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Heterocentrotus

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are polygeminate, with between 9 and 14 pairs of pores per plate, arranged in more or less regular arcs, sometimes almost in bands. All ambulacral plates bear a single, large primary tubercle, occupying almost the entire surface of the plate. The interambulacral plates bear a single, massive nipple,
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These two species are very close visually, and often difficult to distinguish, since they generally share the same distribution area. The main criteria for in situ identification are the radioles, which are generally longer (especially on the aboral side) and more clearly triangular in
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flanked by secondary tubercles on the aboral surface. The radioles are very thick and dense, usually three-faceted (or two or four, or round, mainly in H. mamillatus). They are never modified into scales or plates (unlike
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often has the secondary radioles white or the base of the primary radioles surrounded by a white ring, which never seems to be seen in
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is also usually darker (brown, dark orange...), but the great variability of colorations in this genus makes this criterion delicate.
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The apical disc is dicyclic, with a reduced periproct. The peristome is also reduced and elliptical, with limited mouth notches. The
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has less petalloid aboral ambulacral zones than its sister species, and fewer pore pairs per ambulacral plate (9-12 versus 15-16).
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are brightly colored tropical sea urchins with very thick spines that have given them the nickname "
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Indo-Pacific region (from the east coast of Africa to the Pacific archipelagos),
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The genus consists of sea urchins with rounded (but slightly elliptical)
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at opposite, at the apex of the aboral (upper) face.
457:Radiols (spikes) of a "slate pencil sea urchin" ( 855: 838: 845: 831: 475: 109: 874:Taxa named by Johann Friedrich von Brandt 514: 252: 510: 508: 856: 574: 573: 797: 505: 13: 308: 14: 890: 463:). They are a classical souvenir. 313:This genus contains two species: 801: 529:World Register of Marine Species 450: 434: 416: 356: 330: 241:basin, especially in Reunion or 122: 41: 237:. They are mainly found in the 248: 1: 515:Kroh A, Mooi R, eds. (2023). 468: 817:. You can help Knowledge by 218: 7: 365:Heterocentrotus trigonarius 10: 895: 796: 460:Heterocentrotus mamillatus 443:Heterocentrotus mamillatus 426:Heterocentrotus mamillatus 339:Heterocentrotus mamillatus 263:Heterocentrotus mamillatus 233:is a genus of the familia 582: 525:World Echinoidea Database 215: 210: 119:Scientific classification 117: 108: 23: 554:Natural History Museum 266: 809:This article about a 753:Paleobiology Database 491:Paleobiology Database 377:Indo-Pacific region. 256: 423:Radiols (spikes) of 274:slate pencil urchins 16:Genus of sea urchins 267: 869:Echinoidea genera 826: 825: 791: 790: 740:Open Tree of Life 576:Taxon identifiers 381: 380: 374: 348: 226: 225: 206: 886: 879:Echinoidea stubs 847: 840: 833: 805: 798: 784: 783: 771: 770: 761: 760: 748: 747: 735: 734: 722: 721: 709: 708: 696: 695: 683: 682: 670: 669: 657: 656: 644: 643: 631: 630: 618: 617: 616: 603: 602: 601: 571: 570: 564: 563: 561: 560: 546: 540: 539: 537: 535: 512: 503: 502: 500: 498: 479: 454: 438: 420: 368: 360: 342: 334: 316: 315: 201: 127: 126: 113: 103: 40: 29:Temporal range: 21: 20: 894: 893: 889: 888: 887: 885: 884: 883: 864:Heterocentrotus 854: 853: 852: 851: 794: 792: 787: 779: 774: 766: 764: 756: 751: 743: 738: 730: 725: 717: 712: 704: 699: 691: 686: 678: 673: 665: 660: 652: 647: 639: 634: 628:Heterocentrotus 