25:
128:
103:
478:
would suggest a one- to two-month lactation time, after which the young is weaned and probably leaves the nest. Females have been found carrying either one or two embryos. In a study on the abundance of fruit bats in
Malaysia, two fruit bats, one male, one pregnant female, were found roosting in a hollow limb of a fig tree.
250:
477:
species, with both sexes coming into sexual maturity when they reach 70 grams (approximately one-half adult body weight). While it is unknown how long the females gestate, females have been found carrying embryos from June to July, and have been found lactating from June until
September. This
389:
The dayak bat averages less than 150 grams (5.3 oz). It is gray-brown dorsally with a silver-gray ventral side. Its fur is short. It has a short tail, ranging from ten to twenty percent of its body length. Its wing membranes attach to the second toe on each foot.
411:
seems to prefer the fruits of old-growth
Paleotropical rainforest, or montane rainforest. This makes them particularly susceptible to deforestation. Frugivorous bats are one of the main agents of seed dispersal in the rainforest, making them essential to the ecosystem.
494:. While the reasons the male would lactate are currently unknown, it could be a mechanism to take some of the pressure of lactation off the female. This may confer an evolutionary advantage in lactating males over other non-lactating males.
586:
Hodgkisin R., S. T. Balding, A. Zubaid, T.H. Kunz, Temporal
Variation in the Relative Abundance of Fruit Bat (Megachiroptera:Pteropidae) in Relation to the Availability of Food in a Lowland Malaysian Rain Forest, Biotropica Vol. 36,
573:
Mohd. Azlan J., Ibnu
Maryanto, Agus P. Kartono and M.T. Abdullah. 2003 Diversity, Relative Abundance and Conservation of Chiropterans in Kayan Mentarang National Park, East Kalimantan, Indonesia. Sarawak Museum Journal 79:
406:
The dayak bat is frugivorous, feeding on fruits from trees extending into or past the canopy of the forests of
Malaysia. Figs are a staple of its diet, as are whatever fruits are seasonally available in the high canopy.
398:. It has the smallest skull size, averaging less than 36.1 mm (1.42 in) from the premaxilla to the base of the skull. It has a proportionally wide skull, however, when compared to the other bats in the genus.
430:. Females from Kota Samarahan and Kubah were lactating in September 1994 and August 1996 respectively. Two individuals were caught in the subcanopy (between 10–15 m) by using a vertically-hoisted mist net placed in the
380:
Previous distribution records includes
Selangor and Negeri Sembilan in Peninsular Malaysia, Sumatra; Sepilok and Baturong Caves in Sabah; Baram and Niah areas in Sarawak (Medway 1978, Payne et al. 1985).
420:
A male and five female adults were mist-netted. Two females collected from
Kalimantan Barat in September 1996 were in early and late pregnancy. Both bats were caught in a mist net placed near fruiting
373:
Borneo. Specimens UMS0010, MTA96237, and MTA96238 from
Sarawak and MTA96208 from Sabah are new additional records for the species in the sampling areas. These new sites have extended the range of
567:
Hall LS, Gordon G. Grigg, Craig Moritz, Besar Ketol, Isa Sait, Wahab Marni and M.T. Abdullah. 2004. Biogeography of fruit bats in
Southeast Asia. Sarawak Museum Journal LX(81):191-284.
590:
Helgen K.M., D. Kock, R.K. Comez, N.R. Ingle, M.H. Sinoga, Taxonomy and Natural History of the Southeast Asian Fruit-Bat Genus Dyacopterus, Journal of Mammalogy Vol. 88, p 302-318
609:
2835:
442:
tree above the main canopy (> 30 m from the ground level). However, the two specimens from Kalimantan Barat were taken from the ground level nets in a
593:
Ingle N.F., "Seed Dispersal by wind, birds, and bats between Philippine montane rainforest and successional vegetation" Oecologia. Vol. 134 p 95-104
3567:
3753:
3632:
3606:
431:
345:
is considered a very rare fruit bat species in Peninsular Malaysia and Borneo and none in Thailand. The species were netted in four sites at
606:
3712:
3541:
3199:
3725:
3580:
465:, and no socioeconomic studies have been performed, there is little data available on the reproduction and habits of any of the
3803:
3310:
3175:
3166:
640:
3624:
3092:
3730:
3585:
1504:
1495:
3024:
1549:
3798:
3110:
3060:
1594:
1102:
738:
577:
Hall LS, Richards GC, Abdullah MT. 2002. The bats of Niah National Park, Sarawak. Sarawak Museum Journal. 78: 255-282.
330:
Few specimens of any of the three species exist, due not only to their rarity, but also because they rarely enter the
3042:
1513:
68:
46:
39:
3637:
3301:
3101:
3006:
2997:
2530:
1522:
1120:
774:
3763:
3506:
2707:
1904:
1603:
1567:
1540:
837:
792:
3033:
2698:
1585:
1576:
888:
570:
Karim, C., A.A. Tuen and M.T. Abdullah. 2004. Mammals. Sarawak Museum Journal Special Issue No. 6. 80: 221—234.
