meraspids.
10508: 7981:
Van Roy, Peter; Daley, Allison C.; Briggs, Derek E. G. (2015).
6616: 6554:
Many sponges from the Fezouata biota remain unnamed, including
4962: 4560: 4265: 2605: 2591: 527: 488: 436: 9534: 9036: 9009:, vol. 64, John Wiley & Sons, Ltd, pp. 155–171, 4130:
A common bivalve, tentatively identified as one of the oldest
344:. The fauna of this geological unit is often described as the 9401: 9288:
Drage, Harriet B.; Legg, David A.; Daley, Allison C. (2023).
8343: 7399: 7168: 6642: 154: 9616: 9246: 8482: 7600:
Vinther, Jakob; Van Roy, Peter; Briggs, Derek E. G. (2008).
10469: 9818:
Hunter, Aaron W.; Ortega-Hernández, Javier (January 2021).
9773:
Blake, Daniel B.; Hotchkiss, Frederick H. C. (2022-08-01).
9712: 9574: 8873:
Memoirs of the Association of Australasian Palaeontologists
8672:
Memoirs of the Association of Australasian Palaeontologists
7808:
Jones, N. (2010). "Weird wonders lived past the Cambrian".
3114:
zone. Also abundant in higher-energy deposits of the ?
2486: 1834:
known from multiple life stages, including late meraspids.
865:. Some specimens may have previously been misidentified as 636:. The largest known radiodont and one of the largest known 471:
South Polar paleogeography of the Middle Ordovician, 460 Ma
10588: 10560:
Acta Musei Nationalis Pragae, Series B, Historia Naturalis
10129:
Ebbestad, Jan Ove R.; Lefebvre, Bertrand (November 2015).
9662: 8413: 8299: 10645: 9998: 7145:
A tubular organism likely closely related to conulariids.
3866:, it was likely a deep-water specialist buried in place. 2140:
found in relatively shallow (lower nearshore) sediments.
908:
found in relatively shallow (lower nearshore) sediments.
443:. Other Ordovician oddballs are also present, including 388:, forms that are absent in the Cambrian period. Diverse 8184: 7788:
Van Roy, P; Lefebvre, B; El Hariri, K; Hafid, A (2008).
7734: 6229:
graptoloid with two to four reclined to pendent stipes.
2231:
A small nileid up to 4.5 centimetres (1.8 in) long
1359:
A large asaphid up to 29 centimetres (11 in) long.
10595:
Proceedings of the Royal Society B: Biological Sciences
8046: 7707: 10420: 9817: 5669:
graptoloid, with two subhorizontal to pendent stipes.
10216: 8622: 7340: 4835:"nautiloid" cephalopod in the order Ellesmerocerida. 1805:
A calymenid found in particularly shallow sediments.
7714:(limited access to full article) geoscienceworld.org 7599: 10711: 10318: 8760:"Giant Trilobites from Lower Ordovician of Morocco" 8692:Edgecombe, Gregory D.; Fortey, Richard. A. (2023). 7868: 7741:
geoscienctworld.org (subscription w/ free abstract)
7653: 7651: 475:The fossiliferous strata were deposited just above 9333: 7980: 4563:bivalve. It was likely an infaunal filter-feeder. 4276:, and a single flat calcareous shell on the head. 2737:outnumbers other fossils at any given time. Large 112:from one of the exceptionally preserved intervals. 10652:Palaeogeography, Palaeoclimatology, Palaeoecology 10515:Palaeogeography, Palaeoclimatology, Palaeoecology 10476:Palaeogeography, Palaeoclimatology, Palaeoecology 10427:Palaeogeography, Palaeoclimatology, Palaeoecology 10383:Palaeogeography, Palaeoclimatology, Palaeoecology 10223:Palaeogeography, Palaeoclimatology, Palaeoecology 10179:Palaeogeography, Palaeoclimatology, Palaeoecology 10091:Palaeogeography, Palaeoclimatology, Palaeoecology 9950:Kröger, Björn; Lefebvre, Bertrand (August 2012). 9898:Palaeogeography, Palaeoclimatology, Palaeoecology 9542:Palaeogeography, Palaeoclimatology, Palaeoecology 8590:Palaeogeography, Palaeoclimatology, Palaeoecology 8306:Palaeogeography, Palaeoclimatology, Palaeoecology 7934: 5884:A rare didymograptid with two horizontal stipes. 10825: 10128: 8969:Scripta Musei Geologici Seminarii Barcinonensis 7648: 7602:"Machaeridians are Palaeozoic armoured annelids" 542: 8691: 7559: 6079:) and early graptoloids such as anisograptids. 2295:without clear preferences for specific depths. 1962:without clear preferences for specific depths. 9949: 9772: 9287: 7832:"The First 100 IUGS Geological Heritage Sites" 7743:. Geology (magazine published) 1 February 2019 5353:) may have been transitional between sessile ( 2493:. The name "Furca mauretanica" is an informal 10697: 9077: 8117: 7509:(2011). "A giant Ordovician anomalocaridid". 7504: 6048:A large and common multiramous anisograptid. 5341:) groups, though some mesh-shaped taxa (like 4264:). It had a flattened body with an extensive 8757: 7839:IUGS International Commission on Geoheritage 7336: 7334: 7332: 7186: 7184: 7117: 5357:) graptolites and free-floating graptoloids. 4927:zone (uppermost Tremadocian) up through the 3984: 3539: 2753:are common components of these assemblages. 2644: 2502: 2455: 643: 509: 7330: 7328: 7326: 7324: 7322: 7320: 7318: 7316: 7314: 7312: 4435:. It was likely an infaunal filter-feeder. 3890:A common and endemic eocrinoid, similar to 10704: 10690: 9078:Sumrall, Colin D.; Zamora, Samuel (2011). 8758:Corbacho, Joan; Vela, Joan Antoni (2010). 1199:up to 4.1 centimetres (1.6 in) long. 549:International Union of Geological Sciences 302: 293: 284: 102: 10622: 10454: 10336: 10292: 10034:1983/70368bb0-c2df-490a-8b45-26e7b4397596 10032: 9978: 9891: 9851: 9794: 9746: 9680: 9484: 9466: 9375: 9305: 9264: 8934: 8844: 8385: 8089: 7952: 7684: 7436: 7418: 7347:"Ordovician faunas of Burgess Shale type" 7208: 7181: 6857:, up to 50 centimetres (20 in) tall. 3840:A rare and endemic eocrinoid, similar to 3818:A rare and endemic eocrinoid, similar to 10869:First 100 IUGS Geological Heritage Sites 10319:Muir, L A; GutiĂ©rrez-Marco, J C (2023). 10084: 8962: 8118:Van Roy, Peter; Tetlie, O. Erik (2006). 7309: 6816:Other animals of the Fezouata Formation 6815: 6615: 6290: 5465: 4938: 4028: 2762: 2366: 731: 576: 466: 359: 10376: 9530: 9528: 9526: 9524: 8576: 8574: 8572: 8570: 8568: 8566: 8564: 8562: 8560: 8558: 8556: 8554: 8552: 8550: 8548: 8546: 8544: 8542: 8540: 8538: 8478: 8476: 8474: 8472: 8470: 8468: 8466: 8464: 8295: 8293: 8291: 8289: 8287: 8285: 8283: 8281: 8279: 8277: 8275: 8273: 8271: 8269: 8267: 8265: 8263: 8261: 8259: 8257: 8255: 8253: 8251: 8249: 8180: 8178: 8176: 8174: 8172: 8170: 8168: 8166: 8164: 8162: 8160: 7466: 7171:have been recorded from the formation. 5382:, are also present but much more rare. 640:, at 2 metres (6.6 ft) in length. 462: 10826: 10550: 10546: 10544: 10372: 10370: 10368: 10366: 10364: 10362: 10360: 10358: 10356: 10314: 10312: 10254: 10252: 9708: 9706: 9704: 9702: 9700: 9658: 9656: 9654: 9652: 9612: 9610: 9522: 9520: 9518: 9516: 9514: 9512: 9510: 9508: 9506: 9504: 9397: 9395: 9329: 9327: 9325: 9187: 9185: 9183: 9127: 9125: 9123: 9121: 9073: 9071: 9069: 9067: 8992: 8990: 8988: 8986: 8984: 8982: 8958: 8956: 8954: 8888: 8886: 8870: 8866: 8864: 8783: 8779: 8777: 8753: 8751: 8749: 8747: 8745: 8743: 8741: 8739: 8737: 8735: 8687: 8685: 8669: 8536: 8534: 8532: 8530: 8528: 8526: 8524: 8522: 8520: 8518: 8462: 8460: 8458: 8456: 8454: 8452: 8450: 8448: 8446: 8444: 8409: 8407: 8405: 8339: 8337: 8335: 8247: 8245: 8243: 8241: 8239: 8237: 8235: 8233: 8231: 8229: 8158: 8156: 8154: 8152: 8150: 8148: 8146: 8144: 8142: 8140: 8113: 8111: 8109: 8042: 8040: 8038: 8036: 8034: 8032: 7976: 7974: 7972: 7930: 7928: 7926: 7924: 7922: 7920: 7595: 7593: 7591: 10685: 10416: 10414: 10412: 10212: 10210: 10208: 10168: 10166: 10164: 10124: 10122: 10120: 10085:Ebbestad, Jan Ove R. (October 2016). 10080: 10078: 10076: 10074: 10072: 10070: 10068: 9994: 9992: 9990: 9945: 9943: 9941: 9939: 9937: 9935: 9933: 9931: 9929: 9927: 9887: 9885: 9883: 9881: 9879: 9877: 9875: 9873: 9871: 9768: 9766: 8694:"A novel antennal form in trilobites" 7864: 7862: 7860: 7858: 7856: 7807: 7500: 7498: 7462: 7460: 7458: 7456: 7454: 7452: 7450: 7448: 5391:, Rhabdinopora flabelliformis anglica 2476:Sagenograptus (Araneograptus) murrayi 711:Sagenograptus (Araneograptus) murrayi 696:Sagenograptus (Araneograptus) murrayi 667:Sagenograptus (Araneograptus) murrayi 9191: 6146:Sagenopterus (Araneograptus) murrayi 5450:zone) and the mid-late Floian (late 4921:Oelandodus elongatus-Acodus deltatus 4730:"nautiloid" cephalopod in the order 4667:"nautiloid" cephalopod in the order 4495:"nautiloid" cephalopod in the order 4211:"nautiloid" cephalopod in the order 10541: 10353: 10309: 10249: 9697: 9649: 9607: 9501: 9392: 9322: 9180: 9118: 9084:Journal of Systematic Palaeontology 9064: 8979: 8951: 8883: 8861: 8774: 8732: 8682: 8515: 8441: 8402: 8332: 8226: 8137: 8106: 8029: 7969: 7917: 7588: 6806:, and at least five new species of 6727:but likely belongs to a new genus. 6112:Rhabdinopora flabelliformis anglica 6068:Rhabdinopora flabelliformis anglica 4891:A common tergomyan monoplacophoran. 4370:A common tergomyan monoplacophoran. 2303: 1677:Rhabdinopora flabelliformis anglica 13: 10409: 10325:Estonian Journal of Earth Sciences 10205: 10161: 10117: 10065: 9987: 9924: 9892:Polechová, Marika (October 2016). 9868: 9763: 9669:Frontiers in Ecology and Evolution 9294:Frontiers in Ecology and Evolution 9253:Frontiers in Ecology and Evolution 7941:Frontiers in Ecology and Evolution 7853: 7495: 7445: 7286: 7263: 7240: 7217: 7155: 2543:). Though most closely related to 2165: 2070: 1619: 1587: 1468: 1406: 1334: 1126: 1032: 679: 14: 10880: 7875:Journal of the Geological Society 559:After and subsequent literature: 6781: 6102:R. flabelliformis flabelliformis 3894:in form and preservation style. 