2961:
5093:
2954:
1458:
2283:
2269:
4545:
3749:
4531:
2262:
4538:
1464:
2968:
150:
2276:
5106:
3767:
121:
3755:
374:
3774:
5100:
4525:
3762:
570:
this ridge ran obliquely at an angle of about 35° to the lower edge of the shinbone, covering a vertical distance of about a quarter to a third of the lower shinbone height. At its lower end, this ridge slightly curved upwards, being at this point close to the outer edge of the shinbone, at about a
561:
Apart from the autapomorphy, a unique combination of, in themselves not unique, traits was also demonstrated. The bottom surface of the shinbone was, in its transverse width, about 1.5 times as wide as the longitudinal distance (measured from front to rear). In dinosaurs in general, the lower front
653:
hails from the La Quinta
Formation of northern South America, in what is now Colombia and Venezuela, and was found in the Venezuelan part of the formation. The La Quinta environment is interpreted as a tropical lowland forest. Its location and time suggests eusauropods reached a broad distribution
637:
also contributed to a greater knowledge of evolution by confirming that the equatorial zone of the supercontinent Pangea played an important role in the development of early dinosaurs, as already shown by the discovery of
856:
558:, has a rear edge making a sharp angle with the outer edge. This way a uniquely sharp point is formed which, uniquely also, extends further to behind than the inner projection at the opposite side.
539:
was a small bipedal predator. The shinbone has a length of about 25 cm (9.8 in); from this, the total body length of the animal has been deduced at just over 1.5 m (4.9 ft).
467:, a precise minimum estimate is not known; the actual age could be considerably younger. Both fossils are from the same location, but assumed to represent two individuals. One of these was the
550:. It pertains to the profile, seen from above, of the upper surface of the shinbone. Such surfaces generally have two projections, corners jutting out to behind, left and right. In
412:. However, included in the discoveries were some theropod teeth, indicating a predator must have been present. In 2013, this was affirmed by the discovery of some theropod bones.
774:
5244:
427:
857:"A sauropod from the Lower Jurassic La Quinta Formation (Dept. Cesar, Colombia) and the initial diversification of eusauropods at low latitudes"
399:
5185:
483:
of such a size was present in the La Quinta
Formation. It is specimen IVIC-P-2868, consisting of the damaged upper half of a left
603:, the subgroup encompassing all but the earliest theropods. It thus was placed low in the evolutionary tree of the neotheropods.
5259:
855:
Rincón, Aldo F.; Raad Pájaro, Daniel A.; Jiménez
Velandia, Harold F.; Ezcurra, MartĂn D.; Wilson Mantilla, Jeffrey A. (2022).
830:
2960:
2953:
941:
5092:
571:
fifth of its transverse width. The shinbone extends to below in two bumps, left and right. When seen from the front, with
920:
807:
3748:
2282:
708:
Langer, Max C.; Rincón, Ascanio D.; Ramezani, Jahandar; Solórzano, Andrés; Rauhut, Oliver W.M. (8 October 2014).
2268:
4530:
5254:
4971:
2261:
1457:
3766:
1463:
4544:
495:
The generic name was derived from that of the state of Táchira, where it was discovered, combined with the
2967:
5249:
2275:
595:) by computing an evolutionary tree assuming the fewest evolutionary changes. This analysis showed that
149:
5264:
4537:
3773:
5105:
623:
especially important, because before 2014, there were no unequivocal stem-averostrans known at all.
354:
dinosaurs, with a deduced total body length of just over 1.5 m (4.9 ft). They were likely
1008:
5099:
3754:
913:
521:
506:
380:
5216:
5125:
5211:
5203:
5163:
3479:
3391:
658:
and the
Toarcian faunal turnover. Other dinosaurs from this formation include the eusauropod
274:
5172:
5057:
3168:
3136:
868:
729:
4524:
8:
3492:
3279:
434:
325:
3761:
872:
733:
575:
a line drawn between the bumps made an angle of 80° with the vertical axis of the bone.
5239:
4964:
3468:
3328:
3184:
3124:
2559:
2469:
2352:
1216:
906:
884:
750:
709:
517:
513:
464:
423:
355:
144:
619:
of the
Averostra, it was described as a "stem-averostran". This made the discovery of
542:
The authors established a number of distinguishing traits. One of these is a possible
5198:
4846:
4763:
4737:
4576:
4563:
4441:
3982:
3934:
2766:
2116:
1195:
888:
755:
714:-Averostra) from the earliest Jurassic of the La Quinta formation, Venezuelan Andes"
4950:
4707:
4617:
4406:
4322:
4184:
4103:
4075:
4035:
3916:
3886:
3404:
3157:
3021:
2878:
2778:
2627:
2534:
2492:
2059:
1369:
876:
745:
737:
880:
461:
5148:
5013:
4751:
4714:
4646:
4501:
4494:
4306:
4248:
4221:
4211:
4170:
4005:
3864:
3848:
3666:
3638:
3616:
3503:
3348:
3055:
2829:
2712:
2690:
2611:
2440:
2417:
2410:
2388:
2374:
2367:
2328:
2179:
2026:
1953:
1942:
1878:
1766:
1653:
1230:
1139:
993:
968:
612:
631:(an inferred though yet unproven line of descent) by twenty-five million years.
4978:
4941:
4922:
4884:
4873:
4670:
4383:
4342:
4333:
4315:
4295:
4191:
3905:
3650:
3512:
3339:
3321:
3149:
3116:
3102:
3063:
2837:
2698:
2655:
2648:
2570:
2546:
2499:
2458:
2395:
2381:
2109:
2094:
1921:
1755:
1619:
1587:
1576:
1553:
1515:
1376:
1346:
1270:
1209:
1108:
616:
449:
438:
30:
501:
5233:
5020:
4866:
4827:
4806:
4685:
4678:
4605:
4399:
4372:
4230:
4146:
4124:
4117:
4110:
4053:
3970:
3833:
3805:
3792:
3716:
3593:
3433:
3267:
3198:
3084:
3071:
2816:
2794:
2787:
2751:
2724:
2676:
2600:
2336:
2127:
2101:
2017:
1998:
1991:
1928:
1907:
1886:
1871:
1857:
1816:
1782:
1775:
1694:
1546:
1433:
1426:
1353:
1323:
1312:
1202:
1173:
1166:
1092:
725:
666:
628:
408:
68:
4992:
4930:
4900:
4820:
4813:
4791:
4783:
4663:
4655:
4635:
4350:
4261:
4199:
4089:
4060:
4012:
3841:
3826:
3601:
3533:
3461:
3440:
3419:
3235:
3191:
2999:
2986:
2916:
2869:
2744:
2634:
2586:
2579:
2516:
2485:
2478:
2359:
2186:
2038:
1935:
1837:
1830:
1725:
1709:
1645:
1596:
1528:
1305:
1188:
1181:
1152:
1115:
1084:
759:
660:
600:
566:. A ridge on the front surface demarcates the upper limit of this area. In
543:
480:
416:
233:
120:
5157:
5046:
5027:
4957:
4907:
4893:
4835:
4776:
4721:
4598:
4392:
4268:
4237:
4153:
4139:
3871:
3819:
3687:
3609:
3572:
3454:
3426:
3379:
3299:
3177:
3034:
2885:
2858:
2758:
2705:
2641:
2620:
2593:
2344:
2157:
2150:
2136:
1962:
1809:
1800:
1702:
1673:
1603:
1568:
1561:
1361:
1339:
1332:
1291:
1283:
1259:
1223:
1131:
1123:
43:
854:
741:
373:
5190:
4985:
4914:
4859:
4480:
4279:
4096:
4082:
4067:
4019:
3953:
3879:
3701:
3694:
3680:
3586:
3519:
3447:
3411:
3364:
3306:
3292:
3259:
3251:
3244:
3225:
3048:
3013:
2801:
2402:
2193:
2143:
2080:
2051:
2006:
1984:
1977:
1969:
1893:
1789:
1748:
1741:
1733:
1717:
1637:
1508:
1384:
1237:
1100:
959:
563:
441:
362:
207:
111:
88:
53:
391:
306:
5006:
4999:
4699:
4692:
4365:
4357:
4288:
3941:
3659:
3205:
3109:
2844:
2429:
2314:
2301:
2228:
2164:
2087:
2066:
1823:
1686:
1494:
1481:
1399:
1251:
1070:
1057:
1020:
929:
608:
592:
588:
457:
403:
384:
329:
314:
220:
161:
93:
37:
5119:
5142:
4590:
3998:
3991:
3963:
3856:
3356:
2737:
2073:
1900:
1661:
1298:
977:
953:
525:
468:
395:
358:
321:
317:
194:
181:
129:
83:
78:
63:
58:
48:
433:
The description was based on two fossils, found in a layer of the
5177:
4131:
3893:
3623:
3526:
2851:
2172:
831:"New meat-eating dinosaur lived in the wake of a mass extinction"
655:
484:
453:
445:
394:, remains of dinosaurs have been uncovered at a road-cut between
351:
347:
333:
137:
98:
73:
591:
analysis, establishing its probable evolutionary relationships (
4799:
4487:
1914:
947:
555:
171:
808:"Newfound South American Predator Snacked on Little Dinosaurs"
471:
specimen, IVIC-P-2867. It consists of a nearly complete right
3630:
3371:
3313:
707:
496:
472:
426:, Ascanio D. Rincón, Jahandar Ramezani, Andrés Solórzano and
343:
310:
133:
898:
5063:
3673:
460:
has been confidently established, but due to the limits of
293:
Langer, RincĂłn, Ramezani, SolĂłrzano, & Rauhut, 2014
253:
Langer, RincĂłn, Ramezani, SolĂłrzano, & Rauhut, 2014
4472:
546:, a unique evolutionary innovation shown just by
5231:
4574:
4440:
390:Since the late 1980s in the Venezuelan state of
3914:
805:
475:or shinbone. The second fossil was referred to
2438:
992:
772:
914:
611:, the group all theropods belong to from the
3803:
2997:
2915:
2037:
848:
828:
479:on the assumption that only one species of
921:
907:
119:
5245:Early Jurassic dinosaurs of South America
779:was T-Rex's Tiny Ancestor from Venezuela"
749:
554:the outer projection, at the side of the
2312:
2227:
1492:
1068:
703:
372:
701:
699:
697:
695:
693:
691:
689:
687:
685:
683:
664:, from Colombia, and the ornithischian
444:stage. The region was once part of the
5232:
615:onwards, including the birds. Being a
5124:
5123:
4562:
4561:
3790:
2984:
2299:
1479:
1055:
940:
902:
801:
799:
136:, and IVIC-P-2868, the referred left
680:
607:was part of the stem leading to the
402:. Most of these belonged to a small
645:
13:
861:Journal of Vertebrate Paleontology
822:
806:Charles Q. Choi (7 October 2014).
796:
766:
562:of the shinbone is covered by the
14:
5276:
3791:
773:Hannah Osborne (9 October 2014).
456:. A maximum age of 200.72 ± 0.32
365:like other dinosaurs or lizards.
5104:
5098:
5091:
4543:
4536:
4529:
4523:
3772:
3765:
3760:
3753:
3747:
2985:
2966:
2959:
2952:
2281:
2274:
2267:
2260:
1462:
1456:
148:
41:
512:was chosen in reference to the
829:Sid Perkins (7 October 2014).
775:"Dinosaur Species Discovered:
531:
406:that in 2014 was described as
342:, described from a fossilized
1:
5260:Fossil taxa described in 2014
2300:
1480:
1056:
928:
881:10.1080/02724634.2021.2077112
673:
528:was of strategic importance.
783:International Business Times
578:
490:
378:Tachiraptor admirabilis
368:
7:
422:was named and described by
10:
5281:
718:Royal Society Open Science
710:"New dinosaur (Theropoda,
599:was a basal member of the
16:Extinct genus of dinosaurs
5132:
5088:
5045:
4940:
4883:
4854:
4845:
4771:
4762:
4746:
4735:
4645:
4630:
4626:
4615:
4585:
4570:
4557:
4521:
4468:
4436:
4382:
4332:
4305:
4278:
4256:
4247:
4220:
4209:
4179:
4168:
4048:
4044:
4033:
3981:
3952:
3929:
3925:
3903:
3814:
3799:
3786:
3745:
3715:
3649:
3581:
3571:
3567:
3502:
3487:
3478:
3399:
3390:
3338:
3287:
3278:
3234:
3223:
3167:
3144:
3135:
3097:
3093:
3082:
3043:
3032:
3008:
2993:
2980:
2949:
2911:
2868:
2824:
2815:
2777:
2732:
2723:
2685:
2674:
2610:
2569:
2554:
2545:
2529:
2525:
2514:
2468:
2453:
2449:
2427:
2323:
2308:
2295:
2257:
2223:
2126:
2046:
2016:
1952:
1866:
1855:
1799:
1765:
1681:
1672:
1632:
1628:
1617:
1586:
1541:
1537:
1526:
1503:
1488:
1475:
1453:
1418:
1398:
1322:
1278:
1269:
1161:
1150:
1079:
1064:
1051:
1019:
1001:
988:
936:
280:
273:
145:Scientific classification
143:
127:
118:
23:
487:, a bone of the pelvis.
777:Tachiraptor admirabilis
477:Tachiraptor admirabilis
420:Tachiraptor admirabilis
339:Tachiraptor admirabilis
287:Tachiraptor admirabilis
654:before the breakup of
516:of 1813, conducted by
387:
5212:Paleobiology Database
3480:Carcharodontosaurinae
3392:Carcharodontosauridae
583:The study describing
376:
361:, preying on smaller
5255:Fossils of Venezuela
4473:Dubious coelurosaurs
3169:Metriacanthosaurinae
3137:Metriacanthosauridae
1419:Dubious neotheropods
3493:Carcharodontosaurus
3280:Carcharodontosauria
873:2022JVPal..42E7112R
742:10.1098/rsos.140184
734:2014RSOS....140184L
627:thus reduced their
435:La Quinta Formation
326:La Quinta Formation
320:found in the early
266:T. admirabilis
36:201.46–200.36
5250:Jurassic Venezuela
4965:Archaeornithomimus
3469:Veterupristisaurus
3329:Veterupristisaurus
3185:Metriacanthosaurus
3125:Piatnitzkysauridae
2560:Eustreptospondylus
2470:Piatnitzkysauridae
2353:Chuandongocoelurus
1217:Notatesseraeraptor
670:, from Venezuela.
