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John L. Koprowski

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for the outdoors from the many family fishing trips. He would usually fish for a few hours and then sneaked off to catch turtles, frogs, or salamanders. Squirrels were one of the few species that could be found in an inner, highly urbanized, suburb and became a fascination for the young naturalist; the taxonomic group remained a research interest in his career. He had become fascinated by questions about animal behavior and how animals were able to survive in such challenging environments. Besides biology, John had strong interests in sports, especially baseball, and history; he coauthored a history of the Lakewood, Ohio public school system while a student in Advanced Placement U.S. History at Lakewood High School that was integrated into the curriculum. As a high school student, John was a member of the Latin Club, The Wilderness Society, and the Backpacking and Mountaineering Society that permitted him to combine his interests in science and the outdoors.
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both of his graduate degrees focused on the population, behavioral, and evolutionary ecology of tree squirrels, and continues to often work on squirrels as model organisms. His work has focused on the conservation of biodiversity, particularly mammals, and elucidating patterns in social and mating systems as well as the population response of wildlife to stressors in their environment such as non-native species, human development, disease, fire, and climate change.
270:. John was Professor of wildlife science and Director (2017-2020) in the School of Natural Resources and the Environment and a faculty affiliate of the Institute of the Environment until 2020 when he was granted Professor Emeritus status. In September 2020, he moved to the University of Wyoming as a Wyoming Excellence Chair and Dean of the Haub School of Environment and Natural Resources. 742:.     Empresa de Acueducto, Alcantarillado y Aseo de BogotĂĄ D.C. /EAB-ESP, CorporaciĂłn   AutĂłnoma Regional del Guavio - CORPOGUAVIO, Parques Nacionales Naturales de Colombia        (Parque Nacional Natural Chingaza, DirecciĂłn Territorial OrinoquĂ­a) & Proyectos de ConservaciĂłn de Aguas y Tierras - ProCAT Colombia. BogotĂĄ, D.C. Colombia. 358:
John was raised in the west side suburbs of Cleveland, Ohio spending various amounts of time living in apartments in Cleveland, Lakewood, Westlake, and Rocky River before the family settled into a house in Lakewood while he was in the 6th grade. His parents loved fishing and he obtained his passion
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John Koprowski's research extends from his early experience with the behavior and ecology of wildlife in human impacted environments, often using community-based conservation to enhance success. He conducted and published two papers on ground squirrels from undergraduate course projects, obtained
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John Koprowski is the eldest son of Judith and Anthony Koprowski, a homemaker and a city laborer, who also raised two younger sons, Jerome and Joseph Koprowski. John married Nancy M Cervenak on 21 September 1985. They have two children, Zachary D Koprowski and Emma I Koprowski.
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Mazzamuto, M.V., Merrick, M., Bisi, F., Koprowski, J.L., Wauters, L.A., Martinoli, A., 2020. Timing of resource availability drives divergent social systems and home range dynamics in ecologically similar tree squirrels. Frontiers in Ecology and Evolution 8:174.doi:
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Derbridge, J.J., J.A. Merkle, M.E. Bucci, P. Callahan, J.L. Koprowski, J.L. Polfus, P.R. Krausman. 2015.Experimentally derived δC and δN discrimination factors for gray wolves, and the impact of prior information in Bayesian mixing models. PLoS ONE 10(3):e0119940.
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Koprowski, J.L. 2007. Reproductive strategies and alternative reproductive tactics of tree squirrels. In: Wolff, J. Sherman, P (eds). Rodent Societies: an ecological and evolutionary perspective. Chapter 7: pages 86–95. University of Chicago
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Masoudi, A., U. Bhattarai, J.L. Koprowski, D. Wang. 2018. Elevational distribution and morphological attributes of the entomopathogenic fungi from forests of the Qinling Mountains in China. Applied Microbiology and Biotechnology
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Hale, S.L., Koprowski, J.L., Archer, S.R., 2020. The reintroduction of a small, native herbivore could limit woody plant proliferation in grasslands. Frontiers in Ecology and Evolution 8:233.doi:10.3389/fevo.2020.00233.
