1264:
1655:
1796:
1703:
1027:
1500:
1417:
977:
2537:
2336:
524:
1728:
33:
55:
2108:
1524:
876:
1288:
2043:
1611:
2841:
1918:
1220:
1091:
1002:
2152:
2018:
1333:
671:
1116:
735:
461:
2235:
1753:
626:
436:
412:
1679:
2773:
Pérez-Ben, Celeste M.; Gómez, Raúl O.; Báez, Ana M. (2019-04-04). "A new
Pliocene true toad (Anura: Bufonidae): first record of an extinct species from South America".
1802:
1962:
2401:
652:
2000:
1482:
940:
1637:
1073:
2519:
2441:
1840:
1270:
838:
780:
1943:
1778:
1506:
2500:
2279:
1981:
1379:
799:
2362:
1759:
1182:
741:
717:
568:
2543:
1661:
530:
2995:
1685:
1463:
3021:
3134:
2969:
3008:
3124:
2608:
3013:
3129:
2640:
869:
Colombia, southern
Venezuela (Cerro Neblina and Cerro Marahuaca), eastern Ecuador, northeastern Peru (Loreto), Brazil
2708:
830:
Bolivia (Pando), Brazil (Amazonas, Pará, and Rondônia), Colombia (Amazonas, Caquetá, and
Putumayo) and eastern Peru
3073:
3026:
1795:
1435:
Brazil (between Bujaru in northeastern Pará and Maranhão in the north, south to Bahia and Vitória, Espírito Santo)
2732:
Ferrão, Miquéias; Lima, Albertina
Pimentel; Ron, Santiago; Santos, Sueny Paloma dos; Hanken, James (2020-12-28).
1263:
1654:
2917:
2734:"New Species of Leaf-litter Toad of the Rhinella margaritifera Species Group (Anura: Bufonidae) from Amazonia"
1813:
Brazil(Bahia, Maranhão, Mato Grosso, Mato Grosso do Sul, Minas Gerais, Pará, and Piaui), Bolivia (Santa Cruz)
3091:
1084:
Bolivia (Beni and Pando
Departments), Brazil (Rondônia) and Peru (Departments of Ucayali and Madre de Dios)
2922:
1702:
2935:
2930:
2904:
2657:
2576:. 2007. A new species of arboreal Rhinella (Anura: Bufonidae) from a cloud forest of southeastern Peru.
2125:
northern Peru, eastern
Ecuador (Pastaza), Amazonian Colombia (Amazonas and Caquetá Departments), Brazil
2652:
3078:
3065:
3119:
2681:
54:
1530:
1416:
1026:
2686:
1821:
1499:
607:
3052:
1721:
Panama, Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Peru, Suriname and
Venezuela
2857:
2342:
2260:
2177:
1709:
3000:
2625:
3047:
2694:
2598:
818:
506:
3039:
429:
southeastern Brazil (eastern Paraná, eastern Santa
Catarina and northern Rio Grande do Sul)
2956:
2480:
2049:
1443:
1097:
920:
882:
857:
697:
587:
976:
8:
1617:
1294:
753:
Colombia east of the Andes and
Venezuela north of the Orinoco to French Guiana; Trinidad
442:
2536:
2895:
2845:
2798:
2298:
2158:
2024:
1314:
1226:
1162:
1053:
959:
893:
Brazil (Piauí, Bahia, Goiás, Mato Grosso and
Distrito Federal), Argentina (Corrientes)
486:
266:
229:
169:
49:
3114:
3034:
2943:
2802:
2790:
2755:
2675:
2604:
2460:
2318:
2216:
2133:
2114:
1899:
1879:
1859:
1550:
1339:
1142:
1033:
1008:
983:
632:
549:
467:
308:
2782:
2745:
2421:
2241:
2089:
2069:
1246:
1201:
2948:
2786:
2335:
2662:
2381:
1924:
1734:
1570:
1398:
1351:
northeastern Argentina (Misiones Province), southern Brazil and eastern Paraguay
1122:
901:
677:
523:
2909:
2880:
2197:
1045:
Argentina, Paraguay, Uruguay, eastern Bolivia, and eastern and southern Brazil
761:
709:
Chile between Paposo (Antofagasta Region) and Las Chilcas (Valparaíso Region)
319:
32:
3108:
2794:
2759:
2648:
2107:
1593:
1423:
1359:
418:
1727:
2252:
Paraguay (Amambay and Concepción Departments), Brazil (Mato Grosso do Sul)
1523:
995:
eastern Brazil between the states of Ceará in the north and Rio de Janeiro
139:
1986:Ávila, Morais, Perez, Pansonato, Carvalho, Rojas, Gordo & Farias, 2020
2982:
2889:
221:
644:
Argentina from the Chubut Province northward, Bolivia east of the Andes
541:
Western Panama through Chocoan western Colombia to northwestern Ecuador
287:
2974:
875:
314:
2750:
2733:
2630:(in French). Vol. 3. Bureau Principal de l'Éditeurs. p. 433.
2042:
1610:
1287:
573:(Rojas, Peña, Ávila, Carvalho, Perez, Farias, Gordo & Hrbek, 2022)
2987:
2446:
Rojas, Peña, Ávila, Carvalho, Perez, Farias, Gordo & Hrbek, 2022)
1917:
1219:
1090:
365:
351:
331:
299:
225:
217:
116:
96:
66:
2851:
1001:
619:
Peru (Mendoza, northern Cordillera Central, in the Amazonas Region)
2874:
2840:
2574:
Chaparro, Juan Carlos, Jennifer B. Pramuk and Andrew G. Gluesenkamp
2151:
2017:
1332:
945:(Castillo-Urbina, Glaw, Aguilar-Puntriano, Vences, and Köhler 2021)
86:
2234:
1752:
1115:
734:
670:
625:
460:
435:
3086:
2603:. Vol. 1. The Johns Hopkins University Press. p. 20.