514:Admirable Campaign
465:radiometric dating
424:Max Cardoso Langer
388:
350:. They were small
332:. It includes one
5265:Hettangian genera
5227:
5226:
5199:Open Tree of Life
5126:Taxon identifiers
5117:
5116:
5113:
5112:
5086:
5085:
5082:
5081:
5078:
5077:
5074:
5073:
5041:
5040:
5037:
5036:
4847:Macrocheiriformes
4764:Ornithomimosauria
4738:Maniraptoriformes
4731:
4730:
4577:Maniraptoromorpha
4564:Maniraptoromorpha
4553:
4552:
4519:
4518:
4515:
4514:
4511:
4510:
4464:
4463:
4460:
4459:
4442:Maniraptoromorpha
4432:
4431:
4428:
4427:
4424:
4423:
4420:
4419:
4416:
4415:
4164:
4163:
4029:
4028:
3983:Proceratosauridae
3935:Chingkankousaurus
3782:
3781:
3743:
3742:
3739:
3738:
3735:
3734:
3711:
3710:
3563:
3562:
3559:
3558:
3555:
3554:
3551:
3550:
3547:
3546:
3543:
3542:
3219:
3218:
3215:
3214:
2976:
2975:
2947:
2946:
2943:
2942:
2939:
2938:
2935:
2934:
2907:
2906:
2903:
2902:
2899:
2898:
2895:
2894:
2811:
2810:
2767:Vallibonavenatrix
2670:
2669:
2666:
2665:
2510:
2509:
2291:
2290:
2255:
2254:
2251:
2250:
2247:
2246:
2219:
2218:
2215:
2214:
2211:
2210:
2207:
2206:
2203:
2202:
1851:
1850:
1847:
1846:
1613:
1612:
1471:
1470:
1451:
1450:
1447:
1446:
1443:
1442:
1394:
1393:
1047:
1046:
1043:
1042:
1039:
1038:
458:million years ago
298:
297:
254:
128:IVIC-P-2687, the
5272:
5220:
5219:
5207:
5206:
5194:
5193:
5181:
5180:
5168:
5167:
5166:
5153:
5152:
5151:
5121:
5120:
5108:
5102:
5095:
4951:Aepyornithomimus
4852:
4851:
4769:
4768:
4744:
4743:
4708:Sinocalliopteryx
4628:
4627:
4624:
4623:
4618:Neocoelurosauria
4583:
4582:
4572:
4571:
4559:
4558:
4547:
4540:
4533:
4527:
4470:
4469:
4438:
4437:
4407:Zhuchengtyrannus
4323:Thanatotheristes
4254:
4253:
4218:
4217:
4185:Appalachiosaurus
4177:
4176:
4046:
4045:
4042:
4041:
4036:Pantyrannosauria
3927:
3926:
3923:
3922:
3917:Tyrannosauroidea
3912:
3911:
3887:Xinjiangovenator
3812:
3811:
3801:
3800:
3788:
3787:
3776:
3769:
3764:
3757:
3751:
3579:
3578:
3569:
3568:
3485:
3484:
3405:Acrocanthosaurus
3397:
3396:
3285:
3284:
3232:
3231:
3158:Yangchuanosaurus
3142:
3141:
3095:
3094:
3091:
3090:
3041:
3040:
3022:Lourinhanosaurus
3006:
3005:
2995:
2994:
2982:
2981:
2970:
2963:
2956:
2913:
2912:
2879:Sigilmassasaurus
2822:
2821:
2779:Ceratosuchopsini
2730:
2729:
2683:
2682:
2628:Dubreuillosaurus
2552:
2551:
2535:Streptospondylus
2527:
2526:
2523:
2522:
2493:Piatnitzkysaurus
2451:
2450:
2447:
2446:
2436:
2435:
2321:
2320:
2310:
2309:
2297:
2296:
2285:
2278:
2271:
2264:
2225:
2224:
2060:Ekrixinatosaurus
2044:
2043:
2035:
2034:
1864:
1863:
1679:
1678:
1630:
1629:
1626:
1625:
1539:
1538:
1535:
1534:
1501:
1500:
1490:
1489:
1477:
1476:
1466:
1460:
1370:Procompsognathus
1276:
1275:
1159:
1158:
1077:
1076:
1066:
1065:
1053:
1052:
999:
998:
990:
989:
983:
982:
938:
937:
923:
916:
909:
900:
899:
893:
892:
852:
846:
845:
843:
841:
826:
820:
819:
817:
815:
803:
794:
793:
791:
789:
770:
764:
763:
753:
705:
646:Paleoenvironment
520:, for which the
499:word for thief,
437:dating from the
289:
285:
252:
245:
153:
152:
123:
110:
40:
29:Temporal range:
21:
20:
5280:
5279:
5275:
5274:
5273:
5271:
5270:
5269:
5230:
5229:
5228:
5223:
5215:
5210:
5202:
5197:
5189:
5184:
5176:
5171:
5162:
5161:
5156:
5147:
5146:
5141:
5128:
5118:
5109:
5096:
5070:
5033:
5014:Sinornithomimus
4936:
4879:
4841:
4758:
4752:Compsognathidae
4740:
4727:
4715:Sinosauropteryx
4647:Compsognathidae
4641:
4620:
4611:
4579:
4566:
4549:
4548:
4534:
4507:
4502:Shanyangosaurus
4495:Phaedrolosaurus
4456:
4412:
4378:
4328:
4307:Daspletosaurini
4301:
4274:
4249:Tyrannosaurinae
4243:
4222:Albertosaurinae
4214:
4212:Tyrannosauridae
4205:
4173:
4171:Eutyrannosauria
4160:
4038:
4025:
4006:Proceratosaurus
3977:
3948:
3919:
3908:
3899:
3865:Richardoestesia
3849:Chilantaisaurus
3808:
3795:
3778:
3777:
3758:
3731:
3707:
3667:Australovenator
3645:
3639:Chilantaisaurus
3617:Phuwiangvenator
3539:
3504:Giganotosaurini
3498:
3474:
3386:
3349:Chilantaisaurus
3334:
3274:
3228:
3211:
3163:
3131:
3087:
3078:
3056:Monolophosaurus
3037:
3028:
3002:
2989:
2972:
2971:
2957:
2931:
2891:
2864:
2830:Camarillasaurus
2807:
2773:
2719:
2713:Ostafrikasaurus
2691:Camarillasaurus
2679:
2662:
2612:Afrovenatorinae
2606:
2565:
2541:
2519:
2506:
2464:
2443:
2441:Megalosauroidea
2432:
2423:
2418:Vectaerovenator
2411:Szechuanosaurus
2389:Monolophosaurus
2375:Kaijiangosaurus
2368:Cryolophosaurus
2329:Calamospondylus
2317:
2304:
2287:
2286:
2279:
2272:
2265:
2243:
2199:
2180:Pycnonemosaurus
2122:
2033:
2027:Majungasaurinae
2012:
1954:Majungasaurinae
1948:
1943:Xenotarsosaurus
1879:Dryptosauroides
1860:
1843:
1795:
1767:Elaphrosaurinae
1761:
1668:
1654:Ornithomimoides
1622:
1609:
1582:
1531:
1522:
1497:
1484:
1467:
1461:
1439:
1414:
1390:
1318:
1265:
1231:Shuangbaisaurus
1155:
1146:
1073:
1060:
1035:
1015:
1009:Avemetatarsalia
994:Avemetatarsalia
984:
969:Avemetatarsalia
943:
942:
932:
927:
897:
896:
867:(1): e2077112.
