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Kingdom (biology)

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31: 5109: 5681: 6002: 464: 1601:, of Bacteria, Archaea, and Eukaryota. Combined with the five-kingdom model, this created a six-kingdom model, where the kingdom Monera is replaced by the kingdoms Bacteria and Archaea. This six-kingdom model is commonly used in recent US high school biology textbooks, but has received criticism for compromising the current scientific consensus. But the division of prokaryotes into two kingdoms remains in use with the recent 511:, for "neutral organisms" or "the kingdom of primitive forms", which were neither animal nor plant; he did not include the Regnum Lapideum in his scheme. Haeckel revised the content of this kingdom a number of times before settling on a division based on whether organisms were unicellular (Protista) or multicellular (animals and plants). 5270:
According to genetic data, although eukaryote groups such as plants, fungi, and animals may look different, they are more closely related to each other than they are to either the Eubacteria or Archaea. It was also found that the eukaryotes are more closely related to the Archaea than they are to the
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Following publication of Whittaker's system, the five-kingdom model began to be commonly used in high school biology textbooks. But despite the development from two kingdoms to five among most scientists, some authors as late as 1975 continued to employ a traditional two-kingdom system of animals and
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In 1990, the name "domain" was proposed for the highest rank. This term represents a synonym for the category of dominion (lat. dominium), introduced by Moore in 1974. Unlike Moore, Woese et al. (1990) did not suggest a Latin term for this category, which represents a further argument supporting the
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However, in the same year as the International Society of Protistologists' classification was published (2005), doubts were being expressed as to whether some of these supergroups were monophyletic, particularly the Chromalveolata, and a review in 2006 noted the lack of evidence for several of the
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On this basis, the diagram opposite (redrawn from their article) showed the real "kingdoms" (their quotation marks) of the eukaryotes. A classification which followed this approach was produced in 2005 for the International Society of Protistologists, by a committee which "worked in collaboration
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In 1998, Cavalier-Smith published a six-kingdom model, which has been revised in subsequent papers. The version published in 2009 is shown below. Cavalier-Smith no longer accepted the importance of the fundamental Eubacteria–Archaebacteria divide put forward by Woese and others and supported by
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in 2009 noted that "The deep phylogeny of eukaryotes is an extremely difficult and controversial problem." As of December 2010, there appears to be a consensus that the six supergroup model proposed in 2005 does not reflect the true phylogeny of the eukaryotes and hence how they should be
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The most recent research does not support the classification of the eukaryotes into any of the standard systems. As of April 2010, no set of kingdoms is sufficiently supported by research to attain widespread acceptance. In 2009, Andrew Roger and Alastair Simpson emphasized the need for
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At first, microscopic organisms were classified within the animal and plant kingdoms. However, by the mid–19th century, it had become clear to many that "the existing dichotomy of the plant and animal kingdoms rapidly blurred at its boundaries and outmoded".
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of London a copy of his first observations of microscopic single-celled organisms. Until then, the existence of such microscopic organisms was entirely unknown. Despite this, Linnaeus did not include any microscopic creatures in his original taxonomy.
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The remaining two kingdoms, Protista and Monera, included unicellular and simple cellular colonies. The five kingdom system may be combined with the two empire system. In the Whittaker system, Plantae included some algae. In other systems, such as
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that are not animals, plants or fungi". They held that only monophyletic groups should be accepted as formal ranks in a classification and that – while this approach had been impractical previously (necessitating "literally dozens of eukaryotic
861:. The resulting five-kingdom system, proposed in 1969 by Whittaker, has become a popular standard and with some refinement is still used in many works and forms the basis for new multi-kingdom systems. It is based mainly upon differences in 6129: 852:
and other organisms regarded as plants had long been recognised by some; Haeckel had moved the fungi out of Plantae into Protista after his original classification, but was largely ignored in this separation by scientists of his time.
240:) are the two ranks immediately below kingdom. Superkingdom may be considered as an equivalent of domain or empire or as an independent rank between kingdom and domain or subdomain. In some classification systems the additional rank 5271:
Eubacteria. Although the primacy of the Eubacteria-Archaea divide has been questioned, it has been upheld by subsequent research. There is no consensus on how many kingdoms exist in the classification scheme proposed by Woese.
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Raoult, Didier; Audic, StĂ©phane; Robert, Catherine; Abergel, Chantal; Renesto, Patricia; Ogata, Hiroyuki; La Scola, Bernard; Suzan, Marie; Claverie, Jean-Michel (2004). "The 1.2 megabase genome sequence of Mimivirus".
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In the same way, his paraphyletic kingdom Protozoa includes the ancestors of Animalia, Fungi, Plantae, and Chromista. The advances of phylogenetic studies allowed Cavalier-Smith to realize that all the phyla thought to be
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The IUCN Species Survival Commission calls for the due recognition of fungi as major components of biodiversity in legislation and policy. It fully endorses the Fauna Flora Funga Initiative and asks that the phrases
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outside of a host cell. Another argument is that their placement in the tree would be problematic, since it is suspected that viruses have various evolutionary origins, and they have a penchant for harvesting
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with specialists from many societies". It divided the eukaryotes into the same six "supergroups". The published classification deliberately did not use formal taxonomic ranks, including that of "kingdom".
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Cavalier-Smith and his collaborators revised their classification in 2015. In this scheme they introduced two superkingdoms of Prokaryota and Eukaryota and seven kingdoms. Prokaryota have two kingdoms:
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Ruggiero, Michael A.; Gordon, Dennis P.; Orrell, Thomas M.; Bailly, Nicolas; Bourgoin, Thierry; Brusca, Richard C.; Cavalier-Smith, Thomas; Guiry, Michael D.; Kirk, Paul M.; Thuesen, Erik V. (2015).
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Ruggiero, Michael A.; Gordon, Dennis P.; Orrell, Thomas M.; Bailly, Nicolas; Bourgoin, Thierry; Brusca, Richard C.; Cavalier-Smith, Thomas; Guiry, Michael D.; Kirk, Paul M.; Thuesen, Erik V. (2015).
2215:; the bimembranous-unimembranous transition was thought to be far more fundamental than the long branch of genetic distance of Archaebacteria, viewed as having no particular biological significance. 8079: 696:
promoted and popularized Édouard Chatton's earlier work, particularly in their paper of 1962, "The Concept of a Bacterium"; this created, for the first time, a rank above kingdom—a
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Whittaker, R.H. (January 1969). "New concepts of kingdoms or organisms. Evolutionary relations are better represented by new classifications than by the traditional two kingdoms".
681:, had placed the blue-green algae (or Phycochromacea) in Monera; this would gradually gain acceptance, and the blue-green algae would become classified as bacteria in the phylum 8184:
Burki, Fabien; Shalchian-Tabrizi, Kamran; Minge, Marianne; Skjæveland, Åsmund; Nikolaev, Sergey I.; Jakobsen, Kjetill S. & Pawlowski, Jan (2007). Butler, Geraldine (ed.).
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and colleagues proposed the fundamental subdivision of the prokaryotes into the Eubacteria (later called the Bacteria) and Archaebacteria (later called the Archaea), based on
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While the concept of kingdoms continues to be used by some taxonomists, there has been a movement away from traditional kingdoms, as they are no longer seen as providing a
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plants, dividing the plant kingdom into subkingdoms Prokaryota (bacteria and cyanobacteria), Mycota (fungi and supposed relatives), and Chlorota (algae and land plants).
6130:"Re:wild and IUCN SSC become first global organizations to call for the recognition of fungi as one of three kingdoms of life critical to protecting and restoring Earth" 1826:
was so great (particularly considering the genetic distance of ribosomal genes) that the prokaryotes needed to be separated into two different kingdoms. He then divided
8395:"Analysis of rare genomic changes does not support the unikont–bikont phylogeny, and suggests cyanobacterial symbiosis as the point of primary radiation of eukaryotes" 501:, a third kingdom of life composed of "all the lower creatures, or the primary organic beings"; he retained Regnum Lapideum as a fourth kingdom of minerals. In 1866, 2226:(therefore "monophyletic" in the sense he uses this term) taxa, marking important innovations of biological significance (in regard of the concept of biological 5723:
to their classifications. This can create a "superdomain" called "Acytota", also called "Aphanobionta", of non-cellular life; with the other superdomain being "
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diligence in analyzing new discoveries: "With the current pace of change in our understanding of the eukaryote tree of life, we should proceed with caution."
3309: 8301:"Large-scale phylogenomic analyses reveal that two enigmatic protist lineages, Telonemia and Centroheliozoa, are related to photosynthetic Chromalveolates" 5986:
On the other hand, there are arguments in favor of their inclusion. One of these comes from the discovery of unusually large and complex viruses, such as
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The kingdom-level classification of life is still widely employed as a useful way of grouping organisms, notwithstanding some problems with this approach:
3352: 1866:. Since then, many non-photosynthetic phyla of protists, thought to have secondarily lost their chloroplasts, were integrated into the kingdom Chromista. 4069: 3337: 3289: 3229: 2680: 2620: 1881:. As a result, these amitochondriate protists were separated from the protist kingdom, giving rise to the, at the same time, superkingdom and kingdom 4193: 4064: 3858: 3697: 3304: 3269: 3175: 3150: 3008: 8998: 4168: 4110: 3765: 3397: 3392: 3377: 3342: 3294: 3259: 3140: 3105: 3095: 3090: 3080: 2715: 2635: 2610: 2595: 2585: 2238:(i.e. primitively amitochondriate eukaryotes) had in fact secondarily lost their mitochondria, typically by transforming them into new organelles: 467:
Haeckel's original (1866) conception of the three kingdoms of life, including the new kingdom Protista. Notice the inclusion of the cyanobacterium
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International Code of Botanical Nomenclature (Vienna Code) adopted by the Seventeenth International Botanical Congress, Vienna, Austria, July 2005
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or Eubacteria), while textbooks in other parts of the world, such as the United Kingdom, Pakistan, Bangladesh, India, Greece, Brazil, Spain use
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Parfrey, Laura W.; Barbero, Erika; Lasser, Elyse; Dunthorn, Micah; Bhattacharya, Debashish; Patterson, David J. & Katz, Laura A. (2006).
4279: 4235: 4173: 4115: 4105: 3576: 3367: 3357: 3244: 3130: 3115: 3074: 3013: 2796: 2791: 2700: 2600: 2200: 8299:
Burki, F.; Inagaki, Y.; Brate, J.; Archibald, J. M.; Keeling, P. J.; Cavalier-Smith, T.; Sakaguchi, M.; Hashimoto, T.; et al. (2009).
7006: 5119: 4178: 4120: 3838: 3641: 3636: 3239: 3193: 3160: 2983: 2945: 2940: 2925: 2811: 2755: 5162:
thought life could be divided into three large divisions and referred to them as the "three primary kingdom" model or "urkingdom" model.
4153: 3942: 3823: 3770: 3687: 3586: 3561: 3541: 3215: 3041: 2832: 6723: 6397: 3808: 3621: 3520: 3208: 3066: 2222:("holophyletic" in his terminology) to be valid. He defines Prokaryota, Bacteria, Negibacteria, Unibacteria, and Posibacteria as valid 6354:
Scamardella, Joseph M. (1999). "Not plants or animals: a brief history of the origin of Kingdoms Protozoa, Protista and Protoctista".
9025: 5125: 4158: 3957: 2963: 2842: 8989: 8352:"Phylogenomic analysis supports the monophyly of cryptophytes and haptophytes and the association of Rhizaria with chromalveolates" 6078: 5117: 2902: 5122: 6670:
Balch, W.E.; Magrum, L.J.; Fox, G.E.; Wolfe, C.R. & Woese, C.R. (August 1977). "An ancient divergence among the bacteria".
5127: 6651: 5177:, stressing that there was as much genetic difference between these two groups as between either of them and all eukaryotes. 8980: 5300:") – it had now become possible to divide the eukaryotes into "just a few major groups that are probably all monophyletic". 177:
into biology in 1735, the highest rank was given the name "kingdom" and was followed by four other main or principal ranks:
5700:, so that Rhizaria is not one of the main eukaryote groups. Beyond this, there does not appear to be a consensus. Rogozin 5128: 8028: 5714: 5138: 5120: 2277:, the diagram below is an 'organization chart', not an 'ancestor chart', and does not represent an evolutionary tree. 8565: 7544: 6562: 5126: 299: 7732: 5118: 7621: 6271: 5137: 5692:
As of 2010, there is widespread agreement that the Rhizaria belong with the Stramenopiles and the Alveolata, in a
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can be included in the tree of life. The arguments against include the fact that they are obligate intracellular
5123: 4354:). The Eukaryota have five kingdoms: Protozoa, Chromista, Plantae, Fungi, and Animalia. In this classification a 5139: 5136: 5129: 5121: 2246:, according to the significance of the term given by Cavalier-Smith. Some of the members of the defunct kingdom 1869:
Finally, some protists lacking mitochondria were discovered. As mitochondria were known to be the result of the
7307:; Balch, W.E.; Magrum, L.J.; Fox, G.E.; Wolfe, R.S. (August 1977). "An ancient divergence among the bacteria". 6672: 6164: 5131: 669:
of prokaryotic organisms; as a revised phylum Monera of the Protista, it included organisms now classified as
9018: 8726: 5116: 36: 5130: 9419: 8399: 8305: 7258: 7230: 7192: 7117:
Systemae Naturae, sive regna tria naturae, systematics proposita per classes, ordines, genera & species
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Systemae Naturae, sive regna tria naturae, systematics proposita per classes, ordines, genera & species
4664: 4657: 3179:                Superorder 3174:                Superorder 854: 662: 40: 8958: 8780:
Moreira, David; PurificaciĂłn LĂłpez-GarcĂ­a (2009). "Ten reasons to exclude viruses from the tree of life".
7947: 9414: 8859: 8782: 1877:, it was thought that these amitochondriate eukaryotes were primitively so, marking an important step in 35: 6444: 5114: 39: 38: 6917: 6356: 17: 5135: 5115: 7234: 6516: 5169:
Woese divided the prokaryotes (previously classified as the Kingdom Monera) into two groups, called
5112: 42: 9011: 6593: 338: 8350:
Hackett, J.D.; Yoon, H.S.; Li, S.; Reyes-Prieto, A.; Rummele, S.E. & Bhattacharya, D. (2007).
5132: 5113: 8826: 7989: 1913:, which has since been abandoned; later schemes did not include the Archezoa–Metakaryota divide. 478: 310: 37: 7425:"Towards a natural system of organisms: proposal for the domains Archaea, Bacteria, and Eucarya" 3789:              Infraphylum 3784:              Infraphylum 3774:              Infraphylum 3769:              Infraphylum 3734:              Infraphylum 3729:              Infraphylum 3724:              Infraphylum 3706:              Infraphylum 3701:              Infraphylum 3640:              Infraphylum 3635:              Infraphylum 3590:              Infraphylum 3585:              Infraphylum 3580:              Infraphylum 9265: 8666:"Eocytes: A new ribosome structure indicates a kingdom with a close relationship to eukaryotes" 7671: 6998: 6511: 5134: 1843: 1835: 1605:
scheme of Thomas Cavalier-Smith, although it primarily differs in that Protista is replaced by
458: 5133: 41: 8021:"The new higher-level classification of eukaryotes with emphasis on the taxonomy of protists" 7728: 7667: 7617: 7520: 7482: 6945: 4690: 3169:              Subclass 3164:              Subclass 3154:              Subclass 3149:              Subclass 1855: 1815: 494: 6735: 8908: 8679: 8618: 8463: 8199: 7793: 7436: 7374: 7316: 7270: 6833: 6776: 6731: 6681: 6503: 6216: 6169:(electronic ed.). Vienna: International Association for Plant Taxonomy. Archived from 5980: 5282: 4359: 1129: 1123: 642:
revealed important distinctions between those organisms whose cells do not have a distinct
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Adl SM, Simpson AGB, Farmer MA, Andersen RA, Anderson OR, Barta JR, et al. (2005).