2253:
2104:
2068:
1955:
1558:
1260:
1111:
765:
3783:
3356:
3015:
2988:
2077:
1194:
1066:
819:
810:
3619:
3402:
1664:
1531:
1161:
879:
801:
783:
747:
127:
3788:
3768:
3051:
2602:
2334:
2199:
1727:
1152:
855:
756:
633:
3793:
3773:
3319:
2867:
1786:
2656:
2611:
2406:
2316:
2280:
2095:
2086:
1996:
1627:
1269:
1251:
1218:
1075:
1003:
979:
33:
547:
3778:
3393:
2503:
2235:
2163:
2044:
2020:
1946:
1836:
1745:
1736:
1694:
1382:
1349:
3572:
3468:
2840:
2512:
2415:
2298:
2190:
1703:
1358:
1034:
1013:
846:
362:
50:
469:
species. However, from information gained during studies of Malaysian fruit bats in general,
3435:
2584:
2566:
2226:
1827:
1777:
1463:
1416:
1084:
828:
626:
222:
3663:
3758:
3528:
3411:
2912:
2894:
2593:
2557:
2442:
2370:
2361:
2136:
1987:
358:
3598:
8:
2921:
2783:
2629:
2539:
2424:
2208:
1886:
474:
350:
92:
618:
2858:
2765:
2620:
2548:
2479:
2352:
2262:
2172:
2145:
1818:
1242:
580:
Francis, Charles M., et al.; "Lactation in Male Fruit Bats," Nature, 367:691-692, 1994.
122:
583:
Fackelmann K.A., "Real males that lactate: A batty story." Science News Vol. 145 p 148
439:
3699:
3658:
3515:
3269:
3260:
3134:
2849:
2638:
2494:
2469:
2451:
2397:
2388:
2379:
2343:
2244:
2217:
1913:
1895:
1441:
721:
596:
Konig B., "Cooperative Care of Young in Mammals" Naturwisenschaften. Vol. 84 p 95-104
503:
427:
524:
523:
Csorba, G.; Bumrungsri, S.; Francis, C.; Bates, P.; Gumal, M.; Kingston, T. (2020).
3704:
3690:
3520:
3236:
2876:
2822:
2674:
2665:
2647:
2575:
2521:
2485:
2433:
2307:
2271:
2154:
1922:
1093:
946:
542:
366:
331:
453:
as indicated by Medway (1978), Payne et al. (1985) and Mickleburg et al., (1992).
3645:
2885:
2774:
2460:
2289:
1970:
1636:
1618:
1406:
1317:
1293:
955:
613:
289:
239:
107:
3611:
3491:
3374:
2325:
2181:
686:
491:
354:
3747:
3593:
3426:
3251:
3227:
3149:
2949:
2907:
1185:
994:
870:
533:
112:
522:
3384:
3217:
3190:
3125:
2961:
2689:
1864:
1801:
1718:
1677:
970:
916:
708:
3650:
3554:
3500:
3284:
2971:
2735:
1332:
1284:
1135:
361:, all on Borneo. The specimens, MTA96268 and MTA96269, that were held at
327:
302:
285:
278:
199:
3717:
3546:
2748:
2059:
1937:
1760:
1397:
1049:
692:
249:
3559:
3337:
2798:
2035:
1454:
1373:
1308:
1233:
929:
370:
139:
3462:
3684:
3485:
3347:
2119:
2011:
1877:
1478:
1209:
1025:
680:
674:
668:
662:
323:
319:
159:
3075:
650:
297:
281:
189:
449:
There is no previous information on the biology and ecology for
3533:
2817:
435:
346:
293:
169:
149:
422:
334:
of the forest where they can be caught in scientists' nets.
516:
443:
648:
179:
490:
species is one of the few known natural occurrences of
365:, are the first record for this distinctive species in
3745:
292:south of the Isthmus of Kra, and the islands of
1439:
634:
548:10.2305/IUCN.UK.2020-3.RLTS.T6931A22029918.en
377:to the western and southern parts of Borneo.
1662:
2947:
1862:
914:
706:
641:
627:
248:
101:
2733:
546:
394:is the smallest of the bats in the genus
69:Learn how and when to remove this message
3311:East African little collared fruit bat (
32:This article includes a list of general
3345:
3200:Veldkamp's dwarf epauletted fruit bat (
300:. There are three species in the genus
3746:
607:The incredible milk-producing male bat
415:
3754:IUCN Red List near threatened species
3467:
3466:
3167:Hayman's dwarf epauletted fruit bat (
3025:Lesser Angolan epauletted fruit bat (
2946:
2732:
1861:
1661:
1438:
913:
705:
622:
481:
3176:Peters' dwarf epauletted fruit bat (
3093:Buettikofer's epauletted fruit bat (
1595:New Britain naked-backed fruit bat (
1103:Nusatenggara short-nosed fruit bat (
739:Broad-striped tube-nosed fruit bat (
288:of southeast Asia, specifically the
18:
3043:East African epauletted fruit bat (
1514:Philippine naked-backed fruit bat (
1505:Beaufort's naked-backed fruit bat (
1496:Andersen's naked-backed fruit bat (
534:IUCN Red List of Threatened Species
318:. All are found in the forests of
13:
2531:Little golden-mantled flying fox (
1550:Solomon's naked-backed fruit bat (
1523:Halmahera naked-backed fruit bat (
1121:Indonesian short-nosed fruit bat (
775:Round-eared tube-nosed fruit bat (
38:it lacks sufficient corresponding
14:
3815:
3111:Franquet's epauletted fruit bat (
3061:Wahlberg's epauletted fruit bat (
2708:Sulawesi stripe-faced fruit bat (
1905:Giant golden-crowned flying fox (
1604:Greenish naked-backed fruit bat (
1568:Moluccan naked-backed fruit bat (
1541:Sulawesi naked-backed fruit bat (
838:Philippine tube-nosed fruit bat (
600:
3034:Ethiopian epauletted fruit bat (
2699:Mindoro stripe-faced fruit bat (
1586:Western naked-backed fruit bat (
1577:Panniet naked-backed fruit bat (
889:Steadfast tube-nosed fruit bat (
126:
23:
3007:Peters's epauletted fruit bat (
