124:
102:
428:
279:) have peculiar orange structures along the sides of the abdomen, which may be everted and saclike or withdrawn into the body and inconspicuous. Some melyrids have the two basal antennomeres greatly enlarged. Most adults and larvae are predaceous, but many are common on flowers. The most common North American species belong to the genus
431:
430:
435:
434:
429:
436:
433:
416:
Various authorities have, at times, treated each of the presently-recognized subfamilies as families, and a few tribes have been accorded family status, as well (e.g., "Attalomimidae"). The family
432:
408:
The family
Melyridae contains over 100 genera worldwide, with ~520 species in 48 genera in North America, and 16 genera in Europe; the largest diversity is in tropical rainforests.
859:
312:
1176:
318:
306:
729:
579:
275:
Most are elongate-oval, soft-bodied beetles 10 mm long or less. Many are brightly patterned in black and brown, yellow, or red. Some melyrids (
324:
1139:
1099:
949:
529:
3471:
909:
549:
1166:
1156:
1119:
1009:
969:
919:
849:
679:
629:
519:
388:
3510:
1186:
959:
929:
619:
589:
360:
1277:"A previously missing link in the evolution of dasytine softâwinged flower beetles from Cretaceous Charentese amber (Coleoptera, Melyridae)"
1129:
1069:
1019:
819:
799:
789:
659:
499:
366:
1059:
879:
739:
489:
464:
769:
759:
649:
569:
3256:
1049:
989:
939:
3445:
1221:
Melyrid beetles (Choresine): A putative source for the batrachotoxin alkaloids found in poison-dart frogs and toxic passerine birds
1321:
Gimmel M.L., Bocakova M., Gunter N.L., Leschen R.A.B. (2019) Comprehensive phylogeny of the
Cleroidea (Coleoptera: Cucujiformia).
3484:
1209:
Triplehorn, C. and
Johnson, N.: "Borror and DeLong's Introduction to the Study of Insects, 7th Ed.", page 429. Brooks/Cole, 2005.
1337:
3489:
1369:
3582:
3515:
3424:
3437:
3497:
1218:
John P. Dumbacher, Avit Wako, Scott R. Derrickson, Allan
Samuelson, Thomas F. Spande, John W. Daly (2004):
3615:
3354:
3331:
1275:
Tihelka, Erik; Huang, Diying; Perrichot, Vincent; Cai, Chenyang (2021-04-11). Labandeira, Conrad (ed.).
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8:
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118:
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699:
333:
3093:
2872:
2867:
2790:
2548:
1564:
1288:
1247:
719:
397:
342:
frogs in South
America obtain batrachotoxins from related genera of the Melyridae (
110:
3323:
3385:
3336:
2782:
2735:
2580:
2564:
2240:
2000:
1969:
1884:
1802:
1693:
1661:
371:
346:
does not occur there) has not been tested due to the difficulty of field-work in
3502:
1220:
3432:
3419:
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2255:
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2047:
1992:
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999:
329:
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2658:
2632:
2607:
2503:
2490:
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2347:
2250:
2245:
2126:
2060:
2031:
2015:
1979:
1949:
1902:
1856:
1823:
1739:
1731:
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1540:
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1300:
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417:
59:
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2211:
2191:
2152:
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2010:
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1932:
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1252:
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609:
195:
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2368:
2318:
2310:
2268:
2232:
2203:
2175:
2170:
2086:
2081:
2073:
2023:
1959:
1954:
1868:
1833:
1818:
1810:
1726:
1718:
1535:
709:
599:
458:
338:
284:
276:
34:
101:
3157:
3118:
3110:
3052:
2915:
2877:
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2822:
2642:
2590:
2474:
2405:
2276:
2260:
2216:
2157:
2121:
2101:
2065:
1974:
1964:
1838:
1572:
1556:
1109:
539:
509:
421:
393:
79:
44:
3372:
1342:
1292:
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3406:
3222:
3214:
3206:
3193:
3149:
2948:
2923:
2851:
2679:
2650:
2334:
2165:
2139:
2091:
2005:
1876:
1848:
1681:
1656:
1651:
1632:
1597:
1486:
1469:
1461:
1410:
1398:
1039:
979:
639:
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300:
264:
205:
185:
155:
135:
84:
3450:
3265:
3411:
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3126:
3076:
3020:
2979:
2928:
2585:
2516:
2397:
2109:
1915:
1889:
1747:
1580:
1527:
1506:
1416:
1079:
899:
889:
689:
347:
74:
69:
54:
49:
39:
2843:
2835:
2687:
2421:
2342:
2326:
1146:
1089:
1029:
839:
809:
779:
559:
292:
89:
64:
1224:. The National Academy of Sciences. Vol. 101 no. 45, 15857-15860
2434:
2294:
1927:
1404:
1392:
1379:
869:
332:, which may account for the toxicity of some birds such as the
260:
175:
165:
145:
3559:
3398:
1236:"New cleroid beetles from the Middle-Late Jurassic of China"
1750:(furniture beetles, death watch beetles, spider beetles)
1274:
328:, the latter as yet unnamed) have been found to contain
336:
and hooded pitohui which eat them. The hypothesis that
3602:
2627:(jewel beetles, or metallic wood-boring beetles)
291:is reddish, with two bluish black spots on each
382:in Inner Mongolia, China. The oldest member of
2350:(true weevils, bark beetles, ambrosia beetles)
1517:
1377:
1363:
2593:(ironclad beetles, cylindrical bark beetles)
1233:
1370:
1356:
100:
3257:List of subgroups of the order Coleoptera
2337:(straight snout weevils, New York weevil)
1436:
1251:
2764:(cicada beetle, cicada parasite beetles)
1212:
426:
420:was removed from Melyridae in 1995, and
353:
3603:
2854:(false fireflies, long-lipped beetles)
1679:
1630:
3270:
3269:
1678:
1629:
1504:
1503:
1435:
1351:
1234:Kolibac, Jiri; Huang, Diying (2019).