7797: 7440: 7378: 7320: 7274: 6837: 6780: 6685: 6507: 6220: 6184: 6085: 34: 9194: 9138: 9123: 9105: 8932: 8807: 8641: 8604: 8583: 8538: 8486: 8449: 8421: 8394: 8327: 8300: 8276: 8249: 8222: 8185: 8146: 8117: 8055: 7970: 7928: 7872: 7816: 7781: 7757: 7699: 7649: 7637: 7340: 7212: 7089: 7062: 7028: 6974: 6949: 6909: 6857: 6799: 6762: 6705: 6610: 6469: 6419: 6392: 6302: 6290: 6277: 6030: 5058: 4347: 3656: 3387: 2179: 1542: 1280: 314: 275: 46: 8702: 8665: 8020: 7594: 7569: 7397: 7362: 7174: 7148: 4650: 655: 9255: 9209: 9204: 8924: 8899: 8878: 8799: 8707: 8646: 8571: 8561: 8530: 8491: 8445: 8426: 8375: 8332: 8281: 8227: 8151: 8098: 8047: 8042: 7932: 7920: 7864: 7821: 7762: 7704: 7641: 7599: 7550: 7540: 7502: 7498: 7464: 7459: 7424: 7402: 7332: 7286: 7152: 7094: 7033: 6979: 6901: 6849: 6804: 6697: 6647: 6636: 6558: 6529: 6494: 6461: 6424: 6365: 6325: 6294: 6244: 6239: 6204: 5801: 5777: 5755: 5728: 5720: 5288: 5143: 4752: 4747: 4734: 4729: 4724: 4719: 4351: 3372: 3254: 705: 136: 8936: 8542: 7876: 7344: 7216: 6709: 6473: 5951:
uses the taxonomic rank "kingdom" in the classification of viruses (with the suffix
4283:            Infraphylum 4278:            Infraphylum 3999:            Infraphylum 3994:            Infraphylum 3981:            Infraphylum 3976:            Infraphylum 3956:            Infraphylum 3951:            Infraphylum 3928:            Infraphylum 3923:            Infraphylum 3913:            Infraphylum 3908:            Infraphylum 3857:            Infraphylum 3852:            Infraphylum 3842:            Infraphylum 3837:            Infraphylum 3817:            Infraphylum 3812:            Infraphylum 2679:            Infraphylum 2674:            Infraphylum 2559:            Superphylum 708:
of prokaryotes and eukaryotes. The two-empire system would later be expanded to the
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introduced the terms "prokaryote" and "eukaryote" to differentiate these organisms.
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Hegde, Nagendra; Maddur, Mohan S.; Kaveri, Srini V. & Bayry, Jagadeesh (2009).
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classified, although there is no agreement as to the model which should replace it.
5670: 5514: 5103: 3085: 2906:            Superclass 2901:            Superclass 2227: 709: 218: 206: 74: 54: 8526: 8250:"Phylogenomics reveals a new 'megagroup' including most photosynthetic eukaryotes" 6205:"Towards a natural systs: proposal for the domains Archaea, Bacteria, and Eucarya" 3779:            Subphylum 3764:            Subphylum 3749:            Subphylum 3744:            Subphylum 3719:            Subphylum 3711:            Subphylum 3696:            Subphylum 3655:            Subphylum 3650:            Subphylum 3645:            Subphylum 3630:            Subphylum 3605:            Subphylum 3600:            Subphylum 3575:            Subphylum 3570:            Subphylum 3555:            Subphylum 3550:            Subphylum 3540:            Subphylum 3535:            Subphylum 9355: 9341: 9308: 9297: 9291: 9245: 9240: 9235: 9229: 9223: 9218: 9184: 9179: 9174: 9169: 9163: 9157: 8993: 8502: 8476: 8254: 8212: 8136: 8075:"On the monophyly of chromalveolates using a six-protein phylogeny of eukaryotes" 7901: 7845: 7806: 7282: 6789: 6548: 6525: 3977: 3828: 3760: 3755: 3712: 3396:            Subclass 3391:            Subclass 3381:            Subclass 3376:            Subclass 3371:            Subclass 3234: 2695: 2690: 2615: 1874: 1854:
kingdom. Indeed, the chloroplast of the chromists is located in the lumen of the
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Traditionally, some textbooks from the United States and Canada used a system of
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Lake, James A.; Henderson, Eric; Oakes, Melanie; Clark, Michael W. (June 1984).
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A short history of biology, a general introduction to the study of living things
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Proceedings of the National Academy of Sciences of the United States of America
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Proceedings of the National Academy of Sciences of the United States of America
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Proceedings of the National Academy of Sciences of the United States of America
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In 2004, a review article by Simpson and Roger noted that the Protista were "a
5225: 4863: 4853: 3818: 2988: 2973: 2862: 2826: 2750: 2735: 2725: 2665: 2575: 2318: 2208: 2199:) was subdivided into two sub-kingdoms according to their membrane topologies: 1966: 1878: 1846:). Technological advances in electron microscopy allowed the separation of the 1823: 1667: 1446: 1316: 693: 269:
The classification of living things into animals and plants is an ancient one.
70: 50: 7915: 7896: 7859: 7840: 7536: 3159:            Class 3144:            Class 3134:            Class 3129:            Class 9408: 9384: 8450:"EEF2 analysis challenges the monophyly of Archaeplastida and Chromalveolata" 8122: 7130: 7112: 6020: 5842: 4643: 4636: 4095: 4076: 4000: 3894: 3889: 3884: 3865: 3843: 3803: 3445: 3440: 3435: 3347: 3321: 3284: 3187: 2930: 2914: 2675: 2655: 2650: 2625: 2565: 2251: 2239: 1870: 1863: 1594: 1554: 1510: 1452: 1376: 1334: 1274: 1205: 1141: 882: 689: 682: 502: 482: 306: 253: 193:. Later two further main ranks were introduced, making the sequence kingdom, 170: 8920: 8692: 8631: 8575: 8510: 8370: 8351: 7733:"Kingdoms Protozoa and Chromista and the eozoan root of the eukaryotic tree" 7449: 7387: 6950:"Kingdoms Protozoa and Chromista and the eozoan root of the eukaryotic tree" 6822:
Cavalier-Smith, Thomas (March 26, 1987). "Eucaryotes with no mitochondria".
6229: 8928: 8882: 8873: 8854: 8834: 8803: 8650: 8534: 8495: 8430: 8379: 8336: 8285: 8267: 8231: 8155: 8118:"Evaluating support for the current classification of eukaryotic diversity" 8102: 8051: 7997: 7924: 7868: 7825: 7766: 7748: 7708: 7690: 7244: 7098: 7037: 7019: 6983: 6965: 6905: 6808: 6568: 6465: 6428: 6369: 6329: 5674: 5605: 5535: 5499: 4299: 4203: 4198: 4163: 4143: 3919: 3879: 3790: 3674: 3536: 3497: 3455: 3264: 3120: 2907: 2872: 2867: 2806: 2745: 2740: 2720: 2710: 2705: 2640: 2630: 2590: 2570: 2550: 2212: 2204: 1839: 1831: 1560: 1548: 1536: 1243: 1147: 1105: 1085: 1079: 643: 280: 174: 161:(for fungi) are also used for life present in a particular region or time. 8711: 8093: 8074: 7645: 7603: 7554: 7506: 7468: 7290: 6853: 6533: 6298: 6248: 8412: 8318: 7406: 7336: 7080: 6701: 6643: 6442:
Stanier, R.Y. & Van Neil, C.B. (1962). "The concept of a bacterium".
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contains one or more kingdoms. Intermediate minor rankings are not shown.
8795: 7363:"Phylogenetic structure of the prokaryotic domain: the primary kingdoms" 6614: 6550:
Kingdoms and Domains: An Illustrated Guide to the Phyla of Life on Earth
5677:. Plants are thought to be more distantly related to animals and fungi. 4202:          Infraphylum 4197:          Infraphylum 4187:          Infraphylum 4182:          Infraphylum 4177:          Infraphylum 4172:          Infraphylum 4157:          Infraphylum 4152:          Infraphylum 4119:          Infraphylum 4114:          Infraphylum 4104:          Infraphylum 4099:          Infraphylum 2896:          Infraphylum 2891:          Infraphylum 2754:          Infraphylum 2749:          Infraphylum 2724:          Infraphylum 2719:          Infraphylum 9367: 7974: 7420: 7358: 7328: 7304: 6693: 6457: 6015: 5971: 5847: 5684:
One hypothesis of eukaryotic relationships depicted by Alastair Simpson
5616: 5580: 5576: 5564: 5539: 5460: 5204: 5159: 5083: 5066: 4836: 4826: 4811: 4671: 4551: 4519: 4385: 4322: 4316: 4269: 4127: 4059: 4053: 3990: 3904: 3848: 3750: 3692: 3646: 3631: 3596: 3551: 3424: 3223: 3139:          Superclass 3124:          Superclass 3046: 3003: 2935: 2920: 2786: 2560: 2455: 2423: 2300: 2263: 2196: 1950: 1936: 1827: 1819: 1650: 1636: 1590: 1528: 1486: 1476: 1458: 1419: 1405: 1328: 1322: 1223: 1197: 1191: 1183: 1177: 1159: 1117: 1062: 909: 874: 728: 647: 639: 124: 8248:
Burki, Fabien; Shalchian-Tabrizi, Kamran & Pawlowski, Jan (2008).
5680: 4308:          Subphylum 4303:          Subphylum 4298:          Subphylum 4288:          Subphylum 4273:          Subphylum 4263:          Subphylum 4258:          Subphylum 3989:          Subphylum 3971:          Subphylum 3961:          Subphylum 3946:          Subphylum 3918:          Subphylum 3903:          Subphylum 3898:          Subphylum 3888:          Subphylum 3883:          Subphylum 3847:          Subphylum 3832:          Subphylum 3822:          Subphylum 3807:          Subphylum 3017:          Subphylum 3012:          Subphylum 3007:          Subphylum 2992:          Subphylum 2987:          Subphylum 2982:          Subphylum 2972:          Subphylum 2967:          Subphylum 2684:          Subphylum 2669:          Subphylum 2659:          Subphylum 2654:          Subphylum 2644:          Subphylum 2639:          Subphylum 2634:          Subphylum 2624:          Subphylum 2619:          Subphylum 2599:          Subphylum 2594:          Subphylum 2579:          Subphylum 2574:          Subphylum 885:'s system of five kingdoms, the plants included just the land plants ( 9277: 9148: 9143: 9134: 8183: 8168: 6845: 6606: 6050: 5987: 5967: 5908: 5732: 5588: 5474: 5441: 5292: 5246: 5242: 4934: 4929: 4924: 4895: 4875: 4499: 4495: 4475: 4471: 4453: 4309: 4289: 4284: 4254: 4249: 4222: 4214: 4138: 4032: 3982: 3972: 3936: 3929: 3924: 3909: 3899: 3853: 3780: 3730: 3725: 3681: 3661: 3616: 3611: 3606: 3581: 3546: 3504: 3471: 3341:          Subclass 3336:          Subclass 3308:          Subclass 3303:          Subclass 3293:          Subclass 3288:          Subclass 3278:          Subclass 3273:          Subclass 3268:          Subclass 3253:          Subclass 3248:          Subclass 3094:          Subclass 3089:          Subclass 3025: 2952: 2897: 2877: 2857: 2847: 2801: 2403: 2399: 2379: 2375: 2357: 2274: 2223: 2219: 2089: 1906: 1847: 1706: 1610: 1498: 1470: 1462: 1413: 1387: 1268: 1153: 945: 866: 862: 764: 665:
proposed a four-kingdom classification by creating the novel Kingdom
651: 343: 270: 132: 7966: 6897: 5086:
classification, where there is emphasis in arranging organisms into
1889:
superkingdom, grouping together the five other eukaryotic kingdoms (
9324: 9303: 8454: 8190: 7208: 7063:"Genome Networks Root the Tree of Life between Prokaryotic Domains" 6767: 6587: 6388: 6170: 6054: 6001: 5878: 5873: 5863: 5858: 5620: 5543: 5528: 5482: 5452: 5348: 5331: 5200: 5170: 4917: 4910: 4905: 4900: 4880: 4846: 4841: 4831: 4547: 4523: 4479: 4467: 4399: 4339: 4274: 4259: 3967: 3962: 3952: 3947: 3914: 3813: 3797: 3785: 3759:          Phylum 3740: 3739:          Phylum 3691:          Phylum 3660:          Phylum 3626: 3625:          Phylum 3615:          Phylum 3595:          Phylum 3571: 3566: 3565:          Phylum 3545:          Phylum 3530:          Phylum 3484: 3478: 3466: 3461: 3430: 3408: 3036: 3031: 2978: 2852: 2837: 2780: 2772: 2544: 2451: 2427: 2383: 2371: 2314: 2267: 2259: 2247: 2235: 2192: 2073: 2019: 2004: 1894: 1890: 1882: 1720: 1606: 1566: 1522: 1504: 1431: 1399: 1370: 1348: 1299: 1291: 1135: 1097: 1008: 959: 811: 778: 670: 571: 538: 507: 463: 140: 109: 9003: 3386:          Class 3366:          Class 3356:          Class 3114:          Class 1818:
supported the consensus at that time, that the difference between
9361: 8751: 8393:
Rogozin, I.B.; Basu, M.K.; Csürös, M. & Koonin, E.V. (2009).
8080:
International Journal of Systematic and Evolutionary Microbiology
5963: 5913: 5900: 5895: 5890: 5885: 5736: 5666: 5572: 5403: 5386: 5221: 5174: 4890: 4885: 4868: 4858: 4806: 4515: 4415: 4355: 4343: 4188: 4024: 4013: 4007: 3995: 3735: 3651: 3531: 3526: 3491: 3416: 2419: 1902: 1859: 1851: 1482: 1393: 1359: 1169: 976: 795: 674: 555: 334: 214: 190: 105: 101: 62: 8779: 9116: 9100: 8247: 7157:. Réflexions sur la biologie et la phylogénie des protozoaires" 6665: 6663: 6084:. Fungal Conservation Committee, IUCN SSC. 2021. Archived from 5934: 5920: 5868: 5852: 5815: 5724: 5669:) are descended from the same ancestor as both the unicellular 5510: 5506: 5045: 5040: 5035: 5030: 5025: 5020: 5015: 5010: 5003: 4998: 4993: 4988: 4983: 4821: 4816: 4587: 4571: 4294: 2958: 2887: 2882: 2818: 2491: 2475: 2137: 2121: 1769: 1753: 1051: 923: 858: 742: 666: 469: 374: 322: 194: 117: 97: 89: 78: 7161:
Annales des Sciences Naturelles - Zoologie et Biologie Animale
6049:
Compared to the version Cavalier-Smith published in 2004, the
2218:
Cavalier-Smith does not accept the requirement for taxa to be
9318: 5927: 5830: 5811: 5784: 5693: 5087: 5062: 4976: 4971: 4966: 4961: 4956: 4951: 4946: 4539: 4148: 3058: 2554:        Infrakingdom 2443: 2255: 2105: 1898: 1737: 1257: 992: 849: 330: 248:) can be inserted between subkingdom and infrakingdom, e.g., 210: 186: 157: 151: 145: 93: 8115: 8018: 6660: 6393:"The Prokaryote-Eukaryote Dichotomy: Meanings and Mythology" 6264: 6262: 6260: 6258: 3754:        Superphylum 3686:        Superphylum 3620:        Superphylum 3610:        Superphylum 3560:        Superphylum 3525:        Superphylum 3119:        Infraphylum 3109:        Infraphylum 8775: 8773: 8670: 8609: 6057:
have been moved from Kingdom Protozoa to Kingdom Chromista.
