1034:. They were seen as ideal hunting companions for the elderly who could follow on horseback without exerting themselves, for young hunters who could keep up with them on ponies, and for the poorer hunters who could not afford to maintain a stable of good hunting horses. Before the advent of the fashion for foxhunting in the 19th century, hunting was an all day event where the enjoyment was derived from the chase rather than the kill. In this setting, the tiny Beagle was well matched to the hare, as unlike Harriers they would not quickly finish the hunt, but because of their excellent scent-tracking skills and stamina they were almost guaranteed to eventually catch the hare. The Beagle packs would run closely together ("so close that they might be covered with a sheet") which was useful in a long hunt, as it prevented stray dogs from obscuring the trail. In thick undergrowth they were also preferred to
874:, a condition which causes them to destroy things when left unattended. Not all Beagles will howl, but most will bark when confronted with strange situations, and some will bay (also referred to as "speaking", "giving tongue", or "opening") when they catch the scent of potential quarry. They also generally get along well with cats and other dogs. They are not too demanding with regard to exercise; their inbred stamina means they do not easily tire when exercised, but they also do not need to be worked to exhaustion before they will rest. Regular exercise helps ward off the weight gain to which the breed is prone.
659:, and a black (or occasionally liver) gumdrop nose. The jaw is strong, and the teeth scissor together with the upper teeth fitting perfectly over the lower teeth. Both sets align square to the jaw. The eyes are large, hazel or brown, with a mild, hound-like pleading look. The large ears are long, soft, and low-set, turning towards the cheeks slightly and rounded at the tips. Beagles have a strong, medium-length neck (which is long enough for them to easily bend to the ground to pick up a scent), with little folding in the skin but some evidence of a
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688:. These dogs have mostly white coats with patches of black and brown hair. Tricolor Beagles are almost always born black and white. The white areas are typically set by eight weeks, but the black areas may fade to brown as the puppy matures. (The brown may take between one and two years to fully develop.) Some Beagles gradually change colour during their lives, and may lose their black markings entirely.
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1189:. These dogs are used to detect food items in luggage being taken into the United States. After trialling several breeds, Beagles were chosen because they are relatively small and unintimidating for people who are uncomfortable around dogs, easy to care for, intelligent and work well for rewards. They are also used for this purpose in a number of other countries including by the
720:, the Beagle has one of the best developed senses of smell of any dog. In the 1950s, John Paul Scott and John Fuller began a 13-year study of canine behaviour. As part of this research, they tested the scenting abilities of various breeds by putting a mouse in a one-acre field and timing how long it took the dogs to find it. The Beagles found it in less than a minute while
1209:, due to their size and passive nature. In the United States, as many as 65,000 Beagles are used every year for medical, cosmetic, beauty, and other chemical tests. They are purpose bred and live their lives in cages undergoing experiments. The Rescue + Freedom Project (formerly Beagle Freedom Project) has successfully advocated for Beagles to be released from labs.
870:, as Coren places them among the group with the lowest degree of working/obedience intelligence. Coren's scale, however, does not assess understanding, independence, or creativity. Beagles are excellent with children and this is one of the reasons they have become popular family pets. Beagles are pack animals; they are prone to
563:'s registered breeds; in 2005 and 2006 it ranked 5th out of the 155 breeds registered. In the UK they are not quite so popular, placing 28th and 30th in the rankings of registrations with the Kennel Club in 2005 and 2006 respectively. In the United States the beagle ranked 4th most popular breed in 2012 and 2013, behind the
691:
Two-colour varieties always have a white base colour with areas of the second colour. Tan and white is the most common two-colour variety, but there is a wide range of other colours including lemon, a very light tan; red, a reddish, almost orange, brown; and liver, a darker brown, and black. Liver is
683:
The tricolored Beagle—white with large black areas and light brown shading—is the most common. Tricolored
Beagles occur in a number of shades, from the "Classic Tri" with a jet black saddle (also known as "Blackback"), to the "Dark Tri" (where faint brown markings are intermingled with more prominent
487:
By 1887, the threat of extinction was on the wane: there were 18 beagle packs in
England. The Beagle Club was formed in 1890 and the first standard drawn up at the same time. The following year the Association of Masters of Harriers and Beagles was formed. Both organisations aimed to further the best
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kept a breed known as a Pocket Beagle, which stood 8 to 9 inches (20 to 23 cm) at the shoulder. Small enough to fit in a "pocket" or saddlebag, they rode along on the hunt. The larger hounds would run the prey to ground, then the hunters would release the small dogs to continue the chase through
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provided $ 400,000 in taxpayer money to fund experiments in which 28 Beagles were infected by disease-causing parasites. The White Coat
Project found reports that said dogs taking part in the experiments were "vocalizing in pain" after being injected with foreign substances. Following public outcry,
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Testing of cosmetic products on animals is banned in the member states of the
European Community, although France protested the ban and has made efforts to have it lifted. It is permitted in the United States but is not mandatory if safety can be ascertained by other methods, and the test species is
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gave special status to primates, equids, cats and dogs and in 2005 the Animal
Procedures Committee (set up by the act) ruled that testing on mice was preferable, even though a greater number of individual animals were involved. In 2005 beagles were involved in less than 0.3% of the total experiments
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The traditional foot pack consists of up to 40 Beagles, marshaled by a
Huntsman who directs the pack and who is assisted by a variable number of whippers-in whose job is to return straying hounds to the pack. The Master of the Hunt is in overall day-to-day charge of the pack, and may or may not take
550:
As purebred dogs, Beagles have always been more popular in the United States and Canada than in their native country
England. The National Beagle Club of America was formed in 1888 and by 1901 a Beagle had won a Best in Show title. As in the UK, activity during World War I was minimal, but the breed
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In rabbit-shooting, in gorse and thick cover, nothing can be more cheerful than the beagle. They also are easily heard over long distances and in thick cover. They have been called rabbit-beagles from this employment, for which they are peculiarly qualified, especially those dogs which are somewhat
772:
English and
American varieties are sometimes mentioned. However, there is no official recognition from any Kennel Club for this distinction. Beagles fitting the American Kennel Club standard – which disallows animals over 15 inches (38 cm) – are smaller on average than those fitting the Kennel
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Beagles were in the United States by the 1840s at the latest, but the first dogs were imported strictly for hunting and were of variable quality. Since
Honeywood had only started breeding in the 1830s, it is unlikely these dogs were representative of the modern breed, and the description of them as
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description, since at 22 to 24 inches (56 to 61 cm) it is significantly taller than the modern day Beagle, and in earlier times was even larger. Some writers suggest that the Beagle's scenting ability may have come from cross-breeding earlier strains with the Kerry Beagle. Originally used for
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when all shows were suspended. After the war, the breed was again struggling for survival in the UK: the last of the Pocket
Beagles was probably lost during this time, and registrations fell to an all-time low. A few breeders (notably Reynalton Kennels) managed to revive interest in the dog and by
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and waist and a long, slightly curved tail (known as the "stern") tipped with white. The white tip, known as the flag, was bred for selectively, as the tail remains easily seen when the dog's head is down following a scent. The tail does not curl over the back, but is held upright when the dog is
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The Beagle has an even temper and gentle disposition. Described in several breed standards as "merry", they are amiable and typically neither aggressive nor timid, although this depends on the individual. They enjoy company, and although they may initially be standoffish with strangers, they are
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Roccaro, Mariana; Salini, Romolo; Pietra, Marco; Sgorbini, Micaela; Gori, Eleonora; Dondi, Maurizio; Crisi, Paolo E.; Conte, Annamaria; Dalla Villa, Paolo; Podaliri, Michele; Ciaramella, Paolo; Di Palma, Cristina; Passantino, Annamaria; Porciello, Francesco; Gianella, Paola; Guglielmini, Carlo;
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Although credited with the development of the modern breed, Honeywood concentrated on producing dogs for hunting and it was left to Thomas Johnson to refine the breeding to produce dogs that were both attractive and capable hunters. Two strains were developed: the rough-coated and smooth-coated
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As field dogs they are prone to minor injuries such as cuts and sprains, and, if inactive, obesity is a common problem as they will eat whenever food is available and rely on their owners to regulate their weight. When working or running free they are also likely to pick up parasites such as
474:, was still dividing beagles into four varieties: the medium beagle; the dwarf or lapdog beagle; the fox beagle (a smaller, slower version of the Foxhound); and the rough-coated or terrier beagle, which he classified as a cross between any of the other varieties and one of the Scottish
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Beagles are used in a range of research procedures: fundamental biological research, applied human medicine, applied veterinary medicine, and protection of man, animals, or the environment. Of the 8,018 dogs used in testing in the UK in 2004, 7,799 were Beagles (97.3%). In the UK, the
559:(Uno), won the Best In Show category at the Westminster Kennel Club show for the first time in the competition's history. In North America they have been consistently in the top-ten most-popular breeds for over 30 years. From 1953 to 1959 the beagle was ranked No. 1 on the list of the
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In the 1840s, a standard beagle type was beginning to develop. The distinction between the North Country Beagle and Southern Hound had been lost, but there was still a large variation in size, character, and reliability among the emerging packs. In 1856, "Stonehenge" (the pseudonym of
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Saddle-patterned Beagles that express ancient domino (eA) are called hare-pied. Domino restricts eumelanin production leading to a smaller and faded saddle. Dark hairs in the area of the saddle have a light-coloured hair base. The nose pigment may be lightened down the middle.
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varieties. The rough-coated beagle survived until the beginning of the 20th century, and there were even records of one making an appearance at a dog show as late as 1969, but this variety is now extinct, having probably been absorbed into the standard beagle bloodline.
940:
Their long floppy ears can mean that the inner ear does not receive a substantial air flow or that moist air becomes trapped, and this can lead to ear infections. Beagles may also be affected by a range of eye problems; two common ophthalmic conditions in beagles are
367:
and often entertained guests at her royal table by letting her Pocket Beagles cavort amid their plates and cups. 19th-century sources refer to these breeds interchangeably and it is possible that the two names refer to the same small variety. In George Jesse's
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requests to obtain information about taxpayer funding of animal testing. For example, the White Coat Waste Project, a group of activists that hold that taxpayers should not have to pay $ 20 billion every year for experiments on animals, highlighted that the
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A dog may normally be quite calm but can become very excitable when set off by such things as a ringing doorbell or an owner's movement toward the door. This characteristic may be very annoying to some people. Rank these seven breeds from least to most
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In the United States they appear to have been employed chiefly for hunting rabbits from the earliest imports. Hunting hare with Beagles became popular again in Britain in the mid-19th century and continued until it was made illegal in Scotland by the
1197:, and in Canada, Japan, and the People's Republic of China. Larger breeds are generally used for detection of explosives as this often involves climbing over luggage and on large conveyor belts, work for which the smaller Beagle is not suited.
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occur in Beagles. Two conditions in particular are unique to the breed: "Funny Puppy", in which the puppy is slow to develop and eventually develops weak legs, a crooked back and although normally healthy, is prone to a range of illnesses; and
1000:, in which they sound as if they are choking or gasping for breath, but are actually drawing air in through the mouth and nose. The exact cause of this behaviour is not known, but it can be a common occurrence and is not harmful to the dog.
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A strain known as Patch Hounds was developed by Willet Randall and his family from 1896 specifically for their rabbit hunting ability. They trace their bloodline back to Field Champion Patch, but do not necessarily have a patchwork marking.
429:
formed a good majority of the beagle's bloodline, but the origin of the Harrier is itself obscure. Honeywood's Beagles were small, standing at about 10 inches (25 cm) at the shoulder, and pure white according to John Mills (writing in
1323:, who was subsequently rescued after spending 100 hours buried in the rubble. Beagles were hired by New York City to help with bedbug detection, although some have expressed doubts about the role of such dogs in this type of detection.
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Canine juvenile polyarteritis syndrome, also known as juvenile polyarteritis syndrome of beagle dogs is a multisystemic necrotising vasculitis of the small arteries. The condition is characterised by a reoccurrent acute fever and
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that had started in 1889, and the Beagle Club in the UK held its first show in 1896. The regular showing of the breed led to the development of a uniform type, and the Beagle continued to prove a success up until the outbreak of
420:
in the 1830s; it is believed that this pack formed the basis for the modern breed. Although details of the pack's lineage are not recorded, it is thought that North Country Beagles and Southern Hounds were strongly represented;
404:
and was common in the northern counties. It was smaller than the Southern Hound, less heavy-set, and with a more pointed muzzle. It was faster than its southern counterpart but its scenting abilities were less well-developed.