3565:396787A7-5C70-1B78-FF1D-F9DFFBACF3EC
3526:fa9f206f-e16d-4ad8-ad7e-bd6ad42c7fee
3438:e87bb258-1946-462d-ba23-c9f04a4c51a0
2880:(false metallic wood-boring beetles)
358:The oldest fossil of the family are
13:
471:
14:
3627:
1331:
2491:Perimylopidae, or Promecheilidae
2076:(minute brown scavenger beetles)
392:from the early Late Cretaceous (
122:
32:
2706:(variegated mud-loving beetles)
403:
1315:
1268:
1227:
1203:
411:
270:
1:
2535:(narrow-waisted bark beetles)
2227:(parasitic flat bark beetles)
2178:(short-winged flower beetles)
2034:(lady beetles, or God's cows)
1771:(tooth-necked fungus beetles)
1240:Acta Palaeontologica Polonica
3047:(sand-loving scarab beetles)
2437:(minute tree-fungus beetles)
2263:(silvanid flat bark beetles)
298:Four New Guinean species of
7:
3196:(primitive carrion beetles)
3152:(minute mud-loving beetles)
3015:(bumble bee scarab beetles)
2785:(soft-bodied plant beetles)
2756:(soft bodied plant beetles)
1559:(predaceous diving beetles)
10:
3632:
3031:(scavenger scarab beetles)
3023:(enigmatic scarab beetles)
2196:Lamingtonium binnaberrense
1910:(soft-wing flower beetles)
844:Laporte de Castelnau, 1836
674:Fischer von Waldheim, 1844
455:Laporte de Castelnau, 1840
257:soft-winged flower beetles
3278:
3254:
3184:
3170:(water scavenger beetles)
3135:
3101:
3092:
2970:
2961:
2914:
2903:Rhinorhipus tamborinensis
2889:
2773:
2744:
2682:(long-toed water beetles)
2641:
2615:
2606:
2575:(polypore fungus beetles)
2477:(palm and flower beetles)
2461:(tumbling flower beetles)
2408:(ant-like flower beetles)
2388:
2367:
2293:
2186:(lined flat bark beetles)
2160:(pleasing fungus beetles)
2100:
2055:(handsome fungus beetles)
1991:
1847:
1793:
1784:
1759:
1701:
1692:
1688:
1674:
1639:
1625:
1600:(burrowing water beetles)
1513:
1499:
1445:
1431:
1387:
235:
228:
119:Scientific classification
117:
108:
99:
23:
3217:(feather-winged beetles)
2551:(false longhorn beetles)
2498:(jugular-horned beetles)
2445:(false darkling beetles)
2416:(cryptic fungus beetles)
2235:(shining flower beetles)
1575:(crawling water beetles)
1474:Sikhotealinia zhiltzovae
1197:
2698:(forest stream beetles)
2559:(synchroa bark beetles)
2519:(dead log bark beetles)
2511:(fire-coloured beetles)
2400:(ant-like leaf beetles)
2134:(silken fungus beetles)
2068:(well polished beetles)
2042:(minute fungus beetles)
1861:Acanthocnemus nigricans
1281:Papers in Palaeontology
1134:Mulsant & Rey, 1867
664:Mulsant & Rey, 1867
316:and the two infrequent
3209:(round fungus beetles)
2543:(false flower beetles)
2527:(wedge-shaped beetles)
2485:(false blister beetle)
2469:(hairy fungus beetles)
2424:(conifer bark beetles)
2321:(leaf-rolling weevils)
1982:(bark-gnawing beetles)
1742:(wounded-tree beetles)
1608:(false ground beetles)
1530:(trout-stream beetles)
1253:10.4202/app.00550.2018
1004:Guérin-Méneville, 1830
654:Jacquelin Du Val, 1859
448:
30:Middle JurassicâRecent
3547:Paleobiology Database
3121:(false clown beetles)
2943:(plate-thigh beetles)
2825:(false click beetles)
2730:(water-penny beetles)
2379:(ship-timber beetles)
2358:(pine flower weevils)
2206:(root-eating beetles)
2026:(minute bark beetles)
1937:Phloiophilus edwardsi
1464:(reticulated beetles)
1323:Systematic Entomology
439:
3433:Fauna Europaea (new)
2999:(false stag beetles)
2984:Belohina inexpectata
2846:(net-winged beetles)
2809:(rare click beetles)
2722:(travertine beetles)
2714:(minute mud beetles)
2279:(dry-fungus beetles)
2114:Agapytho foveicollis
1871:(false skin beetles)
1734:(Jacobson's beetles)
1454:Crowsoniella relicta
354:Evolutionary history
310:, the less abundant
2997:Diphyllostomatidae
2329:(primitive weevils)
2142:(flat bark beetles)
1892:(checkered beetles)
1879:(fruitworm beetles)