5788: 5312: 5185: 5147: 4133: 3361:        Superclass 3351:        Superclass 1237: 317:, in 1735. He distinguished two kingdoms of living things: 8605:"The eocyte hypothesis and the origin of eukaryotic cells" 8298: 7779: 6760: 4192:        Subphylum 4167:        Subphylum 4147:        Subphylum 4142:        Subphylum 4137:        Subphylum 4109:        Subphylum 4094:        Subphylum 4068:        Subphylum 4063:        Subphylum 3470:        Subphylum 3465:        Subphylum 3444:        Subphylum 3439:        Subphylum 3434:        Subphylum 3045:        Subphylum 3040:        Subphylum 2944:        Subphylum 2939:        Subphylum 2929:        Subphylum 2924:        Subphylum 2886:        Subphylum 2881:        Subphylum 2866:        Subphylum 2861:        Subphylum 2856:        Subphylum 2846:        Subphylum 2841:        Subphylum 2836:        Subphylum 2759:        Subphylum 2744:        Subphylum 2729:        Subphylum 2714:        Subphylum 8349: 8073:
Harper, J. T.; Waanders, E. & Keeling, P. J. (2005).
7261:(January 1969). "New concepts of kingdoms of organisms". 7061:
Dagan, T.; Roettger, M.; Bryant & Martin, W. (2010).
7060: 6255: 5708: 3202:        Subclass 8895: 8770: 8508: 6634:
Palmer, E. Laurence; Fowler, Seymour H. (January 1975).
5955:); but this is beneath the top level classifications of 3207:        Subclas 650:) and organisms whose cells do have a distinct nucleus ( 8663: 8392: 7782:"A higher level classification of all living organisms" 7679:
Proceedings of the Royal Society B: Biological Sciences
6763:"A higher level classification of all living organisms" 5907: 4293:        Phylum 4268:        Phylum 4253:        Phylum 3966:        Phylum 3941:        Phylum 3893:        Phylum 3878:        Phylum 3827:        Phylum 3802:        Phylum 3673:        Phylum 3002:        Phylum 2977:        Phylum 2962:        Phylum 2694:        Phylum 2664:        Phylum 2649:        Phylum 2629:        Phylum 2614:        Phylum 2589:        Phylum 2569:        Phylum 2564:        Phylum 8852: 7841:"Evolution: Revisiting the Root of the Eukaryote Tree" 6756: 6754: 6752: 3331:        Class 3313:        Class 3298:        Class 3283:        Class 3263:        Class 3243:        Class 3238:        Class 3233:        Class 3099:        Class 3084:        Class 8186:"Phylogenomics reshuffles the eukaryotic supergroups" 8072: 6134:
International Union for Conservation of Nature (IUCN)
5715:
Tree of life (biology) § Developments since 1990
8846: 6940: 6938: 5997: 3985: 3715: 3448: 2242:. This means that all living eukaryotes are in fact 1597:
structure; this would later lead to the proposal of
1379: 481:, often called the "father of microscopy", sent the 8889: 7890: 7888: 7886: 7419: 7303: 6749: 6669: 6202: 6635: 6586: 6270: 5154:. All smaller branches can be considered kingdoms. 164: 77:. Kingdoms are divided into smaller groups called 8443: 7895:Simpson, Alastair G.B.; Roger, Andrew J. (2004). 7660: 6935: 6874: 6349: 6347: 6345: 6343: 6341: 6339: 9406: 8855:"Reasons to include viruses in the tree of life" 8758:. International Committee on Taxonomy of Viruses 8511:"Acytota - associated kingdom of neglected life" 7883: 7056: 7054: 6203:Woese, C.R.; Kandler, O.; Wheelis, M.L. (1990). 4248:      Superphylum 2997:      Superphylum 2957:      Superphylum 2689:      Superphylum 2609:      Superphylum 2584:      Superphylum 131:, noting that some traditional kingdoms are not 6546: 6441: 5111: 33: 7727: 7666: 7616: 7567: 7519: 7485:(1981). "Eukaryote kingdoms: Seven or nine?". 7481: 6996: 6944: 6821: 6336: 6268: 5949:International Committee on Taxonomy of Viruses 5926: 5743:(formerly known as eocytes or Crenarchaeota). 5077: 3346:      Subphylum 3326:      Subphylum 3258:      Subphylum 3228:      Subphylum 3104:      Subphylum 3079:      Subphylum 2810:      Subphylum 2805:      Subphylum 2795:      Subphylum 2790:      Subphylum 273:(384–322 BC) classified animal species in his 135:, meaning that they do not consist of all the 9019: 8827:"New views on the megaclassification of life" 8727:"New views on the megaclassification of life" 8437: 7990:"New views on the megaclassification of life" 7948:"Proposal for the recognition of super ranks" 7251: 7051: 6875:Poole, Anthony; Penny, David (21 June 2007). 6580: 6578: 6487: 6485: 6483: 5919: 5867: 889:), and Protoctista has a broader definition. 8558:Biological systematics: The state of the art 8243: 8241: 8179: 8177: 7894: 7838: 7721: 7223: 7185: 7105: 7007:Proceedings of the Royal Society of London B 6633: 6146: 5846: 309:(1707–1778) laid the foundations for modern 123:Some recent classifications based on modern 8818: 8343: 8162: 7523:(1992). "Origins of secondary metabolism". 7179:Titres et Travaux Scientifiques (1906–1937) 7141: 6353: 4321:      Phylum 4162:      Phylum 4132:      Phylum 4089:      Phylum 4058:      Phylum 4045:      Phylum 3483:      Phylum 3460:      Phylum 3429:      Phylum 3035:      Phylum 3030:      Phylum 2934:      Phylum 2919:      Phylum 2876:      Phylum 2871:      Phylum 2851:      Phylum 2831:      Phylum 2739:      Phylum 2709:      Phylum 505:also proposed a third kingdom of life, the 9026: 9012: 8588:: CS1 maint: location missing publisher ( 8014: 8012: 7981: 7839:Roger, A.J. & Simpson, A.G.B. (2009). 7351: 6575: 6480: 6398:Microbiology and Molecular Biology Reviews 6383: 6381: 6379: 6079:"IUCN SSC acceptance of Fauna Flora Funga" 5837: 5792: 5783: 5665:In this system the multicellular animals ( 4346:. (This was based on the consensus in the 3197:      Class 3192:      Class 2534:Cavalier-Smith's six kingdom system (1998) 1598: 8872: 8701: 8691: 8640: 8630: 8602: 8596: 8485: 8475: 8448:(2008). Redfield, Rosemary Jeanne (ed.). 8420: 8369: 8326: 8292: 8275: 8238: 8221: 8211: 8174: 8145: 8135: 8092: 8041: 7939: 7914: 7858: 7815: 7805: 7756: 7698: 7593: 7458: 7448: 7413: 7396: 7386: 7297: 7257: 7088: 7027: 6973: 6798: 6788: 6515: 6491: 6418: 6238: 6228: 5912: 5824: 5822: 5820: 5807: 5805: 5274: 4348:Taxonomic Outline of Bacteria and Archaea 857:recognized an additional kingdom for the 450: 256:in the classification of Cavalier-Smith. 155:(for animals), and, in the 21st century, 8744: 7570:"Kingdom protozoa and its 18 phyla" 7357: 7229: 7191: 7111: 5933: 5679: 5107: 5093: 4791: 865:; his Plantae were mostly multicellular 462: 390:Regnum Vegetabile ('vegetables'/plants) 116:(Animalia, Plantae, Fungi, Protista and 29: 8981:A Brief History of the Kingdoms of Life 8603:Archibald, John M. (23 December 2008). 8555: 8109: 8009: 7239:. Palo Alto: Pacific Books. p. 6. 7173: 7167: 7147: 7129: 7123: 6588:"Teaching Taxonomy: How Many Kingdoms?" 6435: 6376: 6198: 6196: 6194: 6163:McNeill, J.; et al., eds. (2006). 6162: 5990:, that possess typical cellular genes. 5899: 1885:. This superkingdom was opposed to the 592:  Non‑life   409:  Non‑life   264: 14: 9407: 8955:Empire Biota: a comprehensive taxonomy 8824: 8724: 7987: 7622:"A revised six-kingdom system of life" 6319: 6272:"A revised six-kingdom system of life" 5962:There is ongoing debate as to whether 5709:Comparison of top level classification 2258:. Others were reclassified in kingdom 2185: 1042:Whittaker's five kingdom system (1969) 9007: 7945: 7236:The Classification of Lower Organisms 7195:(1938). "The kingdoms of organisms". 6547:Margulis L, Chapman MJ (2009-03-19). 5166:accurately introduced term dominion. 2734:    Infrakingdom 2704:    Infrakingdom 2604:    Infrakingdom 2207:. Unibacteria was divided into phyla 712:of Archaea, Bacteria, and Eukaryota. 9043: 8657: 7135:Generelle Morphologie der Organismen 6584: 6387: 6191: 5057:Kingdoms such as Protozoa represent 3933:       3870:       3794:       3678:       3665:       3517:       2531: 1039: 173:introduced the rank-based system of 9033: 7897:"The real 'kingdoms' of eukaryotes" 7423:; Kandler, O.; Wheelis, M. (1990). 4796: 4786: 2549:    Subkingdom 1862:. Moreover, only chromists contain 1602: 341:, placing them in a third kingdom, 127:have explicitly abandoned the term 24: 8947: 8029:Journal of Eukaryotic Microbiology 7638:10.1111/j.1469-185X.1998.tb00030.x 6291:10.1111/j.1469-185X.1998.tb00030.x 4773: 4768: 2266:which is now part of infrakingdom 677:. Ernst Haeckel, in his 1904 book 113: 25: 9431: 8974: 8509:Trifonov EN, Kejnovsky E (2016). 5877: 5872: 5862: 5857: 5752:Two superdomains (controversial) 5735:emerged from a phylum within the 4778: 4332: 2254:, were reclassified into kingdom 1810: 85: 9063: 9052: 8043:10.1111/j.1550-7408.2005.00053.x 7527:. Novartis Foundation Symposia. 6000: 5894: 843: 633: 9257: 9196: 8718: 8386: 8357:Molecular Biology and Evolution 8066: 7832: 7773: 7610: 7561: 7513: 7475: 6990: 6868: 6815: 6716: 6627: 6540: 6043: 5889: 5884: 4227:    Phylum 4012:    Phylum 3496:    Phylum 2800:    Phylum 2785:    Phylum 1584: 165:Definition and associated terms 9386: 9375: 9343: 9332: 9125: 9107: 8970:, New York: Worth Publishers. 7586:10.1128/mmbr.57.4.953-994.1993 7309:Journal of Molecular Evolution 6673:Journal of Molecular Evolution 6411:10.1128/MMBR.69.2.292-305.2005 6313: 6155: 6140: 6122: 6071: 5828: 2273:Because Cavalier-Smith allows 873:, and his Fungi multicellular 221:was introduced above kingdom. 13: 1: 8527:10.1080/07391102.2015.1086959 8444:Kim, E.; Graham, L. E. & 7361:; Fox, G.E. (November 1977). 6064: 5673:and the fungi which form the 4369:  Life   2284:  Life   2191:recent research. The kingdom 1920:  Life   1620:  Life   900:  Life   869:, his Animalia multicellular 719:  Life   526:  Life   362:  Life   297:) wrote a parallel work, the 291: 284: 8556:Minelli, Alessandro (1993). 8477:10.1371/journal.pone.0002621 8400:Genome Biology and Evolution 8306:Genome Biology and Evolution 8213:10.1371/journal.pone.0000790 8137:10.1371/journal.pgen.0020220 7807:10.1371/journal.pone.0119248 7499:10.1016/0303-2647(81)90050-2 7283:10.1126/science.163.3863.150 7068:Genome Biology and Evolution 6790:10.1371/journal.pone.0119248 6638:Fieldbook of Natural History 6526:10.1126/science.163.3863.150 1304:Branch Opisthomastigomycota 7: 8953:Pelentier, B. (2007-2015). 8860:Nature Reviews Microbiology 8783:Nature Reviews Microbiology 7672:"Only six kingdoms of life" 7568:Cavalier-Smith, T. (1993). 7197:Quarterly Review of Biology 7181:. E. Sottano, Sète, France. 6999:"Only six kingdoms of life" 6997:Cavalier-Smith, T. (2004). 6324:. Oxford: Clarendon Press. 