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Schizothorax

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Yang, J.; J.X. Yang; and X.Y. Chen (2012). A re-examination of the molecular phylogeny and biogeography of the genus Schizothorax (Teleostei: Cyprinidae) through enhanced sampling, with emphasis on the species in the Yunnan–Guizhou Plateau, China. Journal of Zoological Systematics and Evolutionary
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Qi, D.; Y. Chao; S. Guo; L. Zhao; T. Li; F. Wei; and X. Zhao (2012). Convergent, Parallel and Correlated Evolution of Trophic Morphologies in the Subfamily Schizothoracinae from the Qinghai-Tibetan Plateau. PLoS ONE 7(3): e34070.
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Coad, B.W.; and N.K.d. Ville (2004). On the systematics and distribution of the snow trout Schizothorax pelzami Kessler, 1870, in Iran (Actinopterygii: Cyprinidae). Zoology in the Middle East 32(1): 57-62.
1139: 941: 868: 811: 769: 533: 1067: 571: 1385: 567: 1354: 1291: 562: 1234: 1170: 1312: 1122: 1105: 990: 1299: 1183: 1126: 1053: 1017: 945: 901: 773: 692: 666: 1780: 1412: 1118: 977: 897: 585: 794: 602: 1806: 1345: 798: 598: 391:. The last two were formerly often included in the present genus, and their delimitation is still not entirely clear. In any case, the genus 1341: 1295: 905: 217:, in Turkey. They are primarily found in highland rivers, streams and lakes, although a few species occur in lower-lying locations, like 1754: 1793: 1798: 1629: 1577:
De Graaf, M.; Megens, H.-J.; Samallo, J.; Sibbing, F.A. (2007). "Evolutionary origin of Lake Tana's (Ethiopia) small
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is a part of the schizothoracines (snowtrout and allies), which also includes the genera
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Hammer, U.T. (1986). Saline Lake Ecosystems of the World, p. 458. DR.W.Junk Publishers.
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species: indications of rapid ecological divergence and speciation".
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at least for the largest part (including the marinkas), becoming its
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International Journal of Fauna and Biological Studies, 3 (2): 65-70.
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and they are by no means closely related apart from both being
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Yang, J., Zheng, L.-P., Chen, X.-Y. & Yang, J.-X. (2013):
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fish found in southern and western China, through northern
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Freshwater Ecoregions of the World. Retrieved 18 May 2019.
317:. Their ancestors must thus have diverged as early as the 509:
There are currently 65 recognized species in this genus:
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distributed in the Irrawaddy drainage area in China.
1647:Javed, M.N., Azizullah & Pervaiz, K. (2012): 1852: 286:), while the eastern species are usually called 1630:Description of two new species and revision of 1609:Arunkumar, L. & Alphonsa Moyon, W. (2016): 281: 275: 1603: 1624: 1622: 1492: 1490: 1461: 1459: 1457: 31: 1641: 1570: 496: 484: 472: 1655:Biologia (Pakistan), 58 (1-2): 175-178. 1619: 1487: 1465: 1454: 1853: 457:however and thus here merged with the 1667: 1666: 1637:Zoological Research, 34 (4): 361-367. 1553: 313:, while trouts are in the superorder 13: 213:, to Iran, with a single species, 14: 1882: 1503:10.1111/j.1439-0469.2012.00661.x 51: 1546:Coad, B.; and J. Hales (2008). 446:. They were thus placed in the 306:: Cyprinids are in the teleost 180:McClelland & Griffith, 1842 1540: 1524: 1520:10.1080/09397140.2004.10638044 1507: 1337:Schizothorax rotundimaxillaris 807:Schizothorax heterophysallidos 1: 1447: 1272:Schizothorax pseudoaksaiensis 264:are typically referred to as 1566:10.1371/journal.pone.0034070 841:Schizothorax integrilabiatus 410:and the doubtfully distinct 290:. Although they do resemble 246: 234: 229:means "cloven-breast", from 7: 986:Schizothorax macrophthalmus 276: 256:) 'breast-plate' (see also 10: 1887: 1597:10.1163/157075607780002069 1114:Schizothorax ninglangensis 1036:Schizothorax microcephalus 962:Schizothorax lissolabiatus 914:Schizothorax lantsangensis 728:Schizothorax gongshanensis 625:Schizothorax curvilabiatus 468: 252: 240: 1675: 1407:Schizothorax wangchiachii 1322:Schizothorax richardsonii 1216:Schizothorax plagiostomus 1156:Schizothorax nukiangensis 1026:Schizothorax meridionalis 924:Schizothorax lepidothorax 790:Schizothorax heterochilus 491:Schizothorax plagiostomus 282: 174: 167: 158:Schizothorax plagiostomus 156: 149: 48:Scientific classification 46: 39: 30: 23: 1469:; Pauly, Daniel (eds.). 1421:Schizothorax yunnanensis 1381:Schizothorax skarduensis 1135:Schizothorax nudiventris 1062:Schizothorax molesworthi 1049:Schizothorax microstomus 1013:Schizothorax malacanthus 972:Schizothorax longibarbus 864:Schizothorax kumaonensis 662:Schizothorax dulongensis 649:Schizothorax dolichonema 594:Schizothorax cryptolepis 529:Schizothorax beipanensis 1179:Schizothorax oligolepis 1101:Schizothorax nepalensis 1077:Schizothorax myzostomus 1072:(Blunt-nosed snowtrout) 1000:Schizothorax macropogon 715:Schizothorax eurystomus 611:Schizothorax curvifrons 515:Schizothorax argentatus 479:Schizothorax curvifrons 1388:& A. A. Awan, 1978 1308:Schizothorax raraensis 1258:Schizothorax prophylax 1243:Schizothorax progastus 1211:(Transcaspian marinka) 688:Schizothorax elongatus 550:Schizothorax biddulphi 506: 494: 482: 298:this is merely due to 1871:Cyprinid fish of Asia 1434:Schizothorax zarudnyi 1368:Schizothorax sinensis 1229:Schizothorax prenanti 1166:Schizothorax oconnori 893:Schizothorax labrosus 