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Amborella

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498: 471: 58: 486: 510: 6475: 3214: 40: 3183: 1088:), stability that has permitted the continued survival of tropical forests on New Caledonia. In contrast, drought conditions dominated the Australian climate towards the end of the Tertiary. Current threats to biodiversity in New Caledonia include fires, mining, agriculture, invasion by introduced species, urbanization and global warming. The importance of conserving 1045:
mitochondrial genome revealed that for every gene of its own origin, it contains about six versions from the genomes of an assortment of the plants and algae growing with or upon it. The evolutionary and physiological significance of this is not as yet clear, nor in particular is it clear whether the
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directed downwards. Staminate flowers are approximately 4 to 5 mm in diameter, with 6 to 15 tepals. These flowers bear 10 to 21 spirally arranged stamens, which become progressively smaller toward the center. The innermost may be sterile, amounting to staminodes. The stamens bear triangular
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to the other flowering plants. Comparing characteristics of this basal angiosperm, other flowering plants and fossils may provide clues about how flowers first appeared—what Darwin called the "abominable mystery". This position is consistent with a number of conservative characteristics of its
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Rice, D. W.; Alverson, A. J.; Richardson, A. O.; Young, G. J.; Sanchez-Puerta, M. V.; Munzinger, J.; Barry, K.; Boore, J. L.; Zhang, Y.; dePamphilis, C. W.; Knox, E. B.; Palmer, J. D. (19 December 2013). "Horizontal Transfer of Entire Genomes via Mitochondrial Fusion in the Angiosperm
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develops. Plants may change from one reproductive morphology to the other. In one study, seven cuttings from a staminate plant produced, as expected, staminate flowers at their first flowering, but three of the seven produced carpellate flowers at their second flowering.
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anthers on short broad filaments. An anther consists of four pollen sacs, two on each side, with a small sterile central connective. The anthers have connective tips with small bumps and may be covered with secretions. These features suggest that, as with other
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of the flowering plant lineage. That is, the Amborellaceae represent a line of flowering plants that diverged very early on (more than 130 million years ago) from all the other extant species of flowering plants, and, among extant flowering plants, is the
1836:; Carlson, J. E.; Farmerie, W. G.; Wall, P. K.; Ilut, D. C.; Solow, T. M.; Mueller, L. A.; Landherr, L. L.; Hu, Y.; Buzgo, M.; Kim, S.; Yoo, M.-J.; Frohlich, M. W.; Perl-Treves, R.; Schlarbaum, S. E.; Zhang, X.; Tanksley, S. D.; Oppenheimer, D. G.; 2345:
Zuccolo, A.; Bowers, J. E.; Estill, J. C.; Xiong, Z.; Luo, M.; Sebastian, A.; Goicoechea, J. L.; Collura, K.; Yu, Y.; Jiao, Y.; Duarte, J.; Tang, H.; Ayyampalayam, S.; Rounsley, S.; Kudrna, D.; Paterson, A. H.; Pires, J. C.; Chanderbali, A.;
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Thien, L. B.; Sage, T. L.; Jaffré, T.; Bernhardt, P.; Pontieri, V.; Weston, P. H.; Malloch, D.; Azuma, H.; Graham, S. W.; McPherson, M. A.; Rai, H. S.; Sage, R. F. & Dupre, J.-L. (2003). "The Population Structure and Floral Biology of
967:: one theory is that the Amborellaceae alone are the monophyletic sister to the extant angiosperms; another proposes that the Amborellaceae and Nymphaeales form a clade that is the sister group to all other extant angiosperms. 2199:, p. 55. "La disparition d'Amborella trichopoda impliquerait donc la disparition d'un genre, d'une famille et d'un ordre entier, ainsi que le seul témoin d'au moins 140 millions d'années d'histoire évolutive." 963:
branched off from the main lineage of angiosperms before the ancestors of any other living angiosperms. There is however some uncertainty about the relationship between the Amborellaceae and the
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would imply the disappearance of a genus, a family and an entire order, as well as the only witness to at least 140 million years of evolutionary history." Conservation strategies targeted on
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is apparently basal among the flowering plants, the features of early flowering plants can be inferred by comparing derived traits shared by the main angiosperm lineage but not present in
110: 2292:; Albert, V. A.; Leebens-Mack, J.; Palmer, J. D.; Wing, Rod A.; dePamphilis, Claude W.; Ma, Hong; Carlson, John E.; Altman, Naomi; Kim, Sangtae; Wall, P. K.; Zuccolo, A. & 114: 2475: 1926:(2013). "Angiosperm Phylogeny: A Framework for Studies of Genome Evolution". In Leitch, Ilia J.; Greilhuber, Johann; Doležel, Jaroslav & Wendel, Jonathan F. (eds.). 1011:
characteristic of most flowering plants. The genes responsible for floral traits like scent and colors in other angiosperms, have yet to be found. Further, the female
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plant in the wild, is commonly in intimate contact with shade- and moisture-dependent organisms such as algae, lichens and mosses. In those circumstances, some
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of angiosperms. In systematics the term "basal" describes a lineage that diverges near the base of a phylogeny, and thus earlier than other lineages. Since
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L. Watson and M.J. Dallwitz (1992 onwards). The families of flowering plants: descriptions, illustrations, identification, information retrieval.
350:), but not both. At any one time, a dioecious plant produces only functionally staminate or functionally carpellate flowers. Staminate ("male") 2073: 7393: 7158: 6871: 6646: 3250: 2354:; Albert, V. A.; Ma, H.; Mandoli, D.; Banks, J.; Carlson, J. E.; Tomkins, J.; dePamphilis, C. W.; Wing, R. A. & Leebens-Mack, J. (2011). 699:
arranged in spirals on an elongated, cone-like receptacle rather than the small numbers of parts in distinct whorls of more derived flowers.
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and such associated species is not surprising in principle, but the scale of such transfer has caused considerable surprise. Sequencing the
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Because of its evolutionary position at the base of the flowering plant clade, there was support for sequencing the complete genome of
497: 417: in) in diameter, with 7 or 8 tepals. There are 1 to 3 (or rarely 0) well-differentiated staminodes and a spiral of 4 to 8 free ( 7341: 7080: 6806: 6581: 3298: 2550: 959:
to all other angiosperms, and shows the gymnosperms as a monophyletic group sister to the angiosperms. It supports the theory that
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to serve as a reference for evolutionary studies. In 2010, the US National Science Foundation began a genome sequencing effort in
7506: 7380: 7145: 7093: 6819: 6594: 509: 305:. The leaves are two-ranked, with distinctly serrated or rippled margins, and about 8 to 10 centimetres (3 to 4 inches) long. 1967:
Biodiversité, origine et évolution des Cunoniaceae : implications pour la conservation de la flore de Nouvelle-Calédonie
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can be seen at the tip of the fruit. The skin is papery, surrounding a thin fleshy layer containing a red juice. The inner
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Posluszny, U.; Tomlinson, P.B. (2003), "Aspects of inflorescence and floral development in the putative basal angiosperm
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The islands of New Caledonia are a biodiversity hot-spot, preserving many early diverging lineages of plants, of which
1801:"An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG IV", 7511: 3243: 1656: 1226:"An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG III" 7468: 7398: 1588:
Floyd, S.K. & Friedman, W.E. (2001). "Developmental evolution of endosperm in basal angiosperms: evidence from
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recognized this family, but left it unplaced at order rank due to uncertainty about its relationship to the family
445:(approximately 5 to 7 mm long and 5 mm wide) borne on a short (1 to 2 mm) stalk. The remains of the 7163: 6876: 6651: 470: 7119: 6845: 6620: 2495: 1181: 1133: 7202: 6759: 6745: 6938: 378:, with up to three orders of branching, each branch being terminated by a flower. Each flower is subtended by 6977: 6677: 3277: 2529: 2435: 2412: 1891: 6478: 3303: 3293: 3236: 2555: 2545: 226: 1290: 7289: 7033: 7028: 1191: 1143: 1046:
horizontal gene transfer has anything to do with the apparent stability and conservatism of the species.
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Endress, P. K. & Igersheim, Anton (2000). "The Reproductive Structures of the Basal Angiosperm
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Carlquist, S. J. & Schneider, E. L. (2001). "Vegetative anatomy of the New Caledonian endemic
7019: 7434: 2465: 7280: 7189: 6863: 6495: 990: 737: 260: 6638: 485: 7429: 7184: 7137: 2470: 994: 663: 342:. This means that each plant produces either male flowers (meaning that they have functional 174: 7421: 7372: 7176: 6889: 6664: 1991:
Drew, B. T.; Ruhfel, B. R.; Smith, S. A.; Moore, M. J.; Briggs, B. G.; Gitzendanner, M. A.;
689:" (i.e. ancestral) floral traits in angiosperms, accepted until relatively recently, is the 642:, the Amborellaceae comprise the monotypic order Amborellales at the base of the angiosperm 354:
flowers do not have carpels, whereas the carpellate ("female") flowers have non-functional "
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Endress, P.K.; Igersheim, A. (2000). "The reproductive structures of the basal angiosperm
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is but one. This preservation has been ascribed to climate stability during and since the
271:, meaning it was the earliest group to evolve separately from all other flowering plants. 8: 6533: 1271:(in French). Vol. 11. Paris: Muséum National d’Histoire Naturelle. pp. 157–160. 3228: 2149: 1700: 1613: 7455: 7233: 6959: 6698: 6318: 5536: 5428: 5052: 3539: 3283: 3201: 2535: 2386: 2355: 2328: 2297: 2277: 2248: 2240: 2179: 1868: 1841: 1784: 1753: 1712: 1645: 1625: 1570: 1494: 1444: 1375: 702:
In a study designed to clarify relationships between well-studied model plants such as
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Große-Veldmann, B.; Korotkova, N.; Reinken, B.; Lobin, W. & Barthlott, W. (2011).
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species are recommended, both preserving a diversity of habitats in New Caledonia and
7416: 7302: 7171: 7041: 6912: 6884: 6767: 6659: 6542: 6275: 6074: 5626: 5486: 5415: 5338: 5219: 5105: 5022: 4387: 3378: 3314: 3195: 3010: 2391: 2333: 2289: 2252: 2171: 2102: 2057: 2022: 1996: 1939: 1923: 1873: 1833: 1652: 1470: 1398: 1242: 1225: 1020: 431: 2281: 1757: 1716: 1574: 1498: 1379: 978:, and the draft genome sequence was posted on the project website in December 2013. 7460: 7307: 7238: 6964: 6772: 6703: 6547: 6159: 6149: 6109: 6089: 5865: 5795: 5770: 5717: 5491: 5481: 5244: 5181: 5125: 5100: 4846: 4751: 4718: 4508: 4422: 4417: 4392: 4293: 4255: 4163: 3827: 3690: 3665: 3660: 3453: 3370: 3207: 2792: 2621: 2591: 2381: 2371: 2351: 2323: 2313: 2293: 2269: 2230: 2183: 2161: 2153: 2094: 2049: 2012: 1992: 1931: 1919: 1863: 1853: 1837: 1810: 1780: 1743: 1704: 1629: 1617: 1562: 1532: 1486: 1466: 1436: 1365: 1357: 1247: 1237: 1097: 534: 7294: 6899: 2456:
Ancient plant provides clues to evolutionary mystery (National Science Foundation)
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Dahlgren, R.M.T. (1980). "A revised system of classification of the angiosperms".
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Rudall, Paula (2006). "How many nuclei make an embryo sac in flowering plants?".
1195: 1147: 777: 588: 316: 268: 95: 1935: 1842:"Floral gene resources from basal angiosperms for comparative genomics research" 7004: 6907: 6432: 6422: 6404: 6376: 6338: 6328: 6280: 6247: 6237: 6232: 6154: 6104: 6051: 6036: 5921: 5835: 5775: 5765: 5669: 5571: 5506: 5496: 5456: 5438: 5239: 5166: 4954: 4916: 4871: 4806: 4781: 4756: 4700: 4614: 4382: 4329: 4265: 4122: 4014: 3991: 3953: 3930: 3920: 3723: 3585: 3524: 3388: 3118: 3075: 3020: 2692: 2616: 1008: 765: 635: 328: 82: 1748: 1687:
Thorne, Robert F. (1992). "Classification and geography of flowering plants".
