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Paul D. Boyer

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and mostly the release of tightly bound ATP; that three identical catalytic sites went through compulsory, sequential binding changes; and that the binding changes of the catalytic subunits, circularly arranged on the periphery of the enzyme, were driven by the rotation of a smaller internal
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from 1963 to 1989. He was editor of the classic series, "The Enzymes". When he worked on the series "The Enzymes", he was helped by his wife Lyda as she was a professional editor at UCLA. In 1981, he was faculty research lecturer at UCLA. In that same year, he was awarded the prestigious
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Institute and spearheaded the construction of the building and the organization of an interdepartmental Ph.D. program. This institutional service did not diminish the creativity and originality of his research program, which led to three postulates for the binding mechanism for
1207: 1407: 1216: 358:, Paul married Lyda Whicker in 1939, and they remained married for nearly eighty years until his death in 2018, making him the longest-married Nobel laureate. The Boyers had three children. 517: 3711: 555: 536: 3539: 3141: 3623: 3611: 3527: 3575: 1333: 743: 1173: 1055: 3751: 3736: 3331: 3377: 3261: 3237: 3207: 3437: 3395: 3741: 3533: 3479: 3349: 3431: 2882: 1482: 3401: 3389: 3291: 3273: 3563: 3467: 3551: 366: 405:
on a war-related research project dedicated to stabilization of serum albumin for transfusions. He began his independent research career at the
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and the debating team. He was also his high schools valedictorian and played intramural basketball in high school and college. He received a
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Allchin, Douglas (2002), "To err and win a nobel prize: Paul Boyer, ATP synthase and the emergence of bioenergetics",
537:"Occurrence and Characteristics of a Rapid Exchange of Phosphate Oxygens Catalyzed by Sarcoplasmic Reticulum Vesicles" 3726: 3706: 3671: 3108: 3103: 3098: 3093: 3088: 3078: 3073: 3068: 3063: 3058: 2868: 1986: 1468: 773: 3721: 3150: 1093: 617: 143: 1511: 1393: 3343: 3319: 2936: 2122: 1385: 1329: 837: 1303: 3666: 2760: 2692: 478: 3593: 664: 378: 327:, and grew up in a nonpracticing Mormon family of Dutch, German, French, and English descent. He attended 3701: 1450: 441: 1278: 3661: 1810: 1029: 972: 3545: 3279: 2932: 1923: 1619: 1246: 711: 612: 488: 437: 1119: 2901: 2528: 2018: 1915: 1495: 778: 344: 276: 244: 161: 90: 3052: 2997: 2417: 2309: 1899: 1703: 1603: 807: 758: 679: 429: 406: 390: 229: 223: 2991: 2977: 2800: 2788: 2776: 1759: 1004: 822: 795: 292: 167: 112: 1627: 428:. In 1956, he accepted a Hill Foundation Professorship and moved to the medical campus of the 3497: 3177: 2170: 2102: 1739: 1680: 935:
Boyer, P D (April 1995), "From human serum albumin to rotational catalysis by ATP synthase",
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on June 2, 2018, at the age of 99, less than two months shy of his 100th birthday at his
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Since 1963, he had been a professor in the department of chemistry and biochemistry at
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Boyer, Paul D (October 18, 2002), "A research journey with ATP synthase",
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and introduced kinetic, isotopic, and chemical methods for investigating
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https://www.nobelprize.org/prizes/chemistry/1997/boyer/biographical/
2943: 1307: 433: 280: 473: 374: 32: 1362:. UCLA Department of Chemistry and Biochemistry. Archived from 410: 284: 1386:"Golden Plate Awardees of the American Academy of Achievement" 1490: 1451:
Nobel Prize press release — The 1997 Nobel Prize in Chemistry
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Program for Research and Training in Biotechnology (1985–89)
304: 1147:"In memoriam: Paul Boyer, 99, Nobel laureate in chemistry" 377:. In 2003 he was one of 22 Nobel laureates who signed the 3712:
Members of the United States National Academy of Sciences
