1455:
415:
698:
7767:
6534:
514:
7941:
5336:
150:
733:
7953:
1672:
772:
1584:. The model successfully flew with a combination of soaring and wing flapping. The model was based on a then-current weight estimate of around 80 kg (180 lb), far lower than more modern estimates of over 200 kg (440 lb). The method of flight in these pterosaurs depends largely on their weight, which has been controversial, and widely differing masses have been favored by different scientists. Some researchers have suggested that these animals employed slow,
5342:
950:
4719:
6550:
7774:
4724:
122:
4178:
1601:
7780:
1553:
4186:
1315:
4193:
6542:
674:
1629:. Henderson's work was also further criticized by Witton and Habib in another study, which pointed out that, although Henderson used excellent mass estimations, they were based on outdated pterosaur models, which caused Henderson's mass estimations to be more than double what Habib used in his estimations and that anatomical study of
1509:, catching fish during flight while cleaving the waves with its beak. While this skim-feeding view became widely accepted, it was not subjected to scientific research until 2007 when a study showed that, for such large pterosaurs, it was not a viable method because the energy costs would be too high due to excessive
1662:
had limited walking motion in its wings, with the limbs on each side of the body being moved together and the forelimbs being lifted out of the way of the hindlimbs. This study also suggests that the wings of pterosaurs were only attached to the body, with the legs and feet being tucked underneath,
1496:
In 1996, Lehman and
Langston rejected the scavenging hypothesis, pointing out that the lower jaw bent so strongly downwards that even when it closed completely a gap of over 5 cm (2.0 in) remained between it and the upper jaw, very different from the hooked beaks of specialized scavenging
1657:
was a very powerful flier, it had a large breastbone to support the necessary muscles for creating the flight stroke, and that it would probably have used its powerful hind legs to launch as high as 8 ft (2.4 m) when taking off, allowing it to gain enough height and momentum to begin the
1517:
published an examination of possible feeding habits and ecology of azhdarchids. Witton and Naish noted that most azhdarchid remains are found in inland deposits far from seas or other large bodies of water required for skimming. Additionally, the beak, jaw, and neck anatomy are unlike those of any
709:
More recent estimates based on greater knowledge of azhdarchid proportions place its wingspan at 10–11 m (33–36 ft). Remains found in Texas in 1971 indicate that this pterosaur had a minimum wingspan of about 11 m (36 ft). Generalized height in a bipedal stance, based on its
693:
fossils came from an individual with a wingspan as large as 15.9 m (52 ft). Choosing the middle of three extrapolations from the proportions of other pterosaurs gave an estimate of 11 m, 15.5 m, and 21 m, respectively (36 ft, 50.85 ft, 68.9 ft). In 1981, further advanced
452:
at the
University of Texas, Austin. He would then go on to become a doctoral student at the University of California, Berkeley. The specimen he discovered consisted of a partial wing (in pterosaurs composed of the forearms and elongated fourth finger) from an individual later estimated at over
713:
Body mass estimates for giant azhdarchids are extremely problematic because no existing species shares a similar size or body plan, and in consequence, published results vary widely. Generalized weight, based on some studies that have historically found extremely low weight estimates for
756:
had a very sharp and pointed beak. That is contrary to some earlier reconstructions that showed a blunter snout, based on the inadvertent inclusion of jaw material from another pterosaur species, as this material was named as the holotype of a genus of short-snouted azhdarchid,
525:
At first, it was assumed that the smaller specimens were juvenile or subadult forms of the larger type. Later, when more remains were found, it was realized they could have been a separate species. This possible second species from Texas was provisionally referred to as a
1522:, and that they probably hunted small vertebrates on land or in small streams. Additionally, Quetzalcoatlus had the ability to turn its head 180 degrees which would allow it to be more adept at finding prey and preventing ambushes from foes. Though
1620:, and Mark Witton, a British paleontologist, undertook further investigation into the claims of flightlessness in large pterosaurs. After factoring wingspan, body weight, and aerodynamics, computer modeling led the two researchers to conclude that
1750:'s 350th-anniversary exhibition. The models, which included both flying and standing individuals with wingspans of over 10 m (33 ft), were intended to help build public interest in science. The models were created by scientists from the
1633:
and other big pterosaur forelimbs showed a higher degree of robustness than would be expected if they were purely quadrupedal. This study proposed that large pterosaurs most likely utilized a short burst of powered flight to then transition to
1439:
had precursors in North
America and its apparent rise to widespreadness may represent the expansion of its preferred habitat rather than an immigration event, as some experts have suggested. It co-existed with another azhdarchid known as
1470:. Because the area of the fossil site was 400 km (250 mi) removed from the coastline and there were no indications of large rivers or deep lakes nearby at the end of the Cretaceous, Lawson, in 1975, rejected a
1730:
incorporates a "flight control system/autopilot which processes pilot commands and sensor inputs, implements several feedback loops, and delivers command signals to its various servo-actuators". It is on exhibit at the
1624:
was capable of flight up to 130 km/h (80 mph) for 7 to 10 days at altitudes of 4,600 m (15,000 ft). Habib further suggested a maximum flight range of 13,000–19,000 km (8,000–12,000 mi) for
1592:
had been underestimated, even the highest estimates, and that it was too massive to have achieved powered flight. He estimated it in his 2010 paper as 540 kg (1,190 lb). Henderson argued that it may have been
718:, was as low as 70 kg (150 lb) for a 10 m (32 ft 10 in) individual. A majority of estimates published since the 2000s have been substantially higher, around 200–250 kg (440–550 lb).
1615:
Other flight capability estimates have disagreed with
Henderson's research, suggesting instead an animal superbly adapted to long-range, extended flight. In 2010, Mike Habib, a professor of biomechanics at
654:, the vertebra shows no evidence that it was chewed on by the dinosaur. The bone came from an individual azhdarchid pterosaur estimated to have had a wingspan of 5–5.5 m (16–18 ft).
561:—represent elements which are typically considered undiagnostic to generic or specific level and that this complicates interpretations of azhdarchid taxonomy. For instance, Witton
2994:
1699:. Over the next twenty-five years prior to future discoveries, it would launch similar depictions colloquially known as "paleomemes" in various books, as noted by Darren Naish.
1718:(UAV). They produced a half-scale model weighing 18 kg (40 lb), with a wingspan of 5.5 m (18 ft). Coincidentally, Douglas A. Lawson, who discovered
1454:
609:
differ enough that they cannot be regarded as the same animal, but that the significance of this cannot be ascertained, given uncertainty over the relationships of
3013:
625:, could have made long, transcontinental flights, suggesting that locations as disparate as North America and Europe could have shared giant azhdarchid species.
569:
type material is of generalized enough morphology to be near identical to that of other giant azhdarchids, such as the overlapping elements of the contemporary
2770:
Lawson 1975 (Pterodactyloidea: Azhdarchoidea): A paleoenvironmental reconstruction of the
Javelina Formation (Upper Cretaceous), Big Bend National Park, Texas"
538:
holotype, and include four partial skulls, though they are much less massive, with an estimated wingspan of 5.5 m (18 ft). This species was named
7978:
534:
and
Langston in 1996, indicating that its status was too uncertain to give it a full new species name. The smaller specimens are more complete than the
1493:. Lawson had found the remains of the giant pterosaur while searching for the bones of this dinosaur, which formed an important part of its ecosystem.
3200:
7885:
1568:
and other giant azhdarchids was poorly understood until serious biomechanical studies were conducted in the 21st century. A 1984 experiment by
725:
would have weighed 20 kg (44 lb) for individuals with a wingspan of 4.5 m (15 ft). In 2022, Gregory S. Paul estimated that
460:
Lawson discovered a second site of the same age, about 40 km (25 mi) from the first, where between 1972 and 1974, he and
Professor
1899:
414:
402:, which was known for many years as an unnamed species, before being named by Brian Andres and Wann Langston Jr. (posthumously) in 2021.
2376:"On the size and flight diversity of giant pterosaurs, the use of birds as pterosaur analogues and comments on pterosaur flightlessness"
2640:: paleoenvironmental reconstruction of the Javelina Formation (Upper Cretaceous), Big Bend National Park, Texas". Abstracts of Papers.
1995:
Lawson, Douglas A. (1975). "Pterosaur from the Latest
Cretaceous of West Texas: Discovery of the largest flying creature". Reports.
7993:
7988:
7859:
7998:
2897:
644:. The specimen, BMR P2002.2, was recovered accidentally when it was included in a field jacket prepared to transport part of a
3101:
1588:
flight, while others have concluded that their flight was fast and dynamic. In 2010, Donald
Henderson argued that the mass of
3179:
2326:
729:
had a wingspan of 5 m (16 ft), body length of 3.5 m (11 ft), and body mass of 65 kg (143 lb).
3043:
2587:"Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary"
1518:
known skimming animal. Rather, they concluded that azhdarchids were more likely terrestrial stalkers, similar to modern
1446:, as well as an additional pterosaur taxon, suggesting a relatively high diversity of Late Cretaceous pterosaur genera.
3021:
468:
unearthed three fragmentary skeletons of much smaller individuals. Lawson in 1975 announced the find in an article in
2506:
2355:
1638:. However, a study from 2022 suggests that they would only have flown occasionally and for short distances, like the
137:
7766:
1859:
Lehman, Thomas M.; Mcdowell, Fred W.; Connelly, James N. (2006). "First isotopic (U-Pb) age for the Late
Cretaceous
621:
being investigated. An additional complication to these discussions is the likelihood that huge pterosaurs, such as
474:. That same year, in a subsequent letter to the same journal, he made the original large specimen, TMM 41450-3, the
2975:
2927:"How did extinct giant birds and pterosaurs fly? A comprehensive modeling approach to evaluate soaring performance"
2229:"An azhdarchid pterosaur cervical vertebra from the Hell Creek Formation (Maastrichtian) of southeastern Montana"
1768:
5341:
2247:
1876:
1732:
449:
2669:"Did pterosaurs feed by skimming? Physical modelling and anatomical evaluation of an unusual feeding method"
1482:(which will scavenge, but is more of a terrestrial predator of small animals), but then on the carcasses of
7983:
1534:
mammals than to their smaller cousins, implying that they were uniquely suited to a terrestrial lifestyle.
1292:
1103:
149:
6533:
5335:
1268:
7931:
2570:
Lehman, T. M. (2001). "Late Cretaceous dinosaur provinciality". In Tanke, D. H.; Carpenter, K. (eds.).
4718:
3172:
557:
and colleagues (2010) noted that the type species of the genus—the fragmentary wing bones comprising
2318:
2312:
796:, a group of advanced toothless pterosaurs with unusually long, stiffened necks and giant wingspans
7837:
3276:
1763:
1751:
1544:
is found in great numbers in alkaline lakes, and likely lived like modern gregarious wading birds.
944:
890:
4177:
3073:
4754:
4185:
1715:
1195:
1079:
999:
549:
had not been properly described and diagnosed until 2021 and until then, the status of the genus
491:
4723:
7916:
7799:
7674:
5832:
1127:
441:
7890:
7773:
2104:(Pterosauria, Azhdarchidae) from Late Cretaceous sediments of Big Bend National Park, Texas".
1863:
vertebrate fauna of West Texas, and its significance as a link between two faunal provinces".
697:
8003:
7911:
7903:
7779:
1224:
849:
465:
2995:"Quetzalcoatlus: the evil, pin-headed, toothy nightmare monster that wants to eat your soul"
2818:
1642:(the world's heaviest bird that actively flies) and that they were not able to soar at all.
7973:
7957:
7846:
6541:
5927:
4366:
4276:
4192:
3165:
2862:
2781:
2730:
2453:
2387:
2278:
2157:
2113:
2055:
2004:
1946:
1815:
1608:
637:
499:
6549:
8:
7945:
7431:
7120:
6807:
6106:
6033:
3801:
3521:
2148:
1530:
and other azhdarchids have fore and hind limb proportions more similar to modern running
966:
873:
763:, in 2021. A skull crest was also present but its exact form and size are still unknown.
2866:
2785:
2734:
2457:
2391:
2282:
2206:"Clipping the wings of giant pterosaurs: comments on wingspan estimations and diversity"
2161:
2117:
2059:
2008:
1950:
1819:
689:
When it was first named as a new species in 1975, scientists estimated that the largest
6960:
5434:
5277:
4420:
4223:
4210:
4124:
3966:
3597:
3572:
2951:
2926:
2878:
2799:
2746:
2695:
2668:
2613:
2586:
2547:
2512:
2471:
2410:
2375:
2344:
2294:
2290:
2251:
2181:
2079:
2028:
1969:
1934:
1880:
1841:
1617:
1144:
437:
144:
2847:
2228:
2140:
7898:
7386:
6201:
6130:
5372:
5359:
5302:
5065:
4963:
4865:
4783:
4561:
4538:
4504:
4434:
3975:
3891:
3843:
3817:
3530:
3435:
2956:
2803:
2700:
2618:
2516:
2502:
2475:
2415:
2351:
2322:
2255:
2205:
2173:
2071:
2020:
1974:
1884:
1845:
1833:
1064:
1016:
531:
470:
461:
445:
379:
253:
2882:
2750:
2551:
2185:
2083:
2032:
7615:
7498:
7490:
7282:
7211:
7136:
6678:
6581:
6568:
6500:
6380:
6369:
5619:
5476:
5413:
5188:
5170:
5008:
4832:
4648:
4634:
4592:
4098:
4084:
4016:
3695:
3245:
2946:
2938:
2870:
2789:
2738:
2690:
2680:
2667:
Humphries, Stuart; Bonser, Richard H. C.; Witton, Mark P; Martill, David M (2007).