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has a mixed pollination system, relying on both insect pollinators and wind.
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Water lily genome expands picture of the early evolution of flowering plants
2017: 2000: 7215: 6951: 6690: 6353: 6348: 6333: 6323: 6262: 6144: 6129: 6099: 6056: 6046: 6000: 5977: 5967: 5939: 5840: 5820: 5790: 5732: 5679: 5601: 5556: 5516: 5471: 5461: 5451: 5343: 5312: 5277: 5262: 5176: 5171: 5077: 4831: 4826: 4791: 4771: 4746: 4733: 4723: 4695: 4652: 4647: 4604: 4571: 4556: 4518: 4495: 4432: 4427: 4412: 4377: 4359: 4311: 4250: 4245: 4237: 4112: 4072: 4006: 3986: 3976: 3968: 3935: 3895: 3842: 3837: 3832: 3774: 3769: 3733: 3650: 3529: 3486: 3383: 3360: 3324: 3065: 2920: 2900: 2850: 2782: 2750: 2745: 2698: 2395: 2347: 2337: 2175: 2106: 2061: 2026: 1877: 1267:
Jérémie, J. (1982). "Amborellacées". In A. Aubréville; J. F. Leroy (eds.).
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in the southwest Pacific Ocean. The genus is the only member of the family
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Friedman, W. E. (2009), "The meaning of Darwin's "abominable mystery"",
678:. These traits are presumed to have evolved after the divergence of the 7346: 7085: 6811: 6586: 6391: 6381: 6358: 6242: 6179: 6114: 6041: 6018: 5944: 5890: 5800: 5654: 5641: 5616: 5596: 5348: 5330: 5302: 5272: 5267: 5199: 5158: 5135: 5072: 5062: 5012: 4997: 4926: 4881: 4866: 4776: 4677: 4657: 4619: 4599: 4321: 4227: 4107: 4092: 4082: 3812: 3761: 3700: 3655: 3506: 3463: 3422: 3413: 3113: 3090: 3015: 2968: 2963: 2943: 2905: 2821: 2704: 2643: 2635: 2098: 1708: 1448: 1252: 1030: 877: 295: 212: 39: 7359: 6837: 2166: 1815: 1370: 1083: 7328: 7111: 7067: 6793: 6612: 6568: 6399: 6368: 6300: 6169: 6028: 6010: 5962: 5931: 5860: 5805: 5742: 5727: 5712: 5649: 5631: 5566: 5541: 5378: 5282: 5234: 5214: 4861: 4766: 4741: 4212: 4204: 4102: 4067: 4044: 4034: 4019: 3958: 3915: 3875: 3852: 3622: 3554: 3445: 3143: 3123: 3080: 3070: 2787: 2648: 2235: 2214: 2001:"Another Look at the Root of the Angiosperms Reveals a Familiar Tale" 741: 643: 450: 446: 426: 422: 418: 355: 339: 298: 205: 20: 7124: 6489: 1536: 1440: 7442: 7259: 7210: 6998: 6946: 6724: 6685: 6512: 6252: 6174: 6084: 6066: 5815: 5752: 5704: 5694: 5551: 5368: 5254: 5229: 5191: 5032: 4901: 4886: 4672: 4662: 4637: 4594: 4561: 4470: 4397: 4194: 4177: 4029: 3945: 3799: 3614: 3604: 3516: 3138: 3047: 3042: 3030: 2978: 2948: 2872: 2769: 2735: 2710: 2653: 2360:
genome sheds light on the evolution of angiosperm genome structure"
2273: 1566: 1490: 1288: 1078: 892: 839: 725: 720: 691: 390:. The tepals typically are arranged in a spiral, but sometimes are 383: 324: 222: 7333: 7072: 6850: 6798: 6573: 3182: 1295:— Cultivation of the most ancestral angiosperm in botanic gardens" 1124: 6972: 6625: 6214: 5880: 4979: 4934: 4629: 4581: 4548: 4462: 4186: 4132: 3640: 3105: 3085: 2935: 2925: 2915: 2910: 2797: 2432: 2409: 1895: 1172: 302: 242: 7320: 7059: 6785: 6560: 1732:"An updated phylogenetic classification of the flowering plants" 951:
This hypothesized relationship of the extant seed plants places
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The evolution and classification of flowering plants (2nd ed.)
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has been dramatically stated by Pillon: "The disappearance of
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Typically, 1 to 3 carpels per flower develop into fruit. The
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of foliage leaves. The inflorescences have been described as
371: 312: 283: 215: 208: 69: 1840:; Ma, H.; dePamphilis, C. W. & Leebens-Mack, H. (2005). 1518: 2219:
Baill., a new morphological type of vesselless dicotyledon"
2130: 1831: 733: 461:. The embryo is small and surrounded by copious endosperm. 458: 331:
are absent. Xylem of this form has long been regarded as a
287: 219: 2443: 1973:(PhD) (in French and English). University of New Caledonia 1647:
An integrated system of classification of flowering plants
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Diagram of a female flower with 5 carpels and 2 staminodes
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Megan Scudellari. Genomes Gone Wild, January 1, 2014 |
434:, there is a high degree of developmental plasticity. 2466:
article with detailed photos of plants in cultivation
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genome: an evolutionary reference for plant biology"
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physiology and morphology; for example, the wood of
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blossom model. This envisions flowers with numerous
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is the only genus in the family Amborellaceae. The
989:is of great interest to plant systematists because 421:) carpels. Carpels bear green ovaries; they lack a 397:
Carpellate flowers are roughly 3 to 4 millimetres (
1644: 1343: 1464: 1460: 1458: 7488: 2259: 1548: 366:The small, creamy white flowers are arranged in 4154: 1455: 1057:is typically dioecious, but has been known to 993:analyses consistently place it at or near the 3244: 2496: 1587: 1557:. Current Perspectives on Basal Angiosperms. 1269:Flore de La Nouvelle-Calédonie et Dépendances 358:", structures resembling stamens in which no 346:) or female flowers (flowers with functional 3576: 1918: 1514: 1512: 1510: 1508: 1473:(2004). "Floral Developmental Morphology of 315:tissue that differs from that of most other 2212: 728:, and more derived angiosperms (eudicots), 3251: 3237: 2503: 2489: 1299:The Journal of Botanic Garden Horticulture 38: 2385: 2375: 2327: 2317: 2234: 2213:Bailey, I.W. & Swamy, B.G.L. (1948). 2165: 2016: 1959: 1957: 1955: 1914: 1912: 1867: 1857: 1827: 1825: 1814: 1747: 1671: 1642: 1505: 1369: 1337: 1321: 1319: 1251: 1241: 2039: 2033: 1803:Botanical Journal of the Linnean Society 1796: 1794: 1773:Botanical Journal of the Linnean Society 1770: 1386: 1230:Botanical Journal of the Linnean Society 1220: 1218: 1216: 1019:is even more reduced than normal female 614: 290:up to 8 metres (26 feet) high. It bears 5692: 2410:The Amborella Genome Sequencing Project 2266:International Journal of Plant Sciences 1676:. Bronx, NY: New York Botanical Garden. 1651:. New York: Columbia University Press. 1555:International Journal of Plant Sciences 1542: 1479:International Journal of Plant Sciences 1429:Annals of the Missouri Botanical Garden 1417: 1415: 1325: 1266: 982:Genomic and evolutionary considerations 425:. They contain a single ovule with the 7489: 4461: 2196: 2084: 1963: 1952: 1909: 1822: 1729: 1686: 1392: 1316: 263:analyses consistently place it as the 6494: 6493: 4450: 3232: 2484: 1791: 1581: 1213: 7409:1628de46-45df-41e8-9ae1-692acd3971da 6978:813E27BC-21AF-4BA1-90AE-1500223FF841 6926:17AE28AC-1126-AF08-487D-7C1DE546FDE2 2460:National Tropical Botanical Garden ( 1984: 1412: 1284: 1282: 1280: 1278: 1200:Australian National Botanic Gardens 1152:Australian National Botanic Gardens 13: 7517:Taxa named by Henri Ernest Baillon 7203:urn:lsid:ipni.org:names:77126703-1 2206: 1785:10.1111/j.1095-8339.1980.tb01661.x 537:, of 1981, classified the family: 14: 7533: 2444:Description language for taxonomy 2403: 1275: 503:Buds and staminate (male) flowers 6939:urn:lsid:ipni.org:names:581745-1 6474: 6473: 3213: 3212: 3181: 2471:Nova "First Flower" (transcript) 1243:10.1111/j.1095-8339.2009.00996.x 685:One early 20th century idea of " 508: 496: 484: 469: 56: 6678:urn:lsid:ipni.org:names:27047-1 2223:Journal of the Arnold Arboretum 2124: 2113: 2078: 2067: 1928:Plant Genome Diversity Volume 2 1884: 1764: 1723: 1680: 1665: 1636: 1602:Plant Systematics and Evolution 1182:International Plant Names Index 1134:International Plant Names Index 1068: 382:. The bracts transition into a 7507:Endemic flora of New Caledonia 2268:. 161(supplement): S237–S248. 1260: 1165: 1117: 274: 255:is of great interest to plant 1: 2350:; Chamala, S.; Barbazuk, B.; 1395:The Morphology of Angiosperms 1110: 335:feature of flowering plants. 