6320:Singer, Charles J. (1931). 6269:Cavalier-Smith, T. (1998). 5993: 5158:Based on such RNA studies, 5078:Beyond traditional kingdoms 5067:phylogenetic classification 1287:Subkingdom Dimastigomycota 603:Regnum Lapideum (minerals) 420:Regnum Lapideum (minerals) 10: 9436: 8999:Whittaker's classification 8966:and Helena Curtis (1970), 7531:: 64–80, discussion 80–7. 6585:Case, Emily (2008-10-01). 6357:International Microbiology 6111:animals, fungi, and plants 5942: 5712: 5689:six proposed supergroups. 5280: 5097: 4628: 2178:‡ No longer recognized by 456: 333:). Linnaeus also included 329:('vegetable kingdom', for 259: 9041: 8446:Redfield, Rosemary Jeanne 7916:10.1016/j.cub.2004.08.038 7860:10.1016/j.cub.2008.12.032 7537:10.1002/9780470514344.ch5 7525:Ciba Foundation Symposium 6877:"Engulfed by speculation" 5610: 5562: 5533: 5504: 5479: 5457: 5446: 5434: 5398: 5391: 5379: 5343: 5336: 5324: 5317: 5239: 5218: 5197: 5190: 5065:, and so are rejected by 4950: 4945: 4940: 4933: 4928: 4923: 4916: 4909: 4904: 4899: 4894: 4889: 4884: 4879: 4874: 4840: 4820: 4815: 4810: 4805: 4802: 4584: 4568: 4536: 4492: 4464: 4457: 4446: 4412: 4396: 4389: 4378: 4371: 4358:is any of the eukaryotic 2488: 2472: 2440: 2396: 2368: 2361: 2350: 2311: 2304: 2293: 2286: 2134: 2118: 2102: 2086: 2070: 2063: 2051: 2016: 2009: 1997: 1963: 1947: 1940: 1929: 1922: 1909:). This was known as the 1766: 1750: 1734: 1717: 1710: 1699: 1664: 1647: 1640: 1629: 1622: 1005: 989: 973: 956: 949: 920: 913: 902: 808: 792: 775: 768: 739: 732: 721: 601: 594: 568: 552: 535: 528: 520: 418: 411: 388: 371: 364: 356: 224:Prefixes can be added so 47:biological classification 8987:The five kingdom concept 6594:American Biology Teacher 6445:Archiv fĂĽr Mikrobiologie 6036: 5727:" or cellular life. The 5719:Some authors have added 4313:     4240:     4124:     4081:     4050:     4037:     3862:     3509:     3475:     3452:     3421:     3318:     3220:     3184:     3071:     3022:     2949:     2911:     2823:     2321:as part of a subkingdom 2195:(sole kingdom of empire 848:The differences between 8921:10.1126/science.1101485 8825:Luketa, Stefan (2012). 8725:Luketa, Stefan (2012). 8693:10.1073/pnas.81.12.3786 8632:10.1073/pnas.0811118106 7574:Microbiological Reviews 7450:10.1073/pnas.87.12.4576 7388:10.1073/pnas.74.11.5088 6230:10.1073/pnas.87.12.4576 6115:fauna, flora, and funga 5974:and are not capable of 5587:(or cryptomonads), and 2699:  Subkingdom 1599:three "domains" of life 1264:Subkingdom Gymnomycota 479:Antonie van Leeuwenhoek 313:, now regulated by the 311:biological nomenclature 217:. In 1990, the rank of 8874:10.1038/nrmicro2108-c1 8268:10.1098/rsbl.2008.0224 7749:10.1098/rsbl.2009.0948 7691:10.1098/rspb.2004.2705 7245:10.5962/bhl.title.4474 7020:10.1098/rspb.2004.2705 6966:10.1098/rsbl.2009.0948 6946:Cavalier-Smith, Thomas 5749:Taxonomical root node 5685: 5275:Eukaryotic supergroups 5155: 1844:Gram-positive bacteria 1836:Gram-negative bacteria 1830:into two subkingdoms: 474: 459:Tree of life (biology) 451:Three kingdoms of life 69:is the second highest 58: 8371:10.1093/molbev/msm089 8094:10.1099/ijs.0.63216-0 6147:Linnaeus, C. (1735). 5903:including eukaryotes 5848:Prokaryota / Procarya 5713:Further information: 5683: 5150:data showing Woese's 5141: 5094:Three domains of life 4360:unicellular organisms 1856:endoplasmic reticulum 1816:Thomas Cavalier-Smith 1355:Subkingdom Agnotozoa 1072:Branch Mastigomonera 466: 457:Further information: 339:classification system 108:/Archaebacteria, and 44: 7946:Moore, R.T. (1974). 7155:Pansporella perplexa 6732:Rhode Island College 5981:nucleotide sequences 5542:, and various other 5283:Supergroup (biology) 1541:Phylum Brachiata or 1130:Plasmodiophoromycota 1124:Hyphochytridiomycota 1067:Phylum Myxobacteriae 265:Two kingdoms of life 9420:Biology terminology 9092:Infrakingdom/Branch 8983:at Earthling Nature 8913:2004Sci...306.1344R 8907:(5700): 1344–1350. 8796:10.1038/nrmicro2108 8756:talk.ictvonline.org 8684:1984PNAS...81.3786L 8623:2008PNAS..10520049A 8617:(51): 20049–20050. 8515:J Biomol Struct Dyn 8468:2008PLoSO...3.2621K 8204:2007PLoSO...2..790B 7988:Luketa, S. (2012). 7798:2015PLoSO..1019248R 7685:(1545): 1251–1262. 7441:1990PNAS...87.4576W 7379:1977PNAS...74.5088W 7321:1977JMolE...9..305B 7275:1969Sci...163..150W 7014:(1545): 1251–1262. 6838:1987Natur.326..332C 6781:2015PLoSO..1019248R 6686:1977JMolE...9..305B 6508:1969Sci...163..150W 6221:1990PNAS...87.4576W 5909:Eukaryota / Eukarya 5152:three-domain system 5100:Three-domain system 3310:Loculoascomycetidae 2536: 2186:Six kingdoms (1998) 1911:Archezoa hypothesis 1366:Subkingdom Parazoa 1044: 710:three-domain system 679:The Wonders of Life 663:Herbert F. Copeland 638:The development of 81:(singular phylum). 9415:Kingdoms (biology) 9364: 9321: 9300: 9274: 9232: 9166: 9131: 9119: 9083: 9074: 8992:2021-11-07 at the 8413:10.1093/gbe/evp011 8319:10.1093/gbe/evp022 7729:Cavalier-Smith, T. 7668:Cavalier-Smith, T. 7626:Biological Reviews 7618:Cavalier-Smith, T. 7521:Cavalier-Smith, T. 7483:Cavalier-Smith, T. 7329:10.1007/BF01796092 7081:10.1093/gbe/evq025 6724:"The Six Kingdoms" 6694:10.1007/BF01796092 6458:10.1007/BF00425185 6278:Biological Reviews 6103:animals and plants 5983:from their hosts. 5731:proposes that the 5686: 5156: 4452:Superkingdom  4384:Superkingdom  3388:Homobasidiomycetes 3353:Hemibasidiomycetia 2532: 2250:, like the phylum 2057:Superkingdom  2003:Superkingdom  1935:Superkingdom  1858:instead of in the 1281:Labyrinthulomycota 1075:Phylum Eubacteriae 1058:Branch Myxomonera 1040: 475: 315:Nomenclature Codes 300:Historia Plantarum 279:, while his pupil 276:History of Animals 195:phylum or division 114:five kingdoms only 59: 9402: 9401: 9397: 9396: 9360: 9317: 9296: 9270: 9228: 9162: 9121: 9115: 9079: 9072: 8968:Biology of Plants 8678:(12): 3786–3790. 8169:Burki et al. 2007 8087:(Pt 1): 487–496. 7909:(17): R693–R696. 7435:(12): 4576–4579. 7373:(11): 5088–5090. 7269:(3863): 150–160. 7137:. Reimer, Berlin. 6832:(6111): 332–333. 6653:978-0-070-48425-2 6173:on 6 October 2012 6109:be replaced with 5940: 5939: 5802:non-cellular life 5778:Eocyte hypothesis 5729:eocyte hypothesis 5721:non-cellular life 5671:choanoflagellates 5662: 5661: 5653: 5652: 5644: 5643: 5635: 5634: 5515:choanoflagellates 5423: 5422: 5414: 5413: 5368: 5367: 5359: 5358: 5267: 5266: 5258: 5257: 5144:phylogenetic tree 5051: 5050: 4625: 4624: 4616: 4615: 4607: 4606: 4598: 4597: 4435: 4434: 4426: 4425: 4352:Catalogue of Life 4330: 4329: 4070:Macrorhodophytina 3373:Tremellomycetidae 3338:Sporidiomycetidae 3255:Saccharomycetidae 2529: 2528: 2520: 2519: 2511: 2510: 2502: 2501: 2339: 2338: 2330: 2329: 2175: 2174: 2166: 2165: 2157: 2156: 2148: 2147: 2040: 2039: 2031: 2030: 1986: 1985: 1977: 1976: 1807: 1806: 1798: 1797: 1789: 1788: 1780: 1779: 1688: 1687: 1679: 1678: 1582: 1581: 1037: 1036: 1028: 1027: 1019: 1018: 934: 933: 840: 839: 831: 830: 822: 821: 753: 752: 706:two-empire system 630: 629: 621: 620: 612: 611: 582: 581: 447: 446: 438: 437: 429: 428: 399: 398: 327:Regnum Vegetabile 45:The hierarchy of 16:(Redirected from 9427: 9388: 9377: 9345: 9334: 9259: 9198: 9127: 9109: 9065: 9054: 9044: 9028: 9021: 9014: 9005: 9004: 8941: 8940: 8893: 8887: 8886: 8876: 8850: 8844: 8843: 8831: 8822: 8816: 8815: 8777: 8768: 8767: 8765: 8763: 8748: 8742: 8741: 8731: 8722: 8716: 8715: 8705: 8695: 8661: 8655: 8654: 8644: 8634: 8600: 8594: 8593: 8587: 8579: 8553: 8547: 8546: 8506: 8500: 8499: 8489: 8479: 8441: 8435: 8434: 8424: 8390: 8384: 8383: 8373: 8364:(8): 1702–1713. 8347: 8341: 8340: 8330: 8296: 8290: 8289: 8279: 8245: 8236: 8235: 8225: 8215: 8181: 8172: 8166: 8160: 8159: 8149: 8139: 8113: 8107: 8106: 8096: 8070: 8064: 8063: 8045: 8025: 8016: 8007: 8006: 7994: 7985: 7979: 7978: 7952: 7943: 7937: 7936: 7918: 7892: 7881: 7880: 7862: 7836: 7830: 7829: 7819: 7809: 7777: 7771: 7770: 7760: 7725: 7719: 7718: 7716: 7715: 7702: 7676: 7664: 7658: 7657: 7614: 7608: 7607: 7597: 7565: 7559: 7558: 7517: 7511: 7510: 7493:(3–4): 461–481. 7479: 7473: 7472: 7462: 7452: 7417: 7411: 7410: 7400: 7390: 7355: 7349: 7348: 7301: 7295: 7294: 7255: 7249: 7248: 7227: 7221: 7220: 7189: 7183: 7182: 7171: 7165: 7164: 7163:. 10-VIII: 5–84. 7145: 7139: 7138: 7127: 7121: 7120: 7109: 7103: 7102: 7092: 7058: 7049: 7048: 7046: 7044: 7031: 7003: 6994: 6988: 6987: 6977: 6942: 6933: 6932: 6930: 6928: 6922: 6916:. Archived from 6881: 6872: 6866: 6865: 6846:10.1038/326332a0 6819: 6813: 6812: 6802: 6792: 6758: 6747: 6746: 6744: 6743: 6734:. Archived from 6720: 6714: 6713: 6667: 6658: 6657: 6642:(2nd ed.). 6641: 6631: 6625: 6624: 6622: 6621: 6607:10.2307/30163328 6590: 6582: 6573: 6572: 6544: 6538: 6537: 6519: 6502:(3863): 150–60. 6489: 6478: 6477: 6439: 6433: 6432: 6422: 6385: 6374: 6373: 6351: 6334: 6333: 6317: 6311: 6310: 6274: 6266: 6253: 6252: 6242: 6232: 6200: 6189: 6188: 6181: 6179: 6178: 6159: 6153: 6152: 6144: 6138: 6137: 6136:. 3 August 2021. 6126: 6120: 6119: 6097: 6096: 6090: 6083: 6075: 6058: 6047: 6010: 6005: 6004: 5746: 5745: 5449: 5448: 5437: 5436: 5394: 5393: 5382: 5381: 5339: 5338: 5327: 5326: 5320: 5319: 5308: 5307: 5299: 5193: 5192: 5181: 5180: 5110: 5104:Domain (biology) 4744:3 superkingdoms 4633: 4632: 4460: 4459: 4449: 4448: 4392: 4391: 4381: 4380: 4374: 4373: 4366: 4365: 4325: 4206: 4016: 3987: 3717: 3450: 3400: 3290:Verrucomycetidae 3230:Hemiascomycotina 3086:Chytridiomycetes 3050: 2764: 2681:Chromatibacteria 2621:Chlorobibacteria 2537: 2535: 2364: 2363: 2353: 2352: 2307: 2306: 2296: 2295: 2289: 2288: 2281: 2280: 2066: 2065: 2054: 2053: 2012: 2011: 2000: 1999: 1943: 1942: 1932: 1931: 1925: 1924: 1917: 1916: 1713: 1712: 1702: 1701: 1643: 1642: 1632: 1631: 1625: 1624: 1617: 1616: 1381: 1212:Chlorophycophyta 1045: 1043: 952: 951: 916: 915: 905: 904: 897: 896: 855:Robert Whittaker 771: 770: 735: 734: 724: 723: 716: 715: 597: 596: 531: 530: 523: 522: 515: 514: 414: 413: 367: 366: 359: 358: 351: 350: 296: 293: 289: 286: 21: 9435: 9434: 9430: 9429: 9428: 9426: 9425: 9424: 9405: 9404: 9403: 9398: 9356:Species complex 9106:Superdivision ( 9037: 9035:Taxonomic ranks 9032: 8994:Wayback Machine 8977: 8950: 8948:Further reading 8945: 8944: 8894: 8890: 8851: 8847: 8829: 8823: 8819: 8778: 8771: 8761: 8759: 8750: 8749: 8745: 8729: 8723: 8719: 8662: 8658: 8601: 8597: 8581: 8580: 8568: 8554: 8550: 8507: 8503: 8442: 8438: 8391: 8387: 8348: 8344: 8297: 8293: 8255:Biology Letters 8246: 8239: 8182: 8175: 8167: 8163: 8114: 8110: 8071: 8067: 8023: 8017: 8010: 7992: 7986: 7982: 7967:10.2307/1218807 7950: 7944: 7940: 7902:Current Biology 7893: 7884: 7846:Current Biology 7837: 7833: 7792:(4): e0119248. 