976:, had over 1,100 people come in 2007. Visitor numbers have been increasing at about 10% annually and development of additional bear viewing areas on Kodiak is planned. Also, other bear viewing opportunities exist through air-taxi, charter boat, remote lodge, and trekking operations on the archipelago. Although bear-viewing is often considered a "nonconsumptive" use, it can have serious impacts on bear populations if it is not conducted properly. Most viewing occurs at places where bears congregate because of feeding opportunities that are critical to their survival. If some bears avoid these areas because people are there, those bears may not get the fat and protein they need to make it through the upcoming winter. Consequently, unmanaged bear viewing could affect several bears, especially productive sows with cubs. Often, bear-viewing and bear-hunting are considered incompatible. Even if the bear population is healthy and bear hunting is sustainable, ethical questions arise especially if hunting occurs near viewing areas and either during or soon after the viewing season. Many feel that it is not fair to encourage bears to be close to people during the summer, only to allow them to be shot in the fall. The Kodiak bear plan recognized bear hunting as a legitimate, traditional, and biologically justifiable activity. It recommended that agencies find ways to make bear hunting and bear viewing compatible on the archipelago.
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different areas during two seasons (spring: April 1 β May 15, and fall: October 25 β November 30). Each year, about 4,500 people apply for the 496 permits offered for Kodiak bear hunts (two-thirds to Alaska residents, one-third to non-residents). Nonresidents are required to hire a registered guide who is authorized to hunt in a particular area, and this can cost from $ 10,000 to $ 22,000. All hunters must come into the Alaska
Department of Fish and Game office in Kodiak before going into the field for a brief orientation and must check out before they leave the island. Every bear that is legally killed on the archipelago must be inspected by an ADF&G wildlife biologist before it can be taken from the islands. Pelts receive a stamp from an ADF&G officer if the hunter and guide provide proper documentation to prove licensing. Pelts cannot be transported or legally preserved or sold without the official stamp. Hunting laws are strictly enforced by the ADF&G officers who often have the full support of the local community. Illegal hunting and fishing is frowned upon by the community which maintains a healthy respect for the island's environmental laws, as well. Stiff penalties accompany illegal hunting and fishing. The island's remote location makes trafficking in illegal pelts difficult for would-be poachers.
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Mating season for Kodiak bears is during May and June. They are serially monogamous (having one partner at a time), staying together from two days to two weeks. As soon as the egg is fertilized and divides a few times, it enters a state of suspended animation until autumn when it finally implants on the uterine wall and begins to grow again. Cubs are born in the den during
January or February. Weighing less than 450 g (1 lb) at birth with little hair and closed eyes, they suckle for several months, emerging from the den in May or June, weighing 6.8β9.1 kilograms (15β20 pounds). Typical litter sizes on Kodiak are two or three cubs, with a long-term average of 2.4 cubs per litter. However, Kodiak bears have six functional nipples and can litter up to six cubs have been reported. Sows are sometimes seen with five or six cubs in tow, probably due to adopting cubs from other litters. Most cubs stay with their mothers for three years. Almost half of the cubs die before they leave, with
509:, commonly reaching sizes of 300 to 600 kg (660 to 1,320 lb). The size range for females (sows) is from 181 to 318 kg (399 to 701 lb), and for males (boars), it is 272 to 635 kg (600 to 1,400 lb). Mature males average 477β534 kg (1,052β1,177 lb) over the course of the year, and can weigh up to 680 kg (1,500 lb) at peak times. Females are typically about 20% smaller and 30% lighter than males, and adult sizes are attained when they are six years old. Bears weigh the least when they emerge from their dens in the spring, and can increase their weight by 20β30% during late summer and fall. As with other animals,
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the new regulations seemed to restore bear populations on the Kodiak
Islands. By the 1930s, ranchers in northeast Kodiak reported an increase in bear problems and demanded action. Bears were wrongly seen as a threat to the expanding commercial salmon-fishing industry. To address the dilemma of conserving bears while protecting cattle, salmon, and people, President Franklin D. Roosevelt created the Kodiak National Wildlife Refuge by executive order in 1941. The 7,700 km (1,900,000 acres) refuge roughly encompasses the southwestern two-thirds of Kodiak Island, Uganik Island, the Red Peaks area on northwestern Afognak Island, and all of Ban Island.