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Though the Boyers connected with the Mormon community in
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American Society for Biochemistry and Molecular Biology
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American Society for Biochemistry and Molecular Biology
3149: 354:for graduate studies. Five days before leaving for 1215: 451:. In 1965, he became the founding director of the 347:in 1939 and obtained a Wisconsin Alumni Research 3643: 1273: 1271: 444:of the American Society of Biological Chemists. 1232: 367:the Church of Jesus Christ of Latter-day Saints 263:(July 31, 1918 – June 2, 2018) was an American 2890: 1444:including the Nobel Lecture, December 8, 1997 1079: 3752:Members of the American Philosophical Society 3737:University of California, Los Angeles faculty 3135: 2876: 1476: 1352: 1268: 891:, vol. 35, no. 1, pp. 149–72, 991: 989: 939:, vol. 9, no. 7, pp. 559–61, 923:, vol. 58, no. 1, pp. 107–9, 918: 1296: 3142: 3128: 2883: 2869: 1483: 1469: 1439: 1322: 487:by the Southern California Section of the 241:UCLA Molecular Biology Institute (1965–83) 31: 986: 944: 870: 792:Department of Chemistry and Biochemistry 630:John Simon Guggenheim Memorial Foundation 1171: 1053: 369:(LDS Church). After experimenting with 2961:International Campaign to Ban Landmines 1205: 1153:. University of California, Los Angeles 1144: 886: 3742:University of Wisconsin–Madison alumni 3644: 730:University of California, Los Angeles 318: 3123: 2864: 1464: 934: 856: 790:University of California, Los Angeles 697:University of California, Los Angeles 650:American Academy of Arts and Sciences 449:University of California, Los Angeles 273:University of California, Los Angeles 3161:Paul-Lewis Award in Enzyme Chemistry 1172:McFadden, Robert D. (June 7, 2018). 1054:McFadden, Robert D. (June 7, 2018). 1024: 1022: 618:Paul-Lewis Award in Enzyme Chemistry 585: 575:University of California Los Angeles 560:University of California Los Angeles 541:University of California Los Angeles 522:University of California Los Angeles 1030:"The Nobel Prize in Chemistry 1997" 997:"The Nobel Prize in Chemistry 1997" 436:of the Biochemistry Section of the 413:mechanisms. In 1955, he received a 13: 3747:Annual Reviews (publisher) editors 3682:American people of English descent 1240:"Pfizer Award in Enzyme Chemistry" 579:United States Department of Energy 564:United States Department of Energy 545:United States Department of Energy 526:United States Department of Energy 384: 271:, and a professor of chemistry at 14: 3763: 3692:American people of German descent 3687:American people of French descent 1429: 1019: 969:The nobel prize in chemistry 1997 889:Journal of the History of Biology 774:Royal Swedish Academy of Sciences 3677:American people of Dutch descent 3151:Pfizer Award in Enzyme Chemistry 1713: 1145:Wolpert, Stuart (June 4, 2018). 1094:Freedom From Religion Foundation 547:(through predecessor agency the 535:Kanazawa, T. & P. D. Boyer. 528:(through predecessor agency the 516:Dahms, A. S. & P. D. Boyer. 144:Pfizer Award in Enzyme Chemistry 3732:University of Minnesota faculty 3697:Brigham Young University alumni 1400: 1394:American Academy of Achievement 1378: 1126:. American Humanist Association 859:Journal of Biological Chemistry 823:American Academy of Achievement 510: 399:University of Wisconsin–Madison 96:University of Wisconsin–Madison 1206:Schudel, Matt (June 7, 2018). 1199: 1165: 1138: 1112: 1073: 1047: 919:Lores Arnaiz, G R (1998), "", 838:American Philosophical Society 1: 1080:Boyer, Paul D. (March 2004), 850: 479:Annual Review of Biochemistry 432:. In 1959–1960, he served as 373:, Boyer eventually became an 3717:Nobel laureates in Chemistry 1456:UCLA webpage — Paul D. Boyer 1124:Humanism and Its Aspirations 715:Southern California Section 665:National Academy of Sciences 7: 476:or associate editor of the 401:in 1943, he spent years at 275:(UCLA). He shared the 1997 10: 3768: 1304:"1981 Paul D. Boyer, UCLA" 946:10.1096/fasebj.9.7.7737466 733:Faculty research lecturer 440:(ACS) and in 1969–1970 as 3167: 3157: 3047: 3022: 3007: 2976: 2951: 2931: 2900: 2519: 2220: 1961: 1722: 1711: 1502: 1247:American Chemical Society 712:American Chemical Society 613:American Chemical Society 489:American Chemical Society 438:American