853:
849:
839:
837:
827:
823:
813:
811:
804:
797:
787:
785:
771:
767:
706:
681:
676:
648:
613:Middle Jurassic
581:
534:
493:
448:of the ancient
446:equatorial belt
371:
313:of carnivorous
294:
291:
283:
282:
269:
251:
243:
147:
114:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
27:
17:
12:
11:
5:
5278:
5268:
5267:
5262:
5257:
5252:
5247:
5242:
5225:
5224:
5222:
5221:
5208:
5195:
5182:
5169:
5154:
5138:
5136:
5130:
5129:
5115:
5114:
5111:
5110:
5090:
5089:
5087:
5084:
5083:
5080:
5079:
5076:
5075:
5072:
5071:
5069:
5068:
5067:
5066:
5060:
5051:
5049:
5043:
5042:
5039:
5038:
5035:
5034:
5032:
5031:
5024:
5017:
5010:
5003:
4996:
4989:
4982:
4979:Dromiceiomimus
4975:
4968:
4961:
4954:
4946:
4944:
4942:Ornithomimidae
4938:
4937:
4935:
4934:
4927:
4923:Paraxenisaurus
4919:
4911:
4904:
4897:
4889:
4887:
4885:Deinocheiridae
4881:
4880:
4878:
4877:
4874:Shenzhousaurus
4870:
4863:
4855:
4849:
4843:
4842:
4840:
4839:
4832:
4824:
4817:
4810:
4803:
4796:
4788:
4780:
4772:
4766:
4760:
4759:
4757:
4756:
4747:
4741:
4736:
4733:
4732:
4729:
4728:
4726:
4725:
4718:
4711:
4704:
4696:
4689:
4682:
4675:
4671:Beipiaognathus
4667:
4660:
4651:
4649:
4643:
4642:
4640:
4639:
4631:
4621:
4616:
4613:
4612:
4610:
4609:
4602:
4595:
4586:
4580:
4575:
4568:
4567:
4555:
4554:
4551:
4550:
4542:
4528:
4522:
4520:
4517:
4516:
4513:
4512:
4509:
4508:
4506:
4505:
4498:
4491:
4484:
4476:
4474:
4466:
4465:
4462:
4461:
4458:
4457:
4455:
4454:
4453:
4452:
4446:
4444:
4434:
4433:
4430:
4429:
4426:
4425:
4422:
4421:
4418:
4417:
4414:
4413:
4411:
4410:
4403:
4396:
4388:
4386:
4384:Tyrannosaurini
4380:
4379:
4377:
4376:
4369:
4362:
4354:
4347:
4343:Bistahieversor
4338:
4336:
4334:Teratophoneini
4330:
4329:
4327:
4326:
4319:
4316:Daspletosaurus
4311:
4309:
4303:
4302:
4300:
4299:
4296:Qianzhousaurus
4292:
4284:
4282:
4276:
4275:
4273:
4272:
4265:
4257:
4251:
4245:
4244:
4242:
4241:
4234:
4226:
4224:
4215:
4210:
4207:
4206:
4204:
4203:
4196:
4192:Bistahieversor
4188:
4180:
4174:
4169:
4166:
4165:
4162:
4161:
4159:
4158:
4150:
4143:
4136:
4128:
4121:
4114:
4107:
4100:
4093:
4086:
4079:
4072:
4064:
4057:
4049:
4039:
4034:
4031:
4030:
4027:
4026:
4024:
4023:
4016:
4009:
4002:
3995:
3987:
3985:
3979:
3978:
3976:
3975:
3967:
3959:
3957:
3950:
3949:
3947:
3946:
3938:
3930:
3920:
3915:
3909:
3906:Tyrannoraptora
3904:
3901:
3900:
3898:
3897:
3890:
3883:
3876:
3868:
3861:
3853:
3845:
3838:
3830:
3823:
3815:
3809:
3804:
3797:
3796:
3784:
3783:
3780:
3779:
3771:
3752:
3746:
3744:
3741:
3740:
3737:
3736:
3733:
3732:
3730:
3729:
3728:
3727:
3721:
3719:
3713:
3712:
3709:
3708:
3706:
3705:
3698:
3691:
3684:
3677:
3670:
3663:
3655:
3653:
3651:Megaraptoridae
3647:
3646:
3644:
3643:
3635:
3627:
3620:
3613:
3606:
3598:
3590:
3582:
3576:
3565:
3564:
3561:
3560:
3557:
3556:
3553:
3552:
3549:
3548:
3545:
3544:
3541:
3540:
3538:
3537:
3530:
3523:
3516:
3513:Giganotosaurus
3508:
3506:
3500:
3499:
3497:
3496:
3488:
3482:
3476:
3475:
3473:
3472:
3465:
3458:
3451:
3444:
3437:
3430:
3423:
3416:
3408:
3400:
3394:
3388:
3387:
3385:
3384:
3376:
3368:
3361:
3353:
3344:
3342:
3340:Neovenatoridae
3336:
3335:
3333:
3332:
3325:
3322:Ulughbegsaurus
3318:
3310:
3303:
3296:
3288:
3282:
3276:
3275:
3273:
3272:
3264:
3256:
3248:
3240:
3238:
3229:
3224:
3221:
3220:
3217:
3216:
3213:
3212:
3210:
3209:
3202:
3195:
3188:
3181:
3173:
3171:
3165:
3164:
3162:
3161:
3154:
3150:Xuanhanosaurus
3145:
3139:
3133:
3132:
3130:
3129:
3121:
3117:Xuanhanosaurus
3113:
3106:
3103:Asfaltovenator
3098:
3088:
3083:
3080:
3079:
3077:
3076:
3068:
3064:Megalosauridae
3060:
3052:
3044:
3038:
3033:
3030:
3029:
3027:
3026:
3018:
3009:
3003:
2998:
2991:
2990:
2978:
2977:
2974:
2973:
2965:
2951:
2950:
2948:
2945:
2944:
2941:
2940:
2937:
2936:
2933:
2932:
2930:
2929:
2928:
2927:
2921:
2919:
2909:
2908:
2905:
2904:
2901:
2900:
2897:
2896:
2893:
2892:
2890:
2889:
2882:
2874:
2872:
2866:
2865:
2863:
2862:
2855:
2848:
2841:
2838:Ichthyovenator
2834:
2825:
2819:
2813:
2812:
2809:
2808:
2806:
2805:
2798:
2791:
2783:
2781:
2775:
2774:
2772:
2771:
2763:
2755:
2748:
2741:
2733:
2727:
2721:
2720:
2718:
2717:
2709:
2702:
2699:Cristatusaurus
2695:
2686:
2680:
2675:
2672:
2671:
2668:
2667:
2664:
2663:
2661:
2660:
2656:Poekilopleuron
2652:
2649:Piveteausaurus
2645:
2638:
2631:
2624:
2616:
2614:
2608:
2607:
2605:
2604:
2597:
2590:
2583:
2575:
2573:
2571:Megalosaurinae
2567:
2566:
2564:
2563:
2555:
2549:
2547:Megalosauridae
2543:
2542:
2540:
2539:
2530:
2520:
2515:
2512:
2511:
2508:
2507:
2505:
2504:
2500:Xuanhanosaurus
2496:
2489:
2482:
2474:
2472:
2466:
2465:
2463:
2462:
2459:Yunyangosaurus
2454:
2444:
2439:
2433:
2428:
2425:
2424:
2422:
2421:
2414:
2407:
2399:
2396:Pandoravenator
2392:
2385:
2382:Kayentavenator
2378:
2371:
2364:
2356:
2349:
2341:
2333:
2324:
2318:
2313:
2306:
2305:
2293:
2292:
2289:
2288:
2280:
2273:
2266:
2259:
2258:
2256:
2253:
2252:
2249:
2248:
2245:
2244:
2242:
2241:
2240:
2239:
2233:
2231:
2221:
2220:
2217:
2216:
2213:
2212:
2209:
2208:
2205:
2204:
2201:
2200:
2198:
2197:
2190:
2183:
2176:
2169:
2161:
2154:
2147:
2140:
2132:
2130:
2124:
2123:
2121:
2120:
2113:
2110:Skorpiovenator
2106:
2098:
2091:
2084:
2077:
2070:
2063:
2056:
2047:
2041:
2032:
2031:
2022:
2020:
2014:
2013:
2011:
2010:
2003:
1995:
1988:
1981:
1974:
1966:
1958:
1956:
1950:
1949:
1947:
1946:
1939:
1932:
1925:
1922:Spectrovenator
1918:
1911:
1904:
1897:
1890:
1883:
1875:
1867:
1861:
1856:
1853:
1852:
1849:
1848:
1845:
1844:
1842:
1841:
1834:
1827:
1820:
1813:
1805:
1803:
1797:
1796:
1794:
1793:
1786:
1779:
1771:
1769:
1763:
1762:
1760:
1759:
1756:Spinostropheus
1752:
1745:
1738:
1730:
1722:
1714:
1706:
1699:
1691:
1682:
1676:
1670:
1669:
1667:
1666:
1658:
1650:
1642:
1633:
1623:
1620:Abelisauroidea
1618:
1615:
1614:
1611:
1610:
1608:
1607:
1600:
1592:
1590:
1588:Ceratosauridae
1584:
1583:
1581:
1580:
1577:Saltriovenator
1573:
1565:
1558:
1554:Fosterovenator
1550:
1542:
1532:
1527:
1524:
1523:
1521:
1520:
1516:Lophostropheus
1512:
1504:
1498:
1493:
1486:
1485:
1473:
1472:
1469:
1468:
1455:
1454:
1452:
1449:
1448:
1445:
1444:
1441:
1440:
1438:
1437:
1430:
1422:
1420:
1416:
1415:
1413:
1412:
1411:
1410:
1404:
1402:
1396:
1395:
1392:
1391:
1389:
1388:
1381:
1377:Pterospondylus
1373:
1366:
1358:
1350:
1347:Lucianovenator
1343:
1336:
1328:
1326:
1320:
1319:
1317:
1316:
1309:
1302:
1295:
1288:
1279:
1273:
1271:Coelophysoidea
1267:
1266:
1264:
1263:
1256:
1248:
1241:
1234:
1227:
1220:
1213:
1210:Lophostropheus
1206:
1199:
1192:
1185:
1178:
1170:
1162:
1156:
1151:
1148:
1147:
1145:
1144:
1136:
1128:
1120:
1112:
1109:Erythrovenator
1105:
1097:
1089:
1080:
1074:
1069:
1062:
1061:
1049:
1048:
1045:
1044:
1041:
1040:
1037:
1036:
1034:
1033:
1032:
1031:
1025:
1023:
1017:
1016:
1014:
1013:
1012:
1011:
1002:
996:
986:
985:
981:
980:
971:
962:
956:
950:
934:
933:
926:
925:
918:
911:
903:
895:
894:
847:
821:
795:
765:
678:
677:
675:
672:
647:
644:
617:sister species
580:
577:
533:
530:
492:
489:
450:supercontinent
439:Early Jurassic
370:
367:
296:
295:
292:
278:
277:
271:
270:
262:
260:
256:
255:
241:
237:
236:
231:
224:
223:
218:
211:
210:
205:
198:
197:
192:
185:
184:
179:
175:
174:
169:
165:
164:
159:
155:
154:
141:
140:
125:
124:
116:
115:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
31:Early Jurassic
28:
15:
9:
6:
4:
3:
2:
5277:
5266:
5263:
5261:
5258:
5256:
5253:
5251:
5248:
5246:
5243:
5241:
5238:
5237:
5235:
5218:
5213:
5209:
5205:
5200:
5196:
5192:
5187:
5183:
5179:
5174:
5170:
5165:
5159:
5155:
5150:
5144:
5140:
5139:
5137:
5135:
5131:
5127:
5122:
5107:
5103:
5101:
5094:
5065:
5061:
5059:
5055:
5054:
5053:
5052:
5050:
5048:
5044:
5030:
5029:
5025:
5023:
5022:
5021:Struthiomimus
5018:
5016:
5015:
5011:
5009:
5008:
5004:
5002:
5001:
4997:
4995:
4994:
4990:
4988:
4987:
4983:
4981:
4980:
4976:
4974:
4973:
4972:"Coelosaurus"
4969:
4967:
4966:
4962:
4960:
4959:
4955:
4953:
4952:
4948:
4947:
4945:
4943:
4939:
4933:
4932:
4928:
4925:
4924:
4920:
4917:
4916:
4912:
4910:
4909:
4905:
4903:
4902:
4898:
4896:
4895:
4891:
4890:
4888:
4886:
4882:
4876:
4875:
4871:
4869:
4868:
4867:Pelecanimimus
4864:
4862:
4861:
4857:
4856:
4853:
4850:
4848:
4844:
4838:
4837:
4833:
4830:
4829:
4828:Thecocoelurus
4825:
4823:
4822:
4818:
4816:
4815:
4811:
4809:
4808:
4807:Kinnareemimus
4804:
4802:
4801:
4797:
4794:
4793:
4789:
4786:
4785:
4781:
4779:
4778:
4774:
4773:
4770:
4767:
4765:
4761:
4754:
4753:
4749:
4748:
4745:
4742:
4739:
4734:
4724:
4723:
4719:
4717:
4716:
4712:
4710:
4709:
4705:
4702:
4701:
4697:
4695:
4694:
4690:
4688:
4687:
4686:Huaxiagnathus
4683:
4681:
4680:
4679:Compsognathus
4676:
4673:
4672:
4668:
4666:
4665:
4661:
4658:
4657:
4653:
4652:
4650:
4648:
4644:
4638:
4637:
4633:
4632:
4629:
4625:
4622:
4619:
4614:
4608:
4607:
4606:Ornitholestes
4603:
4601:
4600:
4596:
4593:
4592:
4588:
4587:
4584:
4581:
4578:
4573:
4569:
4565:
4560:
4556:
4546:
4541:
4539:
4532:
4526:
4504:
4503:
4499:
4497:
4496:
4492:
4490:
4489:
4485:
4483:
4482:
4478:
4477:
4475:
4471:
4467:
4450:
4449:
4448:
4447:
4445:
4443:
4439:
4435:
4409:
4408:
4404:
4402:
4401:
4400:Tyrannosaurus
4397:
4395:
4394:
4390:
4389:
4387:
4385:
4381:
4375:
4374:
4373:Teratophoneus
4370:
4368:
4367:
4363:
4360:
4359:
4355:
4353:
4352:
4348:
4345:
4344:
4340:
4339:
4337:
4335:
4331:
4325:
4324:
4320:
4318:
4317:
4313:
4312:
4310:
4308:
4304:
4298:
4297:
4293:
4291:
4290:
4286:
4285:
4283:
4281:
4277:
4271:
4270:
4266:
4264:
4263:
4259:
4258:
4255:
4252:
4250:
4246:
4240:
4239:
4235:
4233:
4232:
4231:Albertosaurus
4228:
4227:
4225:
4223:
4219:
4216:
4213:
4208:
4202:
4201:
4197:
4194:
4193:
4189:
4187:
4186:
4182:
4181:
4178:
4175:
4172:
4167:
4156:
4155:
4151:
4149:
4148:
4147:Xiongguanlong
4144:
4142:
4141:
4137:
4134:
4133:
4129:
4127:
4126:
4125:Suskityrannus
4122:
4120:
4119:
4118:Stokesosaurus
4115:
4113:
4112:
4111:Santanaraptor
4108:
4106:
4105:
4101:
4099:
4098:
4094:
4092:
4091:
4087:
4085:
4084:
4080:
4078:
4077:
4073:
4070:
4069:
4065:
4063:
4062:
4058:
4056:
4055:
4054:Alectrosaurus
4051:
4050:
4047:
4043:
4040:
4037:
4032:
4022:
4021:
4017:
4015:
4014:
4010:
4008:
4007:
4003:
4001:
4000:
3996:
3994:
3993:
3989:
3988:
3986:
3984:
3980:
3973:
3972:
3971:Tanycolagreus
3968:
3966:
3965:
3961:
3960:
3958:
3955:
3951:
3944:
3943:
3939:
3937:
3936:
3932:
3931:
3928:
3924:
3921:
3918:
3913:
3910:
3907:
3902:
3896:
3895:
3891:
3889:
3888:
3884:
3882:
3881:
3877:
3874:
3873:
3869:
3867:
3866:
3862:
3859:
3858:
3854:
3851:
3850:
3846:
3844:
3843:
3839:
3836:
3835:
3834:Bahariasaurus
3831:
3829:
3828:
3824:
3822:
3821:
3817:
3816:
3813:
3810:
3807:
3806:Coelurosauria
3802:
3798:
3794:
3793:Coelurosauria
3789:
3785:
3775:
3770:
3768:
3763:
3756:
3750:
3725:
3724:
3723:
3722:
3720:
3718:
3717:Coelurosauria
3714:
3704:
3703:
3699:
3697:
3696:
3692:
3690:
3689:
3685:
3683:
3682:
3678:
3676:
3675:
3671:
3669:
3668:
3664:
3662:
3661:
3657:
3656:
3654:
3652:
3648:
3642:
3640:
3636:
3634:
3632:
3628:
3626:
3625:
3621:
3619:
3618:
3614:
3612:
3611:
3607:
3604:
3603:
3599:
3596:
3595:
3594:Bahariasaurus
3591:
3589:
3588:
3584:
3583:
3580:
3577:
3574:
3570:
3566:
3536:
3535:
3531:
3529:
3528:
3524:
3522:
3521:
3517:
3515:
3514:
3510:
3509:
3507:
3505:
3501:
3495:
3494:
3490:
3489:
3486:
3483:
3481:
3477:
3471:
3470:
3466:
3464:
3463:
3459:
3457:
3456:
3452:
3450:
3449:
3445:
3443:
3442:
3438:
3436:
3435:
3434:Kelmayisaurus
3431:
3429:
3428:
3424:
3422:
3421:
3417:
3414:
3413:
3409:
3407:
3406:
3402:
3401:
3398:
3395:
3393:
3389:
3382:
3381:
3377:
3374:
3373:
3369:
3367:
3366:
3362:
3359:
3358:
3354:
3351:
3350:
3346:
3345:
3343:
3341:
3337:
3331:
3330:
3326:
3324:
3323:
3319:
3316:
3315:
3311:
3309:
3308:
3304:
3302:
3301:
3297:
3295:
3294:
3290:
3289:
3286:
3283:
3281:
3277:
3270:
3269:
3268:Saurophaganax
3265:
3262:
3261:
3257:
3254:
3253:
3249:
3247:
3246:
3242:
3241:
3239:
3237:
3233:
3230:
3227:
3222:
3208:
3207:
3203:
3201:
3200:
3199:Siamotyrannus
3196:
3194:
3193:
3189:
3187:
3186:
3182:
3180:
3179:
3175:
3174:
3172:
3170:
3166:
3160:
3159:
3155:
3152:
3151:
3147:
3146:
3143:
3140:
3138:
3134:
3127:
3126:
3122:
3119:
3118:
3114:
3112:
3111:
3107:
3105:
3104:
3100:
3099:
3096:
3092:
3089:
3086:
3085:Allosauroidea
3081:
3074:
3073:
3072:Spinosauridae
3069:
3066:
3065:
3061:
3058:
3057:
3053:
3051:
3050:
3046:
3045:
3042:
3039:
3036:
3031:
3024:
3023:
3019:
3016:
3015:
3011:
3010:
3007:
3004:
3001:
2996:
2992:
2988:
2983:
2979:
2969:
2964:
2962:
2955:
2925:
2924:
2923:
2922:
2920:
2918:
2914:
2910:
2888:
2887:
2883:
2881:
2880:
2876:
2875:
2873:
2871:
2867:
2861:
2860:
2856:
2854:
2853:
2849:
2847:
2846:
2842:
2840:
2839:
2835:
2832:
2831:
2827:
2826:
2823:
2820:
2818:
2817:Spinosaurinae
2814:
2804:
2803:
2799:
2797:
2796:
2795:Riparovenator
2792:
2790:
2789:
2788:Ceratosuchops
2785:
2784:
2782:
2780:
2776:
2769:
2768:
2764:
2761:
2760:
2756:
2754:
2753:
2752:Riojavenatrix
2749:
2747:
2746:
2742:
2740:
2739:
2735:
2734:
2731:
2728:
2726:
2725:Baryonychinae
2722:
2715:
2714:
2710:
2708:
2707:
2703:
2701:
2700:
2696:
2693:
2692:
2688:
2687:
2684:
2681:
2678:
2677:Spinosauridae
2673:
2658:
2657:
2653:
2651:
2650:
2646:
2644:
2643:
2639:
2637:
2636:
2632:
2630:
2629:
2625:
2623:
2622:
2618:
2617:
2615:
2613:
2609:
2603:
2602:
2601:Wiehenvenator
2598:
2596:
2595:
2591:
2589:
2588:
2584:
2582:
2581:
2577:
2576:
2574:
2572:
2568:
2562:
2561:
2557:
2556:
2553:
2550:
2548:
2544:
2537:
2536:
2532:
2531:
2528:
2524:
2521:
2518:
2513:
2502:
2501:
2497:
2495:
2494:
2490:
2488:
2487:
2483:
2481:
2480:
2476:
2475:
2473:
2471:
2467:
2461:
2460:
2456:
2455:
2452:
2448:
2445:
2442:
2437:
2434:
2431:
2426:
2420:
2419:
2415:
2413:
2412:
2408:
2405:
2404:
2400:
2398:
2397:
2393:
2391:
2390:
2386:
2384:
2383:
2379:
2377:
2376:
2372:
2370:
2369:
2365:
2362:
2361:
2357:
2355:
2354:
2350:
2347:
2346:
2342:
2339:
2338:
2337:Chienkosaurus
2334:
2331:
2330:
2326:
2325:
2322:
2319:
2316:
2311:
2307:
2303:
2298:
2294:
2284:
2277:
2270:
2263:
2237:
2236:
2235:
2234:
2232:
2230:
2226:
2222:
2196:
2195:
2191:
2189:
2188:
2184:
2182:
2181:
2177:
2175:
2174:
2170:
2167:
2166:
2162:
2160:
2159:
2155:
2153:
2152:
2148:
2146:
2145:
2141:
2139:
2138:
2134:
2133:
2131:
2129:
2128:Furileusauria
2125:
2119:
2118:
2114:
2112:
2111:
2107:
2104:
2103:
2102:Rahiolisaurus
2099:
2097:
2096:
2092:
2090:
2089:
2085:
2083:
2082:
2078:
2076:
2075:
2071:
2069:
2068:
2064:
2062:
2061:
2057:
2054:
2053:
2049:
2048:
2045:
2042:
2040:
2036:
2029:
2028:
2024:
2023:
2021:
2019:
2018:Carnotaurinae
2015:
2009:
2008:
2004:
2001:
2000:
1999:Rahiolisaurus
1996:
1994:
1993:
1992:Majungasaurus
1989:
1987:
1986:
1982:
1980:
1979:
1975:
1972:
1971:
1967:
1965:
1964:
1960:
1959:
1957:
1955:
1951:
1945:
1944:
1940:
1938:
1937:
1933:
1931:
1930:
1929:Tarascosaurus
1926:
1924:
1923:
1919:
1917:
1916:
1912:
1910:
1909:
1905:
1903:
1902:
1898:
1896:
1895:
1891:
1889:
1888:
1887:Eoabelisaurus
1884:
1881:
1880:
1876:
1874:
1873:
1872:Chenanisaurus
1869:
1868:
1865:
1862:
1859:
1858:Abelisauridae
1854:
1840:
1839:
1835:
1833:
1832:
1828:
1826:
1825:
1821:
1819:
1818:
1817:Masiakasaurus
1814:
1812:
1811:
1807:
1806:
1804:
1802:
1798:
1792:
1791:
1787:
1785:
1784:
1783:Huinculsaurus
1780:
1778:
1777:
1776:Elaphrosaurus
1773:
1772:
1770:
1768:
1764:
1758:
1757:
1753:
1751:
1750:
1746:
1744:
1743:
1739:
1736:
1735:
1731:
1728:
1727:
1723:
1720:
1719:
1715:
1712:
1711:
1707:
1705:
1704:
1700:
1697:
1696:
1695:Austrocheirus
1692:
1689:
1688:
1684:
1683:
1680:
1677:
1675:
1671:
1664:
1663:
1659:
1656:
1655:
1651:
1648:
1647:
1643:
1640:
1639:
1635:
1634:
1631:
1627:
1624:
1621:
1616:
1606:
1605:
1601:
1599:
1598:
1594:
1593:
1591:
1589:
1585:
1579:
1578:
1574:
1571:
1570:
1566:
1564:
1563:
1559:
1556:
1555:
1551:
1549:
1548:
1547:Berberosaurus
1544:
1543:
1540:
1536:
1533:
1530:
1525:
1518:
1517:
1513:
1511:
1510:
1506:
1505:
1502:
1499:
1496:
1491:
1487:
1483:
1478:
1474:
1465:
1459:
1436:
1435:
1434:Halticosaurus
1431:
1429:
1428:
1427:Teinurosaurus
1424:
1423:
1421:
1417:
1408:
1407:
1406:
1405:
1403:
1401:
1397:
1387:
1386:
1382:
1379:
1378:
1374:
1372:
1371:
1367:
1364:
1363:
1359:
1356:
1355:
1354:Megapnosaurus
1351:
1349:
1348:
1344:
1342:
1341:
1337:
1335:
1334:
1330:
1329:
1327:
1325:
1324:Coelophysidae
1321:
1315:
1314:
1313:Powellvenator
1310:
1308:
1307:
1303:
1301:
1300:
1296:
1294:
1293:
1289:
1286:
1285:
1281:
1280:
1277:
1274:
1272:
1268:
1262:
1261:
1257:
1254:
1253:
1249:
1247:
1246:
1242:
1240:
1239:
1235:
1233:
1232:
1228:
1226:
1225:
1221:
1219:
1218:
1214:
1212:
1211:
1207:
1205:
1204:
1203:Liliensternus
1200:
1198:
1197:
1193:
1191:
1190:
1186:
1184:
1183:
1179:
1176:
1175:
1174:Dolichosuchus
1171:
1169:
1168:
1167:Dilophosaurus
1164:
1163:
1160:
1157:
1154:
1149:
1142:
1141:
1140:Herrerasauria
1137:
1134:
1133:
1129:
1126:
1125:
1121:
1118:
1117:
1113:
1111:
1110:
1106:
1103:
1102:
1098:
1095:
1094:
1093:Daemonosaurus
1090:
1087:
1086:
1082:
1081:
1078:
1075:
1072:
1067:
1063:
1059:
1054:
1050:
1029:
1028:
1027:
1026:
1024:
1022:
1018:
1010:
1006:
1005:
1004:
1003:
1000:
997:
995:
991:
987:
979:
975:
972:
970:
966:
963:
961:
957:
955:
951:
949:
945:
944:
939:
935:
931:
924:
919:
917:
912:
910:
905:
904:
901:
890:
886:
882:
878:
874:
870:
866:
862:
858:
851:
836:
832:
825:
810:. LiveScience
809:
802:
800:
784:
780:
778:
769:
761:
757:
752:
747:
743:
739:
735:
731:
727:
726:Royal Society
723:
719:
715:
713:
704:
702:
700:
698:
696:
694:
692:
690:
688:
686:
684:
679:
671:
669:
668:
667:Laquintasaura
663:
662:
657:
652:
643:
641:
640:Laquintasaura
636:
632:
630:
629:ghost lineage
626:
622:
618:
614:
610:
606:
602:
598:
594:
590:
586:
576:
574:
569:
565:
559:
557:
553:
549:
545:
540:
538:
529:
527:
523:
522:type locality
519:
518:SimĂłn BolĂvar
515:
511:
508:
507:specific name
504:
503:
498:
488:
486:
482:
478:
474:
470:
466:
463:
459:
455:
451:
447:
443:
440:
436:
431:
429:
428:Oliver Rauhut
425:
421:
418:
415:In 2014, the
413:
411:
410:
409:Laquintasaura
405:
401:
397:
393:
386:
382:
381:type locality
379:
375:
366:
364:
360:
357:
353:
349:
345:
341:
340:
335:
331:
327:
323:
319:
316:
312:
308:
304:
303:
290:
288:
279:
276:
275:Binomial name
272:
268:
267:
261:
258:
257:
250:
249:
242:
239:
238:
235:
232:
229:
226:
225:
222:
219:
216:
213:
212:
209:
206:
203:
200:
199:
196:
193:
190:
187:
186:
183:
180:
177:
176:
173:
170:
167:
166:
163:
160:
157:
156:
151:
146:
142:
139:
135:
131:
126:
122:
117:
113:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
26:
22:
19:
5133:
5097:
5026:
5019:
5012:
5005:
4998:
4993:Ornithomimus
4991:
4984:
4977:
4970:
4963:
4956:
4949:
4931:Tyrannomimus
4929:
4921:
4913:
4906:
4901:Deinocheirus
4899:
4892:
4872:
4865:
4858:
4834:
4826:
4821:Nqwebasaurus
4819:
4814:Nedcolbertia
4812:
4805:
4798:
4792:Calamosaurus
4790:
4784:Aviatyrannis
4782:
4775:
4750:
4720:
4713:
4706:
4698:
4691:
4684:
4677:
4669:
4664:Aristosuchus
4662:
4656:Aniksosaurus
4654:
4636:Aniksosaurus
4634:
4604:
4597:
4589:
4535:
4500:
4493:
4486:
4479:
4405:
4398:
4391:
4371:
4364:
4356:
4351:Dynamoterror
4349:
4341:
4321:
4314:
4294:
4287:
4267:
4262:Asiatyrannus
4260:
4236:
4229:
4200:Dryptosaurus
4198:
4190:
4183:
4152:
4145:
4138:
4130:
4123:
4116:
4109:
4102:
4095:
4090:Jinbeisaurus
4088:
4081:
4074:
4066:
4061:Aviatyrannis
4059:
4052:
4018:
4013:Sinotyrannus
4011:
4004:
3997:
3990:
3969:
3962:
3940:
3933:
3892:
3885:
3878:
3870:
3863:
3855:
3847:
3842:Bicentenaria
3840:
3832:
3827:Asiamericana
3825:
3818:
3759:
3700:
3693:
3686:
3679:
3672:
3665:
3658:
3637:
3629:
3622:
3615:
3608:
3602:Deltadromeus
3600:
3592:
3585:
3534:Tyrannotitan
3532:
3525:
3518:
3511:
3491:
3467:
3462:Taurovenator
3460:
3453:
3446:
3441:Lajasvenator
3439:
3432:
3425:
3420:Concavenator
3418:
3410:
3403:
3378:
3370:
3363:
3355:
3347:
3327:
3320:
3312:
3305:
3298:
3291:
3266:
3258:
3250:
3243:
3236:Allosauridae
3204:
3197:
3192:Shidaisaurus
3190:
3183:
3176:
3156:
3148:
3123:
3115:
3108:
3101:
3070:
3062:
3054:
3047:
3020:
3012:
3000:Avetheropoda
2987:Avetheropoda
2958:
2917:Avetheropoda
2884:
2877:
2870:Spinosaurini
2857:
2850:
2843:
2836:
2828:
2800:
2793:
2786:
2765:
2757:
2750:
2745:Protathlitis
2743:
2736:
2711:
2704:
2697:
2689:
2654:
2647:
2640:
2635:Leshansaurus
2633:
2626:
2619:
2599:
2592:
2587:Megalosaurus
2585:
2580:Duriavenator
2578:
2558:
2533:
2517:Megalosauria
2498:
2491:
2486:Marshosaurus
2484:
2479:Condorraptor
2477:
2457:
2416:
2409:
2401:
2394:
2387:
2380:
2373:
2366:
2360:Cruxicheiros
2358:
2351:
2343:
2335:
2327:
2192:
2187:Quilmesaurus
2185:
2178:
2171:
2163:
2156:
2149:
2142:
2135:
2115:
2108:
2100:
2093:
2086:
2079:
2072:
2065:
2058:
2050:
2039:Brachyrostra
2025:
2005:
1997:
1990:
1983:
1976:
1968:
1961:
1941:
1936:Tralkasaurus
1934:
1927:
1920:
1913:
1906:
1899:
1892:
1885:
1877:
1870:
1838:Vespersaurus
1836:
1831:Velocisaurus
1829:
1822:
1815:
1808:
1788:
1781:
1774:
1754:
1747:
1740:
1732:
1726:Deltadromeus
1724:
1716:
1710:Compsosuchus
1708:
1701:
1693:
1685:
1660:
1652:
1646:Lametasaurus
1644:
1636:
1602:
1597:Ceratosaurus
1595:
1575:
1567:
1560:
1552:
1545:
1529:Ceratosauria
1514:
1507:
1432:
1425:
1383:
1375:
1368:
1360:
1352:
1345:
1338:
1331:
1311:
1306:Podokesaurus
1304:
1297:
1290:
1282:
1258:
1250:
1244:
1243:
1236:
1229:
1222:
1215:
1208:
1201:
1194:
1189:Gojirasaurus
1187:
1182:Dracovenator
1180:
1172:
1165:
1153:Neotheropoda
1138:
1130:
1122:
1116:Guaibasaurus
1114:
1107:
1099:
1091:
1085:Chindesaurus
1083:
973:
964:
864:
860:
850:
838:. Retrieved
835:Science News
834:
824:
812:. Retrieved
786:. Retrieved
782:
776:
768:
721:
717:
711:
665:
661:Perijasaurus
659:
650:
649:
639:
634:
633:
624:
620:
604:
601:Neotheropoda
596:
587:performed a
584:
582:
572:
567:
560:
551:
547:
544:autapomorphy
541:
536:
535:
509:
500:
494:
476:
432:
419:
417:type species
414:
407:
389:
377:
338:
337:
301:
300:
299:
286:
281:
265:
264:
247:
246:
234:Neotheropoda
227:
214:
201:
188:
24:
18:
5164:Tachiraptor
5158:Wikispecies
5134:Tachiraptor
5058:Maniraptora
5047:Maniraptora
5028:Tototlmimus
4958:Anserimimus
4908:Garudimimus
4894:Beishanlong
4836:Valdoraptor
4777:Arkansaurus
4722:Xunmenglong
4599:Juravenator
4393:Tarbosaurus
4269:Nanuqsaurus
4238:Gorgosaurus
4154:Megaraptora
4140:Timurlengia
3872:Sciurumimus
3820:Aratasaurus
3688:Murusraptor
3610:Fukuiraptor
3573:Megaraptora
3455:Shaochilong
3427:Eocarcharia
3380:Megaraptora
3300:Lusovenator
3178:Alpkarakush
3035:Carnosauria
2886:Spinosaurus
2859:Siamosaurus
2759:Suchosaurus
2706:Iberospinus
2642:Magnosaurus
2621:Afrovenator
2594:Torvosaurus
2345:Chilesaurus
2158:Carnotaurus
2151:Caletodraco
2137:Abelisaurus
1963:Arcovenator
1810:Laevisuchus
1801:Noasaurinae
1703:Berthasaura
1674:Noasauridae
1604:Genyodectes
1569:Lukousaurus
1562:Jubbulpuria
1362:Panguraptor
1340:Coelophysis
1333:Camposaurus
1292:Panguraptor
1284:Dracoraptor
1260:Zupaysaurus
1245:Tachiraptor
1224:Sarcosaurus
1124:Nhandumirim
728:: 140–184.