2653:
2649:
2608:
2598:
2539:
2494:
2461:
2405:
2395:
2286:
2243:
2165:
2125:
2121:
2063:
2012:
1964:
1954:
1872:
1823:
1696:
922:
834:
629:
was found to be a valid taxon in 2021 and confirmed to belong to the same genus as
317:
233:
207:
2794:
2765:
2466:
2437:
1828:
1799:
513:
392:, named by Douglas Lawson in 1975 after the tailless fixed-wing aircraft designer
7704:
7547:
7396:
7181:
7085:
7060:
7053:
7040:
6973:
6859:
6757:
6723:
6619:
6599:
6453:
6311:
6252:
6239:
6210:
6086:
5997:
5856:
5729:
5684:
5556:
5485:
5464:
5290:
4951:
4931:
4903:
4879:
4815:
4807:
4800:
4792:
4547:
4527:
4357:
4333:
4283:
4091:
4074:
4064:
4041:
3997:
3989:
3957:
3946:
3925:
3879:
3871:
3810:
3726:
3666:
3585:
3554:
3444:
3423:
3376:
3261:
3236:
2902:
2685:
2603:
2400:
2141:"A new giant pterosaur with a robust skull from the latest Cretaceous of Romania"
2067:
2016:
1959:
1594:
372:
30:
2942:
2721:
Padian, K. (1983). "A functional analysis of flying and walking in pterosaurs".
710:
wingspan, would have been at least 3 m (9.8 ft) high at the shoulder.
7681:
7654:
7632:
7456:
7370:
7355:
7307:
7246:
7231:
7197:
7022:
7006:
6994:
6980:
6933:
6890:
6850:
6764:
6710:
6438:
6424:
6396:
6303:
6224:
6151:
6062:
5818:
5710:
5645:
5586:
5526:
5492:
5212:
5139:
5111:
5030:
4980:
4919:
4888:
4488:
4464:
4442:
4388:
4346:
4290:
4258:
3939:
3856:
3833:
3712:
3638:
3606:
3459:
3408:
3356:
3051:
1663:
much like how modern birds tuck their legs beneath their own bodies in flight.
1635:
1585:
1569:
1510:
1506:
781:
668:
3146:
3134:
3122:
3110:
2874:
2742:
2585:
Longrich, Nicholas R.; Martill, David M.; Andres, Brian; Penny, David (2018).
2543:
2532:
Earth and Environmental Science Transactions of the Royal Society of Edinburgh
2169:
1935:"A Reappraisal of azhdarchid pterosaur functional morphology and paleoecology"
406:
has gained fame as a candidate for the largest flying animal ever discovered.
7967:
7822:
7718:
7689:
7665:
7646:
7604:
7589:
7538:
7527:
7420:
7412:
7362:
7299:
7270:
7254:
7238:
7224:
7165:
7143:
7094:
6987:
6748:
6691:
6610:
6445:
6417:
6389:
6260:
6143:
5934:
5864:
5736:
5719:
5575:
5512:
5456:
5221:
5177:
5058:
5051:
5044:
5037:
5022:
4972:
4896:
4853:
4846:
4772:
4619:
4612:
4605:
4454:
4396:
4381:
4374:
4322:
4105:
3774:
3734:
3684:
3653:
3614:
3509:
3501:
3479:
3383:
3367:
2925:
Goto, Yusuke; Yoda, Ken; Weimerskirch, Henri; Sato, Katsufumi (May 2, 2022).
2346:
Dinosaurs of the Air: The Evolution and Loss of Flight in Dinosaurs and Birds
1837:
1747:
1723:
1707:
1680:
1658:
necessary downstrokes needed for takeoff. This same study also suggests that
1479:
1442:
815:
759:
732:
646:
575:
495:
434:
393:
375:
72:
34:
1726:, a developer of tailless flying wing aircraft in the 1940s. The replica of
1466:
There have been a number of different ideas proposed about the lifestyle of
7711:
7639:
7510:
7405:
7339:
7291:
7101:
7029:
6907:
6875:
6643:
6410:
6338:
6279:
6174:
5811:
5796:
5780:
5771:
5760:
5600:
5563:
5449:
5399:
5242:
5155:
5148:
5128:
5100:
5093:
5079:
5072:
4987:
4944:
4911:
4627:
4569:
4512:
4497:
4477:
4302:
4033:
4025:
4005:
3932:
3911:
3904:
3759:
3547:
3493:
3452:
2960:
2848:"Pterosaur body mass estimates from three-dimensional mathematical slicing"
2704:
2622:
2419:
2177:
2075:
2024:
1978:
1671:
1639:
1514:
1209:
793:
771:
650:
specimen. Despite this association with the remains of a large carnivorous
636:
An azhdarchid neck vertebra, discovered in 2002 from the Maastrichtian age
585:
487:
383:
365:
262:
220:
2976:"Vintage Dinosaur Art: The evolution and ecology of the Dinosaurs: Part 2"
2498:
378:
age of North America. Its name comes from the Aztec feathered serpent god
7872:
7831:
7597:
7557:
7476:
7440:
7174:
6949:
6882:
6825:
6771:
6628:
6296:
6167:
6159:
6119:
6078:
6070:
5945:
5912:
5880:
5872:
5843:
5804:
5788:
5747:
5676:
5668:
5661:
5607:
5548:
5540:
5201:
5163:
5086:
4741:
4674:
4641:
4577:
3918:
3787:
3742:
3719:
3394:
3074:"Pterosaurs Coming to London to Celebrate 350 Years of the Royal Society"
2100:
Kellner, Alexander W. A.; Langston, Wann Jr. (1996). "Cranial remains of
1577:
1423:
554:
503:
47:
2298:
1653:
published in 2021 by Kevin Padian and colleagues instead suggested that
7864:
7565:
7520:
7448:
7323:
7315:
7150:
7129:
7074:
6942:
6917:
6867:
6832:
6778:
6666:
6658:
6651:
6403:
6326:
6318:
6217:
6181:
6019:
5920:
5899:
5888:
5825:
5442:
5235:
5228:
4663:
4554:
3561:
3218:
2636:
Lehman, Thomas M.; Langston, Wann Jr. (1996). "Habitat and behavior of
1743:
1676:
1483:
92:
57:
7877:
1900:"Fleshing out the bones of Quetzalcoatlus, Earth's largest flier ever"
949:
7625:
7067:
7015:
6737:
6730:
6698:
6636:
6362:
6355:
6347:
6267:
6045:
6011:
5961:
5954:
5691:
5629:
5533:
5121:
4520:
4237:
3864:
3704:
3539:
3416:
3338:
3325:
3288:
3227:
3188:
1581:
1486:
1471:
800:
368:
194:
161:
121:
97:
41:
7793:
1691:
as a small-headed scavenger with an extremely long neck in the book
1600:
7816:
7697:
7581:
7573:
7347:
7331:
7262:
6839:
6189:
6094:
6026:
5698:
5653:
5426:
4655:
4315:
3212:
2436:
Padian, K.; Cunningham, J.R.; Langston, W. Jr.; Conway, J. (2021).
1552:
1540:
is found in plains deposits, being likely a solitary hunter, while
1531:
1498:
1314:
651:
475:
454:
181:
87:
82:
67:
62:
52:
1418:
694:
studies lowered these estimates to 11–12 m (36–39 ft).
583:
can be distinguished from other pterosaurs (i.e., if it is not a
570:
419:
102:
77:
7851:
6288:
6055:
5637:
4245:
4112:
3206:
617:
being demonstrated as a valid taxon and its relationships with
483:
171:
3157:
2435:
673:
332:
5386:
3982:
2046:
Lawson, Douglas A. (1975). "Could pterosaurs fly?". Letters.
1703:
1526:, like other pterosaurs, was a quadruped when on the ground,
1519:
430:
2666:
2374:
Witton, Mark P.; Habib, Michael B.; Laudet, Vincent (2010).
1702:
In 1985, the US Defense Advanced Research Projects Agency (
353:
338:
323:
2584:
2220:
2095:
2093:
2530:
Andres, B.; Myers, T. S. (2013). "Lone Star pterosaurs".
2248:
10.1671/0272-4634(2006)26[192:AAPCVF]2.0.CO;2
1877:
10.1671/0272-4634(2006)26[922:FIUAFT]2.0.CO;2
1435:
association probably represents semi-arid inland plains.
344:
2924:
2138:
721:
In 2021, Kevin Padian and his colleagues estimated that
2629:
2090:
16:
Genus of azhdarchid pterosaurs from the Late Cretaceous
4752:
2203:
1928:
1926:
1924:
1922:
1920:
1858:
7929:
2898:"Peerless pterosaur could fly long-distance for days"
2204:
Witton, M.P.; Martill, D.M.; Loveridge, R.F. (2010).
356:
335:
329:
320:
3044:"UAS Advanced Development: Quetzalcoatlus Northropi"
542:
in 2021, named after the genus' original describer.
350:
347:
326:
292:
269:
2369:
2367:
2227:Henderson, Michael D.; Peterson, Joseph E. (2006).
1917:
1576:. MacCready constructed a model flying machine or,
807:within Neoazhdarchia from Andres and Myers (2013).
448:, who was then a geology graduate student from the
341:
3800:
2343:
1572:used practical aerodynamics to test the flight of
444:(dated to around 68 million years ago) in 1971 by
5276:
4221:
2660:
2373:
2226:
2139:Buffetaut, E.; Grigorescu, D.; Csiki, Z. (2002).
7965:
5370:
2841:
2839:
2493:. Princeton University Press. pp. 161–162.
2364:
2132:
613:specimens. These issues can only be resolved by
6579:
6499:
5412:
2350:. Johns Hopkins University Press. p. 472.
2099:
1497:birds. They suggested that, with its long neck
593:should be regarded as a European occurrence of
3260:
2635:
2442:Lawson 1975 (Pterodactyloidea: Azhdarchoidea)"
1804:Lawson 1975 (Pterodactyloidea: Azhdarchoidea)"
1797:
502:, who drove the development of large tailless
396:. The genus also includes the smaller species
3173:
3036:
2836:
2810:
2716:
2714:
2574:. Indiana University Press. pp. 310–328.
2565:
2563:
2561:
2431:
2429:
1714:as the basis for an experimental ornithopter
1513:. In 2008, pterosaur workers Mark Witton and
7164:
2578:
2262:
2199:
2197:
2195:
1990:
1988:
1793:
1791:
1789:
1787:
1785:
1783:
7979:Late Cretaceous pterosaurs of North America
6278:
3005:
2529:
1932:
1580:, with a simple computer functioning as an
3336:
3180:
3166:
2720:
2711:
2558:
2523:
2426:
2039:
1693:The evolution and ecology of the Dinosaurs
1687:In 1975, artist Giovanni Caselli depicted
120:
3066:
2986:
2950:
2845:
2816:
2793:
2694:
2684:
2612:
2602:
2465:
2409:
2399:
2269:Langston, Wann Jr. (1981). "Pterosaurs".
2192:
1985:
1968:
1958:
1852:
1827:
1780:
1607:skeleton mounted in a flying pose at the
3147:"Going out on a limb for Quetzalcoatlus"
3011:
2967:
2335:
2317:. The Jim Pattison Group. 2014. p.
2268:
1670:
1666:
1599:
1551:
1453:
779:cervical vertebrae on the left with the
770:
731:
696:
672:
579:. This being the case and assuming that
512:
413:
409:
2999:Tetrapod Zoology at Scientific American
2973:
2895:
2889:
2491:The Princeton Field Guide to Pterosaurs
1933:Witton, Mark P.; Naish, Darren (2008).
1897:
7966:
4209:
2763:
2569:
2045:
1994:
1738:In 2010, several life-sized models of
1474:lifestyle. Instead, he suggested that
803:showing the phylogenetic placement of
601:also noted that the skull material of
7798:
7797:
6566:
5358:
5357:
4739:
4208:
3323:
3199:
3161:
3050:. AeroVironment. 2011. Archived from
3012:Jacobsen, Annie (February 27, 2011).
2992:
1798:Andres, B.; Langston, W. Jr. (2021).
6567:
2488:
2341:
1898:Sanders, Robert (December 8, 2021).
3014:"When Flying Dinosaurs Were Drones"
2896:Frazier, Reid (November 22, 2010).
1746:as the centerpiece exhibit for the
13:
3123:"The Biggest Thing That Ever Flew"
3117:. Paleo Studio. December 29, 2013.