1567:(whirligig beetles)
304:(the more abundant
263:of the superfamily
3616:Polyphaga families
2862:(glowworm beetles)
2567:(darkling beetles)
2414:Archeocrypticidae
2271:(palmetto beetles)
2018:(dry bark beetles)
1805:(longhorn beetles)
1114:Redtenbacher, 1858
1034:Motschoulsky, 1859
984:Kiesenwetter, 1863
854:Kiesenwetter, 1867
744:Redtenbacher, 1849
624:Motschoulsky, 1859
524:Redtenbacher, 1850
449:
289:C. quadrimaculatus
259:) are a family of
3598:
3597:
3534:Open Tree of Life
3272:Taxon identifiers
3263:
3262:
3250:
3249:
3246:
3245:
3242:
3241:
3234:
3226:
3225:(carrion beetles)
3218:
3210:
3197:
3171:
3153:
3122:
3114:
3088:
3087:
3080:
3072:
3064:
3056:
3048:
3040:
3032:
3024:
3016:
3008:
3000:
2987:
2957:
2956:
2944:
2936:
2906:
2881:
2863:
2855:
2847:
2839:
2826:
2818:
2810:
2802:
2801:(soldier beetles)
2794:
2791:Brachypsectridae
2786:
2765:
2757:
2731:
2723:
2715:
2707:
2699:
2691:
2683:
2670:
2662:
2654:
2628:
2602:
2601:
2594:
2576:
2568:
2560:
2552:
2549:Stenotrachelidae
2544:
2536:
2528:
2520:
2512:
2499:
2486:
2478:
2470:
2462:
2454:
2453:(blister beetles)
2446:
2438:
2425:
2417:
2409:
2401:
2380:
2359:
2351:
2338:
2330:
2322:
2314:
2304:
2280:
2272:
2264:
2236:
2228:
2220:
2207:
2199:
2187:
2179:
2161:
2143:
2135:
2117:
2077:
2069:
2056:
2043:
2035:
2027:
2019:
2001:Akalyptoischiidae
1983:
1940:
1923:
1911:
1893:
1880:
1872:
1864:
1814:
1806:
1780:
1779:
1772:
1751:
1743:
1735:
1722:
1714:
1670:
1669:
1647:
1621:
1620:
1617:
1616:
1609:
1601:
1593:
1576:
1568:
1560:
1552:
1544:
1531:
1495:
1494:
1477:
1465:
1457:
1293:10.1002/spp2.1360
1192:
1182:
1172:
1171:Motschulsky, 1859
1162:
1152:
1135:
1125:
1115:
1105:
1095:
1085:
1075:
1065:
1055:
1045:
1035:
1025:
1015:
1005:
995:
985:
975:
965:
955:
945:
935:
925:
915:
905:
895:
885:
875:
865:
855:
845:
835:
825:
815:
805:
795:
785:
775:
774:Motschulsky, 1854
765:
755:
745:
735:
725:
715:
714:Motschulsky, 1854
705:
695:
685:
675:
665:
655:
645:
635:
625:
615:
605:
604:Motschulsky, 1854
595:
585:
575:
565:
555:
545:
535:
534:Tshernyshev, 2009
525:
515:
505:
495:
485:
437:
334:blue-capped ifrit
250:
249:
224:
16:Family of beetles
3623:
3591:
3590:
3578:
3577:
3568:
3567:
3555:
3554:
3542:
3541:
3529:
3528:
3519:
3518:
3506:
3505:
3503:NHMSYS0001718977
3493:
3492:
3480:
3479:
3467:
3466:
3454:
3453:
3441:
3440:
3428:
3427:
3415:
3414:
3402:
3401:
3389:
3388:
3376:
3375:
3363:
3362:
3350:
3349:
3340:
3339:
3327:
3326:
3314:
3313:
3312:
3299:
3298:
3297:
3267:
3266:
3232:
3224:
3216:
3208:
3195:
3169:
3151:
3120:
3112:
3099:
3098:
3094:Staphyliniformia
3078:
3070:
3062:
3054:
3046:
3038:
3030:
3022:
3014:
3006:
2998:
2981:
2968:
2967:
2942:
2933:Declinia relicta
2930:
2900:
2879:
2873:Sinopyrophoridae
2868:Rhagophthalmidae
2861:
2853:
2845:
2837:
2824:
2816:
2808:
2800:
2792:
2784:
2783:Artematopodidae
2763:
2755:
2729:
2721:
2713:
2705:
2697:
2690:(riffle beetles)
2689:
2681:
2669:(turtle beetles)
2668:
2660:
2652:
2626:
2613:
2612:
2592:
2574:
2566:
2558:
2550:
2542:
2534:
2526:
2518:
2510:
2497:
2484:
2476:
2468:
2460:
2452:
2444:
2436:
2423:
2415:
2407:
2399:
2378:
2357:
2349:
2336:
2328:
2320:
2313:(fungus weevils)
2312:
2303:
2302:
2298:
2285:Tasmosalpingidae
2278:
2270:
2262:
2234:
2226:
2218:
2205:
2193:
2185:
2177:
2159:
2141:
2133:
2111:
2075:
2067:
2054:
2041:
2033:
2025:
2017:
1981:
1934:
1917:
1909:
1891:
1878:
1870:
1858:
1857:Acanthocnemidae
1812:
1804:
1791:
1790:
1770:
1749:
1741:
1733:
1720:
1712:
1699:
1698:
1690:
1689:
1676:
1675:
1645:
1627:
1626:
1607:
1599:
1587:
1574:
1566:
1558:
1550:
1543:(ground beetles)
1542:
1529:
1515:
1514:
1501:
1500:
1471:
1463:
1451:
1450:Crowsoniellidae
1433:
1432:
1372:
1365:
1358:
1349:
1348:
1326:
1319:
1313:
1312:
1287:(4): 1753â1764.