411:
76:
2961:
224:
parts of Mexico, Central and South America. Additionally, the
3060:
2641:
372:
358:
338:
306:
292:
1678:
232:
to Australia, the Caribbean, the Philippines and elsewhere.
2709:"Rhinella Fitzinger, 1826 | Amphibian Species of the World"
241:
106:
386:
There are about 94 living species recognised in the genus
1065:
northeastern Argentina, Uruguay and southeastern Brazil
2367:(Padial, Reichle, McDiarmid, & De la Riva, 2006)
1590:
1468:(Köhler, Vences, Padial, Plewnia & Lötters 2023)
478:
Uruguay and southernmost Brazil (Rio Grande do Sul)
454:
northern Argentina (Córdoba and San Luis provinces)
2731:
2673:
2406:(Ferrão, Souza, Colatreli, Hanken & Lima, 2022)
2824:Bulletin of the American Museum of Natural History
2772:
2674:
1764:Fouquet, Gaucher, Blanc and Velez-Rodriguez, 2007
1511:Fouquet, Gaucher, Blanc and Velez-Rodriguez, 2007
3106:
1078:Ferrão, Lima, Ron, dos Santos & Hanken, 2020
1789:Venezuela (Bolívar), Suriname, Guyana, Brazil
1672:Argentina, Paraguay and Bolivia and in Brazil
1326:Colombia, Venezuela, Trinidad and the Guianas
2623:
2329:Bolivia (Cochabamba, La Paz and Santa Cruz)
2627:Dictionnaire universel d'histoire naturelle
2284:Vaz-Silva, Maciel, Bastos, and Pombal, 2015
1187:Vaz-Silva, Maciel, Bastos, and Pombal, 2015
964:(Vélez-Rodriguez & Ruiz-Carranza, 2002)
423:(Baldissera, Caramaschi & Haddad, 2004)
329:is the Latinised form of the Ancient Greek
297:), the combining form of the Ancient Greek
1653:
785:Sanabria, Quiroga, Arias, and Cortez, 2010
31:
2749:
906:(Lehr, Köhler, Aguilar & Ponce, 2001)
887:Maciel, Brandão, Campos, and Sebben, 2007
1109:Argentina, Brazil, Paraguay and Uruguay
970:Colombia(La Pedrera, Amazonas Province)
2524:Lehr, Pramuk, Hedges, and Córdova, 2007
2265:(Mijares-Urrutia & Arends-R., 2001)
1690:Chaparro, Pramuk, and Gluesenkamp, 2007
925:(McCranie, Wilson & Williams, 1989)
517:Brazil, Colombia, Panama and Venezuela
245:, then they were split into the genera
3107:
2822:(2006). "The Amphibian Tree of Life".
2592:
2590:
1967:Ávila, Pansonato, and Strüssmann, 2010
1535:Cusi, Moravec, Lehr, and Gvoždík, 2017
498:Serranía del Darién in eastern Panama
2856:
2855:
2600:Frogs of the United States and Canada
472:(Maneyro, Arrieta, & de Sá, 2004)
235:Originally, all species of the genus
3092:08C8E3C3-E384-430C-A363-07AC42611E1B
3066:827BAC65-DE1B-F781-8F4A-F059E7DC05C8
2695:participating institution membership
2596:
2246:(Caramaschi & de Niemeyer, 2003)
1384:Vaz-Silva, Valdujo, and Pombal, 2012
2587:
2505:(Brandão, Maciel, and Sebben, 2007)
2310:Argentina, Bolivia, Chile and Peru
2061:Brazil, Colombia, Ecuador and Peru
1826:(Lehr, Pramuk & Lundberg, 2005)
1746:South and mainland Central America
1020:Brazil, Colombia, Ecuador and Peru
13:
2775:Journal of Vertebrate Paleontology
1390:Minas Gerais, southeastern Brazil
14:
3146:
2833:
1598:Pérez-Ben, Gómez & Báez, 2019
349:, meaning “beaked toad”, is from
2839:
2535:
2334:
2233:
2150:
2106:
2041:
2016:
1916:
1794:
1751:
1726:
1701:
1677:
1609:
1522:
1498:
1415:
1331:
1286:
1262:
1218:
1114:
1089:
1025:
1000:
975:
874:
733:
669:
624:
599:Bolivia (Cochabamba Department)
522:
459:
434:
410:
53:
3135:Taxa named by Leopold Fitzinger
2410:Teotonio Falls Leaf-litter Toad
1948:Grant and Bolívar-Garcías, 2014
1275:(Caramaschi & Pombal, 2006)
804:Roberto, Brito, and Thomé, 2014
772:Argentina, Brazil and Paraguay
2766:
2725:
2701:
2667:
2634:
2617:
2567:
2492:Bolivia and southeastern Peru
2465:(Leviton & Duellman, 1978)
612:(Duellman & Schulte, 1992)
1:
3125:Amphibians of Central America
2813:
2787:10.1080/02724634.2019.1576183
1642:Lima, Menin, and Araújo, 2007
2323:(Lötters & Köhler, 2000)
2074:(Myers & Carvalho, 1952)
1851:Venezuela (Bolivar), Guyana
1575:(Rivero & Castaño, 1990)
1539:Lily Rodriguez's beaked toad
1403:(Jiménez de la Espada, 1875)
1306:north-western South America
1058:(Duméril & Bibron, 1841)
1013:(Myers & Carvalho, 1945)
366:
352:
332:
300:
276:
7:
3130:Amphibians of South America
2624:d'Orbigny, Charles (1845).
