135:
111:
932:
Last, P.R.; Naylor, G.J.; Manjaji-Matsumoto, B.M. (2016). "A revised classification of the family
Dasyatidae (Chondrichthyes: Myliobatiformes) based on new morphological and molecular insights".
416:
1297:"Comment on 'Annotated checklist of the living sharks, batoids and chimaeras (Chondrichthyes) of the world, with a focus on biogeographical diversity' by Weigmann (2016)"
470:
1165:
Weigmann, S. (2016). "Annotated checklist of the living sharks, batoids and chimaeras (Chondrichthyes) of the world, with a focus on biogeographical diversity".
726:
693:
371:
899:
1656:
1117:
sp. nov. (Myliobatoidae, Dasyatidae), a new ocellated whipray from the tropical Indo-West
Pacific, described from its cytochrome-oxidase I gene sequence"
439:
520:
480:
459:
1651:
1546:
1507:
1533:
1032:
Fernando, D.; Brown, R.M.K.; Tanna, A.; Gobiraj, R.; Ralicki, H.; Jockusch, E.L.; Ebert, D.A.; Jensen, K.; Caira, J.N. (2019).
1538:
1296:
1112:
1209:
1608:
1011:
1551:
911:
247:
1481:
1468:
971:
1486:
1590:
134:
1613:
1345:
Philippe Borsa; Collin T. Williams; Ashlie J. McIvor; Thierry B. Hoareau; Michael L. Berumen (2021).
1235:
999:
781:
1344:
243:
759:
496:
449:
270:
1577:
1236:"Species boundaries in the Himantura uarnak species complex (Myliobatiformes : Dasyatidae)"
1408:
1346:
867:
1572:
715:
391:
1600:
1564:
1455:
1362:
1311:
1250:
1174:
845:
823:
803:
737:
682:
671:
649:
548:, and subsequently shown to be genetically distinct and reproductively isolated from both
8:
1646:
1641:
1446:
1084:
834:
792:
748:
638:
627:
616:
605:
428:
356:
Twelve extant species are currently recognized as valid, formerly four or five species.
1366:
1315:
1254:
1178:
1111:
Borsa, P.; Durand, J.-D.; Shen, K.-N.; Arlyza, I.S.; Solihin, D.D.; Berrebi, P. (2013).
1636:
1388:
1274:
1234:
Arlyza, I.S.; Shen, K.-N.; Solihin, D.D.; Soedharma D.; Berrebi, P.; Borsa, P. (2013).
1147:
1063:
770:
704:
490:
485:
Borsa, Durand, Shen, Alyza, Solihin & Berrebi, 2013 (Fine-spotted leopard whipray)
361:
305:
264:
129:
118:
1559:
1494:
1392:
1327:
1266:
1190:
1139:
1067:
1055:
949:
878:
856:
594:
533:
504:
401:
216:
1278:
1151:
1378:
1370:
1319:
1258:
1182:
1131:
1045:
941:
294:
1499:
1460:
1473:
411:
206:
1262:
1233:
1374:
1135:
660:
525:
510:
381:
186:
176:
1050:
1033:
945:
1630:
1431:
1110:
1031:
967:
931:
564:
present leopard-like ocellated spots. These are smaller and less numerous in
420:
64:
1331:
1270:
1194:
1143:
1059:
953:
301:
256:
1520:
1440:
1347:"Neotype designation and re-description of Forsskål's reticulate whipray
1034:"New insights into the identities of the elasmobranch fauna of Sri Lanka"
326:
320:
314:
110:
39:
1383:
1512:
1207:
332:
297:
226:
84:
49:
1323:
1186:
1525:
344:
338:
146:
100:
89:
33:
1402:
1425:
980:
290:
196:
166:
79:
74:
59:
54:
44:
94:
69:
156:
1585:
1208:
Eschmeyer, W.N.; R. Fricke; R. van der Laan (24 April 2018).
