2743:
605:
617:
1665:
2738:
572:
425:
135:
111:
596:
specimen include (a) long bones that feature smooth surfaces, which are not marked by the rugosities characteristic of adult bone; (b) the head of the femur is less spherical in shape and is clearly delimited from the adjacent part of the femoral shaft; (c) the distal ends of the tibia and the fibula are not fused to the astragalus and the calcaneum; and (d) the ungual phalanx of the manus is not widest at the proximal articular end as is observed in adults.
2766:
1653:
513:. It was inspired by the fact that compression damage to the specimen had removed the teeth, at the same time shifting various flat round elements below the skull and on top of the left lower jaw, misleading Sternberg into assuming that large "trituration plates" had replaced the normal dentition. The
595:
Coombs supported the assertion that specimen AMNH 5266 represented a juvenile by citing that the vertebral centra were not fused to their neural arches, and that sacral ribs were likewise not fused to vertebrae and to the ilium. Other morphological characters supporting that this is a juvenile
591:
has armor on a majority of the dorsal surfaces of its body. It has a wide, pointed tail club at the end of its armored tail. The skull features postocular caputegulae, which are small polygonal plates of bone that are present on the cranium and are situated to the immediate rear of the eye.
563:, an incomplete left pes, and various other fragments. AMNH 5266 was discovered in 1912 at Red Deer River and was collected by Barnum Brown with assistance from Peter Kaisen, George Olsen, and Charles M Sternberg in the sediments from the Horseshoe Canyon Formation.
586:
was a medium-sized ankylosaurid, measuring up to 4–5 metres (13–16 ft) in length and 2 metric tons (2.2 short tons) in body mass. It was a quadrupedal, ground-dwelling herbivore. Like other ankylosaurs,
1350:
Paul
Penkalski (2018). Revised systematics of the armoured dinosaur Euoplocephalus and its allies. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 287(3): 261-306. doi:
1589:
Arbour VM, Currie PJ (2013) Euoplocephalus tutus and the
Diversity of Ankylosaurid Dinosaurs in the Late Cretaceous of Alberta, Canada, and Montana, USA. PLoS ONE 8(5): e62421.
690:
in distinctive skull and cervical half ring ornamentation, as well as tail club morphology, including the presence of pointed, triangular knob osteoderms in
2903:
714:
2938:
2913:
710:
783:
extending the stratigraphic range of the genus back a few million years. The DPF specimen, however, was later made the holotype of a new species,
2844:
629:
1313:
Arbour, Victoria (2010). "A Cretaceous armoury: Multiple ankylosaurid taxa in the Late
Cretaceous of Alberta, Canada and Montana, USA".
2923:
2928:
2831:
2908:
1697:
2742:
1481:
775:. In a study based on the results of her 2010 SVP abstract, Arbour along with Philip Currie formalized the revalidation of
1718:
548:
1602:
Arbour, V. M.; Currie, P. J. (2015). "Systematics, phylogeny and palaeobiogeography of the ankylosaurid dinosaurs".
2918:
2759:
1247:
1437:(Archosauria: Ornithischia): a foundation for comparative and systematic studies of ankylosaurian dinosaurs",
2933:
2737:
134:
694:. She therefore reassigned all Horseshoe Canyon Formation ankylosaurine specimens previously referred to
537:
1690:
1643:
642:
506:
483:
377:
475:
451:
440:
286:
1555:(Ornithischia: Ankylosauria) from the Upper Cretaceous Dinosaur Park Formation of Alberta, Canada".
1788:
1384:
Arbour, V. M.; Burns, M. E.; Sissons, R. L. (2009). "A redescription of the ankylosaurid dinosaur
514:
397:
2898:
2875:
2786:
2849:
2870:
2862:
1815:
1683:
1367:
C. M. Sternberg (1929) "A toothless armoured dinosaur from the Upper
Cretaceous of Alberta."
792:
522:
1541:
Coombs W. (1971) The
Ankylosauridae. Ph.D. thesis, Columbia University, New York, NY, 487 p.