1272:
1266:
1265:
1255:
1231:
1225:
1216:
1210:
1207:
1190:
1180:
1170:
1160:
1150:
1133:
1123:
1113:
1103:
1093:
1083:
1073:
1063:
1053:
1043:
1033:
1023:
1013:
1003:
993:
983:
973:
963:
953:
943:
933:
923:
913:
903:
893:
883:
873:
863:
853:
843:
833:
823:
813:
803:
793:
783:
773:
763:
753:
743:
733:
723:
713:
703:
693:
683:
673:
663:
653:
643:
633:
623:
613:
603:
593:
583:
573:
563:
553:
543:
533:
523:
513:
503:
493:
484:Waterhouse, 1877
483:
441:Dasytes plumbeus
438:
398:Charentese amber
219:
127:
126:
111:Anthocomus rufus
104:
94:
31:
27:Temporal range:
21:
20:
3631:
3630:
3626:
3625:
3624:
3622:
3621:
3620:
3601:
3600:
3599:
3594:
3586:
3581:
3573:
3571:
3563:
3558:
3550:
3545:
3537:
3532:
3524:
3522:
3514:
3509:
3501:
3496:
3488:
3483:
3475:
3470:
3462:
3457:
3449:
3444:
3436:
3431:
3423:
3418:
3410:
3405:
3397:
3392:
3384:
3379:
3371:
3366:
3358:
3353:
3345:
3343:
3335:
3330:
3322:
3317:
3308:
3307:
3302:
3293:
3292:
3287:
3274:
3264:
3259:
3238:
3180:
3131:
3113:(clown beetles)
3084:
3055:(betsy beetles)
2963:Scarabaeiformia
2953:
2910:
2885:
2817:(click beetles)
2793:(Texas beetles)
2769:
2740:
2736:Ptilodactylidae
2661:(cedar beetles)
2637:
2598:
2581:Trictenotomidae
2467:Mycetophagidae
2384:
2363:
2300:
2299:
2297:
2289:
2241:Phloeostichidae
2192:Lamingtoniidae
2184:Laemophloeidae
2132:Cryptophagidae
2096:
1987:
1970:Thanerocleridae
1933:Phloiophilidae
1920:Metaxina ornata
1885:Chaetosomatidae
1843:
1776:
1755:
1713:(auger beetles)
1694:Bostrichiformia
1684:
1666:
1662:Torridincolidae
1646:(skiff beetles)
1644:Hydroscaphidae
1635:
1613:
1606:Trachypachidae
1551:(tiger beetles)
1509:
1491:
1441:
1427:
1383:
1376:
1334:
1329:
1320:
1316:
1273:
1269:
1232:
1228:
1217:
1213:
1208:
1204:
1200:
1195:
1054:Fabricius, 1775
1044:Fabricius, 1775
1024:Blaisdell, 1924
964:Blaisdell, 1924
694:Blanchard, 1845
474:
472:Selected genera
427:
414:
406:
372:Middle Jurassic
356:
273:
218:
121:
95:
93:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
37:
29:
28:
25:
17:
12:
11:
5:
3629:
3619:
3618:
3613:
3596:
3595:
3593:
3592:
3579:
3569:
3556:
3543:
3530:
3520:
3507:
3494:
3481:
3468:
3455:
3442:
3429:
3420:Fauna Europaea
3416:
3403:
3390:
3377:
3364:
3351:
3341:
3328:
3315:
3300:
3284:
3282:
3276:
3275:
3261:
3260:
3255:
3252:
3251:
3248:
3247:
3244:
3243:
3240:
3239:
3237:
3236:
3233:(rove beetles)
3231:Staphylinidae
3228:
3220:
3212:
3204:
3199:
3190:
3188:
3186:Staphylinoidea
3182:
3181:
3179:
3178:
3173:
3168:Hydrophilidae
3165:
3160:
3155:
3147:
3141:
3139:
3137:Hydrophiloidea
3133:
3132:
3130:
3129:
3124:
3116:
3107:
3105:
3096:
3090:
3089:
3086:
3085:
3083:
3082:
3079:(hide beetles)
3074:
3066:
3063:(rain beetles)
3058:
3050:
3042:
3039:(stag beetles)
3034:
3026:
3018:
3010:
3002:
2994:
2992:Bolboceratidae
2989:
2976:
2974:
2965:
2959:
2958:
2955:
2954:
2952:
2951:
2946:
2938:
2926:
2920:
2918:
2912:
2911:
2909:
2908:
2899:Rhinorhipidae
2895:
2893:
2891:Rhinorhipoidea
2887:
2886:
2884:
2883:
2875:
2870:
2865:
2857:
2849:
2841:
2833:
2828:
2820:
2812:
2804:
2796:
2788:
2779:
2777:
2771:
2770:
2768:
2767:
2759:
2750:
2748:
2742:
2741:
2739:
2738:
2733:
2725:
2717:
2709:
2704:Heteroceridae
2701:
2693:
2685:
2677:
2672:
2667:Chelonariidae
2664:
2659:Callirhipidae
2656:
2653:(pill beetles)
2647:
2645:
2639:
2638:
2636:
2635:
2630:
2621:
2619:
2610:
2604:
2603:
2600:
2599:
2597:
2596:
2588:
2583:
2578:
2570:
2565:Tenebrionidae
2562:
2554:
2546:
2538:
2530:
2522:
2514:
2506:
2501:
2493:
2488:
2480:
2472:
2464:
2456:
2448:
2440:
2432:
2427:
2419:
2411:
2403:
2394:
2392:
2390:Tenebrionoidea
2386:
2385:
2383:
2382:
2373:
2371:
2365:
2364:
2362:
2361:
2353:
2348:Curculionidae
2345:
2340:
2332:
2324:
2316:
2307:
2305:
2295:Curculionoidea
2291:
2290:
2288:
2287:
2282:
2274:
2266:
2258:
2256:Protocucujidae
2253:
2248:
2243:
2238:
2230:
2222:
2214:
2209:
2201:
2189:
2181:
2173:
2168:
2163:
2155:
2150:
2148:Cybocephalidae
2145:
2137:
2129:
2124:
2119:
2106:
2104:
2098:
2097:
2095:
2094:
2089:
2084:
2079:
2071:
2063:
2058:
2050:
2048:Discolomatidae