3844:in form and preservation style. 3822:in form and preservation style. 2559:, it also shows similarities to 301: 292: 283: 276: 258: 35: 10713:Burgess Shale-type preservation 10639: 10582: 10502: 10463: 9811: 9568: 9442: 9281: 9240: 9030: 8824: 8663: 8616: 7824: 7801: 7781: 7755: 7728: 7701: 7553: 5773:A large and common multiramous 5740:A small and rare anisograptid. 482: 16:Geological formation in Morocco 8963:Corbacho, Joan; Vela, Joan A. 7393: 6283: 5332: 2677: 2416:, a short trunk, and no eyes. 1: 10551:Valent, Martin (2015-10-31). 10155:10.1016/j.geobios.2015.09.004 9796:10.1016/j.geobios.2022.07.002 9643:10.1016/j.geobios.2023.08.001 9601:10.1016/j.geobios.2018.11.001 9046:Acta Palaeontologica Polonica 8509:10.1016/j.geobios.2023.06.005 8423:Acta Palaeontologica Polonica 8127:Acta Palaeontologica Polonica 7489:10.1016/j.geobios.2007.02.006 7174: 6810:, including an armored form. 3978:but subsequently referred to 721: 562: 554: 543:IUGS geological heritage site 328:is a geological formation in 10672:10.1016/j.palaeo.2016.03.007 10535:10.1016/j.palaeo.2018.07.012 10496:10.1016/j.palaeo.2016.04.009 10447:10.1016/j.palaeo.2016.03.008 10403:10.1016/j.palaeo.2016.05.018 10243:10.1016/j.palaeo.2016.07.026 10199:10.1016/j.palaeo.2016.03.023 10111:10.1016/j.palaeo.2016.01.003 9918:10.1016/j.palaeo.2015.12.016 9562:10.1016/j.palaeo.2016.02.039 9104:10.1080/14772019.2010.499137 8649:10.1080/10420940.2010.535452 8610:10.1016/j.palaeo.2015.10.048 8326:10.1016/j.palaeo.2016.06.003 7582:10.1016/j.annpal.2007.06.003 6272:"probable late Tremadocian" 6087:R. flabelliformis canadensis 5934:A multiramous anisograptid. 5909:A multiramous dichograptid. 4901: 4469:sulfide-oxidizing bacteria. 2405:Upper Fezouata (mid-Floian) 2278:?Cymatograptus protobalticus 2094:?Cymatograptus protobalticus 1945:?Cymatograptus protobalticus 1794:?Cymatograptus protobalticus 1616:of questionable occurrence. 1497:?Cymatograptus protobalticus 1325:?Cymatograptus protobalticus 1296:?Cymatograptus protobalticus 980:?Cymatograptus protobalticus 898:?Cymatograptus protobalticus 310:Fezouata Formation (Morocco) 206:Approximate paleocoordinates 7: 10839:Ordovician System of Africa 10377:Botting, Joseph P. (2016). 8784:Fortey, Richard A. (2011). 5820:Cymatograptus protobalticus 5761:Cymatograptus protobalticus 5415:Cymatograptus protobalticus 4828:"late early to mid Floian" 4757:Cymatograptus protobalticus 4723:"late early to mid Floian" 4695:Cymatograptus protobalticus 4660:"late early to mid Floian" 4584:Cymatograptus protobalticus 4554:Cymatograptus protobalticus 4522:Cymatograptus protobalticus 4488:"late early to mid Floian" 4456:Cymatograptus protobalticus 4422:Cymatograptus protobalticus 4365:Cymatograptus protobalticus 4336:Cymatograptus protobalticus 4297:Cymatograptus protobalticus 4227:"late early to mid Floian" 4204:"late early to mid Floian" 4164:"late early to mid Floian" 4112:Cymatograptus protobalticus 4021: 3739:Cymatograptus protobalticus 3600:Cymatograptus protobalticus 3399:Cymatograptus protobalticus 3344:Cymatograptus protobalticus 3112:Cymatograptus protobalticus 3093:Cymatograptus protobalticus 2987:An uncommon edrioasteroid. 2975:Cymatograptus protobalticus 771:Cymatograptus protobalticus 342:Burgess shale-type deposits 10: 10885: 10277:10.1038/s42003-023-05377-x 9360:10.1038/s41467-024-48013-w 8370:10.1038/s41598-019-51012-3 8074:10.1038/s41598-022-25000-z 7420:10.1038/s41598-024-71622-w 6619:of the Fezouata Formation 6549: 6294:of the Fezouata Formation 6123:R. flabelliformis socialis 5469:of the Fezouata Formation 5435:?Cymatograptus probalticus 4942:of the Fezouata Formation 4032:of the Fezouata Formation 2766:of the Fezouata Formation 2597:with a large cephalon and 2448:with a shield-like dorsal 2371:of the Fezouata Formation 735:of the Fezouata Formation 580:of the Fezouata Formation 534: 10785: 10719: 9682:10.3389/fevo.2023.1290063 9428:10.1017/S001675682000062X 9307:10.3389/fevo.2023.1226924 9266:10.3389/fevo.2023.1232612 9218:10.1007/s00114-016-1352-5 9158:10.1017/S0016756815000710 9015:10.1002/9781119564249.ch7 8212:10.1017/S0016756820000448 7954:10.3389/fevo.2023.1214109 7142:Lower and Upper Fezouata 6930:Lower and Upper Fezouata 6850:Lower and Upper Fezouata 6502:Lower and Upper Fezouata 6224: 6202: 6072: 6063:R. flabelliformis anglica 6054: 5997: 5973: 5772: 5746: 5664: 5646: 5314: 5302: 5264: 5252: 5150: 5138: 5122: 5110: 5091: 5079: 5063: 5051: 4282: 4206: 4191: 3900: 3761: 3384: 3261: 3149: 2870: 2557:Herefordshire Lagerstätte 2175: 1862: 1722: 1365: 1253: 1176: 1090: 932: 879: 671: 653: 510:Location and stratigraphy 378:Burgess Shale type biotas 271: 253: 243: 204: 165: 160: 150: 145: 137: 127: 117: 101: 26: 21: 7686:10.1016/j.gr.2015.03.009 7562:Annales de PalĂ©ontologie 7293:Upper Fezouata Formation 7270:Lower Fezouata Formation 6167:from the Burgess Shale. 4272:covered in bristle-like 3660:A common minervaecystid 2950:An uncommon and endemic 669:zone and Upper Fezouata 439:as well as the expected 355: 10864:Paleontology in Morocco 10844:Lower Ordovician Series 9719:Journal of Paleontology 9192:Legg, David A. (2016). 8837:Journal of Paleontology 8698:Journal of Paleontology 6128:Anisograptus matanensis 6107:Anisograptus matanensis 6092:Anisograptus matanensis 5576:Anisograptus matanensis 5387:Anisograptus matanensis 5360:Sessile forms, such as 5020:A drepanoistodontid or 3606:An uncommon eocrinoid. 1672:Anisograptus matanensis 1580:arthropod in the order 626:A giant filter-feeding 338:exceptionally preserved 10607:10.1098/rspb.2019.1247 10265:Communications Biology 9980:10.1002/mmng.201200004 9836:10.1098/rsbl.2020.0809 9058:10.4202/app.00582.2018 8999:"The nileid trilobite 8435:10.4202/app.00902.2021 7122: 6606:of diverse and sturdy 5858:A rare didymograptid. 5825:A rare didymograptid. 5103:P. aff. simplicissimus 4886:?Baltograptus jacksoni 4855:?Baltograptus jacksoni 4779:bivalve in the family 4762:?Baltograptus jacksoni 4589:?Baltograptus jacksoni 4527:?Baltograptus jacksoni 4431:bivalve in the family 4302:?Baltograptus jacksoni 4117:?Baltograptus jacksoni 3989: 3544: 2649: 2637:, possibly related to 2507: 2460: 2170: 2130:?Baltograptus jacksoni 2120:P. aff. languedocensis 2075: 1624: 1592: 1570:?Baltograptus jacksoni 1539:?Baltograptus jacksoni 1473: 1411: 1339: 1131: 1058:nearshore) sediments. 1037: 946:?Baltograptus jacksoni 815:A long-spined eyeless 776:?Baltograptus jacksoni 684: 648: 472: 369: 366:Aegirocassis benmoulai 10813:Llanfallteg Formation 10786:Ordovician localities 10568:10.14446/AMNP.2015.51 10338:10.3176/earth.2023.24 9340:Nature Communications 9198:The Science of Nature 7818:10.1038/news.2010.234 7121: 7094:Araneograptus murrayi 7035:Araneograptus murrayi 6250:Baltograptus jacksoni 6244:Araneograptus murrayi 6220:Araneograptus murrayi 6042:Hunnegraptus copiosus 6037:Araneograptus murrayi 6015:Araneograptus murrayi 5992:Hunnegraptus copiosus 5987:Araneograptus murrayi 5961:Hunnegraptus copiosus 5956:Araneograptus murrayi 5929:Hunnegraptus copiosus 5879:Baltograptus jacksoni 5852:Hunnegraptus copiosus 5847:Araneograptus murrayi 5795:Hunnegraptus copiosus 5790:Araneograptus murrayi 5767:Baltograptus jacksoni 5735:Aorograptus victoriae 5683:Baltograptus jacksoni 5601:Araneograptus murrayi 5547:Aorograptus victoriae 5448:Hunnegraptus copiosus 5444:Araneograptus murrayi 5431:Hunnegraptus copiosus 5419:Baltograptus jacksoni 5411:Hunnegraptus copiosus 5407:Araneograptus murrayi 5403:Aorograptus victoriae 5265:A protopanderodontid 5246:A protopanderodontid 5208:A protopanderodontid 4627:Araneograptus murrayi 4330:Araneograptus murrayi 4247:Araneograptus murrayi 4008:Araneograptus murrayi 3988: 3958:Araneograptus murrayi 3933:Baltograptus jacksoni 3914:Araneograptus murrayi 3884:Hunnegraptus copiosus 3879:Araneograptus murrayi 3857:Araneograptus murrayi 3835:Araneograptus murrayi 3813:Araneograptus murrayi 3793:Baltograptus jacksoni 3786:Hunnegraptus copiosus 3780:Araneograptus murrayi 3775:Aorograptus victoriae 3745:Baltograptus jacksoni 3713:Araneograptus murrayi 3688:Araneograptus murrayi 3666:Hunnegraptus copuosus 3651:Araneograptus murrayi 3631:An uncommon mitrate. 3626:Araneograptus murrayi 3594:Araneograptus murrayi 3570:Baltograptus jacksoni 3564:Araneograptus murrayi 3543: 3524:Araneograptus murrayi 3499:Araneograptus murrayi 3477:Araneograptus murrayi 3462:Baltograptus jacksoni 3456:Araneograptus murrayi 3430:Baltograptus jacksoni 3424:Araneograptus murrayi 3405:Baltograptus jacksoni 3373:Araneograptus murrayi 3323:An uncommon cornute. 3318:Araneograptus murrayi 3298:An uncommon cornute. 3293:Araneograptus murrayi 3275:Araneograptus murrayi 3250:Araneograptus murrayi 3230:An uncommon mitrate. 3199:Araneograptus murrayi 3186:An uncommon cornute. 