Chemical Society 389:After Boyer received his 331:, where he was active in 311:for his discovery of the 303:, making Boyer the first 287:mechanism underlying the 254: 210: 184: 177: 136: 128: 104: 83: 64: 42: 30: 23: 3727:Provo High School alumni 3707:Former Latter Day Saints 3672:American Nobel laureates 2637:Venkatraman Ramakrishnan 1916:Wendell Meredith Stanley 1496:Nobel Prize in Chemistry 979: 779:Nobel Prize in Chemistry 549:Atomic Energy Commission 530:Atomic Energy Commission 494: 345:Brigham Young University 277:Nobel Prize in Chemistry 91:Brigham Young University 3722:People from Provo, Utah 3612:Mohammad Seyedsayamdost 3528:Wilfred A. van der Donk 2998:William Daniel Phillips 1330:"William C. Rose Award" 897:10.1023/A:1014583721788 808:University of Wisconsin 759:University of Minnesota 680:University of Stockholm 430:University of Minnesota 407:University of Minnesota 230:University of Minnesota 224:University of Minnesota 3053:Nobel Prize recipients 3040:(Canada/United States) 3009:Physiology or Medicine 2992:Claude Cohen-Tannoudji 2801:Emmanuelle Charpentier 2789:M. Stanley Whittingham 1760:Hans von Euler-Chelpin 872:10.1074/jbc.X200001200 796:Glenn T. Seaborg Medal 748:William C. Rose Award 581:, (December 31, 1993). 293:adenosine triphosphate 232:(Minneapolis, 1956–63) 113:adenosine triphosphate 3498:David W. Christianson 3344:Michael J. Chamberlin 2171:Christian B. Anfinsen 1811:FrĂ©dĂ©ric Joliot-Curie 1446:Energy, Life, and ATP 635:Guggenheim Fellowship 426:alcohol dehydrogenase 415:Guggenheim Fellowship 3667:American biochemists 3516:Karin Musier-Forsyth 3332:Ekkehard K. F. Bautz 3286:Frederic M. Richards 2833:Karl Barry Sharpless 2330:Dudley R. Herschbach 2155:Luis Federico Leloir 1412:search.amphilsoc.org 1408:"APS Member History" 424:on the mechanism of 3378:Frederik C. Hartman 3366:Stephen J. Benkovic 3360:Joseph L. Goldstein 3262:Frank M. Huennekens 3238:G. Robert Greenberg 3208:Bernard L. Horecker 3184:Albert L. Lehninger 3015:Stanley B. Prusiner 2825:Carolyn R. Bertozzi 2737:Jean-Pierre Sauvage 2474:Jens Christian Skou 2442:F. Sherwood Rowland 2354:Charles J. Pedersen 2318:Herbert A. Hauptman 2011:Vincent du Vigneaud 1628:Richard Willstätter 1512:Jacobus van 't Hoff 1390:www.achievement.org 1366:on October 23, 2016 1360:"The Seaborg Medal" 1082:"A Path to Atheism" 827:Golden Plate Award 812:Honorary Doctorate 763:Honorary Doctorate 501:respiratory failure 403:Stanford University 319:Birth and education 309:Jens Christian Skou 226:(St. Paul, 1945–46) 218:Stanford University 16:American biochemist 3702:Chemists from Utah 3438:Kenneth A. Johnson 3396:Richard R. Burgess 3280:Marshall Nirenberg 3268:Jack L. Strominger 2785:John B. Goodenough 2713:William E. Moerner 2593:Richard R. Schrock 2565:Roderick MacKinnon 2537:K. Barry Sharpless 2362:Johann Deisenhofer 2191:Geoffrey Wilkinson 2187:Ernst Otto Fischer 2111:Robert S. Mulliken 2047:Jaroslav HeyrovskĂ˝ 1995:Hermann Staudinger 1815:Irène Joliot-Curie 1438:on Nobelprize.org 1256:on January 1, 2017 1060:The New York Times 921:Medicina (B Aires) 685:Honorary Doctorate 379:Humanist Manifesto 333:student government 323:Boyer was born in 269:analytical chemist 78:, California, U.S. 3662:American atheists 3639: 3638: 3534:Nicole S. Sampson 3480:P. Andrew Karplus 3408:Robert T.N. Tjian 3350:Malcolm L. Gefter 3320:Herbert Weissbach 3314:Robert T. Schimke 3308:William J. Rutter 3244:Eugene P. Kennedy 3117: 3116: 3024:Economic Sciences 2858: 2857: 2841:Moungi G. Bawendi 2761:Richard Henderson 2605:Roger D. Kornberg 2573:Aaron Ciechanover 2406:Rudolph A. Marcus 2310:Robert Merrifield 2246:Peter D. Mitchell 2019:Cyril Hinshelwood 1780:Friedrich Bergius 1704:Richard Zsigmondy 1620:Theodore Richards 1568:Ernest Rutherford 1340:on March 24, 2017 1283:www.nasonline.org 1120:"Notable Signers" 1086:Freethought Today 1007:on March 25, 2017 848: 847: 586:Awards and honors 453:Molecular Biology 329:Provo High School 258: 257: 202:Molecular biology 179:Scientific career 150:Guggenheim Fellow 3759: 3618:Kenichi Yokoyama 3594:Douglas Mitchell 3558:Virginia Cornish 3546:Neil L. Kelleher 3492:Ronald T. Raines 3486:Daniel Herschlag 3432:John W. Kozarich 3384:Thomas A. Steitz 3356:Michael S. Brown 3214:Earl R. Stadtman 3144: 3137: 3130: 3121: 3120: 3055: 3032:Robert C. Merton 