3111:"Quetzalcoatlus - Flying monsters"
2855:Journal of Vertebrate Paleontology
2774:Journal of Vertebrate Paleontology
2642:Journal of Vertebrate Paleontology
2446:Journal of Vertebrate Paleontology
2291:10.1038/scientificamerican0281-122
2236:Journal of Vertebrate Paleontology
2106:Journal of Vertebrate Paleontology
1865:Journal of Vertebrate Paleontology
1808:Journal of Vertebrate Paleontology
1458:Artist's impression of a group of
14:
8015:
3135:"Quetzalcoatlus - Ancient Animal"
3095:
2993:Naish, Darren (August 20, 2013).
1722:in Texas in 1971, named it after
1417:was abundant in Texas during the
766:
705:with a Cessna 172 light aircraft.
138:Houston Museum of Natural Science
7951:
7939:
7778:
7772:
7765:
6548:
6540:
6532:
5340:
5334:
4722:
4717:
4191:
4184:
4176:
2766:"Habitat of the giant pterosaur
1742:were put on display on London's
1313:
948:
316:
148:
135:displayed in quadrupedal stance,
45:
7994:Taxa named by Douglas A. Lawson
7989:Maastrichtian genus extinctions
4740:
3324:
3187:
2980:Love in the Time of Chasmosaurs
2918:
2819:"The Great Pterodactyl Project"
2757:
2482:
2305:
1683:'s 350th anniversary exhibition
1409:
553:was identified as problematic.
482:. The genus name refers to the
3153:. thebrainscoop. June 9, 2020.
2974:Vincent, Mark (May 14, 2012).
2654:10.1080/02724634.1996.10011371
2126:10.1080/02724634.1996.10011310
1891:
1769:Timeline of pterosaur research
677:Size comparison of a human to
657:
1:
7999:Fossil taxa described in 1975
3141:. North 02. October 25, 2019.
2846:Henderson, Donald M. (2010).
2795:10.1080/02724634.2019.1593184
2467:10.1080/02724634.2020.1780247
1829:10.1080/02724634.2021.1907587
1774:
1733:National Air and Space Museum
450:Jackson School of Geosciences
301:Andres and Langston Jr., 2021
3129:. PBS Eons. August 14, 2017.
2686:10.1371/journal.pbio.0050204
2604:10.1371/journal.pbio.2001663
2401:10.1371/journal.pone.0013982
2068:10.1126/science.188.4189.676
2017:10.1126/science.187.4180.947
1960:10.1371/journal.pone.0002271
1800:"Morphology and taxonomy of
1293:Arambourgiania philadelphiae
1104:Shenzhoupterus chaoyangensis
506:aircraft designs resembling
478:of a new genus and species,
7:
2314:Guinness World Records 2014
1757:
1269:Zhejiangopterus linhaiensis
517:Skeletal reconstruction of
429:fossils were discovered in
10:
8020:
3018:Los Angeles Times Magazine
2764:Lehman, Thomas M. (2021).
2438:"Functional morphology of
1695:by British paleontologist
1478:scavenged, similar to the
1449:
666:
565:(2010) suggested that the
453:10 m (33 ft) in
7806:
7762:
7664:
7614:
7546:
7537:
7509:
7485:
7474:
7430:
7395:
7385:
7281:
7219:
7210:
7206:
7195:
7160:
7119:
7084:
7048:
7039:
7005:
6968:
6959:
6932:
6928:
6905:
6849:
6820:
6816:
6805:
6747:
6718:
6709:
6686:
6677:
6618:
6609:
6594:
6590:
6575:
6562:
6529:
6495:
6379:
6337:
6247:
6238:
6200:
6138:
6129:
6114:
6105:
6044:
6006:
5995:
5944:
5907:
5898:
5851:
5842:
5770:
5755:
5746:
5718:
5709:
5618:
5595:
5584:
5573:
5521:
5510:
5475:
5421:
5408:
5397:
5381:
5366:
5353:
5331:
5301:
5285:
5272:
5211:
5196:
5187:
5138:
5110:
5017:
5006:
4962:
4939:
4930:
4878:
4874:
4863:
4841:
4830:
4782:
4767:
4763:
4748:
4735:
4715:
4673:
4600:
4591:
4537:
4487:
4472:
4463:
4452:
4429:
4418:
4356:
4341:
4332:
4310:
4301:
4271:
4267:
4256:
4232:
4217:
4204:
4173:
4123:
4073:
4063:
4015:
3965:
3956:
3899:
3890:
3851:
3842:
3831:
3796:
3785:
3769:
3757:
3694:
3679:
3675:
3664:
3648:
3636:
3596:
3580:
3571:
3529:
3520:
3488:
3477:
3434:
3403:
3393:
3366:
3351:
3347:
3332:
3319:
3287:
3269:
3256:
3195:
2943:10.1093/pnasnexus/pgac023
2875:10.1080/02724631003758334
2826:Engineering & Science
2743:10.1017/S009483730000765X
2544:10.1017/S1755691013000303
2489:Paul, Gregory S. (2022).
2342:Paul, Gregory S. (2002).
2170:10.1007/s00114-002-0307-1
1547:
1501:and long toothless jaws,
1328:
1306:
1289:
1282:
1265:
1258:
1245:
1238:
1221:
1214:
1192:
1185:
1141:
1124:
1117:
1100:
1093:
1076:
1069:
1057:
1013:
996:
989:
963:
941:
934:
927:
887:
870:
863:
846:
839:
827:
820:
545:The holotype specimen of
289:
284:
268:
261:
145:Scientific classification
143:
128:
119:
23:
2572:Mesozoic Vertebrate Life
2210:Acta Geoscientica Sinica
1764:List of pterosaur genera
1752:University of Portsmouth
1712:Quetzalcoatlus northropi
1564:The nature of flight in
1538:Quetzalcoatlus northropi
1421:in a fauna dominated by
1310:Quetzalcoatlus northropi
945:Domeykodactylus ceciliae
891:Tupuxuara longicristatus
743:
736:Skull reconstruction of
480:Quetzalcoatlus northropi
274:Quetzalcoatlus northropi
4755:Archaeopterodactyloidea
1716:unmanned aerial vehicle
1196:Radiodactylus langstoni
1080:Eoazhdarcho liaoxiensis
1000:Noripterus complicidens
662:
486:feathered serpent god,
2817:MacCready, P. (1985).
1684:
1612:
1561:
1463:
1332:Quetzalcoatlus lawsoni
1128:Chaoyangopterus zhangi
792:belongs to the family
786:
740:
706:
686:
522:
442:Big Bend National Park
422:
7912:Paleobiology Database
2499:10.1515/9780691232218
1674:
1667:Cultural significance
1603:
1555:
1462:feeding on the ground
1457:
1225:Azhdarcho lancicollis
850:Thalassodromeus sethi
785:holotype on the right
774:
735:
700:
676:
640:, may also belong to
516:
496:John Knudsen Northrop
466:Texas Memorial Museum
418:Cast of the holotype
417:
410:Discovery and species
4367:Archaeoistiodactylus
4277:Archaeoistiodactylus
3106:at EarthArchives.org
1724:John "Jack" Northrop
1609:Royal Ontario Museum
1505:fed like modern-day
748:Skull material from
638:Hell Creek Formation
500:Northrop Corporation
7984:Ojo Alamo Formation
7432:Targaryendraconidae
7121:Ornithocheiriformes
6808:Ornithocheiromorpha
6107:Neopterodactyloidea
6034:Dsungaripteromorpha
3802:Campylognathoididae
3522:Eudimorphodontoidea
3078:Everything Dinosaur
2867:2010JVPal..30..768H
2786:2021JVPal..41S..21L
2735:1983Pbio....9..218P
2458:2021JVPal..41S.218P
2392:2010PLoSO...513982W
2283:1981SciAm.244b.122L
2271:Scientific American
2162:2002NW.....89..180B
2149:Naturwissenschaften
2118:1996JVPal..16..222K
2060:1975Sci...188..676G
2009:1975Sci...187..947L
1951:2008PLoSO...3.2271W
1820:2021JVPal..41S..46A
1675:Models on London's
967:Dsungaripterus weii
874:Tupuxuara leonardii
6961:Istiodactyliformes
5435:Lonchognathosaurus
5278:Eupterodactyloidea
4421:Pterodactyliformes
4224:Pterodactylomorpha
4211:Pterodactylomorpha
4125:Pterodactylomorpha
3967:Angustinaripterini
3598:Eudimorphodontinae
3573:Eudimorphodontidae
3054:on January 7, 2012
1685:
1618:Chatham University
1613:
1562:
1464:
1145:Jidapterus edentus
787:
741:
707:
687:
597:. However, Witton
523:
438:Javelina Formation
423:
7927:
7926:
7899:Open Tree of Life
7800:Taxon identifiers
7791:
7790:
7787:
7786:
7760:
7759:
7756:
7755:
7752:
7751:
7748:
7747:
7744:
7743:
7740:
7739:
7736:
7735:
7732:
7731:
7728:
7727:
7470:
7469:
7466:
7465:
7387:Targaryendraconia
7381:
7380:
7191:
7190:
7115:
7114:
7111:
7110:
6901:
6900:
6801:
6800:
6797:
6796:
6793:
6792:
6789:
6788:
6558:
6557:
6527:
6526:
6523:
6522:
6519:
6518:
6491:
6490:
6487:
6486:
6483:
6482:
6479:
6478:
6475:
6474:
6471:
6470:
6467:
6466:
6463:
6462:
6234:
6233:
6202:Chaoyangopterinae
6131:Chaoyangopteridae
5991:
5990:
5987:
5986:
5983:
5982:
5979:
5978:
5975:
5974:
5971:
5970:
5506:
5505:
5502:
5501:
5373:Ornithocheiroidea
5360:Ornithocheiroidea
5349:
5348:
5329:
5328:
5325:
5324:
5321:
5320:
5303:Ornithocheiroidea
5268:
5267:
5264:
5263:
5260:
5259:
5256:
5255:
5252:
5251:
5066:Gladocephaloideus
5002:
5001:
4998:
4997:
4964:Aurorazhdarchidae
4866:Ctenochasmatoidea
4826:
4825:
4784:Germanodactylidae
4731:
4730:
4713:
4712:
4709:
4708:
4705:
4704:
4701:
4700:
4697:
4696:
4693:
4692:
4587:
4586:
4562:Dendrorhynchoides
4539:Batrachognathinae
4505:Dendrorhynchoides
4435:Changchengopterus
4414:
4413:
4410:
4409:
4406:
4405:
4200:
4199:
4171:
4170:
4167:
4166:
4163:
4162:
4159:
4158:
4155:
4154:
4151:
4150:
4147:
4146:
4143:
4142:
4059:
4058:
4055:
4054:
4051:
4050:
3976:Angustinaripterus
3892:Rhamphorhynchinae
3844:Rhamphorhynchidae
3827:
3826:
3818:Campylognathoides
3753:
3752:
3632:
3631:
3628:
3627:
3624:
3623:
3531:Raeticodactylidae
3473:
3472:
3469:
3468:
3436:Austriadraconidae
3315:
3314:
3311:
3310:
3307:
3306:
2452:(sup1): 218–251.
2328:978-1-908843-15-9
2054:(4189): 676–677.
2003:(4180): 947–948.
1556:Restoration of a
1406:
1405:
1397:
1396:
1388:
1387:
1379:
1378:
1370:
1369:
1361:
1360:
1352:
1351:
1343:
1342:
1174:
1173:
1165:
1164:
1156:
1155:
1065:Chaoyangopteridae
1046:
1045:
1037:
1036:
1028:
1027:
1017:Noripterus parvus
978:
977:
911:
910:
902:
901:
647:Tyrannosaurus rex
532:Alexander Kellner
498:, the founder of
462:Wann Langston Jr.