642:(active at night) lifestyle. This behavior is especially evident in the bears that live near and within Kodiak City. Kodiak bears do not defend territories, but they do have traditional areas that they use each year (home ranges). Because of the rich variety of foods available on Kodiak, the bears on the archipelago have some of the smallest home ranges of any brown bear populations in North America and a great deal of overlap occurs among the ranges of individual bears. Home ranges of adult sows on Kodiak Island average 130 km (50 sq mi), while boar home ranges average 250 km (97 sq mi).
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hunter being killed by the bear, and occurred on Uganik Island (November 3, 1999), which is part of the Kodiak archipelago. In the other incident, after being attacked by the bear, the hunter stabbed it with a knife, then recovered their rifle and killed the attacking bear. This occurred on
Raspberry Island, home to two full-service wilderness lodges. Prior to that, the last fatality was in 1921. About once every other year, a bear injures a person on Kodiak. In October 2021, a father and son hunting duo survived an attack from a Kodiak bear during an elk registration hunt on Afognak Island.
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impacts of the oil spill and the subsequent cleanup and settlement proved to be beneficial to bears on Kodiak. Bear-safety training exposed thousands of workers to factual information about bears, and money from the settlement fund was used for funding land acquisitions. By the close of the 20th century, over 80% of the refuge lands that had been lost as a result of ANCSA and ANILCA were reinstated into the refuge, either through direct purchase or utilizing conservation easements. Lands were also purchased in
America, Westtown, and
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grass/sedge flats, berry patches, a dead whale, or even an open garbage dump, they often occur in large groups. Along a few streams on Kodiak, up to 60 bears can be seen simultaneously in a 2.6 km (1.0 sq mi) area. To maximize food intake at these ecologically important areas, bears have learned to minimize fighting and fatal interactions by developing a complex communication (both verbal and body posturing) and social structure.
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Archipelago Bear
Conservation and Management Plan was crafted over several months by representatives from 12 diverse user groups, which, after hearing from a variety of experts from agencies and receiving extensive public input, developed more than 270 recommendations for managing and conserving Kodiak bears. Despite the diversity of viewpoints expressed by members of the group, all of the recommendations were by consensus.
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additional nine bears were reported killed annually in defense of life or property during the same time. The number of large, trophy-sized bears (total skull size at least 70 cm ) killed by hunters in recent years has been increasing. In the 1970s, only 2.5% of the bears killed on Kodiak were trophy-sized; in the 1990s and 2000s, the proportion increased to almost 9%.
748:, or cattle, internal organs are eaten first for their high-fat content, however even though there is an abundance of the animals found on the archipelago, few Kodiak bears actively prey on them as other methods of finding food are more energy efficient. Another food source available year-round is the garbage made by the human population of Kodiak Island.
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longer denning periods than bears in the southern areas. Most Kodiak bears dig their dens in hill or mountain sides and they use a wide variety of denning habitats depending on which part of the archipelago they live. Almost a quarter of the adult bears forgo denning, staying somewhat active throughout the winter.
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not compatible on the same ranges and potential solutions included poisons, fences to isolate cattle ranges, and aerial shooting of bears. Again, sport hunters voiced their support for Kodiak bears. Despite public pressure, the state continued actively pursuing and dispatching problem bears until 1970.