3181:Araneograptus murrayi 3163:Araneograptus murrayi 3138:Araneograptus murrayi 3116:Baltograptus jacksoni 3100:Baltograptus jacksoni 3087:Araneograptus murrayi 3062:Baltograptus jacksoni 3036:Araneograptus murrayi 3007:Araneograptus murrayi 2981:Baltograptus jacksoni 2945:Baltograptus jacksoni 2918:Baltograptus jacksoni 2911:Hunnegraptus copiosus 2906:Araneograptus murrayi 2884:Hunnegraptus copuosus 2864:An uncommon cornute. 2859:Araneograptus murrayi 2830:Araneograptus murrayi 2801:Araneograptus murrayi 2731:Araneograptus murrayi 2715:Araneograptus murrayi 2648: 2628:Araneograptus murrayi 2585:Araneograptus murrayi 2531:A small but abundant 2526:Araneograptus murrayi 2506: 2459: 2436:Araneograptus murrayi 2285:Baltograptus jacksoni 2273:Araneograptus murrayi 2244:Araneograptus murrayi 2210:Araneograptus murrayi 2189:Araneograptus murrayi 2169: 2125:Araneograptus murrayi 2074: 2061:Araneograptus murrayi 1952:Baltograptus jacksoni 1940:Araneograptus murrayi 1915:Araneograptus murrayi 1890:Araneograptus murrayi 1825:Araneograptus murrayi 1762:Araneograptus murrayi 1741:Araneograptus murrayi 1643:Araneograptus murrayi 1623: 1591: 1565:Araneograptus murrayi 1534:Araneograptus murrayi 1504:Baltograptus jacksoni 1492:Araneograptus murrayi 1472: 1410: 1379:Araneograptus murrayi 1338: 1320:Araneograptus murrayi 1268:Baltograptus jacksoni 1212:Araneograptus murrayi 1190:Araneograptus murrayi 1130: 1104:Baltograptus jacksoni 1049:Araneograptus murrayi 1036: 1024:Araneograptus murrayi 1006:Araneograptus murrayi 987:Baltograptus jacksoni 975:Araneograptus murrayi 893:Araneograptus murrayi 849:Araneograptus murrayi 810:Baltograptus jacksoni 808:zone up to the ? 806:Araneograptus murrayi 683: 647: 615:Araneograptus murrayi 501:Hunnegraptus copiosus 497:Araneograptus murrayi 470: 363: 6187:Baltograptus minutus 5904:Baltograptus minutus 5713:Baltograptus minutus 5698:Baltograptus minutus 5660:Baltograptus minutus 5452:Baltograptus minutus 5423:Baltograptus minutus 5123:A drepanoistodontid 5064:A drepanoistodontid 4931:zone (lower Floian). 4595:Baltograptus minutus 3655:Baltograptus minutus 3576:A rare diploporite. 2099:Baltograptus minutus 1799:Baltograptus minutus 1736:M. (E.) cf. filacovi 1431:Baltograptus minutus 1301:Baltograptus minutus 1273:Baltograptus minutus 620:Baltograptus minutus 506:zones respectively. 463:Depositional setting 350:Fezouata Lagerstätte 122:Geological formation 33:Tremadocian – Floian 10720:Cambrian localities 10664:2016PPP...460...62N 10527:2018PPP...508...48K 10488:2016PPP...460..130M 10439:2016PPP...460..170V 10395:2016PPP...460...75B 10235:2016PPP...460...35G 10191:2016PPP...460...50L 10147:2015Geobi..48..427E 10103:2016PPP...460...87E 10025:10.1038/nature21055 10017:2017Natur.542..471V 9971:2012FossR..15...61K 9910:2016PPP...460..155P 9787:2022Geobi..72...22B 9731:2017JPal...91..685A 9635:2023Geobi..81...51S 9593:2019Geobi..52...27L 9554:2016PPP...460...97L 9420:2021GeoM..158..509P 9408:Geological Magazine 9352:2024NatCo..15.3808L 9210:2016SciNa.103...21L 9150:2016GeoM..153..524O 9138:Geological Magazine 9096:2011JSPal...9..425S 8911:2017NatSR...739728G 8846:10.1017/jpa.2023.60 8802:2011GeolJ..46..405F 8718:10.1017/jpa.2022.59 8710:2023JPal...97..152E 8641:2010Ichno..17..271G 8602:2016PPP...460....7L 8501:2023Geobi..81...31L 8362:2019NatSR...914941V 8318:2016PPP...460..142M 8204:2021GeoM..158..261S 8192:Geological Magazine 8066:2022NatSR..1220773S 8007:10.1038/nature14256 7999:2015Natur.522...77V 7895:10.1144/jgs2015-017 7887:2015JGSoc.172..541V 7677:2016GondR..34..274M 7626:10.1038/nature06474 7618:2008Natur.451..185V 7574:2007AnPal..93..183L 7531:10.1038/nature09920 7523:2011Natur.473..510V 7481:2007Geobi..40..737B 7371:10.1038/nature09038 7363:2010Natur.465..215V 5480:Graptolite biozone 5425:zone (9), and the “ 4925:Paroistodus proteus 4084:. It was likely an 2601:and sharply angled 2316:(horseshoe crabs), 2249:Several additional 1871:O. forteyi hammondi 656:Pseudoangustidontus 569:Pseudoangustidontus 332:which dates to the 224: /  185: /  28:Stratigraphic range 10834:Fezouata Formation 10803:Fezouata Formation 10777:Zawiszyn Formation 10601:(1907): 20191247. 9739:10.1017/jpa.2017.6 9468:10.7717/peerj.8879 9007:Fossils and Strata 8899:Scientific Reports 8790:Geological Journal 8350:Scientific Reports 8054:Scientific Reports 7407:Scientific Reports 7298:2022-05-18 at the 7275:2022-05-18 at the 7252:2022-05-18 at the 7229:2022-05-18 at the 7224:Fezouata Formation 7123: 5261:S. cf. cornuformis 5119:P. cf. parallellus 5022:protopanderodontid 4728:protocameroceratid 4713:Protocyptendoceras 4260:relatives such as 4256:and their extinct 3990: 3919:A common cornute. 3693:A common cornute. 3545: 3436:A rare eocrinoid. 3411:A rare eocrinoid. 3067:A common mitrate. 2924:A common mitrate. 2650: 2555:from the Silurian 2508: 2461: 2412:with a very large 2171: 2076: 1625: 1593: 1474: 1412: 1340: 1307:A rare calymenid. 1132: 1038: 960:Rusophycus carleyi 685: 649: 473: 370: 322:Fezouata Formation 22:Fezouata Formation 10821: 10820: 10742:Kinzers Formation 10011:(7642): 471–474. 9024:978-1-119-56423-2 8919:10.1038/srep39728 7665:Gondwana Research 7612:(7175): 185–188. 7517:(7348): 510–513. 7210:10.1111/let.12240 7153: 7152: 6794:"dermal plates", 6779: 6778: 6547: 6546: 6281: 6280: 5814:?C. protobalticus 5571:A. cf. matanensis 5330: 5329: 5227:An acanthodontid 5165:P. cf. selenopsis 5060:P. cf. inaequalis 5044:drepanoistodontid 4929:Prioniodus oepiki 4899: 4898: 4719:P. longicameratum 4019: 4018: 3677:Procothurnocystis 3552:Paleosphaeronites 2901:A. trapeziiformis 2675: 2674: 2301: 2300: 1789:N. cf. attenuatus 1084:-like trilobite. 926:A remopleuridid. 719: 718: 318: 317: 10876: 10859:Shale formations 10808:Winneshiek Shale 10752:Poleta Formation 10706: 10699: 10692: 10683: 10682: 10676: 10675: 10643: 10637: 10636: 10626: 10586: 10580: 10579: 10557: 10548: 10539: 10538: 10506: 10500: 10499: 10467: 10461: 10460: 10458: 10418: 10407: 10406: 10374: 10351: 10350: 10340: 10316: 10307: 10306: 10296: 10256: 10247: 10246: 10214: 10203: 10202: 10170: 10159: 10158: 10126: 10115: 10114: 10082: 10063: 10062: 10036: 9996: 9985: 9984: 9982: 9956: 9947: 9922: 9921: 9889: 9866: 9865: 9855: 9815: 9809: 9808: 9798: 9781:. 72–73: 22–36. 9770: 9761: 9760: 9750: 9710: 9695: 9694: 9684: 9660: 9647: 9646: 9614: 9605: 9604: 9572: 9566: 9565: 9532: 9499: 9498: 9488: 9470: 9446: 9440: 9439: 9399: 9390: 9389: 9379: 9331: 9320: 9319: 9309: 9285: 9279: 9278: 9268: 9244: 9238: 9237: 9189: 9178: 9177: 9129: 9116: 9115: 9075: 9062: 9061: 9043: 9034: 9028: 9027: 8994: 8977: 8976: 8960: 8949: 8948: 8938: 8890: 8881: 8880: 8868: 8859: 8858: 8848: 8828: 8822: 8821: 8781: 8772: 8771: 8755: 8730: 8729: 8689: 8680: 8679: 8667: 8661: 8660: 8620: 8614: 8613: 8587: 8578: 8513: 8512: 8480: 8439: 8438: 8420: 8411: 8400: 8399: 8389: 8341: 8330: 8329: 8297: 8224: 8223: 8182: 8135: 8134: 8124: 8115: 8104: 8103: 8093: 8044: 8027: 8026: 7978: 7967: 7966: 7956: 7932: 7915: 7914: 7866: 7851: 7850: 7848: 7846: 7836: 7828: 7822: 7821: 7805: 7799: 7798: 7796: 7785: 7779: 7778: 7776: 7774: 7759: 7753: 7752: 7750: 7748: 7732: 7726: 7725: 7723: 7721: 7705: 7699: 7698: 7688: 7655: 7646: 7645: 7597: 7586: 7585: 7557: 7551: 7550: 7507:Briggs, D. E. G. 7502: 7493: 7492: 7464: 7443: 7442: 7440: 7422: 7397: 7391: 7390: 7343:Briggs, D. E. G. 7338: 7307: 7290: 7284: 7267: 7261: 7244: 7238: 7221: 7215: 7214: 7212: 7188: 6840:Archaeoconularia 6813: 6812: 6613: 6612: 6590:asthenospongiids 6362:angusticardiniid 6288: 6287: 6026:Paratemnograptus 5463: 5462: 5454:zone and early " 5417:zone (7), ? 5413:zone (6), ? 5223:S. aff. gracilis 5204:P. cf. leonardii 5088:P. cf. flexuosus 5000:strachanognathid 4936: 4935: 4622:P. fezouataensis 4026: 4025: 4013:A rare mitrate. 3976:Cantabrigiaster, 3938:A rare cornute. 3751:A rare disparid 3718:A rare cornute. 3504:A rare cornute. 3378:A rare mitrate. 3280:A rare cornute. 3255:A rare cornute. 3204:A rare cornute. 3168:A rare cornute. 3041:A rare mitrate. 2760: 2759: 2692:anomalacystitids 2580:T. tazagurtensis 2364: 2363: 2304:Other arthropods 2216:late meraspids. 2089:P. tazzarinensis 1929:Parabathycheilus 1757:M. (E.) hammondi 1386:dikelokephalinid 1384:A large endemic 970:A. aff. jujuanus 941:A. fezouataensis 920:Lower Fezouata, 729: 728: 574: 573: 334:Early Ordovician 305: 304: 296: 295: 287: 286: 280: 264: 262: 261: 239: 238: 236: 235: 234: 229: 225: 222: 221: 220: 217: 200: 199: 197: 196: 195: 190: 186: 183: 182: 181: 178: 106: 97: 34: 19: 18: 10884: 10883: 10879: 10878: 10877: 10875: 10874: 10873: 10824: 10823: 10822: 10817: 10781: 10757:Qingjiang biota 10715: 10710: 10680: 10679: 10644: 10640: 10587: 10583: 10555: 10549: 10542: 10507: 10503: 10468: 10464: 10419: 10410: 10375: 10354: 10317: 10310: 10257: 10250: 10215: 10206: 10171: 10162: 10127: 10118: 10083: 10066: 9997: 9988: 9954: 9948: 9925: 9890: 9869: 9830:(1): 20200809. 9824:Biology Letters 9816: 9812: 9771: 9764: 9711: 9698: 9661: 9650: 9615: 9608: 9573: 9569: 9533: 9502: 9447: 9443: 9400: 9393: 9332: 9323: 9286: 9282: 9245: 9241: 9190: 9181: 9130: 9119: 9076: 9065: 9041: 9035: 9031: 9025: 8995: 8980: 8961: 8952: 8891: 8884: 8869: 8862: 8829: 8825: 8810:10.1002/gj.1275 8782: 8775: 8756: 8733: 8690: 8683: 8668: 8664: 8621: 8617: 8585: 8579: 8516: 8481: 8442: 8418: 8412: 8403: 8342: 8333: 8298: 8227: 8183: 8138: 8122: 8116: 8107: 8045: 8030: 7993:(7554): 77–80. 