262:, the oldest university in the American west, in 1992 in Salem, Oregon. John was granted tenure and promoted during his time at Willamette. In 2000, John left Oregon to join the faculty of the wildlife program at the 682:
Krishna, M.C, A. Kumar, O.P. Tripathi, J.L. Koprowski. 2016. Diversity, distribution and status of gliding squirrels in protected and non-protected areas of the eastern Himalayas in India. Hystrix, pp. 1–9.
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Posthumus, E.E., J.L. Koprowski, R.S. Steidl. 2015. Red squirrel middens influence abundance but not    diversity of other vertebrates.  PLoS ONE 10(4): e0123633. doi:10.1371/journal.pone.0123633.
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Goldstein, E.A., M.J. Merrick, J.L. Koprowski. 2018. Low survival, high predation pressure present conservation challenges for an endangered endemic forest mammal. Biological Conservation, 221:67-77.
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Chen, H.L., J.L. Koprowski. 2016. Differential effects of roads and traffic on space use and movements of native forest-dependent and introduced edge-tolerant species. PLoS ONE 11(1): e0148121.
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Merrick, M.J., J.L. Koprowski. 2017. Circuit theory to estimate natal dispersal routes and functional landscape connectivity for an endangered small mammal. Landscape Ecology 32:1163–1179.
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Koprowski, J.L., K.E. Munroe, A.J. Edelman. 2016. Gray not grey: ecology of eastern gray squirrels in their native range. pp. 1–17. In: Shuttleworth, C., Lurz, P., Gurnell, J. (eds.)
565: 618: 582:. pp. 33–41 in M.A. Steele, J.F. Merritt, D.A. Zegers (eds.), Ecology and Evolutionary Biology of Tree Squirrels, Special Publication 6, Virginia Museum of Natural History. 310 pp. 926: 638:
Doumas, S.L., J.L. Koprowski. 2013. Return of fire as a restoration tool: long-term effects of burn severity on habitat use by Mexican fox squirrels. Restoration Ecology 21:133-139.
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Goldstein, E.A., M.J. Merrick, J.L. Koprowski. 2017. Functional semelparity drives population dynamics and endangers a peripheral population. Biological Conservation 205:52-59.
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Palmer, RR, Koprowski JL. 2018. Influence of time and flood on diurnal mammal diversity and story level use in IgapĂł Forest in the Peruvian Amazon. In: Myster RW (ed). 2018.
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Koprowski, JL, JF GonzĂĄlez-Maya, DA ZĂĄrrate-Charry, DA, UR Sharma, C Spencer. 2019. Local approaches and community-based conservation. In: Koprowski, JL, PR Krausman (eds).
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Merrick, M.J., J.L. Koprowski. 2017. Altered natal dispersal at the range periphery: the role of behaviour, resources, and maternal condition. Ecology and Evolution 7:58-72.
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Merrick, M.J., J.L. Koprowski. 2017. Should we consider individual behavior differences in applied wildlife conservation studies? Biological Conservation 209:34-44.
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Chen, H.L., J.L. Koprowski. 2016. Barrier effects of roads on an endangered forest obligate: impacts of traffic, edge, and gap. Biological Conservation 199:33-40.
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Mazzella, M., Koprowski, J.L., 2020. Response to fire by a forest specialist in isolated montane forest.     Forest Ecology and Management, 462:117996.
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McColgin, M.E., J.L. Koprowski, P.M. Waser. 2017. White-nosed coatis in Arizona: tropical carnivores in a temperate environment. Journal of Mammalogy 98: 64–74.
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and to serve as the Director of the Mt. Graham Biology Program that includes intensive research on one of the most endangered species in the United States, the
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at Kansas City. This teaching experience and his research on behavior and ecology of tree squirrels permitted John to join the Department of Biology faculty at
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Mendes, C.P., Koprowski, J.L., 2020. Does caching strategy vary with microclimate in endangered Mt.    Graham red squirrels? PLoS ONE 14: e0224947.
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Koprowski, J.L., E.A. Goldstein, K.R. Bennett, C. Pereira-Mendes. 2016. The Sciuridae. pp. 648–837, In: Wilson, D., Lacher, T., Mittermeier, R. (eds.)