286:
524:
has had its validity disputed and has been considered a
1085:"Extinct - valid species | Species | Shark-References"
376:
Last, White & Naylor, 2016 (Australian whipray)
308:revision, many of the species formerly assigned to
1229:
1227:
1628:
1338:
406:(Chabanaud, 1923) (Marbled freshwater whip ray)
1106:
1104:
997:
1224:
1290:
1288:
1164:
1101:
1158:
1657:Taxa named by Friedrich Gustav Jakob Henle
109:
1382:
1294:
1285:
1049:
993:
991:
966:
925:
366:(Annandale, 1909) (Pale-spot whip ray)
1629:
386:(Annandale, 1909) (honeycomb whipray)
1407:
1406:
1243:Molecular Phylogenetics and Evolution
1591:735C87F1-8A63-FF9B-9BF6-FD195F428A3C
1079:
1077:
988:
1652:Taxa named by Johannes Peter Müller
13:
1201:
433:(Blyth, 1860) (Blackedge whipray)
396:(Hamilton, 1822) (Ganges whipray)
14:
1668:
1074:
540:was previously confused not with
454:(Chen, 1948) (Smalleye whip ray)
312:were reassigned to other genera (
1212:. California Academy of Sciences
1012:World Register of Marine Species
518:The former contentious species,
133:
37:
1025:
960:
892:
1:
885:
499:, 1789) (Reticulate whipray)
444:(Sahni & Mehrotra, 1981)
16:Genus of cartilaginous fishes
904:Müller and Henle 1837 (ray)"
7:
1263:10.1016/j.ympev.2012.09.023
571:
10:
1673:
1375:10.1007/s12526-021-01180-1
1136:10.1016/j.crvi.2013.01.004
513:, 1852) (Leopard whipray)
351:
1415:
1051:10.11646/zootaxa.4585.2.1
1004:Müller & Henle, 1837"
946:10.11646/zootaxa.4139.3.2
782:Maculabatis pastinacoides
262:
255:
130:Scientific classification
128:
117:
108:
23:
1124:Comptes Rendus Biologies
998:Bailly, Nicolas (2014).
970:; Pauly, Daniel (eds.).
1304:Journal of Fish Biology
1167:Journal of Fish Biology
777:Himantura pastinacoides
760:Fluvitrygon oxyrhynchus
450:Himantura microphthalma
868:Pateobatis uarnacoides
471:Himantura souarfortuna
300:that is native to the
1573:Paleobiology Database
863:Himantura uarnacoides
727:Brevitrygon javaensis
716:Brevitrygon imbricata
392:Himantura fluviatilis
1089:shark-references.com
846:Fluvitrygon signifer
824:Maculabatis randalli
804:Urogymnus granulatus
799:Himantura ponapensis
755:Himantura oxyrhyncha
738:Pateobatis jenkinsii
694:Brevitrygon heterura
683:Urogymnus granulatus
672:Maculabatis gerrardi
650:Pateobatis jenkinsii
612:Himantura chaophraya
475:Adnet et al., 2020
119:Reticulate whipray (
1367:2021MarBd..51...28B
1355:Marine Biodiversity
1316:2017JFBio..90.1170B
1255:2013MolPE..66..429A
1210:"Catalog of Fishes"
1179:2016JFBio..88..837W
835:Styracura schmardae
830:Himantura schmardae
793:Urogymnus polylepis
788:Himantura polylepis
749:Urogymnus lobistoma
744:Himantura lobistoma
733:Himantura jenkinsii
722:Himantura javaensis
711:Himantura imbricata
689:Himantura heterurus
678:Himantura granulata
639:Urogymnus dalyensis
634:Himantura dalyensis
628:Urogymnus polylepis
623:Himantura chaophrya
617:Urogymnus polylepis
606:Pateobatis bleekeri
429:Himantura marginata
372:Himantura australis
1295:Borsa, P. (2017).