7979: 7970: 7933: 7918: 7867: 7854: 7844: 7842: 7834: 7830: 7829: 7825: 7806: 7802: 7794: 7786: 7782: 7772: 7770: 7761: 7760: 7756: 7746: 7744: 7733: 7729: 7719: 7717: 7706: 7702: 7656: 7649: 7598: 7589: 7558: 7554: 7503: 7496: 7465: 7446: 7398: 7394: 7357:(7295): 215–8. 7339: 7310: 7300:Wayback Machine 7291: 7287: 7277:Wayback Machine 7268: 7264: 7254:Wayback Machine 7245: 7241: 7231:Wayback Machine 7222: 7218: 7189: 7182: 7177: 7158: 7156:Other organisms 7067:A medium-sized 7064:Upper Fezouata 7007:Upper Fezouata 6981:Upper Fezouata 6955:Upper Fezouata 6904:Upper Fezouata 6878:Upper Fezouata 6853:A common large 6784: 6773:A reticulosan. 6746:Upper Fezouata 6717:Upper Fezouata 6653:Upper Fezouata 6600:Pirania auraeum 6552: 6531:Upper Fezouata 6473:Lower Fezouata 6444:Upper Fezouata 6440:P. tadristensis 6415:Upper Fezouata 6386:Upper Fezouata 6357:Upper Fezouata 6347:Angusticardinia 6328:Lower Fezouata 6286: 5976:Paradelograptus 5724:Choristograptus 5446:zone and early 5335: 5198:Protopanderodus 5181:P. cf. gilberti 4923:subzone of the 4904: 4862:bellerophontoid 4802:Upper Fezouata 4516:E. courtessolei 4493:apicrinoceratid 4478:Destombesiceras 4433:Intihuarellidae 4398:afghanodesmatid 4347:monoplacophoran 4313:monoplacophoran 4291:C. courtessolei 4223:B. taichoutense 4213:Ellesmerocerida 4024: 3980:Villebrunaster. 3953:V. fezouatensis 3653:zone up to the 3494:N. cf. dolambii 3002:A. cf. sinuosus 2939:A. moroccoensis 2684:chauvelicystids 2680: 2350:?), a possible 2322:chasmataspidids 2310:synziphosurines 2306: 2193:Upper Fezouata 2056:P. magrebiensis 2034:Upper Fezouata 2008:Lower Fezouata 1981:Upper Fezouata 1771:Cruziana rugosa 1745:Upper Fezouata 1400:Lower Fezouata 1368:Dikelokephalina 1347:Dactylocephalus 1240:Upper Fezouata 1216:Upper Fezouata 1207:B. zemmourensis 1163:Upper Fezouata 1082:Dikelokephalina 1077:Lower Fezouata 1053:Upper Fezouata 922:Upper Fezouata 724: 662:P. duplospineus 628:aegirocassisine 565: 557: 545: 537: 512: 485: 477:storm wave base 465: 372:Over 1,500 non- 358: 314: 313: 312: 311: 308: 307: 306: 298: 297: 289: 288: 259: 257: 232: 230: 226: 223: 218: 215: 213: 211: 210: 193: 191: 187: 184: 179: 176: 174: 172: 171: 113: 96: 95: 90: 85: 80: 75: 70: 65: 60: 55: 50: 45: 40: 32: 31: 17: 12: 11: 5: 10882: 10872: 10871: 10866: 10861: 10856: 10851: 10846: 10841: 10836: 10819: 10818: 10816: 10815: 10810: 10805: 10800: 10795: 10789: 10787: 10783: 10782: 10780: 10779: 10774: 10769: 10764: 10759: 10754: 10749: 10744: 10739: 10734: 10729: 10723: 10721: 10717: 10716: 10709: 10708: 10701: 10694: 10686: 10678: 10677: 10638: 10581: 10540: 10501: 10462: 10408: 10352: 10308: 10248: 10204: 10160: 10141:(6): 427–438. 10116: 10064: 9986: 9923: 9867: 9810: 9762: 9725:(4): 685–714. 9696: 9648: 9606: 9567: 9500: 9441: 9414:(3): 509–517. 9391: 9321: 9280: 9239: 9179: 9144:(3): 524–536. 9117: 9090:(3): 425–454. 9063: 9052:(2): 261–273. 9029: 9023: 8978: 8950: 8882: 8860: 8823: 8796:(5): 405–415. 8773: 8731: 8704:(1): 152–157. 8681: 8662: 8635:(4): 271–283. 8615: 8514: 8440: 8429:(4): 857–877. 8401: 8331: 8225: 8198:(2): 261–270. 8136: 8105: 8028: 7968: 7916: 7881:(5): 541–549. 7852: 7823: 7800: 7780: 7754: 7727: 7700: 7647: 7587: 7552: 7494: 7475:(6): 737–748. 7444: 7392: 7308: 7285: 7262: 7247:Fezouata Shale 7239: 7216: 7203:(2): 296–311. 7179: 7178: 7176: 7173: 7157: 7154: 7151: 7150: 7148: 7143: 7140: 7135: 7127: 7126: 7124: 7115: 7097: 7091: 7086: 7078: 7077: 7075: 7065: 7062: 7057: 7049: 7048: 7046: 7042:palaeoscolecid 7038: 7032: 7027: 7018: 7017: 7015: 7008: 7005: 7000: 6992: 6991: 6989: 6982: 6979: 6974: 6966: 6965: 6963: 6956: 6953: 6948: 6939: 6938: 6936: 6931: 6928: 6923: 6915: 6914: 6912: 6905: 6902: 6897: 6889: 6888: 6886: 6879: 6876: 6871: 6863: 6862: 6860: 6851: 6848: 6843: 6835: 6834: 6831: 6828: 6825: 6822: 6818: 6817: 6783: 6780: 6777: 6776: 6774: 6771: 6769: 6764: 6755: 6754: 6752: 6747: 6744: 6739: 6731: 6730: 6728: 6718: 6715: 6710: 6704: 6703: 6701: 6700:A hamptoniid. 6698: 6696: 6691: 6683: 6682: 6680: 6677: 6675: 6670: 6662: 6661: 6659: 6654: 6651: 6646: 6638: 6637: 6634: 6631: 6628: 6625: 6621: 6620: 6586:hexactinellids 6556:protomonaxonid 6551: 6548: 6545: 6544: 6542: 6532: 6529: 6524: 6516: 6515: 6513: 6503: 6500: 6495: 6487: 6486: 6484: 6474: 6471: 6466: 6458: 6457: 6455: 6445: 6442: 6437: 6429: 6428: 6426: 6416: 6413: 6408: 6400: 6399: 6397: 6387: 6384: 6379: 6371: 6370: 6368: 6358: 6355: 6350: 6342: 6341: 6339: 6329: 6326: 6321: 6313: 6312: 6309: 6306: 6303: 6300: 6296: 6295: 6285: 6282: 6279: 6278: 6276: 6273: 6270: 6265: 6257: 6256: 6254: 6241: 6233: 6232: 6230: 6223: 6217: 6210: 6200: 6199: 6197: 6190: 6184: 6179: 6171: 6170: 6168: 6149: 6143: 6141: 6137:indeterminate 6134: 6133: 6131: 6125: 6119: 6118: 6116: 6104: 6098: 6097: 6095: 6089: 6083: 6082: 6080: 6071: 6065: 6060: 6052: 6051: 6049: 6046: 6034: 6029: 6021: 6020: 6018: 6012: 6006: 6005: 6003: 5998:A multiramous 5996: 5984: 5979: 5971: 5970: 5968: 5965: 5953: 5948: 5938: 5937: 5935: 5932: 5926: 5921: 5913: 5912: 5910: 5907: 5901: 5896: 5888: 5887: 5885: 5882: 5875: 5870: 5867:Expansograptus 5862: 5861: 5859: 5856: 5844: 5839: 5829: 5828: 5826: 5823: 5816: 5811: 5802: 5801: 5799: 5787: 5781: 5780: 5778: 5771: 5757: 5752: 5744: 5743: 5741: 5738: 5732: 5727: 5719: 5718: 5716: 5710: 5704: 5703: 5701: 5695: 5689: 5688: 5686: 5679: 5673: 5672: 5670: 5663: 5657: 5652: 5644: 5643: 5641: 5634: 5627: 5622: 5614: 5613: 5611: 5604: 5598: 5593: 5585: 5584: 5582: 5579: 5573: 5568: 5560: 5559: 5557: 5550: 5544: 5539: 5531: 5530: 5528: 5516:A multiramous 5514: 5503: 5498: 5488: 5487: 5484: 5481: 5478: 5475: 5471: 5470: 5380:rhabdopleurids 5334: 5331: 5328: 5327: 5325: 5319: 5318: 5316: 5313: 5311:T. aff. laevis 5308: 5300: 5299: 5297: 5291: 5286: 5278: 5277: 5275: 5269: 5268: 5266: 5263: 5258: 5255:Semiacontiodus 5250: 5249: 5247: 5244: 5242:S. aff. krummi 5239: 5231: 5230: 5228: 5225: 5220: 5212: 5211: 5209: 5206: 5201: 5193: 5192: 5190: 5189:prioniodontid 5183: 5178: 5170: 5169: 5167: 5161: 5160: 5158: 5149: 5144: 5136: 5135: 5133: 5127: 5126: 5124: 5121: 5116: 5108: 5107: 5105: 5099: 5098: 5096: 5090: 5085: 5077: 5076: 5074: 5068: 5067: 5065: 5062: 5057: 5049: 5048: 5046: 5040: 5038:D. cf. forceps 5035: 5032:Drepanoistodus 5027: 5026: 5024: 5018: 5013: 5005: 5004: 5002: 4996: 4991: 4983: 4982: 4980: 4971: 4966: 4958: 4957: 4954: 4951: 4948: 4944: 4943: 4903: 4900: 4897: 4896: 4894: 4889: 4883: 4878: 4870: 4869: 4867: 4858: 4852: 4847: 4839: 4838: 4836: 4829: 4826: 4821: 4813: 4812: 4810: 4803: 4800: 4795: 4787: 4786: 4784: 4775:A very common 4773: 4753: 4748: 4740: 4739: 4737: 4724: 4721: 4716: 4708: 4707: 4705: 4698: 4691: 4686: 4677: 4676: 4674: 4661: 4658: 4653: 4645: 4644: 4642: 4630: 4624: 4619: 4611: 4610: 4608: 4599: 4580: 4575: 4567: 4566: 4564: 4557: 4550: 4545: 4537: 4536: 4534: 4531: 4518: 4513: 4505: 4504: 4502: 4489: 4486: 4481: 4473: 4472: 4470: 4466: 4452: 4447: 4439: 4438: 4436: 4425: 4418: 4413: 4404: 4403: 4401: 4394: 4387: 4385: 4376: 4375: 4373: 4368: 4361: 4355: 4354: 4352: 4340: 4327: 4321: 4320: 4318: 4306: 4293: 4288: 4285:Carcassonnella 4280: 4279: 4277: 4250: 4244: 4239: 4231: 4230: 4228: 4225: 4219: 4218: 4216: 4205: 4202: 4197: 4189: 4188: 4186: 4165: 4162: 4157: 4149: 4148: 4146: 4128: 4108: 4103: 4095: 4094: 4092: 4089:deposit feeder 4072: 4065: 4060: 4051: 4050: 4047: 4044: 4041: 4038: 4034: 4033: 4023: 4020: 4017: 4016: 4014: 4011: 4005: 4000: 3997:Vizcainocarpus 3992: 3991: 3982: 3965:chinianasterid 3961: 3955: 3950: 3947:Villebrunaster 3942: 3941: 3939: 3936: 3929: 3927:T. zagoraensis 3923: 3922: 3920: 3917: 3911: 3906: 3898: 3897: 3895: 3888: 3876: 3874:R. zagoraensis 3870: 3869: 3867: 3860: 3854: 3848: 3847: 3845: 3838: 3832: 3826: 3825: 3823: 3816: 3810: 3804: 3803: 3801: 3797: 3772: 3767: 3759: 3758: 3756: 3749: 3735: 3730: 3722: 3721: 3719: 3716: 3710: 3705: 3697: 3696: 3694: 3691: 3685: 3680: 3672: 3671: 3669: 3658: 3648: 3643: 3635: 3634: 3632: 3629: 3623: 3618: 3610: 3609: 3607: 3604: 3591: 3589: 3580: 3579: 3577: 3574: 3561: 3556: 3547: 3546: 3537: 3531:minervaecystid 3527: 3521: 3516: 3508: 3507: 3505: 3502: 3496: 3491: 3483: 3482: 3480: 3470:glyptocystitid 3466: 3453: 3448: 3440: 3439: 3437: 3434: 3421: 3415: 3414: 3412: 3409: 3395: 3390: 3382: 3381: 3379: 3376: 3370: 3368:L. pyramidalis 3365: 3357: 3356: 3354: 3347: 3340: 3335: 3327: 3326: 3324: 3321: 3315: 3310: 3302: 3301: 3299: 3296: 3290: 3284: 3283: 3281: 3278: 3272: 3267: 3259: 3258: 3256: 3253: 3247: 3242: 3239:Flabellicarpus 3234: 3233: 3231: 3228: 3221: 3216: 3213:Chinianocarpos 3208: 3207: 3205: 3202: 3196: 3190: 3189: 3187: 3184: 3178: 3172: 3171: 3169: 3166: 3160: 3155: 3152:Chauvelicystis 3147: 3146: 3144: 3141: 3135: 3130: 3127:Bohemiaecystis 3122: 3121: 3119: 3104: 3084: 3079: 3076:Balantiocystis 3071: 3070: 3068: 3065: 3058: 3053: 3050:Balanocystites 3045: 3044: 3042: 3039: 3033: 3028: 3020: 3019: 3017: 3010: 3004: 2999: 2996:Aristocystites 2991: 2990: 2988: 2985: 2971: 2969:A. espilezorum 2966: 2958: 2957: 2955: 2948: 2941: 2936: 2928: 