7778: 7774: 7726: 7722: 7713: 7711: 7674: 7665: 7661: 7615: 7611: 7566: 7562: 7547: 7518: 7514: 7480: 7476: 7418: 7414: 7356: 7352: 7302: 7298: 7259:Whittaker, R.H. 7256: 7252: 7228: 7224: 7190: 7186: 7172: 7168: 7146: 7142: 7128: 7124: 7110: 7106: 7059: 7052: 7042: 7040: 7001: 6995: 6991: 6954:Biology Letters 6943: 6936: 6926: 6924: 6920: 6898:10.1038/447913a 6879: 6873: 6869: 6820: 6816: 6775:(4): e0119248. 6759: 6750: 6741: 6739: 6722: 6721: 6717: 6668: 6661: 6654: 6632: 6628: 6619: 6617: 6583: 6576: 6565: 6545: 6541: 6517:10.1.1.403.5430 6490: 6481: 6440: 6436: 6386: 6377: 6352: 6337: 6318: 6314: 6267: 6256: 6201: 6192: 6183: 6176: 6174: 6160: 6156: 6145: 6141: 6128: 6127: 6123: 6107:fauna and flora 6094: 6092: 6088: 6081: 6077: 6076: 6072: 6067: 6062: 6061: 6048: 6044: 6039: 6006: 5999: 5996: 5945: 5850: 5841: 5800: 5765:Five Dominiums 5717: 5711: 5663: 5654: 5645: 5636: 5604: (or  5424: 5415: 5369: 5360: 5297: 5285: 5277: 5268: 5259: 5140: 5108: 5106: 5098:Main articles: 5096: 5080: 4753:2 superkingdoms 4707: 4698: 4693: 4686: 4678: 4667: 4660: 4653: 4646: 4639: 4631: 4626: 4617: 4608: 4599: 4436: 4427: 4335: 4320: 4312: 4307: 4302: 4297: 4292: 4287: 4282: 4277: 4272: 4267: 4262: 4257: 4252: 4247: 4239: 4231: 4226: 4218: 4211: 4201: 4196: 4194:Spermatophytina 4191: 4186: 4181: 4176: 4171: 4166: 4161: 4156: 4151: 4146: 4141: 4136: 4131: 4123: 4118: 4113: 4108: 4103: 4098: 4093: 4088: 4080: 4072: 4067: 4065:Rhodellophytina 4062: 4057: 4049: 4044: 4036: 4028: 4021: 4011: 4003: 3998: 3993: 3988: 3980: 3978:Cephalochordata 3975: 3970: 3965: 3960: 3955: 3950: 3945: 3940: 3932: 3927: 3922: 3917: 3912: 3907: 3902: 3897: 3892: 3887: 3882: 3877: 3869: 3861: 3859:Cercomeromorpha 3856: 3851: 3846: 3841: 3836: 3831: 3829:Platyhelminthes 3826: 3821: 3816: 3811: 3806: 3801: 3793: 3788: 3783: 3778: 3773: 3768: 3763: 3761:Nemathelminthes 3758: 3756:Nemathelminthes 3753: 3748: 3743: 3738: 3733: 3728: 3723: 3718: 3713:Trilobitomorpha 3710: 3705: 3700: 3698:Cheliceromorpha 3695: 3690: 3685: 3677: 3672: 3664: 3659: 3654: 3649: 3644: 3639: 3634: 3629: 3624: 3619: 3614: 3609: 3604: 3599: 3594: 3589: 3584: 3579: 3574: 3569: 3564: 3559: 3554: 3549: 3544: 3539: 3534: 3529: 3524: 3516: 3508: 3500: 3495: 3487: 3482: 3474: 3469: 3464: 3459: 3451: 3443: 3438: 3433: 3428: 3420: 3412: 3405: 3395: 3390: 3385: 3380: 3375: 3370: 3365: 3360: 3355: 3350: 3345: 3340: 3335: 3330: 3325: 3317: 3312: 3307: 3305:Dendromycetidae 3302: 3297: 3292: 3287: 3282: 3277: 3272: 3270:Calycomycetidae 3267: 3262: 3257: 3252: 3247: 3242: 3237: 3235:Taphrinomycetes 3232: 3227: 3219: 3211: 3206: 3201: 3196: 3191: 3183: 3178: 3176:Trichomycetalia 3173: 3168: 3163: 3158: 3153: 3151:Mucoromycetidae 3148: 3143: 3138: 3133: 3128: 3123: 3118: 3113: 3108: 3103: 3098: 3093: 3088: 3083: 3078: 3070: 3062: 3055: 3049: 3044: 3039: 3034: 3029: 3021: 3016: 3011: 3009:Tubulicorticata 3006: 3001: 2996: 2991: 2986: 2981: 2976: 2971: 2966: 2961: 2956: 2948: 2943: 2938: 2933: 2928: 2923: 2918: 2910: 2905: 2900: 2895: 2890: 2885: 2880: 2875: 2870: 2865: 2860: 2855: 2850: 2845: 2840: 2835: 2830: 2822: 2814: 2809: 2804: 2799: 2794: 2789: 2784: 2776: 2769: 2763: 2758: 2753: 2748: 2743: 2738: 2733: 2728: 2723: 2718: 2713: 2708: 2703: 2698: 2696:Planctobacteria 2693: 2691:Planctobacteria 2688: 2683: 2678: 2673: 2668: 2663: 2658: 2653: 2648: 2643: 2638: 2633: 2628: 2623: 2618: 2616:Sphingobacteria 2613: 2608: 2603: 2598: 2593: 2588: 2583: 2578: 2573: 2568: 2563: 2558: 2553: 2548: 2541: 2533: 2530: 2521: 2512: 2503: 2340: 2331: 2188: 2176: 2167: 2158: 2149: 2041: 2032: 1987: 1978: 1875:proteobacterium 1813: 1808: 1799: 1790: 1781: 1723:or Protoctista 1689: 1680: 1587: 1441:Pseudocoelomata 1434:or Rhynchocoela 1426:Platyhelminthes 1309:Chytridiomycota 1192:Phaeophycophyta 1178:Rhodophycophyta 1041: 1038: 1029: 1020: 962:or Protoctista 935: 846: 841: 832: 823: 781:or Protoctista 754: 656:Édouard Chatton 636: 631: 622: 613: 583: 541:or Protoctista 461: 453: 448: 439: 430: 400: 344:Regnum Lapideum 294: 287: 267: 262: 236:(also known as 167: 51:taxonomic ranks 49:'s eight major 43: 28: 23: 22: 15: 12: 11: 5: 9433: 9423: 9422: 9417: 9400: 9399: 9395: 9394: 9393: 9392: 9381: 9370: 9365: 9358: 9351: 9350: 9349: 9338: 9327: 9322: 9313: 9312: 9311: 9306: 9301: 9294: 9287: 9286: 9285: 9280: 9275: 9268: 9263: 9250: 9249: 9248: 9243: 9238: 9233: 9226: 9221: 9214: 9213: 9212: 9207: 9202: 9189: 9188: 9187: 9182: 9177: 9172: 9167: 9160: 9153: 9152: 9151: 9146: 9141: 9132: 9113: 9096: 9095: 9094: 9089: 9084: 9077: 9069: 9058: 9042: 9039: 9038: 9031: 9030: 9023: 9016: 9008: 9002: 9001: 8996: 8984: 8976: 8975:External links 8973: 8972: 8971: 8964:Peter H. Raven 8961: 8949: 8946: 8943: 8942: 8888: 8845: 8817: 8790:(4): 306–311. 8769: 8743: 8717: 8656: 8595: 8566: 8548: 8501: 8436: 8385: 8342: 8291: 8262:(4): 366–369. 8237: 8173: 8161: 8108: 8065: 8036:(5): 399–451. 8008: 7980: 7961:(4): 650–652. 7938: 7882: 7831: 7772: 7743:(3): 342–345. 7720: 7659: 7632:(3): 203–266. 7609: 7580:(4): 953–994. 7560: 7545: 7512: 7474: 7412: 7350: 7315:(4): 305–311. 7296: 7250: 7231:Copeland, H.F. 7222: 7209:10.1086/394568 7203:(4): 383–420. 7193:Copeland, H.F. 7184: 7166: 7140: 7122: 7104: 7050: 6989: 6960:(3): 342–345. 6934: 6923:on 6 July 2011 6867: 6814: 6748: 6715: 6680:(4): 305–311. 6659: 6652: 6626: 6601:(8): 472–477. 6574: 6563: 6555:Academic Press 6539: 6479: 6434: 6405:(2): 292–305. 6375: 6335: 6312: 6254: 6215:(12): 4576–9. 6190: 6154: 6139: 6121: 6069: 6068: 6066: 6063: 6060: 6059: 6041: 6040: 6038: 6035: 6034: 6033: 6028: 6023: 6018: 6012: 6011: 6008:Biology portal 5995: 5992: 5959:and subrealm. 5944: 5941: 5938: 5937: 5931: 5930: 5924: 5923: 5917: 5916: 5911: 5905: 5904: 5898: 5896:Archaebacteria 5893: 5888: 5882: 5881: 5876: 5871: 5866: 5861: 5856: 5845: 5835: 5834: 5826: 5825: 5823: 5821: 5819: 5808: 5806: 5804: 5791: 5781: 5780: 5775: 5772: 5766: 5763: 5758: 5753: 5750: 5741:Thermoproteota 5710: 5707: 5698:SAR supergroup 5660: 5659: 5656: 5655: 5651: 5650: 5647: 5646: 5642: 5641: 5638: 5637: 5633: 5632: 5629: 5628: 5609: 5602:Archaeplastida 5597: 5596: 5593: 5592: 5561: 5558:Chromalveolata 5553: 5552: 5549: 5548: 5532: 5524: 5523: 5520: 5519: 5503: 5495: 5494: 5491: 5490: 5478: 5470: 5469: 5466: 5465: 5456: 5447: 5445: 5435: 5433: 5430: 5429: 5426: 5425: 5421: 5420: 5417: 5416: 5412: 5411: 5408: 5407: 5397: 5392: 5390: 5380: 5378: 5375: 5374: 5371: 5370: 5366: 5365: 5362: 5361: 5357: 5356: 5353: 5352: 5342: 5337: 5335: 5325: 5323: 5318: 5316: 5306: 5281:Main article: 5276: 5273: 5265: 5264: 5261: 5260: 5256: 5255: 5252: 5251: 5238: 5235: 5234: 5231: 5230: 5226:Archaebacteria 5217: 5214: 5213: 5210: 5209: 5196: 5191: 5189: 5179: 5175:Archaebacteria 5095: 5092: 5088:natural groups 5079: 5076: 5075: 5074: 5070: 5049: 5048: 5043: 5038: 5033: 5028: 5023: 5018: 5013: 5007: 5006: 5001: 4996: 4991: 4986: 4980: 4979: 4974: 4969: 4964: 4959: 4954: 4949: 4944: 4938: 4937: 4932: 4927: 4921: 4920: 4914: 4913: 4908: 4903: 4898: 4893: 4888: 4883: 4878: 4872: 4871: 4866: 4864:Archaebacteria 4861: 4856: 4854:Archaebacteria 4850: 4849: 4844: 4839: 4834: 4829: 4824: 4819: 4814: 4809: 4804: 4800: 4799: 4794: 4789: 4784: 4781: 4776: 4771: 4766: 4763: 4760: 4756: 4755: 4750: 4745: 4742: 4737: 4732: 4727: 4722: 4717: 4714: 4710: 4709: 4700: 4697:Cavalier-Smith 4695: 4691:Cavalier-Smith 4688: 4680: 4669: 4662: 4655: 4648: 4641: 4630: 4627: 4623: 4622: 4619: 4618: 4614: 4613: 4610: 4609: 4605: 4604: 4601: 4600: 4596: 4595: 4592: 4591: 4583: 4580: 4579: 4576: 4575: 4567: 4564: 4563: 4560: 4559: 4535: 4532: 4531: 4528: 4527: 4491: 4488: 4487: 4484: 4483: 4463: 4458: 4456: 4447: 4445: 4442: 4441: 4438: 4437: 4433: 4432: 4429: 4428: 4424: 4423: 4420: 4419: 4411: 4408: 4407: 4404: 4403: 4395: 4390: 4388: 4379: 4377: 4372: 4370: 4364: 4334: 4333:Seven kingdoms 4331: 4328: 4327: 4208: 4169:Pteridophytina 4111:Phragmophytina 4018: 3963:Appendicularia 3819:Acanthocephala 3766:Scalidorhyncha 3402: 3398:Pileomycetidae 3393:Clavomycetidae 3378:Dacrymycetidae 3343:Uredomycetidae 3295:Ostiomycetidae 3260:Euascomycotina 3141:Neozygomycetia 3106:Melanomycotina 3096:Spizomycetidae 3091:Rumpomycetidae 3081:Dictyomycotina 3052: 2989:Coccidiomorpha 2974:Dinoflagellata 2863:Reticulofilosa 2827:Sarcomastigota 2766: 2751:Halomebacteria 2736:Archaebacteria 2726:Actinobacteria 2716:Teichobacteria 2666:Proteobacteria 2636:Sclenobacteria 2611:Pimelobacteria 2596:Euspirochaetae 2586:Endoflagellata 2576:Chlorobacteria 2527: 2526: 2523: 2522: 2518: 2517: 2514: 2513: 2509: 2508: 2505: 2504: 2500: 2499: 2496: 2495: 2487: 2484: 2483: 2480: 2479: 2471: 2468: 2467: 2464: 2463: 2439: 2436: 2435: 2432: 2431: 2395: 2392: 2391: 2388: 2387: 2367: 2362: 2360: 2351: 2349: 2346: 2345: 2342: 2341: 2337: 2336: 2333: 2332: 2328: 2327: 2324: 2323: 2319:Archaebacteria 2310: 2305: 2303: 2294: 2292: 2287: 2285: 2279: 2240:Hydrogenosomes 2209:Archaebacteria 2187: 2184: 2173: 2172: 2169: 2168: 2164: 2163: 2160: 2159: 2155: 2154: 2151: 2150: 2146: 2145: 2142: 2141: 2133: 2130: 2129: 2126: 2125: 2117: 2114: 2113: 2110: 2109: 2101: 2098: 2097: 2094: 2093: 2085: 2082: 2081: 2078: 2077: 2069: 2064: 2062: 2052: 2050: 2047: 2046: 2043: 2042: 2038: 2037: 2034: 2033: 2029: 2028: 2025: 2024: 2015: 2010: 2008: 1998: 1996: 1993: 1992: 1989: 1988: 1984: 1983: 1980: 1979: 1975: 1974: 1971: 1970: 1967:Archaebacteria 1962: 1959: 1958: 1955: 1954: 1946: 1941: 1939: 1930: 1928: 1923: 1921: 1915: 1879:eukaryogenesis 1824:Archaebacteria 1812: 1811:Eight kingdoms 1809: 1805: 1804: 1801: 1800: 1796: 1795: 1792: 1791: 1787: 1786: 1783: 1782: 1778: 1777: 1774: 1773: 1765: 1762: 1761: 1758: 1757: 1749: 1746: 1745: 1742: 1741: 1733: 1730: 1729: 1726: 1725: 1716: 1711: 1709: 1700: 1698: 1695: 1694: 1691: 1690: 1686: 1685: 1682: 1681: 1677: 1676: 1673: 1672: 1668:Archaebacteria 1663: 1660: 1659: 1656: 1655: 1646: 1641: 1639: 1630: 1628: 1623: 1621: 1615: 1603:seven kingdoms 1586: 1583: 1580: 1579: 1578: 1577: 1576: 1575: 1574: 1573: 1572: 1571: 1570: 1569: 1563: 1557: 1551: 1545: 1533: 1532: 1531: 1525: 1519: 1513: 1507: 1501: 1495: 1489: 1467: 1466: 1465: 1455: 1449: 1447:Acanthocephala 1437: 1436: 1435: 1428: 1410: 1409: 1408: 1402: 1384: 1383: 1382: 1373: 