972:"Species in genus
841:Himantura signifer
819:Himantura randalli
810:Himantura punctata
771:Styracura pacifica
766:Himantura pacifica
705:Pateobatis hortlei
667:Himantura gerrardi
601:Himantura bleekeri
585:Himantura alcockii
505:Himantura undulata
412:Himantura leoparda
362:Himantura alcockii
265:Raja sephen uarnak
1624:
1623:
1560:Open Tree of Life
1409:Taxon identifiers
1324:10.1111/jfb.13235
1187:10.1111/jfb.12874
879:Brevitrygon walga
857:Maculabatis toshi
700:Himantura hortlei
595:Maculabatis astra
581:Himantura alcocki
534:Catalog of Fishes
514:
500:
486:
476:
465:
455:
445:
434:
424:
417:Manjaji-Matsumoto
407:
402:Himantura krempfi
397:
387:
377:
367:
278:
277:
251:
1664:
1617:
1616:
1604:
1603:
1594:
1593:
1581:
1580:
1568:
1567:
1555:
1554:
1542:
1541:
1529:
1528:
1516:
1515:
1503:
1502:
1490:
1489:
1477:
1476:
1464:
1463:
1451:
1450:
1449:
1436:
1435:
1434:
1404:
1403:
1397:
1396:
1386:
1349:Himantura uarnak
1342:
1336:
1335:
1310:(4): 1170–1175.
1301:
1292:
1283:
1282:
1240:
1231:
1222:
1221:
1219:
1217:
1205:
1199:
1198:
1162:
1156:
1155:
1121:
1108:
1099:
1098:
1096:
1095:
1081:
1072:
1071:
1053:
1029:
1023:
1022:
1020:
1018:
995:
986:
985:
964:
958:
957:
929:
923:
922:
920:
919:
910:. Archived from
896:
814:Himantura uarnak
508:
494:
491:Himantura uarnak
484:
474:
464:(Bleeker, 1852)
463:
453:
443:
440:Himantura menoni
432:
415:
405:
395:
385:
375:
365:
242:
138:
137:
121:Himantura uarnak
113:
99:
36:
29:Temporal range:
21:
20:
1672:
1671:
1667:
1666:
1665:
1663:
1662:
1661:
1627:
1626:
1625:
1620:
1612:
1607:
1599:
1597:
1589:
1584:
1576:
1571:
1563:
1558:
1550:
1545:
1537:
1532:
1524:
1519:
1511:
1506:
1498:
1493:
1485:
1480:
1472:
1467:
1459:
1454:
1445:
1444:
1439:
1430:
1429:
1424:
1411:
1401:
1400:
1343:
1339:
1299:
1293:
1286:
1238:
1232:
1225:
1215:
1213:
1206:
1202:
1173:(3): 837–1037.
1163:
1159:
1119:
1115:Himantura tutul
1109:
1102:
1093:
1091:
1083:
1082:
1075:
1030:
1026:
1016:
1014:
996:
989:
965:
961:
930:
926:
917:
915:
898:
897:
893:
888:
874:Himantura walga
852:Himantura toshi
645:Himantura draco
590:Himantura astra
574:
521:Himantura tutul
481:Himantura tutul
460:Himantura pareh
354:
274:
268:
241:
207:Myliobatiformes
132:
104:
98:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
31:
30:
27:
17:
12:
11:
5:
1670:
1660:
1659:
1654:
1649:
1644:
1639:
1622:
1621:
1619:
1618:
1605:
1595:
1582:
1569:
1556:
1543:
1530:
1517:
1504:
1491:
1478:
1465:
1452:
1437:
1421:
1419:
1413:
1412:
1399:
1398:
1337:
1284:
1249:(1): 429–435.
1223:
1200:
1157:
1100:
1073:
1044:(2): 201–238.
1024:
987:
968:Froese, Rainer
959:
940:(3): 345–368.