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as surrogates for direct human testing. Minnesota was the first state to enact a Beagle Freedom adoption law in 2014, mandating that dogs and cats are allowed to be adopted once they have completed research testing.
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Snyder, P. W.; Kazacos, E. A.; Scott-Moncrieff, J. C.; HogenEsch, H.; Carlton, W. W.; Glickman, L. T.; Felsburg, P. J. (1995). "Pathologic Features of Naturally Occurring Juvenile Polyarteritis in Beagle Dogs".
647:, but the head is broader and the muzzle shorter, the expression completely different and the legs shorter in proportion to the body. They are generally between 13 and 16 inches (33 and 41 cm) high at the
482:
In size the beagle measures from 10 inches, or even less, to 15. In shape they resemble the old southern hound in miniature, but with more neatness and beauty; and they also resemble that hound in style of
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Peace TA, Goodchild LR, Vasconcelos DY: What's your diagnosis? Fever and leukocytosis in a young beagle. Canine juvenile polyarteritis syndrome (beagle pain syndrome). Lab Anim (NY) 2001 Vol 30 (5) pp.
937:. Affected puppies are slow, have lower co-ordination, fall more often, and do not have a normal gait. It has an estimated carrier rate of 5% and affected rate of 0.1%. A genetic test is available.
776:
Pocket Beagles are sometimes advertised for sale but while the UK Kennel Club originally specified a standard for the pocket Beagle in 1901, the variety is now not recognised by any Kennel Club.
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detection in Australia, and have been mentioned as possible candidates for drug and explosive detection. Because of their gentle nature and unimposing build, they are also frequently used in
812:. Some puppies of this cross are less excitable than a Beagle and with a lower exercise requirement, similar to the Pug parent; but many are highly excitable and require vigorous exercise.
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active. The Beagle has a muscular body and a medium-length, smooth, hard coat. The front legs are straight and carried under the body while the rear legs are muscular and well bent at the
684:
black markings), to the "Faded Tri" (where faint black markings are intermingled with more prominent brown markings). Some tricolored dogs have a broken pattern, sometimes referred to as
350:
was used as a generic description for the smaller hounds, though these dogs differed considerably from the modern breed. Miniature breeds of beagle-type dogs were known from the times of
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when confronted with the unfamiliar makes them good watch dogs. In a 1985 study conducted by Ben and Lynette Hart, the Beagle was given the highest excitability rating, along with the
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imported some dogs from England and began breeding. Rowett's Beagles are believed to have formed the models for the first American standard, drawn up by Rowett, L. H. Twadell, and
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is popular where hunting is no longer permitted or for those owners who do not wish to participate in hunting a live animal, but still wish to exercise their dog's innate skills.
961:, a condition in which eyelashes grow into the eye causing irritation, sometimes exist; both these conditions can be corrected with surgery. They can suffer from several types of
745:
recognises two separate varieties of Beagle: the 13-inch for hounds less than 13 inches (33 cm), and the 15-inch for those between 13 and 15 inches (33 and 38 cm). The
692:
not common and is not permitted in some standards; it tends to occur with yellow eyes. Ticked or mottled varieties may be either white or black with different coloured flecks (
380:
little small mitten-beagle, which may be companion for a ladies kirtle, and in the field will run as cunningly as any hound whatere, only their musick is very small like reeds.
442:
also had Beagle packs around this time, and royal favour no doubt led to some revival of interest in the breed, but Honeywood's pack was regarded as the finest of the three.
1936:
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on animals in the UK, but of the 7670 experiments performed on dogs 7406 involved Beagles (96.6%). Most dogs are bred specifically for this purpose, by companies such as
3116:
3584:
555:'s show and by 1939 a Beagle – Champion Meadowlark Draughtsman – had captured the title of top-winning American-bred dog for the year. On 12 February 2008, a Beagle,
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Standards for the Pocket Beagle were drawn up as late as 1901; these genetic lines are now extinct, although modern breeders have attempted to recreate the variety.
3397:
2634:"Spine – Abstract: Volume 31(10) May 1, 2006 p 1094-1099 Notochord Cells Regulate Intervertebral Disc Chondrocyte Proteoglycan Production and Cell Proliferation"
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965:. Failure of the nasolacrimal drainage system can cause dry eye or leakage of tears onto the face. The cause of primary open angle glaucoma in the Beagle is an
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to be a good worker, silent and obedient, but it had the drawback that it was small and could barely carry a hare. More recently, the trend has been for "
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and weigh between 18 and 35 lb (8.2 and 15.9 kg), with females being slightly smaller than males on average. They have a smooth, somewhat domed
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Fielding: "'What the devil would you have me do?' cries the Squire, turning to Blifil, 'I can no more turn her, than a beagle can turn an old hare.'"
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The Harts posed the following question to a panel of 96 experts, half of which were veterinary surgeons and the other half dog obedience trial judges:
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Beagle hound colours: Two-coloured "tan and white" (older female with fading colour), tricoloured broken, two-coloured "red and white", "tricoloured"
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with weak heads has little resemblance to the standard. Serious attempts at establishing a quality bloodline began in the early 1870s when General
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Although bred for hunting, Beagles are versatile and are nowadays employed for various other roles in detection, therapy, and as family pets.
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groups have reported on abuse of animals inside testing facilities. In 1997 footage secretly filmed by a freelance journalist inside
1125:
In addition to organised beagling, Beagles have been used for hunting or flushing to guns (often in pairs) a wide range of game including
696:), such as the blue-mottled or bluetick Beagle, which has spots that appear to be a midnight-blue colour, similar to the colouring of the
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was credited with saving the life of its owner after using her owner's mobile phone to dial an emergency number. In the aftermath of the
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had several Beagles, and caused an outcry when he picked up one of them by its ears during an official greeting on the White House lawn.
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over a period of 3-7 days. The Beagle is predisposed with most reports concerning the breed — whence the name 'Beagle pain syndrome'.
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McMillan, Kirsten M.; Bielby, Jon; Williams, Carys L.; Upjohn, Melissa M.; Casey, Rachel A.; Christley, Robert M. (1 February 2024).
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With the fashion for faster hunts, the beagle fell out of favour for chasing hare, but was still employed for rabbit hunting. In
392:(or Northern Hound). The Southern Hound, a tall, heavy dog with a square head, and long, soft ears, was common from south of the
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failed to find it at all. The long ears and large lips of the Beagle probably assist in trapping the scents close to the nose.
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interests of the breed, and both were keen to produce a standard type of Beagle. By 1902, the number of packs had risen to 44.
547:, the breed was once again doing well. Registrations dropped again after the end of the war but almost immediately recovered.
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Oliver, James A.C.; Mellersh, Cathryn S. (2020). "Genetics". In Cooper, Barbara; Mullineaux, Elizabeth; Turner, Lynn (eds.).
2015:
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1235:(FDA). When testing toxicity of food additives, food contaminants, and some drugs and chemicals the FDA uses Beagles and
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A 2024 study in the UK found a life expectancy of 12.5 years compared to an average of 12.7 for purebreeds and 12 for
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1222:. In the UK companies breeding animals for research must be licensed under the Animals (Scientific Procedures) Act.
910:, common in Harriers and in some larger breeds, is rarely considered a problem in Beagles. Beagles are considered a
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1422:
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906:(MLS) in which the eyes are slanted and the outer toes are underdeveloped but otherwise development is as normal.
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In this article "Beagle" (with a capital B) is used to distinguish the modern breed from other beagle-type dogs.
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On its formation, the Association of Masters of Harriers and Beagles took over the running of a regular show at
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257:. Possessing a great sense of smell and superior tracking instincts, the beagle is the primary breed used as a
400:. Though slow, it had stamina and an excellent scenting ability. The North Country Beagle was bred chiefly in
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showed a much stronger revival in the U.S. when hostilities ceased. In 1928 it won a number of prizes at the
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3115:. Australian Government: Department of Agriculture, Fisheries and Forestry. 30 November 2000. Archived from
929:(where the immune system attacks the joints) even at a young age. The symptoms can sometimes be relieved by
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Weight gain can be a problem in older or sedentary dogs, which in turn can lead to heart and joint problems.
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1255:, a UK-based breeder of Beagles for testing, closed down in 1997 after pressure from animal rights groups.
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resign effective immediately and that there is a "need to find a new NIH director to replace the outgoing
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affiliated clubs recognise a single type, with a height of between 13 and 16 inches (33 and 41 cm).
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The Kennel Club (UK) standard states the Beagle should give the impression of quality without coarseness.
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Tickell: "Here let me trace beneath the purpled morn, The deep-mouth'd beagle, and the sprightly horn"
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886:. A 2024 Italian study found a life expectancy of 11 years for the breed compared to 10 years overall.
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Pope: "Thus on a roe the well-breath'd beagle flies, And rends his hide fresh-bleeding with the dart"
42:
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3168:. United States Department of Agriculture: Animal and Plant Health Inspection Service. Archived from
3069:. United States Department of Agriculture: Animal and Plant Health Inspection Service. Archived from
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38:
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2148:
1989:
358:, who both had packs of Glove Beagles, so named since they were small enough to fit on a glove, and
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The modern breed was developed in Great Britain around the 1830s from several breeds, including the
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1963:"American Kennel Club 2013 Dog Registration Statistics Historical Comparisons & Notable Trends"
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Dryden: "The rest in shape a beagle's whelp throughout, With broader forehead and a sharper snout"
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use of Beagles for hunting as early as 1902 but the pack is still in existence today. In 2001, the
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2713:"Frequency of the neonatal cerebellar cortical degeneration (NCCD) mutations in Beagles in the UK"
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Beagles have excellent noses; this dog is employed by the US Customs and Border Protection Agency.
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Cowper: "For persevering chase and headlong leaps, True beagle as the staunchest hound he keeps"
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3374:. European Commission — Enterprise and Industry Directorate General. 2007. Archived from
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1079:. Under this legislation, Beagles may still pursue rabbits with the landowner's permission.
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3639:"Experimenters Fed Puppies' Heads to Infected Flies, but That's Not All Fauci's NIH Funded"
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993:, and irritants such as grass seeds can become trapped in their eyes, soft ears, or paws.
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times in literature and paintings and more recently in film, television, and comic books.
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The Beagle has been used for rabbit-hunting since the earliest development of the breed.
161:
Tricolor or white in combination with black & tan/brown or brown/tan or yellow/white
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4328:
4212:
4178:
4157:
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4014:
3605:"WCW Exposé: Fauci Spend $ 424K on Beagle Experiments, Dogs Bitten to Death by Flies"
2949:
2886:
2878:
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2774:
2749:
2653:
2499:
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2281:
2219:
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1348:
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838:
376:
is quoted referring to the beagle as small enough to sit on a man's hand and to the:
2665:
2536:
2519:
930:
700:. Some tricolour Beagles also have ticking of various colours in their white areas.
384:
By the 18th century, two breeds had been developed for hunting hare and rabbit: the
5647:
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4172:
3169:
3166:"USDA's Detector Dogs: Protecting American Agriculture: More Detector Dog Programs"
3070:
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2645:
2541:
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2489:
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1219:
1076:
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911:
793:
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603:
is uncertain, although it has been suggested that the word derives from the French
572:
467:
450:
320:
4822:
4639:
3652:
3510:
2019:
1449:, who in 2008 became the first Beagle to win the Westminster Kennel Club Dog Show
1320:
749:
recognises a single type, with a height not exceeding 15 inches (38 cm). The
5872:
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5688:
5493:
5478:
5473:
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5344:
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4897:
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4832:
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4284:
3848:
2969:
2518:
Alborali, Giovanni L.; Rota Nodari, Sara; Sabatelli, Sonia; Peli, Angelo (2024).
2165:
1277:
1252:
962:
942:
801:
568:
373:
292:
187:
3113:"Old dogs — new tricks Original quarantine K9's on the scent of retirement"
2099:
See various breed standards linked from the breed box at the top of the article.
1090:
As hunting with Beagles was seen as ideal for young people, many of the British
5779:
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4145:
3585:"Should dogs be guinea pigs in government research? A bipartisan group says no"
2874:
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1384:
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385:
336:
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258:
4262:
3738:
2810:
2773:(Sixth ed.). British Small Animal Veterinary Association. p. 131.
2317:
1307:, visiting the sick and elderly in hospital. In June 2006, a trained Beagle
1280:
who will shut down research that violates the dignity of nonhuman animals."