1364: 1363: 1362: 1343: 1342: 1341: 1340: 1339: 1338: 1337: 1331: 1325: 1317:Amastigomycota 1313: 1312: 1311: 1296: 1295: 1294: 1285: 1284: 1283: 1277: 1271: 1252: 1251: 1250: 1249: 1248: 1247: 1246: 1240: 1228: 1227: 1226: 1220: 1202: 1201: 1200: 1188: 1187: 1186: 1164: 1163: 1162: 1156: 1150: 1144: 1138: 1132: 1126: 1120: 1114: 1108: 1092: 1091: 1090: 1089: 1088: 1082: 1076: 1070: 1069: 1068: 1065: 1035: 1034: 1031: 1030: 1026: 1025: 1022: 1021: 1017: 1016: 1013: 1012: 1004: 1001: 1000: 997: 996: 988: 985: 984: 981: 980: 972: 969: 968: 965: 964: 955: 950: 948: 941: 940: 937: 936: 932: 931: 928: 927: 919: 914: 912: 903: 901: 895: 845: 842: 838: 837: 834: 833: 829: 828: 825: 824: 820: 819: 816: 815: 807: 804: 803: 800: 799: 791: 788: 787: 784: 783: 774: 769: 767: 760: 759: 756: 755: 751: 750: 747: 746: 738: 733: 731: 722: 720: 714: 694:C. B. van Niel 688:In the 1960s, 635: 632: 628: 627: 624: 623: 619: 618: 615: 614: 610: 609: 606: 605: 600: 595: 593: 589: 588: 585: 584: 580: 579: 576: 575: 567: 564: 563: 560: 559: 551: 548: 547: 544: 543: 534: 529: 527: 521: 519: 513: 452: 449: 445: 444: 441: 440: 436: 435: 432: 431: 427: 426: 423: 422: 417: 412: 410: 406: 405: 402: 401: 397: 396: 393: 392: 387: 384: 383: 380: 379: 370: 365: 363: 357: 355: 349: 325:kingdom') and 319:Regnum Animale 266: 263: 261: 258: 166: 163: 149:(for plants), 71:taxonomic rank 27:Taxonomic rank 26: 9: 6: 4: 3: 2: 9432: 9421: 9418: 9416: 9413: 9412: 9410: 9391: 9390: 9382: 9380: 9379: 9371: 9369: 9366: 9363: 9359: 9357: 9354: 9353: 9352: 9348: 9347: 9339: 9337: 9336: 9328: 9326: 9323: 9320: 9316: 9315: 9314: 9310: 9307: 9305: 9302: 9299: 9295: 9293: 9290: 9289: 9288: 9284: 9281: 9279: 9276: 9273: 9269: 9267: 9264: 9262: 9261: 9253: 9252: 9251: 9247: 9244: 9242: 9239: 9237: 9234: 9231: 9227: 9225: 9222: 9220: 9217: 9216: 9215: 9211: 9208: 9206: 9203: 9201: 9200: 9192: 9191: 9190: 9186: 9183: 9181: 9178: 9176: 9173: 9171: 9168: 9165: 9161: 9159: 9156: 9155: 9154: 9150: 9147: 9145: 9142: 9140: 9136: 9133: 9130: 9129: 9118: 9114: 9112: 9111: 9102: 9099: 9098: 9097: 9093: 9090: 9088: 9085: 9082: 9078: 9076: 9075:/Superkingdom 9070: 9068: 9067: 9059: 9057: 9056: 9048: 9047: 9046: 9045: 9040: 9036: 9029: 9024: 9022: 9017: 9015: 9010: 9009: 9006: 9000: 8997: 8995: 8991: 8988: 8985: 8982: 8979: 8978: 8969: 8965: 8962: 8959: 8956: 8952: 8951: 8938: 8934: 8930: 8926: 8922: 8918: 8914: 8910: 8906: 8902: 8901: 8892: 8884: 8880: 8875: 8870: 8866: 8862: 8861: 8856: 8849: 8842:(4): 218–237. 8841: 8837: 8836: 8828: 8821: 8813: 8809: 8805: 8801: 8797: 8793: 8789: 8785: 8784: 8776: 8774: 8757: 8753: 8747: 8740:(4): 218–237. 8739: 8735: 8728: 8721: 8713: 8709: 8704: 8699: 8694: 8689: 8685: 8681: 8677: 8673: 8672: 8667: 8660: 8652: 8648: 8643: 8638: 8633: 8628: 8624: 8620: 8616: 8612: 8611: 8606: 8599: 8591: 8585: 8577: 8573: 8569: 8567:0-412-36440-9 8563: 8559: 8552: 8544: 8540: 8536: 8532: 8528: 8524: 8521:(8): 1641–8. 8520: 8516: 8512: 8505: 8497: 8493: 8488: 8483: 8478: 8473: 8469: 8465: 8461: 8457: 8456: 8451: 8447: 8440: 8432: 8428: 8423: 8418: 8414: 8410: 8406: 8402: 8401: 8396: 8389: 8381: 8377: 8372: 8367: 8363: 8359: 8358: 8353: 8346: 8338: 8334: 8329: 8324: 8320: 8316: 8312: 8308: 8307: 8302: 8295: 8287: 8283: 8278: 8273: 8269: 8265: 8261: 8257: 8256: 8251: 8244: 8242: 8233: 8229: 8224: 8219: 8214: 8209: 8205: 8201: 8197: 8193: 8192: 8187: 8180: 8178: 8170: 8165: 8157: 8153: 8148: 8143: 8138: 8133: 8129: 8125: 8124: 8123:PLOS Genetics 8119: 8112: 8104: 8100: 8095: 8090: 8086: 8082: 8081: 8076: 8069: 8061: 8057: 8053: 8049: 8044: 8039: 8035: 8031: 8030: 8022: 8015: 8013: 8005:(4): 218–237. 8004: 8000: 7999: 7991: 7984: 7976: 7972: 7968: 7964: 7960: 7956: 7949: 7942: 7934: 7930: 7926: 7922: 7917: 7912: 7908: 7904: 7903: 7898: 7891: 7889: 7887: 7878: 7874: 7870: 7866: 7861: 7856: 7853:(4): R165–7. 7852: 7848: 7847: 7842: 7835: 7827: 7823: 7818: 7813: 7808: 7803: 7799: 7795: 7791: 7787: 7783: 7776: 7768: 7764: 7759: 7754: 7750: 7746: 7742: 7738: 7734: 7731:(June 2010). 7730: 7724: 7710: 7706: 7701: 7696: 7692: 7688: 7684: 7680: 7673: 7669: 7663: 7655: 7651: 7647: 7643: 7639: 7635: 7631: 7627: 7623: 7619: 7613: 7605: 7601: 7596: 7591: 7587: 7583: 7579: 7575: 7571: 7564: 7556: 7552: 7548: 7546:9780470514344 7542: 7538: 7534: 7530: 7526: 7522: 7516: 7508: 7504: 7500: 7496: 7492: 7488: 7484: 7478: 7470: 7466: 7461: 7456: 7451: 7446: 7442: 7438: 7434: 7430: 7426: 7422: 7416: 7408: 7404: 7399: 7394: 7389: 7384: 7380: 7376: 7372: 7368: 7364: 7360: 7354: 7346: 7342: 7338: 7334: 7330: 7326: 7322: 7318: 7314: 7310: 7306: 7300: 7292: 7288: 7284: 7280: 7276: 7272: 7268: 7264: 7260: 7254: 7246: 7242: 7238: 7237: 7232: 7226: 7218: 7214: 7210: 7206: 7202: 7198: 7194: 7188: 7180: 7176: 7170: 7162: 7158: 7156: 7150: 7144: 7136: 7132: 7126: 7118: 7114: 7108: 7100: 7096: 7091: 7086: 7082: 7078: 7074: 7070: 7069: 7064: 7057: 7055: 7039: 7035: 7030: 7025: 7021: 7017: 7013: 7009: 7008: 7000: 6993: 6985: 6981: 6976: 6971: 6967: 6963: 6959: 6955: 6951: 6947: 6941: 6939: 6919: 6915: 6911: 6907: 6903: 6899: 6895: 6892:(7147): 913. 6891: 6887: 6886: 6878: 6871: 6863: 6859: 6855: 6851: 6847: 6843: 6839: 6835: 6831: 6827: 6826: 6818: 6810: 6806: 6801: 6796: 6791: 6786: 6782: 6778: 6774: 6770: 6769: 6764: 6757: 6755: 6753: 6738:on 2021-05-10 6737: 6733: 6729: 6725: 6719: 6711: 6707: 6703: 6699: 6695: 6691: 6687: 6683: 6679: 6675: 6674: 6666: 6664: 6655: 6649: 6645: 6640: 6639: 6630: 6616: 6612: 6608: 6604: 6600: 6596: 6595: 6589: 6581: 6579: 6570: 6566: 6564:9780080920146 6560: 6556: 6552: 6551: 6543: 6535: 6531: 6527: 6523: 6518: 6513: 6509: 6505: 6501: 6497: 6496: 6488: 6486: 6484: 6475: 6471: 6467: 6463: 6459: 6455: 6451: 6447: 6446: 6438: 6430: 6426: 6421: 6416: 6412: 6408: 6404: 6400: 6399: 6394: 6390: 6384: 6382: 6380: 6371: 6367: 6364:(4): 207–16. 6363: 6359: 6358: 6350: 6348: 6346: 6344: 6342: 6340: 6331: 6327: 6323: 6316: 6308: 6304: 6300: 6296: 6292: 6288: 6285:(3): 203–66. 6284: 6280: 6279: 6273: 6265: 6263: 6261: 6259: 6250: 6246: 6241: 6236: 6231: 6226: 6222: 6218: 6214: 6210: 6206: 6199: 6197: 6195: 6186: 6185:"article 3.1" 6172: 6168: 6167: 6158: 6150: 6143: 6135: 6131: 6125: 6118: 6116: 6112: 6108: 6104: 6091:on 2021-11-11 6087: 6080: 6074: 6070: 6056: 6052: 6046: 6042: 6032: 6029: 6027: 6024: 6022: 6021:Phylogenetics 6019: 6017: 6014: 6013: 6009: 6003: 5998: 5991: 5989: 5984: 5982: 5977: 5973: 5969: 5965: 5960: 5958: 5954: 5950: 5936: 5932: 5929: 5925: 5922: 5918: 5915: 5910: 5906: 5902: 5897: 5892: 5887: 5883: 5880: 5875: 5870: 5865: 5860: 5854: 5849: 5844: 5843:cellular life 5840: 5836: 5832: 5829:Prionobiota ( 5827: 5817: 5813: 5810:Virusobiota ( 5809: 5803: 5799: 5795: 5790: 5786: 5782: 5779: 5776: 5774:Six kingdoms 5773: 5771: 5767: 5764: 5762: 5761:Three domains 5759: 5757: 5754: 5751: 5748: 5747: 5744: 5742: 5738: 5734: 5730: 5726: 5722: 5716: 5706: 5703: 5699: 5695: 5690: 5682: 5678: 5676: 5672: 5668: 5658: 5657: 5649: 5648: 5640: 5639: 5631: 5630: 5627: 5626: 5622: 5618: 5614: 5607: 5603: 5599: 5598: 5595: 5594: 5591: 5590: 5586: 5582: 5578: 5574: 5570: 5566: 5565:Stramenopiles 5560:   5559: 5555: 5554: 5551: 5550: 5547: 5545: 5541: 5537: 5531:   5530: 5526: 5525: 5522: 5521: 5518: 5516: 5512: 5508: 5502:   5501: 5497: 5496: 5493: 5492: 5489: 5488: 5484: 5477:   5476: 5472: 5471: 5468: 5467: 5464: 5462: 5455:   5454: 5451: 5450: 5444:   5443: 5439: 5438: 5432: 5431: 5428: 5427: 5419: 5418: 5410: 5409: 5406: 5405: 5402: 5396: 5395: 5389:   5388: 5384: 5383: 5377: 5376: 5373: 5372: 5364: 5363: 5355: 5354: 5351: 5350: 5347: 5341: 5340: 5334:   5333: 5329: 5328: 5322: 5321: 5315:   5314: 5310: 5309: 5305: 5301: 5294: 5290: 5284: 5279: 5272: 5263: 5262: 5254: 5253: 5250: 5248: 5244: 5237: 5236: 5233: 5232: 5229: 5227: 5223: 5216: 5215: 5212: 5211: 5208: 5206: 5202: 5195: 5194: 5188:   5187: 5183: 5182: 5178: 5176: 5172: 5167: 5163: 5161: 5153: 5149: 5145: 5105: 5101: 5091: 5089: 5085: 5071: 5068: 5064: 5060: 5056: 5055: 5054: 5047: 5044: 5042: 5039: 5037: 5034: 5032: 5029: 5027: 5024: 5022: 5019: 5017: 5014: 5012: 5009: 5008: 5005: 5002: 5000: 4997: 4995: 4992: 4990: 4987: 4985: 4982: 4981: 4978: 4975: 4973: 4970: 4968: 4965: 4963: 4960: 4958: 4955: 4953: 4948: 4943: 4939: 4936: 4931: 4926: 4922: 4919: 4915: 4912: 4907: 4902: 4897: 4892: 4887: 4882: 4877: 4873: 4870: 4867: 4865: 4862: 4860: 4857: 4855: 4852: 4851: 4848: 4845: 4843: 4838: 4835: 4833: 4830: 4828: 4825: 4823: 4818: 4813: 4808: 4801: 4798: 4795: 4793: 4790: 4788: 4785: 4782: 4780: 4777: 4775: 4772: 4770: 4767: 4764: 4761: 4758: 4757: 4754: 4751: 4749: 4746: 4743: 4741: 4738: 4736: 4733: 4731: 4728: 4726: 4723: 4721: 4718: 4715: 4712: 4711: 4705: 4701: 4696: 4692: 4689: 4685: 4681: 4676: 4673: 4670: 4666: 4663: 4659: 4656: 4652: 4649: 4645: 4642: 4638: 4635: 4634: 4621: 4620: 4612: 4611: 4603: 4602: 4594: 4593: 4590: 4589: 4582: 4581: 4578: 4577: 4574: 4573: 4566: 4565: 4562: 4561: 4558: 4557: 4553: 4549: 4545: 4541: 4534: 4533: 4530: 4529: 4526: 4525: 4521: 4517: 4513: 4509: 4505: 4501: 4497: 4490: 4489: 4486: 4485: 4482: 4481: 4477: 4473: 4469: 4462: 4461: 4455: 4451: 4450: 4444: 4443: 4440: 4439: 4431: 4430: 4422: 4421: 4418: 4417: 4410: 4409: 4406: 4405: 4402: 4401: 4394: 4393: 4387: 4383: 4382: 4376: 4375: 4368: 4367: 4363: 4361: 4357: 4353: 4349: 4345: 4341: 4326: 4324: 4319: 4318: 4315:Infrakingdom 4311: 4306: 4301: 4296: 4291: 4286: 4281: 4276: 4271: 4266: 4265:Labyrinthista 4261: 4256: 4251: 4246: 4245: 4242:Infrakingdom 4238: 4237: 4230: 4225: 4224: 4217: 4216: 4209: 4207: 4205: 4200: 4195: 4190: 4185: 4180: 4175: 4170: 4165: 4160: 4155: 4150: 4145: 4140: 4135: 4130: 4129: 4126:Infrakingdom 4122: 4117: 4112: 4107: 4102: 4101:Prasinophytae 4097: 4096:Chlorophytina 4092: 4087: 4086: 4083:Infrakingdom 4079: 4078: 4077:Viridiplantae 4071: 4066: 4061: 4056: 4055: 4052:Infrakingdom 4048: 4043: 4042: 4039:Infrakingdom 4035: 4034: 4027: 4026: 4019: 4017: 4015: 4010: 4009: 4002: 4001:Gnathostomata 3997: 3992: 3984: 3979: 3974: 3969: 3964: 3959: 3954: 3949: 3944: 3939: 3938: 3935:Infrakingdom 3931: 3926: 3921: 3916: 3911: 3906: 3901: 3896: 3895:Echinodermata 3891: 3890:Enteropneusta 3886: 3885:Pterobranchia 3881: 3876: 3875: 3872:Infrakingdom 3868: 3867: 3866:Deuterostomia 3860: 3855: 3850: 3845: 3844:Rhabditophora 3840: 3835: 3830: 3825: 3820: 3815: 3810: 3805: 3804:Acanthognatha 3800: 3799: 3796:Infrakingdom 3792: 3787: 3782: 3777: 3772: 