924:
890:
889:
887:
884:
883:
882:
871:
860:
849:
838:
827:
816:
807:
796:
785:
774:
763:
752:
741:
730:
719:
708:
697:
686:
675:
664:
661:Pateobatis fai
653:
642:
631:
620:
609:
598:
587:
573:
570:
526:junior synonym
516:
515:
501:
487:
477:
466:
456:
446:
435:
425:
408:
398:
388:
382:Himantura fava
378:
368:
353:
350:
276:
275:
269:
260:
259:
253:
252:
234:
230:
229:
224:
220:
219:
214:
210:
209:
204:
200:
199:
194:
190:
189:
187:Elasmobranchii
184:
180:
179:
177:Chondrichthyes
174:
170:
169:
164:
160:
159:
154:
150:
149:
144:
140:
139:
126:
125:
115:
114:
106:
105:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
28:
15:
9:
6:
4:
3:
2:
1669:
1658:
1655:
1653:
1650:
1648:
1645:
1643:
1640:
1638:
1635:
1634:
1632:
1615:
1610:
1606:
1602:
1596:
1592:
1587:
1583:
1579:
1574:
1570:
1566:
1561:
1557:
1553:
1548:
1544:
1540:
1535:
1531:
1527:
1522:
1518:
1514:
1509:
1505:
1501:
1496:
1492:
1488:
1483:
1479:
1475:
1470:
1466:
1462:
1457:
1453:
1448:
1442:
1438:
1433:
1427:
1423:
1422:
1420:
1418:
1414:
1410:
1405:
1394:
1390:
1385:
1380:
1376:
1372:
1368:
1364:
1360:
1356:
1352:
1350:
1341:
1333:
1329:
1325:
1321:
1317:
1313:
1309:
1305:
1298:
1291:
1289:
1280:
1276:
1272:
1268:
1264:
1260:
1256:
1252:
1248:
1244:
1237:
1230:
1228:
1211:
1204:
1196:
1192:
1188:
1184:
1180:
1176:
1172:
1168:
1161:
1153:
1149:
1145:
1141:
1137:
1133:
1129:
1125:
1118:
1116:
1107:
1105:
1090:
1086:
1080:
1078:
1069:
1065:
1061:
1057:
1052:
1047:
1043:
1039:
1035:
1028:
1013:
1009:
1005:
1003:
994:
992:
983:
982:
977:
975:
969:
963:
955:
951:
947:
943:
939:
935:
928:
914:on 2023-03-12
913:
909:
905:
903:
895:
891:
881:
880:
876:: synonym of
875:
872:
870:
869:
865:: synonym of
864:
861:
859:
858:
854:: synonym of
853:
850:
848:
847:
843:: synonym of
842:
839:
837:
836:
832:: synonym of
831:
828:
826:
825:
821:: synonym of
820:
817:
815:
812:: synonym of
811:
808:
806:
805:
801:: synonym of
800:
797:
795:
794:
790:: synonym of
789:
786:
784:
783:
779:: synonym of
778:
775:
773:
772:
768:: synonym of
767:
764:
762:
761:
757:: synonym of
756:
753:
751:
750:
746:: synonym of
745:
742:
740:
739:
735:: synonym of
734:
731:
729:
728:
724:: synonym of
723:
720:
718:
717:
713:: synonym of
712:
709:
707:
706:
702:: synonym of
701:
698:
696:
695:
691:: synonym of
690:
687:
685:
684:
680:: synonym of
679:
676:
674:
673:
669:: synonym of
668:
665:
663:
662:
658:: synonym of
657:
656:Himantura fai
654:
652:
651:
647:: synonym of
646:
643:
641:
640:
636:: synonym of
635:
632:
630:
629:
625:: synonym of
624:
621:
619:
618:
614:: synonym of
613:
610:
608:
607:
603:: synonym of
602:
599:
597:
596:
592:: synonym of
591:
588:
586:
583:: synonym of
582:
579:
578:
577:
569:
567:
563:
559:
556:. Both adult
555:
551:
547:
543:
539:
535:
531:
527:
523:
522:
512:
507:
506:
502:
498:
493:
492:
488:
483:
482:
478:
473:
472:
467:
462:
461:
457:
452:
451:
447:
442:
441:
436:
431:
430:
426:
422:
418:
414:
413:
409:
404:
403:
399:
394:
393:
389:
384:
383:
379:
374:
373:
369:
364:
363:
359:
358:
357:
349:
347:
346:
341:
340:
335:
334:
329:
328:
323:
322:
317:
316:
311:
307:
303:
299:
296:
292:
288:
284:
283:
272:
267:
266:
261:
258:
254:
249:
245:
240:
239:
235:
232:
231:
228:
225:
222:
221:
218:
215:
212:
211:
208:
205:
202:
201:
198:
195:
192:
191:
188:
185:
182:
181:
178:
175:
172:
171:
168:
165:
162:
161:
158:
155:
152:
151:
148:
145:
142:
141:
136:
131:
127:
124:
122:
116:
112:
107:
102:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
35:
26:
22:
19:
1416:
1384:10754/665804
1358:
1354:
1348:
1340:
1307:
1303:
1246:
1242:
1214:. Retrieved
1203:
1170:
1166:
1160:
1130:(2): 82–92.