331:
to give them speed and stamina for deer hunting. Beagles are similar to the
5839:
5809:
5794:
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5637:
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5193:
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1344:
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717:
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544:
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397:
324:
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220:
4338:
3305:"Statistics of Scientific Procedures on Living Animals Great Britain 2004"
3251:"Statistics of Scientific Procedures on Living Animals Great Britain 2005"
2890:
1966:
1914:
1361:, who was billed as "the world's most famous Beagle". The clay animation,
1319:
with a Colombian rescue squad was credited with locating the owner of the
5843:
5799:
5678:
5599:
5316:
5221:
4996:
4971:
4812:
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4754:
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4473:
4408:
4388:
4315:
1288:
1005:
854:
721:
539:
393:
343:
239:
3503:"Seeing Is Believing — cruelty to dogs at Huntingdon Life Sciences"
2718:. Kennel Club Genetics Centre at the Animal Health Trust. Archived from
2546:
1428:
A Canadian bred 15 inch female Beagle with the registered name of Gr Ch
1030:
Beagles were developed primarily for hunting hare, an activity known as
635:
5306:
5286:
5123:
4862:
4644:
4609:
4398:
3041:"Submission to Lord Burns' Committee of Inquiry into Hunting with Dogs"
2748:(3rd ed.). Lippincott, Williams & Wilkins. pp. 656, 718.
1937:"Registration statistics for all recognised dog breeds – 2005 and 2006"
1398:
950:
830:
797:
713:
529:
A uniform type for the breed developed at the start of the 20th century
262:
2367:
1404:
after the breed, and, in turn, lent its name to the ill-fated British
760:
5789:
5724:
5296:
5217:
5188:
5113:
5097:
4961:
4827:
4448:
3067:"USDA's Detector Dogs: Protecting American Agriculture: Why Beagles?"
2859:
1510:
The specific references in each of the author's works are as follows:
1154:
1134:
1062:
498:
401:
328:
327:
to Britain. In Britain, both of these strains were then crossed with
235:
3225:
1516:: She's a beagle, true-bred, and one that adores me: what o' that?"
1417:
An American bred 15 inch male Beagle with the registered name of Ch
1331:
1094:
traditionally maintained Beagle packs. Protests were lodged against
4797:
4749:
4240:
3623:
2454:"Longevity of companion dog breeds: those at risk from early death"
1410:
1142:
1138:
1039:
1031:
1022:
1017:
990:
970:
898:
890:
644:
517:(AKC) in 1885. In the 20th century the breed has spread worldwide.
506:
478:
breeds. Stonehenge also gives the start of a standard description:
254:
243:
3563:
449:
Early images of the beagle (clockwise from top left): 1833, 1835,
5851:
5847:
5673:
5414:
5336:
5326:
5225:
5022:
4976:
4932:
4922:
4872:
4817:
4792:
3677:. Parliament of New South Wales. 13 December 2001. Archived from
1353:
1300:
1162:
1035:
883:
834:
820:
773:
Club standard which allows heights up to 16 inches (41 cm).
664:
648:
475:
303:
1853:"Beagle Breakthrough: Westminster Crowd Favorite Uno Is Top Dog"
675:
5835:
5213:
5047:
4927:
4907:
4807:
4458:
4453:
3398:"Animal-Testing Ban for Cosmetics to Be Upheld, EU Court Rules"
2149:"A NEWLY DISCOVERED "OLD" COLOR: DOMINO AKA PIED IN THE BEAGLE"
1369:
1358:
1157:, and foxes, and have even been recorded as being used to hunt
1150:
1146:
809:
765:
660:
4249:. Translated by Dakyns, H. G. Macmillan and Co. Archived from
3984:"Westminster Crowns 2020 Best in Show, Standard Poodle 'Siba'"
2832:
Rhodes, Karen Helton; Werner, Alexander H. (25 January 2011).
1173:
914:
breed, meaning that they are prone to types of disk diseases.
525:
5273:
5173:
5148:
4937:
4857:
4307:
4126:
The Field Book: Or, Sports and Pastimes of the United Kingdom
3619:"PETA calls for Dr. Fauci to resign: 'Our position is clear'"
3283:"Select Committee on Animals In Scientific Procedures Report"
3139:. Department of Primary Industries and Energy. Archived from
2834:
Blackwell's Five-Minute Veterinary Consult Clinical Companion
2520:"Factors related to longevity and mortality of dogs in Italy"
1158:
917:
656:
652:
417:
83:
3452:. US Department of Health and Human Services. Archived from
2451:
1106:. School and university packs are still maintained by Eton,
445:
315:
The origin of the beagle is not known. In the 11th century,
3132:
Nairn, M.E.; Allen, P.G.; Inglis, A.R.; Tanner, C. (1996).
2516:
2364:
Ranking of Dogs for Obedience/Working Intelligence by Breed
1405:
1225:
982:
978:
643:
The general appearance of the Beagle resembles a miniature
615:
250:
5266:
3897:. Lyndon Baines Johnson Library and Museum. Archived from
1540:, and again: "About her feet were little beagles seen" in
1251:
in the UK showed staff punching and screaming at Beagles.
933:
treatments. Another rare disease in the breed is neonatal
5734:
4947:
4276:
805:
62:
This article is about the dog breed. For other uses, see
4082:
Researches into the History of the British Dog Volume II
792:(Stonehenge) recommended a cross between a Beagle and a
416:
Reverend Phillip Honeywood established a beagle pack in
3426:. US Food and Drug Administration. 1999. Archived from
3765:"In Haiti, a shattered symbol reluctantly yields dead"
3131:
1455:, winner of the 2015 Westminster Kennel Club Dog Show
1161:. In most of these cases, the beagle is employed as a
261:
for prohibited agricultural imports and foodstuffs in
1265:
National Institute of Allergy and Infectious Diseases
3340:"Report of the Animal Procedures Committee for 2004"
2941:
Henry Salt: Humanitarian Reformer and Man of Letters
893:, but this can often be controlled with medication.
833:
for this reason, although their tendency to bark or
768:, a Beagle/pug cross, shows traits from both breeds.
363:
underbrush. Elizabeth I referred to the dogs as her
291:
Beagles have been depicted in popular culture since
3043:. The Mammal Society. 16 March 2000. Archived from
2464:(1). Springer Science and Business Media LLC: 531.
513:in 1887. The beagle was accepted as a breed by the
4204:
4149:
4101:
3093:"A beagle honoured as a defender at national gate"
1988:
1270:People for the Ethical Treatment of Animals (PETA)
1258:There are various examples of activists utilising
804:" and one of the most popular has been the Beagle/
411:
372:from 1866, the early 17th-century poet and writer
5864:
4207:The Encyclopedia of North American Sporting Dogs
3922:"Beagle, a Breed Long Unsung, Wins Best in Show"
1087:on the role of Huntsman on the day of the hunt.
457:'s 1852 "Beagle" image) and Dwarf Beagle (1859).
3476:"Beagle Freedom Law makes history in Minnesota"
3134:"Australian Quarantine a shared responsibility"
2743:
2074:. The Beagle Club of Queensland. Archived from
2768:
2589:
1272:made a call to action that all members of the
1073:Protection of Wild Mammals (Scotland) Act 2002
370:Researches into the History of the British Dog
5252:
4292:
4253:on 13 June 2007 – via eBooks@Adelaide).
3534:"Extremists seek fresh targets close to home"
2632:Erwin, W. Mark; Inman, Robert D. (May 2006).
1855:. Bloomberg. 12 February 2008. Archived from
1205:Beagles are the dog breed most often used in
623:hunting stags, it is today used for hare and
5422:
4008:
3793:. CBS-NY. Associated Press. 17 November 2011
2836:. Ames, Iowa: Wiley-Blackwell. p. 249.
2831:
2213:
2146:
1913:. American Kennel Club. 2006. Archived from
1839:
1789:
1765:
1195:Australian Quarantine and Inspection Service
593:in English literature dates from c. 1475 in
3978:
3976:
3791:"NYC Enlists Pair Of Bedbug-Detecting Dogs"
2687:"Neonatal Cerebellar Cortical Degeneration"
2275:
1391:which provided much of the inspiration for
242:, similar in appearance to the much larger
23:
5259:
5245:
4299:
4285:
4225:
3675:"Police Powers (Drug Detection Dogs) Bill"
3245:
3243:
2631:
2360:"The Intelligence of Dogs. Ranks 70 to 79"
2263:
2241:"Randall. The Patch Era at Beaver Meadows"
1735:
1432:and the pet name of "Miss P" won the 2015
1181:Beagles are used as detection dogs in the
311:is thought to be an ancestor of the beagle
265:around the world. The beagle is a popular
82:
24:
3699:
3500:
2944:. University of Illinois Press. pp.
2809:. Beagle Club of Victoria. Archived from
2545:
2535:
2493:
996:Beagles may exhibit a behaviour known as
246:. The beagle was developed primarily for
4239:
4099:
3986:. American Kennel Club. 11 February 2020
3973:
3919:
3725:
3531:
3400:. Bloomberg. 24 May 2005. Archived from
3027:
2937:
2565:
2563:
2561:
2559:
2557:
2427:
2415:
2276:Mirchandani, Raakhee (4 November 2005).
1750:
1708:
1330:
1287:
1226:Bans and activism against Beagle testing
1215:Animals (Scientific Procedures) Act 1986
1200:
1172:
1061:
1021:
916:
819:
759:
674:
634:
524:
444:
302:
153:Short haired, hard coat of medium length
32:This is an accepted version of this page
4144:
4122:
4039:
3913:
3811:
3334:
3332:
3240:
3023:
3021:
2913:
2439:
2391:"Canine Intelligence—Breed Does Matter"
2218:. Woods N' Water, Inc. pp. 22–23.
2166:"Beagle Coat Colors | Coats and Colors"
2108:
2040:
1827:
1777:
1623:
1621:
1421:and the pet name of "Uno" won the 2008
1367:, also features a beagle – the titular
1187:United States Department of Agriculture
663:; a broad chest narrowing to a tapered
14:
5865:
5267:Dogs originating in the United Kingdom
4257:
4027:
3920:Sandomir, Richard (13 February 2008).
3194:. Beagle Rescue League. Archived from
2925:
2855:
2853:
2573:. American Kennel Club. Archived from
2238:
2185:
2183:
2124:. American Kennel Club. Archived from
2063:
2061:
1965:. American Kennel Club. Archived from
1805:. American Kennel Club. Archived from
1761:
1759:
1731:
1729:
1674:. American Kennel Club. Archived from
1600:
1596:
1594:
1592:
1590:
610:It is not known why the black and tan
339:, though they are smaller and slower.
5550:English Toy Terrier (Black & Tan)
5240:
4280:
4227:Stonehenge (pseudonym of J. H. Walsh)
4202:
4191:
4156:. Weidenfeld & Nicolson History.
4133:
4088:
4079:
4058:
4009:Arnold, David; Arnold, Hazel (1998).
3814:"Doubts Rise on Bedbug-Sniffing Dogs"
3702:"The unlikely enemy of the terrorist"
3275:
2689:. Animal Health Trust. Archived from
2554:
2388:
2346:
2201:
2189:
2069:"What you need to know about Beagles"
1746:
1744:
1720:
1696:
1639:
1627:
1612:
1561:The History of Tom Jones, a Foundling
1326:
1299:Beagles are used as sniffer dogs for
200:Fédération Cynologique Internationale
5740:Dumfriesshire Black and Tan Foxhound
5129:Dumfriesshire Black and Tan Foxhound
4510:Grand Anglo-Français Blanc et Orange
4170:
3329:
3018:
2793:
2771:BSAVA Textbook of Veterinary Nursing
2619:
2609:from the original on 9 October 2022.
2395:Canine Corner. The human-animal bond
2052:
1618:
1191:Ministry of Agriculture and Forestry
396:and probably closely related to the
3532:Woolcock, Nicola (25 August 2005).
2850:
2644:(10). Spinejournal.com: 1094–1099.
2180:
2058:
1911:"AKC Breed Registration Statistics"
1891:from the original on 9 October 2022
1756:
1726:
1587:
1165:, flushing game for hunter's guns.
1026:The Caynsham Foot Beagles (c. 1885)
927:immune mediated polygenic arthritis
925:In rare cases, Beagles may develop
56:
4505:Grand Anglo-Français Blanc et Noir
4123:Maxwell, William Hamilton (1833).
3949:"Canadians triumph at Westminster"
3812:Buckley, Cara (11 November 2010).