2927: 2925: 2922: 2903: 2897: 2896: 2894: 2887: 2881: 2876: 2868: 2867: 2865: 2862: 2856: 2851: 2843: 2842: 2840: 2833: 2827: 2822: 2814: 2813: 2811: 2804: 2798: 2793: 2785: 2784: 2781: 2778: 2775: 2772: 2768: 2767: 2751:Balantiocystis 2696:mitrocystitids 2679: 2676: 2673: 2672: 2670: 2667: 2665: 2660: 2652: 2651: 2642: 2631: 2625: 2620: 2612: 2611: 2609: 2588: 2582: 2577: 2569: 2568: 2566: 2537:synziphosurine 2529: 2523: 2518: 2510: 2509: 2500: 2479: 2473: 2471: 2463: 2462: 2453: 2439: 2433: 2428: 2420: 2419: 2417: 2406: 2403: 2398: 2395:Brachyaglaspis 2390: 2389: 2386: 2383: 2380: 2377: 2373: 2372: 2305: 2302: 2299: 2298: 2296: 2289: 2270: 2265: 2257: 2256: 2254: 2247: 2241: 2235: 2234: 2232: 2229: 2226: 2220: 2219: 2217: 2213: 2207: 2201: 2200: 2198: 2195: 2186: 2181: 2173: 2172: 2163: 2156: 2154: 2152: 2144: 2143: 2141: 2134: 2122: 2117: 2109: 2108: 2106: 2103: 2091: 2086: 2078: 2077: 2068: 2064: 2058: 2053: 2050:Platypeltoides 2045: 2044: 2042: 2035: 2032: 2027: 2019: 2018: 2016: 2009: 2006: 2001: 1992: 1991: 1989: 1982: 1979: 1974: 1966: 1965: 1963: 1956: 1937: 1932: 1924: 1923: 1921: 1918: 1912: 1907: 1899: 1898: 1896: 1893: 1887: 1881: 1880: 1878: 1875: 1873: 1868: 1860: 1859: 1857: 1854: 1852: 1847: 1838: 1837: 1835: 1828: 1822: 1817: 1809: 1808: 1806: 1803: 1791: 1786: 1778: 1777: 1775: 1765: 1759: 1753: 1752: 1750: 1747: 1738: 1733: 1720: 1719: 1717: 1710: 1708: 1703: 1695: 1694: 1692: 1681: 1669: 1664: 1661:Leptoplastides 1656: 1655: 1653: 1646: 1640: 1635: 1627: 1626: 1617: 1610: 1608: 1603: 1595: 1594: 1585: 1574: 1562: 1557: 1549: 1548: 1546: 1543: 1531: 1526: 1518: 1517: 1515: 1508: 1489: 1484: 1476: 1475: 1466: 1459: 1457: 1452: 1444: 1443: 1441: 1434: 1428: 1423: 1414: 1413: 1404: 1401: 1398: 1392: 1391: 1389: 1382: 1376: 1371: 1363: 1362: 1360: 1357: 1355: 1350: 1342: 1341: 1332: 1329: 1317: 1311: 1310: 1308: 1305: 1293: 1287: 1286: 1284: 1277: 1264: 1262:C. pradesensis 1259: 1251: 1250: 1248: 1241: 1238: 1233: 1224: 1223: 1221: 1218: 1209: 1203: 1202: 1200: 1193: 1187: 1182: 1174: 1173: 1171: 1164: 1161: 1156: 1148: 1147: 1145: 1142: 1140: 1134: 1133: 1124: 1121: 1119: 1113: 1112: 1110: 1107: 1101: 1096: 1088: 1087: 1085: 1078: 1075: 1070: 1062: 1061: 1059: 1055: 1046: 1040: 1039: 1030: 1027: 1021: 1015: 1014: 1012: 1009: 1003: 997: 996: 994: 991: 972: 966: 965: 963: 949: 943: 938: 930: 929: 927: 924: 918: 912: 911: 909: 902: 890: 888:A. cf. incisus 885: 877: 876: 874: 852: 846: 841: 833: 832: 830: 813: 803: 798: 790: 789: 787: 780: 767: 762: 754: 753: 750: 747: 744: 741: 737: 736: 723: 720: 717: 716: 714: 708: 702: 701: 699: 693: 687: 686: 677: 670: 664: 659: 651: 650: 641: 624: 612: 607: 599: 598: 595: 592: 589: 586: 582: 581: 564: 561: 556: 553: 544: 541: 536: 533: 511: 508: 484: 481: 464: 461: 459:in abundance. 425:marrellomorphs 357: 354: 346:Fezouata biota 326:Fezouata Shale 316: 315: 309: 300: 299: 291: 290: 282: 281: 275: 274: 273: 272: 269: 268: 255: 251: 250: 248:Drâa-Tafilalet 245: 241: 240: 228:73.9°S 108.1°E 208: 202: 201: 169: 163: 162: 158: 157: 152: 148: 147: 143: 142: 139: 135: 134: 132:Zini Formation 129: 125: 124: 119: 115: 114: 107: 99: 98: 91: 86: 81: 76: 71: 66: 61: 56: 51: 46: 41: 36: 24: 23: 15: 9: 6: 4: 3: 2: 10881: 10870: 10867: 10865: 10862: 10860: 10857: 10855: 10852: 10850: 10847: 10845: 10842: 10840: 10837: 10835: 10832: 10831: 10829: 10814: 10811: 10809: 10806: 10804: 10801: 10799: 10796: 10794: 10791: 10790: 10788: 10784: 10778: 10775: 10773: 10772:Wheeler Shale 10770: 10768: 10765: 10763: 10762:Sirius Passet 10760: 10758: 10755: 10753: 10750: 10748: 10745: 10743: 10740: 10738: 10737:Emu Bay Shale 10735: 10733: 10730: 10728: 10727:Burgess Shale 10725: 10724: 10722: 10718: 10714: 10707: 10702: 10700: 10695: 10693: 10688: 10687: 10684: 10673: 10669: 10665: 10661: 10657: 10653: 10649: 10642: 10634: 10630: 10625: 10620: 10616: 10612: 10608: 10604: 10600: 10596: 10592: 10585: 10577: 10573: 10569: 10565: 10561: 10554: 10547: 10545: 10536: 10532: 10528: 10524: 10520: 10516: 10512: 10505: 10497: 10493: 10489: 10485: 10481: 10477: 10473: 10466: 10457: 10452: 10448: 10444: 10440: 10436: 10432: 10428: 10424: 10417: 10415: 10413: 10404: 10400: 10396: 10392: 10388: 10384: 10380: 10373: 10371: 10369: 10367: 10365: 10363: 10361: 10359: 10357: 10348: 10344: 10339: 10334: 10330: 10326: 10322: 10315: 10313: 10304: 10300: 10295: 10290: 10286: 10282: 10278: 10274: 10270: 10266: 10262: 10255: 10253: 10244: 10240: 10236: 10232: 10228: 10224: 10220: 10213: 10211: 10209: 10200: 10196: 10192: 10188: 10184: 10180: 10176: 10169: 10167: 10165: 10156: 10152: 10148: 10144: 10140: 10136: 10132: 10125: 10123: 10121: 10112: 10108: 10104: 10100: 10096: 10092: 10088: 10081: 10079: 10077: 10075: 10073: 10071: 10069: 10060: 10056: 10052: 10048: 10044: 10040: 10035: 10030: 10026: 10022: 10018: 10014: 10010: 10006: 10002: 9995: 9993: 9991: 9981: 9976: 9972: 9968: 9964: 9960: 9959:Fossil Record 9953: 9946: 9944: 9942: 9940: 9938: 9936: 9934: 9932: 9930: 9928: 9919: 9915: 9911: 9907: 9903: 9899: 9895: 9888: 9886: 9884: 9882: 9880: 9878: 9876: 9874: 9872: 9863: 9859: 9854: 9849: 9845: 9841: 9837: 9833: 9829: 9825: 9821: 9814: 9806: 9802: 9797: 9792: 9788: 9784: 9780: 9776: 9769: 9767: 9758: 9754: 9749: 9744: 9740: 9736: 9732: 9728: 9724: 9720: 9716: 9709: 9707: 9705: 9703: 9701: 9692: 9688: 9683: 9678: 9674: 9670: 9666: 9659: 9657: 9655: 9653: 9644: 9640: 9636: 9632: 9628: 9624: 9620: 9613: 9611: 9602: 9598: 9594: 9590: 9586: 9582: 9578: 9571: 9563: 9559: 9555: 9551: 9547: 9543: 9539: 9531: 9529: 9527: 9525: 9523: 9521: 9519: 9517: 9515: 9513: 9511: 9509: 9507: 9505: 9496: 9492: 9487: 9482: 9478: 9474: 9469: 9464: 9460: 9456: 9452: 9445: 9437: 9433: 9429: 9425: 9421: 9417: 9413: 9409: 9405: 9398: 9396: 9387: 9383: 9378: 9373: 9369: 9365: 9361: 9357: 9353: 9349: 9345: 9341: 9337: 9330: 9328: 9326: 9317: 9313: 9308: 9303: 9299: 9295: 9291: 9284: 9276: 9272: 9267: 9262: 9258: 9254: 9250: 9243: 9235: 9231: 9227: 9223: 9219: 9215: 9211: 9207: 9203: 9199: 9195: 9188: 9186: 9184: 9175: 9171: 9167: 9163: 9159: 9155: 9151: 9147: 9143: 9139: 9135: 9128: 9126: 9124: 9122: 9113: 9109: 9105: 9101: 9097: 9093: 9089: 9085: 9081: 9074: 9072: 9070: 9068: 9059: 9055: 9051: 9047: 9040: 9033: 9026: 9020: 9016: 9012: 9008: 9004: 9002: 8993: 8991: 8989: 8987: 8985: 8983: 8974: 8970: 8966: 8959: 8957: 8955: 8946: 8942: 8937: 8932: 8928: 8924: 8920: 8916: 8912: 8908: 8904: 8900: 8896: 8889: 8887: 8878: 8874: 8867: 8865: 8856: 8852: 8847: 8842: 8838: 8834: 8827: 8819: 8815: 8811: 8807: 8803: 8799: 8795: 8791: 8787: 8780: 8778: 8769: 8765: 8761: 8754: 8752: 8750: 8748: 8746: 8744: 8742: 8740: 8738: 8736: 8727: 8723: 8719: 8715: 8711: 8707: 8703: 8699: 8695: 8688: 8686: 8677: 8673: 8666: 8658: 8654: 8650: 8646: 8642: 8638: 8634: 8630: 8626: 8619: 8611: 8607: 8603: 8599: 8595: 8591: 8584: 8577: 8575: 8573: 8571: 8569: 8567: 8565: 8563: 8561: 8559: 8557: 8555: 8553: 8551: 8549: 8547: 8545: 8543: 8541: 8539: 8537: 8535: 8533: 8531: 8529: 8527: 8525: 8523: 8521: 8519: 8510: 8506: 8502: 8498: 8494: 8490: 8486: 8479: 8477: 8475: 8473: 8471: 8469: 8467: 8465: 8463: 8461: 8459: 8457: 8455: 8453: 8451: 8449: 8447: 8445: 8436: 8432: 8428: 8424: 8417: 8410: 8408: 8406: 8397: 8393: 8388: 8383: 8379: 8375: 8371: 8367: 8363: 8359: 8355: 8351: 8347: 8340: 8338: 8336: 8327: 8323: 8319: 8315: 8311: 8307: 8303: 8296: 8294: 8292: 8290: 8288: 8286: 8284: 8282: 8280: 8278: 8276: 8274: 8272: 8270: 8268: 8266: 8264: 8262: 8260: 8258: 8256: 8254: 8252: 8250: 8248: 8246: 8244: 8242: 8240: 8238: 8236: 8234: 8232: 8230: 8221: 8217: 8213: 8209: 8205: 8201: 8197: 8193: 8189: 8181: 8179: 8177: 8175: 8173: 8171: 8169: 8167: 8165: 8163: 8161: 8159: 8157: 8155: 8153: 8151: 8149: 8147: 8145: 8143: 8141: 8133:(2): 239–246. 8132: 8128: 8121: 8114: 8112: 8110: 8101: 8097: 8092: 8087: 8083: 8079: 8075: 8071: 8067: 8063: 8059: 8055: 8051: 8043: 8041: 8039: 8037: 8035: 8033: 8024: 8020: 8016: 8012: 8008: 8004: 8000: 7996: 7992: 7988: 7984: 7977: 7975: 7973: 7964: 7960: 7955: 7950: 7946: 7942: 7938: 7931: 7929: 7927: 7925: 7923: 7921: 7912: 7908: 7904: 7900: 7896: 7892: 7888: 7884: 7880: 7876: 7872: 7865: 7863: 7861: 7859: 7857: 7840: 7833: 7827: 7819: 7815: 7811: 7804: 7793: 7792: 7784: 7768: 7767:The Economist 7764: 7758: 7742: 7738: 7731: 7715: 7711: 7704: 7696: 7692: 7687: 7682: 7678: 7674: 7670: 7666: 7662: 7654: 7652: 7643: 7639: 7635: 7631: 7627: 7623: 7619: 7615: 7611: 7607: 7603: 7596: 7594: 7592: 7583: 7579: 7575: 7571: 7567: 7563: 7556: 7548: 7544: 7540: 7536: 7532: 7528: 7524: 7520: 7516: 7512: 7508: 7505:Van Roy, P.; 7501: 7499: 7490: 7486: 7482: 7478: 7474: 7470: 7463: 7461: 7459: 7457: 7455: 7453: 7451: 7449: 7439: 7434: 7430: 7426: 7421: 7416: 7412: 7408: 7404: 7396: 7388: 7384: 7380: 7376: 7372: 7368: 7364: 7360: 7356: 7352: 7348: 7344: 7337: 7335: 7333: 7331: 7329: 7327: 7325: 7323: 7321: 7319: 7317: 7315: 7313: 7305: 7301: 7297: 7294: 7289: 7282: 7278: 7274: 7271: 7266: 7259: 7255: 7251: 7248: 7243: 7236: 7232: 7228: 7225: 7220: 7211: 7206: 7202: 7198: 7194: 7187: 7185: 7180: 7172: 7170: 7166: 7162: 7149: 7147: 7144: 7141: 7139: 7136: 7134: 7133: 7132:Sphenothallus 7129: 