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In 2001, a citizens advisory committee was established to work closely with the Alaska
Department of Fish and Game (ADF&G), with the cooperation of Kodiak NWR, to develop a management plan addressing several problems that affect bears, including hunting, habitat, and viewing. The resulting Kodiak
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and
Raspberry Islands and on the Kodiak National Wildlife Refuge, but liberalized bear seasons on nonrefuge lands on Kodiak. During the 1960s, state biologists worked with ranchers along the Kodiak road system to examine and reduce the predation problem. Biologists reported that cattle and bears were
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return. Salmon runs extend from May through
September on most of the archipelago and bears consume the five species of Pacific salmon that spawn in local streams and lakes. Bears often prioritize the brain, flesh, and eggs of salmon for their high nutritional value. In the late summer and early fall,
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Kodiak bears that have recently left their mothers, at ages 3β5 years, have high mortality rates with only 56% of males and 89% of females surviving. Most young female bears stay within or near their mother's home range, while most males move farther away. Most adult sows die of natural causes (56%),
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Kodiak bears have interacted with humans for centuries, especially hunters and other people in the rural coastal regions of the archipelago. The bears are hunted for sport and are encountered by hunters pursuing other species. Less frequently, Kodiak bears are killed by people whose property (such as
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Kodiak bear research and habitat protection is done cooperatively by the ADF&G and Kodiak
National Wildlife Refuge. Bear hunting is managed by the ADF&G, and hunting regulations are established by the Alaska Board of Game. Currently, a finely tuned management system distributes hunters in 32
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The underlying themes of the recommendations were continued conservation of the bear population at its current level, increased education programs to teach people how to live with bears on Kodiak, and protection of bear habitat with allowances for continued human use of the archipelago. Although the
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and transferred into state ownership. The Kodiak Brown Bear Trust coordinated a coalition of sportsmen and other wildlife conservation groups from around the nation to lobby for the use of settlement funds to acquire Kodiak lands. The groups also directly contributed funding to protect small parcels
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drainage. The hydro project was the first significant invasion of inland bear habitat on Kodiak Island. To address the opposition encountered from the public and agencies, a mitigation settlement was negotiated in 1981 which included brown bear research and the establishment of the Kodiak Brown Bear
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Professional interest in guided Kodiak bear hunts and concern for unregulated resource use in frontier lands such as Alaska prompted the territorial government's newly established Alaska Game Commission to abolish commercial bear hunting (selling the hides) on the archipelago in 1925. The impacts of
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looked to the sea for their sustenance. At that time, Kodiak Natives (Alutiiqs) occasionally hunted bears, using their meat for food, hides for clothing and bedding, and teeth for adornment. Traditional stories often revolved around the similarity between bears and humans, and the mystical nature of
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filmed a television program about brown bears, which included a segment on two brown bear attacks. Both incidents involved hunters who were hunting by themselves and were returning to game they had killed previously, and left alone in order to continue hunting. One of the attacks was fatal, with the
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Usually, Kodiak bears attempt to avoid encounters with people. The most notable exceptions to this behavior pattern occur when bears are surprised, threatened, or attracted by human food, garbage, or hunter-killed game. However, there has been an increase in Kodiak encounters due to increases in the
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on the northern islands, to steep, glaciated mountains rising to Koniag Peak's 1,360 m (4,470 ft) along the central spine of Kodiak Island, to rolling hills and flat tundra on the south end of the archipelago. About 14,000 people live on the archipelago, primarily in and around the city of
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In 1971, the Alaska Native Claims Settlement Act (ANCSA) resolved many long-standing land issues with Aboriginal Alaskans statewide. The impacts were felt strongly on the archipelago as large areas were conveyed to the Native corporations. Federal management of the National Forest lands on Afognak
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pelts. The number of bears harvested increased substantially when sea otter populations declined and after the United States acquired Alaska in 1867, bear harvests on Kodiak increased, peaking at as many as 250 bears per year. Commercial fishing activities increased in the late 1880s and canneries
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Kodiak bears reach sexual maturity at age five, but most sows are over nine years old when they successfully wean their first litter. The average time between litters is four years. Sows continue to produce cubs throughout their lives, but their productivity diminishes after they are 20 years old.