3739:"Dog praised for life-saving call"
3002:. 20 November 2006. Archived from
2650:10.1097/01.brs.0000216593.97157.dd
2318:"Stars fuel designer dogs fashion"
1741:
1660:. Baldwin and Craddock: 164. 1833.
1652:"The fox-hound, and fox-hunting".
1504:
1485:
1473:
1440:
1075:, and in England and Wales by the
953:", a prolapse of the gland of the
736:
57:
5909:
5883:Dog breeds originating in England
4272:
3352:from the original on 10 July 2007
3263:from the original on 10 July 2007
2996:"Directory of UK hunts 2006/2007"
2869:(4). SAGE Publications: 337–345.
2328:from the original on 16 July 2007
1939:. The Kennel Club. Archived from
1702:
1664:
1260:Freedom of Information Act (FOIA)
707:
655:with a medium-length, square-cut
269:due to its size and good temper.
135:22–25 lb (10.0–11.3 kg)
4334:Anglo-Français de Petite Vénerie
3941:
3317:from the original on 2 July 2007
2304:from the original on 1 July 2007
2278:"Designer Dogs: Meet the Puggle"
1556:(published posthumously in 1749)
1434:Westminster Kennel Club Dog Show
1423:Westminster Kennel Club Dog Show
935:cerebellar cortical degeneration
829:easily won over. They make poor
145:20–23 lb (9.1–10.4 kg)
3887:
3862:
3831:
3805:
3783:
3757:
3731:
3719:
3700:Geoghegan, Tom (13 July 2005).
3693:
3667:
3645:
3631:
3611:
3597:
3577:
3556:
3525:
3494:
3468:
3442:
3416:
3390:
3364:
3297:
3210:
3184:
3158:
3085:
3059:
3033:
2988:
2962:
2931:
2919:
2907:
2897:
2825:
2799:
2787:
2762:
2737:
2705:
2679:
2625:
2613:
2537:10.1016/j.prevetmed.2024.106155
2510:
2445:
2433:
2421:
2409:
2389:Coren, Stanley (15 July 2009).
2382:
2352:
2340:
2269:
2257:
2232:
2207:
2195:
2158:
2140:
2114:
2102:
2093:
2046:
2034:
2018:. Breeds of Dog. Archived from
2008:
1981:
1955:
1929:
1903:
1871:
1845:
1833:
1821:
1795:
1783:
1771:
412:Development of the modern breed
4680:Schweizerischer Niederlaufhund
4515:Grand Anglo-Français Tricolore
4434:Chien Français Blanc et Orange
4235:. London: G. Routledge and Co.
4232:Manual of British Rural Sports
4152:The History of the Countryside
4011:A New Owner's Guide to Beagles
3851:, p. 54, 15 December 1974
2602:. The Beagle Club. June 2019.
2597:"The Beagle – A Healthy Breed"
2524:Preventive Veterinary Medicine
1884:. American Kennel Club. 2006.
1714:
1690:
1645:
1633:
1606:
1505:
1486:
1474:
1283:
815:
783:
472:Manual of British Rural Sports
13:
1:
5684:Cavalier King Charles Spaniel
3501:Broughton, Zoe (March 2001).
3450:"How to do Business with FDA"
2970:"Activists steal beagle pack"
2744:Gelatt, Kirk N., ed. (1999).
1879:"AKC Registration Statistics"
1459:
1116:Royal Agricultural University
796:as a retriever. He found the
731:
630:
520:
497:looking like straight-legged
124:13–16 in (33–41 cm)
4615:Petit Basset Griffon Vendéen
4520:Grand Basset Griffon Vendéen
4429:Chien Français Blanc et Noir
4349:Austrian Black and Tan Hound
4013:. T.F.H. Publications, Inc.
3839:"World's Most Famous Beagle"
3372:"Cosmetics and animal tests"
1274:National Institute of Health
1233:Food and Drug Administration
1168:
7:
5529:West Highland White Terrier
4720:Styrian Coarse-Haired Hound
4580:Istrian Coarse-haired Hound
4409:Bosnian Broken-haired Hound
4100:Kraeuter, Kristine (2001).
4084:. London: Robert Hardwicke.
2147:Beagle Brief (April 2020).
1522:(c. 1600) Act II Scene III
1430:Tashtins Lookin For Trouble
851:West Highland White Terrier
614:, present in Ireland since
607:which means "gate throat".
10:
5914:
5586:Staffordshire Bull Terrier
4605:Montenegrin Mountain Hound
4306:
4195:The Sportsman's Repository
4040:Daglish, E. Fitch (1961).
4035:. George Routledge and Co.
3895:"President Johnson's Dogs"
3564:"White Coat Waste Project"
3218:"Rescue + Freedom Project"
2875:10.1177/030098589503200401
2478:10.1038/s41598-023-50458-w
2280:. Fox News. Archived from
1672:"What is a Pocket Beagle?"
1547:Fables, Ancient and Modern
1528:: You are a sweet beagle"
1015:
1011:
298:
61:
59:Breed of small scent hound
5828:
5707:
5666:
5598:
5563:
5542:
5413:
5335:
5272:
5202:
5106:
5010:
5002:Treeing Tennessee Brindle
4946:
4773:
4585:Istrian Shorthaired Hound
4545:Griffon Fauve de Bretagne
4491:a/k/a Florida Cracker Cur
4314:
3874:National Purebred Dog Day
3769:Twin Cities Pioneer Press
3192:"Questions & Answers"
2938:Hendrick, George (1977).
1995:Oxford English Dictionary
1840:Arnold, & Arnold 1998
1790:Arnold, & Arnold 1998
1766:Arnold, & Arnold 1998
1654:The New Sporting Magazine
1554:To a Lady before Marriage
1534:(1607) Act III Scene IV:2
1351:since the 1950s with the
897:and a number of types of
877:
824:Beagles are happy to rest
599:. The origin of the word
586:Oxford English Dictionary
491:
453:'s Medium (1859, reusing
214:
203:
198:
191:
186:
181:
177:
168:
165:
160:
157:
152:
149:
139:
128:
123:
120:
115:
111:
103:
93:
81:
76:
5365:English Springer Spaniel
4735:Treeing Walker Coonhound
4540:Griffon Bleu de Gascogne
4439:Chien Français Tricolore
4364:Basset Fauve de Bretagne
4246:On Hunting (Cynegeticus)
3653:"Termite Detection Dogs"
3540:. London. Archived from
3222:Rescue + Freedom Project
2746:Veterinary Ophthalmology
2571:"Beagle Health Problems"
1803:"Get to know the Beagle"
1576:(1715–20) Book XV:697–8
1419:K-Run's Park Me In First
1394:On the Origin of Species
1249:Huntingdon Life Sciences
889:Beagles may be prone to
867:The Intelligence of Dogs
860:They are ranked 72nd in
596:The Squire of Low Degree
557:K-Run's Park Me In First
39:latest accepted revision
5770:Old Welsh Grey Sheepdog
5164:Old Croatian Sighthound
5134:Grand Fauve de Bretagne
4765:Westphalian Dachsbracke
4705:Serbian Tricolour Hound
4419:Briquet Griffon Vendéen
4394:Black and Tan Coonhound
4374:Bavarian Mountain Hound
4359:Basset Bleu de Gascogne
4354:Basset Artésien Normand
4137:The Sportsman's Library
2298:"Designing A Cuter Dog"
1104:Animal Liberation Front
1100:Wye College beagle pack
904:Musladin-Lueke syndrome
589:, the first mention of
578:
553:Westminster Kennel Club
432:The Sportsman's Library
64:Beagle (disambiguation)
5581:Miniature Bull Terrier
5519:Sporting Lucas Terrier
5484:Parson Russell Terrier
5454:Dandie Dinmont Terrier
5405:Welsh Springer Spaniel
5355:English Cocker Spaniel
5350:Curly-Coated Retriever
5018:American Leopard Hound
4685:Segugio dell'Appennino
4620:Petit Bleu de Gascogne
4525:Grand Bleu de Gascogne
4211:. Willow Creek Press.
4089:Jesse, Edward (1858).
4080:Jesse, George (1866).
3870:"Gromit: Purebred Dog"
3521:– via dbsst.org.
3480:Minnesota Star Tribune
3285:. House of Lords. 2002
2300:. CBS. 6 August 2006.
1499:
1336:
1293:
1292:Red & white Beagle
1178:
1067:
1060:
1027:
922:
825:
769:
680:
640:
530:
485:
458:
382:
312:
5750:English White Terrier
5745:English Water Spaniel
5715:Black and Tan Terrier
5375:Flat-Coated Retriever
4967:Catahoula Leopard Dog
4655:Rastreador Brasileiro
4530:Grand Griffon Vendéen
4203:Smith, Steve (2002).
4065:. Howell Book House.
4059:Fogle, Bruce (1990).
3844:The Victoria Advocate
3345:. Home Office. 2004.
3310:. Home Office. 2004.
3256:. Home Office. 2005.
3097:People's Daily Online
2214:Fisher, Dave (2002).
1568:The Progress of Error
1494:
1378:Lyndon Baines Johnson
1334:
1317:search and rescue dog
1313:2010 Haiti earthquake
1291:
1231:not specified by the
1201:Use in animal testing
1176:
1120:Christ Church, Oxford
1065:
1055:
1025:
920:
823:
763:
678:
638:
528:
480:
448:
378:
317:William the Conqueror
306:
182:Kennel club standards
5760:North Country Beagle
5694:King Charles Spaniel
5633:Pembroke Welsh Corgi
5628:Old English Sheepdog
5618:Cardigan Welsh Corgi
5459:Jack Russell Terrier
5322:West Country Harrier
5159:North Country Beagle
4760:West Country Harrier
4444:Colombian Fino Hound
4267:. Blanchard and Lea.
4192:Scott, John (1845).
4134:Mills, John (1845).
4104:Training Your Beagle
3681:on 29 September 2007
3404:on 30 September 2007
2863:Veterinary Pathology
2253:on 14 November 2011.
2081:on 25 September 2011
1538:The Cock and the Fox
1526:Mistress Tenterhook'
1376:Former US President
1343:Beagles appeared in
1193:in New Zealand, the
747:Canadian Kennel Club
743:American Kennel Club
724:took 15 minutes and
561:American Kennel Club
515:American Kennel Club
390:North Country Beagle
278:North Country Beagle
5893:Hunting with hounds
5815:Tweed Water Spaniel
5805:Toy Trawler Spaniel
5765:Old English Bulldog
5730:Cumberland Sheepdog
5210:Anglo-French hounds
5083:Rhodesian Ridgeback
4730:Transylvanian Hound
4495:Gascon Saintongeois
4414:Briquet de Provence
4174:The Beagle Handbook
3961:on 18 February 2015
3593:. 15 November 2016.
3590:The Washington Post
2470:2024NatSR..14..531M
2324:. 6 February 2006.
2168:. 25 September 2023
1465:Informational notes
967:autosomal recessive
847:Miniature Schnauzer
284:, and possibly the
29:Page version status
5699:Northern Inuit Dog
5514:Smooth Fox Terrier
5489:Patterdale Terrier
5469:Manchester Terrier
5439:Bedlington Terrier
5390:Labrador Retriever
5312:Scottish Deerhound
5139:King's White Hound
5093:Xarnego Valenciano
5078:Portuguese Podengo
5063:Phu Quoc Ridgeback
5038:Haofa Tangkhul Hui
4913:Scottish Deerhound
4788:American Staghound
4675:Schweizer Laufhund
4640:Polish Hunting Dog
4404:Bluetick Coonhound
4324:Alpine Dachsbracke
4171:Rice, Dan (2000).
4044:. London: Foyles.
3927:The New York Times
3876:, 13 December 2018
3641:. 25 October 2021.
3627:. 5 November 2021.
3544:on 12 January 2008
3073:on 2 February 2007
2458:Scientific Reports
2397:. Psychology Today
2370:on 24 October 2011
1642:, pp. 223–232
1574:The Iliad of Homer
1542:Palamon and Arcite
1383:The ship on which
1364:Wallace and Gromit
1337:
1327:In popular culture
1294:
1179:
1131:cottontail rabbits
1068:
1028:
923:
872:separation anxiety
826:
770:
698:Bluetick Coonhound
681:
641:
565:Labrador Retriever
531:
470:), writing in the
459:
313:
35:
5860:
5859:
5720:Blue Paul Terrier
5643:Shetland Sheepdog
5623:Lancashire Heeler
5594:
5593:
5555:Yorkshire Terrier
5234:
5233:
5169:Potsdam Greyhound
5028:Cirneco dell'Etna
4843:Italian Greyhound
4725:Tarsus çatalburun
4695:Segugio Maremmano
4660:Redbone Coonhound
4550:Griffon Nivernais
4489:Florida Brown Dog
4464:English Coonhound
4329:American Foxhound
4092:Anecdotes of Dogs
3771:. 25 January 2010
3706:BBC News Magazine
3172:on 10 August 2007
3099:. 6 December 2006
3030:, pp. 97–104
3006:on 6 October 2007
3000:Horse & Hound
2843:978-0-8138-1596-1
2796:, pp. 167–74
2780:978-1-910-44339-2
2725:on 6 October 2014
2022:on 13 August 2007
1564:(1749) Chapter 7.