708:, and the basal angiosperms 649: 7: 7497:Monotypic angiosperm genera 1936:10.1007/978-3-7091-1160-4_1 1930:. Springer. pp. 1–11. 1892:"Amborella Genome Database" 1192:Harvard University Herbaria 1144:Harvard University Herbaria 744:shown below was generated. 523: 10: 7538: 2042:American Journal of Botany 1730:Thorne, Robert F. (1992). 1525:Canadian Journal of Botany 1188:Royal Botanic Gardens, Kew 1140:Royal Botanic Gardens, Kew 1049: 528: 18: 7249: 6988: 6714: 6502: 6469: 6431: 6413: 6390: 6367: 6299: 6261: 6213: 6202: 6065: 6027: 6009: 5991: 5953: 5930: 5912: 5889: 5878: 5751: 5703: 5640: 5437: 5414: 5403: 5329: 5311: 5253: 5190: 5157: 5134: 5086: 4988: 4977: 4925: 4732: 4709: 4686: 4628: 4580: 4547: 4494: 4483: 4479: 4358: 4347: 4320: 4302: 4284: 4236: 4203: 4185: 4176: 4145: 4058: 4005: 3967: 3944: 3861: 3798: 3760: 3732: 3714: 3631: 3613: 3598: 3567: 3515: 3472: 3444: 3421: 3412: 3401: 3369: 3341: 3323: 3312: 3272: 3190: 3179: 3104: 3056: 3038: 3029: 3006: 2995: 2934: 2881: 2868: 2859: 2846: 2835: 2817: 2806: 2778: 2767: 2726: 2676: 2667: 2634: 2630: 2612: 2601: 2575: 2564: 2524: 1749:10.5642/aliso.19921302.08 1332:(2nd ed.). Elsevier. 890: 875: 860: 853: 836: 829: 812: 805: 788: 781: 763: 756: 457:and surrounds the single 180: 173: 53:Scientific classification 51: 46: 37: 30: 7512:Plants described in 1873 2377:10.1186/gb-2011-12-5-r48 2356:"A physical map for the 1035:horizontal gene transfer 601:Subclass Magnoliideae , 19:Not to be confused with 7223:Watson & Dallwitz: 2319:10.1186/gb-2008-9-3-402 2158:10.1126/science.1246275 738:expressed sequence tags 598:Superorder Magnolianae 574:Subclass Magnoliideae 571:Superorder Magnolianae 550:Division Magnoliophyta 1672:Cronquist, A. (1988). 1643:Cronquist, A. (1981). 1397:. London: Hutchinson. 1326:Simpson, M.G. (2010). 1021:angiosperm gametophyte 991:molecular phylogenetic 740:for floral genes, the 261:molecular phylogenetic 241:and contains a single 7430:Paleobiology Database 7185:Paleobiology Database 2476:NCBI Taxonomy Browser 2018:10.1093/sysbio/syt108 1859:10.1186/1471-2229-5-5 1622:10.1007/s006060170026 1393:Sporne, K.R. (1974). 1362:10.1353/psc.2001.0020 630:. In the more recent 615:Modern classification 604:Class Magnoliopsida . 547:Class Magnoliopsida 544:Subclass Magnoliidae 6900:Amborella_trichopoda 6760:amborella-trichopoda 6746:Amborella trichopoda 6716:Amborella trichopoda 2358:Amborella trichopoda 2262:Amborella trichopoda 2217:Amborella trichopoda 1551:Amborella trichopoda 1521:Amborella trichopoda 1475:Amborella trichopoda 1425:Amborella trichopoda 1346:Amborella trichopoda 1293:Amborella trichopoda 1236:(2): 105–121. 2009. 1127:Amborella trichopoda 1094:Amborella trichopoda 972:Amborella trichopoda 705:Arabidopsis thaliana 660:Amborella trichopoda 577:Class Magnoliopsida 562:Thorne system (1992) 478:Berkeley, California 386:of undifferentiated 248:Amborella trichopoda 225:to the main island, 184:Amborella trichopoda 2150:2013Sci...342.1468R 2144:(6165): 1468–1473. 2054:10.3732/ajb.0800150 1964:Pillon, Y. (2008). 1701:1992BotRv..58..225T 1614:2001PSyEv.228..153F 730:chloroplast genomes 6319:Pentaphragmataceae 5537:Stegnospermataceae 5429:Berberidopsidaceae 5053:Pentadiplandraceae 3540:Atherospermataceae 2464:, United States), 2449:2007-01-03 at the 2438:2007-01-03 at the 2427:2005-02-12 at the 2415:2020-01-29 at the 2312:(3): 402.1–402.6. 2264:(Amborellaceae)". 2099:10.1002/bies.20488 2005:Systematic Biology 1709:10.1007/BF02858611 1596:(Nymphaceae), and 1553:(Amborellaceae)". 1523:(Amborellaceae)", 1477:(Amborellaceae)". 1427:(Amborellaceae)". 568:Order Magnoliales 394:at the periphery. 166:A. trichopoda 7484: 7483: 7417:Open Tree of Life 7172:Open Tree of Life 6885:Open Tree of Life 6660:Open Tree of Life 6496:Taxon identifiers 6487: 6486: 6465: 6464: 6461: 6460: 6457: 6456: 6453: 6452: 6449: 6448: 6276:Cardiopteridaceae 6198: 6197: 6075:Plocospermataceae 5874: 5873: 5688: 5687: 5627:Anacampserotaceae 5487:Ancistrocladaceae 5416:Berberidopsidales 5399: 5398: 5395: 5394: 5391: 5390: 5387: 5386: 5339:Biebersteiniaceae 5220:Sphaerosepalaceae 5106:Strasburgeriaceae 5023:Setchellanthaceae 4973: 4972: 4446: 4445: 4388:Cercidiphyllaceae 4343: 4342: 4172: 4171: 4141: 4140: 3594: 3593: 3563: 3562: 3397: 3396: 3379:Austrobaileyaceae 3226: 3225: 3196:Basal angiosperms 3177: 3176: 3173: 3172: 3169: 3168: 3165: 3164: 3161: 3160: 3157: 3156: 3011:Berberidopsidales 2991: 2990: 2987: 2986: 2763: 2762: 2759: 2758: 2093:(11): 1067–1071. 1945:978-3-7091-1160-4 1846:BMC Plant Biology 1816:10.1111/boj.12385 1809:(1): 1–20, 2016, 1592:(Amborellaceae), 1561:(S6): S237–S248. 1404:978-0-09-120611-6 1329:Plant Systematics 1086:million years ago 1084:66 to 3 948: 947: 939: 938: 930: 929: 921: 920: 912: 911: 903: 902: 432:basal angiosperms 197: 196: 155: 137: 121: 7529: 7522:Dioecious plants 7477: 7476: 7464: 7463: 7451: 7450: 7438: 7437: 7425: 7424: 7412: 7411: 7402: 7401: 7389: 7388: 7376: 7375: 7363: 7362: 7350: 7349: 7337: 7336: 7324: 7323: 7311: 7310: 7298: 7297: 7285: 7284: 7283: 7270: 7269: 7268: 7242: 7241: 7229: 7228: 7219: 7218: 7206: 7205: 7193: 7192: 7180: 7179: 7167: 7166: 7154: 7153: 7141: 7140: 7128: 7127: 7115: 7114: 7102: 7101: 7089: 7088: 7076: 7075: 7063: 7062: 7050: 7049: 7037: 7036: 7024: 7023: 7022: 7009: 7008: 7007: 6981: 6980: 6968: 6967: 6955: 6954: 6942: 6941: 6929: 6928: 6916: 6915: 6903: 6902: 6893: 6892: 6880: 6879: 6867: 6866: 6854: 6853: 6841: 6840: 6828: 6827: 6815: 6814: 6802: 6801: 6789: 6788: 6776: 6775: 6763: 6762: 6750: 6749: 6748: 6735: 6734: 6733: 6707: 6706: 6694: 6693: 6681: 6680: 6668: 6667: 6655: 6654: 6642: 6641: 6629: 6628: 6616: 6615: 6603: 6602: 6590: 6589: 6577: 6576: 6564: 6563: 6551: 6550: 6538: 6537: 6536: 6523: 6522: 6521: 6491: 6490: 6477: 6476: 6211: 6210: 6160:Thomandersiaceae 6150:Lentibulariaceae 6110:Scrophulariaceae 6090:Tetrachondraceae 5887: 5886: 5876: 5875: 5866:Mitrastemonaceae 5796:Pentaphylacaceae 5771:Tetrameristaceae 5718:Hydrostachyaceae 5701: 5700: 5690: 5689: 5492:Rhabdodendraceae 5482:Dioncophyllaceae 5412: 5411: 5245:Dipterocarpaceae 5182:Dipentodontaceae 5126:Crossosomataceae 5101:Geissolomataceae 4986: 4985: 4847:Chrysobalanaceae 4752:Ctenolophonaceae 4719:Lepidobotryaceae 4509:Anisophylleaceae 4492: 4491: 4481: 4480: 4459: 4458: 4448: 4447: 4423:Tetracarpaeaceae 4418:Aphanopetalaceae 4393:Daphniphyllaceae 4356: 4355: 4345: 4344: 4294:Trochodendraceae 4256:Circaeasteraceae 4183: 4182: 4164:Ceratophyllaceae 4152: 4151: 3828:Alstroemeriaceae 3691:Potamogetonaceae 3666:Scheuchzeriaceae 3661:Hydrocharitaceae 3611: 3610: 3574: 3573: 3454:Aristolochiaceae 3419: 3418: 3410: 3409: 3371:Austrobaileyales 3321: 3320: 3317: 3315:Basalangiosperms 3299:Acrogymnospermae 3261:flowering plants 3253: 3246: 3239: 3230: 3229: 3216: 3215: 3185: 3036: 3035: 3004: 3003: 2866: 2865: 2844: 2843: 2815: 2814: 2776: 2775: 2674: 2673: 2632: 2631: 2610: 2609: 2592:Austrobaileyales 2573: 2572: 2551:Acrogymnospermae 2513:flowering plants 2505: 2498: 2491: 2482: 2481: 2399: 2389: 2379: 2341: 2331: 2321: 2285: 2256: 2238: 2236:10.5962/p.324625 2200: 2194: 2188: 2187: 2169: 2128: 2122: 2117: 2111: 2110: 2082: 2076: 2071: 2065: 2064: 2037: 2031: 2030: 2020: 1988: 1982: 1981: 1979: 1978: 1972: 1961: 1950: 1949: 1916: 1907: 1906: 1904: 1903: 1894:. Archived from 1888: 1882: 1881: 1871: 1861: 1829: 1820: 1819: 1818: 1798: 1789: 1788: 1768: 1762: 1761: 1751: 1727: 1721: 1720: 1689:Botanical Review 1684: 1678: 1677: 1669: 1663: 1662: 1650: 1640: 1634: 1633: 1608:(3–4): 153–169. 1600:(Illiciaceae)". 1585: 1579: 1578: 1546: 1540: 1539: 1516: 1503: 1502: 1462: 1453: 1452: 1419: 1410: 1408: 1390: 1384: 1383: 1373: 1341: 1335: 1333: 1323: 1314: 1313: 1311: 1310: 1286: 1273: 1272: 1264: 1258: 1257: 1255: 1245: 1222: 1211: 1210: 1208: 1206: 1169: 1163: 1162: 1160: 1158: 1121: 1107:in cultivation. 1087: 1061:in cultivation. 