7128: 7125: 7120: 7116: 7113: 7109: 7106: 7105:aphroditiform 7102: 7098: 7095: 7092: 7090: 7087: 7085: 7084: 7080: 7079: 7076: 7073: 7070: 7066: 7063: 7061: 7058: 7056: 7055: 7051: 7050: 7047: 7043: 7039: 7036: 7033: 7031: 7028: 7026: 7024: 7020: 7019: 7016: 7013: 7009: 7006: 7004: 7001: 6999: 6998: 6994: 6993: 6990: 6987: 6983: 6980: 6978: 6975: 6973: 6972: 6968: 6967: 6964: 6961: 6957: 6954: 6952: 6949: 6947: 6945: 6941: 6940: 6937: 6935: 6932: 6929: 6927: 6924: 6922: 6921: 6917: 6916: 6913: 6910: 6906: 6903: 6901: 6898: 6896: 6895: 6894:Elegantilites 6891: 6890: 6887: 6884: 6880: 6877: 6875: 6872: 6870: 6869: 6865: 6864: 6861: 6859: 6856: 6852: 6849: 6847: 6844: 6842: 6841: 6837: 6836: 6832: 6829: 6827:Stratigraphy 6826: 6823: 6820: 6819: 6814: 6811: 6809: 6805: 6801: 6797: 6793: 6789: 6782:Other animals 6775: 6772: 6770: 6768: 6765: 6763: 6761: 6757: 6756: 6753: 6751: 6748: 6745: 6743: 6740: 6738: 6737: 6733: 6732: 6729: 6726: 6723: 6719: 6716: 6714: 6711: 6709: 6706: 6705: 6702: 6699: 6697: 6695: 6692: 6690: 6689: 6685: 6684: 6681: 6678: 6676: 6674: 6671: 6669: 6668: 6664: 6663: 6660: 6658: 6655: 6652: 6650: 6647: 6645: 6644: 6640: 6639: 6635: 6632: 6630:Stratigraphy 6629: 6626: 6623: 6622: 6618: 6614: 6611: 6609: 6603: 6601: 6597: 6594: 6591: 6587: 6584: 6580: 6576: 6572: 6568: 6564: 6560: 6557: 6543: 6540: 6537: 6533: 6530: 6528: 6525: 6523: 6522: 6518: 6517: 6514: 6511: 6508: 6504: 6501: 6499: 6496: 6494: 6493: 6489: 6488: 6485: 6482: 6479: 6475: 6472: 6470: 6467: 6465: 6464: 6460: 6459: 6456: 6453: 6450: 6446: 6443: 6441: 6438: 6436: 6435: 6431: 6430: 6427: 6424: 6421: 6417: 6414: 6412: 6409: 6407: 6406: 6402: 6401: 6398: 6395: 6392: 6388: 6385: 6383: 6380: 6378: 6377: 6373: 6372: 6369: 6366: 6363: 6359: 6356: 6354: 6351: 6349: 6348: 6344: 6343: 6340: 6337: 6334: 6330: 6327: 6325: 6322: 6320: 6319: 6315: 6314: 6310: 6307: 6305:Stratigraphy 6304: 6301: 6298: 6297: 6293: 6289: 6277: 6274: 6271: 6269: 6266: 6264: 6263: 6259: 6258: 6255: 6253: 6251: 6245: 6242: 6239: 6235: 6234: 6231: 6228: 6221: 6218: 6215: 6211: 6208: 6207: 6201: 6198: 6195: 6191: 6188: 6185: 6183: 6180: 6178: 6177: 6176:Schizograptus 6173: 6172: 6169: 6166: 6165: 6160: 6156: 6155: 6150: 6147: 6144: 6142: 6140: 6139:rhabdopleurid 6136: 6135: 6132: 6129: 6126: 6124: 6121: 6120: 6117: 6115: 6113: 6108: 6105: 6103: 6100: 6099: 6096: 6093: 6090: 6088: 6085: 6084: 6081: 6078: 6077: 6069: 6066: 6064: 6061: 6059: 6058: 6053: 6050: 6047: 6045: 6043: 6038: 6035: 6033: 6032:P. magnificus 6030: 6028: 6027: 6023: 6022: 6019: 6016: 6013: 6011: 6008: 6007: 6004: 6001: 5995: 5993: 5988: 5985: 5983: 5982:P. norvegicus 5980: 5978: 5977: 5972: 5969: 5966: 5964: 5962: 5957: 5954: 5952: 5951:"K." supremus 5949: 5947: 5945: 5944:Kiaerograptus 5940: 5939: 5936: 5933: 5930: 5927: 5925: 5922: 5920: 5919: 5915: 5914: 5911: 5908: 5905: 5902: 5900: 5897: 5895: 5894: 5890: 5889: 5886: 5883: 5880: 5876: 5874: 5871: 5869: 5868: 5864: 5863: 5860: 5857: 5855: 5853: 5848: 5845: 5843: 5840: 5838: 5836: 5835:Didymograptus 5831: 5830: 5827: 5824: 5821: 5817: 5815: 5812: 5810: 5809: 5808:Cymatograptus 5804: 5803: 5800: 5798: 5796: 5791: 5788: 5786: 5783: 5782: 5779: 5776: 5770: 5768: 5762: 5758: 5756: 5753: 5751: 5750: 5745: 5742: 5739: 5736: 5733: 5731: 5728: 5726: 5725: 5721: 5720: 5717: 5714: 5711: 5709: 5706: 5705: 5702: 5699: 5696: 5694: 5691: 5690: 5687: 5684: 5680: 5678: 5675: 5674: 5671: 5668: 5667:didymograptid 5661: 5658: 5656: 5653: 5651: 5650: 5645: 5642: 5639: 5635: 5632: 5628: 5626: 5625:A. eivionicus 5623: 5621: 5620: 5616: 5615: 5612: 5609: 5608:Sagenograptus 5605: 5602: 5599: 5597: 5594: 5592: 5591: 5590:Araneograptus 5587: 5586: 5583: 5580: 5577: 5574: 5572: 5569: 5567: 5566: 5562: 5561: 5558: 5555: 5551: 5548: 5545: 5543: 5540: 5538: 5537: 5536:Ancoragraptus 5533: 5532: 5529: 5526: 5523: 5519: 5515: 5512: 5508: 5504: 5502: 5501:"A." tenellus 5499: 5497: 5495: 5490: 5489: 5485: 5482: 5479: 5476: 5473: 5472: 5468: 5464: 5461: 5460: 5457: 5453: 5449: 5445: 5440: 5439: 5436: 5432: 5428: 5424: 5420: 5416: 5412: 5408: 5404: 5400: 5396: 5392: 5388: 5383: 5381: 5377: 5376: 5371: 5370: 5365: 5364: 5363:Didymograptus 5359: 5356: 5352: 5351: 5346: 5345: 5344:Araneograptus 5340: 5326: 5324: 5323:T. cf. sweeti 5321: 5320: 5317: 5315:An acodontid 5312: 5309: 5307: 5306: 5301: 5298: 5296: 5292: 5290: 5287: 5285: 5284: 5280: 5279: 5276: 5274: 5271: 5270: 5267: 5262: 5259: 5257: 5256: 5251: 5248: 5245: 5243: 5240: 5238: 5237: 5233: 5232: 5229: 5226: 5224: 5221: 5219: 5218: 5214: 5213: 5210: 5207: 5205: 5202: 5200: 5199: 5195: 5194: 5191: 5188: 5184: 5182: 5179: 5177: 5176: 5172: 5171: 5168: 5166: 5163: 5162: 5159: 5157: 5154: 5148: 5147:P. cf. primus 5145: 5143: 5142: 5137: 5134: 5132: 5129: 5128: 5125: 5120: 5117: 5115: 5114: 5109: 5106: 5104: 5101: 5100: 5097: 5095: 5094:acanthodontid 5089: 5086: 5084: 5083: 5078: 5075: 5073: 5070: 5069: 5066: 5061: 5058: 5056: 5055: 5050: 5047: 5045: 5041: 5039: 5036: 5034: 5033: 5029: 5028: 5025: 5023: 5019: 5017: 5014: 5012: 5011: 5007: 5006: 5003: 5001: 4997: 4995: 4994:C. longibasis 4992: 4990: 4989: 4985: 4984: 4981: 4979: 4978:prioniodontid 4976: 4972: 4970: 4967: 4965: 4964: 4960: 4959: 4955: 4952: 4949: 4946: 4945: 4941: 4937: 4934: 4933: 4930: 4926: 4922: 4918: 4913: 4912: 4908: 4895: 4893: 4890: 4887: 4884: 4882: 4879: 4877: 4876: 4872: 4871: 4868: 4866: 4863: 4859: 4856: 4853: 4851: 4848: 4846: 4845: 4841: 4840: 4837: 4834: 4830: 4827: 4825: 4822: 4820: 4819: 4815: 4814: 4811: 4808: 4804: 4801: 4799: 4796: 4794: 4793: 4789: 4788: 4785: 4782: 4778: 4774: 4772: 4770: 4765: 4763: 4758: 4754: 4752: 4749: 4747: 4746: 4742: 4741: 4738: 4736: 4733: 4729: 4725: 4722: 4720: 4717: 4715: 4714: 4710: 4709: 4706: 4703: 4699: 4696: 4692: 4690: 4687: 4684: 4683: 4679: 4678: 4675: 4673: 4670: 4669:Dissidocerida 4666: 4662: 4659: 4657: 4654: 4652: 4651: 4647: 4646: 4643: 4641: 4638: 4637:paragastropod 4635: 4631: 4628: 4625: 4623: 4620: 4618: 4617: 4613: 4612: 4609: 4607: 4604: 4600: 4598: 4596: 4592: 4590: 4585: 4581: 4579: 4576: 4574: 4573: 4569: 4568: 4565: 4562: 4558: 4555: 4551: 4549: 4546: 4544: 4543: 4539: 4538: 4535: 4532: 4530: 4528: 4523: 4519: 4517: 4514: 4512: 4511: 4507: 4506: 4503: 4501: 4498: 4494: 4490: 4487: 4485: 4482: 4480: 4479: 4475: 4474: 4471: 4467: 4465: 4463: 4457: 4453: 4451: 4448: 4446: 4445: 4441: 4440: 4437: 4434: 4430: 4426: 4423: 4419: 4417: 4414: 4411: 4410: 4406: 4405: 4402: 4399: 4395: 4392: 4388: 4386: 4383: 4382: 4378: 4377: 4374: 4372: 4369: 4366: 4362: 4360: 4357: 4356: 4353: 4351: 4348: 4345: 4341: 4339: 4337: 4331: 4328: 4326: 4323: 4322: 4319: 4317: 4314: 4311: 4307: 4305: 4303: 4298: 4294: 4292: 4289: 4287: 4286: 4281: 4278: 4275: 4271: 4267: 4263: 4259: 4255: 4251: 4248: 4245: 4243: 4240: 4238: 4237: 4233: 4232: 4229: 4226: 4224: 4221: 4220: 4217: 4214: 4210: 4209:bathmoceratid 4203: 4201: 4198: 4196: 4195: 4190: 4187: 4185: 4182: 4179:in the order 4178: 4174: 4170: 4166: 4163: 4161: 4158: 4156: 4155: 4151: 4150: 4147: 4144: 4140: 4137: 4136:endosymbiotic 4133: 4129: 4127: 4125: 4120: 4118: 4113: 4109: 4107: 4104: 4102: 4101: 4097: 4096: 4093: 4090: 4087: 4083: 4080: 4077: 4073: 4070: 4066: 4064: 4061: 4058: 4057: 4053: 4052: 4048: 4045: 4043:Stratigraphy 4042: 4039: 4036: 4035: 4031: 4027: 4015: 4012: 4009: 4006: 4004: 4001: 3999: 3998: 3994: 3993: 3987: 3983: 3981: 3977: 3973: 3969: 3966: 3962: 3959: 3956: 3954: 3951: 3949: 3948: 3944: 3943: 3940: 3937: 3934: 3930: 3928: 3925: 3924: 3921: 3918: 3915: 3912: 3910: 3907: 3905: 3904: 3903:Thoralicystis 3899: 3896: 3893: 3892:R. destombesi 3889: 3887: 3885: 3880: 3877: 3875: 3872: 3871: 3868: 3865: 3864:R. destombesi 3861: 3858: 3855: 3853: 3850: 3849: 3846: 3843: 3839: 3836: 3833: 3831: 3828: 3827: 3824: 3821: 3820:R. destombesi 3817: 3814: 3811: 3809: 3806: 3805: 3802: 3798: 3796: 3794: 3789: 3787: 3783: 3781: 3776: 3773: 3771: 3770:R. destombesi 3768: 3766: 3765: 3764:Rhopalocystis 3760: 3757: 3754: 3750: 3748: 3746: 3740: 3736: 3734: 3731: 3729: 3728: 3727:Ramseyocrinus 3724: 3723: 3720: 3717: 3714: 3711: 3709: 3706: 3704: 3703: 3702:Prokopicystis 3699: 3698: 3695: 3692: 3689: 3686: 3684: 3681: 3679: 3678: 3674: 3673: 3670: 3667: 3663: 3659: 3656: 3652: 3649: 3647: 3644: 3642: 3641: 3637: 3636: 3633: 3630: 3627: 3624: 3622: 3619: 3617: 3616: 3612: 3611: 3608: 3605: 3603: 3601: 3595: 3592: 3590: 3588: 3587: 3582: 3581: 3578: 3575: 3573: 3571: 3565: 3562: 3560: 3557: 3555: 3553: 3549: 3548: 3542: 3538: 3535: 3532: 3528: 3525: 3522: 3520: 3517: 3515: 3514: 3510: 3509: 3506: 3503: 3500: 3497: 3495: 3492: 3490: 3489: 3485: 3484: 3481: 3478: 3474: 3471: 3467: 3465: 3463: 3457: 3454: 3452: 3449: 3447: 3446: 3445:Macrocystella 3442: 3441: 3438: 3435: 3433: 3431: 3425: 3422: 3420: 3417: 3416: 3413: 3410: 3408: 3406: 3400: 3396: 3394: 3393:L. aff. deani 3391: 3389: 3388: 3387:Lingulocystis 3383: 3380: 3377: 3374: 3371: 3369: 3366: 3364: 3363: 3359: 3358: 3355: 3352: 3348: 3345: 3341: 3339: 3336: 3334: 3333: 3329: 3328: 3325: 3322: 3319: 3316: 3314: 3311: 3309: 3308: 3304: 3303: 3300: 3297: 3294: 3291: 3289: 3286: 3285: 3282: 3279: 3276: 3273: 3271: 3268: 3266: 3265: 3264:Galliaecystis 