2426:(Lynch & Renjifo, 1990)
2386:(Lynch & Renjifo, 1990)
1864:(Duellman & Toft, 1979)
1666:(Müller and Hellmich, 1936)
843:Narvaes and Rodrigues, 2009
706:Vallenar toad; Atacama toad
641:Common toad; Argentine toad
239:were included in the genus
10:
3151:
2642:
2036:Ecuador, Peru and Bolivia
1884:(Cochran & Goin, 1970)
1718:South American common toad
1448:(Harvey & Smith, 1994)
1131:Valle Santiago beaked toad
592:(Harvey & Smith, 1993)
554:(Pramuk and Kadivar, 2003)
381:
373:
359:
339:
307:
293:
285:means ‘little nose’, from
2864:
2682:Oxford English Dictionary
2597:Dodd, C. Kenneth (2013).
313:, ‘nose’) and the Latin
175:
168:
161:
156:
145:
138:
50:Scientific classification
48:
39:
30:
23:
2560:
2511:Brazil(Piauí and Bahia)
2307:Warty toad; Huanuco toad
1803:Rhinella mirandaribeiroi
1531:Rhinella lilyrodriguezae
1517:French Guiana, Suriname
1042:Cope's toad; cururu toad
399:Binomial name and author
2687:Oxford University Press
2658:A Greek–English Lexicon
2354:Colombia and Venezuela
2142:Mesopotamia beaked toad
1822:Rhinella multiverrucosa
686:Arunco; Concepcion toad
608:Rhinella arborescandens
560:Loja Province, Ecuador
2548:(Mordavec, Lehr, 2014)
2343:Rhinella sternosignata
2261:Rhinella sclerocephala
2178:Rhinella rubropunctata
1963:Rhinella paraguayensis
1710:Rhinella margaritifera
1626:Santa Rita beaked toad
495:Cerro Mali beaked toad
163:95 species; see table
3048:Paleobiology Database
2649:Liddell, Henry George
2402:Rhinella teotoniensis
2390:Antioquia Beaked Toad
2208:Colombia (Antioquia)
2145:Colombia (Antioquia)
1891:Colombia (Antioquia)
1888:Colombian beaked toad
819:Rhinella castaneotica
653:Rhinella arequipensis
507:Rhinella acutirostris
2481:Rhinella veraguensis
2189:Chile and Argentina
2050:Rhinella proboscidea
2001:Rhinella pleuroptera
1483:Rhinella leptoscelis
1444:Rhinella justinianoi
1098:Rhinella fernandezae
988:(Wied-Neuwied, 1821)
941:Rhinella chullachaki
921:Rhinella chrysophora
883:Rhinella cerradensis
858:Rhinella ceratophrys
698:Rhinella atacamensis
588:Rhinella amboroensis
343:, ‘porous, spongy’).
206:South American toads
204:, commonly known as
147:Rhinella proboscidea
2685:(Online ed.).
2430:Trueb's Beaked Toad
1638:Rhinella magnussoni
1618:Rhinella macrorhina
1295:Rhinella horribilis
1074:Rhinella exostosica
443:Rhinella achalensis
265:are now considered
16:Genus of amphibians
2520:Rhinella yanachaga
2442:Rhinella unapensis
2299:Rhinella spinulosa
2159:Rhinella rubescens
2025:Rhinella poeppigii
1841:Rhinella nattereri
1348:Yellow Cururu toad
1315:Rhinella humboldti
1271:Rhinella hoogmoedi
1235:Common lesser toad
1227:Rhinella granulosa
1163:Rhinella gallardoi
1054:Rhinella dorbignyi
960:Rhinella cristinae
849:Panama (Chiriquí)
839:Rhinella centralis
781:Rhinella bernardoi
487:Rhinella acrolopha
3102:
3101:
3035:Open Tree of Life
2858:Taxon identifiers
2751:10.1643/CH2020043
2713:research.amnh.org
2693:(Subscription or
2610:978-1-4214-0633-6
2558:
2557:
2469:Alto Maranon toad
2461:Rhinella vellardi
2373:Bolivia (La Paz)
2319:Rhinella stanlaii
2217:Rhinella rumbolli
2182:(Guichenot, 1848)
2134:Rhinella rostrata
2115:Rhinella roqueana
2078:Rio Parahyba toad
1954:Colombia (Chocó)
1944:Rhinella paraguas
1900:Rhinella ocellata
1880:Rhinella nicefori
1860:Rhinella nesiotes
1845:(Bokermann, 1967)
1779:Rhinella merianae
1579:Murri beaked toad
1551:Rhinella limensis
1507:Rhinella lescurei
1487:(Boulenger, 1912)
1364:(Stejneger, 1913)
1340:Rhinella icterica
1154:Bolivia and Peru
1147:(Boulenger, 1903)
1143:Rhinella fissipes
1134:Ecuador and Peru
1034:Rhinella diptycha
1009:Rhinella dapsilis
984:Rhinella crucifer
862:(Boulenger, 1882)
633:Rhinella arenarum
616:Mendoza Pass toad
550:Rhinella amabilis
468:Rhinella achavali
197:
196:
134:
3142:
3120:Amphibian genera
3095:
3094:
3082:
3081:
3069:
3068:
3056:
3055:
3043:
3042:
3030:
3029:
3017:
3016:
3004:
3003:
2991:
2990:
2978:
2977:
2965:
2964:
2952:
2951:
2939:
2938:
2926:
2925:
2913:
2912:
2900:
2899:
2898:
2885:
2884:
2883:
2853:
2852:
2844:Data related to
2843:
2821:
2807:
2806:
2770:
2764:
2763:
2753:
2729:
2723:
2722:
2720:
2719:
2705:
2699:
2698:
2690:
2678:
2671:
2665:
2645:
2644:
2638:
2632:
2631:
2621:
2615:
2614:
2594:
2585:
2575:
2571:
2549:
2539:
2525:
2506:
2501:Rhinella veredas
2486:
2466:
2447:
2427:
2422:Rhinella truebae
2413:Bolivia, Brazil
2407:
2387:
2368:
2348:
2338:
2324:
2304:
2303:(Wiegmann, 1834)
2285:
2280:Rhinella sebbeni
2266:
2247:
2242:Rhinella scitula
2237:
2222:
2203:
2183:
2164:
2154:
2139:
2120:
2110:
2095:
2094:(Gallardo, 1961)
2090:Rhinella quechua
2075:
2070:Rhinella pygmaea
2055:
2045:
2030:
2020:
2006:
1987:
1982:Rhinella parecis
1968:
1949:
1930:
1920:
1905:
1885:
1865:
1846:
1827:
1808:
1807:(Gallardo, 1965)
1798:
1784:
1783:(Gallardo, 1965)
1765:
1755:
1740:
1739:(Linnaeus, 1758)
1730:
1715:
1714:(Laurenti, 1768)
1705:
1691:
1681:
1667:
1657:
1643:
1623:
1613:
1599:
1592:
1576:
1556:
1536:
1526:
1512:
1502:
1488:
1469:
1449:
1429:
1419:
1404:
1385:
1380:Rhinella inopina
1365:
1345:
1335:
1320:
1319:(Gallardo, 1965)
1300:
1299:(Wiegmann, 1833)
1290:
1276:
1266:
1252:
1247:Rhinella henseli
1232:
1222:
1207:
1206:(Gallardo, 1967)
1202:Rhinella gnustae
1188:
1168:
1148:
1128:
1118:
1106:Bella Vista toad
1103:
1102:(Gallardo, 1957)
1093:
1079:
1059:
1039:
1029:
1014:
1004:
989:
979:
965:
946:
926:
907:
888:
878:
863:
844:
824:
823:(Caldwell, 1991)
805:
800:Rhinella casconi
786:
767:
766:(Céspedez, 2000)
747:
746:(Gallardo, 1965)
737:
723:
722:(Gallardo, 1965)
703:
683:
673:
658:
638:
628:
613:
593:
574:
555:
536:
535:(Thominot, 1884)
526:
512:
492:
473:
463:
448:
438:
424:
414:
393:
392:
376:
375:
369:
362:
361:
355:
342:
341:
335:
312:
311:
303:
296:
295:
216:, is a genus of
191:
183:
132:
58:
57:
35:
21:
20:
3150:
3149:
3145:
3144:
3143:
3141:
3140:
3139:
3105:
3104:
3103:
3098:
3090:
3085:
3077:
3072:
3064:
3059:
3051:
3046:
3038:
3033:
3025:
3020:
3012:
3007:
2999:
2994:
2986:
2981:
2973:
2968:
2960:
2955:
2947:
2942:
2934:
2929:
2921:
2916:
2908:
2903:
2894:
2893:
2888:
2879:
2878:
2873:
2860:
2836:
2819:
2816:
2811:
2810:
2771:
2767:
2730:
2726:
2717:
2715:
2707:
2706:
2702:
2692:
2672:
2668:
2663:Perseus Project
2639:
2635:
2622:
2618:
2611:
2595:
2588:
2573:
2572:
2568:
2563:
2547:
2523:
2504:
2485:(Schmidt, 1857)
2484:
2464:
2445:
2425:
2405:
2385:
2382:Rhinella tenrec
2366:
2363:Rhinella tacana
2347:(Günther, 1858)
2346:
2322:
2302:
2283:
2264:
2245:
2221:(Carrizo, 1992)
2220:
2201:
2181:
2162:
2137:
2118:
2093:
2073:
2053:
2029:(Tschudi, 1845)
2028:
2004:
1985:
1966:
1947:
1928:
1925:Rhinella ornata
1904:(Günther, 1858)
1903:
1883:
1863:
1844:
1825:
1806:
1782:
1763:
1760:Rhinella martyi
1738:
1735:Rhinella marina
1713:
1689:
1665:
1641:
1621:
1597:
1574:
1571:Rhinella lindae
1559:Peru Coast toad
1554:
1534:
1510:
1486:
1467:
1447:
1428:(Stevaux, 2002)
1427:
1407:Rio Perene toad
1402:
1399:Rhinella iserni
1383:
1363:
1343:
1318:
1298:
1274:
1250:
1230:
1210:Rio Grande toad
1205:
1186:
1183:Rhinella gildae
1171:Gallardo's toad
1167:(Carrizo, 1992)
1166:
1146:
1127:(Peracca, 1904)
1126:
1123:Rhinella festae
1101:
1077:
1062:Dorbigny's toad
1057:
1037:
1017:Bom Jardim toad
1012:
987:
963:
951:Peru (Huánuco)
944:
924:
912:Peru (Huánuco)
905:
902:Rhinella chavin
886:
861:
842:
822:
810:Brazil (Ceará)
803:
784:
765:
745:
742:Rhinella beebei
721:
718:Rhinella azarai
701:
681:
678:Rhinella arunco
657:(Vellard, 1959)
656:
636:
611:
596:Cochabamba toad
591:
572:
569:Rhinella angeli
553:
534:
510:
490:
471:
446:
422:
384:
279:
214:Rio Viejo toads
189:
181:
152:
149:
133:Fitzinger, 1826
131:
52:
42:Rhinella marina
17:
12:
11:
5:
3148:
3138:
3137:
3132:
3127:
3122:
3117:
3100:
3099:
3097:
3096:
3083:
3070:
3057:
3044:
3031:
3018:
3005:
2992:
2979:
2966:
2953:
2940:
2927:
2914:
2901:
2886:
2870:
2868:
2862:
2861:
2850:
2849:
2848:at Wikispecies
2835:
2834:External links
2832:
2831:
2830:
2815:
2812:
2809:
2808:
2765:
2744:(4): 967–986.