878:Schizothorax labiatus 828:Schizothorax huegelii 701:Schizothorax esocinus 675:Schizothorax edeniana 629:(Wu & Tsao, 1992) 500: 488: 476: 1394:Schizothorax waltoni 1355:Schizothorax saltans 1286:Schizothorax ramzani 1253:(Dinnawah snowtrout) 1202:Schizothorax pelzami 995:(Nepalese snowtrout) 851:Schizothorax kozlovi 752:Schizothorax griseus 738:Schizothorax grahami 503:Schizothorax waltoni 300:convergent evolution 1611:Schizothorax chivae 1483:. May 2019 version. 1192:Schizothorax parvus 937:Schizothorax leukus 765:Schizothorax heteri 710:(Chirruh snowtrout) 635:Schizothorax davidi 580:Schizothorax chongi 563:Schizothorax chivae 315:Protacanthopterygii 1471:"Species in genus 1332:(Common snowtrout) 1087:Schizothorax nasus 873:(Kumaon snowtrout) 620:(Sattar snowtrout) 524:(Balkhash marinka) 507: 495: 483: 244:) 'to cleave' and 1866:Cyprinidae genera 1848: 1847: 1820:Open Tree of Life 1669:Taxon identifiers 1548:Helmand - Sistan. 1443: 1429: 1416: 1402: 1389: 1376: 1363: 1349: 1331: 1316: 1303: 1280: 1267:(Eğirdir marinka) 1266: 1252: 1238: 1224: 1210: 1197: 1187: 1174: 1161: 1151: 1130: 1109: 1096:(Dongu snowtrout) 1095: 1082: 1071: 1057: 1044: 1031: 1021: 1008: 994: 981: 967: 957: 932: 919: 909: 888:(Kunar snowtrout) 887: 872: 859: 846: 845:(Wu et al., 1992) 836: 823: 802: 785: 760: 747: 733: 723: 709: 696: 683: 670: 657: 644: 630: 619: 606: 589: 575: 558: 545: 523: 453:, which is quite 323:million years ago 221:and lakes of the 186: 185: 181: 145: 1878: 1841: 1840: 1828: 1827: 1815: 1814: 1802: 1801: 1789: 1788: 1776: 1775: 1763: 1762: 1750: 1749: 1737: 1736: 1724: 1723: 1711: 1710: 1709: 1696: 1695: 1694: 1664: 1663: 1657: 1645: 1639: 1626: 1617: 1607: 1601: 1600: 1574: 1568: 1557: 1551: 1544: 1538: 1528: 1522: 1511: 1505: 1497:Research 50(3). 1494: 1485: 1484: 1463: 1437: 1424: 1410: 1397: 1384: 1371: 1358: 1340: 1325: 1317:(Rara snowtrout) 1311: 1289: 1275: 1261: 1246: 1232: 1219: 1205: 1196:W. H. Tsao, 1964 1195: 1182: 1169: 1160:W. H. Tsao, 1964 1159: 1138: 1117: 1104: 1090: 1081:W. H. Tsao, 1964 1080: 1065: 1052: 1039: 1030:W. H. Tsao, 1964 1029: 1016: 1003: 989: 975: 966:W. H. Tsao, 1964 965: 940: 927: 918:W. H. Tsao, 1964 917: 896: 881: 867: 854: 844: 831: 810: 793: 768: 755: 741: 732:W. H. Tsao, 1964 731: 718: 704: 691: 678: 665: 652: 638: 628: 614: 597: 583: 566: 553: 532: 518: 465:in the process. 321:, more than 200 285: 284: 279: 255: 254: 249: 243: 242: 237: 179: 140: 56: 55: 41:Schizothorax sp. 35: 21: 20: 1886: 1885: 1881: 1880: 1879: 1877: 1876: 1875: 1851: 1850: 1849: 1844: 1836: 1831: 1823: 1818: 1810: 1805: 1797: 1792: 1784: 1779: 1771: 1766: 1758: 1753: 1745: 1740: 1732: 1727: 1719: 1714: 1705: 1704: 1699: 1690: 1689: 1684: 1671: 1661: 1660: 1646: 1642: 1627: 1620: 1608: 1604: 1575: 1571: 1558: 1554: 1545: 1541: 1529: 1525: 1512: 1508: 1495: 1488: 1464: 1455: 1450: 1440:A. M. Nikolskii 1068:B. L. Chaudhuri 