2818:
2769:
1611:
1564:
1397:
1252:
654:
498:
444:
339:
290:
8:
2175:
2072:
648:
1615:
1568:
1401:
1911:
1627:
1413:
1330:
129:
2857:
2668:
1932:
1631:
1487:
1477:
1334:
1039:
779:, and one specimen from the Dinosaur Park Formation, TMP 1997.132.1, was referred to
502:
1417:
1283:"A new ankylosaurid from the late Cretaceous Two Medicine Formation of Montana, USA"
2133:
2080:
1918:
1657:
1619:
1572:
1522:
1405:
1326:
1322:
1290:
1623:
2598:
2562:
2502:
2468:
2461:
2354:
2295:
2167:
2064:
1980:
1940:
1773:
857:
679:
495:
479:
374:
30:
604:
424:
2809:
2661:
2633:
2541:
2514:
2410:
2391:
2306:
2252:
2117:
2087:
2057:
2020:
1996:
1590:
1122:
802:
552:
370:
253:
240:
556:
474:
remains. The badly crushed skeleton was collected by
Sternberg in 1916 from a
2892:
2624:
2534:
2527:
2479:
2423:
2370:
2336:
2237:
2183:
2158:
2140:
2043:
1989:
1962:
1954:
1669:
1527:
1506:
1491:
1456:
Coombs, W.P., Jr. (1986, June). A Juvenile
Ankylosaur Referable to the Genus
977:
816:
740:
560:
533:
510:
491:
385:
266:
68:
1460:(Reptilia, Ornithischia). Journal of Vertebrate Paleontology, 6(2), 162-173.
2654:
2640:
2569:
2453:
2377:
2322:
2273:
2245:
2230:
2217:
2190:
2109:
2094:
2050:
2030:
1897:
1879:
1857:
1827:
1800:
1748:
1706:
1576:
1433:
Vickaryous, M.K. and
Russell, A.P., 2003, "A redescription of the skull of
1079:
991:
874:
736:
616:
543:
In 1986 Coombs examined specimen AMNH 5266, at the time by him referred to
363:
298:
227:
195:
1351:
1295:
1282:
2689:
2606:
2555:
2495:
2446:
2288:
2266:
2198:
2147:
1947:
1925:
1757:
1409:
1369:
Canada
Department of Mines Geological Survey Bulletin (Geological Series)
1146:
840:
471:
211:
43:
731:
on the basis of remains that were originally thought to be referable to
636:
age of the Late
Cretaceous of North America. He synonymised the species
571:
547:, and determined that it was a juvenile. It consists of five vertebral
2836:
2682:
2675:
2613:
2488:
2439:
2431:
2384:
2329:
2315:
2281:
2259:
2125:
1969:
1904:
1388:
Parks, 1924 (Ornithischia: Ankylosauria) and a revision of the genus".
946:
723:
88:
53:
1507:"Unusual cranial and postcranial anatomy in the archetypal ankylosaur
2696:
2576:
2363:
2343:
2205:
2102:
2004:
1471:
1022:
905:
788:
633:
632:
concluded that there was only one species of ankylosaurid during the
529:
487:
463:
381:
146:
93:
37:
2780:
721:
as well. The second study, by Penkalski (2013), named and described
2803:
2703:
1739:
1730:
1163:
752:
455:
448:
366:
179:
166:
121:
83:
78:
63:
58:
48:
2823:
2591:
2583:
929:
728:
401:
389:
356:
110:
98:
73:
1724:
1675:
526:
393:
156:
747:
specimen that was included in this analysis was its holotype.
666:
was generally accepted and thus CMN 8530 was assigned to
2548:
678:(2009), In an abstract presented at the SVP 2010 conference,
467:
459:
359:
2710:
1601:
709:
was formalized in three studies. The first, published by
1595:
795:
of the Ankylosaurinae conducted by Arbour and Currie:
1641:
767:, and specimens that are thought to represent either
482:. It was collected from the upper part of the Lower
328:
305:
1383:
759:and some specimens that are referred to it, while
2890:
1429:
1427:
1363:
1361:
1359:
1772:
1550:
1544:
1691:
1424:
1356:
1591:https://doi.org/10.1371/journal.pone.0062421
1504:
1452:
1450:
1308:
1306:
1276:
1274:
1272:
1270:
1268:
555:, one dorsal and two sacral ribs, the right
1877:
1476:. Princeton University Press. p. 264.
2904:Late Cretaceous dinosaurs of North America
1698:
1684:
1535:
1346:
1344:
670:. However, following the redescription of
373:. It is known from the entire span of the
109:
1526:
1447:
1439:Zoological Journal of the Linnean Society
1303:
1294:
1280:
1265:
559:, the complete right hindlimb, the right
388:stage, about 72.8-67 Ma ago) of southern
615:
603:
570:
423:
2939:Multispecific non-avian dinosaur genera
2914:Taxa named by Charles Mortram Sternberg
1379:
1377:
1352:https://doi.org/10.1127/njgpa/2018/0717
1341:
2891:
1473:The Princeton Field Guide to Dinosaurs
1312:
2785:
2784:
1717:
1679:
1551:Penkalski, P.; Blows, W. T. (2013). "
735:. Penkalski (2013) performed a small
1469:
1374:
1604:Journal of Systematic Palaeontology
1505:Arbour, V.M.; Mallon, J.C. (2017).