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Kodiak bears begin entering their dens in late October. Pregnant sows are usually the first to go to dens; males are the last. Males begin emerging from their dens in early April, while sows with new cubs may stay in dens until late June. Bears living on the north end of Kodiak Island tend to have
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male weighed 751 kg (1,656 lb), and had a hind foot measurement of 46 cm (18 in). A large male Kodiak bear stands up to 1.5 m (4.9 ft) tall at the shoulder when it is standing on all four legs. When standing fully upright on its hind legs, a large male could reach a
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Since statehood, the reported number of Kodiak bears killed by hunters has ranged from 77 (1968β1969) to 206 (1965β1966). From 2000 to 2006, an average of 173 Kodiak bears were killed by hunters each year (118 during the fall season and 55 in the spring season). Over 75% of those were males. An
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in 1989, although some were displaced from traditional feeding and traveling areas by cleanup crews. No one was injured by a bear, and no Kodiak bears were killed. To mitigate the adverse impacts of the spill, Exxon reached a settlement with the state and federal governments. Paradoxically, the
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Bears live throughout the archipelago, adapting to local resources and retaining relatively small home ranges and comparable densities in most habitats. With such a variety and abundance of food sources, bears are surprisingly intelligent in their eating habits. The first foods bears eat in the
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Hair colors range from pale blonde to orange (typically females or bears from southern parts of the archipelago) to dark brown. Cubs will often retain a white "natal ring" around their neck for the first few years of life. The Kodiak bear's color is similar to that of its close relatives, the
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The Kodiak bear is much like other brown bears in intelligence, although its tendency to feed in large dense groups leads to more complex social behaviors. Kodiak bears are generally solitary in nature; however, when food is concentrated in small areas, such as along salmon spawning streams,
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Trust. The hydroelectric project was completed in 1985. Human alteration of bear habitat on Kodiak and Afognak Islands spurred renewed interest and funding for bear research on the archipelago, resulting in a surge of baseline and applied bear research on Kodiak through the 1980s and 1990s.
533:. Nicknamed "Clyde", he weighed 966 kg (2,130 lb) when he died in June 1987 at the age of 22. According to zoo director Terry Lincoln, Clyde probably weighed close to 1,089 kg (2,400 lb) a year earlier. He still had a fat layer of 9 in (23 cm) when he died..
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proliferated throughout the archipelago. Bears were viewed as competitors for the salmon resource and were routinely shot when seen on streams or coasts. At the same time, sportsmen and scientists had recognized the Kodiak bear as the largest in the world, and they voiced concerns about
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with cool temperatures, overcast skies, fog, windy conditions, and moderate to heavy precipitation throughout most of the year. Although the archipelago only covers about 13,000 km (5,000 sq mi), a rich variety of topography and vegetation ranges from dense forests of
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was transferred to Native Corporation ownership with the passage of the Alaska National Interest Lands Conservation Act in 1980 (ANILCA), and the Kodiak National Wildlife Refuge lost control of 130,000 hectares (310,000 acres) of prime bear habitat (more than 17% of refuge lands).
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spring are emerging vegetation (such as grasses and forbs) and animals that may have died during the winter. This allows the bear to quickly replace the weight that was lost during hibernation. As summer progresses, a wide variety of vegetation supplies nutritional needs until
626:, and adjacent islands). The Kodiak bear population was estimated to include 3,526 bears in 2005, yielding an estimated archipelago-wide population density of 270 bears per 1000 km (700 per 1000 sq. mi). During the past decade, the population has been slowly increasing.
571:β "cheek bones"). The total skull size is the sum of these two measurements. The largest bear ever killed in North America was from Kodiak Island, with a total skull size of 78.1 cm (30.7 in), and eight of the top 10 brown bears listed in the
391:), with the main difference being size, as Kodiak bears are on average 1.5 to 2 times larger than their cousins. Despite this large variation in size, the diet and lifestyle of the Kodiak bear do not differ greatly from those of other brown bears.
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to ripen earlier, berry season is now overlapping with salmon season and some bears are abandoning salmon runs to focus on the berries. Bears also feed on wind-rowed seaweed and invertebrates on some beaches throughout the year. When eating
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local population as well as increased hunting of Kodiak bears. Bear safety precautions aim at avoiding such situations, understanding bear needs and behavior, and learning how to recognize the warning signs bears give when stressed.
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record book are from Kodiak. The average skull size of Kodiak bears that were killed by hunters in the first five years of the 21st century was 63.8 cm (25.1 in) for boars, and 55.4 cm (21.8 in) for sows.
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on Kodiak Island. That project included an earthen dam on Terror Lake with Kodiak National Wildlife Refuge and a 10 km (6 mi) tunnel through a mountain ridge to a penstock and powerhouse in the
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Russian hunters came to the area in the late 18th century to capitalize on the abundant fur resources. Bear hides were considered a "minor fur" and sold for about the same price as river
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group's role is merely advisory, government management agencies expressed a commitment to implement all of the regulations that were feasible and within their legal jurisdictions.
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While there is generally much variation in size between brown bears in different areas, most usually weigh between 115 and 360 kg (254 and 794 lb); the Kodiak bear illustrates
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McLoughlin, P. D.; Ferguson, S. H.; Messier, F. (2000). "Intraspecific variation in home range overlap with habitat quality: a comparison among brown bear populations".