3767: 3762: 3757: 3752: 3747: 3742: 3737: 3732: 3727: 3722: 3714: 3709: 3704: 3699: 3694: 3689: 3684: 3683: 3680:Infrakingdom 3676: 3671: 3670: 3667:Infrakingdom 3663: 3658: 3653: 3648: 3643: 3638: 3633: 3628: 3623: 3618: 3613: 3608: 3603: 3598: 3593: 3588: 3583: 3578: 3573: 3568: 3563: 3558: 3553: 3548: 3543: 3538: 3533: 3528: 3523: 3522: 3519:Infrakingdom 3515: 3514: 3507: 3506: 3499: 3494: 3493: 3486: 3481: 3480: 3477:Infrakingdom 3473: 3468: 3463: 3458: 3457: 3454:Infrakingdom 3447: 3446:Archaeocyatha 3442: 3441:Calcispongiae 3437: 3436:Hyalospongiae 3432: 3427: 3426: 3423:Infrakingdom 3419: 3418: 3411: 3410: 3403: 3401: 3399: 3394: 3389: 3384: 3383:Auromycetidae 3379: 3374: 3369: 3364: 3363:Hymenomycetia 3359: 3354: 3349: 3348:Orthomycotina 3344: 3339: 3334: 3329: 3328:Septomycotina 3324: 3323: 3322:Basidiomycota 3316: 3315:Plectomycetes 3311: 3306: 3301: 3300:Loculomycetes 3296: 3291: 3286: 3285:Pyrenomycetes 3281: 3280:Pezomycetidae 3276: 3275:Lecomycetidae 3271: 3266: 3261: 3256: 3251: 3250:Dipomycetidae 3246: 3241: 3236: 3231: 3226: 3225: 3218: 3217: 3210: 3205: 3204:Pleistophorea 3200: 3195: 3190: 3189: 3188:Microsporidia 3182: 3181:Pyxomycetalia 3177: 3172: 3171:Pedomycetidae 3167: 3166:Entomycetidae 3162: 3157: 3156:Meromycetidae 3152: 3147: 3142: 3137: 3132: 3127: 3126:Eozygomycetia 3122: 3117: 3112: 3107: 3102: 3101:Enteromycetes 3097: 3092: 3087: 3082: 3077: 3076: 3069: 3068: 3061: 3060: 3053: 3051: 3048: 3043: 3038: 3033: 3028: 3027: 3024:Infrakingdom 3020: 3019:Filocorticata 3015: 3010: 3005: 3000: 2999:Heterokaryota 2995: 2990: 2985: 2980: 2975: 2970: 2969:Protalveolata 2965: 2960: 2955: 2954: 2951:Infrakingdom 2947: 2942: 2937: 2932: 2931:Pseudociliata 2927: 2922: 2917: 2916: 2915:Discicristata 2913:Infrakingdom 2909: 2904: 2899: 2894: 2889: 2884: 2879: 2874: 2869: 2864: 2859: 2854: 2849: 2844: 2839: 2834: 2829: 2828: 2825:Infrakingdom 2821: 2820: 2813: 2808: 2803: 2798: 2793: 2788: 2783: 2782: 2775: 2774: 2767: 2765: 2762: 2761:Sulfobacteria 2757: 2752: 2747: 2742: 2737: 2732: 2727: 2722: 2717: 2712: 2707: 2702: 2697: 2692: 2687: 2682: 2677: 2676:Alphabacteria 2672: 2671:Rhodobacteria 2667: 2662: 2661:Phycobacteria 2657: 2656:Gloeobacteria 2652: 2651:Cyanobacteria 2647: 2646:Fibrobacteria 2642: 2637: 2632: 2627: 2626:Flavobacteria 2622: 2617: 2612: 2607: 2606:Glycobacteria 2602: 2597: 2592: 2587: 2582: 2581:Deinobacteria 2577: 2572: 2567: 2566:Heliobacteria 2562: 2557: 2552: 2547: 2546: 2539: 2538: 2525: 2524: 2516: 2515: 2507: 2506: 2498: 2497: 2494: 2493: 2486: 2485: 2482: 2481: 2478: 2477: 2470: 2469: 2466: 2465: 2462: 2461: 2457: 2453: 2449: 2445: 2438: 2437: 2434: 2433: 2430: 2429: 2425: 2421: 2417: 2413: 2409: 2405: 2401: 2394: 2393: 2390: 2389: 2386: 2385: 2381: 2377: 2373: 2366: 2365: 2359: 2355: 2354: 2348: 2347: 2344: 2343: 2335: 2334: 2326: 2325: 2322: 2320: 2316: 2309: 2308: 2302: 2298: 2297: 2291: 2290: 2283: 2282: 2278: 2276: 2271: 2269: 2265: 2261: 2257: 2253: 2252:Microsporidia 2249: 2245: 2241: 2237: 2231: 2229: 2225: 2221: 2216: 2214: 2210: 2206: 2202: 2198: 2194: 2183: 2181: 2171: 2170: 2162: 2161: 2153: 2152: 2144: 2143: 2140: 2139: 2132: 2131: 2128: 2127: 2124: 2123: 2116: 2115: 2112: 2111: 2108: 2107: 2100: 2099: 2096: 2095: 2092: 2091: 2084: 2083: 2080: 2079: 2076: 2075: 2068: 2067: 2060: 2056: 2055: 2049: 2048: 2045: 2044: 2036: 2035: 2027: 2026: 2023: 2021: 2014: 2013: 2006: 2002: 2001: 1995: 1994: 1991: 1990: 1982: 1981: 1973: 1972: 1969: 1968: 1961: 1960: 1957: 1956: 1953: 1952: 1945: 1944: 1938: 1934: 1933: 1927: 1926: 1919: 1918: 1914: 1912: 1908: 1904: 1900: 1896: 1892: 1888: 1884: 1880: 1876: 1872: 1871:endosymbiosis 1867: 1865: 1864:chlorophyll c 1861: 1857: 1853: 1849: 1845: 1841: 1837: 1833: 1829: 1825: 1821: 1817: 1803: 1802: 1794: 1793: 1785: 1784: 1776: 1775: 1772: 1771: 1764: 1763: 1760: 1759: 1756: 1755: 1748: 1747: 1744: 1743: 1740: 1739: 1732: 1731: 1728: 1727: 1724: 1722: 1715: 1714: 1708: 1704: 1703: 1697: 1696: 1693: 1692: 1684: 1683: 1675: 1674: 1671: 1669: 1662: 1661: 1658: 1657: 1654: 1652: 1645: 1644: 1638: 1634: 1633: 1627: 1626: 1619: 1618: 1614: 1612: 1608: 1604: 1600: 1596: 1595:ribosomal RNA 1592: 1568: 1564: 1562: 1558: 1556: 1555:Echinodermata 1552: 1550: 1546: 1544: 1540: 1539: 1538: 1534: 1530: 1526: 1524: 1520: 1518: 1514: 1512: 1511:Sipunculoidea 1508: 1506: 1502: 1500: 1496: 1494: 1490: 1488: 1484: 1480: 1479: 1478: 1474: 1473: 1472: 1468: 1464: 1460: 1456: 1454: 1453:Aschelminthes 1450: 1448: 1444: 1443: 1442: 1438: 1433: 1429: 1427: 1423: 1422: 1421: 1417: 1416: 1415: 1411: 1407: 1403: 1401: 1397: 1396: 1395: 1391: 1390: 1389: 1385: 1378: 1377:Archaeocyatha 1374: 1372: 1368: 1367: 1365: 1361: 1357: 1356: 1354: 1353: 1352: 1351: 1350: 1344: 1336: 1335:Basidiomycota 1332: 1330: 1326: 1324: 1320: 1319: 1318: 1314: 1310: 1306: 1305: 1303: 1302: 1301: 1297: 1293: 1289: 1288: 1286: 1282: 1278: 1276: 1275:Acrasiomycota 1272: 1270: 1266: 1265: 1263: 1262: 1261: 1260: 1259: 1253: 1245: 1241: 1239: 1235: 1234: 1233: 1229: 1225: 1221: 1219: 1215: 1214: 1213: 1209: 1208: 1207: 1206:Euchlorophyta 1203: 1199: 1195: 1194: 1193: 1189: 1185: 1181: 1180: 1179: 1175: 1174: 1173: 1172: 1171: 1165: 1161: 1157: 1155: 1151: 1149: 1145: 1143: 1142:Cnidosporidia 1139: 1137: 1133: 1131: 1127: 1125: 1121: 1119: 1115: 1113: 1109: 1107: 1103: 1102: 1101: 1100: 1099: 1093: 1087: 1083: 1081: 1077: 1074: 1073: 1071: 1066: 1064: 1060: 1059: 1057: 1056: 1055: 1054: 1053: 1047: 1046: 1033: 1032: 1024: 1023: 1015: 1014: 1011: 1010: 1003: 1002: 999: 998: 995: 994: 987: 986: 983: 982: 979: 978: 971: 970: 967: 966: 963: 961: 954: 953: 947: 943: 942: 939: 938: 930: 929: 926: 925: 918: 917: 911: 907: 906: 899: 898: 894: 890: 888: 884: 883:Lynn Margulis 878: 876: 872: 868: 864: 860: 856: 851: 844:Five kingdoms 836: 835: 827: 826: 818: 817: 814: 813: 806: 805: 802: 801: 798: 797: 790: 789: 786: 785: 782: 780: 773: 772: 766: 762: 761: 758: 757: 749: 748: 745: 744: 737: 736: 730: 726: 725: 718: 717: 713: 711: 707: 703: 699: 695: 691: 690:Roger Stanier 686: 684: 683:Cyanobacteria 680: 676: 672: 668: 664: 659: 657: 653: 649: 645: 641: 634:Four kingdoms 626: 625: 617: 616: 608: 607: 604: 599: 598: 591: 590: 587: 586: 578: 577: 574: 573: 566: 565: 562: 561: 558: 557: 550: 549: 546: 545: 542: 540: 533: 532: 525: 524: 517: 516: 512: 510: 509: 504: 503:Ernst Haeckel 500: 497:proposed the 496: 491: 487: 484: 483:Royal Society 480: 472: 471: 465: 460: 455: 443: 442: 434: 433: 425: 424: 421: 416: 415: 408: 407: 404: 403: 395: 394: 391: 386: 385: 382: 381: 378: 376: 369: 368: 361: 360: 353: 352: 348: 346: 345: 340: 336: 332: 328: 324: 320: 316: 312: 308: 307:Carl Linnaeus 304: 303:, on plants. 302: 301: 295: 287 BC 282: 278: 277: 272: 257: 255: 254:Deuterostomia 251: 247: 243: 239: 235: 231: 227: 222: 220: 216: 212: 208: 204: 200: 196: 192: 188: 184: 180: 176: 172: 171:Carl Linnaeus 162: 160: 159: 154: 153: 148: 147: 142: 138: 134: 130: 126: 121: 119: 115: 111: 107: 103: 99: 95: 91: 87: 82: 80: 76: 73:, just below 72: 68: 64: 56: 52: 48: 32: 19: 9383: 9372: 9340: 9329: 9254: 9193: 9122: 9104: 9091: 9086: 9080: 9060: 9049: 8967: 8954: 8904: 8898: 8891: 8864: 8858: 8848: 8839: 8835:Protistology 8833: 8820: 8787: 8781: 8760:. Retrieved 8755: 8746: 8737: 8734:Protistology 8733: 8720: 8675: 8669: 8659: 8614: 8608: 8598: 8557: 8551: 8518: 8514: 8504: 8462:(7): e2621. 8459: 8453: 8439: 8404: 8398: 8388: 8361: 8355: 8345: 8310: 8304: 8294: 8259: 8253: 8195: 8189: 8164: 8130:(12): e220. 8127: 8121: 8111: 8084: 8078: 8068: 8033: 8027: 8002: 7998:Protistology 7996: 7983: 7958: 7954: 7941: 7906: 7900: 7850: 7844: 7834: 7789: 7785: 7775: 7740: 7736: 7723: 7712:. Retrieved 7682: 7678: 7662: 7629: 7625: 7612: 7577: 7573: 7563: 7528: 7524: 7515: 7490: 7486: 7477: 7432: 7428: 7415: 7370: 7366: 7353: 7312: 7308: 7299: 7266: 7262: 7253: 7235: 7225: 7200: 7196: 7187: 7178: 7169: 7160: 7154: 7143: 7134: 7125: 7116: 7113:Linnaeus, C. 7107: 7072: 7066: 7041:. Retrieved 7011: 7005: 6992: 6957: 6953: 6925:. Retrieved 6918:the original 6889: 6883: 6870: 6829: 6823: 6817: 6772: 6766: 6740:. Retrieved 6736:the original 6727: 6718: 6677: 6671: 6637: 6629: 6618:. Retrieved 6598: 6592: 6569:Google Books 6567:– via 6549: 6542: 6499: 6493: 6452:(1): 17–35. 6449: 6443: 6437: 6402: 6396: 6361: 6355: 6321: 6315: 6282: 6276: 6212: 6208: 6175:. Retrieved 6171:the original 6165: 6157: 6148: 6142: 6133: 6124: 6114: 6110: 6106: 6102: 6099: 6093:. Retrieved 6086:the original 6073: 6045: 5985: 5961: 5952: 5946: 5838: 5798:Aphanobionta 5797: 5793: 5769: 5718: 5701: 5691: 5687: 5675:Opisthokonta 5664: 5625:glaucophytes 5611: 5606:Primoplantae 5600:   5563: 5556:   5536:Foraminifera 5534: 5527:   5505: 5500:Opisthokonta 5498:   5487:slime moulds 5481:most lobose 5480: 5473:   5458: 5440:Domain  5399: 5385:Domain  5344: 5330:Domain  5311:   5302: 5286: 5278: 5269: 5240: 5219: 5198: 5184:   5168: 5164: 5157: 5081: 5061:rather than 5052: 4703: 4683: 4674: 4585: 4569: 4544:glaucophytes 4537: 4504:cryptophytes 4493: 4465: 4413: 4397: 4336: 4314: 4300:Bigyromonada 4241: 4233: 4232:   4220: 4219:   4212: 4210: 4204:Angiospermae 4199:Gymnospermae 4184:Sphenophytae 4164:Tracheophyta 4144:Anthocerotae 4125: 4082: 4074: 4073:   4051: 4038: 4030: 4029:   4022: 4020: 4005: 4004:   3934: 3920:Eleutherozoa 3880:Hemichordata 3871: 3863: 3795: 3791:Nematomorpha 3679: 3675:Chaetognatha 3669:Chaetognathi 3666: 3537:Gymnolaemata 3518: 3510: 3502: 3501:   3498:Myxosporidia 3489: 3488:   3476: 3456:Coelenterata 3453: 3422: 3414: 3413:   3406: 3404: 3333:Septomycetes 3319: 3265:Discomycetes 3221: 3213: 3212:   3185: 3136:Glomomycetes 3121:Zygomycotina 3111:Allomycotina 3072: 3064: 3063:   3056: 3054: 3023: 2994:Manubrispora 2950: 2912: 2908:Dictyostelia 2873:Foraminifera 2868:Monadofilosa 2824: 2816: 2815:   2807:Anaeromonada 2778: 2777:   2770: 2768: 2746:Euryarcheota 2741:Mendosicutes 2731:Togobacteria 2721:Endobacteria 2711:Posibacteria 2706:Posibacteria 2686:Thiobacteria 2641:Fusobacteria 2631:Eurybacteria 2591:Spirochaetae 2571:Hadobacteria 2556:Lipobacteria 2551:Negibacteria 2542: 2540: 2489: 2473: 2448:glaucophytes 2441: 2408:cryptophytes 2397: 2369: 2356:Empire  2312: 2299:Empire  2272: 2244:metakaryotes 2232: 2220:monophyletic 2217: 2213:Posibacteria 2205:Negibacteria 2189: 2177: 2135: 2119: 2103: 2087: 2071: 2017: 1964: 1948: 1868: 1840:Posibacteria 1832:Negibacteria 1814: 1767: 1751: 1735: 1718: 1705:Empire  1665: 1648: 1635:Empire  1588: 1585:Six kingdoms 1561:Hemichordata 1549:Chaetognatha 1346: 1345: 1255: 1254: 1244:Tracheophyta 1167: 1166: 1148:Zoomastigina 1106:Euglenophyta 1095: 1094: 1086:Spirochaetae 1080:Actinomycota 1049: 1048: 1006: 990: 974: 957: 944:Empire  921: 908:Empire  891: 879: 871:heterotrophs 847: 809: 793: 776: 763:Empire  740: 727:Empire  701: 698:superkingdom 697: 687: 678: 660: 637: 602: 569: 553: 536: 506: 498: 492: 488: 476: 473:with plants. 