856: 855: 832: 831: 808: 807: 784: 783: 759: 758: 753:seed plants 748: 747: 718:(Nymphaeaceae), 654:Currently plant 535:Cronquist system 512: 500: 488: 473: 441:is an ovoid red 416: 415: 411: 406: 405: 401: 317:flowering plants 269:flowering plants 186: 151: 133: 109: 61: 60: 42: 28: 27: 16:Species of shrub 7537: 7536: 7532: 7531: 7530: 7528: 7527: 7526: 7487: 7486: 7485: 7480: 7472: 7467: 7459: 7454: 7446: 7441: 7433: 7428: 7420: 7415: 7407: 7405: 7397: 7392: 7384: 7379: 7371: 7366: 7358: 7353: 7345: 7340: 7332: 7327: 7319: 7314: 7306: 7301: 7293: 7288: 7279: 7278: 7273: 7264: 7263: 7258: 7245: 7237: 7232: 7224: 7222: 7214: 7209: 7201: 7196: 7188: 7183: 7175: 7170: 7162: 7157: 7149: 7144: 7136: 7131: 7123: 7118: 7110: 7105: 7097: 7092: 7084: 7079: 7071: 7066: 7058: 7053: 7045: 7040: 7032: 7027: 7018: 7017: 7012: 7003: 7002: 6997: 6984: 6976: 6971: 6963: 6958: 6950: 6945: 6937: 6932: 6924: 6919: 6911: 6906: 6898: 6896: 6888: 6883: 6875: 6870: 6862: 6857: 6849: 6844: 6836: 6831: 6823: 6818: 6810: 6805: 6797: 6792: 6784: 6779: 6771: 6766: 6758: 6753: 6744: 6743: 6738: 6729: 6728: 6723: 6710: 6702: 6697: 6689: 6684: 6676: 6671: 6663: 6658: 6650: 6645: 6637: 6632: 6624: 6619: 6611: 6606: 6598: 6593: 6585: 6580: 6572: 6567: 6559: 6554: 6546: 6541: 6532: 6531: 6526: 6517: 6516: 6511: 6498: 6488: 6483: 6445: 6441:Paracryphiaceae 6427: 6409: 6386: 6363: 6295: 6257: 6228:Torricelliaceae 6207: 6194: 6095:Calceolariaceae 6080:Carlemanniaceae 6061: 6023: 6005: 5987: 5949: 5926: 5908: 5883: 5870: 5747: 5697: 5684: 5660:Balanophoraceae 5636: 5547:Lophiocarpaceae 5527:Achatocarpaceae 5522:Caryophyllaceae 5512:Macarthuriaceae 5477:Drosophyllaceae 5433: 5408: 5383: 5325: 5307: 5293:Crypteroniaceae 5288:Melastomataceae 5249: 5186: 5153: 5130: 5088:Crossosomatales 5082: 5058:Gyrostemonaceae 5028:Koeberliniaceae 4982: 4969: 4921: 4912:Picrodendraceae 4877:Lacistemataceae 4837:Dichapetalaceae 4812:Centroplacaceae 4802:Lophopyxidaceae 4762:Erythroxylaceae 4728: 4705: 4682: 4624: 4576: 4543: 4514:Corynocarpaceae 4488: 4475: 4455: 4442: 4403:Grossulariaceae 4352: 4339: 4316: 4298: 4286:Trochodendrales 4280: 4261:Lardizabalaceae 4232: 4199: 4168: 4156:Ceratophyllales 4147:Ceratophyllidae 4137: 4128:Ecdeiocoleaceae 4118:Flagellariaceae 4054: 4001: 3963: 3940: 3911:Tecophilaeaceae 3881:Blandfordiaceae 3857: 3823:Petermanniaceae 3808:Campynemataceae 3794: 3756: 3728: 3710: 3671:Aponogetonaceae 3627: 3602: 3590: 3559: 3511: 3497:Himantandraceae 3468: 3440: 3406: 3404:Mesangiospermae 3393: 3365: 3337: 3313: 3308: 3289:Marchantiophyta 3268: 3257: 3227: 3222: 3186: 3153: 3100: 3052: 3025: 3000: 2983: 2954:Crossosomatales 2930: 2877: 2855: 2840: 2831: 2811: 2802: 2793:Trochodendrales 2772: 2755: 2722: 2663: 2626: 2622:Ceratophyllales 2606: 2597: 2569: 2560: 2541:Marchantiophyta 2520: 2509: 2451:Wayback Machine 2440:Wayback Machine 2429:Wayback Machine 2417:Wayback Machine 2406: 2209: 2207:Further reading 2204: 2203: 2195: 2191: 2129: 2125: 2118: 2114: 2083: 2079: 2072: 2068: 2038: 2034: 1989: 1985: 1976: 1974: 1970: 1962: 1953: 1946: 1917: 1910: 1901: 1899: 1890: 1889: 1885: 1832:Albert, V. A.; 1830: 1823: 1800: 1799: 1792: 1769: 1765: 1728: 1724: 1685: 1681: 1670: 1666: 1659: 1641: 1637: 1586: 1582: 1547: 1543: 1537:10.1139/b03-004 1517: 1506: 1463: 1456: 1441:10.2307/3298537 1420: 1413: 1405: 1391: 1387: 1350:Pacific Science 1342: 1338: 1324: 1317: 1308: 1306: 1287: 1276: 1265: 1261: 1224: 1223: 1214: 1204: 1202: 1171: 1170: 1166: 1156: 1154: 1123: 1122: 1118: 1113: 1082: 1071: 1052: 984: 949: 940: 931: 922: 913: 904: 766:Acrogymnosperms 666:lineage in the 652: 617: 595:Order Laurales 591:classified it: 589:Dahlgren system 564:classified it: 541:Order Laurales 531: 526: 521: 520: 519: 516: 513: 504: 501: 492: 489: 480: 474: 413: 409: 408: 403: 399: 398: 338:The species is 329:vessel elements 319:. The xylem of 301:leaves without 282:is a sprawling 277: 193: 188: 182: 169: 150: 132: 108: 55: 24: 17: 12: 11: 5: 7535: 7525: 7524: 7519: 7514: 7509: 7504: 7499: 7482: 7481: 7479: 7478: 7465: 7461:wfo-9000000014 7452: 7439: 7426: 7413: 7403: 7390: 7377: 7364: 7351: 7338: 7325: 7312: 7299: 7286: 7271: 7255: 7253: 7247: 7246: 7244: 7243: 7239:wfo-7000000020 7230: 7220: 7207: 7194: 7181: 7168: 7155: 7142: 7129: 7116: 7103: 7090: 7077: 7064: 7051: 7038: 7025: 7010: 6994: 6992: 6986: 6985: 6983: 6982: 6969: 6965:wfo-0000961029 6956: 6943: 6930: 6917: 6904: 6894: 6881: 6868: 6855: 6842: 6829: 6816: 6803: 6790: 6777: 6764: 6751: 6736: 6720: 6718: 6712: 6711: 6709: 6708: 6704:wfo-4000001555 6695: 6682: 6669: 6656: 6643: 6630: 6617: 6604: 6591: 6578: 6565: 6552: 6539: 6524: 6508: 6506: 6500: 6499: 6485: 6484: 6482: 6481: 6470: 6467: 6466: 6463: 6462: 6459: 6458: 6455: 6454: 6451: 6450: 6447: 6446: 6444: 6443: 6437: 6435: 6433:Paracryphiales 6429: 6428: 6426: 6425: 6423:Escalloniaceae 6419: 6417: 6411: 6410: 6408: 6407: 6405:Caprifoliaceae 6402: 6396: 6394: 6388: 6387: 6385: 6384: 6379: 6377:Columelliaceae 6373: 6371: 6365: 6364: 6362: 6361: 6356: 6351: 6346: 6341: 6339:Argophyllaceae 6336: 6331: 6329:Alseuosmiaceae 6326: 6321: 6316: 6311: 6305: 6303: 6297: 6296: 6294: 6293: 6288: 6283: 6281:Phyllonomaceae 6278: 6273: 6267: 6265: 6259: 6258: 6256: 6255: 6250: 6248:Myodocarpaceae 6245: 6240: 6238:Pittosporaceae 6235: 6233:Griseliniaceae 6230: 6225: 6219: 6217: 6208: 6203: 6200: 6199: 6196: 6195: 6193: 6192: 6187: 6182: 6177: 6172: 6167: 6162: 6157: 6155:Schlegeliaceae 6152: 6147: 6142: 6137: 6132: 6127: 6122: 6117: 6112: 6107: 6105:Plantaginaceae 6102: 6097: 6092: 6087: 6082: 6077: 6071: 6069: 6063: 6062: 6060: 6059: 6054: 6052:Sphenocleaceae 6049: 6044: 6039: 6037:Convolvulaceae 6033: 6031: 6025: 6024: 6022: 6021: 6015: 6013: 6007: 6006: 6004: 6003: 5997: 5995: 5989: 5988: 5986: 5985: 5980: 5975: 5970: 5965: 5959: 5957: 5951: 5950: 5948: 5947: 5942: 5936: 5934: 5928: 5927: 5925: 5924: 5922:Metteniusaceae 5918: 5916: 5910: 5909: 5907: 5906: 5901: 5895: 5893: 5884: 5879: 5872: 5871: 5869: 5868: 5863: 5858: 5853: 5848: 5843: 5838: 5836:Sarraceniaceae 5833: 5828: 5823: 5818: 5813: 5808: 5803: 5798: 5793: 5788: 5783: 5778: 5776:Fouquieriaceae 5773: 5768: 5766:Marcgraviaceae 5763: 5757: 5755: 5749: 5748: 5746: 5745: 5740: 5735: 5730: 5725: 5720: 5715: 5709: 5707: 5698: 5693: 5686: 5685: 5683: 5682: 5677: 5672: 5670:Misodendraceae 5667: 5662: 5657: 5652: 5646: 5644: 5638: 5637: 5635: 5634: 5629: 5624: 5619: 5614: 5609: 5604: 5599: 5594: 5589: 5584: 5579: 5574: 5572:Phytolaccaceae 5569: 5564: 5559: 5554: 5549: 5544: 5539: 5534: 5529: 5524: 5519: 5514: 5509: 5507:Asteropeiaceae 5504: 5499: 5497:Simmondsiaceae 5494: 5489: 5484: 5479: 5474: 5469: 5464: 5459: 5457:Plumbaginaceae 5454: 5449: 5443: 5441: 5439:Caryophyllales 5435: 5434: 5432: 5431: 5426: 5420: 5418: 5409: 5404: 5401: 5400: 5397: 5396: 5393: 5392: 5389: 5388: 5385: 5384: 5382: 5381: 5376: 5371: 5366: 5361: 5356: 5351: 5346: 5341: 5335: 5333: 5327: 5326: 5324: 5323: 5317: 5315: 5309: 5308: 5306: 5305: 5300: 5295: 5290: 5285: 5280: 5275: 5270: 5265: 5259: 5257: 5251: 5250: 5248: 5247: 5242: 5240:Sarcolaenaceae 5237: 5232: 5227: 5222: 5217: 5212: 5207: 5202: 5196: 5194: 5188: 5187: 5185: 5184: 5179: 5174: 5169: 5167:Gerrardinaceae 5163: 5161: 5155: 5154: 5152: 5151: 5146: 5140: 5138: 5132: 5131: 5129: 5128: 5123: 5118: 5113: 5108: 5103: 5098: 5092: 5090: 5084: 5083: 5081: 5080: 5075: 5070: 5065: 5060: 5055: 5050: 5045: 5040: 5035: 5030: 5025: 5020: 5015: 5010: 5005: 5000: 4994: 4992: 4983: 4978: 4975: 4974: 4971: 4970: 4968: 4967: 4962: 4957: 4955:Elaeocarpaceae 4952: 4947: 4942: 4937: 4931: 4929: 4923: 4922: 4920: 4919: 4917:Phyllanthaceae 4914: 4909: 4904: 4899: 4894: 4889: 4884: 4879: 4874: 4872:Passifloraceae 4869: 4864: 4859: 4854: 4849: 4844: 4839: 4834: 4829: 4824: 4819: 4814: 4809: 4807:Putranjivaceae 4804: 4799: 4794: 4789: 4784: 4782:Calophyllaceae 4779: 4774: 4769: 4764: 4759: 4757:Rhizophoraceae 4754: 4749: 4744: 4738: 4736: 4730: 4729: 4727: 4726: 4721: 4715: 4713: 4707: 4706: 4704: 4703: 4701:Zygophyllaceae 4698: 4692: 4690: 4684: 4683: 4681: 4680: 4675: 4670: 4665: 4660: 4655: 4650: 4645: 4640: 4634: 4632: 4626: 4625: 4623: 4622: 4617: 4615:Ticodendraceae 4612: 4607: 4602: 