2998:Ansell's epauletted fruit bat (
2254:New Ireland masked flying fox (
2105:New Georgian monkey-faced Bat (
2069:Bougainville monkey-faced Bat (
1956:Small white-winged flying fox (
1559:Lesser naked-backed fruit bat (
1112:Greater short-nosed fruit bat (
766:Mountain tube-nosed fruit bat (
456:
337:
3016:Gambian epauletted fruit bat (
2989:Angolan epauletted fruit bat (
2078:Guadalcanal monkey-faced Bat (
1067:Lesser short-nosed fruit bat (
820:Demonic tube-nosed fruit bat (
811:Malaita tube-nosed fruit bat (
793:Keast's tube-nosed fruit bat (
434:at Kubah. The individual from
384:
1:
3804:Taxa named by Oldfield Thomas
3302:São Tomé collared fruit bat (
1532:Biak naked-backed fruit bat (
880:Lesser tube-nosed fruit bat (
802:Island tube-nosed fruit bat (
784:Dragon tube-nosed fruit bat (
748:Common tube-nosed fruit bat (
509:
3292:(Little collared fruit bats)
3052:Minor epauletted fruit bat (
2603:Philippine gray flying fox (
2200:Bismarck masked flying fox (
1261:Ratanaworabhan's fruit bat (
856:Umboi tube-nosed fruit bat (
461:As there are few samples of
7:
3320:Little collared fruit bat (
1978:(Straw-coloured fruit bats)
1787:New Caledonia blossom bat (
1195:Fischer's pygmy fruit bat (
497:
10:
3820:
2657:New Caledonia flying fox (
2407:Black-bearded flying fox (
2096:Montane monkey-faced Bat (
2087:Greater monkey-faced Bat (
1997:Straw-coloured fruit bat (
1628:Sulawesi harpy fruit bat (
1270:White-collared fruit bat (
1252:Javan tailless fruit bat (
1162:Rickart's dyak fruit bat (
1004:Spotted-winged fruit bat (
980:Mindanao pygmy fruit bat (
284:species found only on the
3799:Mammals described in 1890
3674:
3475:
3424:
3394:Short-palated fruit bat (
3382:
3373:
3336:
3282:
3249:
3225:
3216:
3188:
3147:
3123:
3073:
2969:
2960:
2956:
2942:
2796:
2746:
2742:
2728:
2687:
2504:Geelvink Bay flying fox (
2335:Livingstone's fruit bat (
2236:Large Samoan flying fox (
2164:Small Samoan flying fox (
2117:
2057:
2045:Small-toothed fruit bat (
2033:
2021:Fijian monkey-faced Bat (
2009:
1968:
1947:White-winged flying fox (
1935:
1875:
1871:
1857:
1837:Moss-forest blossom bat (
1799:
1758:
1737:Black-bellied fruit bat (
1728:Fardoulis's blossom bat (
1716:
1695:Long-tongued nectar bat (
1685:(Long-tongued fruit bats)
1675:
1671:
1657:
1616:
1476:
1452:
1448:
1434:
1395:
1371:
1350:Greater musky fruit bat (
1330:
1306:
1282:
1231:
1207:
1183:
1153:Brooks's dyak fruit bat (
1133:
1047:
1023:
992:
968:
927:
923:
909:
868:
757:Pallas's tube-nosed bat (
719:
715:
701:
657:
649:Extant species of family
256:
247:
228:
221:
123:Scientific classification
121:
99:
90:
85:
3357:D'Anchieta's fruit bat (
2868:Leschenault's rousette (
2513:Grey-headed flying fox (
2416:Black-eared flying fox (
2299:Ontong Java flying fox (
2191:Ashy-headed flying fox (
1768:(Long-tailed fruit bats)
1704:Long-tongued fruit bat (
1486:(Bare-backed fruit bats)
1359:Lesser musky fruit bat (
1035:Black-capped fruit bat (
847:Eastern tube-nosed Bat (
3403:Campo-Ma’an fruit bat (
2612:Temminck's flying fox (
2585:Little red flying fox (
2567:Madagascan flying fox (
2317:Andersen's flying fox (
2281:Gilliard's flying fox (
2227:Spectacled flying fox (
1828:Halmahera blossom bat (
1778:Long-tailed fruit bat (
1219:Salim Ali's fruit bat (
1076:Horsfield's fruit bat (
829:Lesser tube-nosed bat (
729:(tube-nosed fruit bats)
401:
277:) is a relatively rare
53:more precise citations.
3764:Bats of Southeast Asia
3157:(Dwarf epauleted bats)
2979:(Epauleted fruit bats)
2913:Long-haired rousette (
2895:Bare-backed rousette (
2594:Seychelles fruit bat (
2558:Rodrigues flying fox (
2443:Mauritian flying fox (
2371:Big-eared flying fox (
2137:Admiralty flying fox (
1988:Madagascan fruit bat (
1746:Woodford's fruit bat (
1383:Blanford's fruit bat (
1057:(Dog-faced fruit bats)
1014:Balionycteris seimundi
363:Tanjungpura University
257:Dayak fruit bat range
3507:Dyacopterus spadiceus
3477:Dyacopterus spadiceus
2922:Madagascan rousette (
2841:Geoffroy's rousette (
2806:(rousette fruit bats)
2784:Philippine dawn bat (
2630:Vanikoro flying fox (
2540:Solomons flying fox (
2425:Caroline flying fox (
2209:Moluccan flying fox (
1887:Sulawesi flying fox (
527:Dyacopterus spadiceus
463:Dyacopterus spadiceus
274:Dyacopterus spadiceus
232:Dyacopterus spadiceus
3676:Cynopterus spadiceus
3436:Zenker's fruit bat (
3102:Dobson's fruit bat (
2766:Greater nectar bat (
2621:Insular flying fox (
2549:Rennell flying fox (
2353:Okinawa flying fox (
2263:Nicobar flying fox (
2173:Vanuatu flying fox (
2146:Aldabra flying fox (
1819:Common blossom bat (
1464:Bulmer's fruit bat (
1243:Tailless fruit bat (
1085:Peters's fruit bat (
541:: e.T6931A22029918.