13:
1557:Canadian Journal of Earth Sciences
1390:Journal of Vertebrate Paleontology
1315:Journal of Vertebrate Paleontology
540:where the holotype was reposited.
454: 8530, a partially preserved
14:
2950:
2924:Campanian genus first appearances
599:
2765:
2764:
2741:
2736:
1705:
1663:
1651:
133:
41:
2929:Maastrichtian genus extinctions
1583:
1248:Timeline of ankylosaur research
486:(unit 2), dating to the latest
400:(75.6 Ma ago). It contains two
1498:
1463:
1327:10.1080/02724634.2010.10411819
566:
1:
2909:Fossil taxa described in 1929
1624:10.1080/14772019.2015.1059985
1287:Acta Palaeontologica Polonica
1258:
478:quarry, 8 miles southwest of
396:, and is also known from the
509:means "toothless lizard" in
419:
7:
1241:
763:was placed in a clade with
674:as a valid genus by Arbour
538:Geological Survey of Canada
10:
2955:
1386:Dyoplosaurus acutosquameus
643:Dyoplosaurus acutosquameus
484:Horseshoe Canyon Formation
378:Horseshoe Canyon Formation
16:Extinct genus of dinosaurs
2793:
2754:
2734:
2623:
2522:
2513:
2478:
2418:
2409:
2353:
2305:
2225:
2216:
2157:
2038:
2029:
2018:
1979:
1892:
1888:
1873:
1854:
1826:
1808:
1799:
1781:
1768:
1713:
1509:Ankylosaurus magniventris
1470:Paul, Gregory S. (2016).
1160:
1143:
1136:
1119:
1093:
1076:
1069:
1062:
1036:
1019:
1012:
1005:
988:
981:
943:
926:
919:
902:
895:
888:
871:
854:
837:
830:
813:
806:
476:Canadian Museum of Nature
441:Charles Mortram Sternberg
325:
320:
304:
297:
130:Scientific classification
128:
117:
108:
23:
1528:10.1139/facets-2017-0063
658:. The synonymisation of
2919:Paleontology in Alberta
398:Dinosaur Park Formation
1577:10.1139/cjes-2012-0098
1281:Penkalski, P. (2013).
717:in 2013, re-validated
625:
613:
580:
433:
2871:Paleobiology Database
1321:(Supplement 2): 55A.
1296:10.4202/app.2012.0125
793:phylogenetic analysis
755:with the holotype of
660:Anodontosaurus lambei
638:Anodontosaurus lambei
619:
607:
574:
523:Lawrence Morris Lambe
501:, about 71-70.2
427:
369:within the subfamily
310:Anodontosaurus lambei
2934:Ornithischian genera
1435:Euoplocephalus tutus
1410:10.1671/039.029.0405
1253:2018 in paleontology
743:specimens. The only
664:Euoplocephalus tutus
655:Euoplocephalus tutus
1616:2016JSPal..14..385A
1569:2013CaJES..50..171P
1553:Scolosaurus cutleri
1402:2009JVPal..29.1117A
791:is based on a 2015
649:Scolosaurus cutleri
432:tail club AMNH 5245
626:
614:
581:
575:Holotype skull of
434:
2886:
2885:
2858:Open Tree of Life
2787:Taxon identifiers
2778:
2777:
2750:
2749:
2732:
2731:
2728:
2727:
2724:
2723:
2720:
2719:
2669:Nodocephalosaurus
2405:
2404:
2401:
2400:
2014:
2013:
1933:Serendipaceratops
1869:
1868:
1865:
1864:
1850:
1849:
1846:
1845:
1483:978-1-78684-190-2
1238:
1237:
1229:
1228:
1220:
1219:
1211:
1210:
1202:
1201:
1193:
1192:
1184:
1183:
1175:
1174:
1108:
1107:
1051:
1050:
1040:Nodocephalosaurus
967:
966:
958:
957:
739:analysis of some
503:million years ago
428:Cast of referred
348:
347:
342:
293:
2946:
2879:
2878:
2866:
2865:
2853:
2852:
2840:
2839:
2827:
2826:
2814:
2813:
2812:
2782:
2781:
2768:
2767:
2745:
2740:
2520:
2519:
2416:
2415:
2223:
2222:
2134:Propanoplosaurus
2081:Glyptodontopelta
2036:
2035:
2027:
2026:
1919:Priodontognathus
1890:
1889:
1886:
1885:
1875:
1874:
1806:
1805:
1779:
1778:
1770:
1769:
1763:
1762:
1715:
1714:
1700:
1693:
1686:
1677:
1676:
1668:
1667:
1666:
1656:
1655:
1654:
1647:
1636:
1635:
1599:
1593:
1587:
1581:
1580:
1548:
1542:
1539:
1533:
1532:
1530:
1502:
1496:
1495:
1467:
1461:
1454:
1445:
1431:
1422:
1421:
1381:
1372:
1365:
1354:
1348:
1339:
1338:
1310:
1301:
1300:
1298:
1278:
1139:
1138:
1072:
1071:
1065:
1064:
1015:
1014:
1008:
1007:
984:
983:
922:
921:
898:
897:
891:
890:
833:
832:
809:
808:
799:
798:
787:. The following
751:was placed in a
715:William T. Blows
705:The validity of
579:, TMP 1997.132.1
490:to the earliest
337:
330:
307:
285:
278:
265:
252:
239:
226:
210:
194:
138:
137:
113:
103:
40:
29:Temporal range:
21:
20:
2954:
2953:
2949:
2948:
2947:
2945:
2944:
2943:
2889:
2888:
2887:
2882:
2874:
2869:
2861:
2856:
2848:
2843:
2835:
2830:
2822:
2817:
2808:
2807:
2802:
2789:
2779:
2774:
2746:
2716:
2619:
2599:Tianzhenosaurus
2563:Minotaurasaurus
2509:
2503:Zhongyuansaurus
2474:
2469:Zhejiangosaurus
2462:Liaoningosaurus
2397:
2355:Struthiosaurini
2349:
2301:
2296:Tatankacephalus
2212:
2168:Gargoyleosaurus
2153:
2065:Gargoyleosaurus
2023:
2010:
1981:Parankylosauria
1975:
1941:Sinankylosaurus
1882:
1861:
1842:
1822:
1795:
1789:Avemetatarsalia
1774:Avemetatarsalia
1764:
1720:
1719:
1709:
1704:
1674:
1664:
1662:
1652:
1650:
1642:
1640:
1639:
1600:
1596:
1588:
1584:
1549:
1545:
1540:
1536:
1503:
1499:
1484:
1468:
1464:
1455:
1448:
1432:
1425:
1382:
1375:
1366:
1357:
1349:
1342:
1311:
1304:
1279:
1266:
1261:
1244:
1239:
1230:
1221:
1212:
1203:
1194:
1185:
1176:
1109:
1052:
968:
959:
858:Zhejiangosaurus
680:Victoria Arbour
624:skull AMNH 5238
602:
569:
534:palaeontologist
496:Late Cretaceous
422:
375:Late Cretaceous
336:
316:
315:Sternberg, 1929
313:
284:
276:
263:
250:
237:
224:
208:
192:
132:
124:skull CMN 8530
104:
102:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
31:Late Cretaceous
27:
17:
12:
11:
5:
2952:
2942:
2941:
2936:
2931:
2926:
2921:
2916:
2911:
2906:
2901:
2884:
2883:
2881:
2880:
2867:
2854:
2841:
2828:
2815:
2799:
2797:
2795:Anodontosaurus
2791:
2790:
2776:
2775:
2773:
2772:
2762:
2755:
2752:
2751:
2748:
2747:
2735:
2733:
2730:
2729:
2726:
2725:
2722:
2721:
2718:
2717:
2715:
2714:
2707:
2700:
2693:
2686:
2679:
2672:
2665:
2662:Euoplocephalus
2658:
2651:
2648:Anodontosaurus
2644:
2637:
2634:Akainacephalus
2629:
2627:
2621:
2620:
2618:
2617:
2610:
2603:
2595:
2588:
2580:
2573:
2566:
2559:
2552:
2545:
2542:Crichtonsaurus
2538:
2531:
2523:
2517:
2515:Ankylosaurinae
2511:
2510:
2508:
2507:
2499:
2492:
2484:
2482:
2476:
2475:
2473:
2472:
2465:
2458:
2450:
2443:
2436:
2428:
2419:
2413:
2411:Ankylosauridae
2407:
2406:
2403:
2402:
2399:
2398:
2396:
2395:
2392:Struthiosaurus
2388:
2381:
2374:
2367:
2359:
2357:
2351:
2350:
2348:
2347:
2340:
2333:
2326:
2319:
2311:
2309:
2307:Panoplosaurini
2303:
2302:
2300:
2299:
2292:
2285:
2278:
2270:
2263:
2256:
2253:Niobrarasaurus
2249:
2242:
2234:
2226:
2220:
2214:
2213:
2211:
2210:
2202:
2195:
2187:
2180:
2172:
2163:
2161:
2155:
2154:
2152:
2151:
2144:
2137:
2130:
2122:
2118:Polacanthoides
2114:
2106:
2099:
2091:
2088:Horshamosaurus
2084:
2077:
2069:
2061:
2058:Dongyangopelta
2054:
2047:
2039:
2033:
2024:
2021:Euankylosauria
2019:
2016:
2015:
2012:
2011:
2009:
2008:
2001:
1997:Kunbarrasaurus
1993:
1985:
1983:
1977:
1976:
1974:
1973:
1966:
1959:
1951:
1944:
1937:
1929:
1922:
1915:
1908:
1901:
1893:
1883:
1878:
1871:
1870:
1867:
1866:
1863:
1862:
1855:
1852:
1851:
1848:
1847:
1844:
1843:
1841:
1840:
1839:
1838:
1832:
1830:
1824:
1823:
1821:
1820:
1819:
1818:
1809:
1803:
1797:
1796:
1794:
1793:
1792:
1791:
1782:
1776:
1766:
1765:
1761:
1760:
1751:
1742:
1733:
1727:
1711:
1710:
1703:
1702:
1695:
1688:
1680:
1673:
1672:
1660:
1638:
1637:
1594:
1582:
1563:(2): 171–182.