4597: 4592: 4586: 4584: 4578: 4577: 4575: 4574: 4569: 4564: 4559: 4553: 4551: 4545: 4544: 4542: 4541: 4536: 4531: 4526: 4521: 4516: 4511: 4506: 4500: 4498: 4489: 4484: 4477: 4476: 4474: 4473: 4467: 4465: 4456: 4451: 4444: 4443: 4441: 4440: 4435: 4430: 4425: 4420: 4415: 4410: 4405: 4400: 4395: 4390: 4385: 4383:Hamamelidaceae 4380: 4375: 4370: 4364: 4362: 4353: 4348: 4341: 4340: 4338: 4337: 4332: 4330:Myrothamnaceae 4326: 4324: 4318: 4317: 4315: 4314: 4308: 4306: 4300: 4299: 4297: 4296: 4290: 4288: 4282: 4281: 4279: 4278: 4273: 4268: 4266:Menispermaceae 4263: 4258: 4253: 4248: 4242: 4240: 4234: 4233: 4231: 4230: 4225: 4220: 4215: 4209: 4207: 4201: 4200: 4198: 4197: 4191: 4189: 4180: 4174: 4173: 4170: 4169: 4167: 4166: 4160: 4158: 4149: 4143: 4142: 4139: 4138: 4136: 4135: 4130: 4125: 4123:Joinvilleaceae 4120: 4115: 4110: 4105: 4100: 4095: 4090: 4085: 4080: 4075: 4070: 4064: 4062: 4056: 4055: 4053: 4052: 4047: 4042: 4037: 4032: 4027: 4022: 4017: 4015:Strelitziaceae 4011: 4009: 4003: 4002: 4000: 3999: 3994: 3992:Pontederiaceae 3989: 3984: 3979: 3973: 3971: 3965: 3964: 3962: 3961: 3956: 3954:Dasypogonaceae 3950: 3948: 3942: 3941: 3939: 3938: 3933: 3931:Amaryllidaceae 3928: 3923: 3921:Xeronemataceae 3918: 3913: 3908: 3903: 3898: 3893: 3888: 3883: 3878: 3873: 3867: 3865: 3859: 3858: 3856: 3855: 3850: 3845: 3840: 3835: 3830: 3825: 3820: 3815: 3810: 3804: 3802: 3796: 3795: 3793: 3792: 3787: 3782: 3777: 3772: 3766: 3764: 3758: 3757: 3755: 3754: 3749: 3744: 3738: 3736: 3730: 3729: 3727: 3726: 3724:Petrosaviaceae 3720: 3718: 3712: 3711: 3709: 3708: 3703: 3698: 3693: 3688: 3683: 3678: 3673: 3668: 3663: 3658: 3653: 3648: 3643: 3637: 3635: 3629: 3628: 3626: 3625: 3619: 3617: 3608: 3596: 3595: 3592: 3591: 3589: 3588: 3586:Chloranthaceae 3582: 3580: 3571: 3565: 3564: 3561: 3560: 3558: 3557: 3552: 3547: 3542: 3537: 3532: 3527: 3525:Calycanthaceae 3521: 3519: 3513: 3512: 3510: 3509: 3504: 3499: 3494: 3489: 3484: 3478: 3476: 3470: 3469: 3467: 3466: 3461: 3456: 3450: 3448: 3442: 3441: 3439: 3438: 3433: 3427: 3425: 3416: 3407: 3402: 3399: 3398: 3395: 3394: 3392: 3391: 3389:Schisandraceae 3386: 3381: 3375: 3373: 3367: 3366: 3364: 3363: 3358: 3353: 3347: 3345: 3339: 3338: 3336: 3335: 3329: 3327: 3318: 3310: 3309: 3307: 3306: 3301: 3296: 3294:Polypodiophyta 3291: 3286: 3281: 3273: 3270: 3269: 3256: 3255: 3248: 3241: 3233: 3224: 3223: 3221: 3220: 3210: 3204: 3198: 3191: 3188: 3187: 3180: 3178: 3175: 3174: 3171: 3170: 3167: 3166: 3163: 3162: 3159: 3158: 3155: 3154: 3152: 3151: 3146: 3141: 3136: 3131: 3126: 3121: 3116: 3110: 3108: 3102: 3101: 3099: 3098: 3096:Paracryphiales 3093: 3088: 3083: 3078: 3073: 3068: 3062: 3060: 3054: 3053: 3051: 3050: 3045: 3039: 3033: 3027: 3026: 3024: 3023: 3021:Caryophyllales 3018: 3013: 3007: 3001: 2996: 2993: 2992: 2989: 2988: 2985: 2984: 2982: 2981: 2976: 2971: 2966: 2961: 2956: 2951: 2946: 2940: 2938: 2932: 2931: 2929: 2928: 2923: 2918: 2913: 2908: 2903: 2898: 2893: 2887: 2885: 2879: 2878: 2876: 2875: 2869: 2863: 2857: 2856: 2854: 2853: 2847: 2841: 2836: 2833: 2832: 2830: 2829: 2824: 2818: 2812: 2807: 2804: 2803: 2801: 2800: 2795: 2790: 2785: 2779: 2773: 2768: 2765: 2764: 2761: 2760: 2757: 2756: 2754: 2753: 2748: 2743: 2738: 2732: 2730: 2724: 2723: 2721: 2720: 2714: 2708: 2702: 2696: 2690: 2684: 2677: 2671: 2665: 2664: 2662: 2661: 2656: 2651: 2646: 2640: 2638: 2628: 2627: 2625: 2624: 2619: 2613: 2607: 2604:Mesangiosperms 2602: 2599: 2598: 2596: 2595: 2589: 2583: 2576: 2570: 2565: 2562: 2561: 2559: 2558: 2553: 2548: 2546:Polypodiophyta 2543: 2538: 2533: 2525: 2522: 2521: 2508: 2507: 2500: 2493: 2485: 2479: 2478: 2473: 2468: 2458: 2453: 2419: 2405: 2404:External links 2402: 2401: 2400: 2364:Genome Biology 2342: 2306:Genome Biology 2286: 2274:10.1086/317571 2257: 2229:(3): 245–254. 2208: 2205: 2202: 2201: 2189: 2123: 2112: 2077: 2066: 2032: 2011:(3): 368–382. 1983: 1951: 1944: 1908: 1883: 1821: 1790: 1763: 1742:(2): 365–389. 1722: 1695:(3): 225–348. 1679: 1664: 1657: 1635: 1580: 1567:10.1086/317571 1541: 1504: 1491:10.1086/424024 1485:(6): 925–947. 1454: 1435:(3): 466–490. 1411: 1403: 1385: 1356:(3): 305–312. 1336: 1315: 1274: 1259: 1212: 1164: 1115: 1114: 1112: 1109: 1070: 1067: 1051: 1048: 983: 980: 946: 945: 942: 941: 937: 936: 933: 932: 928: 927: 924: 923: 919: 918: 915: 914: 910: 909: 906: 905: 901: 900: 897: 896: 889: 886: 885: 882: 881: 874: 871: 870: 867: 866: 859: 854: 852: 849: 848: 845: 844: 835: 830: 828: 825: 824: 821: 820: 811: 806: 804: 801: 800: 797: 796: 787: 782: 780: 774: 773: 770: 769: 762: 757: 755: 746: 651: 648: 616: 613: 612: 611: 610: 609: 608: 607: 606: 605: 585: 584: 583: 582: 581: 580: 579: 578: 558: 557: 556: 555: 554: 553: 552: 551: 530: 527: 525: 522: 518: 517: 514: 507: 505: 502: 495: 493: 490: 483: 481: 475: 468: 465: 464: 463: 368:inflorescences 323:contains only 276: 273: 237:and the order 195: 194: 189: 178: 177: 171: 170: 163: 161: 157: 156: 143: 139: 138: 127: 123: 122: 103: 99: 98: 93: 86: 85: 80: 73: 72: 67: 63: 62: 49: 48: 47:Male specimen 44: 43: 35: 34: 15: 9: 6: 4: 3: 2: 7534: 7523: 7520: 7518: 7515: 7513: 7510: 7508: 7505: 7503: 7500: 7498: 7495: 7494: 7492: 7475: 7470: 7466: 7462: 7457: 7453: 7449: 7444: 7440: 7436: 7431: 7427: 7423: 7418: 7414: 7410: 7404: 7400: 7395: 7391: 7387: 7382: 7378: 7374: 7369: 7365: 7361: 7356: 7352: 7348: 7343: 7339: 7335: 7330: 7326: 7322: 7317: 7313: 7309: 7304: 7300: 7296: 7291: 7287: 7282: 7276: 7272: 7267: 7261: 7257: 7256: 7254: 7252: 7248: 7240: 7235: 7231: 7227: 7221: 7217: 7212: 7208: 7204: 7199: 7195: 7191: 7186: 7182: 7178: 7173: 7169: 7165: 7160: 7156: 7152: 7147: 7143: 7139: 7134: 7130: 7126: 7121: 7117: 7113: 7108: 7104: 7100: 7095: 7091: 7087: 7082: 7078: 7074: 7069: 7065: 7061: 7056: 7052: 7048: 7043: 7039: 7035: 7030: 7026: 7021: 7020:Amborellaceae 7015: 7011: 7006: 7000: 6996: 6995: 6993: 6991: 6990:Amborellaceae 6987: 6979: 6974: 6970: 6966: 6961: 6957: 6953: 6948: 6944: 6940: 6935: 6931: 6927: 6922: 6918: 6914: 6909: 6905: 6901: 6895: 6891: 6886: 6882: 6878: 6873: 6869: 6865: 6860: 6856: 6852: 6847: 6843: 6839: 6834: 6830: 6826: 6821: 6817: 6813: 6808: 6804: 6800: 6795: 6791: 6787: 6782: 6778: 6774: 6769: 6765: 6761: 6756: 6752: 6747: 6741: 6737: 6732: 6726: 6722: 6721: 6719: 6717: 6713: 6705: 6700: 6696: 6692: 6687: 6683: 6679: 6674: 6670: 6666: 6661: 6657: 6653: 6648: 6644: 6640: 6635: 6631: 6627: 6622: 6618: 6614: 6609: 6605: 6601: 6596: 6592: 6588: 6583: 6579: 6575: 6570: 6566: 6562: 6557: 6553: 6549: 6544: 6540: 6535: 6529: 6525: 6520: 6514: 6510: 6509: 6507: 6505: 6501: 6497: 6492: 6480: 6472: 6471: 6468: 6442: 6439: 6438: 6436: 6434: 6430: 6424: 6421: 6420: 6418: 6416: 6415:Escalloniales 6412: 6406: 6403: 6401: 6398: 6397: 6395: 6393: 6389: 6383: 6380: 6378: 6375: 6374: 6372: 6370: 6366: 6360: 6357: 6355: 6352: 6350: 6347: 6345: 6344:Menyanthaceae 6342: 6340: 6337: 6335: 6332: 6330: 6327: 6325: 6322: 6320: 6317: 6315: 6314:Campanulaceae 6312: 6310: 6307: 6306: 6304: 6302: 6298: 6292: 6291:Aquifoliaceae 6289: 6287: 6286:Helwingiaceae 6284: 6282: 6279: 6277: 6274: 6272: 6271:Stemonuraceae 6269: 6268: 6266: 6264: 6260: 6254: 6251: 6249: 6246: 6244: 6241: 6239: 6236: 6234: 6231: 6229: 6226: 6224: 6223:Pennantiaceae 6221: 6220: 6218: 6216: 6212: 6209: 6206: 6201: 6191: 6190:Orobanchaceae 6188: 6186: 6185:Paulowniaceae 6183: 6181: 6178: 6176: 6173: 6171: 6168: 6166: 6163: 6161: 6158: 6156: 6153: 6151: 6148: 6146: 6143: 6141: 6138: 6136: 6133: 6131: 6128: 6126: 6123: 6121: 6120:Linderniaceae 6118: 6116: 6113: 6111: 6108: 6106: 6103: 6101: 6098: 6096: 6093: 6091: 6088: 6086: 6083: 6081: 6078: 6076: 6073: 6072: 6070: 6068: 6064: 6058: 6055: 6053: 6050: 6048: 6045: 6043: 6040: 6038: 6035: 6034: 6032: 6030: 6026: 6020: 6017: 6016: 6014: 6012: 6008: 6002: 5999: 5998: 5996: 5994: 5990: 5984: 5981: 5979: 5976: 5974: 5971: 5969: 5966: 5964: 5961: 5960: 5958: 5956: 5952: 5946: 5943: 5941: 5938: 5937: 5935: 5933: 5929: 5923: 5920: 5919: 5917: 5915: 5914:Metteniusales 5911: 5905: 5902: 5900: 5899:Oncothecaceae 5897: 5896: 5894: 5892: 5888: 5885: 5882: 5877: 5867: 5864: 5862: 5859: 5857: 