2724:
2700:
2666:
2633:
2616:
2609:
2586:
2565:
2564:
2562:
2559:
2556:
2555:
2554:Peru (Yungas)
2552:
2550:
2544:Rhinella yunga
2540:
2532:
2531:
2528:
2526:
2516:
2513:
2512:
2509:
2507:
2497:
2494:
2493:
2490:
2487:
2477:
2474:
2473:
2470:
2467:
2457:
2454:
2453:
2450:
2448:
2438:
2435:
2434:
2431:
2428:
2418:
2415:
2414:
2411:
2408:
2398:
2395:
2394:
2391:
2388:
2378:
2375:
2374:
2371:
2369:
2359:
2356:
2355:
2352:
2349:
2339:
2331:
2330:
2327:
2325:
2315:
2312:
2311:
2308:
2305:
2295:
2292:
2291:
2288:
2286:
2276:
2273:
2272:
2269:
2267:
2257:
2254:
2253:
2250:
2248:
2238:
2230:
2229:
2226:
2223:
2213:
2210:
2209:
2206:
2204:
2198:Rhinella ruizi
2194:
2191:
2190:
2187:
2184:
2174:
2171:
2170:
2167:
2165:
2155:
2147:
2146:
2143:
2140:
2130:
2127:
2126:
2123:
2121:
2111:
2103:
2102:
2099:
2098:Incachaca toad
2096:
2086:
2083:
2082:
2079:
2076:
2066:
2063:
2062:
2059:
2056:
2046:
2038:
2037:
2034:
2031:
2021:
2013:
2012:
2009:
2007:
1997:
1994:
1993:
1990:
1988:
1978:
1975:
1974:
1971:
1969:
1959:
1956:
1955:
1952:
1950:
1940:
1937:
1936:
1933:
1931:
1921:
1913:
1912:
1909:
1908:Ocellated toad
1906:
1896:
1893:
1892:
1889:
1886:
1876:
1873:
1872:
1869:
1866:
1856:
1853:
1852:
1849:
1847:
1837:
1834:
1833:
1830:
1828:
1818:
1815:
1814:
1811:
1809:
1799:
1791:
1790:
1787:
1785:
1775:
1772:
1771:
1768:
1766:
1756:
1748:
1747:
1744:
1741:
1731:
1723:
1722:
1719:
1716:
1706:
1698:
1697:
1694:
1692:
1682:
1674:
1673:
1670:
1668:
1662:Rhinella major
1658:
1650:
1649:
1646:
1644:
1634:
1631:
1630:
1627:
1624:
1614:
1606:
1605:
1602:
1600:
1587:
1584:
1583:
1580:
1577:
1567:
1564:
1563:
1560:
1557:
1555:(Werner, 1901)
1547:
1544:
1543:
1540:
1537:
1527:
1519:
1518:
1515:
1513:
1503:
1495:
1494:
1491:
1489:
1479:
1476:
1475:
1472:
1470:
1460:
1457:
1456:
1453:
1450:
1440:
1437:
1436:
1433:
1430:
1420:
1412:
1411:
1408:
1405:
1395:
1392:
1391:
1388:
1386:
1376:
1373:
1372:
1369:
1366:
1356:
1353:
1352:
1349:
1346:
1336:
1328:
1327:
1324:
1321:
1311:
1308:
1307:
1304:
1301:
1291:
1283:
1282:
1279:
1277:
1267:
1259:
1258:
1255:
1253:
1243:
1240:
1239:
1236:
1233:
1223:
1215:
1214:
1211:
1208:
1198:
1195:
1194:
1191:
1189:
1179:
1176:
1175:
1172:
1169:
1159:
1156:
1155:
1152:
1149:
1139:
1136:
1135:
1132:
1129:
1119:
1111:
1110:
1107:
1104:
1094:
1086:
1085:
1082:
1080:
1070:
1067:
1066:
1063:
1060:
1050:
1047:
1046:
1043:
1040:
1030:
1022:
1021:
1018:
1015:
1005:
997:
996:
993:
990:
980:
972:
971:
968:
966:
956:
953:
952:
949:
947:
937:
934:
933:
930:
929:Rio Viejo toad
927:
917:
914:
913:
910:
908:
898:
895:
894:
891:
889:
879:
871:
870:
867:
864:
854:
851:
850:
847:
845:
835:
832:
831:
828:
825:
815:
812:
811:
808:
806:
796:
793:
792:
789:
787:
777:
774:
773:
770:
768:
762:Rhinella bergi
758:
755:
754:
751:
748:
738:
730:
729:
726:
724:
714:
711:
710:
707:
704:
694:
691:
690:
687:
684:
682:(Molina, 1782)
674:
666:
665:
662:
661:Rio Chili toad
659:
649:
646:
645:
642:
639:
637:(Hensel, 1867)
629:
621:
620:
617:
614:
604:
601:
600:
597:
594:
584:
581:
580:
577:
575:
565:
562:
561:
558:
556:
546:
543:
542:
539:
537:
531:Rhinella alata
527:
519:
518:
515:
513:
503:
500:
499:
496:
493:
483:
480:
479:
476:
474:
464:
456:
455:
452:
449:
439:
431:
430:
427:
425:
415:
407:
406:
403:
400:
397:
383:
380:
379:
378:
363:, ‘beak’) and
344:
324:
278:
275:
195:
194:
193:
192:
184:
173:
172:
166:
165:
159:
158:
154:
153:
150:
143:
142:
136:
135:
124:
120:
119:
114:
110:
109:
104:
100:
99:
94:
90:
89:
84:
80:
79:
74:
70:
69:
64:
60:
59:
46:
45:
37:
36:
28:
27:
15:
9:
6:
4:
3:
2:
3147:
3136:
3133:
3131:
3128:
3126:
3123:
3121:
3118:
3116:
3113:
3112:
3110:
3093:
3088:
3084:
3080:
3075:
3071:
3067:
3062:
3058:
3054:
3049:
3045:
3041:
3036:
3032:
3028:
3023:
3019:
3015:
3010:
3006:
3002:
2997:
2993:
2989:
2984:
2980:
2976:
2971:
2967:
2963:
2958:
2954:
2950:
2945:
2941:
2937:
2932:
2928:
2924:
2919:
2915:
2911:
2906:
2902:
2897:
2891:
2887:
2882:
2876:
2872:
2871:
2869:
2867:
2863:
2859:
2854:
2847:
2842:
2838:
2837:
2828:
2825:
2820:Frost, Darrel
2818:
2817:
2804:
2800:
2796:
2792:
2788:
2784:
2780:
2776:
2769:
2761:
2757:
2752:
2747:
2743:
2739:
2735:
2728:
2714:
2710:
2704:
2696:
2688:
2684:
2683:
2677:
2670:
2664:
2660:
2659:
2654:
2653:Scott, Robert
2650:
2646:
2637:
2629:
2628:
2620:
2612:
2606:
2602:
2601:
2593:
2591:
2584:(2): 203-212.
2583:
2579:
2578:Herpetologica
2570:
2566:
2553:
2551:
2546:
2545:
2541:
2538:
2534:
2533:
2530:Peru (Pasco)
2529:
2527:
2522:
2521:
2517:
2515:
2514:
2510:
2508:
2503:
2502:
2498:
2496:
2495:
2491:
2488:
2483:
2482:
2478:
2476:
2475:
2471:
2468:
2463:
2462:
2458:
2456:
2455:
2451:
2449:
2444:
2443:
2439:
2437:
2436:
2432:
2429:
2424:
2423:
2419:
2417:
2416:
2412:
2409:
2404:
2403:
2399:
2397:
2396:
2392:
2389:
2384:
2383:
2379:
2377:
2376:
2372:
2370:
2365:
2364:
2360:
2358:
2357:
2353:
2350:
2345:
2344:
2340:
2337:
2333:
2332:
2328:
2326:
2321:
2320:
2316:
2314:
2313:
2309:
2306:
2301:
2300:
2296:
2294:
2293:
2289:
2287:
2282:
2281:
2277:
2275:
2274:
2270:
2268:
2263:
2262:
2258:
2256:
2255:
2251:
2249:
2244:
2243:
2239:
2236:
2232:
2231:
2227:
2224:
2219:
2218:
2214:
2212:
2211:
2207:
2205:
2202:(Grant, 2000)
2200:
2199:
2195:
2193:
2192:
2188:
2185:
2180:
2179:
2175:
2173:
2172:
2168:
2166:
2161:
2160:
2156:
2153:
2149:
2148:
2144:
2141:
2138:(Noble, 1920)
2136:
2135:
2131:
2129:
2128:
2124:
2122:
2119:(Melin, 1941)
2117:
2116:
2112:
2109:
2105:
2104:
2100:
2097:
2092:
2091:
2087:
2085:
2084:
2080:
2077:
2072:
2071:
2067:
2065:
2064:
2060:
2057:
2052:
2051:
2047:
2044:
2040:
2039:
2035:
2032:
2027:
2026:
2022:
2019:
2015:
2014:
2010:
2008:
2005:Schmidt, 1857
2003:
2002:
1998:
1996:
1995:
1991:
1989:
1984:
1983:
1979:
1977:
1976:
1972:
1970:
1965:
1964:
1960:
1958:
1957:
1953:
1951:
1946:
1945:
1941:
1939:
1938:
1934:
1932:
1927:
1926:
1922:
1919:
1915:
1914:
1910:
1907:
1902:
1901:
1897:
1895:
1894:
1890:
1887:
1882:
1881:
1877:
1875:
1874:
1870:
1867:
1862:
1861:
1857:
1855:
1854:
1850:
1848:
1843:
1842:
1838:
1836:
1835:
1832:Peru (Pasco)
1831:
1829:
1824:
1823:
1819:
1817:
1816:
1812:
1810:
1805:
1804:
1800:
1797:
1793:
1792:
1788:
1786:
1781:
1780:
1776:
1774:
1773:
1769:
1767:
1762:
1761:
1757:
1754:
1750:
1749:
1745:
1742:
1737:
1736:
1732:
1729:
1725:
1724:
1720:
1717:
1712:
1711:
1707:
1704:
1700:
1699:
1695:
1693:
1688:
1687:
1686:Rhinella manu
1683:
1680:
1676:
1675:
1671:
1669:
1664:
1663:
1659:
1656:
1652:
1651:
1647:
1645:
1640:
1639:
1635:
1633:
1632:
1628:
1625:
1622:(Trueb, 1971)
1620:
1619:
1615:
1612:
1608:
1607:
1603:
1601:
1596:
1595:
1594:Rhinella loba
1588:
1586:
1585:
1581:
1578:
1573:
1572:
1568:
1566:
1565:
1561:
1558:
1553:
1552:
1548:
1546:
1545:
1541:
1538:
1533:
1532:
1528:
1525:
1521:
1520:
1516:
1514:
1509:
1508:
1504:
1501:
1497:
1496:
1492:
1490:
1485:
1484:
1480:
1478:
1477:
1473:
1471:
1466:
1465:
1464:Rhinella kuka
1461:
1459:
1458:
1454:
1452:El Chape toad
1451:
1446:
1445:
1441:
1439:
1438:
1434:
1431:
1426:
1425:
1424:Rhinella jimi
1421:
1418:
1414:
1413:
1409:
1406:
1401:
1400:
1396:
1394:
1393:
1389:
1387:
1382:
1381:
1377:
1375:
1374:
1370:
1367:
1362:
1361:
1360:Rhinella inca
1357:
1355:
1354:
1350:
1347:
1342:
1341:
1337:
1334:
1330:
1329:
1325:
1323:Rivero's toad
1322:
1317:
1316:
1312:
1310:
1309:
1305:
1302:
1297:
1296:
1292:
1289:
1285:
1284:
1280:
1278:
1273:
1272:
1268:
1265:
1261:
1260:
1256:
1254:
1249:
1248:
1244:
1242:
1241:
1237:
1234:
1229:
1228:
1224:
1221:
1217:
1216:
1212:
1209:
1204:
1203:
1199:
1197:
1196:
1192:
1190:
1185:
1184:
1180:
1178:
1177:
1173:
1170:
1165:
1164:
1160:
1158:
1157:
1153:
1151:Carabaya toad
1150:
1145:
1144:
1140:
1138:
1137:
1133:
1130:
1125:
1124:
1120:
1117:
1113:
1112:
1108:
1105:
1100:
1099:
1095:
1092:
1088:
1087:
1083:
1081:
1076:
1075:
1071:
1069:
1068:
1064:
1061:
1056:
1055:
1051:
1049:
1048:
1044:
1041:
1036:
1035:
1031:
1028:
1024:
1023:
1019:
1016:
1011:
1010:
1006:
1003:
999:
998:
994:
991:
986:
985:
981:
978:
974:
973:
969:
967:
962:
961:
957:
955:
954:
950:
948:
943:
942:
938:
936:
935:
931:
928:
923:
922:
918:
916:
915:
911:
909:
904:
903:
899:
897:
896:
892:
890:
885:
884:
880:
877:
873:
872:
868:
865:
860:
859:
855:
853:
852:
848:
846:
841:
840:
836:
834:
833:
829:
826:
821:
820:
816:
814:
813:
809:
807:
802:
801:
797:
795:
794:
790:
788:
783:
782:
778:
776:
775:
771:
769:
764:
763:
759:
757:
756:
752:
750:Rivero's Toad
749:
744:
743:
739:
736:
732:
731:
727:
725:
720:
719:
715:
713:
712:
708:
705:
700:
699:
695:
693:
692:
688:
685:
680:
679:
675:
672:
668:
667:
663:
660:
655:
654:
650:
648:
647:
643:
640:
635:
634:
630:
627:
623:
622:
618:
615:
610:
609:
605:
603:
602:
598:
595:
590:
589:
585:
583:
582:
578:
576:
571:
570:
566:
564:
563:
559:
557:
552:
551:
547:
545:
544:
540:
538:
533:
532:
528:
525:
521:
520:
516:
514:
509:
508:
504:
502:
501:
497:
494:
491:(Trueb, 1971)
489:
488:
484:
482:
481:
477:
475:
470:
469:
465:
462:
458:
457:
453:
450:
445:
444:
440:
437:
433:
432:
428:
426:
421:
420:
419:Rhinella abei
416:
413:
409:
408:
405:Distribution
404:
401:
398:
395:
394:
391:
389:
370:
368:
356:
354:
348:
347:Rhamphophryne
345:
336:
334:
328:
325:
322:
321:
316:
310:
304:
302:
290:
289:
284:
281:
280:
274:
272:
268:
264:
263:Rhamphophryne
260:
256:
255:
254:Rhamphophryne
250:
249:
244:
243:
238:
233:
231:
227:
223:
219:
215:
211:
207:
203:
202:
188:
187:Rhamphophryne
185:
180:
177:
176:
174:
171:
167:
164:
160:
155:
148:
144:
141:
137:
130:
129:
125:
122:
121:
118:
115:
112:
111:
108:
105:
102:
101:
98:
95:
92:
91:
88:
85:
82:
81:
78:
75:
72:
71:
68:
65:
62:
61:
56:
51:
47:
43:
38:
34:
29:
26:
22:
19:
2865:
2826:
2823:
2781:: e1576183.