2045:
2037:
2032:Coccinellidae
2029:
2021:
2016:Bothrideridae
2013:
2008:
2003:
1997:
1995:
1993:Coccinelloidea
1989:
1988:
1986:
1985:
1980:Trogossitidae
1977:
1972:
1967:
1962:
1957:
1952:
1947:
1942:
1930:
1925:
1913:
1905:
1900:
1898:Lophocateridae
1895:
1887:
1882:
1874:
1866:
1853:
1851:
1845:
1844:
1842:
1841:
1836:
1831:
1826:
1821:
1816:
1813:(leaf beetles)
1811:Chrysomelidae
1808:
1799:
1797:
1795:Chrysomeloidea
1788:
1782:
1781:
1778:
1777:
1775:
1774:
1765:
1763:
1757:
1756:
1754:
1753:
1745:
1740:Nosodendridae
1737:
1732:Jacobsoniidae
1729:
1724:
1721:(skin beetles)
1716:
1707:
1705:
1696:
1686:
1685:
1672:
1671:
1668:
1667:
1665:
1664:
1659:
1654:
1649:
1640:
1637:
1636:
1623:
1622:
1619:
1618:
1615:
1614:
1612:
1611:
1603:
1595:
1590:Meru phyllisae
1583:
1578:
1570:
1562:
1554:
1546:
1538:
1533:
1524:
1522:
1511:
1510:
1497:
1496:
1493:
1492:
1490:
1489:
1484:
1482:Micromalthidae
1479:
1467:
1459:
1446:
1443:
1442:
1429:
1428:
1426:
1425:
1419:
1413:
1407:
1401:
1395:
1388:
1385:
1384:
1375:
1374:
1367:
1360:
1352:
1346:
1345:
1343:Coleoptera.org
1340:
1338:Fauna Europaea
1333:
1332:External links
1330:
1328:
1327:
1314:
1267:
1226:
1211:
1201:
1199:
1196:
1194:
1193:
1183:
1173:
1163:
1153:
1143:
1140:Spinapalochrus
1136:
1126:
1116:
1106:
1100:Pagurodactylus
1096:
1094:Marshall, 1951
1086:
1076:
1066:
1056:
1046:
1036:
1026:
1016:
1006:
996:
986:
976:
966:
956:
950:Haplomalachius
946:
936:
926:
916:
906:
904:Stephens, 1830
896:
886:
876:
874:Erichson, 1840
866:
864:Stephens, 1830
856:
846:
836:
834:Champion, 1921
826:
824:Schilsky, 1896
816:
806:
796:
786:
776:
766:
756:
746:
736:
726:
724:Erichson, 1840
716:
706:
704:Erichson, 1840
696:
686:
676:
666:
656:
646:
644:Erichson, 1840
636:
634:Champion, 1923
626:
616:
606:
596:
586:
584:Erichson, 1840
576:
566:
556:
546:
544:Erichson, 1840
536:
530:Anthomalachius
526:
516:
514:Erichson, 1840
506:
496:
494:Westwood, 1839
486:
475:
473:
470:
469:
468:
462:
456:
413:
410:
405:
402:
370:from the late
355:
352:
330:batrachotoxins
272:
269:
255:(common name:
248:
247:
246:
245:
242:
239:
233:
232:
226:
225:
213:
209:
208:
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199:
198:
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105:
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96:
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73:
68:
63:
58:
53:
48:
43:
38:
33:
26:
15:
9:
6:
4:
3:
2:
3628:
3617:
3614:
3612:
3609:
3608:
3606:
3589:
3584:
3580:
3576:
3570:
3566:
3561:
3557:
3553:
3548:
3544:
3540:
3535:
3531:
3527:
3521:
3517:
3512:
3508:
3504:
3499:
3495:
3491:
3486:
3482:
3478:
3473:
3469:
3465:
3460:
3456:
3452:
3447:
3443:
3439:
3434:
3430:
3426:
3421:
3417:
3413:
3408:
3404:
3400:
3395:
3391:
3387:
3382:
3378:
3374:
3369:
3365:
3361:
3356:
3352:
3348:
3342:
3338:
3333:
3329:
3325:
3320:
3316:
3311:
3305:
3301:
3296:
3290:
3286:
3285:
3283:
3281:
3277:
3273:
3268:
3258:
3253:
3235:
3229:
3227:
3221:
3219:
3213:
3211:
3205:
3203:
3200:
3198:
3192:
3191:
3189:
3187:
3183:
3177:
3174:
3172:
3166:
3164:
3161:
3159:
3156:
3154:
3148:
3146:
3143:
3142:
3140:
3138:
3134:
3128:
3125:
3123:
3119:Sphaeritidae
3117:
3115:
3109:
3108:
3106:
3104:
3100:
3097:
3095:
3091:
3081:
3075:
3073:
3069:Scarabaeidae
3067:
3065:
3059:
3057:
3051:
3049:
3043:
3041:
3035:
3033:
3027:
3025:
3019:
3017:
3011:
3009:
3007:(dor beetles)
3003:
3001:
2995:
2993:
2990:
2988:
2985:
2978:
2977:
2975:
2973:
2972:Scarabaeoidea
2969:
2966:
2964:
2960:
2950:
2947:
2945:
2939:
2937:
2934:
2927:
2925:
2922:
2921:
2919:
2917:
2913:
2907:
2904:
2897:
2896:
2894:
2892:
2888:
2882:
2876:
2874:
2871:
2869:
2866:
2864:
2858:
2856:
2850:
2848:
2842:
2840:
2834:
2832:
2829:
2827:
2821:
2819:
2813:
2811:
2807:Cerophytidae
2805:
2803:
2797:
2795:
2789:
2787:
2781:
2780:
2778:
2776:
2772:
2766:
2762:Rhipiceridae
2760:
2758:
2752:
2751:
2749:
2747:
2743:
2737:
2734:
2732:
2726:
2724:
2718:
2716:
2710:
2708:
2702:
2700:
2696:Eulichadidae
2694:
2692:
2686:
2684:
2678:
2676:
2675:Cneoglossidae
2673:
2671:
2665:
2663:
2657:
2655:
2649:
2648:
2646:
2644:
2640:
2634:
2633:Schizopodidae
2631:
2629:
2623:
2622:
2620:
2618:
2614:
2611:
2609:
2608:Elateriformia
2605:
2595:
2589:
2587:
2584:
2582:
2579:
2577:
2573:Tetratomidae
2571:
2569:
2563:
2561:
2555:
2553:
2547:
2545:
2539:
2537:
2531:
2529:
2525:Ripiphoridae
2523:
2521:
2515:
2513:
2509:Pyrochroidae
2507:
2505:
2504:Pterogeniidae
2502:
2500:
2494:
2492:
2489:
2487:
2481:
2479:
2473:
2471:
2465:
2463:
2457:
2455:
2449:
2447:
2443:Melandryidae
2441:
2439:
2433:
2431:
2430:Chalcodryidae
2428:
2426:
2420:
2418:
2412:
2410:
2404:
2402:
2396:
2395:
2393:
2391:
2387:
2381:
2375:
2374:
2372:
2370:
2366:
2360:
2356:Nemonychidae
2354:
2352:
2346:
2344:
2341:
2339:
2333:
2331:
2325:
2323:
2317:
2315:
2309:
2308:
2306:
2296:
2292:
2286:
2283:
2281:
2275:
2273:
2267:
2265:
2259:
2257:
2254:
2252:
2251:Propalticidae
2249:
2247:
2246:Priasilphidae
2244:
2242:
2239:
2237:
2231:
2229:
2225:Passandridae
2223:
2221:
2219:(sap beetles)
2215:
2213:
2210:
2208:
2202:
2200:
2197:
2190:
2188:
2182:
2180:
2174:
2172:
2169:
2167:
2164:
2162:
2156:
2154:
2151:
2149:
2146:
2144:
2138:
2136:
2130:
2128:
2127:Cavognathidae
2125:
2123:
2120:
2118:
2115:
2108:
2107:
2105:
2103:
2099:
2093:
2090:
2088:
2085:
2083:
2080:
2078:
2072:
2070:
2064:
2062:
2061:Eupsilobiidae
2059:
2057:
2053:Endomychidae
2051:
2049:
2046:
2044:
2040:Corylophidae
2038:
2036:
2030:
2028:
2022:
2020:
2014:
2012:
2009:
2007:
2004:
2002:
1999:
1998:
1996:
1994:
1990:
1984:
1978:
1976:
1973:
1971:
1968:
1966:
1963:
1961:
1958:
1956:
1955:Protopeltidae
1953:
1951:
1950:Prionoceridae
1948:
1946:
1943:
1941:
1938:
1931:
1929:
1926:
1924:
1921:
1914:
1912:
1906:
1904:
1903:Mauroniscidae
1901:
1899:
1896:
1894:
1888:
1886:
1883:
1881:
1875:
1873:
1867:
1865:
1862:
1855:
1854:
1852:
1850:
1846:
1840:
1837:
1835:
1832:
1830:
1827:
1825:
1824:Megalopodidae
1822:
1820:
1817:
1815:
1809:
1807:
1803:Cerambycidae
1801:
1800:
1798:
1796:
1792:
1789:
1787:
1783:
1773:
1769:Derodontidae
1767:
1766:
1764:
1762:
1761:Derodontoidea
1758:
1752:
1746:
1744:
1738:
1736:
1730:
1728:
1727:Endecatomidae
1725:
1723:
1717:
1715:
1711:Bostrichidae
1709:
1708:
1706:
1704:
1703:Bostrichoidea
1700:
1697:
1695:
1691:
1687:
1683:
1677:
1673:
1663:
1660:
1658:
1657:Sphaeriusidae
1655:
1653:
1650:
1648:
1642:
1641:
1638:
1634:
1628:
1624:
1610:
1604:
1602:
1596:
1594:
1591:
1584:
1582:
1579:
1577:
1571:
1569:
1563:
1561:
1555:
1553:
1549:Cicindelidae
1547:
1545:
1539:
1537:
1534:
1532:
1526:
1525:
1523:
1520:
1516:
1512:
1508:
1502:
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1424:
1420:
1418:
1414:
1412:
1408:
1406:
1402:
1400:
1396:
1394:
1390:
1389:
1386:
1381:
1373:
1368:
1366:
1361:
1359:
1354:
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1344:
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1339:
1336:
1335:
1324:
1318:
1310:
1306:
1302:
1298:
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1278:
1271:
1263:
1259:
1254:
1249:
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1237:
1230:
1223:
1222:
1215:
1206:
1202:
1189:
1188:
1184:
1179:
1178:
1174:
1169:
1168:
1164:
1159:
1158:
1154:
1151:Leconte, 1859
1149:
1148:
1144:
1142:
1141:
1137:
1132:
1131:
1127:
1122:
1121:
1117:
1112:
1111:
1107:
1102:
1101:
1097:
1092:
1091:
1087:
1084:Thomson, 1859
1082:
1081:
1077:
1074:Leconte, 1852
1072:
1071:
1067:
1062:
1061:
1057:
1052:
1051:
1047:
1042:
1041:
1037:
1032:
1031:
1027:
1022:
1021:
1017:
1012:
1011:
1007:
1002:
1001:
997:
992:
991:
987:
982:
981:
977:
972:
971:
967:
962:
961:
957:
952:
951:
947:
942:
941:
937:
932:
931:
927:
922:
921:
917:
914:Leconte, 1861
912:
911:
910:Eschatocrepis
907:
902:
901:
897:
894:LeConte, 1859
892:
891:
887:
884:Wolcott, 1944
882:
881:
877:
872:
871:
867:
862:
861:
857:
852:
851:
847:
842:
841:
837:
832:
831:
827:
822:
821:
817:
812:
811:
807:
802:
801:
797:
792:
791:
787:
784:Laporte, 1838
782:
781:
777:
772:
771:
767:
762:
761:
757:
752:
751:
750:Cordylepherus
747:
742:
741:
737:
734:Erichson 1840
732:
731:
727:
722:
721:
717:
712:
711:
707:
702:
701:
697:
692:
691:
687:
684:Leconte, 1880
682:
681:
677:
672:
671:
667:
662:
661:
657:
652:
651:
647:
642:
641:
637:
632:
631:
627:
622:
621:
617:
614:MacLeay, 1872
612:
611:
607:
602:
601:
597:
592:
591:
587:
582:
581:
577:
572:
571:
567:
564:Laporte, 1836
562:
561:
557:
552:
551:
550:Arthrobrachus
547:
542:
541:
537:
532:
531:
527:
522:
521:
517:
512:
511:
507:
502:
501:
497:
492:
491:
487:
482:
481:
477:
476:
466:
463:
461:Fleming, 1821
460:
457:
454:
451:
450:
446:
442:
425:
423:
419:
418:Mauroniscidae
409:
401:
399:
395:
391:
390:
385:
381:
377:
373:
369:
368:
363:
362:
351:
349:
345:
341:
340:
335:
331:
327:
326:
321:
320:
315:
314:
309:
308:
303:
302:
296:
294:
290:
286:
282:
278:
268:
266:
262:
258:
254:
243:
240:
238:Attalomimidae
237:
236:
234:
231:
227:
222:
217:
214:
211:
210:
207:
204:
202:Superfamily:
201:
200:
197:
194:
191:
190:
187:
184:
181:
180:
177:
174:
171:
170:
167:
164:
161:
160:
157:
154:
151:
150:
147:
144:
141:
140:
137:
134:
131:
130:
125:
120:
116:
113:
112:
107:
103:
98:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
36:
22:
19:
3279:
3158:Helophoridae
3150:Georissidae
3145:Epimetopidae
3061:Pleocomidae
3045:Ochodaeidae
3029:Hybosoridae
3013:Glaphyridae
3005:Geotrupidae
2983:
2980:Belohinidae
2941:Eucinetidae
2932:
2929:Decliniidae
2902:
2860:Phengodidae
2799:Cantharidae
2754:Dascillidae
2746:Dascilloidea
2728:Psephenidae
2720:Lutrochidae
2712:Limnichidae
2625:Buprestidae
2617:Buprestoidea
2557:Synchroidae
2541:Scraptiidae
2533:Salpingidae
2496:Prostomidae
2483:Oedemeridae
2459:Mordellidae
2377:Lymexylidae
2369:Lymexyloidea
2319:Attelabidae
2311:Anthribidae
2269:Smicripidae
2233:Phalacridae
2217:Nitidulidae
2212:Myraboliidae
2204:Monotomidae
2195:
2176:Kateretidae
2153:Cyclaxyridae
2113:
2110:Agapythidae
2074:Latridiidae
2024:Cerylonidae
2011:Anamorphidae
1945:Phycosecidae
1936:
1919:
1916:Metaxinidae
1907:
1869:Biphyllidae
1860:
1829:Orsodacnidae
1786:Cucujiformia
1719:Dermestidae
1589:
1528:Amphizoidae
1473:
1453:
1439:Archostemata
1423:Holometabola
1421:Superorder:
1415:Infraclass:
1325:44: 527-558.
1322:
1317:
1284:
1280:
1270:
1243:
1239:
1229:
1219:
1214:
1205:
1185:
1175:
1167:Trichochrous
1165:
1157:Temnopsophus
1155:
1145:
1138:
1128:
1120:Scelopristis
1118:
1108:
1104:Gorham, 1900
1098:
1088:
1078:
1068:
1064:Gorham, 1882
1058:
1048:
1038:
1028:
1018:
1010:Leptovectura
1008:
998:
994:Pascoe, 1866
988:
978:
970:Hylodanacaea
968:
958:
948:
938:
934:Kraatz, 1895
928:
920:Falsomelyris
918:
908:
898:
888:
878:
868:
858:
850:Dolichophron
848:
838:
830:Dicranolaius
828:
818:
808:
798:
788:
778:
768:
758:
748:
738:
728:
718:
708:
698:
688:
680:Chaetocoelus
678:
668:
658:
648:
638:
630:Carphuroides
628:
618:
610:Balanophorus
608:
598:
588:
578:
568:
558:
554:Solier, 1849
548:
538:
528:
520:Anthodromius
518:
508:
498:
488:
478:
440:
415:
407:
404:Distribution
389:Protodasytes
387:
380:Daohugou bed
365:
359:
357:
343:
337:
323:
317:
313:C. semiopaca
311:
305:
299:
297:
288:
280:
274:
256:
252:
251:
215:
196:Cucujiformia
192:Infraorder:
109:
18:
3459:iNaturalist
3304:Wikispecies
3202:Hydraenidae
3176:Spercheidae
3163:Hydrochidae
3127:Synteliidae
3111:Histeridae
3103:Histeroidea
3053:Passalidae
3021:Glaresidae
2878:Throscidae
2838:(fireflies)
2836:Lampyridae
2823:Eucnemidae
2815:Elateridae
2775:Elateroidea
2591:Zopheridae
2475:Mycteridae
2406:Anthicidae
2277:Sphindidae
2261:Silvanidae
2171:Hobartiidae
2158:Erotylidae
2087:Mycetaeidae
2082:Murmidiidae
2066:Euxestidae
1960:Rentoniidae
1834:Oxypeltidae
1819:Disteniidae
1652:Lepiceridae
1581:Hygrobiidae
1573:Haliplidae
1557:Dytiscidae
1536:Aspidytidae
1187:Vecturoides
1124:Mayor, 2004
1014:Casey, 1895
960:Hoppingiana
954:Evers, 1985
944:Broun, 1883
930:Hadrocnemus
860:Dolichosoma
814:Casey, 1895
804:Casey, 1895
794:Casey, 1895
764:Casey, 1895
754:Evers, 1985
710:Clanoptilus
620:Byturosomus
600:Axinotarsus
590:Attalusinus
574:Casey, 1895
504:Solier 1849
467:Leach, 1815
459:Malachiinae
412:Subfamilies
400:of France.