446:Douglas A. Lawson
308:
307:
302:
257:
8011:
7956:
7955:
7954:
7944:
7943:
7942:
7935:
7920:
7919:
7907:
7906:
7894:
7893:
7881:
7880:
7868:
7867:
7855:
7854:
7842:
7841:
7840:
7827:
7826:
7825:
7795:
7794:
7782:
7776:
7770:
7769:
7616:Coloborhynchinae
7544:
7543:
7499:Ornithocheiridae
7491:Brasileodactylus
7483:
7482:
7393:
7392:
7283:Ornithocheirinae
7217:
7216:
7212:Ornithocheiridae
7208:
7207:
7204:
7203:
7162:
7161:
7158:
7157:
7137:Brasileodactylus
7046:
7045:
6966:
6965:
6934:Lonchodraconidae
6930:
6929:
6926:
6925:
6818:
6817:
6814:
6813:
6716:
6715:
6684:
6683:
6679:Nyctosauromorpha
6616:
6615:
6592:
6591:
6588:
6587:
6582:Pteranodontoidea
6577:
6576:
6569:Pteranodontoidea
6564:
6563:
6553:
6552:
6545:
6544:
6537:
6536:
6501:Pteranodontoidea
6497:
6496:
6381:Quetzalcoatlinae
6370:Mistralazhdarcho
6335:
6334:
6276:
6275:
6245:
6244:
6136:
6135:
6112:
6111:
6004:
6003:
5905:
5904:
5849:
5848:
5768:
5767:
5753:
5752:
5716:
5715:
5620:Thalassodromidae
5593:
5592:
5582:
5581:
5519:
5518:
5477:Dsungaripterinae
5419:
5418:
5414:Dsungaripteridae
5410:
5409:
5406:
5405:
5379:
5378:
5368:
5367:
5355:
5354:
5344:
5339:
5338:
5283:
5282:
5274:
5273:
5194:
5193:
5189:Ctenochasmatinae
5171:Plataleorhynchus
5015:
5014:
5009:Ctenochasmatidae
4937:
4936:
4876:
4875:
4872:
4871:
4839:
4838:
4833:Euctenochasmatia
4765:
4764:
4761:
4760:
4750:
4749:
4737:
4736:
4726:
4721:
4649:Propterodactylus
4635:Ningchengopterus
4598:
4597:
4593:Pterodactyloidea
4470:
4469:
4461:
4460:
4427:
4426:
4339:
4338:
4308:
4307:
4269:
4268:
4265:
4264:
4230:
4229:
4219:
4218:
4206:
4205:
4195:
4189:
4188:
4181:
4180:
4099:Jianchangopterus
4085:Fenghuangopterus
4071:
4070:
4017:Rhamphorhynchini
3963:
3962:
3897:
3896:
3849:
3848:
3840:
3839:
3798:
3797:
3794:
3793:
3767:
3766:
3696:Dimorphodontidae
3677:
3676:
3673:
3672:
3646:
3645:
3578:
3577:
3527:
3526:
3486:
3485:
3401:
3400:
3349:
3348:
3345:
3344:
3334:
3333:
3321:
3320:
3267:
3266:
3258:
3257:
3251:
3250:
3246:Pterosauromorpha
3197:
3196:
3182:
3175:
3168:
3159:
3158:
3154:
3142:
3130:
3118:
3090:
3089:
3087:
3085:
3070:
3064:
3063:
3061:
3059:
3040:
3034:
3033:
3031:
3029:
3024:on March 8, 2013
3020:. Archived from
3009:
3003:
3002:
2990:
2984:
2983:
2971:
2965:
2964:
2954:
2922:
2916:
2915:
2913:
2911:
2893:
2887:
2886:
2852:
2843:
2834:
2833:
2823:
2814:
2808:
2807:
2797:
2761:
2755:
2754:
2718:
2709:
2708:
2698:
2688:
2664:
2658:
2657:
2633:
2627:
2626:
2616:
2606:
2582:
2576:
2575:
2567:
2556:
2555:
2527:
2521:
2520:
2486:
2480:
2479:
2469:
2433:
2424:
2423:
2413:
2403:
2371:
2362:
2361:
2349:
2339:
2333:
2332:
2309:
2303:
2302:
2266:
2260:
2259:
2233:
2224:
2218:
2217:
2201:
2190:
2189:
2145:
2136:
2130:
2129:
2097:
2088:
2087:
2043:
2037:
2036:
1992:
1983:
1982:
1972:
1962:
1930:
1915:
1914:
1912:
1910:
1895:
1889:
1888:
1856:
1850:
1849:
1831:
1795:
1697:Beverly Halstead
1317:
1285:
1284:
1261:
1260:
1241:
1240:
1217:
1216:
1188:
1187:
1120:
1119:
1096:
1095:
1072:
1071:
1060:
1059:
992:
991:
952:
937:
936:
930:
929:
923:Dsungaripteridae
866:
865:
842:
841:
835:Thalassodrominae
830:
829:
823:
822:
811:
810:
775:A comparison of
573:giant azhdarchid
363:
362:
359:
358:
355:
352:
349:
346:
343:
340:
337:
334:
331:
328:
325:
322:
300:
294:
271:
252:
245:
234:Quetzalcoatlinae
232:
219:
208:Pterodactyloidea
206:
193:
153:
152:
124:
114:
44:
29:Temporal range:
21:
20:
8019:
8018:
8014:
8013:
8012:
8010:
8009:
8008:
7964:
7963:
7962:
7952:
7950:
7940:
7938:
7930:
7928:
7923:
7915:
7910:
7902:
7897:
7889:
7884:
7876:
7871:
7863:
7858:
7850:
7845:
7836:
7835:
7830:
7821:
7820:
7815:
7802:
7792:
7783:
7764:
7724:
7705:Liaoningopterus
7660:
7610:
7548:Tropeognathinae
7533:
7505:
7479:
7462:
7426:
7397:Cimoliopteridae
7377:
7277:
7200:
7187:
7182:Zhenyuanopterus
7156:
7107:
7086:Istiodactylinae
7080:
7061:Longchengpterus
7054:Lingyuanopterus
7041:Istiodactylidae
7035:
7001:
6974:Hongshanopterus
6955:
6924:
6910:
6897:
6860:Hongshanopterus
6845:
6810:
6785:
6758:Barbaridactylus
6743:
6705:
6673:
6620:Pteranodontidae
6605:
6600:Santanadactylus
6584:
6571:
6554:
6547:
6539:
6531:
6515:
6459:
6454:Zhejiangopterus
6375:
6333:
6312:Palaeocursornis
6274:
6253:Montanazhdarcho
6240:Azhdarchiformes
6230:
6211:Chaoyangopterus
6196:
6125:
6101:
6087:Montanazhdarcho
6040:
6000:
5998:Azhdarchomorpha
5967:
5940:
5894:
5857:Aymberedactylus
5838:
5766:
5742:
5730:Aymberedactylus
5705:
5685:Thalassodromeus
5614:
5589:
5578:
5569:
5557:Montanazhdarcho
5515:
5498:
5486:Domeykodactylus
5471:
5465:Tendaguripterus
5402:
5393:
5375:
5362:
5345:
5333:
5317:
5297:
5291:Altmuehlopterus
5248:
5207:
5183:
5134:
5106:
5011:
4994:
4958:
4952:Gallodactylidae
4932:Aurorazhdarchia
4926:
4904:Normannognathus
4880:Gallodactylidae
4868:
4859:
4835:
4822:
4816:Tendaguripterus
4808:Normannognathus
4801:Germanodactylus
4793:Altmuehlopterus
4778:
4757:
4744:
4727:
4689:
4669:
4583:
4548:Batrachognathus
4533:
4528:Vesperopterylus
4483:
4457:
4448:
4423:
4402:
4358:Wukongopterinae
4352:
4334:Wukongopteridae
4328:
4297:
4284:Normannognathus
4261:
4252:
4226:
4213:
4196:
4183:
4175:
4139:
4119:
4092:Jianchangnathus
4075:Scaphognathidae
4065:Digibrevisauria
4047:
4042:Rhamphorhynchus
4011:
3998:Qinglongopterus
3990:Harpactognathus
3958:Rhamphorhynchae
3952:
3947:Qinglongopterus
3926:Harpactognathus
3886:
3880:Scaphognathinae
3872:Parapsicephalus
3836:
3823:
3811:Bergamodactylus
3790:
3781:
3763:
3749:
3727:Parapsicephalus
3690:
3669:
3667:Macronychoptera
3660:
3642:
3620:
3592:
3586:Arcticodactylus
3567:
3555:Raeticodactylus
3516:
3482:
3465:
3445:Arcticodactylus
3430:
3424:Raeticodactylus
3389:
3377:Austriadactylus
3362:
3341:
3328:
3303:
3283:
3277:Avemetatarsalia
3262:Avemetatarsalia
3252:
3237:Avemetatarsalia
3202:
3201:
3191:
3186:
3145:
3133:
3121:
3109:
3098:
3093:
3083:
3081:
3080:. June 22, 2010
3072:
3071:
3067:
3057:
3055:
3042:
3041:
3037:
3027:
3025:
3010:
3006:
2991:
2987:
2972:
2968:
2923:
2919:
2909:
2907:
2903:Morning Edition
2894:
2890:
2850:
2844:
2837:
2821:
2815:
2811:
2780:(sup1): 21–45.
2762:
2758:
2719:
2712:
2665:
2661:
2648:(Sup. 3): 48A.
2634:
2630:
2597:(3): e2001663.
2583:
2579:
2568:
2559:
2528:
2524:
2509:
2487:
2483:
2434:
2427:
2372:
2365:
2358:
2340:
2336:
2329:
2311:
2310:
2306:
2267:
2263:
2231:
2225:
2221:
2202:
2193:
2143:
2137:
2133:
2098:
2091:
2044:
2040:
1993:
1986:
1931:
1918:
1908:
1906:
1896:
1892:
1857:
1853:
1796:
1781:
1777:
1760:
1669:
1636:thermal soaring
1550:
1452:
1412:
1407:
1398:
1389:
1380:
1371:
1362:
1353:
1344:
1175:
1166:
1157:
1047:
1038:
1029:
979:
912:
903:
769:
746:
671:
665:
660:
412:
373:Late Cretaceous
371:known from the
319:
315:
280:
277:
251:
243:
230:
217:
204:
191:
147:
136:
134:
115:
113:
112:
111:
110:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
39:
38:
31:Late Cretaceous
27:
17:
12:
11:
5:
8017:
8007:
8006:
8001:
7996:
7991:
7986:
7981:
7976:
7961:
7960:
7948:
7925:
7924:
7922:
7921:
7908:
7895:
7882:
7869:
7856:
7843:
7838:Quetzalcoatlus
7828:
7812:
7810:
7808:Quetzalcoatlus
7804:
7803:
7789:
7788:
7785:
7784:
7763:
7761:
7758:
7757:
7754:
7753:
7750:
7749:
7746:
7745:
7742:
7741:
7738:
7737:
7734:
7733:
7730:
7729:
7726:
7725:
7723:
7722:
7715:
7708:
7701:
7694:
7686:
7682:Caulkicephalus
7678:
7670:
7668:
7662:
7661:
7659:
7658:
7655:Uktenadactylus
7651:
7643:
7636:
7633:Coloborhynchus
7629:
7621:
7619:
7612:
7611:
7609:
7608:
7601:
7594:
7586:
7578:
7570:
7562:
7553:
7551:
7541:
7535:
7534:
7532:
7531:
7524:
7516:
7514:
7507:
7506:
7504:
7503:
7495:
7486:
7480:
7475:
7472:
7471:
7468:
7467:
7464:
7463:
7461:
7460:
7457:Targaryendraco
7453:
7445:
7436:
7434:
7428:
7427:
7425:
7424:
7417:
7409:
7401:
7399:
7390:
7383:
7382:
7379:
7378:
7376:
7375:
7371:Uktenadactylus
7367:
7359:
7356:Ornithocheirus
7352:
7344:
7336:
7328:
7320:
7312:
7308:Coloborhynchus
7304:
7296:
7287:
7285:
7279:
7278:
7276:
7275:
7267:
7259:
7251:
7247:Caulkicephalus
7243:
7235:
7232:Arthurdactylus
7228:
7220:
7214:
7201:
7198:Ornithocheirae
7196:
7193:
7192:
7189:
7188:
7186:
7185:
7178:
7170:
7168:
7155:
7154:
7147:
7140:
7133:
7125:
7123:
7117:
7116:
7113:
7112:
7109:
7108:
7106:
7105:
7098:
7090:
7088:
7082:
7081:
7079:
7078:
7071:
7064:
7057:
7049:
7043:
7037:
7036:
7034:
7033:
7026:
7023:Linlongopterus
7019:
7011:
7009:
7007:Mimodactylidae
7003:
7002:
7000:
6999:
6995:Lonchodectidae
6991:
6984:
6981:Linlongopterus
6977:
6969:
6963:
6957:
6956:
6954:
6953:
6946:
6938:
6936:
6923:
6922:
6913:
6911:
6906:
6903:
6902:
6899:
6898:
6896:
6895:
6891:Targaryendraco
6887:
6879:
6872:
6864:
6855:
6853:
6851:Lonchodectidae
6847:
6846:
6844:
6843:
6836:
6829:
6821:
6811:
6806:
6803:
6802:
6799:
6798:
6795:
6794:
6791:
6790:
6787:
6786:
6784:
6783:
6775:
6768:
6765:Muzquizopteryx
6761:
6753:
6751:
6745:
6744:
6742:
6741:
6734:
6727:
6719:
6713:
6711:Aponyctosauria
6707:
6706:
6704:
6703:
6695:
6687:
6681:
6675:
6674:
6672:
6671:
6663:
6655:
6648:
6640:
6633:
6624:
6622:
6613:
6607:
6606:
6604:
6603:
6595:
6585:
6580:
6573:
6572:
6560:
6559:
6556:
6555:
6530:
6528:
6525:
6524:
6521:
6520:
6517:
6516:
6514:
6513:
6512:
6511:
6505:
6503:
6493:
6492:
6489:
6488:
6485:
6484:
6481:
6480:
6477:
6476:
6473:
6472:
6469:
6468:
6465:
6464:
6461:
6460:
6458:
6457:
6450:
6442:
6439:Thanatosdrakon
6435:
6432:Quetzalcoatlus
6428:
6425:Phosphatodraco
6421:
6414:
6407:
6400:
6397:Arambourgiania
6393:
6385:
6383:
6377:
6376:
6374:
6373:
6366:
6359:
6352:
6343:
6341:
6332:
6331:
6323:
6315:
6308:
6304:Navajodactylus
6300:
6293:
6284:
6282:
6273:
6272:
6264:
6257:
6248:
6242:
6236:
6235:
6232:
6231:
6229:
6228:
6225:Shenzhoupterus
6221:
6214:
6206:
6204:
6198:
6197:
6195:
6194:
6186:
6178:
6171:
6164:
6156:
6152:Argentinadraco
6148:
6139:
6133:
6127:
6126:
6124:
6123:
6115:
6109:
6103:
6102:
6100:
6099:
6091:
6083:
6075:
6067:
6063:Argentinadraco
6059:
6051:
6049:
6042:
6041:
6039:
6038:
6030:
6023:
6016:
6007:
6001:
5996:
5993:
5992:
5989:
5988:
5985:
5984:
5981:
5980:
5977:
5976:
5973:
5972:
5969:
5968:
5966:
5965:
5958:
5950:
5948:
5942:
5941:
5939:
5938:
5931:
5924:
5917:
5908:
5902:
5896:
5895:
5893:
5892:
5885:
5877:
5869:
5861:
5852:
5846:
5840:
5839:
5837:
5836:
5829:
5822:
5819:Nemicolopterus
5815:
5808:
5801:
5793:
5785:
5776:
5774:
5765:
5764:
5756:
5750:
5744:
5743:
5741:
5740:
5733:
5725:
5723:
5713:
5711:Tapejariformes
5707:
5706:
5704:
5703:
5695:
5688:
5681:
5673:
5665:
5658:
5650:
5646:Argentinadraco
5642:
5634:
5625:
5623:
5616:
5615:
5613:
5612:
5604:
5596:
5590:
5587:Tapejaromorpha
5585:
5579:
5574:
5571:
5570:
5568:
5567:
5560:
5553:
5545:
5537:
5530:
5527:Argentinadraco
5522:
5516:
5511:
5508:
5507:
5504:
5503:
5500:
5499:
5497:
5496:
5493:Dsungaripterus
5489:
5481:
5479:
5473:
5472:
5470:
5469:
5461:
5453:
5446:
5439:
5431:
5422:
5416:
5403:
5398:
5395:
5394:
5392:
5391:
5382:
5376:
5371:
5364:
5363:
5351:
5350:
5347:
5346:
5332:
5330:
5327:
5326:
5323:
5322:
5319:
5318:
5316:
5315:
5314:
5313:
5307:
5305:
5299:
5298:
5296:
5295:
5286:
5280:
5270:
5269:
5266:
5265:
5262:
5261:
5258:
5257:
5254:
5253:
5250:
5249:
5247:
5246:
5239:
5232:
5225:
5217:
5215:
5213:Pterodaustrini
5209:
5208:
5206:
5205:
5197:
5191:
5185:
5184:
5182:
5181:
5174:
5167:
5160:
5152:
5144:
5142:
5140:Gnathosaurinae
5136:
5135:
5133:
5132:
5125:
5117:
5115:
5112:Moganopterinae
5108:
5107:
5105:
5104:
5097:
5090:
5083:
5076:
5069:
5062:
5055:
5048:
5041:
5034:
5031:Balaenognathus
5027:
5018:
5012:
5007:
5004:
5003:
5000:
4999:
4996:
4995:
4993:
4992:
4984:
4981:Aurorazhdarcho
4977:
4968:
4966:
4960:
4959:
4957:
4956:
4948:
4940:
4934:
4928:
4927:
4925:
4924:
4920:Moganopterinae
4916:
4908:
4900:
4893:
4889:Aurorazhdarcho
4884:
4882:
4869:
4864:
4861:
4860:
4858:
4857:
4850:
4842:
4836:
4831:
4828:
4827:
4824:
4823:
4821:
4820:
4812:
4804:
4797:
4788:
4786:
4780:
4779:
4777:
4776:
4768:
4758:
4753:
4746:
4745:
4733:
4732:
4729:
4728:
4716:
4714:
4711:
4710:
4707:
4706:
4703:
4702:
4699:
4698:
4695:
4694:
4691:
4690:
4688:
4687:
4686:
4685:
4679:
4677:
4671:
4670:
4668:
4667:
4660:
4652:
4645:
4638:
4631:
4624:
4616:
4609:
4601:
4595:
4589:
4588:
4585:
4584:
4582:
4581:
4574:
4566:
4558:
4551:
4543:
4541:
4535:
4534:
4532:
4531:
4524:
4517:
4509:
4501:
4493:
4491:
4489:Anurognathinae
4485:
4484:
4482:
4481:
4473:
4467:
4465:Anurognathidae
4458:
4453:
4450:
4449:
4447:
4446:
4443:Douzhanopterus
4439:
4430:
4424:
4419:
4416:
4415:
4412:
4411:
4408:
4407:
4404:
4403:
4401:
4400:
4393:
4389:Douzhanopterus
4385:
4378:
4371:
4362:
4360:
4354:
4353:
4351:
4350:
4347:Kunpengopterus
4342:
4336:
4330:
4329:
4327:
4326:
4319:
4311:
4305:
4299:
4298:
4296:
4295:
4291:Anurognathidae
4287:
4280:
4272:
4262:
4259:Monofenestrata
4257:
4254:
4253:
4251:
4250:
4242:
4233:
4227:
4222:
4215:
4214:
4202:
4201:
4198:
4197:
4174:
4172:
4169:
4168:
4165:
4164:
4161:
4160:
4157:
4156:
4153:
4152:
4149:
4148:
4145:
4144:
4141:
4140:
4138:
4137:
4136:
4135:
4129:
4127:
4121:
4120:
4118:
4117:
4109:
4102:
4095:
4088:
4080:
4078:
4068:
4061:
4060:
4057:
4056:
4053:
4052:
4049:
4048:
4046:
4045:
4038:
4030:
4021:
4019:
4013:
4012:
4010:
4009:
4002:
3994:
3986:
3979:
3971:
3969:
3960:
3954:
3953:
3951:
3950:
3943:
3940:Orientognathus
3936:
3929:
3922:
3915:
3908:
3900:
3894:
3888:
3887:
3885:
3884:
3876:
3868:
3861:
3857:Dolicorhamphus
3852:
3846:
3837:
3834:Breviquartossa
3832:
3829:
3828:
3825:
3824:
3822:
3821:
3814:
3806:
3804:
3791:
3786:
3783:
3782:
3780:
3779:
3770:
3764:
3758:
3755:
3754:
3751:
3750:
3748:
3747:
3739:
3731:
3723:
3716:
3713:Caelestiventus
3709:
3700:
3698:
3692:
3691:
3689:
3688:
3680:
3670:
3665:
3662:
3661:
3659:
3658:
3649:
3643:
3639:Zambellisauria
3637:
3634:
3633:
3630:
3629:
3626:
3625:
3622:
3621:
3619:
3618:
3611:
3607:Carniadactylus
3602:
3600:
3594:
3593:
3591:
3590:
3581:
3575:
3569:
3568:
3566:
3565:
3558:
3551:
3544:
3535:
3533:
3524:
3518:
3517:
3515:
3514:
3506:
3498:
3489:
3483:
3478:
3475:
3474:
3471:
3470:
3467:
3466:
3464:
3463:
3460:Seazzadactylus
3456:
3449:
3440:
3438:
3432:
3431:
3429:
3428:
3420:
3413:
3409:Carniadactylus
3404:
3398:
3391:
3390:
3388:
3387:
3380:
3372:
3370:
3364:
3363:
3361:
3360:
3357:Daohugoupterus
3352:
3342:
3337:
3330:
3329:
3317:
3316:
3313:
3312:
3309:
3308:
3305:
3304:
3302:
3301:
3300:
3299:
3293:
3291:
3285:
3284:
3282:
3281:
3280:
3279:
3270:
3264:
3254:
3253:
3249:
3248:
3239:
3230:
3221:
3215:
3209:
3193:
3192:
3185:
3184:
3177:
3170:
3162:
3156:
3155:
3143:
3131:
3119:
3107:
3103:Quetzalcoatlus
3097:
3096:External links
3094:
3092:
3091:
3065:
3035:
3004:
2985:
2966:
2937:(1): pgac023.
2917:
2888:
2861:(3): 768–785.
2835:
2809:
2768:Quetzalcoatlus
2756:
2729:(3): 218–239.
2710:
2659:
2638:Quetzalcoatlus
2628:
2577:
2557:
2522:
2507:
2481:
2440:Quetzalcoatlus
2425:
2386:(11): e13982.
2363:
2356:
2334:
2327:
2304:
2277:(2): 122–137.
2261:
2242:(1): 192–195.
2219:
2191:
2156:(4): 180–184.
2131:
2112:(2): 222–231.
2102:Quetzalcoatlus
2089:
2038:
1984:
1916:
1890:
1871:(4): 922–928.
1851:
1802:Quetzalcoatlus
1778:
1776:
1773:
1772:
1771:
1766:
1759:
1756:
1689:Quetzalcoatlus
1668:
1665:
1660:Quetzalcoatlus
1655:Quetzalcoatlus
1605:Quetzalcoatlus
1574:Quetzalcoatlus
1570:Paul MacCready
1566:Quetzalcoatlus
1558:Quetzalcoatlus
1549:
1546:
1528:Quetzalcoatlus
1524:Quetzalcoatlus
1503:Quetzalcoatlus
1476:Quetzalcoatlus
1468:Quetzalcoatlus
1460:Quetzalcoatlus
1451:
1448:
1437:Quetzalcoatlus
1433:Quetzalcoatlus
1415:Quetzalcoatlus
1411:
1408:
1404:
1403:
1400:
1399:
1395:
1394:
1391:
1390:
1386:
1385:
1382:
1381:
1377:
1376:
1373:
1372:
1368:
1367:
1364:
1363:
1359:
1358:
1355:
1354:
1350:
1349:
1346:
1345:
1341:
1340:
1337:
1336:
1327:
1324:
1323:
1320:
1319:
1305:
1302:
1301:
1298:
1297:
1288:
1283:
1281:
1278:
1277:
1274:
1273:
1264:
1259:
1257:
1254:
1253:
1250:
1249:
1244:
1239:
1237:
1234:
1233:
1230:
1229:
1220:
1215:
1213:
1205:
1204:
1201:
1200:
1191:
1186:
1184:
1181:
1180:
1177:
1176:
1172:
1171:
1168:
1167:
1163:
1162:
1159:
1158:
1154:
1153:
1150:
1149:
1140:
1137:
1136:
1133:
1132:
1123:
1118:
1116:
1113:
1112:
1109:
1108:
1099:
1094:
1092:
1089:
1088:
1085:
1084:
1075:
1070:
1068:
1058:
1056:
1053:
1052:
1049:
1048:
1044:
1043:
1040:
1039:
1035:
1034:
1031:
1030:
1026:
1025:
1022:
1021:
1012:
1009:
1008:
1005:
1004:
995:
990:
988:
985:
984:
981:
980:
976:
975:
972:
971:
962:
959:
958:
955:
954:
940:
935:
933:
928:
926:
918:
917:
914:
913:
909:
908:
905:
904:
900:
899:
896:
895:
886:
883:
882:
879:
878:
869:
864:
862:
859:
858:
855:
854:
845:
840:
838:
828:
826:
821:
819:
809:
805:Quetzalcoatlus
790:Quetzalcoatlus
782:Phosphatodraco
777:Quetzalcoatlus
768:
767:Classification
765:
754:Quetzalcoatlus
745:
742:
716:Quetzalcoatlus
701:Comparison of
691:Quetzalcoatlus
669:Pterosaur size
664:
661:
659:
656:
642:Quetzalcoatlus
611:Quetzalcoatlus
595:Quetzalcoatlus
551:Quetzalcoatlus
528:Quetzalcoatlus
508:Quetzalcoatlus
427:Quetzalcoatlus
411:
408:
364:is a genus of
312:Quetzalcoatlus
306:
305:
304:
303:
287:
286:
285:Other species
282:
281:
278:
266:
265:
259:
258:
248:Quetzalcoatlus
241:
237:
236:
228:
224:
223:
215:
211:
210:
202:
198:
197:
189:
185:
184:
179:
175:
174:
169:
165:
164:
159:
155:
154:
141:
140:
131:Quetzalcoatlus
126:
125:
117:
116:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
28:
25:Quetzalcoatlus
15:
9:
6:
4:
3:
2:
8016:
8005:
8002:
8000:
7997:
7995:
7992:
7990:
7987:
7985:
7982:
7980:
7977:
7975:
7972:
7971:
7969:
7959:
7958:United States
7949:
7947:
7937:
7936:
7933:
7918:
7913:
7909:
7905:
7900:
7896:
7892:
7887:
7883:
7879:
7874:
7870:
7866:
7861:
7857:
7853:
7848:
7844:
7839:
7833:
7829:
7824:
7818:
7814:
7813:
7811:
7809:
7805:
7801:
7796:
7781:
7777:
7775:
7768:
7721:
7720:
7719:Maaradactylus
7716:
7714:
7713:
7709:
7707:
7706:
7702:
7700:
7699:
7695:
7692:
7691:
7690:Cearadactylus
7687:
7684:
7683:
7679:
7677:
7676:
7672:
7671:
7669:
7667:
7666:Anhanguerinae
7663:
7657:
7656:
7652:
7649:
7648:
7647:Siroccopteryx
7644:
7642:
7641:
7637:
7635:
7634:
7630:
7628:
7627:
7623:
7622:
7620:
7617:
7613:
7607:
7606:
7605:Tropeognathus
7602:
7600:
7599:
7595:
7592:
7591:
7590:Siroccopteryx
7587:
7584:
7583:
7579:
7576:
7575:
7571:
7568:
7567:
7563:
7560:
7559:
7555:
7554:
7552:
7549:
7545:
7542:
7540:
7539:Anhangueridae
7536:
7530:
7529:
7528:Iberodactylus
7525:
7523:
7522:
7518:
7517:
7515:
7512:
7508:
7501:
7500:
7496:
7493:
7492:
7488:
7487:
7484:
7481:
7478:
7473:
7459:
7458:
7454:
7451:
7450:
7446:
7443:
7442:
7438:
7437:
7435:
7433:
7429:
7423:
7422:
7421:Cimoliopterus
7418:
7415:
7414:
7413:Camposipterus
7410:
7408:
7407:
7403:
7402:
7400:
7398:
7394:
7391:
7388:
7384:
7373:
7372:
7368:
7365:
7364:
7363:Siroccopteryx
7360:
7358:
7357:
7353:
7350:
7349:
7345:
7342:
7341:
7337:
7334:
7333:
7329:
7326:
7325:
7321:
7318:
7317:
7313:
7310:
7309:
7305:
7302:
7301:
7300:Camposipterus
7297:
7294:
7293:
7289:
7288:
7286:
7284:
7280:
7273:
7272:
7271:Tropeognathus
7268:
7265:
7264:
7260:
7257:
7256:
7255:Cimoliopterus
7252:
7249:
7248:
7244:
7241:
7240:
7239:Camposipterus
7236:
7234:
7233:
7229:
7227:
7226:
7225:Araripesaurus
7222:
7221:
7218:
7215:
7213:
7209:
7205:
7202:
7199:
7194:
7184:
7183:
7179:
7177:
7176:
7172:
7171:
7169:
7167:
7166:Boreopteridae
7163:
7159:
7153:
7152:
7148:
7146:
7145:
7144:Cearadactylus
7141:
7139:
7138:
7134:
7132:
7131:
7127:
7126:
7124:
7122:
7118:
7104:
7103:
7099:
7097:
7096:
7095:Istiodactylus
7092:
7091:
7089:
7087:
7083:
7077:
7076:
7072:
7070:
7069:
7065:
7063:
7062:
7058:
7056:
7055:
7051:
7050:
7047:
7044:
7042:
7038:
7032:
7031:
7027:
7025:
7024:
7020:
7018:
7017:
7013:
7012:
7010:
7008:
7004:
6997:
6996:
6992:
6990:
6989:
6988:Yixianopterus
6985:
6983:
6982:
6978:
6976:
6975:
6971:
6970:
6967:
6964:
6962:
6958:
6952:
6951:
6947:
6945:
6944:
6940:
6939:
6937:
6935:
6931:
6927:
6920:
6919:
6915:
6914:
6912:
6909:
6904:
6893:
6892:
6888:
6885:
6884:
6880:
6878:
6877:
6873:
6870:
6869:
6865:
6862:
6861:
6857:
6856:
6854:
6852:
6848:
6842:
6841:
6837:
6835:
6834:
6830:
6828:
6827:
6823:
6822:
6819:
6815:
6812:
6809:
6804:
6781:
6780:
6776:
6774:
6773:
6769:
6767:
6766:
6762:
6760:
6759:
6755:
6754:
6752:
6750:
6749:Nyctosauridae
6746:
6740:
6739:
6735:
6733:
6732:
6728:
6726:
6725:
6721:
6720:
6717:
6714:
6712:
6708:
6701:
6700:
6696:
6694:
6693:
6692:Alamodactylus
6689:
6688:
6685:
6682:
6680:
6676:
6669:
6668:
6664:
6661:
6660:
6656:
6654:
6653:
6649:
6646:
6645:
6641:
6639:
6638:
6634:
6631:
6630:
6626:
6625:
6623:
6621:
6617:
6614:
6612:
6611:Pteranodontia
6608:
6602:
6601:
6597:
6596:
6593:
6589:
6586:
6583:
6578:
6574:
6570:
6565:
6561:
6551:
6546:
6543:
6535:
6509:
6508:
6507:
6506:
6504:
6502:
6498:
6494:
6456:
6455:
6451:
6448:
6447:
6446:Wellnhopterus
6443:
6441:
6440:
6436:
6434:
6433:
6429:
6427:
6426:
6422:
6420:
6419:
6418:Hatzegopteryx
6415:
6413:
6412:
6408:
6406:
6405:
6401:
6399:
6398:
6394:
6392:
6391:
6390:Aralazhdarcho
6387:
6386:
6384:
6382:
6378:
6372:
6371:
6367:
6365:
6364:
6360:
6358:
6357:
6353:
6350:
6349:
6345:
6344:
6342:
6340:
6336:
6329:
6328:
6324:
6321:
6320:
6316:
6314:
6313:
6309:
6306:
6305:
6301:
6299:
6298:
6294:
6291:
6290:
6286:
6285:
6283:
6281:
6277:
6270:
6269:
6265:
6263:
6262:
6261:Radiodactylus
6258:
6255:
6254:
6250:
6249:
6246:
6243:
6241:
6237:
6227:
6226:
6222:
6220:
6219:
6215:
6213:
6212:
6208:
6207:
6205:
6203:
6199:
6192:
6191:
6187:
6184:
6183:
6179:
6177:
6176:
6172:
6170:
6169:
6165:
6162:
6161:
6157:
6154:
6153:
6149:
6146:
6145:
6144:Apatorhamphus
6141:
6140:
6137:
6134:
6132:
6128:
6122:
6121:
6117:
6116:
6113:
6110:
6108:
6104:
6097:
6096:
6092:
6089:
6088:
6084:
6081:
6080:
6076:
6073:
6072:
6068:
6065:
6064:
6060:
6058:
6057:
6053:
6052:
6050:
6047:
6043:
6036:
6035:
6031:
6029:
6028:
6024:
6022:
6021:
6017:
6014:
6013:
6009:
6008:
6005:
6002:
5999:
5994:
5964:
5963:
5959:
5957:
5956:
5952:
5951:
5949:
5947:
5943:
5937:
5936:
5935:Tupandactylus
5932:
5930:
5929:
5925:
5923:
5922:
5918:
5915:
5914:
5910:
5909:
5906:
5903:
5901:
5897:
5891:
5890:
5886:
5883:
5882:
5878:
5875:
5874:
5870:
5867:
5866:
5865:Caupedactylus
5862:
5859:
5858:
5854:
5853:
5850:
5847:
5845:
5841:
5835:
5834:
5830:
5828:
5827:
5823:
5821:
5820:
5816:
5814:
5813:
5809:
5807:
5806:
5802:
5799:
5798:
5794:
5791:
5790:
5786:
5783:
5782:
5778:
5777:
5775:
5773:
5769:
5763:
5762:
5758:
5757:
5754:
5751:
5749:
5745:
5739:
5738:
5737:Caupedactylus
5734:
5732:
5731:
5727:
5726:
5724:
5721:
5720:Caupedactylia
5717:
5714:
5712:
5708:
5701:
5700:
5696:
5694:
5693:
5689:
5687:
5686:
5682:
5679:
5678:
5674:
5671:
5670:
5666:
5664:
5663:
5659:
5656:
5655:
5651:
5648:
5647:
5643:
5640:
5639:
5635:
5632:
5631:
5627:
5626:
5624:
5621:
5617:
5610:
5609:
5605:
5603:
5602:
5598:
5597:
5594:
5591:
5588:
5583:
5580:
5577:
5576:Neoazhdarchia
5572:
5566:
5565:
5561:
5559:
5558:
5554:
5551:
5550:
5546:
5543:
5542:
5538:
5536:
5535:
5531:
5529:
5528:
5524:
5523:
5520:
5517:
5514:
5513:Azhdarchoidea
5509:
5495:
5494:
5490:
5488:
5487:
5483:
5482:
5480:
5478:
5474:
5467:
5466:
5462:
5459:
5458:
5457:Puntanipterus
5454:
5452:
5451:
5447:
5445:
5444:
5440:
5437:
5436:
5432:
5429:
5428:
5424:
5423:
5420:
5417:
5415:
5411:
5407:
5404:
5401:
5396:
5389:
5388:
5384:
5383:
5380:
5377:
5374:
5369:
5365:
5361:
5356:
5352:
5343:
5337:
5311:
5310:
5309:
5308:
5306:
5304:
5300:
5293:
5292:
5288:
5287:
5284:
5281:
5279:
5275:
5271:
5245:
5244:
5240:
5238:
5237:
5233:
5231:
5230:
5226:
5224:
5223:
5222:Beipiaopterus
5219:
5218:
5216:
5214:
5210:
5204:
5203:
5199:
5198:
5195:
5192:
5190:
5186:
5180:
5179:
5178:Tacuadactylus
5175:
5173:
5172:
5168:
5166:
5165:
5161:
5158:
5157:
5153:
5151:
5150:
5146:
5145:
5143:
5141:
5137:
5131:
5130:
5126:
5124:
5123:
5119:
5118:
5116:
5113:
5109:
5103:
5102:
5098:
5096:
5095:
5091:
5089:
5088:
5084:
5082:
5081:
5077:
5075:
5074:
5070:
5068:
5067:
5063:
5061:
5060:
5059:Forfexopterus
5056:
5054:
5053:
5052:Elanodactylus
5049:
5047:
5046:
5045:Cratonopterus
5042:
5040:
5039:
5038:Cathayopterus
5035:
5033:
5032:
5028:
5025:
5024:
5023:Ardeadactylus
5020:
5019:
5016:
5013:
5010:
5005:
4990:
4989:
4985:
4983:
4982:
4978:
4975:
4974:
4973:Ardeadactylus
4970:
4969:
4967:
4965:
4961:
4954:
4953:
4949:
4947:
4946:
4942:
4941:
4938:
4935:
4933:
4929:
4922:
4921:
4917:
4914:
4913:
4909:
4906:
4905:
4901:
4899:
4898:
4897:Cycnorhamphus
4894:
4891:
4890:
4886:
4885:
4883:
4881:
4877:
4873:
4870:
4867:
4862:
4856:
4855:
4854:Pterodactylus
4851:
4849:
4848:
4847:Diopecephalus
4844:
4843:
4840:
4837:
4834:
4829:
4818:
4817:
4813:
4810:
4809:
4805:
4803:
4802:
4798:
4795:
4794:
4790:
4789:
4787:
4785:
4781:
4775:
4774:
4773:Prejanopterus
4770:
4769:
4766:
4762:
4759:
4756:
4751:
4747:
4743:
4738:
4734:
4725:
4720:
4683:
4682:
4681:
4680:
4678:
4676:
4672:
4666:
4665:
4661:
4658:
4657:
4653:
4651:
4650:
4646:
4644:
4643:
4639:
4637:
4636:
4632:
4630:
4629:
4625:
4622:
4621:
4620:Herbstosaurus
4617:
4615:
4614:
4613:Eurolimnornis
4610:
4608:
4607:
4606:Dermodactylus
4603:
4602:
4599:
4596:
4594:
4590:
4580:
4579:
4575:
4572:
4571:
4567:
4564:
4563:
4559:
4557:
4556:
4552:
4550:
4549:
4545:
4544:
4542:
4540:
4536:
4530:
4529:
4525:
4523:
4522:
4518:
4515:
4514:
4510:
4507:
4506:
4502:
4500:
4499:
4495:
4494:
4492:
4490:
4486:
4480:
4479:
4475:
4474:
4471:
4468:
4466:
4462:
4459:
4456:
4455:Caelidracones
4451:
4445:
4444:
4440:
4437:
4436:
4432:
4431:
4428:
4425:
4422:
4417:
4399:
4398:
4397:Wukongopterus
4394:
4391:
4390:
4386:
4384:
4383:
4382:Darwinopterus
4379:
4377:
4376:
4375:Cuspicephalus
4372:
4369:
4368:
4364:
4363:
4361:
4359:
4355:
4349:
4348:
4344:
4343:
4340:
4337:
4335:
4331:
4325:
4324:
4323:Pterorhynchus
4320:
4318:
4317:
4313:
4312:
4309:
4306:
4304:
4300:
4293:
4292:
4288:
4286:
4285:
4281:
4279:
4278:
4274:
4273:
4270:
4266:
4263:
4260:
4255:
4248:
4247:
4243:
4240:
4239:
4235:
4234:
4231:
4228:
4225:
4220:
4216:
4212:
4207:
4203:
4194:
4190:
4187:
4179:
4133:
4132:
4131:
4130:
4128:
4126:
4122:
4115:
4114:
4110:
4108:
4107:
4106:Scaphognathus
4103:
4101:
4100:
4096:
4094:
4093:
4089:
4087:
4086:
4082:
4081:
4079:
4076:
4072:
4069:
4066:
4062:
4044:
4043:
4039:
4036:
4035:
4031:
4028:
4027:
4023:
4022:
4020:
4018:
4014:
4008:
4007:
4003:
4000:
3999:
3995:
3992:
3991:
3987:
3985:
3984:
3980:
3978:
3977:
3973:
3972:
3970:
3968:
3964:
3961:
3959:
3955:
3949:
3948:
3944:
3942:
3941:
3937:
3935:
3934:
3930:
3928:
3927:
3923:
3921:
3920:
3916:
3914:
3913:
3909:
3907:
3906:
3902:
3901:
3898:
3895:
3893:
3889:
3882:
3881:
3877:
3874:
3873:
3869:
3867:
3866:
3862:
3859:
3858:
3854:
3853:
3850:
3847:
3845:
3841:
3838:
3835:
3830:
3820:
3819:
3815:
3813:
3812:
3808:
3807:
3805:
3803:
3799:
3795:
3792:
3789:
3784:
3777:
3776:
3775:Eudimorphodon
3772:
3771:
3768:
3765:
3761:
3756:
3745:
3744:
3740:
3737:
3736:
3735:Peteinosaurus
3732:
3729:
3728:
3724:
3722:
3721:
3717:
3715:
3714:
3710:
3707:
3706:
3702:
3701:
3699:
3697:
3693:
3687:
3686:
3685:Herbstosaurus
3682:
3681:
3678:
3674:
3671:
3668:
3663:
3656:
3655:
3654:Peteinosaurus
3651:
3650:
3647:
3644:
3640:
3635:
3617:
3616:
3615:Eudimorphodon
3612:
3609:
3608:
3604:
3603:
3601:
3599:
3595:
3588:
3587:
3583:
3582:
3579:
3576:
3574:
3570:
3564:
3563:
3559:
3557:
3556:
3552:
3550:
3549:
3545:
3542:
3541:
3537:
3536:
3534:
3532:
3528:
3525:
3523:
3519:
3512:
3511:
3510:Preondactylia
3507:
3504:
3503:
3502:Peteinosaurus
3499:
3496:
3495:
3491:
3490:
3487:
3484:
3481:
3480:Eopterosauria
3476:
3462:
3461:
3457:
3455:
3454:
3450:
3447:
3446:
3442:
3441:
3439:
3437:
3433:
3426:
3425:
3421:
3419:
3418:
3414:
3411:
3410:
3406:
3405:
3402:
3399:
3396:
3392:
3386:
3385:
3384:Preondactylus
3381:
3379:
3378:
3374:
3373:
3371:
3369:
3368:Preondactylia
3365:
3359:
3358:
3354:
3353:
3350:
3346:
3343:
3340:
3335:
3331:
3327:
3322:
3318:
3297:
3296:
3295:
3294:
3292:
3290:
3286:
3278:
3274:
3273:
3272:
3271:
3268:
3265:
3263:
3259:
3255:
3247:
3243:
3240:
3238:
3234:
3231:
3229:
3225:
3222:
3220:
3216:
3214:
3210:
3208:
3204:
3203:
3198:
3194:
3190:
3183:
3178:
3176:
3171:
3169:
3164:
3163:
3160:
3152:
3148:
3144:
3140:
3136:
3132:
3128:
3124:
3120:
3116:
3112:
3108:
3105:
3104:
3100:
3099:
3079:
3075:
3069:
3053:
3049:
3045:
3039:
3023:
3019:
3015:
3008:
3000:
2996:
2989:
2981:
2977:
2970:
2962:
2958:
2953:
2948:
2944:
2940:
2936:
2932:
2928:
2921:
2905:
2904:
2899:
2892:
2884:
2880:
2876:
2872:
2868:
2864:
2860:
2856:
2849:
2842:
2840:
2831:
2827:
2820:
2813:
2805:
2801:
2796:
2791:
2787:
2783:
2779:
2775:
2771:
2769:
2760:
2752:
2748:
2744:
2740:
2736:
2732:
2728:
2724:
2717:
2715:
2706:
2702:
2697:
2692:
2687:
2682:
2678:
2674:
2670:
2663:
2655:
2651:
2647:
2643:
2639:
2632:
2624:
2620:
2615:
2610:
2605:
2600:
2596:
2592:
2588:
2581:
2573:
2566:
2564:
2562:
2553:
2549:
2545:
2541:
2537:
2533:
2526:
2518:
2514:
2510:
2508:9780691232218
2504:
2500:
2496:
2492:
2485:
2477:
2473:
2468:
2463:
2459:
2455:
2451:
2447:
2443:
2441:
2432:
2430:
2421:
2417:
2412:
2407:
2402:
2397:
2393:
2389:
2385:
2381:
2377:
2370:
2368:
2359:
2357:0-8018-6763-0
2353:
2348:
2347:
2338:
2330:
2324:
2320:
2316:
2315:
2308:
2300:
2296:
2292:
2288:
2284:
2280:
2276:
2272:
2265:
2257:
2253:
2249:
2245:
2241:
2237:
2230:
2223:
2215:
2211:
2207:
2200:
2198:
2196:
2187:
2183:
2179:
2175:
2171:
2167:
2163:
2159:
2155:
2151:
2150:
2142:
2135:
2127:
2123:
2119:
2115:
2111:
2107:
2103:
2096:
2094:
2085:
2081:
2077:
2073:
2069:
2065:
2061:
2057:
2053:
2049:
2042:
2034:
2030:
2026:
2022:
2018:
2014:
2010:
2006:
2002:
1998:
1991:
1989:
1980:
1976:
1971:
1966:
1961:
1956:
1952:
1948:
1944:
1940:
1936:
1929:
1927:
1925:
1923:
1921:
1905:
1904:Berkeley News
1901:
1894:
1886:
1882:
1878:
1874:
1870:
1866:
1862:
1855:
1847:
1843:
1839:
1835:
1830:
1825:
1821:
1817:
1814:(sup1): 142.
1813:
1809:
1805:
1803:
1794:
1792:
1790:
1788:
1786:
1784:
1779:
1770:
1767:
1765:
1762:
1761:
1755:
1753:
1749:
1748:Royal Society
1745:
1741:
1736:
1734:
1729:
1725:
1721:
1717:
1713:
1709:
1708:AeroVironment
1705:
1700:
1698:
1694:
1690:
1682:
1681:Royal Society
1678:
1673:
1664:
1661:
1656:
1652:
1648:
1643:
1641:
1637:
1632:
1628:
1623:
1619:
1610:
1606:
1602:
1598:
1596:
1591:
1587:
1583:
1579:
1575:
1571:
1567:
1559:
1554:
1545:
1543:
1539:
1535:
1533:
1529:
1525:
1521:
1516:
1512:
1508:
1504:
1500:
1494:
1492:
1488:
1485:
1481:
1480:marabou stork
1477:
1473:
1469:
1461:
1456:
1447:
1445:
1444:
1443:Wellnhopterus
1438:
1434:
1430:
1426:
1425:
1420:
1416:
1402:
1401:
1393:
1392:
1384:
1383:
1375:
1374:
1366:
1365:
1357:
1356:
1348:
1347:
1339:
1338:
1335:
1334:
1333:
1326:
1325:
1322:
1321:
1318:
1316:
1312:
1311:
1304:
1303:
1300:
1299:
1296:
1295:
1294:
1287:
1286:
1280:
1279:
1276:
1275:
1272:
1271:
1270:
1263:
1262:
1256:
1255:
1252:
1251:
1248:
1243:
1242:
1236:
1235:
1232:
1231:
1228:
1227:
1226:
1219:
1218:
1211:
1207:
1206:
1203:
1202:
1199:
1198:
1197:
1190:
1189:
1183:
1182:
1179:
1178:
1170:
1169:
1161:
1160:
1152:
1151:
1148:
1147:
1146:
1139:
1138:
1135:
1134:
1131:
1130:
1129:
1122:
1121:
1115:
1114:
1111:
1110:
1107:
1106:
1105:
1098:
1097:
1091:
1090:
1087:
1086:
1083:
1082:
1081:
1074:
1073:
1066:
1062:
1061:
1055:
1054:
1051:
1050:
1042:
1041:
1033:
1032:
1024:
1023:
1020:
1019:
1018:
1011:
1010:
1007:
1006:
1003:
1002:
1001:
994:
993:
987:
986:
983:
982:
974:
973:
970:
969:
968:
961:
960:
957:
956:
953:
951:
947:
946:
939:
938:
932:
931:
924:
920:
919:
916:
915:
907:
906:
898:
897:
894:
893:
892:
885:
884:
881:
880:
877:
876:
875:
868:
867:
861:
860:
857:
856:
853:
852:
851:
844:
843:
836:
832:
831:
825:
824:
817:
816:Neoazhdarchia
813:
812:
808:
806:
802:
797:
795:
791:
784:
783:
778:
773:
764:
762:
761:
760:Wellnhopterus
755:
751:
739:
734:
730:
728:
724:
719:
717:
711:
704:
699:
695:
692:
684:
680:
675:
670:
655:
653:
649:
648:
643:
639:
634:
632:
628:
624:
620:
616:
612:
608:
604:
603:Hatzegopteryx
600:
596:
592:
591:Hatzegopteryx
588:
587:
582:
578:
577:
576:Hatzegopteryx
572:
568:
564:
560:
556:
552:
548:
543:
541:
537:
533:
529:
520:
515:
511:
509:
505:
501:
497:
493:
492:specific name
489:
485:
481:
477:
473:
472:
467:
463:
458:
456:
451:
447:
443:
439:
436:
435:Maastrichtian
432:
428:
421:
416:
407:
405:
401:
400:
395:
394:Jack Northrop
391:
390:
385:
381:
377:
376:Maastrichtian
374:
370:
367:
361:
314:
313:
299:
298:
291:
290:
288:
283:
276:
275:
267:
264:
260:
255:
250:
249:
242:
239:
238:
235:
229:
226:
225:
222:
216:
213:
212:
209:
203:
200:
199:
196:
190:
187:
186:
183:
180:
177:
176:
173:
170:
167:
166:
163:
160:
157:
156:
151:
146:
142:
139:
132:
127:
123:
118:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
43:
36:
35:Maastrichtian
32:
26:
22:
19:
8004:Quetzalcoatl
7946:Paleontology
7807:
7771:
7717:
7712:Ludodactylus
7710:
7703:
7696:
7688:
7680:
7673:
7653:
7645:
7640:Nicorhynchus
7638:
7631:
7624:
7603:
7596:
7588:
7580:
7572:
7564:
7556:
7526:
7519:
7511:Hamipteridae
7497:
7489:
7455:
7447:
7439:
7419:
7411:
7406:Aetodactylus
7404:
7369:
7361:
7354:
7346:
7340:Ludodactylus
7338:
7330:
7322:
7314:
7306:
7298:
7292:Aetodactylus
7290:
7269:
7261:
7253:
7245:
7237:
7230:
7223:
7180:
7173:
7149:
7142:
7135:
7128:
7102:Liaoxipterus
7100:
7093:
7073:
7066:
7059:
7052:
7030:Mimodactylus
7028:
7021:
7014:
6993:
6986:
6979:
6972:
6948:
6941:
6916:
6908:Lanceodontia
6889:
6881:
6876:Lonchodectes
6874:
6866:
6858:
6838:
6831:
6824:
6777:
6770:
6763:
6756:
6736:
6729:
6722:
6697:
6690:
6665:
6657:
6650:
6644:Ornithostoma
6642:
6635:
6627:
6598:
6538:
6452:
6444:
6437:
6431:
6430:
6423:
6416:
6411:Eurazhdarcho
6409:
6402:
6395:
6388:
6368:
6361:
6354:
6346:
6339:Azhdarchinae
6325:
6317:
6310:
6302:
6295:
6287:
6280:Azhdarchidae
6266:
6259:
6251:
6223:
6216:
6209:
6188:
6180:
6175:Meilifeilong
6173:
6166:
6158:
6150:
6142:
6118:
6093:
6085:
6077:
6069:
6061:
6054:
6032:
6025:
6018:
6010:
5960:
5953:
5933:
5926:
5919:
5911:
5887:
5879:
5871:
5863:
5855:
5831:
5824:
5817:
5812:Huaxiapterus
5810:
5803:
5797:Eopteranodon
5795:
5787:
5781:Afrotapejara
5779:
5772:Sinopterinae
5761:Afrotapejara
5759:
5735:
5728:
5697:
5690:
5683:
5675:
5667:
5660:
5652:
5644:
5636:
5628:
5606:
5601:Bennettazhia
5599:
5564:Ornithostoma
5562:
5555:
5547:
5539:
5532:
5525:
5491:
5484:
5463:
5455:
5450:Ordosipterus
5448:
5441:
5433:
5425:
5400:Tapejaroidea
5385:
5289:
5243:Pterodaustro
5241:
5234:
5227:
5220:
5200:
5176:
5169:
5162:
5156:Huanhepterus
5154:
5149:Gnathosaurus
5147:
5129:Moganopterus
5127:
5120:
5101:Pterofiltrus
5099:
5094:Petrodactyle
5092:
5085:
5080:Liaodactylus
5078:
5073:Kepodactylus
5071:
5064:
5057:
5050:
5043:
5036:
5029:
5021:
4988:Huanhepterus
4986:
4979:
4971:
4950:
4945:Aerodactylus
4943:
4918:
4912:Petrodactyle
4910:
4902:
4895:
4887:
4852:
4845:
4814:
4806:
4799:
4791:
4771:
4662:
4654:
4647:
4640:
4633:
4628:Kryptodrakon
4626:
4618:
4611:
4604:
4576:
4570:Jeholopterus
4568:
4560:
4553:
4546:
4526:
4519:
4513:Jeholopterus
4511:
4503:
4498:Anurognathus
4496:
4478:Mesadactylus
4476:
4441:
4433:
4395:
4387:
4380:
4373:
4365:
4345:
4321:
4314:
4303:Darwinoptera
4289:
4282:
4275:
4244:
4236:
4182:
4111:
4104:
4097:
4090:
4083:
4040:
4034:Nesodactylus
4032:
4026:Cacibupteryx
4024:
4006:Sericipterus
4004:
3996:
3988:
3981:
3974:
3945:
3938:
3933:Nesodactylus
3931:
3924:
3917:
3912:Cacibupteryx
3910:
3905:Bellubrunnus
3903:
3878:
3870:
3863:
3855:
3816:
3809:
3773:
3760:Lonchognatha
3741:
3733:
3725:
3718:
3711:
3703:
3683:
3652:
3613:
3605:
3584:
3560:
3553:
3548:Pachagnathus
3546:
3538:
3508:
3500:
3494:Austriadraco
3492:
3458:
3453:Austriadraco
3451:
3443:
3422:
3415:
3407:
3382:
3375:
3355:
3241:
3232:
3223:
3150:
3138:
3126:
3114:
3102:
3082:. Retrieved
3077:
3068:
3056:. Retrieved
3052:the original
3047:
3038:
3026:. Retrieved
3022:the original
3017:
3007:
2998:
2988:
2979:
2969:
2934:
2930:
2920:
2908:. Retrieved
2901:
2891:
2858:
2854:
2829:
2825:
2812:
2777:
2773:
2767:
2759:
2726:
2723:Paleobiology
2722:
2676:
2673:PLOS Biology
2672:
2662:
2645:
2641:
2637:
2631:
2594:
2591:PLOS Biology
2590:
2580:
2571:
2535:
2531:
2525:
2490:
2484:
2449:
2445:
2439:
2383:
2379:
2345:
2337:
2313:
2307:
2274:
2270:
2264:
2239:
2235:
2222:
2213:
2209:
2153:
2147:
2134:
2109:
2105:
2101:
2051:
2047:
2041:
2000:
1996:
1945:(5): e2271.