2778:
2774:
2768:
2741:
2737:
2727:
2716:. Retrieved
2712:
2703:
2680:
2669:
2656:
2636:
2626:
2619:
2599:
2581:
2577:
2569:
2542:
2518:
2499:
2489:Veragua toad
2479:
2459:
2440:
2420:
2400:
2380:
2361:
2341:
2317:
2297:
2278:
2259:
2240:
2215:
2196:
2176:
2163:(Lutz, 1925)
2157:
2132:
2113:
2088:
2068:
2054:(Spix, 1824)
2048:
2023:
1999:
1980:
1961:
1942:
1923:
1898:
1878:
1858:
1839:
1820:
1801:
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1569:
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1397:
1378:
1358:
1344:(Spix, 1824)
1338:
1313:
1293:
1269:
1251:(Lutz, 1934)
1245:
1231:(Spix, 1824)
1225:
1200:
1181:
1161:
1141:
1121:
1096:
1072:
1052:
1038:(Cope, 1862)
1032:
1007:
992:Striped toad
982:
958:
939:
919:
900:
881:
856:
837:
817:
798:
779:
760:
740:
716:
696:
676:
651:
631:
606:
586:
567:
548:
529:
511:(Spix, 1824)
505:
485:
466:
451:Cordoba toad
441:
417:
387:
385:
364:
350:
346:
330:
326:
318:
298:
286:
282:
270:
262:
258:
253:
252:
247:
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240:
236:
234:
213:
210:beaked toads
209:
205:
200:
199:
198:
186:
182:Wagler, 1828
178:
162:
146:
140:Type species
127:
126:
41:
24:
18:
2983:iNaturalist
2890:Wikispecies
2676:"rhamphoid"
2351:Falcon toad
2058:Beaked toad
1868:Laguna toad
1432:Cururu toad
866:Horned toad
702:(Cei, 1962)
447:(Cei, 1972)
402:Common name
257:. However,
222:Neotropical
190:Trueb, 1971
40:Cane toad (
3109:Categories
2814:References
2718:2019-10-26
2697:required.)
2271:Venezuela
2225:Salta toad
2186:Rusty toad
1929:Spix, 1824
1604:Argentina
1213:Argentina
1174:Argentina
791:Argentina
377:, ‘toad’).
315:diminutive
230:Introduced
220:native to
218:true toads
151:Spix, 1824
2803:133110392
2795:0272-4634
2760:0045-8511
2433:Colombia
2393:Colombia
2033:Gray toad
1973:Paraguay
1770:Suriname
1743:Cane toad
1629:Colombia
1582:Colombia
1368:Inca toad
1303:Cane toad
932:Honduras
827:Para toad
728:Paraguay
277:Etymology
228:has been
226:cane toad
117:Bufonidae
73:Kingdom:
67:Eukaryota
3115:Rhinella
2923:Rhinella
2910:Rhinella
2896:Rhinella
2881:Q2363065
2875:Wikidata
2866:Rhinella
2846:Rhinella
2829:: 1–371.
2228:Bolivia
2101:Bolivia
1474:Bolivia
1455:Bolivia
388:Rhinella
353:rhamphos
283:Rhinella
271:Rhinella
267:synonyms
237:Rhinella
201:Rhinella
170:Synonyms
157:Species
128:Rhinella
113:Family:
97:Amphibia
87:Chordata
83:Phylum:
77:Animalia
63:Domain:
25:Rhinella
3087:ZooBank
3079:1432724
3001:1408174
2975:2422625
2661:at the
2290:Brazil
2169:Brazil
2081:Brazil
1992:Brazil
1935:Brazil
1911:Brazil
1648:Brazil
1281:Brazil
1257:Brazil
1238:Brazil
1193:Brazil
382:Species
333:chaûnos
327:Chaunus
317:suffix
259:Chaunus
248:Chaunus
179:Chaunus
123:Genus:
103:Order:
93:Class:
3053:379331
3040:909159
3027:651667
3014:773237
2936:112082
2801:
2793:
2758:
2738:Copeia
2643:χαῦνος
2607:
689:Chile
367:phrunē
360:ῥάµϕος
340:χαῦνος
288:rhino-
3074:WoRMS
3061:Plazi
2996:IRMNG
2988:64819
2962:43313
2799:S2CID
2691:
2561:Notes
2472:Peru
2452:Peru
2011:Peru
1871:Peru
1696:Peru
1562:Peru
1542:Peru
1493:Peru
1410:Peru
1371:Peru
664:Peru
579:Peru
396:Image
374:φρύνη
320:-ella
294:ῥῑνο-
107:Anura
3022:NCBI
3009:ITIS
2970:GBIF
2949:78MK
2931:BOLD
2791:ISSN
2756:ISSN
2605:ISBN
301:rhis
261:and
251:and
242:Bufo
2957:EoL
2944:CoL
2918:ASW
2905:AFD
2827:297
2783:doi
2746:doi
2742:108
309:ῥίς
269:of
212:or
3111::
3089::
3076::
3063::
3050::
3037::
3024::
3011::
2998::
2985::
2972::
2959::
2946::
2933::
2920::
2907::
2892::
2877::
2797:.
2789:.
2779:39
2777:.
2754:.
2740:.
2736:.
2711:.
2679:.
2655:;
2651:;
2647:.
2589:^
2582:63
2580:.
390::
273:.
208:,
44:)
2805:.
2785::
2762:.
2748::
2721:.
2689:.
2613:.
1591:†
371:(
357:(
337:(
323:.
305:(
291:(
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