1349:animated cartoons
1102:was taken by the
1047:Anecdotes of Dogs
947:corneal dystrophy
912:chondrodystrophic
839:Yorkshire Terrier
808:cross known as a
726:Scottish Terriers
583:According to the
360:Queen Elizabeth I
228:
227:
210:
209:
173:
172:
47:27 September 2024
26:
16:(Redirected from
5905:
5571:Bull and terrier
5534:Wire Fox Terrier
5504:Sealyham Terrier
5499:Scottish Terrier
5464:Lakeland Terrier
5434:Airedale Terrier
5420:
5419:
5380:Golden Retriever
5292:English Foxhound
5261:
5254:
5247:
5238:
5237:
5053:Pampas Deerhound
4918:Silken Windhound
4903:Rampur Greyhound
4893:Polish Greyhound
4690:Segugio Italiano
4600:Lithuanian Hound
4469:English Foxhound
4424:Bruno Jura Hound
4301:
4294:
4287:
4278:
4277:
4268:
4254:
4236:
4222:
4210:
4199:
4198:. Henry G. Bohn.
4188:
4167:
4155:
4141:
4130:
4119:
4107:
4096:
4085:
4076:
4055:
4036:
4024:
3996:
3995:
3993:
3991:
3980:
3971:
3970:
3968:
3966:
3957:. Archived from
3945:
3939:
3938:
3936:
3934:
3917:
3911:
3910:
3908:
3906:
3891:
3885:
3884:
3883:
3881:
3866:
3860:
3859:
3858:
3856:
3835:
3829:
3828:
3826:
3824:
3809:
3803:
3802:
3800:
3798:
3787:
3781:
3780:
3778:
3776:
3761:
3755:
3754:
3752:
3750:
3735:
3729:
3728:, pp. 89–92
3723:
3717:
3716:
3714:
3712:
3697:
3691:
3690:
3688:
3686:
3671:
3665:
3664:
3662:
3660:
3649:
3643:
3642:
3635:
3629:
3628:
3615:
3609:
3608:
3601:
3595:
3594:
3581:
3575:
3574:
3572:
3570:
3560:
3554:
3553:
3551:
3549:
3529:
3523:
3522:
3520:
3518:
3509:. Archived from
3498:
3492:
3491:
3489:
3487:
3472:
3466:
3465:
3463:
3461:
3456:on 3 August 2005
3446:
3440:
3439:
3437:
3435:
3424:"Animal Testing"
3420:
3414:
3413:
3411:
3409:
3394:
3388:
3387:
3385:
3383:
3368:
3362:
3361:
3359:
3357:
3351:
3344:
3336:
3327:
3326:
3324:
3322:
3316:
3309:
3301:
3295:
3294:
3292:
3290:
3279:
3273:
3272:
3270:
3268:
3262:
3255:
3247:
3238:
3237:
3235:
3233:
3228:on 4 August 2018
3224:. Archived from
3214:
3208:
3207:
3205:
3203:
3198:on 4 August 2018
3188:
3182:
3181:
3179:
3177:
3162:
3156:
3155:
3153:
3151:
3145:
3138:
3128:
3126:
3124:
3119:on 31 March 2012
3108:
3106:
3104:
3089:
3083:
3082:
3080:
3078:
3063:
3057:
3056:
3054:
3052:
3037:
3031:
3025:
3016:
3015:
3013:
3011:
2992:
2986:
2985:
2983:
2981:
2976:. 5 January 2001
2966:
2960:
2959:
2935:
2929:
2923:
2917:
2911:
2905:
2901:
2895:
2894:
2857:
2848:
2847:
2829:
2823:
2822:
2820:
2818:
2813:on 4 August 2007
2803:
2797:
2791:
2785:
2784:
2766:
2760:
2759:
2741:
2735:
2734:
2732:
2730:
2724:
2717:
2709:
2703:
2702:
2700:
2698:
2693:on 23 March 2013
2683:
2677:
2676:
2674:
2672:
2629:
2623:
2617:
2611:
2610:
2608:
2601:
2593:
2587:
2586:
2584:
2582:
2567:
2552:
2551:
2549:
2539:
2514:
2508:
2507:
2497:
2449:
2443:
2437:
2431:
2425:
2419:
2413:
2407:
2406:
2404:
2402:
2386:
2380:
2379:
2377:
2375:
2366:. Archived from
2356:
2350:
2349:, pp. 176–7
2344:
2338:
2337:
2335:
2333:
2313:
2311:
2309:
2293:
2291:
2289:
2273:
2267:
2261:
2255:
2254:
2252:
2246:. Archived from
2245:
2236:
2230:
2229:
2211:
2205:
2199:
2193:
2187:
2178:
2177:
2175:
2173:
2162:
2156:
2155:
2153:
2144:
2138:
2137:
2135:
2133:
2118:
2112:
2106:
2100:
2097:
2091:
2090:
2088:
2086:
2080:
2073:
2065:
2056:
2050:
2044:
2038:
2032:
2031:
2029:
2027:
2012:
2006:
2005:
2003:
2001:
1992:
1985:
1979:
1978:
1976:
1974:
1959:
1953:
1952:
1950:
1948:
1933:
1927:
1926:
1924:
1922:
1907:
1901:
1900:
1898:
1896:
1890:
1883:
1875:
1869:
1868:
1866:
1864:
1859:on 13 April 2008
1849:
1843:
1837:
1831:
1830:, pp. 10–12
1825:
1819:
1818:
1816:
1814:
1809:on 30 March 2014
1799:
1793:
1787:
1781:
1775:
1769:
1763:
1754:
1748:
1739:
1733:
1724:
1718:
1712:
1706:
1700:
1694:
1688:
1687:
1685:
1683:
1668:
1662:
1661:
1649:
1643:
1637:
1631:
1630:, pp. 438–9
1625:
1616:
1610:
1604:
1598:
1509:
1490:
1478:
1077:Hunting Act 2004
998:reverse sneezing
969:mutation in the
794:Scottish Terrier
790:John Henry Walsh
751:Kennel Club (UK)
573:Golden Retriever
468:John Henry Walsh
335:and the extinct
321:St. Hubert Hound
223:
217:
179:
178:
113:
112:
96:
86:
74:
73:
21:
5913:
5912:
5908:
5907:
5906:
5904:
5903:
5902:
5863:
5862:
5861:
5856:
5824:
5775:Paisley Terrier
5755:Norfolk Spaniel
5703:
5689:English Mastiff
5662:
5590:
5559:
5538:
5494:Plummer Terrier
5479:Norwich Terrier
5474:Norfolk Terrier
5426:
5409:
5345:Clumber Spaniel
5331:
5268:
5265:
5235:
5230:
5198:
5102:
5073:Podenco Canario
5068:Podenco Andaluz
5006:
4957:Black Mouth Cur
4942:
4838:Irish Wolfhound
4833:Hortaya borzaya
4769:
4710:SlovenskĂ˝ Kopov
4670:Schillerstövare
4665:Sabueso Español
4560:Hamiltonstövare
4535:Greek Harehound
4310:
4305:
4275:
4259:Youatt, William
4219:
4185:
4164:
4146:Rackham, Oliver
4116:
4073:
4052:
4021:
4000:
3999:
3989:
3987:
3982:
3981:
3974:
3964:
3962:
3947:
3946:
3942:
3932:
3930:
3918:
3914:
3904:
3902:
3901:on 11 July 2007
3893:
3892:
3888:
3879:
3877:
3868:
3867:
3863:
3854:
3852:
3849:Victoria, Texas
3837:
3836:
3832:
3822:
3820:
3810:
3806:
3796:
3794:
3789:
3788:
3784:
3774:
3772:
3763:
3762:
3758:
3748:
3746:
3737:
3736:
3732:
3724:
3720:
3710:
3708:
3698:
3694:
3684:
3682:
3673:
3672:
3668:
3658:
3656:
3655:. K9 Centre.com
3651:
3650:
3646:
3637:
3636:
3632:
3617:
3616:
3612:
3607:. 30 July 2021.
3603:
3602:
3598:
3583:
3582:
3578:
3568:
3566:
3562:
3561:
3557:
3547:
3545:
3530:
3526:
3516:
3514:
3499:
3495:
3485:
3483:
3474:
3473:
3469:
3459:
3457:
3448:
3447:
3443:
3433:
3431:
3422:
3421:
3417:
3407:
3405:
3396:
3395:
3391:
3381:
3379:
3378:on 29 June 2007
3370:
3369:
3365:
3355:
3353:
3349:
3342:
3338:
3337:
3330:
3320:
3318:
3314:
3307:
3303:
3302:
3298:
3288:
3286:
3281:
3280:
3276:
3266:
3264:
3260:
3253:
3249:
3248:
3241:
3231:
3229:
3216:
3215:
3211:
3201:
3199:
3190:
3189:
3185:
3175:
3173:
3164:
3163:
3159:
3149:
3147:
3146:on 10 July 2007
3143:
3136:
3129:
3122:
3120:
3111:
3109:
3102:
3100:
3091:
3090:
3086:
3076:
3074:
3065:
3064:
3060:
3050:
3048:
3039:
3038:
3034:
3026:
3019:
3009:
3007:
2994:
2993:
2989:
2979:
2977:
2968:
2967:
2963:
2956:
2936:
2932:
2924:
2920:
2912:
2908:
2902:
2898:
2858:
2851:
2844:
2830:
2826:
2816:
2814:
2805:
2804:
2800:
2792:
2788:
2781:
2767:
2763:
2756:
2742:
2738:
2728:
2726:
2722:
2715:
2711:
2710:
2706:
2696:
2694:
2685:
2684:
2680:
2670:
2668:
2630:
2626:
2618:
2614:
2606:
2599:
2595:
2594:
2590:
2580:
2578:
2577:on 11 June 2007
2569:
2568:
2555:
2515:
2511:
2450:
2446:
2438:
2434:
2426:
2422:
2418:, pp. 77–8
2414:
2410:
2400:
2398:
2387:
2383:
2373:
2371:
2358:
2357:
2353:
2345:
2341:
2331:
2329:
2316:
2314:
2307:
2305:
2296:
2294:
2287:
2285:
2284:on 29 June 2007
2274:
2270:
2264:Stonehenge 1856
2262:
2258:
2250:
2243:
2237:
2233:
2226:
2212:
2208:
2200:
2196:
2188:
2181:
2171:
2169:
2164:
2163:
2159:
2151:
2145:
2141:
2131:
2129:
2128:on 29 June 2007
2122:"Beagle Colors"
2120:
2119:
2115:
2107:
2103:
2098:
2094:
2084:
2082:
2078:
2071:
2067:
2066:
2059:
2051:
2047:
2039:
2035:
2025:
2023:
2014:
2013:
2009:
1999:
1997:
1987:
1986:
1982:
1972:
1970:
1969:on 5 April 2014
1961:
1960:
1956:
1946:
1944:
1943:on 28 June 2007
1935:
1934:
1930:
1920:
1918:
1917:on 16 July 2007
1909:
1908:
1904:
1894:
1892:
1888:
1881:
1877:
1876:
1872:
1862:
1860:
1851:
1850:
1846:
1842:, pp. 14–5
1838:
1834:
1826:
1822:
1812:
1810:
1801:
1800:
1796:
1788:
1784:
1776:
1772:
1764:
1757:
1749:
1742:
1738:, pp. 98–9
1736:Stonehenge 1856
1734:
1727:
1723:, pp. 75–8
1719:
1715:
1707:
1703:
1695:
1691:
1681:
1679:
1670:
1669:
1665:
1651:
1650:
1646:
1638:
1634:
1626:
1619:
1611:
1607:
1599:
1588:
1578:
1571:
1565:
1557:
1551:
1535:
1523:
1511:
1492:
1462:
1443:
1441:Notable Beagles
1373:– in its cast.