2918:(United Kingdom) 2885: 2878: 2871: 2862: 2861: 2851: 2849:Alexei I. Ekimov 2835: 2829:Morten P. Meldal 2819: 2807: 2795: 2779: 2763: 2753:Jacques Dubochet 2747: 2731: 2715: 2699: 2683: 2677:Robert Lefkowitz 2671: 2663: 2647: 2641:Thomas A. Steitz 2631: 2615: 2607: 2599: 2589:Robert H. Grubbs 2583: 2567: 2555: 2539: 2512: 2510:Hideki Shirakawa 2496: 2488: 2476: 2460: 2444: 2428: 2420: 2408: 2400: 2398:Richard R. Ernst 2392: 2384: 2372: 2356: 2340: 2324: 2312: 2304: 2296: 2288: 2276: 2274:Frederick Sanger 2260: 2254:Herbert C. Brown 2248: 2240: 2232: 2230:William Lipscomb 2213: 2201: 2193: 2181: 2165: 2163:Gerhard Herzberg 2157: 2149: 2137: 2129: 2113: 2105: 2097: 2089: 2077: 2065: 2057: 2049: 2041: 2039:Frederick Sanger 2033: 2025: 2023:Nikolay Semyonov 2013: 2005: 1997: 1989: 1977: 1975:Glenn T. Seaborg 1954: 1942: 1934: 1926: 1918: 1902: 1900:Artturi Virtanen 1894: 1886: 1884:George de Hevesy 1878: 1877: 1871: 1870: 1864: 1863: 1857: 1845: 1837: 1825: 1817: 1805: 1797: 1796: 1790: 1782: 1770: 1762: 1750: 1742: 1740:Heinrich Wieland 1734: 1732:Theodor Svedberg 1717: 1716: 1706: 1698: 1697: 1691: 1683: 1675: 1667: 1659: 1658: 1652: 1644: 1643: 1637: 1636: 1630: 1622: 1614: 1606: 1594: 1586: 1578: 1570: 1562: 1554: 1546: 1544:Adolf von Baeyer 1538: 1530: 1528:Svante Arrhenius 1522: 1514: 1485: 1478: 1471: 1462: 1461: 1443: 1423: 1422: 1420: 1418: 1404: 1398: 1397: 1382: 1376: 1375: 1373: 1371: 1356: 1350: 1349: 1347: 1345: 1336:. Archived from 1326: 1320: 1319: 1317: 1315: 1310:on April 4, 2016 1300: 1294: 1293: 1291: 1289: 1275: 1266: 1265: 1263: 1261: 1255: 1249:. Archived from 1244: 1236: 1230: 1229: 1227: 1225: 1219: 1214:. Archived from 1203: 1197: 1196: 1194: 1192: 1169: 1163: 1162: 1160: 1158: 1142: 1136: 1135: 1133: 1131: 1116: 1110: 1109: 1107: 1105: 1096:, archived from 1077: 1071: 1070: 1068: 1066: 1051: 1045: 1044: 1042: 1040: 1026: 1017: 1016: 1014: 1012: 1003:. Archived from 993: 965: 948: 931: 915: 883: 874: 865:(42): 39045–61, 639:study in Sweden 590: 589: 417:and worked with 261:Paul Delos Boyer 71: 52: 50: 35: 21: 20: 3767: 3766: 3762: 3761: 3760: 3758: 3757: 3756: 3642: 3641: 3640: 3635: 3462:Michael H. Gelb 3402:Paul L. Mordich 3390:Daniel V. Santi 3338:Howard M. Temin 3292:Samuel B. Weiss 3274:Charles Gilvarg 3232:Merton F. Utter 3202:Arthur Kornberg 3163: 3153: 3148: 3118: 3113: 3051: 3043: 3034:(United States) 3018: 3017:(United States) 3003: 3000:(United States) 2988:(United States) 2972: 2969:(United States) 2947: 2927: 2912:(United States) 2896: 2889: 2859: 2854: 2838: 2822: 2813:David MacMillan 2810: 2805:Jennifer Doudna 2798: 2782: 2766: 2750: 2741:Fraser Stoddart 2734: 2725:Paul L. Modrich 2718: 2702: 2686: 2674: 2666: 2661:Ei-ichi Negishi 2653:Richard F. Heck 2650: 2634: 2621:Osamu Shimomura 2618: 2610: 2602: 2586: 2570: 2558: 2542: 2529:William Knowles 2526: 2515: 2506:Alan MacDiarmid 2499: 2491: 2479: 2463: 2458:Richard Smalley 2447: 2434:Paul J. Crutzen 2431: 2423: 2411: 2403: 2395: 2387: 2375: 2359: 2350:Jean-Marie Lehn 2343: 2327: 2315: 2307: 2299: 2291: 2279: 2263: 2251: 2243: 2235: 2227: 2216: 2211:Vladimir Prelog 2204: 2196: 2184: 2168: 2160: 2152: 2140: 2132: 2116: 2108: 2103:Robert Woodward 2100: 2095:Dorothy Hodgkin 2092: 2080: 2068: 2060: 2052: 2044: 2036: 2028: 2016: 2008: 2000: 1992: 1980: 1968: 1957: 1945: 1940:William Giauque 1937: 1929: 1924:Robert Robinson 1921: 1908:James B. Sumner 1905: 1897: 1889: 1881: 1875: 1874: 1868: 1867: 1861: 1860: 1855:Leopold RuĹľiÄŤka 1851:Adolf Butenandt 1848: 1840: 1828: 1820: 1808: 1800: 1794: 1793: 1788:Irving Langmuir 1785: 1773: 1765: 1753: 1745: 1737: 1729: 1718: 1714: 1709: 1701: 1695: 1694: 1686: 1678: 1673:Frederick Soddy 1670: 1662: 1656: 1655: 1647: 1641: 1640: 1634: 1633: 1625: 1617: 1609: 1600:Victor Grignard 1597: 1589: 1581: 1576:Wilhelm Ostwald 1573: 1565: 1557: 1549: 1541: 1533: 1525: 1517: 1509: 1498: 1489: 1432: 1427: 1426: 1416: 1414: 1406: 1405: 1401: 1384: 1383: 1379: 1369: 1367: 1358: 1357: 1353: 1343: 1341: 1328: 1327: 1323: 1313: 1311: 1302: 1301: 1297: 1287: 1285: 1279:"Paul D. Boyer" 1277: 1276: 1269: 1259: 1257: 1253: 1242: 1238: 1237: 1233: 1223: 1221: 1220:on June 7, 2018 1212:Washington Post 1204: 1200: 1190: 1188: 1170: 1166: 1156: 1154: 1143: 1139: 1129: 1127: 1118: 1117: 1113: 1103: 1101: 1100:on June 