651:Tachiraptor
635:Tachiraptor
625:Tachiraptor
621:Tachiraptor
605:Tachiraptor
597:Tachiraptor
585:Tachiraptor
573:Tachiraptor
568:Tachiraptor
552:Tachiraptor
548:Tachiraptor
537:Tachiraptor
532:Description
510:admirabilis
481:neotheropod
363:vertebrates
305:("thief of
302:Tachiraptor
248:Tachiraptor
25:Tachiraptor
5234:Categories
4986:Gallimimus
4915:Harpymimus
4860:Harpymimus
4481:Iliosuchus
4451:see below↓
4280:Alioramini
4097:Juratyrant
4083:Eotyrannus
4068:Bagaraatan
4020:Yutyrannus
3954:Coeluridae
3880:Vayuraptor
3726:see below↓
3702:Tratayenia
3695:Orkoraptor
3681:Megaraptor
3587:Aoniraptor
3520:Mapusaurus
3448:Sauroniops
3412:Altispinax
3365:Neovenator
3307:Siamraptor
3293:Datanglong
3260:Epanterias
3252:Antrodemus
3245:Allosaurus
3226:Allosauria
3049:Altispinax
3014:Gasosaurus
2926:see below↓
2802:Suchomimus
2403:Sinosaurus
2238:see below↓
2194:Viavenator
2144:Aucasaurus
2081:Ilokelesia
2052:Dahalokely
2007:Rajasaurus
1985:Indosaurus
1978:Genusaurus
1970:Dahalokely
1894:Indosuchus
1790:Limusaurus
1749:Ligabueino
1742:Kiyacursor
1734:Genusaurus
1718:Dahalokely
1638:Betasuchus
1509:Dornraptor
1409:see below↓
1385:Segisaurus
1238:Sinosaurus
1101:Eodromaeus
1030:see below↓
978:Dinosauria
960:Sauropsida
840:8 December
814:8 December
788:8 December
674:References
564:talus bone
442:Hettangian
356:generalist
208:Saurischia
195:Dinosauria
112:Hettangian
5240:Theropods
5149:Q18204180
5062:includes
5007:Rativates
5000:Qiupalong
4700:Scipionyx
4693:Mirischia
4366:Lythronax
4358:Labocania
4289:Alioramus
3942:Labocania
3660:Aerosteon
3206:Sinraptor
3110:Erectopus
2845:Irritator
2430:Orionides
2315:Tetanurae
2302:Tetanurae
2229:Tetanurae
2165:Elemgasem
2088:Llukalkan
2067:Elemgasem
1824:Noasaurus
1687:Afromimus
1495:Averostra
1482:Averostra
1400:Averostra
1252:Velocipes
1071:Theropoda
1058:Theropoda
1021:Theropoda
946:Kingdom:
930:Theropoda
889:251501541
609:Averostra
593:phylogeny
589:cladistic
579:Phylogeny
491:Etymology
404:herbivore
385:Venezuela
369:Discovery
359:predators
330:Venezuela
318:dinosaurs
259:Species:
221:Theropoda
168:Kingdom:
162:Eukaryota
5178:47459440
5143:Wikidata
4591:Coelurus
3999:Kileskus
3992:Guanlong
3964:Coelurus
3857:Gualicho
3357:Gualicho
2738:Baryonyx
2074:Guemesia
1901:Kryptops
1662:Ozraptor
1299:Pendraig
954:Chordata
952:Phylum:
948:Animalia
760:26064540
526:La Grita
469:holotype
400:Seboruco
396:La Grita
322:Jurassic
315:theropod
309:") is a
182:Chordata
178:Phylum:
172:Animalia
158:Domain:
130:holotype
5204:6158268
5191:8662438
4132:Timimus
3894:Zuolong
3624:Rapator
3527:Meraxes
2852:Oxalaia
2173:Koleken
1196:Lepidus
958:Class:
869:Bibcode
751:4448901
730:Bibcode
656:Pangaea
505:. The
485:ischium
454:Pangaea
392:Táchira
352:bipedal
348:ischium
334:species
324:period
307:Táchira
284:†
263:†
240:Genus:
138:ischium
105:↓
5217:332598
4800:Hexing
4488:Kakuru
4076:Dilong
2117:Thanos
2095:Niebla
1915:Rugops
1908:Kurupi
887:
758:
748:
556:fibula
502:raptor
462:zircon
132:right
5064:birds
4104:Moros
3631:Siats
3372:Siats
3314:Siats
974:Clade
965:Clade
885:S2CID
724:(2).
497:Latin
473:tibia
344:tibia
311:genus
228:Clade
215:Clade
202:Clade
189:Clade
134:tibia
5186:GBIF
5056:see
3674:Maip
1132:Tawa
1007:see
842:2014
816:2014
790:2014
756:PMID
712:stem
398:and
346:and
44:Preęž’
5173:EoL
877:doi
746:PMC
738:doi
524:of
452:of
383:in
328:of
5236::
5214::
5201::
5188::
5175::
5160::
5145::
976::
967::
883:.
875:.
865:42
863:.
859:.
833:.
798:^
781:.
754:.
744:.
736:.
720:.
716:.
682:^
642:.
430:.
336:,
230::
217::
204::
191::
94:Pg
38:Ma
33:,
4926:?
4918:?
4831:?
4795:?
4787:?
4755:?
4703:?
4674:?
4659:?
4594:?
4361:?
4346:?
4195:?
4157:?
4135:?
4071:?
3974:?
3956:?
3945:?
3875:?
3860:?
3852:?
3837:?
3641:?
3633:?
3605:?
3597:?
3575:?
3415:?
3383:?
3375:?
3360:?
3352:?
3317:?
3271:?
3263:?
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3153:?
3128:?
3120:?
3075:?
3067:?
3059:?
3025:?
3017:?
2833:?
2770:?
2762:?
2716:?
2694:?
2659:?
2538:?
2503:?
2406:?
2363:?
2348:?
2340:?
2332:?
2168:?
2105:?
2055:?
2030:?
2002:?
1973:?
1882:?
1737:?
1729:?
1721:?
1713:?
1698:?
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1665:?
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1572:?
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1519:?
1380:?
1365:?
1357:?
1287:?
1255:?
1177:?
1143:?
1135:?
1127:?
1119:?
1104:?
1096:?
1088:?
922:e
915:t
908:v
891:.
879::
871::
844:.
818:.
792:.
762:.
740::
732::
722:1
244:â€
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:ęž’
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