856:A. M. Nikolskii 471: 260:). The western 227:scientific name 163: 160: 139: 50: 17: 16:Genus of fishes 12: 11: 5: 1884: 1874: 1873: 1868: 1863: 1846: 1845: 1843: 1842: 1829: 1816: 1803: 1790: 1777: 1764: 1751: 1738: 1725: 1712: 1697: 1681: 1679: 1673: 1672: 1659: 1658: 1650:Racoma ramzani 1640: 1618: 1602: 1584:Animal Biology 1569: 1552: 1539: 1523: 1506: 1486: 1467:Froese, Rainer 1452: 1451: 1449: 1446: 1445: 1444: 1430: 1417: 1403: 1390: 1377: 1364: 1350: 1333: 1318: 1304: 1282: 1268: 1254: 1239: 1225: 1212: 1198: 1188: 1175: 1162: 1152: 1131: 1110: 1097: 1083: 1073: 1058: 1045: 1032: 1022: 1009: 996: 982: 968: 958: 933: 920: 910: 889: 874: 860: 847: 837: 824: 803: 786: 761: 748: 734: 724: 711: 697: 684: 671: 658: 645: 631: 621: 607: 590: 576: 572:Alphonsa Moyon 559: 546: 525: 470: 467: 463:junior synonym 388:Schizopygopsis 364:Oxygymnocypris 352:Gymnodiptychus 184: 183: 172: 171: 165: 164: 161: 154: 153: 147: 146: 132: 128: 127: 122: 118: 117: 112: 108: 107: 102: 98: 97: 95:Actinopterygii 92: 88: 87: 82: 78: 77: 72: 68: 67: 62: 58: 57: 44: 43: 37: 36: 28: 27: 15: 9: 6: 4: 3: 2: 1883: 1872: 1869: 1867: 1864: 1862: 1859: 1858: 1856: 1839: 1834: 1830: 1826: 1821: 1817: 1813: 1808: 1804: 1800: 1795: 1791: 1787: 1782: 1778: 1774: 1769: 1765: 1761: 1756: 1752: 1748: 1743: 1739: 1735: 1730: 1726: 1722: 1717: 1713: 1708: 1702: 1698: 1693: 1687: 1683: 1682: 1680: 1678: 1674: 1670: 1665: 1656: 1653: 1651: 1644: 1638: 1635: 1633: 1625: 1623: 1616: 1612: 1606: 1598: 1594: 1590: 1586: 1585: 1580: 1573: 1567: 1563: 1556: 1549: 1543: 1537: 1536:90-6193-535-0 1533: 1527: 1521: 1517: 1510: 1504: 1500: 1493: 1491: 1482: 1481: 1476: 1474: 1468: 1462: 1460: 1458: 1453: 1441: 1436: 1435: 1431: 1427: 1423: 1422: 1418: 1414: 1409: 1408: 1404: 1400: 1396: 1395: 1391: 1387: 1383: 1382: 1378: 1374: 1370: 1369: 1365: 1361: 1357: 1356: 1351: 1347: 1343: 1339: 1338: 1334: 1329: 1324: 1323: 1319: 1314: 1310: 1309: 1305: 1301: 1297: 1293: 1288: 1287: 1283: 1281:(Ili marinka) 1278: 1274: 1273: 1269: 1264: 1260: 1259: 1255: 1250: 1245: 1244: 1240: 1236: 1231: 1230: 1226: 1222: 1218: 1217: 1213: 1208: 1204: 1203: 1199: 1194: 1193: 1189: 1185: 1181: 1180: 1176: 1172: 1168: 1167: 1163: 1158: 1157: 1153: 1149: 1145: 1141: 1137: 1136: 1132: 1128: 1124: 1120: 1116: 1115: 1111: 1107: 1103: 1102: 1098: 1093: 1089: 1088: 1084: 1079: 1078: 1074: 1069: 1064: 1063: 1059: 1055: 1051: 1050: 1046: 1042: 1038: 1037: 1033: 1028: 1027: 1023: 1019: 1015: 1014: 1010: 1006: 1002: 1001: 997: 992: 988: 987: 983: 979: 974: 973: 969: 964: 963: 959: 955: 951: 947: 943: 939: 938: 934: 930: 926: 925: 921: 916: 915: 911: 907: 903: 899: 895: 894: 890: 885: 880: 879: 875: 870: 866: 865: 861: 857: 853: 852: 848: 843: 842: 838: 834: 830: 829: 825: 821: 817: 813: 809: 808: 804: 800: 796: 792: 791: 787: 783: 779: 775: 771: 767: 766: 762: 758: 754: 753: 749: 745: 740: 739: 735: 730: 729: 725: 721: 717: 716: 712: 