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Fitak, R., J.L. Koprowski, M. Culver. 2013.  Severe reduction in genetic variation in a montane isolate: the endangered Mount Graham red squirrel (
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Koprowski, J.L. 2017. Introduction. In: GonzĂĄlez-Maya JF, Galindo-Tarazona R, Urquijo Collazos MM, ZĂĄrate Vanegas M & Parra-Romero A (eds). 2017.
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Merrick, M.J., J.L. Koprowski. 2016. Evidence of natal habitat preference induction within one habitat type. Proc. Royal Society London B 283:20152106.
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Chen, H.L., J.L. Koprowski. 2015. Animal occurrence and space use change in the landscape of anthropogenic noise. Biological Conservation 192: 315–322.
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Jo, Y.-S., J.T. Baccus, J.L. Koprowski. 2018. Mammals of Korea: a review of their taxonomy, distribution and conservation status. Zootaxa 4522:1-216.
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Hale, S.L., J.L. Koprowski. 2018. Ecosystem‐level effects of keystone species reintroduction: a literature review. Restoration Ecology 26:439-445.
214:, mammalogist, conservation biologist, and leading expert on the ecology and conservation of wildlife, especially squirrels, was born in 1961 in 745:
Koprowski, J.L., K.E. Munroe. 2017. The professional interview: preparing for success. pp. 158–165, In:Henke, S.E., Krausman, P.R. (eds.)
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Minor, R., J.L. Koprowski. 2016. Seed removal increases through scramble competition with an invasive species. PLoS ONE 10(12): e0143927.
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Koprowski, J.L. 1989. Burrow distribution of thirteen-lined ground squirrels in relation to tree canopies. Prairie Naturalist 21:185-188.
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While a doctoral and postdoctoral student at the University of Kansas, John began teaching night courses in introductory biology at
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Outstanding Mentor of Graduate and Professional Students, University of Arizona Graduate & Professional Student Council (2011)
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Paudel, S., P. Pal, M.V. Cove, S.R. Jnawali, G. Abel, J.L. Koprowski, R. Ranabhat. 2015. The endangered Ganges River dolphin
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Western National Parks Association's Emil W. Haury Award for Outstanding Achievement in Science in the National Park (1999)
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to earn his M.A. in zoology in August, 1985 with advisor Dr. Willard D. Klimstra. His Ph.D. was completed in May, 1991 at
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Koprowski, J.L. 1987. Nocturnal activity of the woodchuck in an urban Ohio park. Canadian Field-Naturalist 101:606-607.
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Kellam, J.O., D.K. Jansen, A.T. Johnson, R.W. Arwood, M.J. Merrick, J.L. Koprowski. 2016. Big Cypress fox squirrel (
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Nowhere to run and nowhere to hide: response of endemic Mt. Graham red squirrels to catastrophic forest damage
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Jo, YS, Baccus, JT, Koprowski, JL. 2018. Mammals of Korea, National Institute of Biological Sciences, 573 pp.
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Paudel, S., Koprowski, J.L., 2020. Factors affecting the persistence of endangered Ganges River dolphins (
699:) ecology and habitat use in cypress dome swamp-pine forest mosaic.  Journal of Mammalogy 97:200-210. 542: 676:
in Nepal: abundance, habitat    and conservation threats. Endangered Species Research 29:59-68.
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The use of GIS and modelling approaches in squirrel population management and conservation: a review
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Outstanding Scholarly Publication, School of Natural Resources, University of Arizona (2007, 2002)
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Dorothy Haglund Outstanding Dissertation Award, The Graduate School, University of Kansas (1991)
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Report on Research: A Journal of Creative Activity from the University of Arizona 20(2): 28–31.
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Koprowski, J.L., W.S. Fairbanks. 2013. Wildlife behavior. In: Krausman, P.R. Cain, J. (ed.)
906: 669:) use seasonally flooded forests in the Peruvian Amazon? Journal of Mammalogy 96:1295-1304. 573:
Natal philopatry, communal nesting, and kinship in fox squirrels and eastern gray squirrels
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where he graduated with honors in Biology studying with advisor Dr. Kenneth B. Armitage.