468: 454: 419: 389: 372: 342: 326: 318: 305: 298: 281:Theophrastus 274: 268: 245: 241: 237: 234:infrakingdom 233: 229: 225: 223: 175:nomenclature 168: 156: 150: 144: 143:. The terms 139:of a common 133:monophyletic 128: 122: 86:six kingdoms 83: 66: 60: 9283:Infrafamily 9266:Superfamily 9180:Subterclass 9149:Microphylum 9144:Infraphylum 9139:Subdivision 9101:Superphylum 8752:"ICTV Code" 8313:: 231–238. 8198:(8): e790. 8171:, p. 4 7487:Bio Systems 7359:Woese, C.R. 7305:Woese, C.R. 7175:Chatton, É. 7149:Chatton, É. 7131:Haeckel, E. 6728:www.ric.edu 6644:McGraw-Hill 6026:Systematics 5976:replication 5756:Two empires 5739:called the 5696:dubbed the 5617:green algae 5613:Land plants 5585:Cryptophyta 5569:Brown Algae 5401:prokaryotic 5346:prokaryotic 4942:Vegetabilia 4762:3 kingdoms 4759:2 kingdoms 4556:land plants 4552:green algae 4512:Brown Algae 4508:Heterokonta 4305:Pseudofungi 4280:Hypogyrista 4244:Heterokonta 4236:Chromobiota 4234:Subkingdom 4229:Cryptophyta 4221:Subkingdom 4174:Psilophytae 4116:Charophytae 4106:Tetraphytae 4091:Chlorophyta 4085:Chlorophyta 4075:Subkingdom 4047:Glaucophyta 4041:Glaucophyta 4031:Subkingdom 4006:Subkingdom 3874:Coelomopora 3834:Turbellaria 3776:Kinorhyncha 3746:Onychophora 3721:Mandibulata 3708:Chelicerata 3703:Pycnogonida 3657:Pogonophora 3602:Brachiopoda 3592:Cephalopoda 3577:Glossophora 3557:Cycliophora 3513:Protostomia 3503:Subkingdom 3490:Subkingdom 3415:Subkingdom 3368:Gelimycetes 3358:Ustomycetes 3245:Endomycetes 3214:Subkingdom 3199:Microsporea 3146:Zygomycetes 3131:Bolomycetes 3116:Allomycetes 3075:Archemycota 3065:Subkingdom 3014:Epiplasmata 2893:Archamoebae 2817:Subkingdom 2797:Axostylaria 2792:Eopharyngia 2779:Subkingdom 2701:Unibacteria 2601:Leptospirae 2460:land plants 2456:green algae 2416:Brown Algae 2412:Heterokonta 2317:— includes 2201:Unibacteria 2180:taxonomists 2059:Metakaryota 1887:Metakaryota 1653:(Bacteria) 1543:Pogonophora 1537:Enterocoela 1517:Echiuroidea 1493:Brachiopoda 1477:Schizocoela 1386:Subkingdom 1298:Subkingdom 1218:Chlorophyta 1204:Subkingdom 1190:Subkingdom 1176:Subkingdom 1112:Chrysophyta 887:Embryophyta 875:saprotrophs 654:). In 1937 648:prokaryotes 499:Protoctista 250:Protostomia 238:infraregnum 137:descendants 9409:Categories 9368:Subspecies 9309:Infratribe 9292:Supertribe 9241:Infraorder 9224:Superorder 9195:Division ( 9175:Infraclass 9158:Superclass 9124:Division ( 9087:Subkingdom 9062:Subrealm ( 8867:(8): 615. 8560:. London. 8407:: 99–113. 7737:Biol. Lett 7714:2010-04-29 7075:: 379–92. 6742:2020-07-25 6620:2020-07-28 6177:2011-02-20 6095:2022-03-04 6065:References 6055:rhizarians 6051:alveolates 6016:Cladistics 5972:metabolism 5970:that lack 5874:Eubacteria 5787:/ Vitae / 5733:eukaryotes 5581:Haptophyta 5540:Radiolaria 5461:flagellate 5293:eukaryotes 5205:Eubacteria 5171:Eubacteria 5160:Carl Woese 4837:Eubacteria 4827:Eubacteria 4812:Prokaryota 4797:7 kingdoms 4792:6 kingdoms 4787:8 kingdoms 4779:6 kingdoms 4774:5 kingdoms 4769:4 kingdoms 4520:Haptophyta 4386:Prokaryota 4350:, and the 4323:Haptophyta 4317:Haptophyta 4270:Ochrophyta 4179:Lycophytae 4128:Cormophyta 4121:Rudophytae 4060:Rhodophyta 4054:Rhodophyta 3991:Vertebrata 3905:Pelmatozoa 3849:Neodermata 3839:Mucorhabda 3751:Tardigrada 3693:Arthropoda 3647:Clitellata 3642:Pharyngata 3637:Operculata 3632:Polychaeta 3597:Brachiozoa 3552:Entoprocta 3425:Spongiaria 3240:Geomycetes 3224:Ascomycota 3194:Minisporea 3161:Zoomycetes 3047:Radiolaria 3026:Actinopoda 3004:Ciliophora 2984:Gregarinae 2946:Saccostoma 2941:Plicostoma 2936:Euglenozoa 2926:Tetramitia 2921:Percolozoa 2812:Parabasala 2787:Metamonada 2756:Eurytherma 2561:Eobacteria 2424:Haptophyta 2301:Prokaryota 2264:Metamonada 2236:archezoans 2197:Prokaryota 1951:Eubacteria 1937:Prokaryota 1828:Eubacteria 1820:Eubacteria 1670:(Archaea) 1651:Eubacteria 1637:Prokaryota 1591:Carl Woese 1529:Arthropoda 1487:Ectoprocta 1459:Entoprocta 1420:Acoelomata 1406:Ctenophora 1329:Ascomycota 1323:Zygomycota 1269:Myxomycota 1224:Charophyta 1198:Phaeophyta 1184:Rhodophyta 1160:Ciliophora 1118:Pyrrophyta 1063:Cyanophyta 910:Prokaryota 867:autotrophs 729:Prokaryota 704:—with the 652:eukaryotes 640:microscopy 377:(animals) 288: 371 226:subkingdom 125:cladistics 18:Subkingdom 9374:Variety ( 9331:Section ( 9278:Subfamily 9256:Section ( 9246:Parvorder 9219:Magnorder 9185:Parvclass 9135:Subphylum 8584:cite book 7933:207051421 7421:Woese, C. 6512:CiteSeerX 6161:See e.g. 5988:Mimivirus 5968:parasites 5621:red algae 5589:Alveolata 5546:protozoa 5483:amoeboids 5475:Amoebozoa 5463:protozoa 5442:Eukaryota 5404:Archaeans 5296:'kingdoms 5247:Eukaryota 5146:based on 5084:cladistic 4935:Chromista 4930:Chromista 4925:Chromista 4876:Eukaryota 4748:2 empires 4740:3 domains 4735:2 empires 4730:2 empires 4725:2 empires 4720:2 empires 4702:Ruggiero 4665:Whittaker 4500:Alveolata 4496:Chromista 4476:Choanozoa 4472:Amoebozoa 4454:Eukaryota 4310:Opalinata 4290:Diatomeae 4285:Chrysista 4255:Sagenista 4250:Sagenista 4223:Cryptista 4215:Chromista 4154:Sphagneae 4139:Hepaticae 4134:Bryophyta 4033:Biliphyta 3983:Conodonta 3973:Acraniata 3943:Urochorda 3937:Chordonia 3930:Echinozoa 3925:Asterozoa 3910:Blastozoa 3900:Homalozoa 3854:Trematoda 3824:Monokonta 3781:Nematoida 3771:Priapozoa 3731:Myriapoda 3726:Crustacea 3688:Haemopoda 3682:Ecdysozoa 3662:Nemertina 3617:Sipuncula 3612:Sipuncula 3607:Phoronida 3587:Spiculata 3582:Univalvia 3562:Conchozoa 3547:Kamptozoa 3542:Lophopoda 3505:Bilateria 3472:Medusozoa 3216:Neomycota 3042:Spasmaria 2953:Alveolata 2898:Mycetozoa 2878:Amoebozoa 2858:Phytomyxa 2848:Choanozoa 2833:Neomonada 2802:Trichozoa 2404:Alveolata 2400:Chromista 2380:Choanozoa 2376:Amoebozoa 2358:Eukaryota 2275:paraphyly 2224:paraphyla 2090:Chromista 1907:Chromista 1850:from the 1848:Chromista 1707:Eukaryota 1611:Chromista 1589:In 1977, 1535:Subgrade 1499:Phoronida 1475:Subgrade 1471:Coelomata 1463:Kamptozoa 1414:Bilateria 1388:Eumetazoa 1238:Bryophyta 1232:Metaphyta 1154:Sarcodina 946:Eukaryota 863:nutrition 765:Eukaryota 661:In 1938, 495:John Hogg 477:In 1674, 271:Aristotle 230:subregnum 9342:Series ( 9325:Subgenus 9304:Subtribe 9236:Suborder 9170:Subclass 8990:Archived 8937:84298461 8929:15486256 8883:19561628 8804:19270719 8762:26 April 8651:19091952 8576:27895507 8543:38178747 8535:26305806 8496:18612431 8455:PLOS ONE 8431:20333181 8380:17488740 8337:20333193 8286:18522922 8232:17726520 8191:PLOS ONE 8156:17194223 8103:15653923 8052:16248873 7925:15341755 7877:13172971 7869:19243692 7826:25923521 7786:PLOS ONE 7767:20031978 7709:15306349 7670:(2004). 7620:(1998). 7345:27788891 7233:(1956). 7217:84634277 7177:(1937). 7151:(1925). 7133:(1866). 7115:(1735). 7099:20624742 7043:29 April 7038:15306349 6984:20031978 6948:(2009). 6927:15 March 6906:17581566 6809:25923521 6768:PLOS ONE 6710:27788891 6615:30163328 6474:29859498 6466:13916221 6429:15944457 6391:(2005). 6389:Sapp, J. 6370:10943416 6053:and the 6031:Taxonomy 5994:See also 5935:Animalia 5914:Protista 5879:Bacteria 5864:Bacteria 5859:Bacteria 5770:kingdoms 5608:)  5544:amoeboid 5529:Rhizaria 5459:various 5453:Excavata 5349:Bacteria 5332:Bacteria 5291:for all 5289:grab-bag 5201:Bacteria 5069:systems. 5046:Animalia 5041:Animalia 5036:Animalia 5031:Animalia 5026:Animalia 5021:Animalia 5016:Animalia 5011:Animalia 4918:Protozoa 4911:Protozoa 4906:Protozoa 4901:Archezoa 4891:Protista 4886:Protista 4881:Protista 4847:Bacteria 4842:Bacteria 4832:Bacteria 4807:Protista 4658:Copeland 4637:Linnaeus 4588:Animalia 4586:Kingdom 4570:Kingdom 4538:Kingdom 4524:Rhizaria 4494:Kingdom 4480:Excavata 4468:Protozoa 4466:Kingdom 4414:Kingdom 4400:Bacteria 4398:Kingdom 4340:Bacteria 4275:Phaeista 4260:Bicoecia 4213:Kingdom 4023:Kingdom 3968:Chordata 3953:Ascidiae 3948:Tunicata 3915:Crinozoa 3814:Rotifera 3809:Trochata 3798:Platyzoa 3786:Nematoda 3741:Lobopoda 3627:Annelida 3622:Vermizoa 3572:Bivalvia 3567:Mollusca 3521:Lophozoa 3485:Placozoa 3479:Placozoa 3467:Anthozoa 3462:Cnidaria 3431:Porifera 3409:Animalia 3407:Kingdom 3209:Disporea 3067:Eomycota 3057:Kingdom 3037:Radiozoa 3032:Heliozoa 2979:Sporozoa 2853:Cercozoa 2838:Apusozoa 2781:Archezoa 2773:Protozoa 2771:Kingdom 2545:Bacteria 2543:Kingdom 2492:Animalia 2490:Kingdom 2474:Kingdom 2442:Kingdom 2428:Rhizaria 2398:Kingdom 2384:Excavata 2372:Protozoa 2370:Kingdom 2315:Bacteria 2313:Kingdom 2268:Excavata 2260:Protozoa 2248:Archezoa 2193:Bacteria 2138:Animalia 2136:Kingdom 2120:Kingdom 2104:Kingdom 2088:Kingdom 2074:Protozoa 2072:Kingdom 2020:Archezoa 2018:Kingdom 2005:Archezoa 1965:Kingdom 1949:Kingdom 1895:Protozoa 1891:Animalia 1883:Archezoa 1770:Animalia 1768:Kingdom 1752:Kingdom 1736:Kingdom 1721:Protista 1719:Kingdom 1666:Kingdom 1649:Kingdom 1607:Protozoa 1567:Chordata 1523:Annelida 1505:Mollusca 1432:Nemertea 1400:Cnidaria 1371:Porifera 1349:Animalia 1347:Kingdom 1300:Eumycota 1292:Oomycota 1256:Kingdom 1168:Kingdom 1136:Sporozoa 1098:Protista 1096:Kingdom 1050:Kingdom 1009:Animalia 1007:Kingdom 991:Kingdom 975:Kingdom 960:Protista 958:Kingdom 922:Kingdom 812:Animalia 810:Kingdom 794:Kingdom 779:Protista 777:Kingdom 741:Kingdom 671:Bacteria 572:Animalia 570:Kingdom 554:Kingdom 539:Protista 537:Kingdom 518:  508:Protista 493:In 1860 354:  335:minerals 244:(Latin: 141:ancestor 110:Bacteria 102:Protista 90:Animalia 9362:Species 9081:Kingdom 9051:Realm ( 8909:Bibcode 8900:Science 8812:3907750 8712:6587394 8680:Bibcode 8642:2629348 8619:Bibcode 8487:2440802 8464:Bibcode 8422:2817406 8328:2817417 8277:2610160 8223:1949142 8200:Bibcode 8147:1713255 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Index

Subkingdom

biological classification
taxonomic ranks
domain
biology
taxonomic rank
domain
phyla
six kingdoms
Animalia
Plantae
Fungi
Protista
Archaea
Bacteria
five kingdoms only
Monera
cladistics
monophyletic
descendants
ancestor
flora
fauna
funga
Carl Linnaeus
nomenclature
class
order
genus

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