1341:Anthropomorphic
1335:Tricolor Beagle
1329:
1286:
1278:Francis Collins
1253:Consort Kennels
1228:
1203:
1171:
1020:
1014:
963:retinal atrophy
880:
818:
786:
739:
737:Breed varieties
734:
710:
633:
618:times, has the
581:
569:German Shepherd
523:
494:
425:suspected that
414:
374:Gervase Markham
365:singing beagles
301:
219:
215:
188:The Kennel Club
94:
89:
88:Tricolor Beagle
72:
67:
60:
55:
54:
53:
52:
51:
50:
34:
22:
15:
12:
11:
5:
5911:
5901:
5900:
5895:
5890:
5885:
5880:
5878:Companion dogs
5875:
5858:
5857:
5855:
5854:
5829:
5826:
5825:
5823:
5822:
5817:
5812:
5807:
5802:
5797:
5792:
5787:
5782:
5780:Southern Hound
5777:
5772:
5767:
5762:
5757:
5752:
5747:
5742:
5737:
5732:
5727:
5722:
5717:
5711:
5709:
5705:
5704:
5702:
5701:
5696:
5691:
5686:
5681:
5676:
5670:
5668:
5664:
5663:
5661:
5660:
5658:Welsh Sheepdog
5655:
5650:
5645:
5640:
5635:
5630:
5625:
5620:
5615:
5610:
5608:Bearded Collie
5604:
5602:
5596:
5595:
5592:
5591:
5589:
5588:
5583:
5578:
5573:
5567:
5565:
5561:
5560:
5558:
5557:
5552:
5546:
5544:
5540:
5539:
5537:
5536:
5531:
5526:
5521:
5516:
5511:
5506:
5501:
5496:
5491:
5486:
5481:
5476:
5471:
5466:
5461:
5456:
5451:
5446:
5444:Border Terrier
5441:
5436:
5430:
5428:
5417:
5411:
5410:
5408:
5407:
5402:
5400:Sussex Spaniel
5397:
5392:
5387:
5382:
5377:
5372:
5367:
5362:
5360:English Setter
5357:
5352:
5347:
5341:
5339:
5333:
5332:
5330:
5329:
5324:
5319:
5314:
5309:
5304:
5299:
5294:
5289:
5284:
5278:
5276:
5270:
5269:
5264:
5263:
5256:
5249:
5241:
5232:
5231:
5229:
5228:
5203:
5200:
5199:
5197:
5196:
5191:
5186:
5184:Southern Hound
5181:
5176:
5171:
5166:
5161:
5156:
5151:
5146:
5141:
5136:
5131:
5126:
5121:
5116:
5110:
5108:
5104:
5103:
5101:
5100:
5095:
5090:
5088:Thai Ridgeback
5085:
5080:
5075:
5070:
5065:
5060:
5055:
5050:
5045:
5040:
5035:
5030:
5025:
5020:
5014:
5012:
5008:
5007:
5005:
5004:
4999:
4994:
4992:Stephens Stock
4989:
4987:Mountain Feist
4984:
4979:
4974:
4969:
4964:
4959:
4953:
4951:
4950:hunting breeds
4944:
4943:
4941:
4940:
4935:
4930:
4925:
4920:
4915:
4910:
4905:
4900:
4895:
4890:
4885:
4883:Mahratta Hound
4880:
4875:
4870:
4868:Levriero Sardo
4865:
4860:
4855:
4850:
4845:
4840:
4835:
4830:
4825:
4820:
4815:
4810:
4805:
4800:
4795:
4790:
4785:
4779:
4777:
4771:
4770:
4768:
4767:
4762:
4757:
4755:Welsh Foxhound
4752:
4747:
4745:Tyrolean Hound
4742:
4737:
4732:
4727:
4722:
4717:
4715:Smalandstõvare
4712:
4707:
4702:
4697:
4692:
4687:
4682:
4677:
4672:
4667:
4662:
4657:
4652:
4647:
4642:
4637:
4632:
4627:
4622:
4617:
4612:
4607:
4602:
4597:
4592:
4587:
4582:
4577:
4572:
4567:
4562:
4557:
4552:
4547:
4542:
4537:
4532:
4527:
4522:
4517:
4512:
4507:
4502:
4497:
4492:
4486:
4481:
4479:Estonian Hound
4476:
4471:
4466:
4461:
4456:
4451:
4446:
4441:
4436:
4431:
4426:
4421:
4416:
4411:
4406:
4401:
4396:
4391:
4386:
4384:Beagle-Harrier
4381:
4376:
4371:
4366:
4361:
4356:
4351:
4346:
4341:
4336:
4331:
4326:
4320:
4318:
4312:
4311:
4304:
4303:
4296:
4289:
4281:
4274:
4273:External links
4271:
4270:
4269:
4255:
4237:
4223:
4217:
4200:
4189:
4183:
4168:
4162:
4142:
4140:. W. Paterson.
4131:
4120:
4114:
4097:
4086:
4077:
4071:
4062:The Dog's Mind
4056:
4050:
4037:
4029:Blakey, Robert
4025:
4019:
3998:
3997:
3972:
3940:
3912:
3886:
3861:
3830:
3818:New York Times
3804:
3782:
3756:
3745:. 20 June 2006
3730:
3718:
3692:
3666:
3644:
3630:
3610:
3596:
3576:
3555:
3524:
3513:on 1 July 2015
3493:
3467:
3441:
3430:on 23 May 2007
3415:
3389:
3363:
3328:
3296:
3274:
3239:
3209:
3183:
3157:
3084:
3058:
3047:on 12 May 2010
3032:
3017:
2987:
2961:
2954:
2930:
2918:
2906:
2896:
2849:
2842:
2824:
2798:
2786:
2779:
2761:
2754:
2736:
2704:
2678:
2624:
2612:
2588:
2553:
2509:
2444:
2432:
2420:
2408:
2381:
2351:
2339:
2268:
2256:
2231:
2224:
2216:Rabbit hunting
2206:
2194:
2179:
2157:
2139:
2113:
2101:
2092:
2057:
2045:
2033:
2016:"Kerry Beagle"
2007:
1980:
1954:
1928:
1902:
1870:
1844:
1832:
1820:
1794:
1782:
1770:
1755:
1740:
1725:
1713:
1701:
1689:
1678:on 9 June 2007
1663:
1644:
1632:
1617:
1605:
1585:
1584:
1514:Sir Toby Belch
1512:Shakespeare: "
1468:
1461:
1458:
1457:
1456:
1450:
1442:
1439:
1438:
1437:
1426:
1415:
1385:Charles Darwin
1381:
1374:
1328:
1325:
1309:assistance dog
1285:
1282:
1237:miniature pigs
1227:
1224:
1207:animal testing
1202:
1199:
1183:Beagle Brigade
1170:
1167:
1092:public schools
1016:Main article:
1013:
1010:
895:hypothyroidism
879:
876:
817:
814:
788:In the 1850s,
785:
782:
738:
735:
733:
730:
712:Alongside the
709:
708:Sense of smell
706:
632:
629:
580:
577:
522:
519:
511:Norman Ellmore
503:Richard Rowett
493:
490:
440:Lord Winterton
423:William Youatt
413:
410:
386:Southern Hound
337:Southern Hound
309:Southern Hound
300:
297:
282:Southern Hound
226:
225:
212:
211:
208:
207:
202:
196:
195:
190:
184:
183:
175:
174:
171:
170:
167:
166:Life span
163:
162:
159:
155:
154:
151:
147:
146:
143:
140:
137:
136:
133:
130:
126:
125:
122:
118:
117:
109:
108:
105:
101:
100:
99:English Beagle
97:
91:
90:
87:
79:
78:
70:
58:
36:
30:
27:
25:
9:
6:
4:
3:
2:
5910:
5899:
5896:
5894:
5891:
5889:
5886:
5884:
5881:
5879:
5876:
5874:
5871:
5870:
5868:
5853:
5849:
5845:
5841:
5840:Fell Terriers
5837:
5834:
5831:
5830:
5827:
5821:
5820:Welsh Hillman
5818:
5816:
5813:
5811:
5808:
5806:
5803:
5801:
5798:
5796:
5793:
5791:
5788:
5786:
5783:
5781:
5778:
5776:
5773:
5771:
5768:
5766:
5763:
5761:
5758:
5756:
5753:
5751:
5748:
5746:
5743:
5741:
5738:
5736:
5733:
5731:
5728:
5726:
5723:
5721:
5718:
5716:
5713:
5712:
5710:
5706:
5700:
5697:
5695:
5692:
5690:
5687:
5685:
5682:
5680:
5677:
5675:
5672:
5671:
5669:
5665:
5659:
5656:
5654:
5653:Smooth Collie
5651:
5649:
5646:
5644:
5641:
5639:
5636:
5634:
5631:
5629:
5626:
5624:
5621:
5619:
5616:
5614:
5613:Border Collie
5611:
5609:
5606:
5605:
5603:
5601:
5597:
5587:
5584:
5582:
5579:
5577:
5574:
5572:
5569:
5568:
5566:
5562:
5556:
5553:
5551:
5548:
5547:
5545:
5541:
5535:
5532:
5530:
5527:
5525:
5524:Welsh Terrier
5522:
5520:
5517:
5515:
5512:
5510:
5507:
5505:
5502:
5500:
5497:
5495:
5492:
5490:
5487:
5485:
5482:
5480:
5477:
5475:
5472:
5470:
5467:
5465:
5462:
5460:
5457:
5455:
5452:
5450:
5449:Cairn Terrier
5447:
5445:
5442:
5440:
5437:
5435:
5432:
5431:
5429:
5425:
5421:
5418:
5416:
5412:
5406:
5403:
5401:
5398:
5396:
5393:
5391:
5388:
5386:
5385:Gordon Setter
5383:
5381:
5378:
5376:
5373:
5371:
5370:Field Spaniel
5368:
5366:
5363:
5361:
5358:
5356:
5353:
5351:
5348:
5346:
5343:
5342:
5340:
5338:
5334:
5328:
5325:
5323:
5320:
5318:
5315:
5313:
5310:
5308:
5305:
5303:
5300:
5298:
5295:
5293:
5290:
5288:
5285:
5283:
5280:
5279:
5277:
5275:
5271:
5262:
5257:
5255:
5250:
5248:
5243:
5242:
5239:
5227:
5223:
5219:
5215:
5211:
5208:
5205:
5204:
5201:
5195:
5192:
5190:
5187:
5185:
5182:
5180:
5177:
5175:
5172:
5170:
5167:
5165:
5162:
5160:
5157:
5155:
5152:
5150:
5147:
5145:
5142:
5140:
5137:
5135:
5132:
5130:
5127:
5125:
5122:
5120:
5117:
5115:
5112:
5111:
5109:
5105:
5099:
5096:
5094:
5091:
5089:
5086:
5084:
5081:
5079:
5076:
5074:
5071:
5069:
5066:
5064:
5061:
5059:
5058:Pharaoh Hound
5056:
5054:
5051:
5049:
5046:
5044:
5041:
5039:
5036:
5034:
5031:
5029:
5026:
5024:
5021:
5019:
5016:
5015:
5013:
5011:Miscellaneous
5009:
5003:
5000:
4998:
4995:
4993:
4990:
4988:
4985:
4983:
4980:
4978:
4975:
4973:
4970:
4968:
4965:
4963:
4960:
4958:
4955:
4954:
4952:
4949:
4945:
4939:
4936:
4934:
4931:
4929:
4926:
4924:
4921:
4919:
4916:
4914:
4911:
4909:
4906:
4904:
4901:
4899:
4896:
4894:
4891:
4889:
4886:
4884:
4881:
4879:
4876:
4874:
4871:
4869:
4866:
4864:
4861:
4859:
4856:
4854:
4851:
4849:
4846:
4844:
4841:
4839:
4836:
4834:
4831:
4829:
4826:
4824:
4823:Galgo Español
4821:
4819:
4816:
4814:
4811:
4809:
4806:
4804:
4803:Banjara Hound
4801:
4799:
4796:
4794:
4791:
4789:
4786:
4784:
4781:
4780:
4778:
4776:
4772:
4766:
4763:
4761:
4758:
4756:
4753:
4751:
4748:
4746:
4743:
4741:
4738:
4736:
4733:
4731:
4728:
4726:
4723:
4721:
4718:
4716:
4713:
4711:
4708:
4706:
4703:
4701:
4700:Serbian Hound
4698:
4696:
4693:
4691:
4688:
4686:
4683:
4681:
4678:
4676:
4673:
4671:
4668:
4666:
4663:
4661:
4658:
4656:
4653:
4651:
4650:Posavac Hound
4648:
4646:
4643:
4641:
4638:
4636:
4633:
4631:
4628:
4626:
4623:
4621:
4618:
4616:
4613:
4611:
4608:
4606:
4603:
4601:
4598:
4596:
4595:Liangshan Dog
4593:
4591:
4588:
4586:
4583:
4581:
4578:
4576:
4573:
4571:
4568:
4566:
4565:Hanover Hound
4563:
4561:
4558:
4556:
4553:
4551:
4548:
4546:
4543:
4541:
4538:
4536:
4533:
4531:
4528:
4526:
4523:
4521:
4518:
4516:
4513:
4511:
4508:
4506:
4503:
4501:
4498:
4496:
4493:
4490:
4487:
4485:
4484:Finnish Hound
4482:
4480:
4477:
4475:
4472:
4470:
4467:
4465:
4462:
4460:
4457:
4455:
4452:
4450:
4447:
4445:
4442:
4440:
4437:
4435:
4432:
4430:
4427:
4425:
4422:
4420:
4417:
4415:
4412:
4410:
4407:
4405:
4402:
4400:
4397:
4395:
4392:
4390:
4387:
4385:
4382:
4380:
4377:
4375:
4372:
4370:
4367:
4365:
4362:
4360:
4357:
4355:
4352:
4350:
4347:
4345:
4342:
4340:
4337:
4335:
4332:
4330:
4327:
4325:
4322:
4321:
4319:
4317:
4313:
4309:
4302:
4297:
4295:
4290:
4288:
4283:
4282:
4279:
4266:
4265:
4260:
4256:
4252:
4248:
4247:
4242:
4238:
4234:
4233:
4228:
4224:
4220:
4218:1-57223-501-2
4214:
4209:
4208:
4201:
4197:
4196:
4190:
4186:
4184:0-7641-1464-6
4180:
4176:
4175:
4169:
4165:
4163:1-84212-440-4
4159:
4154:
4153:
4147:
4143:
4139:
4138:
4132:
4128:
4127:
4121:
4117:
4115:0-7641-1648-7
4111:
4106:
4105:
4098:
4095:. H. G. Bohn.