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Brady, M.J., J.L. Koprowski, R.N. Gwinn, Y.S. Jo, K. Young. 2017. Eastern fox squirrel (
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Global hotspots, centers of diversity, and conservation of the tree and flying squirrels
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Conflict between the sexes: a review of social and mating systems of the tree squirrels
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Koprowski, J.L. 1993. Alternative reproductive tactics in male eastern gray squirrels:
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Emil W. Haury Award for Outstanding Achievement in Science in the National Park (1999)
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Dean, Haub School of Environment & Natural Resources at University of Wyoming
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Ramos‐Lara, N., J.L. Koprowski. 2015. Spacing behavior of a non‐larder‐hoarding
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Outstanding Faculty Member, School of Natural Resources (2017, 2013, 2005, 2003)
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International Wildlife Management: Conservation Challenges in a Changing World.
735:, Linnaeus 1758) introduction to the Sonoran Desert.  Mammalia 81:221-223. 376:"Haub School of Environment and Natural Resources | University of Wyoming" 215: 191: 171: 544:
International Wildlife Management: Conservation Challenges in a Changing World
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Aldo Leopold Memorial Award/Aldo Leopold Medal, The Wildlife Society (2022)
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Palmer, R.R., J.L. Koprowski. 2015. How do Neotropical pygmy squirrels (
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The response of tree squirrels to fragmentation: a review and synthesis
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The Last Refuge of the Mt. Graham Red Squirrel: Ecology of Endangerment
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and also advanced courses in vertebrate biology and evolution at the
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Graduate Teaching Assistant of the Year, University of Kansas (1990)
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Severely insect-damaged forest: A temporary trap for red squirrels?
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Fellows of the American Association for the Advancement of Science
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Ecology, Conservation & Management of Grey Squirrels in Europe
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in Lakewood, Ohio, in June 1979 and earned his B.S. in zoology at
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Fellow, American Association for the Advancement of Science (2018)
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News release, University of Newcastle upon Tyne, April 25, 2006.
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Extensive expertise in the ecology and conservation of wildlife
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Thorington, RW, Koprowski, JL, Steele, MA, Whatton, J. 2012.
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Handling tree squirrels with an efficient and safe restraint
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Wildlife Management: Contemporary Principles and Practices
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Ohio State University College of Arts and Sciences alumni
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Aldo Leopold Memorial Award, The Wildlife Society (2022)
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Mount Graham Biological Program at University of Arizona
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IgapĂł (Black-water flooded forests) of the Amazon Basin
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The Habits of Highly Effective Squirrel Researchers.
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Removal of copulatory plugs by female tree squirrels
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IUCN North American Coordinator-Small Mammals (2010)
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American Association for the Advancement of Science