626: 621: 614:Heterocentrotus 612: 611: 606: 597: 596: 591: 584:Heterocentrotus 578: 568: 567: 558: 556: 548: 547: 543: 533: 531: 519:Heterocentrotus 513: 506: 496: 494: 485:Heterocentrotus 481: 480: 476: 471: 464: 455: 446: 439: 430: 421: 407:Skeletal-wise, 322:Scientific name 311: 309:List of species 302:Colobocentrotus 276:", named after 270:Heterocentrotus 251: 230:Heterocentrotus 217:2 species (see 200: 197:Heterocentrotus 121: 104: 102: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 51: 46: 35: 34: 27: 25:Heterocentrotus 17: 12: 11: 5: 892: 882: 881: 876: 871: 866: 850: 849: 842: 835: 827: 824: 823: 806: 789: 788: 786: 785: 772: 762: 749: 736: 723: 710: 697: 684: 671: 658: 645: 632: 619: 604: 588: 586: 580: 579: 566: 565: 541: 504: 473: 472: 470: 467: 466: 465: 456: 449: 447: 440: 433: 431: 422: 415: 402:H. trigonarius 398:H. trigonarius 390:H. trigonarius 386:H. trigonarius 379: 378: 375: 361: 353: 352: 349: 335: 327: 326: 323: 320: 310: 307: 250: 247: 235:Echinometridae 224: 223: 213: 212: 208: 207: 193: 189: 188: 186:Echinometridae 183: 179: 178: 173: 169: 168: 163: 159: 158: 153: 149: 148: 143: 139: 138: 133: 129: 128: 115: 114: 106: 105: 97: 92: 87: 82: 77: 72: 67: 62: 57: 52: 47: 42: 28: 15: 9: 6: 4: 3: 2: 891: 880: 877: 875: 872: 870: 867: 865: 862: 861: 859: 848: 843: 841: 836: 834: 829: 828: 822: 820: 816: 812: 807: 804: 800: 799: 795: 782: 777: 773: 769: 763: 759: 754: 750: 746: 741: 737: 733: 728: 724: 720: 715: 711: 707: 702: 698: 694: 689: 685: 681: 676: 672: 668: 663: 659: 655: 650: 646: 642: 637: 633: 629: 624: 620: 615: 609: 605: 600: 594: 590: 589: 587: 585: 581: 577: 572: 555: 551: 545: 530: 526: 522: 521:Brandt, 1835" 520: 511: 509: 493:. Fossilworks 492: 488: 486: 478: 474: 462: 461: 453: 448: 445: 444: 437: 432: 428: 427: 419: 414: 413: 412: 410: 409:H. mamillatus 405: 403: 399: 395: 394:H. mamillatus 391: 387: 376: 372: 367: 366: 362: 359: 355: 354: 350: 346: 341: 340: 336: 333: 329: 328: 325:Distribution 324: 321: 318: 317: 314: 306: 304: 303: 297: 292: 290: 286: 281: 279: 275: 271: 265: 264: 259: 255: 246: 244: 240: 236: 232: 231: 222: 220: 214: 209: 204: 199: 198: 194: 191: 190: 187: 184: 181: 180: 177: 174: 171: 170: 167: 164: 161: 160: 157: 156:Echinodermata 154: 151: 150: 147: 144: 141: 140: 137: 134: 131: 130: 125: 120: 116: 112: 107: 100: 95: 90: 85: 80: 75: 70: 65: 60: 55: 50: 45: 39: 36:15.97โ€“0  32: 26: 22: 19: 819:expanding it 808: 793: 583: 557:. Retrieved 553: 550:"Acrocladia" 544: 532:. Retrieved 524: 518: 495:. Retrieved 490: 487:Brandt 1835" 484: 477: 458: 441: 424: 408: 406: 401: 397: 393: 389: 385: 382: 363: 337: 312: 300: 293: 282: 278:slate pencil 269: 268: 261: 239:Indo-Pacific 229: 228: 227: 216: 196: 195: 24: 18: 688:iNaturalist 608:Wikispecies 249:Description 176:Camarodonta 858:Categories 811:sea urchin 559:2022-11-13 534:22 October 497:22 October 469:References 296:ambulacrum 166:Echinoidea 289:periproct 142:Kingdom: 136:Eukaryota 33:โ€“recent, 593:Wikidata 429:sculpted 404:either. 345:Linnaeus 211:Species 182:Family: 152:Phylum: 146:Animalia 132:Domain: 768:4193306 706:1015335 680:2278865 599:Q508348 373:, 1816) 371:Lamarck 347:, 1758) 192:Genus: 172:Order: 162:Class: 31:Miocene 781:205582 765:uBio: 745:631179 719:656015 243:Hawaii 205:, 1835 203:Brandt 813:is a 776:WoRMS 758:33055 732:31179 701:IRMNG 693:47614 667:61037 654:7P5VB 641:84355 319:Image 815:stub 727:NCBI 714:ITIS 675:GBIF 636:BOLD 536:2023 499:2023 285:test 280:.. 258:test 219:text 44:Pre๊ž’ 662:EoL 649:CoL 623:AFD 260:of 860:: 778:: 755:: 742:: 729:: 716:: 703:: 690:: 677:: 664:: 651:: 638:: 625:: 610:: 595:: 552:. 527:. 523:. 507:^ 489:. 221:) 94:Pg 38:Ma 846:e 839:t 832:v 821:. 562:. 538:. 517:" 501:. 483:" 369:( 343:( 99:N 89:K 84:J 79:T 74:P 69:C 64:D 59:S 54:O 49:๊ž’

Index

Miocene
Ma
Pre๊ž’
๊ž’
O
S
D
C
P
T
J
K
Pg
N

Scientific classification
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Eukaryota
Animalia
Echinodermata
Echinoidea
Camarodonta
Echinometridae
Heterocentrotus
Brandt
text
Echinometridae
Indo-Pacific
Hawaii

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