4094:
4093:
4087:
4083:
4078:
4074:
4072:0-87605-513-7
4068:
4064:
4063:
4057:
4053:
4051:0-7071-0631-1
4047:
4043:
4038:
4034:
4030:
4026:
4022:
4020:0-7938-2785-X
4016:
4012:
4007:
4006:
4005:
4004:
3985:
3979:
3977:
3960:
3956:
3955:
3950:
3944:
3929:
3928:
3923:
3916:
3900:
3896:
3890:
3875:
3871:
3865:
3850:
3846:
3845:
3840:
3834:
3819:
3815:
3808:
3792:
3786:
3770:
3766:
3760:
3744:
3740:
3734:
3727:
3726:Kraeuter 2001
3722:
3707:
3703:
3696:
3680:
3676:
3670:
3654:
3648:
3640:
3634:
3626:
3625:
3620:
3614:
3606:
3600:
3592:
3591:
3586:
3580:
3565:
3559:
3543:
3539:
3535:
3528:
3512:
3508:
3507:The Ecologist
3504:
3497:
3482:. 21 May 2014
3481:
3477:
3471:
3455:
3451:
3445:
3429:
3425:
3419:
3403:
3399:
3393:
3377:
3373:
3367:
3348:
3341:
3335:
3333:
3313:
3306:
3300:
3284:
3278:
3259:
3252:
3246:
3244:
3227:
3223:
3219:
3213:
3197:
3193:
3187:
3171:
3167:
3161:
3142:
3135:
3118:
3114:
3098:
3094:
3088:
3072:
3068:
3062:
3046:
3042:
3036:
3029:
3028:Kraeuter 2001
3024:
3022:
3005:
3001:
2997:
2991:
2975:
2971:
2965:
2957:
2955:0-252-00611-9
2951:
2947:
2943:
2942:
2934:
2927:
2922:
2915:
2910:
2900:
2892:
2888:
2884:
2880:
2876:
2872:
2868:
2864:
2856:
2854:
2845:
2839:
2835:
2828:
2812:
2808:
2802:
2795:
2790:
2782:
2776:
2772:
2765:
2757:
2755:0-683-30076-8
2751:
2747:
2740:
2721:
2714:
2708:
2692:
2688:
2682:
2667:
2663:
2659:
2655:
2651:
2647:
2643:
2639:
2635:
2628:
2622:, p. 161
2621:
2616:
2605:
2598:
2592:
2576:
2572:
2566:
2564:
2562:
2560:
2558:
2548:
2543:
2538:
2533:
2529:
2525:
2521:
2513:
2505:
2501:
2496:
2491:
2487:
2483:
2479:
2475:
2471:
2467:
2463:
2459:
2455:
2448:
2441:
2436:
2429:
2428:Kraeuter 2001
2424:
2417:
2416:Kraeuter 2001
2412:
2396:
2392:
2385:
2369:
2365:
2361:
2355:
2348:
2343:
2327:
2323:
2319:
2303:
2299:
2283:
2279:
2272:
2265:
2260:
2249:
2242:
2235:
2227:
2225:0-9707493-6-8
2221:
2217:
2210:
2204:, p. 173
2203:
2198:
2191:
2186:
2184:
2167:
2161:
2150:
2143:
2127:
2123:
2117:
2110:
2105:
2096:
2077:
2070:
2064:
2062:
2055:, p. 147
2054:
2049:
2042:
2037:
2021:
2017:
2011:
1996:
1991:
1984:
1968:
1964:
1958:
1942:
1938:
1932:
1916:
1912:
1906:
1887:
1880:
1874:
1858:
1854:
1848:
1841:
1836:
1829:
1824:
1808:
1804:
1798:
1791:
1786:
1779:
1774:
1767:
1762:
1760:
1752:
1751:Kraeuter 2001
1747:
1745:
1737:
1732:
1730:
1722:
1717:
1710:
1709:Kraeuter 2001
1705:
1699:, p. 172
1698:
1693:
1677:
1673:
1667:
1659:
1655:
1648:
1641:
1636:
1629:
1624:
1622:
1615:, p. 209
1614:
1609:
1603:, p. 110
1602:
1597:
1595:
1593:
1591:
1586:
1583:
1582:
1577:
1575:
1569:
1563:
1562:
1555:
1549:
1548:
1543:
1539:
1533:
1532:
1527:
1521:
1520:
1519:Twelfth Night
1515:
1508:
1507:
1503:
1498:
1493:
1489:
1488:
1484:
1480:
1477:
1476:
1472:
1467:
1466:
1454:
1451:
1448:
1445:
1444:
1435:
1431:
1427:
1424:
1420:
1416:
1413:
1412:
1407:
1403:
1402:
1396:
1395:
1390:
1386:
1382:
1379:
1375:
1372:
1371:
1366:
1365:
1360:
1356:
1355:
1350:
1346:
1342:
1339:
1338:
1333:
1324:
1322:
1321:HĂ´tel Montana
1318:
1314:
1310:
1306:
1302:
1297:
1290:
1281:
1279:
1275:
1271:
1266:
1261:
1256:
1254:
1250:
1246:
1241:
1238:
1234:
1223:
1221:
1216:
1210:
1208:
1198:
1196:
1192:
1188:
1184:
1175:
1166:
1164:
1160:
1156:
1152:
1148:
1144:
1140:
1136:
1132:
1128:
1127:snowshoe hare
1123:
1121:
1117:
1113:
1109:
1105:
1101:
1097:
1093:
1088:
1084:
1082:
1078:
1074:
1064:
1059:
1054:
1052:
1048:
1043:
1041:
1038:when hunting
1037:
1033:
1024:
1019:
1009:
1007:
1001:
999:
994:
992:
988:
987:harvest mites
984:
980:
974:
972:
968:
964:
960:
956:
952:
948:
944:
938:
936:
932:
928:
919:
915:
913:
909:
908:Hip dysplasia
905:
900:
896:
892:
887:
885:
875:
873:
869:
868:
863:
862:Stanley Coren
858:
856:
852:
848:
844:
843:Cairn Terrier
840:
836:
832:
822:
813:
811:
807:
803:
802:designer dogs
799:
795:
791:
781:
777:
774:
767:
762:
758:
756:
752:
748:
744:
729:
727:
723:
719:
715:
705:
701:
699:
695:
689:
687:
677:
673:
671:
666:
662:
658:
654:
650:
646:
637:
628:
626:
621:
617:
613:
608:
606:
602:
598:
597:
592:
588:
587:
576:
574:
570:
566:
562:
558:
554:
548:
546:
541:
536:
527:
518:
516:
512:
508:
504:
500:
489:
484:
479:
477:
473:
469:
463:
456:
452:
447:
443:
441:
437:
436:Prince Albert
433:
428:
424:
419:
409:
406:
403:
399:
395:
391:
387:
381:
377:
375:
371:
366:
361:
357:
353:
349:
345:
340:
338:
334:
330:
326:
322:
318:
310:
305:
296:
294:
289:
287:
283:
279:
275:
270:
268:
264:
260:
259:detection dog
256:
252:
249:
245:
241:
237:
233:
222:
213:
206:
201:
197:
194:
189:
185:
180:
176:
164:
156:
148:
144:
141:
138:
134:
131:
127:
119:
114:
110:
106:
102:
98:
92:
85:
80:
75:
69:
65:
48:
44:
40:
33:
28:
19:
5898:Scent hounds
5844:Fox Terriers
5832:
5810:Turnspit dog
5795:Talbot Hound
5785:Sleuth hound
5638:Rough Collie
5600:Herding dogs
5576:Bull Terrier
5509:Skye Terrier
5281:
5206:
5194:Talbot Hound
5179:Sleuth hound
5154:Norman Hound
5119:Celtic Hound
5043:Ibizan Hound
5033:Cretan Hound
4982:Mountain Cur
4888:Mudhol Hound
4853:Kangaroo dog
4783:Afghan Hound
4635:Polish Hound
4590:Kerry Beagle
4555:Halden Hound
4500:German Hound
4378:
4369:Basset Hound
4344:Artois Hound
4263:
4251:the original
4245:
4231:
4206:
4194:
4177:. Barron's.
4173:
4151:
4136:
4129:. E. Wilson.
4125:
4108:. Barron's.
4103:
4091:
4081:
4061:
4041:
4032:
4010:
4003:Bibliography
4002:
4001:
3988:. Retrieved
3963:. Retrieved
3959:the original
3952:
3943:
3931:. Retrieved
3925:
3915:
3903:. Retrieved
3899:the original
3889:
3878:, retrieved
3873:
3864:
3853:, retrieved
3842:
3833:
3823:19 September
3821:. Retrieved
3817:
3807:
3797:19 September
3795:. Retrieved
3785:
3773:. Retrieved
3768:
3759:
3747:. Retrieved
3742:
3733:
3721:
3709:. Retrieved
3705:
3695:
3683:. Retrieved
3679:the original
3669:
3657:. Retrieved
3647:
3633:
3622:
3613:
3599:
3588:
3579:
3567:. Retrieved
3558:
3546:. Retrieved
3542:the original
3537:
3527:
3515:. Retrieved
3511:the original
3506:
3496:
3484:. Retrieved
3479:
3470:
3458:. Retrieved
3454:the original
3444:
3432:. Retrieved
3428:the original
3418:
3406:. Retrieved
3402:the original
3392:
3380:. Retrieved
3376:the original
3366:
3354:. Retrieved
3319:. Retrieved
3299:
3287:. Retrieved
3277:
3265:. Retrieved
3230:. Retrieved
3226:the original
3221:
3212:
3200:. Retrieved
3196:the original
3186:
3174:. Retrieved
3170:the original
3160:
3148:. Retrieved
3141:the original
3121:. Retrieved
3117:the original
3101:. Retrieved
3096:
3087:
3075:. Retrieved
3071:the original
3061:
3049:. Retrieved
3045:the original
3035:
3008:. Retrieved
3004:the original
2999:
2990:
2978:. Retrieved
2973:
2964:
2940:
2933:
2928:, p. 77
2921:
2916:, p. 42
2914:Maxwell 1833
2909:
2899:
2866:
2862:
2833:
2827:
2815:. Retrieved
2811:the original
2801:
2789:
2770:
2764:
2745:
2739:
2727:. Retrieved
2720:the original
2707:
2695:. Retrieved
2691:the original
2681:
2669:. Retrieved
2641:
2637:
2627:
2615:
2591:
2579:. Retrieved
2575:the original
2547:11585/961937
2527:
2523:
2512:
2461:
2457:
2447:
2442:, p. 55
2440:Daglish 1961
2435:
2430:, p. 96
2423:
2411:
2399:. Retrieved
2394:
2384:
2372:. Retrieved
2368:the original
2363:
2354:
2342:
2330:. Retrieved
2321:
2306:. Retrieved
2286:. Retrieved
2282:the original
2271:
2266:, p. 46
2259:
2248:the original
2239:Libby, Ray.