361:Sinomelyris
339:Phyllobates
319:C. rugiceps
285:Malachiinae
277:Malachiinae
271:Description
244:Malachiidae
3605:Categories
3223:Silphidae
3215:Ptiliidae
3207:Leiodidae
3194:Agyrtidae
3037:Lucanidae
2916:Scirtoidea
2852:Omethidae
2831:Jurasaidae
2680:Dryopidae
2651:Byrrhidae
2643:Byrrhoidea
2335:Brentidae
2140:Cucujidae
2122:Boganiidae
2102:Cucujoidea
1975:Thymalidae
1965:Rhadalidae
1908:Melyridae
1877:Byturidae
1839:Vesperidae
1748:Ptiniidae
1598:Noteridae
1565:Gyrinidae
1541:Carabidae
1470:Jurodidae
1462:Cupedidae
1409:Subclass:
1399:Arthropoda
1380:Coleoptera
1191:Fall, 1930
1181:Horn, 1870
1161:Horn, 1872
1130:Sphinginus
1110:Psilothrix
1070:Microlipus
1020:Listropsis
820:Dasytidius
800:Dasytellus
790:Dasytastes
670:Ceratistes
660:Cerapheles
594:Leng, 1918
540:Apalochrus
510:Anthocomus
500:Amecocerus
422:Rhadalidae
394:Cenomanian
367:Juraniscus
307:C. pulchra
182:Suborder:
176:Coleoptera
156:Arthropoda
3611:Melyridae
3337:Melyridae
3324:Melyridae
3310:Melyridae
3280:Melyridae
3077:Trogidae
3071:(scarabs)
2949:Scirtidae
2924:Clambidae
2517:Pythidae
2451:Meloidae
2398:Aderidae
2301:(weevils)
2166:Helotidae
2092:Teredidae
2006:Alexiidae
1890:Cleridae
1849:Cleroidea
1682:Polyphaga
1680:Suborder
1633:Myxophaga
1631:Suborder
1586:Meruidae
1505:Suborder
1487:Ommatidae
1437:Suborder
1411:Pterygota
1391:Kingdom:
1309:234862770
1301:2056-2799
1262:0567-7920
1177:Trophimus
1060:Melyrodes
1040:Malachius
980:Hypebaeus
974:Pic, 1926
924:Pic, 1913
880:Enallonyx
740:Condylops
640:Carphurus
490:Amauronia
480:Ablechrus
465:Melyrinae
453:Dasytinae
445:Dasytinae
424:in 2019.
384:Dasytinae
376:Callovian
344:Choresine
301:Choresine
265:Cleroidea
253:Melyridae
241:Dasytidae
216:Melyridae
206:Cleroidea
186:Polyphaga
142:Kingdom:
136:Eukaryota
24:Melyridae
3368:BugGuide
3344:BioLib:
3295:Q1189209
3289:Wikidata
2844:Lycidae
2688:Elmidae
2586:Ulodidae
2422:Boridae
2327:Belidae
1928:Peltidae
1521:families
1507:Adephaga
1417:Neoptera
1397:Phylum:
1393:Animalia
1382:families
1080:Nepachys
900:Enicopus
890:Endeodes
770:Cyrtosus
760:Cradytes
700:Charopus
690:Chalchas
650:Cerallus
570:Asydates
348:Colombia
325:C. sp. A
230:Synonyms
212:Family:
152:Phylum:
146:Animalia
132:Domain:
3575:4779681
2435:Ciidae
2343:Caridae
1405:Insecta
1403:Class:
1378:Extant
1147:Tanaops
1090:Nodopus
1050:Melyris
1030:Listrus
990:Intybia
940:Halyles
840:Divales
810:Dasytes
780:Danacea
730:Colotes
720:Collops
580:Attalus
560:Astylus
396:) aged
293:elytron
281:Collops
261:beetles
172:Order:
166:Insecta
162:Class:
3588:150723
3572:uBio:
3552:346380
3523:NZOR:
3516:186093
3490:113869
3477:101016
3412:1MELYF
1519:Extant
1307:
1299:
1260:
870:Ebaeus
223:, 1815
3583:WoRMS
3560:Plazi
3539:58189
3472:IRMNG
3464:82067
3425:11036
3347:10063
1305:S2CID
1198:Notes
1000:Laius
221:Leach
3511:NCBI
3485:ITIS
3451:7838
3446:GBIF
3407:EPPO
3373:7482
3360:1085
3355:BOLD
1297:ISSN
1258:ISSN
364:and
322:and
35:Preê
3498:NBN
3399:389
3394:EoL
3386:CL4
3381:CoL
3332:AFD
3319:ADW
1289:doi
1248:doi
386:is
287:);
3607::
3585::
3562::
3549::
3536::
3513::
3500::
3487::
3474::
3461::
3448::
3435::
3422::
3409::
3396::
3383::
3370::
3357::
3334::
3321::
3306::
3291::
1303:.
1295:.
1283:.
1279:.
1256:.
1246:.
1244:64
1242:.
1238:.
378:)
350:.
295:.
267:.
85:Pg
2986:)
2982:(
2935:)
2931:(
2905:)
2901:(
2198:)
2194:(
2116:)
2112:(
1939:)
1935:(
1922:)
1918:(
1863:)
1859:(
1592:)
1588:(
1476:)
1472:(
1456:)
1452:(
1371:e
1364:t
1357:v
1311:.
1291::
1285:7
1264:.
1250::
447:)
443:(
374:(
283:(
90:N
80:K
75:J
70:T
65:P
60:C
55:D
50:S
45:O
40:ê
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