5854: 5852: 5849: 5847: 5846:Actinidiaceae 5844: 5842: 5839: 5837: 5834: 5832: 5829: 5827: 5826:Diapensiaceae 5824: 5822: 5819: 5817: 5814: 5812: 5809: 5807: 5804: 5802: 5799: 5797: 5794: 5792: 5789: 5787: 5786:Lecythidaceae 5784: 5782: 5781:Polemoniaceae 5779: 5777: 5774: 5772: 5769: 5767: 5764: 5762: 5761:Balsaminaceae 5759: 5758: 5756: 5754: 5750: 5744: 5741: 5739: 5736: 5734: 5731: 5729: 5726: 5724: 5723:Hydrangeaceae 5721: 5719: 5716: 5714: 5711: 5710: 5708: 5706: 5702: 5699: 5696: 5691: 5681: 5678: 5676: 5675:Schoepfiaceae 5673: 5671: 5668: 5666: 5663: 5661: 5658: 5656: 5653: 5651: 5648: 5647: 5645: 5643: 5639: 5633: 5630: 5628: 5625: 5623: 5622:Portulacaceae 5620: 5618: 5615: 5613: 5612:Halophytaceae 5610: 5608: 5605: 5603: 5600: 5598: 5595: 5593: 5592:Molluginaceae 5590: 5588: 5587:Nyctaginaceae 5585: 5583: 5582:Sarcobataceae 5580: 5578: 5577:Petiveriaceae 5575: 5573: 5570: 5568: 5565: 5563: 5560: 5558: 5555: 5553: 5550: 5548: 5545: 5543: 5540: 5538: 5535: 5533: 5532:Amaranthaceae 5530: 5528: 5525: 5523: 5520: 5518: 5515: 5513: 5510: 5508: 5505: 5503: 5500: 5498: 5495: 5493: 5490: 5488: 5485: 5483: 5480: 5478: 5475: 5473: 5470: 5468: 5465: 5463: 5460: 5458: 5455: 5453: 5450: 5448: 5447:Frankeniaceae 5445: 5444: 5442: 5440: 5436: 5430: 5427: 5425: 5424:Aextoxicaceae 5422: 5421: 5419: 5417: 5413: 5410: 5407: 5406:Superasterids 5402: 5380: 5377: 5375: 5374:Simaroubaceae 5372: 5370: 5367: 5365: 5362: 5360: 5359:Anacardiaceae 5357: 5355: 5352: 5350: 5347: 5345: 5342: 5340: 5337: 5336: 5334: 5332: 5328: 5322: 5321:Picramniaceae 5319: 5318: 5316: 5314: 5310: 5304: 5301: 5299: 5296: 5294: 5291: 5289: 5286: 5284: 5281: 5279: 5276: 5274: 5271: 5269: 5266: 5264: 5261: 5260: 5258: 5256: 5252: 5246: 5243: 5241: 5238: 5236: 5233: 5231: 5228: 5226: 5225:Thymelaeaceae 5223: 5221: 5218: 5216: 5213: 5211: 5208: 5206: 5205:Muntingiaceae 5203: 5201: 5198: 5197: 5195: 5193: 5189: 5183: 5180: 5178: 5175: 5173: 5170: 5168: 5165: 5164: 5162: 5160: 5156: 5150: 5147: 5145: 5142: 5141: 5139: 5137: 5133: 5127: 5124: 5122: 5121:Stachyuraceae 5119: 5117: 5116:Guamatelaceae 5114: 5112: 5111:Staphyleaceae 5109: 5107: 5104: 5102: 5099: 5097: 5094: 5093: 5091: 5089: 5085: 5079: 5076: 5074: 5071: 5069: 5066: 5064: 5061: 5059: 5056: 5054: 5051: 5049: 5046: 5044: 5043:Emblingiaceae 5041: 5039: 5038:Salvadoraceae 5036: 5034: 5031: 5029: 5026: 5024: 5021: 5019: 5018:Limnanthaceae 5016: 5014: 5011: 5009: 5006: 5004: 5003:Tropaeolaceae 5001: 4999: 4996: 4995: 4993: 4991: 4987: 4984: 4981: 4976: 4966: 4965:Brunelliaceae 4963: 4961: 4960:Cephalotaceae 4958: 4956: 4953: 4951: 4948: 4946: 4943: 4941: 4938: 4936: 4933: 4932: 4930: 4928: 4924: 4918: 4915: 4913: 4910: 4908: 4907:Ixonanthaceae 4905: 4903: 4900: 4898: 4897:Euphorbiaceae 4895: 4893: 4892:Rafflesiaceae 4890: 4888: 4885: 4883: 4880: 4878: 4875: 4873: 4870: 4868: 4865: 4863: 4860: 4858: 4855: 4853: 4850: 4848: 4845: 4843: 4842:Euphroniaceae 4840: 4838: 4835: 4833: 4830: 4828: 4825: 4823: 4822:Malpighiaceae 4820: 4818: 4815: 4813: 4810: 4808: 4805: 4803: 4800: 4798: 4797:Caryocaraceae 4795: 4793: 4790: 4788: 4787:Podostemaceae 4785: 4783: 4780: 4778: 4775: 4773: 4770: 4768: 4765: 4763: 4760: 4758: 4755: 4753: 4750: 4748: 4745: 4743: 4740: 4739: 4737: 4735: 4731: 4725: 4722: 4720: 4717: 4716: 4714: 4712: 4708: 4702: 4699: 4697: 4694: 4693: 4691: 4689: 4688:Zygophyllales 4685: 4679: 4676: 4674: 4671: 4669: 4666: 4664: 4661: 4659: 4656: 4654: 4651: 4649: 4646: 4644: 4641: 4639: 4636: 4635: 4633: 4631: 4627: 4621: 4618: 4616: 4613: 4611: 4610:Casuarinaceae 4608: 4606: 4603: 4601: 4598: 4596: 4593: 4591: 4590:Nothofagaceae 4588: 4587: 4585: 4583: 4579: 4573: 4570: 4568: 4565: 4563: 4560: 4558: 4555: 4554: 4552: 4550: 4546: 4540: 4537: 4535: 4532: 4530: 4529:Tetramelaceae 4527: 4525: 4524:Cucurbitaceae 4522: 4520: 4517: 4515: 4512: 4510: 4507: 4505: 4504:Apodanthaceae 4502: 4501: 4499: 4497: 4493: 4490: 4487: 4482: 4478: 4472: 4469: 4468: 4466: 4464: 4460: 4457: 4454: 4449: 4439: 4438:Cynomoriaceae 4436: 4434: 4431: 4429: 4426: 4424: 4421: 4419: 4416: 4414: 4411: 4409: 4408:Saxifragaceae 4406: 4404: 4401: 4399: 4396: 4394: 4391: 4389: 4386: 4384: 4381: 4379: 4376: 4374: 4371: 4369: 4368:Peridiscaceae 4366: 4365: 4363: 4361: 4357: 4354: 4351: 4346: 4336: 4333: 4331: 4328: 4327: 4325: 4323: 4319: 4313: 4310: 4309: 4307: 4305: 4301: 4295: 4292: 4291: 4289: 4287: 4283: 4277: 4276:Ranunculaceae 4274: 4272: 4271:Berberidaceae 4269: 4267: 4264: 4262: 4259: 4257: 4254: 4252: 4249: 4247: 4244: 4243: 4241: 4239: 4235: 4229: 4226: 4224: 4221: 4219: 4218:Nelumbonaceae 4216: 4214: 4211: 4210: 4208: 4206: 4202: 4196: 4193: 4192: 4190: 4188: 4184: 4181: 4179: 4175: 4165: 4162: 4161: 4159: 4157: 4153: 4150: 4148: 4144: 4134: 4131: 4129: 4126: 4124: 4121: 4119: 4116: 4114: 4111: 4109: 4106: 4104: 4101: 4099: 4096: 4094: 4091: 4089: 4088:Eriocaulaceae 4086: 4084: 4081: 4079: 4076: 4074: 4071: 4069: 4066: 4065: 4063: 4061: 4057: 4051: 4050:Zingiberaceae 4048: 4046: 4043: 4041: 4038: 4036: 4033: 4031: 4028: 4026: 4025:Heliconiaceae 4023: 4021: 4018: 4016: 4013: 4012: 4010: 4008: 4004: 3998: 3997:Haemodoraceae 3995: 3993: 3990: 3988: 3985: 3983: 3982:Commelinaceae 3980: 3978: 3975: 3974: 3972: 3970: 3966: 3960: 3957: 3955: 3952: 3951: 3949: 3947: 3943: 3937: 3934: 3932: 3929: 3927: 3926:Asphodelaceae 3924: 3922: 3919: 3917: 3914: 3912: 3909: 3907: 3906:Ixioliriaceae 3904: 3902: 3901:Doryanthaceae 3899: 3897: 3894: 3892: 3889: 3887: 3884: 3882: 3879: 3877: 3874: 3872: 3869: 3868: 3866: 3864: 3860: 3854: 3851: 3849: 3846: 3844: 3841: 3839: 3836: 3834: 3831: 3829: 3826: 3824: 3821: 3819: 3818:Melanthiaceae 3816: 3814: 3811: 3809: 3806: 3805: 3803: 3801: 3797: 3791: 3788: 3786: 3785:Cyclanthaceae 3783: 3781: 3778: 3776: 3773: 3771: 3768: 3767: 3765: 3763: 3759: 3753: 3752:Dioscoreaceae 3750: 3748: 3747:Burmanniaceae 3745: 3743: 3742:Nartheciaceae 3740: 3739: 3737: 3735: 3731: 3725: 3722: 3721: 3719: 3717: 3716:Petrosaviales 3713: 3707: 3706:Cymodoceaceae 3704: 3702: 3699: 3697: 3696:Posidoniaceae 3694: 3692: 3689: 3687: 3684: 3682: 3679: 3677: 3676:Juncaginaceae 3674: 3672: 3669: 3667: 3664: 3662: 3659: 3657: 3654: 3652: 3649: 3647: 3646:Tofieldiaceae 3644: 3642: 3639: 3638: 3636: 3634: 3630: 3624: 3621: 3620: 3618: 3616: 3612: 3609: 3606: 3601: 3597: 3587: 3584: 3583: 3581: 3579: 3578:Chloranthales 3575: 3572: 3570: 3569:Chloranthidae 3566: 3556: 3553: 3551: 3548: 3546: 3545:Hernandiaceae 3543: 3541: 3538: 3536: 3535:Gomortegaceae 3533: 3531: 3528: 3526: 3523: 3522: 3520: 3518: 3514: 3508: 3505: 3503: 3502:Eupomatiaceae 3500: 3498: 3495: 3493: 3492:Degeneriaceae 3490: 3488: 3485: 3483: 3482:Myristicaceae 3480: 3479: 3477: 3475: 3471: 3465: 3462: 3460: 3457: 3455: 3452: 3451: 3449: 3447: 3443: 3437: 3434: 3432: 3429: 3428: 3426: 3424: 3420: 3417: 3415: 3411: 3408: 3405: 3400: 3390: 3387: 3385: 3382: 3380: 3377: 3376: 3374: 3372: 3368: 3362: 3359: 3357: 3354: 3352: 3351:Hydatellaceae 3349: 3348: 3346: 3344: 3340: 3334: 3333:Amborellaceae 3331: 3330: 3328: 3326: 3322: 3319: 3316: 3311: 3305: 3302: 3300: 3297: 3295: 3292: 3290: 3287: 3285: 3282: 3279: 3275: 3274: 3271: 3266: 3262: 3254: 3249: 3247: 3242: 3240: 3235: 3234: 3231: 3219: 3211: 3209: 3205: 3203: 3199: 3197: 3193: 3192: 3189: 3184: 3150: 3147: 3145: 3142: 3140: 3137: 3135: 3132: 3130: 3127: 3125: 3122: 3120: 3119:Metteniusales 3117: 3115: 3112: 3111: 3109: 3107: 3103: 3097: 3094: 3092: 3089: 3087: 3084: 3082: 3079: 3077: 3076:Escalloniales 3074: 3072: 3069: 3067: 3064: 3063: 3061: 3059: 3055: 3049: 3046: 3044: 3041: 3040: 3037: 3034: 3032: 3028: 3022: 3019: 3017: 3014: 3012: 3009: 3008: 3005: 3002: 2999: 2998:Superasterids 2994: 2980: 2977: 2975: 2972: 2970: 2967: 2965: 2962: 2960: 2957: 2955: 2952: 2950: 2947: 2945: 2942: 2941: 2939: 2937: 2933: 2927: 2924: 2922: 2919: 2917: 2914: 2912: 2909: 2907: 2904: 2902: 2899: 2897: 2894: 2892: 2891:Zygophyllales 2889: 2888: 2886: 2884: 2880: 2874: 2871: 2870: 2867: 2864: 2862: 2858: 2852: 2849: 2848: 2845: 2842: 2839: 2834: 2828: 2825: 2823: 2820: 2819: 2816: 2813: 2810: 2809:Core