1543:
1534:
1521:(2): 764–794.
1497:
1482:
1462:
1458:Euoplocephalus
1446:
1423:
1373:
1355:
1340:
1302:
1263:
1262:
1260:
1257:
1256:
1255:
1250:
1243:
1240:
1236:
1235:
1232:
1231:
1227:
1226:
1223:
1222:
1218:
1217:
1214:
1213:
1209:
1208:
1205:
1204:
1200:
1199:
1196:
1195:
1191:
1190:
1187:
1186:
1182:
1181:
1178:
1177:
1173:
1172:
1169:
1168:
1159:
1156:
1155:
1152:
1151:
1142:
1137:
1135:
1132:
1131:
1128:
1127:
1123:Euoplocephalus
1118:
1115:
1114:
1111:
1110:
1106:
1105:
1102:
1101:
1097:Anodontosaurus
1092:
1089:
1088:
1085:
1084:
1075:
1070:
1068:
1063:
1061:
1058:
1057:
1054:
1053:
1049:
1048:
1045:
1044:
1035:
1032:
1031:
1028:
1027:
1018:
1013:
1011:
1006:
1004:
1001:
1000:
997:
996:
987:
982:
980:
974:
973:
970:
969:
965:
964:
961:
960:
956:
955:
952:
951:
942:
939:
938:
935:
934:
925:
920:
918:
915:
914:
911:
910:
901:
896:
894:
889:
887:
884:
883:
880:
879:
870:
867:
866:
863:
862:
853:
850:
849:
846:
845:
836:
831:
829:
826:
825:
822:
821:
812:
807:
805:
803:Ankylosaurinae
797:
781:Anodontosaurus
777:Anodontosaurus
757:Euoplocephalus
749:Anodontosaurus
745:Anodontosaurus
733:Euoplocephalus
711:Paul Penkalski
707:Anodontosaurus
700:Anodontosaurus
696:Euoplocephalus
692:Anodontosaurus
688:Euoplocephalus
686:distinct from
684:Anodontosaurus
601:
600:Classification
598:
589:Anodontosaurus
584:Anodontosaurus
568:
565:
545:Euoplocephalus
458:including the
437:Anodontosaurus
421:
418:
371:Ankylosaurinae
352:Anodontosaurus
346:
345:
344:
343:
323:
322:
321:Other species
318:
317:
314:
302:
301:
295:
294:
281:Anodontosaurus
274:
270:
269:
261:
257:
256:
254:Ankylosaurinae
248:
244:
243:
241:Ankylosauridae
235:
231:
230:
222:
215:
214:
206:
199:
198:
190:
183:
182:
177:
170:
169:
164:
160:
159:
154:
150:
149:
144:
140:
139:
126:
125:
115:
114:
106:
105:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
28:
25:Anodontosaurus
15:
9:
6:
4:
3:
2:
2951:
2940:
2937:
2935:
2932:
2930:
2927:
2925:
2922:
2920:
2917:
2915:
2912:
2910:
2907:
2905:
2902:
2900:
2899:Ankylosaurids
2897:
2896:
2894:
2877:
2872:
2868:
2864:
2859:
2855:
2851:
2846:
2842:
2838:
2833:
2829:
2825:
2820:
2816:
2811:
2805:
2801:
2800:
2798:
2796:
2792:
2788:
2783:
2771:
2763:
2761:
2757:
2756:
2753:
2744:
2739:
2713:
2712:
2708:
2706:
2705:
2701:
2699:
2698:
2694:
2692:
2691:
2687:
2685:
2684:
2680:
2678:
2677:
2673:
2671:
2670:
2666:
2664:
2663:
2659:
2657:
2656:
2652:
2650:
2649:
2645:
2643:
2642:
2638:
2636:
2635:
2631:
2630:
2628:
2626:
2625:Ankylosaurini
2622:
2616:
2615:
2611:
2609:
2608:
2604:
2601:
2600:
2596:
2594:
2593:
2589:
2586:
2585:
2581:
2579:
2578:
2574:
2572:
2571:
2567:
2565:
2564:
2560:
2558:
2557:
2553:
2551:
2550:
2546:
2544:
2543:
2539:
2537:
2536:
2535:Crichtonpelta
2532:
2530:
2529:
2528:Bissektipelta
2525:
2524:
2521:
2518:
2516:
2512:
2505:
2504:
2500:
2498:
2497:
2493:
2491:
2490:
2486:
2485:
2483:
2481:
2480:Shamosaurinae
2477:
2471:
2470:
2466:
2464:
2463:
2459:
2456:
2455:
2451:
2449:
2448:
2444:
2442:
2441:
2437:
2434:
2433:
2429:
2426:
2425:
2424:Ahshislepelta
2421:
2420:
2417:
2414:
2412:
2408:
2394:
2393:
2389:
2387:
2386:
2382:
2380:
2379:
2375:
2373:
2372:
2371:Hungarosaurus
2368:
2366:
2365:
2361:
2360:
2358:
2356:
2352:
2346:
2345:
2341:
2339:
2338:
2337:Panoplosaurus
2334:
2332:
2331:
2327:
2325:
2324:
2320:
2318:
2317:
2313:
2312:
2310:
2308:
2304:
2298:
2297:
2293:
2291:
2290:
2286:
2284:
2283:
2279:
2276:
2275:
2271:
2269:
2268:
2264:
2262:
2261:
2257:
2255:
2254:
2250:
2248:
2247:
2243:
2240:
2239:
2238:Ahshislepelta
2235:
2233:
2232:
2228:
2227:
2224:
2221:
2219:
2215:
2208:
2207:
2203:
2201:
2200:
2196:
2193:
2192:
2188:
2186:
2185:
2184:Hoplitosaurus
2181:
2178:
2177:
2173:
2170:
2169:
2165:
2164:
2162:
2160:
2159:Polacanthinae
2156:
2150:
2149:
2145:
2143:
2142:
2141:Rhadinosaurus
2138:
2136:
2135:
2131:
2128:
2127:
2123:
2120:
2119:
2115:
2112:
2111:
2107:
2105:
2104:
2100:
2097:
2096:
2092:
2090:
2089:
2085:
2083:
2082:
2078:
2075:
2074:
2070:
2067:
2066:
2062:
2060:
2059:
2055:
2053:
2052:
2048:
2046:
2045:
2044:Acanthopholis
2041:
2040:
2037:
2034:
2032:
2028:
2025:
2022:
2017:
2007:
2006:
2002:
1999:
1998:
1994:
1992:
1991:
1990:Antarctopelta
1987:
1986:
1984:
1982:
1978:
1972:
1971:
1967:
1965:
1964:
1963:Tianchisaurus
1960:
1957:
1956:
1955:Stegosaurides
1952:
1950:
1949:
1945:
1943:
1942:
1938:
1935:
1934:
1930:
1928:
1927:
1923:
1921:
1920:
1916:
1914:
1913:
1909:
1907:
1906:
1902:
1900:
1899:
1895:
1894:
1891:
1887:
1884:
1881:
1876:
1872:
1860:
1859:
1853:
1836:
1835:
1834:
1833:
1831:
1829:
1825:
1817:
1813:
1812:
1811:
1810:
1807:
1804:
1802:
1798:
1790:
1786:
1785:
1784:
1783:
1780:
1777:
1775:
1771:
1767:
1759:
1755:
1752:
1750:
1746:
1743:
1741:
1737:
1734:
1732:
1728:
1726:
1722:
1721:
1716:
1712:
1708:
1701:
1696:
1694:
1689:
1687:
1682:
1681:
1678:
1671:
1661:
1659:
1649:
1648:
1645:
1633:
1629:
1625:
1621:
1617:
1613:
1609:
1605:
1598:
1592:
1586:
1578:
1574:
1570:
1566:
1562:
1558:
1554:
1547:
1538:
1529:
1524:
1520:
1516:
1512:
1510:
1501:
1493:
1489:
1485:
1479:
1475:
1474:
1466:
1459:
1453:
1451:
1443:
1440:
1436:
1430:
1428:
1419:
1415:
1411:
1407:
1403:
1399:
1395:
1391:
1387:
1380:
1378:
1370:
1364:
1362:
1360:
1353:
1347:
1345:
1336:
1332:
1328:
1324:
1320:
1316:
1309:
1307:
1297:
1292:
1288:
1284:
1277:
1275:
1273:
1271:
1269:
1264:
1254:
1251:
1249:
1246:
1245:
1234:
1233:
1225:
1224:
1216:
1215:
1207:
1206:
1198:
1197:
1189:
1188:
1180:
1179:
1171:
1170:
1167:
1166:
1165:
1158:
1157:
1154:
1153:
1150:
1149:
1148:
1141:
1140:
1134:
1133:
1130:
1129:
1126:
1125:
1124:
1117:
1116:
1113:
1112:
1104:
1103:
1100:
1099:
1098:
1091:
1090:
1087:
1086:
1083:
1082:
1081:
1074:
1073:
1067:
1066:
1060:
1059:
1056:
1055:
1047:
1046:
1043:
1042:
1041:
1034:
1033:
1030:
1029:
1026:
1025:
1024:
1017:
1016:
1010:
1009:
1003:
1002:
999:
998:
995:
994:
993:
986:
985:
979:
978:Ankylosaurini
976:
975:
972:
971:
963:
962:
954:
953:
950:
949:
948:
941:
940:
937:
936:
933:
932:
931:
924:
923:
917:
916:
913:
912:
909:
908:
907:
900:
899:
893:
892:
886:
885:
882:
881:
878:
877:
876:
869:
868:
865:
864:
861:
860:
859:
852:
851:
848:
847:
844:
843:
842:
835:
834:
828:
827:
824:
823:
820:
819:
818:
817:Crichtonpelta
811:
810:
804:
801:
800:
796:
794:
790:
786:
782:
778:
774:
770:
766:
762:
758:
754:
750:
746:
742:
741:ankylosaurine
738:
734:
730:
726:
725:
720:
716:
712:
708:
703:
701:
697:
693:
689:
685:
681:
677:
673:
669:
665:
661:
657:
656:
651:
650:
645:
644:
639:
635:
631:
630:Walter Coombs
623:
618:
611:
606:
597:
593:
590:
585:
578:
573:
564:
562:
558:
554:
550:
546:
541:
539:
535:
531:
528:
524:
520:
516:
515:specific name
512:
511:Ancient Greek
508:
504:
500:
497:
494:stage of the
493:
492:Maastrichtian
489:
485:
481:
477:
473:
469:
465:
461:
457:
453:
450:
446:
442:
439:was named by
438:
431:
426:
417:
415:
414:
409:
408:
403:
399:
395:
391:
387:
386:Maastrichtian
383:
379:
376:
372:
368:
365:
361:
358:
354:
353:
341:
335:
334:
327:
326:
324:
319:
312:
311:
303:
300:
296:
292:
288:
283:
282:
275:
272:
271:
268:
267:Ankylosaurini
262:
259:
258:
255:
249:
246:
245:
242:
236:
233:
232:
229:
223:
220:
217:
216:
213:
207:
204:
201:
200:
197:
191:
188:
185:
184:
181:
178:
175:
172:
171:
168:
165:
162:
161:
158:
155:
152:
151:
148:
145:
142:
141:
136:
131:
127:
123:
120:
116:
112:
107:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
36:75.6–67
32:
26:
22:
19:
2794:
2709:
2702:
2695:
2688:
2681:
2674:
2667:
2660:
2655:Dyoplosaurus
2653:
2647:
2646:
2641:Ankylosaurus
2639:
2632:
2612:
2605:
2597:
2590:
2582:
2575:
2570:Pinacosaurus
2568:
2561:
2554:
2547:
2540:
2533:
2526:
2501:
2494:
2487:
2467:
2460:
2454:Hylaeosaurus
2452:
2445:
2438:
2430:
2422:
2390:
2383:
2378:Pawpawsaurus
2376:
2369:
2362:
2342:
2335:
2328:
2323:Denversaurus
2321:
2314:
2294:
2287:
2280:
2274:Peloroplites
2272:
2265:
2258:
2251:
2246:Borealopelta
2244:
2236:
2231:Acantholipan
2229:
2218:Nodosaurinae
2204:
2197:
2191:Peloroplites
2189:
2182:
2174:
2166:
2146:
2139:
2132:
2124:
2116:
2110:Mymoorapelta
2108:
2101:
2095:Hylaeosaurus
2093:
2086:
2079:
2071:
2063:
2056:
2051:Anoplosaurus
2049:
2042:
2031:Nodosauridae
2003:
1995:
1988:
1968:
1961:
1953:
1946:
1939:
1931:
1924:
1917:
1910:
1903:
1898:Cryptosaurus
1896:
1880:Ankylosauria
1858:Ankylosauria
1856:
1828:Ankylosauria
1816:Ornithischia
1801:Ornithischia
1753:
1749:Ornithischia
1744:
1735:
1707:Ankylosauria
1607:
1603:
1597:
1585:
1560:
1556:
1552:
1546:
1537:
1518:
1514:
1508:
1500:
1472:
1465:
1457:
1441:
1438:
1434:
1393:
1389:
1385:
1371:54(49):28-33
1368:
1318:
1314:
1286:
1162:
1161:
1145:
1144:
1121:
1120:
1096:
1095:
1094:
1080:Ankylosaurus
1078:
1077:
1038:
1037:
1021:
1020:
992:Dyoplosaurus
990:
989:
945:
944:
928:
927:
904:
903:
875:Pinacosaurus
873:
872:
856:
855:
839:
838:
815:
814:
784:
780:
776:
772:
769:Dyoplosaurus
768:
765:Dyoplosaurus
764:
760:
756:
748:
744:
737:phylogenetic
732:
722:
718:
706:
704:
699:
695:
691:
687:
683:
675:
672:Dyoplosaurus
671:
667:
663:
659:
653:
647:
641:
637:
627:
621:
609:
594:
588:
583:
582:
576:
544:
542:
518:
507:generic name
436:
435:
429:
412:
411:
406:
405:
384:to "middle"
364:ankylosaurid
351:
350:
349:
332:
331:
309:
308:
299:Type species
280:
279:
228:Ankylosauria
218:
202:
196:Ornithischia
186:
173:
118:
24:
18:
2690:Scolosaurus
2607:Tsagantegia
2556:Jinyunpelta
2496:Shamosaurus
2447:Chuanqilong
2289:Silvisaurus
2267:Patagopelta
2199:Polacanthus
2148:Sauroplites
1948:Spicomellus
1926:Sarcolestes
1758:Thyreophora
1610:(5): 1–60.
1396:(4): 1117.
1147:Scolosaurus
841:Tsagantegia
785:A. inceptus
773:Scolosaurus
719:Scolosaurus
682:considered
622:A. inceptus
577:A. inceptus
567:Description
553:neural arch
472:postcranial
447:, based on
413:A. inceptus
338:Penkalski,
333:A. inceptus
247:Subfamily:
212:Thyreophora
2893:Categories
2758:See also:
2683:Platypelta
2676:Oohkotokia
2614:Zaraapelta
2489:Gobisaurus
2440:Cedarpelta
2432:Aletopelta
2385:Stegopelta
2330:Edmontonia
2316:Animantarx
2282:Sauropelta
2260:Nodosaurus
2126:Priconodon
1970:Vectipelta
1905:Dracopelta
1837:see below↓
1740:Dinosauria
1259:References
947:Zaraapelta
761:Oohkotokia
724:Oohkotokia
521:, honours
470:and other
380:(mid Late
180:Dinosauria
2697:Talarurus
2577:Saichania
2364:Europelta
2344:Texasetes
2206:Taohelong
2103:Invictarx
2005:Stegouros
1723:Kingdom:
1658:Dinosaurs
1632:214625754
1492:985402380
1444:: 157–186
1335:220429286
1023:Talarurus
906:Saichania
789:cladogram
634:Campanian
628:In 1971,
620:Referred
610:A. lambei
608:Referred
536:from the
530:geologist
488:Campanian
464:half ring
430:A. lambei
420:Discovery
407:A. lambei
382:Campanian
367:dinosaurs
287:Sternberg
153:Kingdom:
147:Eukaryota
119:A. lambei
2824:24179445
2810:Q4769855
2804:Wikidata
2770:Category
2760:Timeline
2704:Ziapelta
2176:Gastonia
2073:Gastonia
1731:Chordata
1729:Phylum:
1725:Animalia
1418:85665879
1242:See also
1164:Ziapelta
753:polytomy
668:E. tutus
527:Canadian
456:skeleton
449:holotype
234:Family:
167:Chordata
163:Phylum:
157:Animalia
143:Domain:
122:holotype
2863:4128597
2850:1235636
2837:4821857
2592:Tarchia
2584:Shanxia
1644:Portals
1612:Bibcode
1565:Bibcode
1398:Bibcode
930:Tarchia
729:Montana
557:ischium
402:species
390:Alberta
357:extinct
273:Genus:
260:Tribe:
1670:Canada
1630:
1515:FACETS
1490:
1480:
1416:
1333:
676:et al.
646:, and
549:centra
525:, the
519:lambei
505:. The
499:period
480:Morrin
394:Canada
355:is an
2876:63439
2845:IRMNG
2549:Datai
1912:Minmi
1754:Clade
1745:Clade
1736:Clade
1628:S2CID
1414:S2CID
1331:S2CID
727:from
652:with
612:skull
468:armor
460:skull
360:genus
219:Clade
203:Clade
187:Clade
174:Clade
2832:GBIF
2711:Zuul
1814:see
1787:see
1488:OCLC
1478:ISBN
713:and
662:and
551:, a
532:and
445:1928
410:and
340:2018
291:1929
44:PreꞒ
2819:EoL
1620:doi
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