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does not list subspecies. The brown bear species, of which the Kodiak subspecies is a member, is listed as Lower Risk or Least Concern. The Kodiak is not listed as an
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An average adult male measures 244 cm (8 ft 0 in) in length, and stands 133 cm (4 ft 4 in) tall at the shoulder. The largest recorded
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In the past 20 years, bear viewing has become increasingly popular on Kodiak and other parts of Alaska. The most accessible bear-viewing location on Kodiak,
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Appearance. The polar bear is the largest member of the bear family, with the exception of the Alaska's Kodiak brown bears, which equal polar bears in size.
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Alaska achieved statehood in 1959 and assumed responsibility for managing the state's wildlife. The Alaska Board of Game reduced bear-hunting seasons on
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while most adult male bears are killed by hunters (91%). The oldest known male bear in the wild was 27 years old, and the oldest female was 35.
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bears consume several types of berries when they reach their ripest point, and have the highest levels of sugar. As climate change causes
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Buckner, E. L. and J. Reneau. 2005. Records of North American Big Game. Boone & Crockett Club; 12.00 edition (October 1, 2005).
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456:). Genetic samples from bears on Kodiak have shown that they are most closely related to brown bears on the Alaska Peninsula and
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In 1975, construction of a logging road began on Afognak Island, and timber harvesting began in 1977. In 1979, work began on an
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For comparison, the record for the Californian grizzly and polar bears in the wilderness is about 998 kg (2,200 lb).
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exists within the population. Although the current population is healthy, productive, and has shown no overt adverse signs of
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Physiologically and physically, the Kodiak bear is very similar to the other brown bear subspecies, such as the mainland
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to a lesser extent, closely monitor the size and health of the population and the number of bears hunted in the state.
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Smith, R. B.; Van Daele, L. J. (1990). "Impacts of Hydroelectric Development on Brown Bears, Kodiak Island, Alaska".
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1719:"Kodiak Archipelago Bear. Conservation and Management Plan. Part 2. Biology, History, and Management of Kodiak Bears"
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2109:"Kodiak Archipelago Bear Conservation and Management Plan. Alaska Department of Fish and Game, Anchorage, U.S."
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1671:. Job completion report for Challenge Cost Share Project. Kodiak National Wildlife Refuge, Kodiak, Alaska, U.S.
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was the first to recognize the Kodiak bear as a unique subspecies of the brown bear, and he named it "
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Talbot, S. L. J. R. Gust, G. K. Sage, A. Fischbach, K. Amstrup, W. Leacock, and L.Vav Daele. (2006).
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height of 3 m (9.8 ft). The largest verified size for a captive Kodiak bear was for a
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and the northeastern part of Kodiak Island. About half of the archipelago is included in the
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livestock) or person are threatened. In recent history there has been an increasing focus on
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and protection of the Kodiak bear population as human activity in its range increases. The
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1551:. Alaska Department of Fish and Game, Juneau, Alaska, U.S. pp. 74β101. Archived from
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452:. Subsequent taxonomic work merged all North American brown bears into a single species (
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1724:. Alaska Department of Fish and Game, Division of Wildlife Conservation. Archived from
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Servheen, C.; Herrero, S.; Peyton, B.; Pelletier, K.; Moll, K.; Moll, J., eds. (1999),
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on the back of the skull to the front tooth), and the width (maximum width between the
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1922:. Anchorage, Alaska, the U.S.A.: Great Northwest Publishing and Distributing Company.
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and six outlying villages. Roads and other human alterations are generally limited to
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Alaska Department of Fish and Game files, 211 Mission Road, Kodiak, Alaska 99615 U.S.
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One fatal bear attack on a person on the Kodiak archipelago occurred in 1999. The
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Kodiak bears can sometimes weigh considerably more than their wild counterparts.
2006:"Status of Alaska Sea Otter Populations and Developing Conflicts With Fisheries"
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996:(Bear), with the pronunciation varying between Northern and Southern dialects.