eudicots 2805: 2799: 2796: 2794: 2791: 2789: 2786: 2784: 2781: 2780: 2777: 2774: 2771: 2766: 2752: 2749: 2747: 2744: 2742: 2739: 2737: 2734: 2733: 2731: 2729: 2725: 2718: 2715: 2712: 2709: 2706: 2703: 2700: 2697: 2694: 2693:Petrosaviales 2691: 2688: 2685: 2682: 2679: 2678: 2675: 2672: 2670: 2666: 2660: 2657: 2655: 2652: 2650: 2647: 2645: 2642: 2641: 2639: 2637: 2633: 2629: 2623: 2620: 2618: 2617:Chloranthales 2615: 2614: 2611: 2608: 2605: 2600: 2593: 2590: 2587: 2584: 2581: 2578: 2577: 2574: 2571: 2568: 2563: 2557: 2554: 2552: 2549: 2547: 2544: 2542: 2539: 2537: 2534: 2531: 2527: 2526: 2523: 2518: 2514: 2506: 2501: 2499: 2494: 2492: 2487: 2486: 2483: 2477: 2474: 2472: 2469: 2467: 2463: 2459: 2457: 2454: 2452: 2448: 2445: 2441: 2437: 2434: 2430: 2426: 2423: 2422:Amborellaceae 2420: 2418: 2414: 2411: 2408: 2407: 2397: 2393: 2388: 2383: 2378: 2373: 2369: 2365: 2361: 2359: 2353: 2352:Soltis, P. S. 2349: 2348:Soltis, D. E. 2343: 2339: 2335: 2330: 2325: 2320: 2315: 2311: 2307: 2303: 2301: 2295: 2294:Soltis, P. S. 2291: 2290:Soltis, D. E. 2287: 2283: 2279: 2275: 2271: 2267: 2263: 2258: 2254: 2250: 2246: 2242: 2237: 2232: 2228: 2224: 2220: 2218: 2211: 2210: 2198: 2193: 2185: 2181: 2177: 2173: 2168: 2163: 2159: 2155: 2151: 2147: 2143: 2139: 2135: 2127: 2121: 2116: 2108: 2104: 2100: 2096: 2092: 2088: 2081: 2075: 2070: 2063: 2059: 2055: 2051: 2047: 2043: 2036: 2028: 2024: 2019: 2014: 2010: 2006: 2002: 1998: 1997:Soltis, D. E. 1994: 1993:Soltis, P. S. 1987: 1969: 1968: 1960: 1958: 1956: 1947: 1941: 1937: 1933: 1929: 1925: 1924:Soltis, D. E. 1921: 1920:Soltis, P. S. 1915: 1913: 1898:on 2020-01-29 1897: 1893: 1887: 1879: 1875: 1870: 1865: 1860: 1855: 1851: 1847: 1843: 1839: 1838:Soltis, P. S. 1835: 1834:Soltis, D. E. 1828: 1826: 1817: 1812: 1808: 1804: 1797: 1795: 1786: 1782: 1779:(2): 91–124. 1778: 1774: 1767: 1759: 1755: 1750: 1745: 1741: 1737: 1733: 1726: 1718: 1714: 1710: 1706: 1702: 1698: 1694: 1690: 1683: 1675: 1668: 1660: 1658:9780231038805 1654: 1649: 1648: 1639: 1631: 1627: 1623: 1619: 1615: 1611: 1607: 1603: 1599: 1595: 1591: 1584: 1576: 1572: 1568: 1564: 1560: 1556: 1552: 1545: 1538: 1534: 1530: 1526: 1522: 1515: 1513: 1511: 1509: 1500: 1496: 1492: 1488: 1484: 1480: 1476: 1472: 1471:Soltis, D. E. 1468: 1467:Soltis, P. S. 1461: 1459: 1450: 1446: 1442: 1438: 1434: 1430: 1426: 1418: 1416: 1406: 1400: 1396: 1389: 1381: 1377: 1372: 1367: 1363: 1359: 1355: 1351: 1347: 1340: 1331: 1330: 1322: 1320: 1304: 1300: 1296: 1294: 1285: 1283: 1281: 1279: 1270: 1263: 1254: 1249: 1244: 1239: 1235: 1231: 1227: 1221: 1219: 1217: 1201: 1197: 1193: 1189: 1185: 1183: 1178: 1176: 1168: 1153: 1149: 1145: 1141: 1137: 1135: 1130: 1128: 1120: 1116: 1108: 1106: 1104: 1099: 1095: 1091: 1085: 1080: 1076: 1066: 1064: 1060: 1056: 1047: 1044: 1040: 1036: 1032: 1028: 1024: 1022: 1018: 1014: 1010: 1006: 1001: 996: 992: 988: 979: 977: 973: 968: 966: 962: 958: 954: 944: 943: 935: 934: 926: 925: 917: 916: 908: 907: 899: 898: 895: 894: 888: 887: 884: 883: 880: 879: 873: 872: 869: 868: 865: 864: 858: 857: 851: 850: 847: 846: 843: 842: 841: 834: 833: 827: 826: 823: 822: 819: 818: 817: 810: 809: 803: 802: 799: 798: 795: 794: 793: 786: 785: 779: 776: 775: 772: 771: 768: 767: 761: 760: 754: 750: 749: 745: 743: 739: 735: 731: 727: 723: 722: 717: 716: 711: 707: 706: 700: 698: 694: 693: 688: 683: 681: 677: 673: 669: 665: 661: 657: 647: 645: 641: 637: 633: 629: 625: 624:APG II system 621: 603: 602: 600: 599: 597: 596: 594: 593: 592: 590: 576: 575: 573: 572: 570: 569: 567: 566: 565: 563: 549: 548: 546: 545: 543: 542: 540: 539: 538: 536: 511: 506: 499: 494: 487: 482: 479: 472: 467: 466: 462: 460: 456: 452: 448: 444: 440: 435: 433: 428: 424: 420: 395: 393: 389: 385: 381: 377: 373: 370:borne in the 369: 364: 361: 357: 353: 349: 345: 341: 336: 334: 330: 326: 322: 318: 314: 310: 306: 304: 300: 297: 293: 289: 285: 281: 272: 270: 267:to all other 266: 262: 258: 254: 250: 249: 244: 240: 236: 235:Amborellaceae 232: 231:New Caledonia 228: 224: 221: 217: 214: 210: 207: 203: 202: 192: 187: 185: 179: 176: 175:Binomial name 172: 168: 167: 162: 159: 158: 154: 149: 148: 144: 141: 140: 136: 131: 130:Amborellaceae 128: 125: 124: 120: 116: 112: 107: 104: 101: 100: 97: 94: 91: 88: 87: 84: 83:Tracheophytes 81: 78: 75: 74: 71: 68: 65: 64: 59: 54: 50: 45: 41: 36: 33: 29: 26: 22: 7281:Amborellales 7251:Amborellales 7250: 6989: 6715: 6503: 6354:Calyceraceae 6349:Goodeniaceae 6334:Phellinaceae 6324:Stylidiaceae 6263:Aquifoliales 6145:Bignoniaceae 6130:Martyniaceae 6100:Gesneriaceae 6057:Hydroleaceae 6047:Montiniaceae 6001:Boraginaceae 5978:Gelsemiaceae 5968:Gentianaceae 5940:Eucommiaceae 5841:Roridulaceae 5821:Symplocaceae 5791:Sladeniaceae 5733:Curtisiaceae 5680:Loranthaceae 5602:Didiereaceae 5557:Barbeuiaceae 5517:Microteaceae 5472:Nepenthaceae 5462:Polygonaceae 5452:Tamaricaceae 5344:Nitrariaceae 5313:Picramniales 5278:Vochysiaceae 5263:Combretaceae 5177:Tapisciaceae 5172:Petenaeaceae 5078:Brassicaceae 4832:Trigoniaceae 4827:Balanopaceae 4792:Hypericaceae 4772:Bonnetiaceae 4747:Irvingiaceae 4734:Malpighiales 4724:Celastraceae 4696:Krameriaceae 4653:Elaeagnaceae 4648:Dirachmaceae 4605:Juglandaceae 4572:Polygalaceae 4557:Quillajaceae 4519:Coriariaceae 4496:Cucurbitales 4433:Haloragaceae 4428:Penthoraceae 4413:Crassulaceae 4378:Altingiaceae 4360:Saxifragales 4312:Dilleniaceae 4251:Papaveraceae 4246:Eupteleaceae 4238:Ranunculales 4146: 4113:Restionaceae 4073:Bromeliaceae 4007:Zingiberales 3987:Philydraceae 3977:Hanguanaceae 3969:Commelinales 3936:Asparagaceae 3896:Hypoxidaceae 3843:Ripogonaceae 3838:Philesiaceae 3833:Colchicaceae 3775:Velloziaceae 3770:Triuridaceae 3734:Dioscoreales 3651:Alismataceae 3599: 3568: 3530:Siparunaceae 3487:Magnoliaceae 3384:Trimeniaceae 3361:Nymphaeaceae 3325:Amborellales 3304:Angiospermae 3259:Families of 3066:Aquifoliales 2959:Picramniales 2921:Cucurbitales 2901:Malpighiales 2851:Saxifragales 2783:Ranunculales 2751:Zingiberales 2746:Commelinales 2699:Dioscoreales 2580:Amborellales 2579: 2556:Angiospermae 2367: 2363: 2357: 2309: 2305: 2299: 2265: 2261: 2226: 2222: 2216: 2192: 2141: 2137: 2133: 2126: 2115: 2090: 2086: 2080: 2069: 2045: 2041: 2035: 2008: 2004: 1986: 1975:. Retrieved 1966: 1927: 1900:. Retrieved 1896:the original 1886: 1849: 1845: 1806: 1802: 1776: 1772: 1766: 1739: 1735: 1725: 1692: 1688: 1682: 1673: 1667: 1646: 1638: 1605: 1601: 1597: 1593: 1589: 1583: 1558: 1554: 1550: 1544: 1531:(1): 28–39, 1528: 1524: 1520: 1482: 1478: 1474: 1432: 1428: 1424: 1394: 1388: 1353: 1349: 1345: 1339: 1328: 1307:. Retrieved 1302: 1298: 1292: 1268: 1262: 1233: 1229: 1205:23 September 1203:. Retrieved 1180: 1174: 1167: 1157:23 September 1155:. Retrieved 1132: 1126: 1119: 1105:conservation 1102: 1093: 1089: 1074: 1072: 1069:Conservation 1062: 1054: 1053: 1042: 1038: 1026: 1025: 1016: 1004: 1000:sister group 986: 985: 975: 971: 969: 960: 957:sister taxon 952: 950: 891: 876: 861: 838: 837: 814: 813: 791: 790: 789: 764: 751:extant  719: 713: 709: 703: 701: 690: 684: 679: 675: 671: 662:as the most 659: 656:systematists 653: 628:Nymphaeaceae 619: 618: 586: 559: 532: 491:Young leaves 476:Specimen in 436: 396: 365: 351: 337: 320: 308: 307: 279: 278: 265:sister group 257:systematists 252: 247: 246: 239:Amborellales 238: 234: 227:Grande Terre 200: 199: 198: 183: 181: 165: 164: 146: 145: 129: 106:Amborellales 105: 89: 76: 31: 25: 7502:Angiosperms 7355:iNaturalist 7275:Wikispecies 7107:iNaturalist 7014:Wikispecies 6833:iNaturalist 6740:Wikispecies 6608:iNaturalist 6528:Wikispecies 6309:Rousseaceae 6205:Campanulids 6165:Verbenaceae 6140:Acanthaceae 6135:Pedaliaceae 6125:Byblidaceae 5993:Boraginales 5983:Apocynaceae 5973:Loganiaceae 5955:Gentianales 5904:Icacinaceae 5856:Cyrillaceae 5851:Clethraceae 5831:Styracaceae 5811:Primulaceae 5738:Grubbiaceae 5665:Santalaceae 5607:Basellaceae 5562:Gisekiaceae 5502:Physenaceae 5467:Droseraceae 5364:Sapindaceae 5354:Burseraceae 5298:Alzateaceae 5210:Neuradaceae 5149:Francoaceae 5144:Geraniaceae 5096:Aphloiaceae 5068:Capparaceae 5048:Tovariaceae 5008:Moringaceae 4990:Brassicales 4950:Cunoniaceae 