3, 2011 1078: 1074: 1064: 1062: 1052: 1048: 1038: 1036: 1028: 1027: 1020: 1010: 1008: 995: 994: 987: 982: 853: 791: 714: 638: 600: 588: 513: 497: 472:Paul Boyer was 387: 385:Academic career 321: 250: 206: 173: 124: 100: 84:Alma mater 79: 73: 69: 60: 54: 48: 46: 38: 26: 17: 12: 11: 5: 3765: 3755: 3754: 3749: 3744: 3739: 3734: 3729: 3724: 3719: 3714: 3709: 3704: 3699: 3694: 3689: 3684: 3679: 3674: 3669: 3664: 3659: 3654: 3637: 3636: 3634: 3633: 3627: 3621: 3615: 3609: 3603: 3600:Michelle Chang 3597: 3591: 3585: 3579: 3573: 3570:Sarah O'Connor 3567: 3564:Vahe Bandarian 3561: 3555: 3549: 3543: 3537: 3531: 3525: 3519: 3513: 3507: 3501: 3495: 3489: 3483: 3477: 3471: 3468:Donald Hilvert 3465: 3459: 3453: 3447: 3444:James A. Wells 3441: 3435: 3429: 3426:Gregory Petsko 3423: 3417: 3411: 3405: 3399: 3393: 3387: 3381: 3375: 3369: 3363: 3353: 3347: 3341: 3335: 3329: 3323: 3317: 3311: 3305: 3302:Salih J. Wakil 3298:P. Roy Vagelos 3295: 3289: 3283: 3277: 3271: 3265: 3259: 3253: 3247: 3241: 3235: 3229: 3223: 3217: 3211: 3205: 3199: 3196:Britton Chance 3193: 3190:Henry A. Lardy 3187: 3181: 3175: 3172:David E. Green 3168: 3165: 3164: 3158: 3155: 3154: 3147: 3146: 3139: 3132: 3124: 3115: 3114: 3112: 3111: 3106: 3101: 3096: 3091: 3086: 3081: 3076: 3071: 3066: 3061: 3056: 3048: 3045: 3044: 3042: 3041: 3035: 3028: 3026: 3020: 3019: 3013: 3011: 3005: 3004: 3002: 3001: 2995: 2989: 2982: 2980: 2974: 2973: 2971: 2970: 2964: 2957: 2955: 2949: 2948: 2942: 2940: 2929: 2928: 2926: 2925: 2919: 2916:John E. Walker 2913: 2906: 2904: 2898: 2897: 2888: 2887: 2880: 2873: 2865: 2856: 2855: 2853: 2852: 2836: 2820: 2808: 2796: 2780: 2773:Gregory Winter 2769:Frances Arnold 2764: 2748: 2732: 2716: 2700: 2693:Michael Levitt 2689:Martin Karplus 2684: 2672: 2664: 2648: 2632: 2629:Roger Y. Tsien 2625:Martin Chalfie 2616: 2608: 2600: 2584: 2568: 2556: 2540: 2523: 2521: 2517: 2516: 2514: 2513: 2502:Alan J. Heeger 2497: 2489: 2477: 2470:John E. Walker 2461: 2445: 2429: 2421: 2409: 2401: 2393: 2385: 2373: 2370:Hartmut Michel 2357: 2346:Donald J. Cram 2341: 2325: 2313: 2305: 2297: 2289: 2286:Roald Hoffmann 2277: 2270:Walter Gilbert 2261: 2249: 2241: 2238:Ilya Prigogine 2233: 2224: 2222: 2218: 2217: 2215: 2214: 2207:John Cornforth 2202: 2194: 2182: 2175:Stanford Moore 2166: 2158: 2150: 2138: 2130: 2123:Ronald Norrish 2114: 2106: 2098: 2090: 2078: 2066: 2058: 2050: 2042: 2034: 2031:Alexander Todd 2026: 2014: 2006: 1998: 1990: 1978: 1971:Edwin McMillan 1965: 1963: 1959: 1958: 1956: 1955: 1943: 1935: 1927: 1919: 1903: 1895: 1887: 1879: 1872: 1865: 1858: 1846: 1838: 1831:Norman Haworth 1826: 1818: 1806: 1798: 1791: 1783: 1771: 1763: 1751: 1743: 1735: 1726: 1724: 1720: 1719: 1712: 1710: 1708: 1707: 1699: 1692: 1684: 1676: 1668: 1665:Walther Nernst 1660: 1653: 1645: 1638: 1631: 1623: 1615: 1607: 1595: 1587: 1579: 1571: 1563: 1560:Eduard Buchner 1555: 1547: 1539: 1536:William Ramsay 1531: 1523: 1515: 1506: 1504: 1500: 1499: 1488: 1487: 1480: 1473: 1465: 1459: 1458: 1453: 1448: 1431: 1430:External links 1428: 1425: 1424: 1399: 1377: 1351: 1321: 1295: 1267: 1231: 1198: 1178:New York Times 1164: 1137: 1111: 1072: 1046: 1034:NobelPrize.org 1018: 1001:Nobelprize.org 984: 983: 981: 978: 977: 976: 966: 932: 916: 884: 852: 849: 846: 845: 843: 840: 835: 831: 830: 828: 825: 820: 816: 815: 813: 810: 805: 801: 800: 798: 793: 788: 784: 783: 781: 776: 771: 767: 766: 764: 761: 756: 752: 751: 749: 746: 741: 737: 736: 734: 731: 728: 724: 723: 721: 716: 709: 705: 704: 702: 699: 694: 690: 689: 687: 682: 677: 673: 672: 670: 667: 662: 658: 657: 655: 652: 647: 643: 642: 640: 632: 627: 623: 622: 620: 615: 610: 606: 605: 602: 597: 594: 587: 584: 583: 582: 567: 552: 533: 512: 509: 499:Boyer died of 496: 493: 386: 383: 320: 317: 301:John E. Walker 256: 255: 252: 251: 249: 248: 242: 239: 233: 227: 221: 214: 212: 208: 207: 205: 204: 199: 194: 188: 186: 182: 181: 175: 174: 172: 171: 165: 159: 153: 147: 140: 138: 134: 133: 130: 126: 125: 123: 122: 115: 108: 106: 105:Known for 102: 101: 99: 98: 93: 87: 85: 81: 80: 74: 72:(aged 99) 66: 62: 61: 55: 44: 40: 39: 36: 28: 27: 24: 15: 9: 6: 4: 3: 2: 3764: 3753: 3750: 3748: 3745: 3743: 3740: 3738: 3735: 3733: 3730: 3728: 3725: 3723: 3720: 3718: 3715: 3713: 3710: 3708: 3705: 3703: 3700: 3698: 3695: 3693: 3690: 3688: 3685: 3683: 3680: 3678: 3675: 3673: 3670: 3668: 