2234:
2215:
2209:
2197:
2192:, p. 40
2172:25 September
2170:. Retrieved
2160:
2142:
2130:. Retrieved
2126:the original
2116:
2111:, p. 44
2109:Daglish 1961
2104:
2095:
2083:. Retrieved
2076:the original
2048:
2043:, p. 37
2041:Daglish 1961
2036:
2024:. Retrieved
2020:the original
2010:
1998:. Retrieved
1994:
1983:
1971:. Retrieved
1967:the original
1957:
1945:. Retrieved
1941:the original
1931:
1919:. Retrieved
1915:the original
1905:
1893:. Retrieved
1873:
1861:. Retrieved
1857:the original
1847:
1835:
1828:Daglish 1961
1823:
1811:. Retrieved
1807:the original
1797:
1792:, p. 14
1785:
1778:Daglish 1961
1773:
1768:, p. 12
1716:
1704:
1692:
1680:. Retrieved
1676:the original
1666:
1657:
1653:
1647:
1635:
1608:
1580:
1579:
1573:
1567:
1559:
1553:
1545:
1541:
1537:
1529:
1525:
1517:
1513:
1506:
1501:
1500:
1495:
1487:
1482:
1481:
1475:
1470:
1469:
1464:
1463:
1409:
1400:
1392:
1368:
1362:
1352:
1345:comic strips
1298:
1295:
1257:
1242:
1229:
1211:
1204:
1180:
1124:
1089:
1085:
1081:Drag hunting
1069:
1058:wire-haired.
1056:
1051:Edward Jesse
1046:
1044:
1029:
1002:
995:
975:
959:distichiasis
955:third eyelid
939:
924:
888:
881:
865:
859:
827:
787:
778:
775:
771:
740:
722:Fox Terriers
718:Basset Hound
711:
702:
693:
690:
685:
682:
642:
625:drag hunting
619:
612:Kerry Beagle
609:
604:
600:
594:
590:
584:
582:
549:
545:World War II
535:Peterborough
532:
495:
486:
481:
471:
464:
460:
431:
415:
407:
398:Talbot Hound
383:
379:
369:
364:
347:
341:
325:Talbot hound
319:brought the
314:
290:
274:Talbot Hound
271:
231:
229:
221:domestic dog
68:
46:
37:This is the
31:
5800:Toy Bulldog
5679:Bullmastiff
5317:Welsh Hound
4997:Treeing Cur
4898:Rajapalayam
4878:Magyar agár
4813:Chippiparai
4775:Sighthounds
4740:Trigg Hound
4625:Plott Hound
4575:Hygen Hound
4474:Erbi Txakur
4316:Scenthounds
3965:18 February
3933:18 February
3051:11 December
2926:Blakey 1854
1863:13 February
1780:, p. 9
1753:, p. 9
1711:, p. 7
1601:Youatt 1852
1531:Westward Ho
1315:, a Beagle
1305:pet therapy
1284:Other roles
1245:vivisection
1108:Marlborough
1006:cervicalgia
884:crossbreeds
855:Fox Terrier
816:Temperament
784:Crossbreeds
540:World War I
394:River Trent
344:Middle Ages
293:Elizabethan
253:, known as
240:scent hound
95:Other names
5888:FCI breeds
5867:Categories
5648:Smithfield
5564:Bull types
5543:Toy breeds
5307:Otterhound
5287:Bloodhound
5218:Coonhounds
5124:Chien-gris
4645:Porcelaine
4610:Otterhound
4399:Bloodhound
3775:26 January
3711:8 November
3659:8 November
3123:9 November
3077:2 February
2980:4 November
2671:3 November
2530:: 106155.
2401:26 October
2374:26 October
2347:Fogle 1990
2202:Fogle 1990
2190:Fogle 1990
2085:14 October
1721:Scott 1845
1697:Mills 1845
1640:Jesse 1866
1628:Jesse 1858
1613:Smith 2002
1544:both from
1524:Webster: "
1497:excitable.
1460:References
1397:was named
1389:the voyage
1357:character
1135:game birds
951:Cherry eye
831:guard dogs
798:crossbreed
732:Variations
714:Bloodhound
631:Appearance
521:Popularity
499:Dachshunds
451:Stonehenge
434:in 1845).
329:Greyhounds
263:quarantine
169:12.5 years
5790:Staghound
5725:Buckhound
5297:Greyhound
5189:Staghound
5114:Buckhound
5098:Xiasi Dog
4962:Blue Lacy
4828:Greyhound
4449:Dachshund
4339:Ariégeois
4243:(1897) .
3954:Dog World
3538:The Times
2883:0300-9858
2794:Rice 2000
2620:Rice 2000
2486:2045-2322
2053:Rice 2000
2000:3 October
1581:Citations
1169:Detection
1155:wild boar
991:tapeworms
402:Yorkshire
356:Henry VII
352:Edward II
342:From the
238:of small
71:Dog breed
5852:Lurchers
5848:Longdogs
5415:Terriers
5337:Gun dogs
5226:Lurchers
5144:Laconian
4948:Cur-type
4798:Bakhmull
4750:Valdueza
4630:Poitevin
4261:(1852).
4241:Xenophon
4229:(1856).
4148:(2000).
4033:Shooting
4031:(1854).
3880:12 March
3743:BBC News
3624:Fox News
3347:Archived
3312:Archived
3258:Archived
3232:4 August
3202:4 August
2974:BBC News
2666:23470450
2658:16648742
2604:Archived
2504:38302530
2495:10834484
2326:Archived
2322:BBC News
2302:Archived
1990:"Beagle"
1973:29 April
1886:Archived
1813:29 April
1411:Beagle 2
1143:red deer
1139:roe deer
1049:(1846),
1040:pheasant
1036:spaniels
1032:beagling
1018:Beagling
971:ADAMTS10
943:glaucoma
899:dwarfism
891:epilepsy
645:Foxhound
605:begueule
507:Illinois
483:hunting.
427:Harriers
388:and the
323:and the
255:beagling
244:foxhound
205:standard
193:standard
43:reviewed
5836:Collies
5708:Extinct
5674:Bulldog
5424:Working
5395:Pointer
5327:Whippet
5302:Harrier
5214:Bassets
5107:Extinct
5023:Basenji
4977:Lurcher
4933:Whippet
4923:Sloughi
4873:Longdog
4848:Kaikadi
4818:Chortai
4793:Azawakh
4570:Harrier
4264:The Dog
4042:Beagles
3569:8 March
3460:11 July
2891:7483207
2466:Bibcode
1921:11 July
1895:11 July
1408:lander
1406:Martian
1354:Peanuts
1301:termite
1185:of the
1163:gun dog
1012:Hunting
931:steroid
694:ticking
670:stifles
665:abdomen
649:withers
476:terrier
333:Harrier
299:History
286:Harrier
248:hunting
142:Females
107:England
18:Beagles
5873:Hounds
5850:&
5833:Types:
5427:breeds
5282:Beagle
5274:Hounds
5222:Feists
5207:Types:
5048:Maneto
4928:Taigan
4908:Saluki
4863:Kombai
4808:Borzoi
4459:Dunker
4454:Drever
4379:Beagle
4308:Hounds
4215:
4181:
4160:
4112:
4069:
4048:
4017:
3990:22 May
3905:9 July
3855:31 May
3749:9 July
3685:9 July
3548:9 July
3517:1 July
3486:22 May
3434:9 July
3408:9 July
3382:9 July
3356:9 July
3321:9 July
3289:9 July
3267:9 July
3176:9 July
3150:9 July
3103:9 July
3010:9 July
2952:
2904:23-26.
2889:
2881:
2840:
2817:9 July
2777:
2752:
2729:13 May
2697:13 May
2664:
2656:
2581:9 July
2502:
2492:
2484:
2332:9 July
2308:9 July
2288:9 July
2222:
2132:9 July
2026:9 July
1947:9 July
1682:9 July
1570:(1782)
1550:(1700)
1453:Miss P
1401:Beagle
1370:Gromit
1359:Snoopy
1220:Harlan
1151:coyote
1147:bobcat
1118:, and
1114:, the
1112:Radley
1096:Eton's
1053:says:
989:, and
973:gene.
957:, and
878:Health
853:, and
810:puggle
766:Puggle
661:dewlap
657:muzzle
620:beagle
616:Celtic
601:beagle
591:beagle
571:, and
492:Export
455:Youatt
348:beagle
280:, the
276:, the
232:beagle
158:Colour
129:Weight
121:Height
116:Traits
104:Origin
77:Beagle
5667:Other
5174:Rache
5149:Limer
4972:Feist
4938:Xigou
4858:Kanni
4389:Billy
3350:(PDF)
3343:(PDF)
3315:(PDF)
3308:(PDF)
3261:(PDF)
3254:(PDF)
3144:(PDF)
3137:(PDF)
2946:68–71
2807:"FAQ"
2723:(PDF)
2716:(PDF)
2662:S2CID
2638:Spine
2607:(PDF)
2600:(PDF)
2251:(PDF)
2244:(PDF)
2152:(PDF)
2079:(PDF)
2072:(PDF)
1889:(PDF)
1882:(PDF)
1387:made
1243:Anti-
1159:stoat
983:ticks
979:fleas
653:skull
505:from
418:Essex
236:breed
234:is a
132:Males
4213:ISBN
4179:ISBN
4158:ISBN
4110:ISBN
4067:ISBN
4046:ISBN
4015:ISBN
3992:2020
3967:2015
3935:2015
3907:2007
3882:2024
3857:2013
3825:2015
3799:2015
3777:2010
3751:2007
3713:2011
3687:2007
3661:2011
3571:2022
3550:2007
3519:2015
3488:2014
3462:2007
3436:2007
3410:2007
3384:2007
3358:2007
3323:2007
3291:2007
3269:2007
3234:2018
3204:2018
3178:2007
3152:2007
3125:2011
3105:2007
3079:2007
3053:2011
3012:2007
2982:2011
2950:ISBN
2887:PMID
2879:ISSN
2838:ISBN
2819:2007
2775:ISBN
2750:ISBN
2731:2013
2699:2013
2673:2008
2654:PMID
2583:2007
2500:PMID
2482:ISSN
2403:2011
2376:2011
2334:2007
2310:2007
2290:2007
2220:ISBN
2174:2023
2134:2007
2087:2011
2028:2007
2002:2016
1975:2014
1949:2007
1923:2007
1897:2007
1865:2008
1815:2014
1684:2007
1399:HMS
1347:and
945:and
835:howl
753:and
741:The
716:and
686:pied
579:Name
438:and
354:and
307:The
251:hare
230:The
150:Coat
5735:Cur
2871:doi
2646:doi
2542:hdl
2532:doi
2528:225
2490:PMC
2474:doi
1447:Uno
949:. "
864:'s
806:Pug
755:FCI
267:pet
216:Dog
45:on
5869::
5846:,
5842:,
5838:,
5224:,
5220:,
5216:,
5212:,
3975:^
3951:.
3924:.
3872:,
3847:,
3841:,
3816:.
3767:.
3741:.
3704:.
3621:.
3587:.
3536:.
3505:.
3478:.
3331:^
3242:^
3220:.
3130:*
3110:*
3095:.
3020:^
2998:.
2972:.
2948:.
2885:.
2877:.
2867:32
2865:.
2852:^
2660:.
2652:.
2642:31
2640:.
2636:.
2556:^
2540:.
2526:.
2522:.
2498:.
2488:.
2480:.
2472:.
2462:14
2460:.
2456:.
2393:.
2362:.
2320:.
2315:*
2295:*
2182:^
2060:^
1993:.
1758:^
1743:^
1728:^
1656:.
1620:^
1589:^
1502:d.
1483:c.
1471:a.
1153:,
1149:,
1145:,
1141:,
1137:,
1133:,
1129:,
1122:.
1110:,
1042:.
985:,
981:,
857:.
849:,
845:,
841:,
764:A
672:.
627:.
575:.
567:,
346:,
288:.
41:,
5260:e
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