4945:Oxalidaceae 4940:Connaraceae 4857:Achariaceae 4852:Humiriaceae 4817:Elatinaceae 4711:Celastrales 4668:Cannabaceae 4643:Barbeyaceae 4567:Surianaceae 4539:Begoniaceae 4534:Datiscaceae 4373:Paeoniaceae 4350:Superrosids 4335:Gunneraceae 4304:Dilleniales 4223:Platanaceae 4098:Thurniaceae 4078:Rapateaceae 4040:Marantaceae 3891:Lanariaceae 3886:Asteliaceae 3871:Orchidaceae 3863:Asparagales 3848:Smilacaceae 3790:Pandanaceae 3780:Stemonaceae 3686:Zosteraceae 3681:Maundiaceae 3633:Alismatales 3550:Monimiaceae 3474:Magnoliales 3459:Saururaceae 3436:Winteraceae 3431:Canellaceae 3414:Magnoliidae 3356:Cabombaceae 3343:Nymphaeales 3276:Supergroup 3134:Gentianales 3129:Boraginales 3058:Campanulids 2974:Brassicales 2896:Celastrales 2838:Superrosids 2827:Dilleniales 2728:Commelinids 2717:Asparagales 2687:Alismatales 2659:Magnoliales 2586:Nymphaeales 2567:Angiosperms 2528:Supergroup 2197:Pillon 2008 2048:(1): 5–21, 1465:Buzgo, M.; 1253:10654/18083 1029:, being an 1013:gametophyte 965:Nymphaeales 778:angiosperms 275:Description 96:Angiosperms 7491:Categories 7125:77126703-1 6908:Plant List 6392:Dipsacales 6382:Bruniaceae 6359:Asteraceae 6243:Araliaceae 6180:Phrymaceae 6115:Stilbaceae 6042:Solanaceae 6019:Vahliaceae 5945:Garryaceae 5891:Icacinales 5801:Sapotaceae 5655:Opiliaceae 5642:Santalales 5617:Talinaceae 5597:Montiaceae 5349:Kirkiaceae 5331:Sapindales 5303:Penaeaceae 5273:Onagraceae 5268:Lythraceae 5200:Cytinaceae 5159:Huerteales 5136:Geraniales 5073:Cleomaceae 5063:Resedaceae 5013:Caricaceae 4998:Akaniaceae 4927:Oxalidales 4882:Salicaceae 4867:Goupiaceae 4777:Clusiaceae 4678:Urticaceae 4658:Rhamnaceae 4620:Betulaceae 4600:Myricaceae 4322:Gunnerales 4228:Proteaceae 4108:Cyperaceae 4093:Mayacaceae 4083:Xyridaceae 3813:Corsiaceae 3762:Pandanales 3701:Ruppiaceae 3656:Butomaceae 3507:Annonaceae 3464:Piperaceae 3423:Canellales 3202:Alismatids 3114:Icacinales 3091:Dipsacales 3016:Santalales 2969:Huerteales 2964:Sapindales 2944:Geraniales 2906:Oxalidales 2822:Gunnerales 2705:Pandanales 2644:Canellales 2636:Magnoliids 2511:Orders of 2370:(5): R48. 2167:11336/2616 1977:2013-06-22 1902:2017-12-21 1371:10125/2455 1309:2016-10-21 1111:References 1059:change sex 1031:understory 1007:lacks the 878:magnoliids 419:apocarpous 356:staminodes 213:understory 115:A.V.Bobrov 7448:100352497 6864:126024907 6534:Amborella 6519:Q13418082 6504:Amborella 6400:Adoxaceae 6369:Bruniales 6301:Asterales 6170:Lamiaceae 6029:Solanales 6011:Vahliales 5963:Rubiaceae 5932:Garryales 5861:Ericaceae 5806:Ebenaceae 5743:Cornaceae 5728:Loasaceae 5713:Nyssaceae 5650:Olacaceae 5632:Cactaceae 5567:Aizoaceae 5542:Limeaceae 5379:Meliaceae 5283:Myrtaceae 5235:Cistaceae 5215:Malvaceae 4862:Violaceae 4767:Ochnaceae 4742:Pandaceae 4213:Sabiaceae 4205:Proteales 4103:Juncaceae 4068:Typhaceae 4045:Costaceae 4035:Cannaceae 4020:Lowiaceae 3959:Arecaceae 3916:Iridaceae 3876:Boryaceae 3853:Liliaceae 3623:Acoraceae 3555:Lauraceae 3446:Piperales 3284:Bryophyta 3149:Vahliales 3144:Solanales 3124:Garryales 3081:Bruniales 3071:Asterales 2788:Proteales 2649:Piperales 2536:Bryophyta 2300:Amborella 2253:240346942 2134:Amborella 2087:BioEssays 1590:Amborella 1305:: 143–155 1196:Libraries 1175:Amborella 1148:Libraries 1090:Amborella 1075:Amborella 1063:Amborella 1055:Amborella 1043:Amborella 1039:Amborella 1027:Amborella 1017:Amborella 1005:Amborella 987:Amborella 976:Amborella 961:Amborella 953:Amborella 792:Amborella 742:cladogram 710:Amborella 687:primitive 682:lineage. 680:Amborella 676:Amborella 672:Amborella 650:Phylogeny 644:phylogeny 634:systems, 620:Amborella 455:lignified 427:micropyle 352:Amborella 340:dioecious 333:primitive 325:tracheids 321:Amborella 309:Amborella 299:evergreen 292:alternate 286:or small 280:Amborella 253:Amborella 218:or small 206:monotypic 201:Amborella 160:Species: 147:Amborella 66:Kingdom: 32:Amborella 21:Ambarella 7443:Tropicos 7260:Wikidata 7226:amborell 7216:42000010 7211:Tropicos 7005:Q1142499 6999:Wikidata 6952:21200356 6947:Tropicos 6913:kew-7867 6897:PalDat: 6851:581745-1 6725:Wikidata 6691:40011664 6686:Tropicos 6513:Wikidata 6479:Category 6253:Apiaceae 6175:Mazaceae 6085:Oleaceae 6067:Lamiales 5816:Theaceae 5753:Ericales 5705:Cornales 5695:Asterids 5552:Kewaceae 5369:Rutaceae 5255:Myrtales 5230:Bixaceae 5192:Malvales 5033:Bataceae 4902:Linaceae 4887:Peraceae 4673:Moraceae 4663:Ulmaceae 4638:Rosaceae 4595:Fagaceae 4562:Fabaceae 4471:Vitaceae 4398:Iteaceae 4195:Buxaceae 4178:Eudicots 4030:Musaceae 3946:Arecales 3800:Liliales 3615:Acorales 3605:Monocots 3517:Laurales 3218:Category 3208:Lilioids 3139:Lamiales 3048:Ericales 3043:Cornales 3031:Asterids 2979:Malvales 2949:Myrtales 2770:Eudicots 2736:Arecales 2711:Liliales 2681:Acorales 2669:Monocots 2654:Laurales 2447:Archived 2436:Archived 2425:Archived 2413:Archived 2396:21619600 2338:18341710 2296:(2008). 2282:84820330 2245:43781302 2176:24357311 2107:17041880 2062:21628174 2027:24391149 1999:(2014). 1878:15799777 1758:85738663 1717:40348158 1598:Illicium 1575:84820330 1499:84793812 1380:35832198 1079:Tertiary 1037:between 893:eudicots 863:monocots 840:Illicium 726:monocots 721:Illicium 692:Magnolia 524:Taxonomy 451:pericarp 384:perianth 303:stipules 259:because 126:Family: 119:Zaytzeva 117:, & 111:Melikyan 7347:7941101 7321:6003628 7266:Q689001 7086:3667900 7060:2497097 6973:ZooBank 6812:3667902 6786:5403756 6731:Q310470 6639:1294247 6626:27047-1 6587:3667901 6561:5403754 6215:Apiales 5881:Lamiids 4980:Malvids 4935:Huaceae 4630:Rosales 4582:Fagales 4549:Fabales 4463:Vitales 4187:Buxales 4133:Poaceae 3641:Araceae 3600:Lilidae 3106:Lamiids 3086:Apiales 2936:Malvids 2926:Fagales 2916:Rosales 2911:Fabales 2873:Vitales 2798:Buxales 2387:3219971 2329:2397498 2184:2499045 2146:Bibcode 2138:Science 1869:1083416 1697:Bibcode 1630:2142920 1610:Bibcode 1449:3298537 1103:ex situ 1050:Ecology 1009:vessels 955:as the 658:accept 636:APG III 529:History 412:⁄ 402:⁄ 392:whorled 348:carpels 344:stamens 243:species 223:endemic 142:Genus: 102:Order: 70:Plantae 7474:886389 7435:182968 7422:927960 7406:NZOR: 7399:261009 7386:846604 7360:152141 7295:216171 7177:295910 7138:112153 7034:413498 6890:303950 6838:129826 6825:450322 6755:ARKive 6665:303952 4486:Fabids 4453:Rosids 4060:Poales 3265:APG IV 2883:Fabids 2861:Rosids 2741:Poales 2517:APG IV 2462:Hawaii 2394:  2384:  2336:  2326:  2280:  2251:  2243:  2182:  2174:  2105:  2060:  2025:  1942:  1922:& 1876:  1866:  1756:  1715:  1655:  1628:  1594:Nuphar 1573:  1497:  1469:& 1447:  1409:p. 98. 1401:  1378:  1334:p. 186 1194:& 1184:(IPNI) 1146:& 1136:(IPNI) 1098:relict 816:Nuphar 732:using 724:, the 715:Nuphar 640:APG IV 447:stigma 388:tepals 380:bracts 360:pollen 296:simple 216:shrubs 191:Baill. 153:Baill. 135:Pichon 7469:WoRMS 7373:12564 7368:IRMNG 7334:1AMBO 7308:924HW 7190:55385 7164:22097 7151:18132 7133:IRMNG 7112:71436 7073:1AMBF 6921:Plazi 6877:13333 6799:BMBTR 6773:65ZQY 6652:13332 6634:IRMNG 6613:71271 6574:1BMBG 6548:628VH 3278:Plant 2530:Plant 2298:"The 2278:S2CID 2249:S2CID 2241:JSTOR 2180:S2CID 1971:(PDF) 1852:: 5. 1754:S2CID 1736:Aliso 1713:S2CID 1626:S2CID 1571:S2CID 1495:S2CID 1445:JSTOR 1376:S2CID 697:parts 668:clade 664:basal 443:drupe 439:fruit 423:style 376:cymes 372:axils 313:xylem 284:shrub 229:, of 220:trees 209:genus 204:is a 90:Clade 77:Clade 7394:NCBI 7381:ITIS 7342:GBIF 7329:EPPO 7290:APNI 7198:POWO 7159:NCBI 7146:ITIS 7120:IPNI 7094:GRIN 7081:GBIF 7068:EPPO 7029:BOLD 6934:POWO 6872:NCBI 6859:IUCN 6846:IPNI 6820:GRIN 6807:GBIF 6794:EPPO 6673:POWO 6647:NCBI 6621:IPNI 6595:GRIN 6582:GBIF 6569:EPPO 2442:via 2392:PMID 2334:PMID 2172:PMID 2103:PMID 2058:PMID 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Index

Ambarella

Scientific classification
Edit this classification
Plantae
Tracheophytes
Angiosperms
Amborellales
Melikyan
A.V.Bobrov
Zaytzeva
Amborellaceae
Pichon
Amborella
Baill.
Binomial name
Baill.
monotypic
genus
understory
shrubs
trees
endemic
Grande Terre
New Caledonia
species
systematists
molecular phylogenetic
sister group
flowering plants

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