811:Haub School of Environment and Natural Resources, 208:Haub School of Environment & Natural Resources 852:Computers May Help Save Mount Graham Red Squirrel 873: 834:, Columbus Dispatch, published February 2, 2008. 592:Koprowski, J.L., M.I. Alanen, A.M. Lynch. 2005. 776:Johns Hopkins University Press, Baltimore, USA. 127:North American Coordinator-Small Mammals (2010) 947:Southern Illinois University Carbondale alumni 819:Nut? What Nut? The Squirrel Outwits to Survive 617:Lurz, P.W.W., J.L. Koprowski, D.J.Wood. 2008. 551: 839:Safely Studying Endangered Animals in Arizona 390:"The University of Wyoming, Laramie, Wyoming" 845:Can Squirrels Spy for People? Iran Thinks So 841:UANews, University of Arizona, June 4, 2007. 826:Interview for Ohio State Alumni Association. 749:, Johns Hopkins University Press, Baltimore. 704:Handbook of Mammals of the World, Volume 6, 628:Forest Ecology and Management 257: 464–470. 520:Sanderson, HR, Koprowski, JL. (eds). 2009. 407: 405: 29: 868:, The Associated Press, updated 7/8/2004. 866:Endangered squirrel faces wildfire threat 547:, Johns Hopkins University Press, 248 pp. 535:, Johns Hopkins University Press, 472 pp. 882:Fellows of the Linnean Society of London 589:. Wildlife Society Bulletin, 30:101-103. 517:, Smithsonian Institution Press. 224 pp. 412:Steele, M.A., and Koprowski, J.L. 2001. 120:Fellow, Linnean Society of London (2019) 832:Buckeye Chuck says spring is in the air 596:. Biological Conservation, 127:491-498. 402: 874: 783:). Ecology and Evolution 10:3138-3148. 740:El Oso Andino en el Macizo de Chingaza 645:). Conservation Genetics 14:1233-1241. 624:Zugmeyer, C.A., J.L. Koprowski. 2009. 575:. Journal of Mammalogy, 77: 1006–1016. 428:"Kansas City Kansas Community College" 417:Smithsonian Institution Press. 224 pp. 769:.  Springer, Bern, Switzerland. 610:Koprowski, J.L., N. Rajamani. 2008. 252:Kansas City Kansas Community College 130:Western National Parks Association’s 942:Southern Illinois University alumni 643:Tamiasciurus hudsonicus grahamensis 122:Fellow, The Wildlife Society (2016) 13: 821:NY Times, published: July 5, 2010. 568:. Journal of Mammalogy 73:572-576. 541:Koprowski, JL, Krausman PR. 2019. 256:University of Missouri–Kansas City 14: 958: 800: 655:doi:10.1371/journal.pone.01111940 635:, Johns Hopkins University Press. 603:. Animal Conservation 8: 369–376. 511:Steele, MA, Koprowski, JL. 2001. 116:Fellow, The Explorers Club (2023) 897:21st-century American biologists 747:Becoming a Wildlife Professional 234:in June, 1983. He then attended 561:. Behavioral Ecology 4:165-171. 781:Platanista gangetica gangetica 683:doi:10.4404/hystrix-27.2-11688 674:Platanista gangetica gangetica 621:. Current Science 95: 918–922. 614:. Current Science 95: 851–856. 559:"Making the best of a bad job" 526:, University of Arizona Press. 480: 471: 462: 448: 434: 420: 382: 368: 226:John Koprowski graduated from 1: 912:University of Wyoming faculty 902:University of Arizona faculty 887:Fellows of the Explorers Club 857:Hutchinson, Jan McCoy (2004) 514:North American Tree Squirrels 414:North American Tree Squirrels 353: 922:Ohio State University people 807:John L. Koprowski's Web Page 362: 236:Southern Illinois University 221: 84:Southern Illinois University 7: 937:University of Kansas alumni 932:University of Kansas people 864:Rotstein, Arthur H. (2004) 837:Littlefield, Joanne (2007) 813:Professor John L. Koprowski 552:Representative Publications 10: 963: 706:Lynx Ediciones, Barcelona. 344: 292:Linnean Society of London 277: 268:Mount Graham Red Squirrel 245: 197: 143: 136: 110: 93: 75: 67: 59: 37: 28: 21: 830:Riepenhoff, Jill (2008) 796:10.3389/fevo.2020.00174. 505: 71:United States of America 817:Angier, Natalie (2010) 697:Sciurus niger avicennia 442:"Willamette University" 850:Jordan, Claire (2006) 585:Koprowski, J.L. 2002. 578:Koprowski, J.L. 1998. 571:Koprowski, J.L. 1996. 564:Koprowski, J.L. 1992. 532:Squirrels of the World 290:Elected Fellow of the 283:Elected Fellow of the 206:, Dean and Professor, 599:Koprowski, JL. 2005. 264:University of Arizona 260:Willamette University 232:Ohio State University 212:University of Wyoming 80:Ohio State University 299:The Wildlife Society 240:University of Kansas 228:Lakewood High School 186:and conservation of 148:Conservation biology 88:University of Kansas 667:Sciurillus pusillus 824:Battenberg, Erik. 306:the Explorers Club 304:Elected Fellow of 297:Elected Fellow of 180:endangered species 160:population ecology 23:John Lad Koprowski 16:American biologist 204:John L. Koprowski 201: 200: 168:forest management 138:Scientific career 48:29 September 1961 954: 500: 499: 494:. 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Index


Ohio State University
Southern Illinois University
University of Kansas

Mount Graham Biological Program at University of Arizona
IUCN
Western National Parks Association’s
Conservation biology
vertebrates
behavioral
population ecology
social behavior
forest management
urban wildlife
threatened
endangered species
ecology
squirrels
climate change
Haub School of Environment & Natural Resources
University of Wyoming
Lakewood, Ohio
Lakewood High School
Ohio State University
Southern Illinois University
University of Kansas
Kansas City Kansas Community College
University of Missouri–Kansas City
Willamette University

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