707: 703: 702: 698: 694: 690: 689: 685: 681: 677: 676: 672: 668: 664: 663: 659: 655: 651: 650: 646: 642: 637: 636: 632: 627: 626: 622: 617: 613: 612: 608: 604: 600: 596: 595: 591: 587: 582: 581: 577: 573: 569: 565: 564: 560: 556: 552: 551: 547: 543: 539: 535: 531: 530: 526: 521: 517: 516: 512: 511: 510: 505: 504: 499: 493: 492: 487: 481: 480: 475: 466: 464: 460: 456: 452: 449: 445: 444: 439: 438: 433: 432: 427: 426: 421: 420: 419:Messinobarbus 415: 414: 409: 408: 407:sensu stricto 404: 403: 398: 394: 390: 389: 384: 383: 378: 377: 372: 371: 370:Platypharodon 366: 365: 360: 359: 354: 353: 348: 347: 342: 341: 336: 335: 334:Aspiorhynchus 330: 326: 324: 320: 316: 312: 309: 305: 301: 297: 293: 289: 278: 273: 269: 268: 263: 259: 248: 236: 232: 231:Ancient Greek 228: 224: 220: 219:Lake Balkhash 216: 212: 208: 204: 200: 196: 192: 191: 182: 178: 173: 170: 166: 159: 155: 152: 148: 143: 138: 137: 133: 130: 129: 126: 123: 120: 119: 116: 113: 110: 109: 106: 105:Cypriniformes 103: 100: 99: 96: 93: 90: 89: 86: 83: 80: 79: 76: 73: 70: 69: 66: 63: 60: 59: 54: 49: 45: 42: 38: 34: 29: 26: 22: 19: 1861:Schizothorax 1721:Schizothorax 1707:Schizothorax 1677:Schizothorax 1676: 1654: 1649: 1643: 1636: 1632:Schizothorax 1631: 1614: 1610: 1605: 1591:(1): 39–48. 1588: 1582: 1578: 1572: 1555: 1542: 1526: 1509: 1478: 1473:Schizothorax 1472: 1432: 1419: 1405: 1392: 1379: 1366: 1353: 1335: 1320: 1306: 1284: 1270: 1256: 1241: 1235:T. L. Tchang 1227: 1214: 1200: 1190: 1177: 1164: 1154: 1133: 1127:D. D. Zhuang 1112: 1099: 1085: 1075: 1060: 1047: 1034: 1024: 1011: 998: 984: 970: 960: 935: 922: 912: 902:D. D. Zhuang 891: 876: 862: 849: 839: 826: 805: 788: 763: 750: 736: 726: 713: 699: 686: 673: 660: 647: 633: 623: 609: 592: 578: 561: 548: 527: 513: 508: 501: 489: 477: 455:paraphyletic 441: 435: 429: 425:Carasobarbus 423: 417: 411: 405: 400: 393:Schizothorax 392: 386: 380: 376:Ptychobarbus 374: 368: 362: 356: 350: 344: 338: 332: 329:Schizothorax 328: 327: 311:Ostariophysi 287: 266: 265: 223:Sistan Basin 215:S. prophylax 214: 211:Central Asia 190:Schizothorax 189: 188: 187: 176: 175: 162:Heckel, 1838 157: 151:Type species 136:Schizothorax 135: 134: 40: 25:Schizothorax 24: 18: 1768:iNaturalist 1701:Wikispecies 1373:Herzenstein 1277:Herzenstein 1263:Pietschmann 1184:S. Y. Huang 1123:K. X. Zhang 1054:S. Y. Huang 1018:S. Y. Huang 946:L. P. Zheng 774:L. P. Zheng 693:S. Y. Huang 667:S. Y. Huang 654:Herzenstein 431:Labeobarbus 413:Luciobarbus 358:Gymnocypris 270:from their 121:Subfamily: 1855:Categories 1448:References 1413:P. W. Fang 1328:J. E. Gray 1249:McClelland 1148:J. X. Yang 1144:X. Y. Chen 1119:Y. H. Wang 978:P. W. Fang 954:J. X. Yang 950:X. Y. Chen 929:J. X. Yang 898:Y. H. Wang 884:McClelland 820:J. X. Yang 816:X. Y. Chen 782:J. X. Yang 778:X. Y. Chen 680:McClelland 586:P. W. Fang 542:J. X. Yang 538:X. Y. Chen 459:Cyprininae 382:Schizopyge 308:superorder 203:South Asia 115:Cyprinidae 