3665: 3663: 3660: 3658: 3655: 3653: 3650: 3649: 3647: 3631: 3628: 3625: 3622: 3619: 3616: 3613: 3610: 3607: 3606:Emily Balskus 3604: 3601: 3598: 3595: 3592: 3589: 3586: 3583: 3580: 3577: 3574: 3571: 3568: 3565: 3562: 3559: 3556: 3553: 3552:Carsten Krebs 3550: 3547: 3544: 3541: 3538: 3535: 3532: 3529: 3526: 3523: 3522:Dorothee Kern 3520: 3517: 3514: 3511: 3510:Ruma Banerjee 3508: 3505: 3502: 3499: 3496: 3493: 3490: 3487: 3484: 3481: 3478: 3475: 3472: 3469: 3466: 3463: 3460: 3457: 3454: 3451: 3448: 3445: 3442: 3439: 3436: 3433: 3430: 3427: 3424: 3421: 3420:JoAnne Stubbe 3418: 3415: 3412: 3409: 3406: 3403: 3400: 3397: 3394: 3391: 3388: 3385: 3382: 3379: 3376: 3373: 3372:Paul Schimmel 3370: 3367: 3364: 3361: 3357: 3354: 3351: 3348: 3345: 3342: 3339: 3336: 3333: 3330: 3327: 3324: 3321: 3318: 3315: 3312: 3309: 3306: 3303: 3299: 3296: 3293: 3290: 3287: 3284: 3281: 3278: 3275: 3272: 3269: 3266: 3263: 3260: 3257: 3256:Arthur Pardee 3254: 3251: 3250:Minor J. Coon 3248: 3245: 3242: 3239: 3236: 3233: 3230: 3227: 3226:Paul D. Boyer 3224: 3221: 3220:Alton Meister 3218: 3215: 3212: 3209: 3206: 3203: 3200: 3197: 3194: 3191: 3188: 3185: 3182: 3179: 3178:Van R. Potter 3176: 3173: 3170: 3169: 3166: 3162: 3159:Formerly the 3156: 3152: 3145: 3140: 3138: 3133: 3131: 3126: 3125: 3122: 3110: 3107: 3105: 3102: 3100: 3097: 3095: 3092: 3090: 3087: 3085: 3082: 3080: 3077: 3075: 3072: 3070: 3067: 3065: 3062: 3060: 3057: 3054: 3050: 3049: 3046: 3039: 3038:Myron Scholes 3036: 3033: 3030: 3029: 3027: 3025: 3021: 3016: 3012: 3010: 3006: 2999: 2996: 2993: 2990: 2987: 2984: 2983: 2981: 2979: 2975: 2968: 2967:Jody Williams 2965: 2963:(Switzerland) 2962: 2959: 2958: 2956: 2954: 2950: 2945: 2941: 2938: 2934: 2930: 2923: 2920: 2917: 2914: 2911: 2910:Paul D. Boyer 2908: 2907: 2905: 2903: 2899: 2894: 2886: 2881: 2879: 2874: 2872: 2867: 2866: 2863: 2850: 2846: 2845:Louis E. Brus 2842: 2837: 2834: 2830: 2826: 2821: 2818: 2817:Benjamin List 2814: 2809: 2806: 2802: 2797: 2794: 2793:Akira Yoshino 2790: 2786: 2781: 2778: 2774: 2770: 2765: 2762: 2758: 2757:Joachim Frank 2754: 2749: 2746: 2742: 2738: 2733: 2730: 2726: 2722: 2721:Tomas Lindahl 2717: 2714: 2710: 2706: 2701: 2698: 2697:Arieh Warshel 2694: 2690: 2685: 2682: 2681:Brian Kobilka 2678: 2673: 2670: 2669:Dan Shechtman 2665: 2662: 2658: 2654: 2649: 2646: 2645:Ada E. Yonath 2642: 2638: 2633: 2630: 2626: 2622: 2617: 2614: 2609: 2606: 2601: 2598: 2594: 2590: 2585: 2582: 2578: 2577:Avram Hershko 2574: 2569: 2566: 2562: 2557: 2554: 2553:Kurt WĂĽthrich 2550: 2549:Koichi Tanaka 2546: 2541: 2538: 2534: 2530: 2525: 2524: 2522: 2518: 2511: 2507: 2503: 2498: 2495: 2490: 2487: 2483: 2478: 2475: 2471: 2467: 2466:Paul D. Boyer 2462: 2459: 2455: 2451: 2446: 2443: 2439: 2435: 2430: 2427: 2422: 2419: 2418:Michael Smith 2415: 2410: 2407: 2402: 2399: 2394: 2391: 2386: 2383: 2379: 2378:Sidney Altman 2374: 2371: 2367: 2363: 2358: 2355: 2351: 2347: 2342: 2339: 2335: 2331: 2326: 2323: 2319: 2314: 2311: 2306: 2303: 2298: 2295: 2290: 2287: 2283: 2282:Kenichi Fukui 2278: 2275: 2271: 2267: 2262: 2259: 2255: 2250: 2247: 2242: 2239: 2234: 2231: 2226: 2225: 2223: 2219: 2212: 2208: 2203: 2200: 2195: 2192: 2188: 2183: 2180: 2179:William Stein 2176: 2172: 2167: 2164: 2159: 2156: 2151: 2148: 2144: 2139: 2136: 2131: 2128: 2127:George Porter 2124: 2120: 2119:Manfred Eigen 2115: 2112: 2107: 2104: 2099: 2096: 2091: 2088: 2084: 2079: 2076: 2072: 2067: 2064: 2063:Melvin Calvin 2059: 2056: 2055:Willard Libby 2051: 2048: 2043: 2040: 2035: 2032: 2027: 2024: 2020: 2015: 2012: 2007: 2004: 2003:Linus Pauling 1999: 1996: 1991: 1988: 1987:Richard Synge 1984: 1983:Archer Martin 1979: 1976: 1972: 1967: 1966: 1964: 1960: 1953: 1949: 1944: 1941: 1936: 1933: 1932:Arne Tiselius 1928: 1925: 1920: 1917: 1913: 1912:John Northrop 1909: 1904: 1901: 1896: 1893: 1888: 1885: 1880: 1873: 1866: 1859: 1856: 1852: 1847: 1844: 1839: 1836: 1832: 1827: 1824: 1819: 1816: 1812: 1807: 1804: 1799: 1792: 1789: 1784: 1781: 1777: 1772: 1769: 1764: 1761: 1757: 1756:Arthur Harden 1752: 1749: 1748:Adolf Windaus 1744: 1741: 1736: 1733: 1728: 1727: 1725: 1721: 1705: 1700: 1693: 1690: 1685: 1682: 1681:Francis Aston 1677: 1674: 1669: 1666: 1661: 1654: 1651: 1646: 1639: 1632: 1629: 1624: 1621: 1616: 1613: 1612:Alfred Werner 1608: 1605: 1604:Paul Sabatier 1601: 1596: 1593: 1588: 1585: 1580: 1577: 