3784:Mammals of Thailand
3412:Pohle's fruit bat (
3261:Azagnyi fruit bat (
3135:Hammer-headed Bat (
2924:R. madagascariensis
2859:Egyptian rousette (
2850:Sulawesi rousette (
2639:Kosrae flying fox (
2495:Masked flying fox (
2470:Ornate flying fox (
2452:Temotu flying fox (
2398:Indian flying fox (
2389:Mariana fruit bat (
2380:Lesser flying fox (
2362:Lyle's flying fox (
2344:Lombok flying fox (
2245:Ryukyu flying fox (
2218:Makira flying fox (
1958:D. microleucopterus
1914:Palawan fruit bat (
1896:Talaud flying fox (
416:Biology and ecology
93:Conservation status
3237:Angolan rousette (
2877:Linduan Rousette (
2843:R. amplexicaudatus
2823:Manado fruit bat (
2675:Dwarf flying fox (
2666:Pemba flying fox (
2648:Large flying fox (
2576:Samoa flying fox (
2522:Bonin flying fox (
2486:Pelew flying fox (
2434:Great flying fox (
2308:Small flying fox (
2272:Banks flying fox (
2155:Black flying fox (
1923:Sunda flying fox (
1340:(Musky fruit bats)
1143:(Dayak fruit bats)
1094:Minute fruit bat (
947:Borneo fruit bat (
937:(Pygmy fruit bats)
612:2010-09-23 at the
492:paternal lactation
482:Paternal lactation
438:was caught in the
426:tree in disturbed
3789:Mammals of Brunei
3769:Bats of Indonesia
3741:
3740:
3659:Open Tree of Life
3469:Taxon identifiers
3460:
3459:
3456:
3455:
3452:
3451:
3448:
3447:
3405:C. campomaanensis
3369:
3368:
3332:
3331:
3295:
3212:
3211:
3160:
3086:
2982:
2938:
2937:
2934:
2933:
2886:Comoro rousette (
2809:
2775:Cave nectar bat (
2759:
2756:(Dawn fruit bats)
2724:
2723:
2720:
2719:
2461:Ceram fruit bat (
2290:Gray flying fox (
2229:P. conspicillatus
2130:
1981:
1853:
1852:
1849:
1848:
1812:
1771:
1688:
1653:
1652:
1649:
1648:
1637:Harpy fruit bat (
1489:
1442:Harpyionycterinae
1430:
1429:
1426:
1425:
1407:Swift fruit bat (
1343:
1318:Dusky fruit bat (
1296:O. cartilagonodus
1294:Luzon fruit bat (
1171:Dayak fruit bat (
1146:
1123:C. titthaecheilus
1060:
1037:C. melanocephalus
956:Pygmy fruit bat (
940:
905:
904:
901:
900:
732:
504:Mammals of Borneo
488:D. spadiceus
471:D. spadiceus
451:D. spadiceus
428:peat swamp forest
409:D. spadiceus
392:D. spadiceus
375:D. spadiceus
308:D. spadiceus
261:
260:
214:D. spadiceus
116:
79:
78:
71:
3811:
3794:Fauna of Sumatra
3774:Bats of Malaysia
3734:
3733:
3721:
3720:
3708:
3707:
3695:
3694:
3693:
3667:
3666:
3654:
3653:
3641:
3640:
3628:
3627:
3615:
3614:
3602:
3601:
3589:
3588:
3576:
3575:
3563:
3562:
3550:
3549:
3537:
3536:
3524:
3523:
3511:
3510:
3509:
3496:
3495:
3494:
3464:
3463:
3380:
3379:
3343:
3342:
3304:M. brachycephala
3294:
3293:
3289:
3270:Woermann's bat (
3223:
3222:
3159:
3158:
3154:
3085:
3084:
3083:(Epauleted bats)
3080:
2981:
2980:
2976:
2967:
2966:
2958:
2957:
2944:
2943:
2808:
2807:
2803:
2758:
2757:
2753:
2744:
2743:
2730:
2729:
2596:P. seychellensis
2515:P. poliocephalus
2436:P. neohibernicus
2326:Kei flying fox (
2211:P. chrysoproctus
2182:Aru flying fox (
2139:P. admiralitatum
2129:
2128:
2124:
1980:
1979:
1975:
1873:
1872:
1859:
1858:
1811:
1810:
1806:
1789:N. neocaledonica
1770:
1769:
1765:
1687:
1686:
1682:
1673:
1672:
1659:
1658:
1488:
1487:
1483:
1450:
1449:
1436:
1435:
1342:
1341:
1337:
1145:
1144:
1140:
1059:
1058:
1054:
939:
938:
934:
925:
924:
911:
910:
731:
730:
726:
717:
716:
703:
702:
643:
636:
629:
620:
619:
560:
559:
557:
555:
550:
520:
486:The male of the
367:Kalimantan Barat
328:the Philippines.
316:D. rickarti
252:
234:
131:
130:
110:
105:
104:
86:Dayak fruit bat
83:
82:
74:
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3290:
3288:
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3143:
3119:
3095:E. buettikoferi
3082:
3081:
3079:
3069:
2978:
2977:
2975:
2952:
2930:
2870:R. leschenaulti
2805:
2804:
2802:
2792:
2755:
2754:
2752:
2738:
2716:
2683:
2632:P. tuberculatus
2454:P. nitendiensis
2337:P. livingstonii
2283:P. gilliardorum
2126:
2125:
2123:
2113:
2053:
2029:
2005:
1977:
1976:
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1767:
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1684:
1683:
1681:
1667:
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1619:Harpyionycteris
1612:
1579:D. pannietensis
1485:
1484:
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1472:
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1422:
1391:
1367:
1339:
1338:
1336:
1326:
1302:
1278:
1227:
1203:
1179:
1142:
1141:
1139:
1129:
1105:C. nusatenggara
1056:
1055:
1053:
1043:
1019:
988:
982:A. paucidentata
964:
936:
935:
933:
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614:Wayback Machine
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587:p. 522-533
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446:swamp forest.