3260: 3257: 3254: 3251: 3248: 3246: 3243: 3241: 3240: 3236: 3235: 3232: 3229: 3226: 3222: 3220: 3217: 3215: 3214: 3210: 3209: 3206: 3203: 3200: 3197: 3195: 3192: 3191: 3188: 3185: 3182: 3179: 3177: 3174: 3173: 3170: 3167: 3164: 3161: 3159: 3156: 3154: 3153: 3148: 3145: 3142: 3139: 3136: 3134: 3131: 3129: 3128: 3124: 3123: 3120: 3117: 3113: 3109: 3105: 3103: 3101: 3096: 3094: 3088: 3085: 3083: 3080: 3078: 3077: 3073: 3072: 3069: 3066: 3063: 3059: 3057: 3054: 3052: 3051: 3047: 3046: 3043: 3040: 3037: 3034: 3032: 3029: 3027: 3026: 3022: 3021: 3018: 3015: 3011: 3008: 3005: 3003: 3000: 2998: 2997: 2993: 2992: 2989: 2986: 2984: 2982: 2976: 2972: 2970: 2967: 2965: 2964: 2960: 2959: 2956: 2953: 2952:edrioasteroid 2949: 2946: 2942: 2940: 2937: 2935: 2934: 2930: 2929: 2926: 2923: 2921: 2919: 2914: 2912: 2907: 2904: 2902: 2899: 2898: 2895: 2892: 2888: 2885: 2882: 2880: 2877: 2875: 2874: 2869: 2866: 2863: 2860: 2857: 2855: 2852: 2850: 2849: 2848:Amygdalotheca 2845: 2844: 2841: 2838: 2834: 2831: 2828: 2826: 2823: 2821: 2820: 2816: 2815: 2812: 2809: 2805: 2802: 2799: 2797: 2796:A. cf. moorei 2794: 2792: 2791: 2787: 2786: 2782: 2779: 2777:Stratigraphy 2776: 2773: 2770: 2769: 2765: 2761: 2758: 2754: 2752: 2748: 2744: 2743:Macrocystella 2740: 2739:Rhopalocystis 2734: 2732: 2728: 2724: 2723:Rhopalocystis 2720: 2719:Rhopalocystis 2716: 2711: 2709: 2705: 2701: 2697: 2693: 2689: 2685: 2671: 2668: 2666: 2664: 2661: 2659: 2658: 2654: 2653: 2647: 2643: 2640: 2636: 2632: 2629: 2626: 2624: 2621: 2619: 2618: 2614: 2613: 2610: 2607: 2604: 2600: 2596: 2593: 2589: 2586: 2583: 2581: 2578: 2576: 2575: 2571: 2570: 2567: 2564: 2563: 2558: 2554: 2553: 2548: 2547: 2542: 2541:euchelicerate 2538: 2534: 2530: 2527: 2524: 2522: 2521:S. abundantis 2519: 2517: 2516: 2512: 2511: 2505: 2501: 2498: 2497: 2492: 2488: 2484: 2480: 2477: 2474: 2472: 2469: 2465: 2464: 2458: 2454: 2451: 2447: 2446:marrellomorph 2444: 2443:acercostracan 2440: 2437: 2434: 2432: 2429: 2427: 2426: 2422: 2421: 2418: 2415: 2411: 2407: 2404: 2402: 2401:B. singularis 2399: 2397: 2396: 2392: 2391: 2387: 2384: 2382:Stratigraphy 2381: 2378: 2375: 2374: 2370: 2365: 2362: 2360: 2357: 2353: 2349: 2348: 2343: 2339: 2335: 2331: 2327: 2323: 2319: 2315: 2311: 2297: 2294: 2290: 2288: 2286: 2281: 2279: 2274: 2271: 2269: 2268:T. aff. borni 2266: 2264: 2263: 2259: 2258: 2255: 2252: 2248: 2245: 2242: 2240: 2237: 2236: 2233: 2230: 2227: 2225: 2222: 2221: 2218: 2214: 2211: 2208: 2206: 2203: 2202: 2199: 2196: 2194: 2190: 2187: 2185: 2184:S. angustatus 2182: 2180: 2179: 2174: 2168: 2164: 2161: 2160:odontopleurid 2157: 2155: 2153: 2151: 2150: 2146: 2145: 2142: 2139: 2138:pharostomatid 2135: 2133: 2131: 2126: 2123: 2121: 2118: 2116: 2115: 2114:Prionocheilus 2111: 2110: 2107: 2105:A calymenid. 2104: 2102: 2100: 2095: 2092: 2090: 2087: 2085: 2084: 2080: 2079: 2073: 2069: 2065: 2062: 2059: 2057: 2054: 2052: 2051: 2047: 2046: 2043: 2040: 2036: 2033: 2031: 2028: 2026: 2025: 2021: 2020: 2017: 2014: 2013:pharostomatid 2010: 2007: 2005: 2002: 2000: 1998: 1994: 1993: 1990: 1987: 1983: 1980: 1978: 1975: 1973: 1972: 1968: 1967: 1964: 1961: 1957: 1955: 1953: 1948: 1946: 1941: 1938: 1936: 1933: 1931: 1930: 1926: 1925: 1922: 1919: 1916: 1913: 1911: 1908: 1906: 1905: 1901: 1900: 1897: 1894: 1891: 1888: 1886: 1883: 1882: 1879: 1876: 1874: 1872: 1869: 1867: 1866: 1861: 1858: 1855: 1853: 1851: 1848: 1846: 1844: 1843:Nobiliasaphus 1840: 1839: 1836: 1833: 1829: 1826: 1823: 1821: 1818: 1816: 1815: 1811: 1810: 1807: 1804: 1802: 1800: 1795: 1792: 1790: 1787: 1785: 1784: 1780: 1779: 1776: 1773: 1772: 1766: 1763: 1760: 1758: 1755: 1754: 1751: 1748: 1746: 1742: 1739: 1737: 1734: 1731: 1727: 1726: 1721: 1718: 1715: 1714:lichakephalid 1711: 1709: 1707: 1704: 1702: 1701: 1700:Lichakephalus 1697: 1696: 1693: 1690: 1686: 1682: 1680: 1678: 1673: 1670: 1668: 1665: 1663: 1662: 1658: 1657: 1654: 1651: 1647: 1644: 1641: 1639: 1636: 1634: 1633: 1629: 1628: 1622: 1618: 1615: 1611: 1609: 1607: 1604: 1602: 1601: 1597: 1596: 1590: 1586: 1583: 1579: 1575: 1573: 1571: 1566: 1563: 1561: 1558: 1556: 1555: 1551: 1550: 1547: 1545:A cheirurid. 1544: 1542: 1540: 1535: 1532: 1530: 1527: 1525: 1524: 1520: 1519: 1516: 1513: 1509: 1507: 1505: 1500: 1498: 1493: 1490: 1488: 1485: 1483: 1482: 1478: 1477: 1471: 1467: 1464: 1460: 1458: 1456: 1453: 1451: 1450: 1446: 1445: 1442: 1439: 1435: 1432: 1429: 1427: 1424: 1421: 1420: 1416: 1415: 1409: 1405: 1402: 1399: 1397: 1394: 1393: 1390: 1387: 1383: 1380: 1377: 1375: 1374:D. brenchleyi 1372: 1370: 1369: 1364: 1361: 1358: 1356: 1354: 1351: 1349: 1348: 1344: 1343: 1337: 1333: 1330: 1328: 1326: 1321: 1318: 1316: 1313: 1312: 1309: 1306: 1304: 1302: 1297: 1294: 1292: 1289: 1288: 1285: 1282: 1278: 1276: 1274: 1269: 1265: 1263: 1260: 1258: 1257: 1252: 1249: 1246: 1242: 1239: 1237: 1234: 1232: 1230: 1226: 1225: 1222: 1219: 1217: 1213: 1210: 1208: 1205: 1204: 1201: 1198: 1194: 1191: 1188: 1186: 1183: 1181: 1180: 1175: 1172: 1169: 1165: 1162: 1160: 1157: 1155: 1154: 1150: 1149: 1146: 1143: 1141: 1139: 1136: 1135: 1129: 1125: 1122: 1120: 1118: 1115: 1114: 1111: 1108: 1105: 1102: 1100: 1099:B. destombesi 1097: 1095: 1094: 1089: 1086: 1083: 1079: 1076: 1074: 1071: 1069: 1068: 1064: 1063: 1060: 1056: 1054: 1050: 1047: 1045: 1042: 1041: 1035: 1031: 1028: 1025: 1022: 1020: 1017: 1016: 1013: 1010: 1007: 1004: 1002: 999: 998: 995: 992: 990: 988: 983: 981: 976: 973: 971: 968: 967: 964: 961: 957: 954: 950: 947: 944: 942: 939: 937: 936: 931: 928: 925: 923: 919: 917: 914: 913: 910: 907: 906:remopleuridid 903: 901: 899: 894: 891: 889: 886: 884: 883: 882:Apatokephalus 878: 875: 872: 868: 864: 860: 857: 853: 850: 847: 845: 842: 840: 839: 835: 834: 831: 828: 824: 823: 818: 814: 811: 807: 804: 802: 799: 797: 796: 792: 791: 788: 785: 781: 779: 777: 772: 768: 766: 763: 761: 760: 756: 755: 751: 748: 746:Stratigraphy 745: 742: 739: 738: 734: 730: 727: 715: 712: 709: 707: 704: 703: 700: 697: 694: 692: 689: 688: 682: 678: 675: 668: 665: 663: 660: 658: 657: 652: 646: 642: 639: 638:panarthropods 635: 632: 629: 625: 623: 621: 616: 613: 611: 608: 606: 605: 601: 600: 596: 593: 591:Stratigraphy 590: 587: 584: 583: 579: 575: 572: 570: 560: 552: 550: 540: 532: 529: 525: 521: 517: 507: 505: 502: 498: 494: 490: 480: 478: 469: 460: 458: 454: 453:cheloniellids 450: 449:machaeridians 446: 442: 438: 434: 430: 429:paleoscolecid 426: 422: 418: 417: 412: 408: 404: 400: 396: 391: 387: 383: 379: 375: 367: 362: 353: 351: 347: 343: 339: 335: 331: 327: 323: 279: 270: 267: 256: 252: 249: 246: 242: 237: 209: 207: 203: 198: 170: 168: 164: 159: 156: 153: 149: 144: 140: 136: 133: 130: 126: 123: 120: 116: 111: 110:marrellomorph 105: 100: 94: 89: 84: 79: 74: 69: 64: 59: 54: 49: 44: 39: 29: 25: 20: 10802: 10767:Spence Shale 10747:Pioche Shale 10655: 10651: 10641: 10598: 10594: 10584: 10559: 10518: 10514: 10504: 10479: 10475: 10465: 10456:11449/161915 10430: 10426: 10386: 10382: 10328: 10324: 10268: 10264: 10226: 10222: 10182: 10178: 10138: 10134: 10094: 10090: 10008: 10004: 9965:(2): 61–75. 9962: 9958: 9901: 9897: 9827: 9823: 9813: 9778: 9722: 9718: 9672: 9668: 9626: 9622: 9584: 9580: 9570: 9545: 9541: 9458: 9454: 9444: 9411: 9407: 9343: 9339: 9297: 9293: 9283: 9256: 9252: 9242: 9201: 9197: 9141: 9137: 9087: 9083: 9049: 9045: 9032: 9006: 9000: 8972: 8968: 8905:(1): 39728. 8902: 8898: 8876: 8872: 8836: 8826: 8793: 8789: 8767: 8763: 8701: 8697: 8675: 8671: 8665: 8632: 8628: 8618: 8593: 8589: 8492: 8488: 8426: 8422: 8356:(1): 14941. 8353: 8349: 8309: 8305: 8195: 8191: 8130: 8126: 8060:(1): 20773. 8057: 8053: 7990: 7986: 7944: 7940: 7878: 7874: 7843:. Retrieved 7838: 7826: 7809: 7803: 7790: 7783: 7771:. Retrieved 7766: 7757: 7745:. Retrieved 7740: 7730: 7718:. Retrieved 7713: 7703: 7668: 7664: 7609: 7605: 7565: 7561: 7555: 7514: 7510: 7472: 7468: 7413:(1): 20807. 7410: 7406: 7395: 7354: 7350: 7288: 7265: 7242: 7219: 7200: 7196: 7161:Chitinozoans 7159: 7146: 7137: 7130: 7101:machaeridian 7093: 7089:P. bengtsoni 7088: 7081: 7059: 7052: 7034: 7030:P.? tenensis 7029: 7023:Palaeoscolex 7021: 7002: 6995: 6976: 6969: 6950: 6942: 6934: 6925: 6918: 6899: 6892: 6873: 6868:Cavernolites 6866: 6858: 6845: 6838: 6804:problematica 6785: 6766: 6758: 6750: 6741: 6734: 6725: 6721: 6713:'H.' christi 6712: 6707: 6693: 6686: 6679:A "choiid". 6672: 6665: 6657: 6648: 6641: 6604: 6599: 6596: 6593: 6553: 6526: 6519: 6497: 6490: 6468: 6461: 6439: 6432: 6410: 6403: 6381: 6374: 6352: 6345: 6323: 6316: 6267: 6260: 6249: 6247: 6243: 6238:Tetragraptus 6237: 6227:tetragraptid 6219: 6213: 6206:Tetragraptus 6204: 6193: 6186: 6181: 6174: 6162: 6159:pterobranchs 6154:Rhabdopleura 6152: 6145: 6127: 6122: 6111: 6110: 6106: 6101: 6091: 6086: 6074: 6067: 6062: 6057:Rhabdinopora 6055: 6041: 6040: 6036: 6031: 6024: 6014: 6009: 6002:graptoloid. 