1942:
1938:
1907:. Retrieved
1903:
1893:
1868:
1864:
1860:
1854:
1811:
1807:
1801:
1740:Q. northropi
1739:
1737:
1728:Q. northropi
1727:
1720:Q. northropi
1719:
1711:
1701:
1692:
1688:
1686:
1659:
1654:
1650:
1647:Q. northropi
1646:
1644:
1640:Kori bustard
1631:Q. northropi
1630:
1627:Q. northropi
1626:
1622:Q. northropi
1621:
1614:
1604:
1590:Q. northropi
1589:
1573:
1565:
1563:
1557:
1541:
1537:
1536:
1527:
1523:
1515:Darren Naish
1502:
1495:
1490:
1475:
1467:
1465:
1459:
1441:
1436:
1432:
1428:
1422:
1414:
1413:
1410:Paleobiology
1331:
1330:
1329:
1309:
1308:
1307:
1291:
1290:
1267:
1266:
1246:
1223:
1222:
1210:Azhdarchidae
1194:
1193:
1143:
1142:
1126:
1125:
1102:
1101:
1078:
1077:
1015:
1014:
998:
997:
965:
964:
943:
942:
889:
888:
872:
871:
848:
847:
804:
798:
794:Azhdarchidae
789:
788:
780:
776:
758:
753:
749:
747:
737:
726:
722:
720:
715:
712:
708:
703:Q. northropi
702:
690:
688:
682:
681:(green) and
679:Q. northropi
678:
645:
641:
635:
631:Q. northropi
630:
626:
623:Q. northropi
622:
618:
615:Q. northropi
614:
610:
606:
602:
598:
594:
590:
586:nomen dubium
584:
581:Q. northropi
580:
574:
567:Q. northropi
566:
562:
559:Q. northropi
558:
550:
547:Q. northropi
546:
544:
539:
536:Q. northropi
535:
527:
524:
518:
507:
488:Quetzalcoatl
479:
469:
459:
426:
424:
404:Q. northropi
403:
398:
397:
389:Q. northropi
388:
387:
384:type species
380:Quetzalcoatl
311:
310:
309:
296:
295:
279:Lawson, 1975
273:
272:
263:Type species
247:
246:
221:Azhdarchidae
130:
24:
18:
7974:Azhdarchids
7873:iNaturalist
7832:Wikispecies
7598:Thapunngaka
7558:Amblydectes
7477:Anhangueria
7441:Aussiedraco
7175:Boreopterus
6950:Lonchodraco
6883:Lonchodraco
6826:Aussiedraco
6772:Nyctosaurus
6629:Bogolubovia
6297:Bogolubovia
6168:Lacusovagus
6160:Eoazhdarcho
6120:Eoazhdarcho
6079:Leptostomia
6071:Keresdrakon
5946:Caiuajarina
5913:Bakonydraco
5881:Lacusovagus
5873:Keresdrakon
5844:Tapejarinae
5805:Huaxiadraco
5789:Bakonydraco
5748:Tapejaridae
5677:Leptostomia
5669:Lacusovagus
5662:Kariridraco
5608:Keresdrakon
5549:Leptostomia
5541:Keresdrakon
5202:Ctenochasma
5164:Lusognathus
5087:Otogopterus
4742:Lophocratia
4675:Lophocratia
4642:Pangupterus
4578:Sinomacrops
3919:Dorygnathus
3788:Novialoidea
3743:Rhamphinion
3720:Dimorphodon
3395:Caviramidae
3339:Pterosauria
3326:Pterosauria
3289:Pterosauria
3228:Archosauria
3189:Pterosauria
3028:November 1,
2910:November 1,
2832:(2): 18–24.
2679:(8): e204.
1909:December 7,
1861:Alamosaurus
1645:Studies of
1578:ornithopter
1491:Alamosaurus
1472:fish-eating
1429:Alamosaurus
1424:Alamosaurus
799:Below is a
752:shows that
658:Description
589:), perhaps
555:Mark Witton
504:flying wing
227:Subfamily:
195:Pterosauria
40:68–66
7968:Categories
7675:Anhanguera
7566:Ferrodraco
7521:Hamipterus
7449:Barbosania
7324:Ferrodraco
7316:Draigwenia
7151:Hamipterus
7130:Barbosania
7075:Nurhachius
6943:Ikrandraco
6918:Draigwenia
6868:Ikrandraco
6833:Serradraco
6779:Volgadraco
6667:Volgadraco
6659:Tethydraco
6652:Pteranodon
6510:see below↓
6404:Cryodrakon
6327:Volgadraco
6319:Tethydraco
6218:Jidapterus
6182:Microtuban
6020:Microtuban
5921:Europejara
5900:Tapejarini
5889:Vectidraco
5826:Sinopterus
5443:Noripterus
5312:see below↓
5236:Gegepterus
5229:Eosipterus
4684:see below↓
4664:Wenupteryx
4555:Cascocauda
4134:see below↓
3562:Yelaphomte
3298:see below↓
3219:Sauropsida
3084:August 20,
2931:PNAS Nexus
2538:(3–4): 1.
1775:References
1744:South Bank
1677:South Bank
1651:Q. lawsoni
1595:flightless
1542:Q. lawsoni
1489:, such as
1484:titanosaur
1247:TMM 42489
750:Q. lawsoni
738:Q. lawsoni
727:Q. lawsoni
723:Q. lawsoni
683:Q. lawsoni
667:See also:
627:Q. lawsoni
619:Q. lawsoni
607:Q. lawsoni
540:Q. lawsoni
519:Q. lawsoni
425:The first
399:Q. lawsoni
366:azhdarchid
297:Q. lawsoni
201:Suborder:
7626:Aerodraco
7068:Luchibang
7016:Haopterus
6738:Simurghia
6731:Epapatelo
6699:Cretornis
6637:Dawndraco
6363:Azhdarcho
6356:Albadraco
6348:Aerotitan
6268:Alanqidae
6046:Alanqidae
6012:Cretornis
5962:Torukjara
5955:Caiuajara
5692:Tupuxuara
5630:Aerotitan
5534:Inabtanin
5122:Feilongus
4521:Luopterus
4238:Allkaruen
3865:Klobiodon
3705:Allkaruen
3540:Caviramus
3417:Caviramus
3205:Kingdom:
3048:avinc.com
2804:245009158
2517:249332375
2476:245125427
2256:130751879
1885:130280606
1846:245125409
1838:0272-4634
1582:autopilot
1560:in flight
1499:vertebrae
1487:sauropods
801:cladogram
433:from the
369:pterosaur
168:Kingdom:
162:Eukaryota
129:Restored
7817:Wikidata
7698:Guidraco
7582:Mythunga
7574:Haliskia
7348:Mythunga
7332:Guidraco
7263:Haliskia
6840:Unwindia
6190:Xericeps
6095:Xericeps
6027:Xericeps
5928:Tapejara
5699:Xericeps
5654:Banguela
5427:Banguela
4656:Samrukia
4316:Ceoptera
3213:Chordata
3211:Phylum:
3207:Animalia
3058:July 28,
2961:36712794
2883:85679231
2751:88434056
2705:17676976
2623:29534059
2552:84617119
2420:21085624
2380:PLOS ONE
2299:24964287
2216:: 79–81.
2186:15423666
2178:12061403
2084:44400704
2076:17755159
2033:46396417
2025:17745279
1979:18509539
1939:PLOS ONE
1758:See also
1679:for the
1532:ungulate
1507:skimmers
652:dinosaur
571:Romanian
476:holotype
455:wingspan
214:Family:
182:Chordata
178:Phylum:
172:Animalia
158:Domain:
133:skeleton
7932:Portals
7904:4946961
7891:1042058
7865:4818447
7852:4532239
7823:Q131205
6724:Alcione
5833:Wightia
3217:Class:
3151:YouTube
3139:YouTube
3127:YouTube
3115:YouTube
2952:9802081
2863:Bibcode
2782:Bibcode
2731:Bibcode
2696:1925135
2614:5849296
2454:Bibcode
2411:2981443
2388:Bibcode
2279:Bibcode
2158:Bibcode
2114:Bibcode
2056:Bibcode
2048:Science
2005:Bibcode
1997:Science
1970:2386974
1947:Bibcode
1816:Bibcode
1586:soaring
1450:Feeding
1419:Lancian
1212:
1067:
925:
837:
818:
530:sp. by
494:honors
471:Science
464:of the
420:humerus
240:Genus:
188:Order:
109:↓
7878:523016
6289:Alanqa
6056:Alanqa
5638:Alanqa
4246:Sordes
4113:Sordes
2959:
2949:
2881:
2802:
2749:
2703:
2693:
2621:
2611:
2550:
2515:
2505:
2474:
2418:
2408:
2354:
2325:
2297:
2254:
2184:
2176:
2082:
2074:
2031:
2023:
1977:
1967:
1883:
1844:
1836:
1706:) and
1548:Flight
1520:storks
1427:. The
1208:
1063:
921:
833:
814:
685:(blue)
599:et al.
563:et al.
490:. The
382:. The
256:, 1975
254:Lawson
7917:38504
7886:IRMNG
5387:Piksi
3983:Dearc
3242:Clade
3233:Clade
3224:Clade
2906:. NPR
2879:S2CID
2851:(PDF)
2822:(PDF)
2800:S2CID
2747:S2CID
2548:S2CID
2513:S2CID
2472:S2CID
2295:JSTOR
2252:S2CID
2232:(PDF)
2182:S2CID
2144:(PDF)
2080:S2CID
2029:S2CID
1881:S2CID
1842:S2CID
1710:used
1704:DARPA
744:Skull
484:Aztec
431:Texas
7860:GBIF
3275:see
3086:2020
3060:2011
3030:2017
2957:PMID
2912:2017
2701:PMID
2619:PMID
2503:ISBN
2416:PMID
2352:ISBN
2323:ISBN
2174:PMID
2072:PMID
2021:PMID
1975:PMID
1911:2023
1834:ISSN
1649:and
1511:drag
663:Size
605:and
48:PreꞒ
7847:EoL
2947:PMC
2939:doi
2871:doi
2790:doi
2739:doi
2691:PMC
2681:doi
2650:doi
2609:PMC
2599:doi
2540:doi
2536:103
2495:doi
2462:doi
2406:PMC
2396:doi
2287:doi
2275:244
2244:doi
2166:doi
2122:doi
2064:doi
2052:188
2013:doi
2001:187
1965:PMC
1955:doi
1873:doi
1824:doi
633:.
440:at
386:is
37:),
7970::
7914::
7901::
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3076:.
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2955:.
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2933:.
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2877:.
2869:.
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2857:.
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2830:49
2828:.
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