1572: 1569: 1564: 1561: 1556: 1553: 1552:Henri Moissan 1548: 1545: 1540: 1537: 1532: 1529: 1524: 1521: 1516: 1513: 1508: 1507: 1505: 1501: 1497: 1493: 1486: 1481: 1479: 1474: 1472: 1467: 1466: 1463: 1457: 1454: 1452: 1449: 1447: 1442: 1437: 1436:Paul D. Boyer 1434: 1433: 1413: 1409: 1403: 1395: 1391: 1387: 1381: 1365: 1361: 1355: 1339: 1335: 1331: 1325: 1309: 1305: 1299: 1284: 1280: 1274: 1272: 1252: 1248: 1241: 1235: 1218: 1213: 1209: 1202: 1187: 1183: 1179: 1175: 1168: 1152: 1151:UCLA Newsroom 1148: 1141: 1130:September 15, 1125: 1121: 1115: 1099: 1095: 1091: 1087: 1083: 1076: 1065:September 17, 1061: 1057: 1050: 1035: 1031: 1025: 1023: 1006: 1002: 998: 992: 990: 985: 974: 970: 967: 964: 960: 956: 952: 947: 942: 938: 933: 930: 926: 922: 917: 914: 910: 906: 902: 898: 894: 890: 885: 882: 878: 873: 868: 864: 860: 855: 854: 844: 841: 839: 836: 833: 832: 829: 826: 824: 821: 818: 817: 814: 811: 809: 806: 803: 802: 799: 797: 794: 789: 786: 785: 782: 780: 777: 775: 772: 769: 768: 765: 762: 760: 757: 754: 753: 750: 747: 745: 742: 739: 738: 735: 732: 729: 726: 725: 722: 720: 717: 713: 710: 707: 706: 703: 700: 698: 695: 692: 691: 688: 686: 683: 681: 678: 675: 674: 671: 668: 666: 663: 660: 659: 656: 653: 651: 648: 645: 644: 641: 636: 633: 631: 628: 625: 624: 621: 619: 616: 614: 611: 608: 607: 603: 598: 596:Organization 595: 592: 591: 580: 576: 572: 569:Boyer, P. D. 568: 565: 561: 557: 554:Boyer, P. D. 553: 550: 546: 542: 538: 534: 531: 527: 523: 519: 515: 514: 508: 506: 502: 492: 490: 486: 481: 480: 475: 470: 468: 463: 459: 458:ATP synthesis 454: 450: 445: 443: 439: 435: 431: 427: 423: 422:Hugo Theorell 420: 416: 412: 408: 404: 400: 396: 392: 382: 380: 376: 372: 368: 364: 359: 357: 353: 350: 346: 342: 338: 334: 330: 326: 316: 314: 313:Na+/K+-ATPase 310: 306: 302: 298: 294: 290: 286: 282: 278: 274: 270: 266: 262: 253: 246: 243: 240: 237: 234: 231: 228: 225: 222: 219: 216: 215: 213: 209: 203: 200: 198: 195: 193: 190: 189: 187: 183: 180: 176: 169: 168:Seaborg Medal 166: 163: 160: 157: 154: 151: 148: 145: 142: 141: 139: 135: 131: 127: 120: 116: 114: 110: 109: 107: 103: 97: 94: 92: 89: 88: 86: 82: 77: 67: 63: 58: 53:July 31, 1918 45: 41: 37:Boyer in 2016 34: 29: 25:Paul D. Boyer 22: 19: 3630:Amie K. Boal 3582:Kate Carroll 3540:James Berger 3504:Eric T. Kool 3474:Gerald Joyce 3456:Carl O. Pabo 3225: 3160: 3083: 2922:Jens C. Skou 2909: 2777:George Smith 2657:Akira Suzuki 2613:Gerhard Ertl 2597:Yves Chauvin 2545:John B. Fenn 2533:Ryoji Noyori 2520:2001–present 2494:Ahmed Zewail 2465: 2454:Harold Kroto 2438:Mario Molina 2366:Robert Huber 2338:John Polanyi 2322:Jerome Karle 2258:Georg Wittig 2143:Derek Barton 2135:Lars Onsager 2087:Giulio Natta 2083:Karl Ziegler 2075:John Kendrew 1843:Richard Kuhn 1768:Hans Fischer 1584:Otto Wallach 1520:Emil Fischer 1445: 1415:. Retrieved 1411: 1402: 1389: 1380: 1368:. Retrieved 1364:the original 1354: 1342:. Retrieved 1338:the original 1324: 1312:. Retrieved 1308:the original 1298: 1286:. Retrieved 1282: 1258:. Retrieved 1251:the original 1234: 1222:. Retrieved 1217:the original 1211: 1201: 1189:. Retrieved 1177: 1167: 1155:. Retrieved 1150: 1140: 1128:. Retrieved 1123: 1114: 1102:, retrieved 1098:the original 1089: 1085: 1075: 1063:. Retrieved 1059: 1049: 1037:. Retrieved 1033: 1009:. Retrieved 1005:the original 968: 936: 920: 888: 862: 858: 719:Tolman Award 701:McCoy Award 599:Award title, 511:Publications 498: 485:Tolman Medal 477: 471: 446: 395:biochemistry 388: 371:Unitarianism 360: 322: 297:ATP synthase 289:biosynthesis 260: 259: 211:Institutions 197:Biochemistry 178: 156:Tolman Award 132:Lyda Whicker 118: 111:Researching 70:(2018-06-02) 68:June 2, 2018 18: 3657:2018 deaths 3652:1918 births 3624:Rahul Kohli 3450:Ronald Vale 3414:Thomas Cech 3326:Jack Preiss 2893:Nobel Prize 2745:Ben Feringa 2729:Aziz Sancar 2709:Stefan Hell 2705:Eric Betzig 2482:Walter Kohn 2450:Robert Curl 2426:George Olah 2414:Kary Mullis 2390:Elias Corey 2382:Thomas Cech 2334:Yuan T. Lee 2302:Henry Taube 1835:Paul Karrer 1823:Peter Debye 1803:Harold Urey 1689:Fritz Pregl 1650:Fritz Haber 1592:Marie Curie 1417:December 3, 1288:December 3, 505:Los Angeles 352:Scholarship 325:Provo, Utah 162:Nobel Prize 119:The Enzymes 76:Los Angeles 57:Provo, Utah 3646:Categories 3588:Hening Lin 2986:Steven Chu 2933:Literature 2581:Irwin Rose 2561:Peter Agre 2486:John Pople 2294:Aaron Klug 2199:Paul Flory 2147:Odd Hassel 2071:Max Perutz 1952:Kurt Alder 1948:Otto Diels 1776:Carl Bosch 851:References 551:), (1972). 