6000:sigmagraptid 5991: 5990: 5986: 5981: 5974: 5960: 5959: 5955: 5950: 5941: 5928: 5923: 5918:Hunnegraptus 5916: 5903: 5898: 5891: 5878: 5872: 5865: 5851: 5850: 5846: 5841: 5832: 5819: 5813: 5805: 5794: 5793: 5789: 5784: 5775:dichograptid 5766: 5764: 5760: 5755:C. multiplex 5754: 5749:Clonograptus 5747: 5734: 5729: 5722: 5712: 5707: 5697: 5692: 5682: 5677:?B. jacksoni 5676: 5659: 5654: 5649:Baltograptus 5647: 5638:monograptids 5631:Azygograptus 5630: 5624: 5619:Azygograptus 5617: 5607: 5600: 5595: 5588: 5575: 5570: 5565:Anisograptus 5563: 5546: 5541: 5534: 5518:anisograptid 5510: 5507:Adelograptus 5506: 5500: 5494:Adelograptus 5491: 5459: 5456:Azygograptus 5455: 5451: 5447: 5443: 5441: 5438: 5434: 5430: 5427:Azygograptus 5426: 5422: 5418: 5414: 5410: 5406: 5402: 5398: 5395:Adelograptus 5394: 5390: 5386: 5384: 5373: 5367: 5361: 5358: 5350:Rhabdinopora 5348: 5342: 5336: 5322: 5310: 5303: 5289:S. cf. stola 5288: 5281: 5272: 5260: 5253: 5241: 5234: 5222: 5217:Scalpellodus 5215: 5203: 5196: 5180: 5173: 5164: 5146: 5139: 5130: 5118: 5111: 5102: 5087: 5082:Parapaltodus 5080: 5071: 5059: 5052: 5037: 5030: 5015: 5008: 4993: 4986: 4968: 4961: 4932: 4928: 4924: 4920: 4914: 4911: 4907: 4905: 4892: 4885: 4880: 4875:Thoralispira 4873: 4865: 4854: 4849: 4842: 4831:A very rare 4823: 4816: 4797: 4790: 4777:actinodontid 4769:Azygograptus 4768: 4766: 4761: 4760: 4756: 4750: 4743: 4735: 4726:A very rare 4718: 4711: 4700:A very rare 4694: 4688: 4680: 4672: 4655: 4648: 4640: 4634:onychochilid 4626: 4621: 4614: 4606: 4594: 4593: 4588: 4587: 4583: 4577: 4570: 4553: 4547: 4540: 4526: 4525: 4521: 4515: 4510:Ekaterodonta 4508: 4500: 4484:D. zagorense 4483: 4476: 4462:Azygograptus 4461: 4459: 4455: 4449: 4442: 4429:actinodontid 4427:A very rare 4421: 4415: 4407: 4391:Azygograptus 4390: 4379: 4371: 4364: 4359:C. vizcainoi 4358: 4350: 4335: 4333: 4329: 4324: 4316: 4301: 4300: 4296: 4290: 4283: 4246: 4241: 4234: 4222: 4207:A very rare 4199: 4192: 4184: 4169:baltoceratid 4167:A very rare 4159: 4152: 4124:Azygograptus 4123: 4121: 4116: 4115: 4111: 4105: 4098: 4069:Azygograptus 4068: 4062: 4054: 4007: 4002: 3995: 3979: 3975: 3957: 3952: 3945: 3932: 3926: 3913: 3908: 3901: 3891: 3883: 3882: 3878: 3873: 3863: 3856: 3852:R. havliceki 3851: 3842:R. havliceki 3841: 3834: 3829: 3819: 3812: 3807: 3792: 3790: 3785: 3784: 3779: 3778: 3774: 3769: 3762: 3744: 3742: 3738: 3732: 3725: 3712: 3707: 3700: 3687: 3682: 3675: 3665: 3654: 3650: 3645: 3640:Plasiacystis 3638: 3625: 3620: 3613: 3599: 3597: 3593: 3583: 3569: 3567: 3563: 3558: 3550: 3529:An uncommon 3523: 3519:N. agterbosi 3518: 3513:Nimchacystis 3511: 3498: 3493: 3486: 3476: 3461: 3459: 3455: 3450: 3443: 3429: 3427: 3423: 3418: 3404: 3402: 3398: 3392: 3385: 3372: 3367: 3360: 3343: 3337: 3330: 3317: 3312: 3305: 3292: 3287: 3274: 3269: 3262: 3249: 3244: 3237: 3225:Azygograptus 3224: 3218: 3211: 3198: 3194:C. vizcainoi 3193: 3180: 3175: 3162: 3157: 3150: 3137: 3132: 3125: 3115: 3111: 3099: 3097: 3092: 3090: 3086: 3081: 3074: 3061: 3055: 3048: 3035: 3030: 3025:Aspidocarpus 3023: 3006: 3001: 2994: 2980: 2978: 2974: 2968: 2961: 2944: 2938: 2931: 2917: 2915: 2910: 2909: 2905: 2900: 2883: 2879:A. escandrei 2878: 2871: 2858: 2853: 2846: 2829: 2824: 2819:Ampelocarpus 2817: 2800: 2795: 2788: 2755: 2750: 2747:Plasiacystis 2746: 2742: 2738: 2735: 2730: 2727:diploporites 2722: 2718: 2714: 2712: 2708:somasteroids 2704:rhenopyrgids 2681: 2662: 2657:Tremaglaspis 2655: 2639:chelicerates 2627: 2622: 2615: 2584: 2579: 2572: 2560: 2550: 2544: 2525: 2520: 2513: 2494: 2475: 2435: 2430: 2423: 2400: 2393: 2345: 2342:cheloniellid 2338:leanchoiliid 2334:canadaspidid 2326:phyllocarids 2307: 2284: 2282: 2277: 2276: 2272: 2267: 2262:Toletanaspis 2260: 2250: 2243: 2238: 2223: 2209: 2205:S. ebbestadi 2204: 2192: 2188: 2183: 2176: 2149:Selenopeltis 2147: 2129: 2128: 2124: 2119: 2112: 2098: 2097: 2093: 2088: 2081: 2060: 2055: 2048: 2029: 2024:Platycoryphe 2022: 2003: 1997:Pharostomina 1995: 1976: 1969: 1960:bathycheilid 1951: 1949: 1944: 1943: 1939: 1934: 1927: 1914: 1909: 1902: 1889: 1884: 1870: 1863: 1849: 1841: 1824: 1819: 1812: 1798: 1797: 1793: 1788: 1781: 1769: 1761: 1756: 1744: 1740: 1735: 1729: 1723: 1705: 1698: 1688: 1676: 1675: 1671: 1666: 1659: 1650:ptychopariid 1642: 1637: 1630: 1605: 1598: 1576:A very rare 1569: 1568: 1564: 1559: 1552: 1538: 1537: 1533: 1529:F. peregrina 1528: 1521: 1503: 1501: 1496: 1495: 1491: 1486: 1479: 1454: 1447: 1430: 1425: 1417: 1395: 1378: 1373: 1366: 1352: 1345: 1324: 1323: 1319: 1314: 1300: 1299: 1295: 1290: 1272: 1271: 1267: 1261: 1256:Colpocoryphe 1254: 1235: 1229:Ceraurinella 1227: 1215: 1211: 1206: 1189: 1184: 1177: 1168:bathycheilid 1158: 1153:Bathycheilus 1151: 1137: 1116: 1103: 1098: 1091: 1081: 1072: 1065: 1052: 1048: 1044:A. tataensis 1043: 1023: 1018: 1005: 1000: 986: 984: 979: 978: 974: 969: 959: 945: 940: 933: 921: 915: 904:An uncommon 897: 896: 892: 887: 880: 870: 866: 848: 843: 838:Anacheirurus 836: 826: 820: 817:raphiophorid 809: 805: 800: 793: 775: 774: 770: 764: 757: 725: 710: 705: 695: 690: 674:Aegirocassis 673: 666: 661: 654: 619: 618: 614: 610:A. benmoulae 609: 604:Aegirocassis 602: 568: 566: 558: 546: 538: 513: 500: 496: 486: 483:Preservation 474: 441:problematica 440: 419:), possible 416:Aegirocassis 414: 371: 365: 349: 345: 325: 321: 319: 233:-73.9; 108.1 189:30.6°N 5.7°W 108:Fossil of a 10849:Tremadocian 10793:Castle Bank 10482:: 130–141. 10433:: 170–178. 10271:(1): 1002. 9904:: 155–169. 9748:11336/44871 9346:(1): 3808. 9204:(3–4): 21. 9001:Symphysurus 8312:: 142–154. 7845:13 November 7671:: 274–283. 7304:Fossilworks 7281:Fossilworks 7258:Fossilworks 7235:Fossilworks 7108:polychaetes 7069:pauxillitid 7054:Pauxillites 6997:Nephrotheca 6971:Gompholites 6920:Eoconularia 6760:Valospongia 6749:A piraniid. 6708:'Hamptonia' 6656:A "choiid". 6583:reticulosan 6571:hamptoniids 6563:leptomitids 6559:demosponges 6536:heterorthid 6527:T. marocana 6498:R. fasciata 6478:plectorthid 6463:Plectorthis 6423:linguliform 6394:linguliform 6336:linguliform 6292:Brachiopods 6284:Brachiopods 6194:Holograptus 5924:H. copiosus 5893:Holograptus 5655:B. deflexus 5467:Graptolites 5458:interval"). 5393:zone (2), “ 5333:Graptolites 5187:balognathid 5156:ozarkodinid 5153:periodontid 5113:Paroistodus 5016:D. arcuatus 4969:A. deltatus 4807:rostroconch 4751:R. michelae 4702:praenuculid 4616:Pelecyogyra 4603:lesuerillid 4572:Lesueurilla 4497:Discosorida 4450:C. guiraudi 4444:Coxiconchia 4381:Cardiolaria 4262:halkieriids 4254:aculiferans 4242:C. kroegeri 4236:Calvapilosa 4200:B. australe 4194:Bathmoceras 4181:Orthocerida 4154:Bactroceras 4141:-oxidizing 4079:protobranch 4076:ctenodontid 4056:Alococoncha 3968:somasteroid 3615:Peltocystis 3586:Pareocrinus 3559:P.? prokopi 3473:rhombiferan 3451:M. bohemica 3419:L. elongata 3362:Lagnocystis 3245:F. rushtoni 3082:B. regnelli 3014:diploporite 2933:Anedriophus 2873:Anatifopsis 2790:Aethocrinus 2764:Echinoderms 2678:Echinoderms 2635:mollisoniid 2546:Dibasterium 2515:Setapedites 2496:nomen nudum 2431:E. hrungnir 2356:lepadomorph 2318:eurypterids 2314:xiphosurans 2251:Symphysurus 2178:Symphysurus 2039:homalonotid 1977:P. marochii 1971:Parvilichas 1935:P. gallicus 1820:N. deynouxi 1725:Megistaspis 1554:Geragnostus 1419:Ectillaenus 1159:B. gallicus 871:Parapilekia 784:leiostegiid 765:A. quadrata 524:Tremadocian 518:, north of 516:Draa Valley 457:xiphosurans 421:halkieriids 395:Demosponges 390:echinoderms 386:graptolites 374:mineralized 231: / 192: / 167:Coordinates 10828:Categories 10798:Soom Shale 10732:Chengjiang 9548:: 97–121. 8764:Batalleria 7568:(3): 183. 7175:References 7165:acritarchs 7114:and kin). 7112:scaleworms 7083:Plumulites 7060:P. thaddei 6855:conulariid 6800:priapulids 6742:P. auraeum 6608:lithistids 6469:P. simplex 6449:paurorthid 6434:Paurorthis 6420:acrothelid 6382:L. salteri 6333:acrothelid 6214:T. bulmani 6076:Dictyonema 5785:C. rigidus 5708:B. minutus 5693:?B. kurcki 5633:interval” 5596:A. murrayi 5542:A. bulmani 5525:graptoloid 5421:zone (8), 5409:zone (5), 5405:zone (4), 5401:zone (3), 5369:Dictyonema 5339:graptoloid 5295:belodellid 5236:Scolopodus 5175:Prioniodus 5131:P. proteus 5010:Drepanodus 4864:gastropod. 4833:rioceratid 4781:Redoniidae 4771:interval" 4732:Endocerida 4682:Praenucula 4605:gastropod. 4464:interval" 4409:Cienagomya 4393:interval" 4258:stem group 4177:cephalopod 4126:interval" 4071:interval" 3830:R. grandis 3646:P. mobilis 3621:P. cornuta 3488:Nanocarpus 3270:G. ubaghsi 3227:interval" 3176:C. ubaghsi 3158:C. spinosa 2963:Argodiscus 2854:A. griffei 2700:eocrinoids 2425:Enosiaspis 2410:aglaspidid 2369:arthropods 2352:retifaciid 2293:dalmanitid 2224:S. sicardi 1904:Orometopus 1783:Neseuretus 1730:Ekeraspsis 1706:L. stubbsi 1667:L. salteri 1632:Indiligens 1578:artiopodan 1291:C. thorali 1197:bavarillid 1067:Asaphopsis 1019:A. stubbsi 935:Asaphellus 844:A. adserai 801:A. priscus 733:Trilobites 722:Trilobites 691:P. izdigua 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1728:( 1502:? 1266:? 1231:? 985:? 769:? 93:N 83:K 78:J 73:T 68:P 63:C 58:D 53:S 48:O 43:ęž’

Index

Stratigraphic range
Preęž’
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O
S
D
C
P
T
J
K
Pg
N

marrellomorph
Geological formation
Zini Formation
Shale
Coordinates
30°36′N 5°42′W / 30.6°N 5.7°W / 30.6; -5.7
Approximate paleocoordinates
73°54′S 108°06′E / 73.9°S 108.1°E / -73.9; 108.1
Drâa-Tafilalet
Morocco
Fezouata Formation is located in Morocco
Morocco
Early Ordovician
exceptionally preserved
Burgess shale-type deposits

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