418:
404:
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340:
312:D. brooksi
290:Malay Peninsula
265:dayak fruit bat
243:
236:
230:
217:
125:
117:
108:Near Threatened
106:
102:
95:
75:
64:
58:
55:
45:Please help to
44:
28:
24:
17:
12:
11:
5:
3817:
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3796:
3791:
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3779:Bats of Borneo
3776:
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3390:
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3375:Scotonycterini
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3210:
3209:
3207:
3206:
3196:
3194:
3186:
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3173:
3169:M. intermedius
3163:
3161:
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3141:
3131:
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2932:
2931:
2929:
2928:
2919:
2902:
2901:
2892:
2883:
2874:
2865:
2861:R. aegyptiacus
2856:
2847:
2830:
2829:
2812:
2810:
2794:
2793:
2791:
2790:
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2772:
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2760:
2740:
2739:
2726:
2725:
2722:
2721:
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2705:
2701:S. mindorensis
2695:
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2663:
2654:
2645:
2636:
2627:
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2609:
2600:
2591:
2582:
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2564:
2560:P. rodricensis
2555:
2546:
2537:
2528:
2519:
2510:
2501:
2492:
2483:
2476:
2467:
2458:
2449:
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2431:
2422:
2413:
2409:P. melanopogon
2404:
2395:
2386:
2377:
2368:
2359:
2355:P. loochoensis
2350:
2346:P. lombocensis
2341:
2332:
2323:
2319:P. intermedius
2314:
2310:P. hypomelanus
2305:
2296:
2287:
2278:
2269:
2260:
2251:
2242:
2233:
2224:
2215:
2206:
2202:P. capistratus
2197:
2188:
2179:
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2152:
2148:P. aldabrensis
2143:
2133:
2131:
2127:(flying foxes)
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2017:
2015:
2007:
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1982:
1966:
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1949:D. leucopterus
1943:
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1809:(blossom bats)
1797:
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1665:Macroglossinae
1655:
1654:
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1634:
1624:
1622:
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1601:
1592:
1583:
1574:
1570:D. moluccensis
1565:
1556:
1547:
1538:
1529:
1520:
1511:
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1492:
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1168:
1159:
1149:
1147:
1131:
1130:
1128:
1127:
1118:
1109:
1100:
1091:
1087:C. luzoniensis
1082:
1078:C. horsfieldii
1073:
1063:
1061:
1045:
1044:
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1041:
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1029:
1021:
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895:
886:
876:
874:
866:
865:
863:
862:
853:
844:
835:
826:
817:
813:N. malaitensis
808:
799:
790:
781:
772:
763:
754:
745:
735:
733:
713:
712:
699:
698:
696:
695:
689:
687:Laurasiatheria
683:
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601:External links
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473:seems to be a
458:
455:
417:
414:
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386:
383:
355:Kota Samarahan
339:
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269:dyak fruit bat
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119:
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77:
76:
31:
29:
22:
16:Species of bat
15:
9:
6:
4:
3:
2:
3816:
3805:
3802:
3800:
3797:
3795:
3792:
3790:
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3604:
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3530:
3526:
3522:
3517:
3513:
3508:
3502:
3498:
3493:
3487:
3483:
3482:
3480:
3478:
3474:
3470:
3465:
3441:
3439:
3434:
3433:
3431:
3429:
3428:
3427:Scotonycteris
3423:
3417:
3415:
3410:
3408:
3406:
3401:
3399:
3397:
3392:
3391:
3389:
3387:
3386:
3381:
3378:
3376:
3372:
3362:
3360:
3355:
3354:
3352:
3350:
3349:
3344:
3341:
3339:
3335:
3325:
3323:
3318:
3316:
3314:
3309:
3307:
3305:
3300:
3299:
3297:
3287:
3286:
3281:
3275:
3273:
3268:
3266:
3264:
3259:
3258:
3256:
3254:
3253:
3252:Megaloglossus
3248:
3242:
3240:
3239:L. angolensis
3235:
3234:
3232:
3230:
3229:
3228:Lissonycteris
3224:
3221:
3219:
3215:
3205:
3203:
3202:N. veldkampii
3198:
3197:
3195:
3193:
3192:
3187:
3181:
3179:
3174:
3172:
3170:
3165:
3164:
3162:
3152:
3151:
3150:Micropteropus
3146:
3140:
3138:
3137:H. monstrosus
3133:
3132:
3130:
3128:
3127:
3122:
3116:
3114:
3109:
3107:
3105:
3100:
3098:
3096:
3091:
3090:
3088:
3078:
3077:
3072:
3066:
3064:
3059:
3057:
3055:
3050:
3048:
3046:
3041:
3039:
3037:
3032:
3030:
3028:
3023:
3021:
3019:
3014:
3012:
3010:
3005:
3003:
3001:
2996:
2994:
2992:
2991:E. angolensis
2987:
2986:
2984:
2974:
2973:
2968:
2965:
2963:
2959:
2955:
2951:
2950:Epomophorinae
2945:
2941:
2927:
2925:
2920:
2918:
2916:
2910:
2909:
2908:Stenonycteris
2904:
2903:
2900:
2898:
2897:R. spinalatus
2893:
2891:
2889:
2888:R. obliviosus
2884:
2882:
2880:
2879:R. linduensis
2875:
2873:
2871:
2866:
2864:
2862:
2857:
2855:
2853:
2852:R. celebensis
2848:
2846:
2844:
2838:
2837:
2832:
2831:
2828:
2826:
2820:
2819:
2814:
2813:
2811:
2801:
2800:
2795:
2789:
2787:
2782:
2780:
2778:
2773:
2771:
2769:
2764:
2763:
2761:
2751:
2750:
2745:
2741:
2737:
2731:
2727:
2713:
2711:
2706:
2704:
2702:
2697:
2696:
2694:
2692:
2691:
2686:
2680:
2678:
2673:
2671:
2669:
2668:P. voeltzkowi
2664:
2662:
2660:
2655:
2653:
2651:
2646:
2644:
2642:
2637:
2635:
2633:
2628:
2626:
2624:
2619:
2617:
2615:
2614:P. temminckii
2610:
2608:
2606:
2601:
2599:
2597:
2592:
2590:
2588:
2587:P. scapulatus
2583:
2581:
2579:
2574:
2572:
2570:
2565:
2563:
2561:
2556:
2554:
2552:
2547:
2545:
2543:
2538:
2536:
2534:
2529:
2527:
2525:
2520:
2518:
2516:
2511:
2509:
2507:
2502:
2500:
2498:
2497:P. personatus
2493:
2491:
2489:
2488:P. pelewensis
2484:
2482:
2481:
2477:
2475:
2473:
2468:
2466:
2464:
2459:
2457:
2455:
2450:
2448:
2446:
2441:
2439:
2437:
2432:
2430:
2428:
2427:P. molossinus
2423:
2421:
2419:
2414:
2412:
2410:
2405:
2403:
2401:
2396:
2394:
2392:
2387:
2385:
2383:
2378:
2376:
2374:
2369:
2367:
2365:
2360:
2358:
2356:
2351:
2349:
2347:
2342:
2340:
2338:
2333:
2331:
2329:
2324:
2322:
2320:
2315:
2313:
2311:
2306:
2304:
2302:
2297:
2295:
2293:
2288:
2286:
2284:
2279:
2277:
2275:
2270:
2268:
2266:
2261:
2259:
2257:
2252:
2250:
2248:
2247:P. dasymallus
2243:
2241:
2239:
2234:
2232:
2230:
2225:
2223:
2221:
2216:
2214:
2212:
2207:
2205:
2203:
2198:
2196:
2194:
2189:
2187:
2185:
2180:
2178:
2176:
2171:
2169:
2167:
2162:
2160:
2158:
2153:
2151:
2149:
2144:
2142:
2140:
2135:
2134:
2132:
2122:
2121:
2116:
2110:
2108:
2103:
2101:
2099:
2094:
2092:
2090:
2085:
2083:
2081:
2076:
2074:
2072:
2067:
2066:
2064:
2062:
2061:
2056:
2050:
2048:
2043:
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2032:
2026:
2024:
2019:
2018:
2016:
2014:
2013:
2008:
2002:
2000:
1995:
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1991:
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1983:
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1972:
1967:
1961:
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1954:
1952:
1950:
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1944:
1942:
1940:
1939:
1934:
1928:
1926:
1921:
1919:
1917:
1912:
1910:
1908:
1903:
1901:
1899:
1894:
1892:
1890:
1889:A. celebensis
1885:
1884:
1882:
1880:
1879:
1874:
1870:
1866:
1860:
1856:
1842:
1840:
1835:
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1824:
1822:
1817:
1816:
1814:
1804:
1803:
1798:
1792:
1790:
1785:
1783:
1781:
1780:N. macdonaldi
1776:
1775:
1773:
1763:
1762:
1757:
1751:
1749:
1744:
1742:
1740:
1735:
1733:
1731:
1730:M. fardoulisi
1726:
1725:
1723:
1721:
1720:
1715:
1709:
1707:
1702:
1700:
1698:
1693:
1692:
1690:
1680:
1679:
1674:
1670:
1666:
1660:
1656:
1642:
1640:
1639:H. whiteheadi
1635:
1633:
1631:
1630:H. celebensis
1626:
1625:
1623:
1621:
1620:
1615:
1609:
1607:
1602:
1600:
1598:
1597:D. praedatrix
1593:
1591:
1589:
1584:
1582:
1580:
1575:
1573:
1571:
1566:
1564:
1562:
1557:
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1548:
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1519:
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1501:
1499:
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1469:
1467:
1462:
1461:
1459:
1457:
1456:
1451:
1447:
1443:
1437:
1433:
1419:
1418:
1417:T. suhaniahae
1414:
1412:
1410:
1409:T. nigrescens
1405:
1404:
1402:
1400:
1399:
1394:
1388:
1386:
1381:
1380:
1378:
1376:
1375:
1370:
1364:
1362:
1357:
1355:
1353:
1348:
1347:
1345:
1335:
1334:
1329:
1323:
1321:
1316:
1315:
1313:
1311:
1310:
1305:
1299:
1297:
1292:
1291:
1289:
1287:
1286:
1281:
1275:
1273:
1268:
1266:
1264:
1259:
1257:
1255:
1250:
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1240:
1238:
1236:
1235:
1230:
1224:
1222:
1217:
1216:
1214:
1212:
1211:
1206:
1200:
1198:
1193:
1192:
1190:
1188:
1187:
1186:Haplonycteris
1182:
1176:
1174:
1169:
1167:
1165:
1160:
1158:
1156:
1151:
1150:
1148:
1138:
1137:
1132:
1126:
1124:
1119:
1117:
1115:
1110:
1108:
1106:
1101:
1099:
1097:
1092:
1090:
1088:
1083:
1081:
1079:
1074:
1072:
1070:
1069:C. brachyotis
1065:
1064:
1062:
1052:
1051:
1046:
1040:
1038:
1033:
1032:
1030:
1028:
1027:
1022:
1016:
1015:
1011:
1009:
1007:
1002:
1001:
999:
997:
996:
995:Balionycteris
991:
985:
983:
978:
977:
975:
973:
972:
967:
961:
959:
954:
952:
950:
945:
944:
942:
932:
931:
926:
922:
918:
912:
908:
894:
892:
887:
885:
883:
878:
877:
875:
873:
872:
871:Paranyctimene
867:
861:
859:
854:
852:
850:
845:
843:
841:
836:
834:
832:
827:
825:
823:
818:
816:
814:
809:
807:
805:
800:
798:
796:
791:
789:
787:
786:N. draconilla
782:
780:
778:
773:
771:
769:
764:
762:
760:
759:N. cephalotes
755:
753:
751:
750:N. albiventer
746:
744:
742:
737:
736:
734:
724:
723:
718:
714:
710:
704:
700:
694:
690:
688:
684:
682:
678:
676:
672:
670:
666:
664:
660:
659:
656:
652:
644:
639:
637:
632:
630:
625:
624:
621:
615:
611:
608:
605:
604:
595:
592:
589:
585:
582:
579:
576:
572:
569:
566:
565:
549:
544:
540:
536:
535:
530:
528:
519:
515:
505:
502:
501:
495:
493:
489:
479:
476:
472:
468:
464:
454:
452:
447:
445:
441:
437:
433:
429:
425:
424:
413:
410:
399:
397:
393:
382:
378:
376:
372:
368:
364:
360:
356:
352:
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344:
335:
333:
329:
325:
321:
317:
313:
309:
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304:
299:
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291:
287:
283:
280:
276:
275:
270:
266:
255:
251:
246:
241:
235:
233:
227:
224:
223:Binomial name
220:
216:
215:
210:
207:
206:
203:
202:
198:
195:
194:
191:
188:
185:
184:
181:
178:
175:
174:
171:
168:
165:
164:
161:
158:
155:
154:
151:
148:
145:
144:
141:
138:
135:
134:
129:
124:
120:
114:
109:
98:
94:
89:
84:
81:
73:
70:
62:
52:
48:
42:
41:
35:
30:
21:
20:
3675:
3476:
3437:
3425:
3413:
3404:
3395:
3385:Casinycteris
3383:
3359:P. anchietae
3358:
3346:
3321:
3312:
3303:
3283:
3272:M. woermanni
3271:
3262:
3250:
3238:
3226:
3218:Myonycterini
3201:
3191:Nanonycteris
3189:
3177:
3168:
3148:
3136:
3126:Hypsignathus
3124:
3113:E. franqueti
3112:
3103:
3094:
3074:
3063:E. wahlbergi
3062:
3053:
3044:
3035:
3026:
3018:E. gambianus
3017:
3009:E. crypturus
3008:
2999:
2990:
2970:
2962:Epomophorini
2923:
2914:
2905:
2896:
2887:
2878:
2869:
2860:
2851:
2842:
2833:
2824:
2815:
2797:
2785:
2776:
2767:
2747:
2709:
2700:
2690:Styloctenium
2688:
2677:P. woodfordi
2676:
2667:
2658:
2649:
2640:
2631:
2622:
2613:
2605:P. speciosus
2604:
2595:
2586:
2578:P. samoensis
2577:
2568:
2559:
2550:
2541:
2532:
2524:P. pselaphon
2523:
2514:
2505:
2496:
2487:
2480:P. pelagicus
2478:
2471:
2462:
2453:
2444:
2435:
2426:
2418:P. melanotus
2417:
2408:
2399:
2391:P. mariannus
2390:
2382:P. mahaganus
2381:
2372:
2363:
2354:
2345:
2336:
2327:
2318:
2309:
2300:
2291:
2282:
2273:
2264:
2255:
2246:
2237:
2228:
2219:
2210:
2201:
2192:
2183:
2175:P. anetianus
2174:
2166:P. allenorum
2165:
2156:
2147:
2138:
2118:
2106:
2097:
2089:P. flanneryi
2088:
2079:
2070:
2058:
2046:
2034:
2023:M. acrodonta
2022:
2010:
1998:
1990:E. dupreanum
1989:
1969:
1957:
1948:
1936:
1924:
1915:
1906:
1897:
1888:
1876:
1865:Pteropodinae
1838:
1830:S. carolinae
1829:
1821:S. australis
1820:
1802:Syconycteris
1800:
1788:
1779:
1759:
1748:M. woodfordi
1747:
1738:
1729:
1719:Melonycteris
1717:
1705:
1696:
1678:Macroglossus
1676:
1638:
1629:
1617:
1605:
1596:
1587:
1578:
1569:
1560:
1551:
1542:
1533:
1525:D. crenulata
1524:
1515:
1507:D. beauforti
1506:
1498:D. anderseni
1497:
1477:
1465:
1453:
1415:
1408:
1396:
1385:S. blanfordi
1384:
1372:
1360:
1351:
1331:
1319:
1307:
1295:
1283:
1271:
1262:
1253:
1245:M. ecaudatus
1244:
1232:
1221:L. salimalii
1220:
1208:
1196:
1184:
1173:D. spadiceus
1172:
1170:
1163:
1154:
1134:
1122:
1113:
1104:
1095:
1086:
1077:
1068:
1048:
1036:
1024:
1012:
1005:
993:
981:
971:Alionycteris
969:
957:
948:
928:
917:Cynopterinae
890:
881:
869:
858:N. vizcaccia
857:
849:N. robinsoni
848:
839:
830:
821:
812:
803:
794:
785:
776:
767:
758:
749:
740:
720:
709:Nyctimeninae
685:Superorder:
679:Infraclass:
651:Pteropodidae
552:. Retrieved
538:
532:
526:
518:
487:
485:
470:
466:
462:
460:
457:Reproduction
450:
448:
421:
419:
408:
405:
395:
391:
388:
379:
374:
343:D. spadiceus
342:
341:
338:Distribution
315:
311:
307:
301:
273:
272:
268:
264:
262:
231:
229:
213:
212:
200:
190:Pteropodidae
80:
65:
56:
37:
3759:Dyacopterus
3555:iNaturalist
3501:Wikispecies
3414:C. ophiodon
3396:C. argynnis
3322:M. torquata
3285:Myonycteris
3178:M. pusillus
3036:E. labiatus
2972:Epomophorus
2736:Rousettinae
2710:S. wallacei
2650:P. vampyrus
2623:P. tonganus
2551:P. rennelli
2463:P. ocularis
2373:P. macrotis
2328:P. keyensis
2301:P. howensis
2274:P. fundatus
2265:P. faunulus
2220:P. cognatus
2193:P. caniceps
2184:P. aruensis
1938:Desmalopex
1925:A. mackloti
1916:A. leucotis
1739:M. melanops
1706:M. sobrinus
1516:D. chapmani
1466:A. bulmerae
1333:Ptenochirus
1285:Otopteropus
1272:M. wetmorei
1263:M. niphanae
1254:M. kusnotoi
1197:H. fischeri
1164:D. rickarti
1136:Dyacopterus
1006:B. maculata
949:A. aequalis
777:N. cyclotis
554:19 November
467:Dyacopterus
432:Sungai Rayu
396:Dyacopterus
385:Description
303:Dyacopterus
286:Sunda Shelf
279:frugivorous
201:Dyacopterus
51:introducing
3748:Categories
3691:Q109647213
3438:S. zenkeri
3313:M. relicta
3263:M. azagnyi
3104:E. dobsoni
3045:E. minimus
3027:E. grandis
3000:E. anselli
2948:Subfamily
2915:R. lanosus
2786:E. robusta
2777:E. spelaea
2749:Eonycteris
2734:Subfamily
2659:P. vetulus
2641:P. ualanus
2542:P. rayneri
2533:P. pumilus
2472:P. ornatus
2292:P. griseus
2256:P. ennisae
2098:P. pulchra
2060:Pteralopex
1907:A. jubatus
1898:A. humilis
1863:Subfamily
1761:Notopteris
1697:M. minimus
1663:Subfamily
1606:D. viridis
1588:D. peronii
1552:D. inermis
1543:D. exoleta
1440:Subfamily
1398:Thoopterus
1155:D. brooksi
1096:C. minutus
1050:Cynopterus
915:Subfamily
831:N. minutus
822:N. masalai
768:N. certans
707:Subfamily
693:Chiroptera
510:References
475:monogamous
371:Indonesian
332:sub-canopy
180:Chiroptera
59:March 2013
34:references
3338:Plerotini
2906:Subgenus
2836:Rousettus
2834:Subgenus
2825:R. bidens
2816:Subgenus
2799:Rousettus
2506:P. pohlei
2400:P. medius
2157:P. alecto
2080:P. atrata
2071:P. anceps
2047:N. frosti
2036:Neopteryx
1999:E. helvum
1839:S. hobbit
1534:D. emersa
1455:Aproteles
1374:Sphaerias
1352:P. jagori
1320:P. lucasi
1309:Penthetor
1234:Megaerops
1114:C. sphinx
958:A. alecto
930:Aethalops
882:P. raptor
840:N. rabori
795:N. keasti
722:Nyctimene
661:Kingdom:
359:Pontianak
208:Species:
146:Kingdom:
140:Eukaryota
3685:Wikidata
3625:13800094
3573:10457086
3492:Q1765328
3486:Wikidata
3348:Plerotes
3054:E. minor
2768:E. major
2569:P. rufus
2445:P. niger
2364:P. lylei
2120:Pteropus
2012:Mirimiri
1878:Acerodon
1561:D. minor
1479:Dobsonia
1361:P. minor
1210:Latidens
1026:Chironax
891:P. tenax
804:N. major
741:N. aello
681:Eutheria
675:Mammalia
669:Chordata
667:Phylum:
663:Animalia
610:Archived
574:251-265.
498:See also
440:emergent
324:Thailand
320:Malaysia
186:Family:
170:Mammalia
160:Chordata
156:Phylum:
150:Animalia
136:Domain:
113:IUCN 3.1
3718:8013088
3612:1004366
3547:2432776
3076:Epomops
2238:P. coxi
2107:P. taki
1971:Eidolon
691:Order:
673:Class:
298:Sumatra
282:megabat
242:, 1890)
196:Genus:
176:Order:
166:Class:
111: (
47:improve
3731:946856
3664:606019
3638:326081
3586:631568
3534:327334
2818:Boneia
436:Poring
347:Poring
326:, and
294:Borneo
240:Thomas
36:, but
3705:6C7YN
3651:83214
3568:IRMNG
3560:40975
3521:385HV
423:Ficus
351:Kubah
3726:ITIS
3713:GBIF
3633:NCBI
3599:6931
3594:IUCN
3581:ITIS
3542:GBIF
556:2021
539:2020
444:peat
402:Diet
357:and
314:and
296:and
263:The
3700:CoL
3620:MSW
3607:MDD
3529:EoL
3516:CoL
543:doi
267:or
3750::
3728::
3715::
3702::
3687::
3661::
3648::
3635::
3622::
3609::
3596::
3583::
3570::
3557::
3544::
3531::
3518::
3503::
3488::
2911::
2839::
2821::
537:.
531:.
369:,
353:,
349:,
322:,
310:,
306::
3440:)
3416:)
3407:)
3398:)
3361:)
3324:)
3315:)
3306:)
3274:)
3265:)
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3180:)
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2670:)
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2001:)
1992:)
1960:)
1951:)
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1918:)
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1900:)
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1116:)
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806:)
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788:)
779:)
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761:)
752:)
743:)
642:e
635:t
628:v
558:.
545::
529:"
525:"
271:(
238:(
115:)
72:)
66:(
61:)
57:(
43:.
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