1313:Terashima 1106:Terashima 991:Terashima 906:L. C. Gao 757:Pellegrin 568:Arunkumar 448:subfamily 443:Cyprinion 346:Diptychus 340:Chuanchia 304:Teleostei 288:snowtrout 71:Kingdom: 65:Eukaryota 1686:Wikidata 1480:FishBase 1346:C. Z. Wu 1342:Y. F. Wu 799:T. Y. Fu 795:M. R. Ye 603:M. R. Ye 599:T. Y. Fu 451:Barbinae 437:Aulopyge 319:Triassic 267:marinkas 235:schízeïn 225:. Their 207:Himalaya 199:cyprinid 169:Synonyms 125:Barbinae 111:Family: 85:Chordata 81:Phylum: 75:Animalia 61:Domain: 1786:1369042 1760:2363554 1692:Q310753 1442:, 1897) 1415:, 1936) 1330:, 1832) 1302:, 2012) 1300:Pervaiz 1251:, 1839) 1237:, 1930) 1207:Kessler 1140:J. Yang 1070:, 1913) 980:, 1936) 942:J. Yang 886:, 1842) 812:J. Yang 770:J. Yang 746:, 1904) 720:Kessler 643:, 1880) 641:Sauvage 588:, 1936) 555:Günther 534:J. Yang 520:Kessler 469:Species 397:barbels 296:habitus 283:маринка 277:marinka 272:Russian 262:species 241:σχίζειν 131:Genus: 101:Order: 91:Class: 1838:270830 1825:211540 1799:638910 1579:Barbus 1534:  1428:, 1923 1426:Norman 1401:, 1905 1375:, 1889 1362:, 1905 1348:, 1992 1344:& 1315:, 1984 1298:& 1279:, 1889 1265:, 1933 1223:, 1838 1221:Heckel 1209:, 1870 1186:, 1985 1173:, 1908 1150:, 2009 1146:& 1129:, 1981 1125:& 1108:, 1984 1094:, 1838 1092:Heckel 1056:, 1982 1043:, 1877 1041:F. Day 1020:, 1985 1007:, 1905 993:, 1984 956:, 2013 952:& 931:, 1991 908:, 1981 904:& 871:, 1971 858:, 1903 835:, 1838 833:Heckel 822:, 2009 818:& 801:, 1986 797:& 784:, 2013 780:& 759:, 1931 722:, 1872 708:, 1838 706:Heckel 695:, 1985 682:, 1842 669:, 1985 656:, 1889 618:, 1838 616:Heckel 605:, 1984 601:& 574:, 2016 570:& 557:, 1876 544:, 2009 540:& 522:, 1874 402:Barbus 292:trouts 258:thorax 247:thórax 209:) and 177:Racoma 144:, 1838 142:Heckel 1833:WoRMS 1812:75361 1781:IRMNG 1773:91921 1734:71166 1399:Regan 1386:Mirza 1360:Regan 1296:Ullah 1292:Javed 1171:Lloyd 1005:Regan 869:Menon 744:Regan 274:name 253:θώραξ 195:genus 193:is a 1807:NCBI 1794:ITIS 1755:GBIF 1747:7DQR 1729:BOLD 1532:ISBN 440:and 416:and 385:and 1742:CoL 1716:ADW 1593:doi 1562:doi 1516:doi 1499:doi 422:), 294:in 197:of 1857:: 1835:: 1822:: 1809:: 1796:: 1783:: 1770:: 1757:: 1744:: 1731:: 1718:: 1703:: 1688:: 1621:^ 1589:57 1587:. 1489:^ 1477:. 1456:^ 1294:, 1142:, 1121:, 948:, 944:, 900:, 814:, 776:, 772:, 536:, 434:, 428:, 379:, 373:, 367:, 361:, 355:, 349:, 343:, 337:, 325:. 1599:. 1595:: 1564:: 1518:: 1501:: 1475:" 1438:( 1411:( 1352:† 1326:( 1290:( 1247:( 1233:( 1066:( 976:( 882:( 742:( 639:( 584:( 399:( 280:( 250:( 238:( 205:(

Index


Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Actinopterygii
Cypriniformes
Cyprinidae
Barbinae
Schizothorax
Heckel
Type species
Synonyms
genus
cyprinid
South Asia
Himalaya
Central Asia
Lake Balkhash
Sistan Basin
scientific name
Ancient Greek
thorax
species
Russian
trouts
habitus
convergent evolution
Teleostei

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