532:), (1972). 393:degree in 349:Foundation 265:biochemist 117:Editor of 49:1918-07-31 3576:Jin Zhang 2924:(Denmark) 2902:Chemistry 2895:laureates 2266:Paul Berg 2221:1976–2000 1962:1951–1975 1892:Otto Hahn 1723:1926–1950 1503:1901–1925 1492:Laureates 1370:March 23, 1344:March 23, 1314:March 23, 1260:March 23, 1224:August 3, 1186:0362-4331 1104:March 16, 1039:April 12, 1011:March 23, 601:Category 462:phosphate 442:President 419:Professor 397:from the 363:Wisconsin 356:Wisconsin 341:chemistry 285:enzymatic 238:(1963–90) 220:(1943–45) 192:Chemistry 121:(1971–90) 2994:(France) 2944:Dario Fo 963:31505304 937:FASEB J. 913:18445197 905:12068893 881:12181328 577:(UCLA), 562:(UCLA), 543:(UCLA), 524:(UCLA), 434:Chairman 295:(ATP)" ( 283:on the " 281:research 2978:Physics 2946:(Italy) 1494:of the 1191:June 9, 1157:June 5, 955:7737466 929:9674216 842:Member 669:Member 654:Fellow 467:subunit 375:atheist 299:) with 3632:(2021) 3626:(2020) 3620:(2019) 3614:(2018) 3608:(2017) 3602:(2016) 3596:(2015) 3590:(2014) 3584:(2013) 3578:(2012) 3572:(2011) 3566:(2010) 3560:(2009) 3554:(2008) 3548:(2007) 3542:(2006) 3536:(2005) 3530:(2004) 3524:(2003) 3518:(2002) 3512:(2001) 3506:(2000) 3500:(1999) 3494:(1998) 3488:(1997) 3482:(1996) 3476:(1995) 3470:(1994) 3464:(1993) 3458:(1992) 3452:(1991) 3446:(1990) 3440:(1989) 3434:(1988) 3428:(1987) 3422:(1986) 3416:(1985) 3410:(1984) 3404:(1983) 3398:(1982) 3392:(1981) 3386:(1980) 3380:(1979) 3374:(1978) 3368:(1977) 3362:(1976) 3358:& 3352:(1975) 3346:(1974) 3340:(1973) 3334:(1972) 3328:(1971) 3322:(1970) 3316:(1969) 3310:(1968) 3304:(1967) 3300:& 3294:(1966) 3288:(1965) 3282:(1964) 3276:(1963) 3270:(1962) 3264:(1961) 3258:(1960) 3252:(1959) 3246:(1958) 3240:(1957) 3234:(1956) 3228:(1955) 3222:(1954) 3216:(1953) 3210:(1952) 3204:(1951) 3198:(1950) 3192:(1949) 3186:(1948) 3180:(1947) 3174:(1946) 2839:2023: 2823:2022: 2811:2021: 2799:2020: 2783:2019: 2767:2018: 2751:2017: 2735:2016: 2719:2015: 2703:2014: 2687:2013: 2675:2012: 2667:2011: 2651:2010: 2635:2009: 2619:2008: 2611:2007: 2603:2006: 2587:2005: 2571:2004: 2559:2003: 2543:2002: 2527:2001: 2500:2000: 2492:1999: 2480:1998: 2464:1997: 2448:1996: 2432:1995: 2424:1994: 2412:1993: 2404:1992: 2396:1991: 2388:1990: 2376:1989: 2360:1988: 2344:1987: 2328:1986: 2316:1985: 2308:1984: 2300:1983: 2292:1982: 2280:1981: 2264:1980: 2252:1979: 2244:1978: 2236:1977: 2228:1976: 2205:1975: 2197:1974: 2185:1973: 2169:1972: 2161:1971: 2153:1970: 2141:1969: 2133:1968: 2117:1967: 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676:1974 661:1970 646:1968 626:1955 609:1955 604:Refs 593:Year 495:Death 391:Ph.D. 343:from 3109:2002 3104:2001 3099:2000 3094:1999 3089:1998 3079:1996 3074:1995 3069:1994 3064:1993 3059:1992 2937:1997 1876:1942 1869:1941 1862:1940 1795:1933 1696:1924 1657:1919 1642:1917 1635:1916 1419:2021 1372:2017 1346:2017 1316:2017 1290:2021 1262:2017 1226:2019 1193:2018 1182:ISSN 1159:2018 1132:2012 1106:2010 1067:2018 1041:2023 1013:2017 951:PMID 925:PMID 901:PMID 877:PMID 337:B.S. 305:Utah 279:for 236:UCLA 65:Died 43:Born 941:doi 893:doi 867:doi 863:277 339:in 291:of 3648:: 2847:/ 2843:/ 2831:/ 2827:/ 2815:/ 2803:/ 2791:/ 2787:/ 2775:/ 2771:/ 2759:/ 2755:/ 2743:/ 2739:/ 2727:/ 2723:/ 2711:/ 2707:/ 2695:/ 2691:/ 2679:/ 2659:/ 2655:/ 2643:/ 2639:/ 2627:/ 2623:/ 2595:/ 2591:/ 2579:/ 2575:/ 2563:/ 2551:/ 2547:/ 2535:/ 2531:/ 2508:/ 2504:/ 2484:/ 2472:/ 2468:/ 2456:/ 2452:/ 2440:/ 2436:/ 2416:/ 2380:/ 2368:/ 2364:/ 2352:/ 2348:/ 2336:/ 2332:/ 2320:/ 2284:/ 2272:/ 2268:/ 2256:/ 2209:/ 2189:/ 2177:/ 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Index


Provo, Utah
Los Angeles
Brigham Young University
University of Wisconsin–Madison
adenosine triphosphate
Pfizer Award in Enzyme Chemistry
Guggenheim Fellow
Tolman Award
Nobel Prize
Seaborg Medal
Chemistry
Biochemistry
Molecular biology
Stanford University
University of Minnesota
University of Minnesota
UCLA
UC
biochemist
analytical chemist
University of California, Los Angeles
Nobel Prize in Chemistry
research
enzymatic
biosynthesis
adenosine triphosphate
ATP synthase
John E. Walker
Utah

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