1127:
1123:
1114:
1092:. Retrieved
1088:
1041:
1037:
1027:
1015:. Retrieved
1007:
1001:
979:
973:
962:
937:
933:
927:
916:. Retrieved
912:the original
907:
901:
894:
877:
873:
866:
862:
855:
851:
844:
840:
833:
829:
822:
818:
813:
809:
802:
798:
791:
787:
780:
776:
769:
765:
758:
754:
747:
743:
736:
732:
725:
721:
714:
710:
703:
699:
692:
688:
681:
677:
670:
666:
659:
655:
648:
644:
637:
633:
626:
622:
615:
611:
604:
600:
593:
589:
584:
580:
576:Data from:
575:
565:
561:
557:
553:
549:
545:
541:
537:
529:
519:
517:
503:
497:J. F. Gmelin
489:
479:
469:
458:
448:
438:
427:
410:
400:
390:
380:
370:
360:
355:
343:
337:
331:
325:
319:
313:
309:
304:. In a 2016
302:Indo-Pacific
281:
280:
279:
271:J. F. Gmelin
263:
257:Type species
244:J. P. Müller
237:
236:
193:Superorder:
120:
32:37.2–0
24:
18:
1521:iNaturalist
1441:Wikispecies
908:Fossilworks
558:H. leoparda
554:H. leoparda
546:H. leoparda
544:, but with
536:. However,
327:Maculabatis
321:Fluvitrygon
315:Brevitrygon
227:Urogymninae
223:Subfamily:
1647:Ray genera
1642:Dasyatidae
1631:Categories
1094:2023-07-06
918:2023-07-06
886:References
333:Pateobatis
298:Dasyatidae
217:Dasyatidae
183:Subclass:
103:to Present
1637:Himantura
1474:Himantura
1461:Himantura
1447:Himantura
1417:Himantura
1393:255610805
1361:(2): 28.
1068:146082424
1002:Himantura
974:Himantura
902:Himantura
550:H. uarnak
542:H. uarnak
530:H. uarnak
345:Urogymnus
339:Styracura
310:Himantura
306:taxonomic
282:Himantura
238:Himantura
153:Kingdom:
147:Eukaryota
101:Oligocene
25:Himantura
1426:Wikidata
1332:28026866
1279:17310045
1271:23023209
1216:24 April
1195:26860638
1152:13204782
1144:23608177
1060:31716166
981:FishBase
954:27470808
572:Synonyms
566:H. tutul
562:H. tutul
538:H. tutul
423:, 2008
291:stingray
213:Family:
197:Batoidea
167:Chordata
163:Phylum:
157:Animalia
143:Domain:
1513:2419609
1432:Q135505
1363:Bibcode
1312:Bibcode
1251:Bibcode
1175:Bibcode
1038:Zootaxa
934:Zootaxa
532:by the
511:Bleeker
352:Species
293:in the
233:Genus:
203:Order:
173:Class:
1614:105752
1601:684642
1598:uBio:
1578:354629
1565:674419
1539:160971
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1066:
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1017:2 July
952:
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295:family
273:, 1789
250:, 1837
246:&
1609:WoRMS
1586:Plazi
1552:86362
1526:47278
1389:S2CID
1300:(PDF)
1275:S2CID
1239:(PDF)
1148:S2CID
1120:(PDF)
1064:S2CID
1008:WoRMS
287:genus
285:is a
248:Henle
1547:NCBI
1534:ITIS
1508:GBIF
1500:4XB4
1487:2975
1482:BOLD
1328:PMID
1267:PMID
1218:2018
1191:PMID
1140:PMID
1056:PMID
1042:4585
1019:2023
950:PMID
938:4139
560:and
552:and
421:Last
342:and
40:PreꞒ
1495:CoL
1469:AFD
1456:ADW
1379:hdl
1371:doi
1320:doi
1259:doi
1183:doi
1132:doi
1128:336
1046:doi
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528:of
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289:of
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95:N
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75:T
70:P
65:C
60:D
55:S
50:O
45:Ꞓ
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