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1865:. Alaska Department of Fish and Game, Division of Wildlife Conservation. 2008
1776:"Climate change is luring Kodiak bears away from their iconic salmon streams"
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1265:"Brown Bear-Human Interactions Associated with Deer Hunting on Kodiak Island"
1132:. Alaska Department of Fish and Game, Division of Wildlife Conservation. 2008
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1592:"Population dynamics and management of brown bears on Kodiak Island, Alaska"
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Also, an individual named Teddy, which portrayed a killer bear in the movie
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Early human occupants of the archipelago when the land was locked into the
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Kodiak bears are the largest brown bear and are even comparable in size to
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1313:. Final report to the Kodiak National Wildlife Refuge, Kodiak Alaska, U.S.
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The standard method of evaluating the size of bears is by measuring their
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Genetic tests demonstrate this population to have mixed brown bear and
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2238:"Kodiak Alutiiq Language Level I & II Teaching Resource Workbook"
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2157:"Endangered Species program, list of mammals endangered on the US"
1752:"Kodiak Brown Bear Fact Sheet, Alaska Department of Fish and Game"
1798:"The Social Behaviour of Brown Bears on an Alaskan Salmon Stream"
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Genetic characterization of brown bears of the Kodiak Archipelago
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Brown bear management report of survey and inventory activities
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Egbert, Allan L.; Stokes, Allen W.; Egbert, Allen L. (1976).
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This brown bear population only occurs on the islands of the
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54:"Alaskan brown bear" redirects here. Not to be confused with
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may be in need of reorganization to comply with Knowledge's
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Productivity of female brown bears on Kodiak Island, Alaska
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2185:. Alaska Department of Fish and Game. 2008. Archived from
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Alaska Department of Fish & Game. 2002. Archived from
1247:"Kodiak Archipelago Bear Conservation and Management Plan"
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368:. They are also considered by some to be a population of
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Kodiak Archipelago Bear Conservation and Management Plan
1517:(in Latin). Sumptibus C. Salfeld. pp. 138β39.
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by adult males being one of the major causes of death.
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bears because of their proximity to the spirit world.
2137:. IUCN International Union for Conservation of Nature
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2183:"Hunting Kodiak bears β a question and answer guide"
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1252:. Alaska Department of Fish and Game. February 2002.
448:" in honor of the celebrated Baltic naturalist, Dr.
407:, of which the Kodiak is a subspecies, as being of "
1419:. Enfield, Middlesex : Guinness Superlatives.
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Proceedings of the Biological Society of Washington
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1203:Bears: status survey and conservation action plan
472:than other, more diverse brown bear populations.
2895:
1993:, the U.S.A.: Alaska Natural History Association
1980:
1978:
1976:
1974:
1159:. U.S. Fish and Wildlife service. Archived from
468:, it may be more susceptible to new diseases or
2254:
1913:
1911:
1589:
1539:
1193:
895:Taxidermized bear in a sporting-goods store in
51:Largest subspecies of brown bears/grizzly bears
2159:. US Fish and Wildlife service. Archived from
1987:The History of Bears on the Kodiak Archipelago
1834:
1540:Van Daele, L.J. (1 July 2002 β 30 June 2004).
918:International Union for Conservation of Nature
883:of important bear habitat around the islands.
542:Smithsonian National Museum of Natural History
89:to make improvements to the overall structure.
2329:
1971:
1663:
1661:
1659:
1585:
1583:
1581:
1579:
1577:
1575:
1573:
1069:
1067:
684:The islands of the Kodiak Archipelago have a
1908:
1886:"Wounds, not cold, killed hunter exam finds"
1680:
1410:
1387:"Bear Facts Kodiak National Wildlife Refuge"
1381:
1379:
1121:
1076:"Preliminary Synopsis of the American Bears"
831:Guided hunters and competition for resources
654:
563:to measure the length of the skull (back of
2103:
2101:
1951:
1535:
1533:
1504:
1415:The Guinness Book of Animal Facts and Feats
1119:
1117:
1115:
1113:
1111:
1109:
1107:
1105:
1103:
1101:
771:
593:
15:
2336:
2322:
1883:
1656:
1570:
1064:
315:
155:
127:
16:
1984:
1702:
1667:Barnes, V.G. and Van Daele, L.J. (2006).
1462:
1376:
847:
788:
105:Learn how and when to remove this message
2475:East Siberian brown bear (provisionally
2098:
1530:
1098:
979:
964:takes guests from a wilderness lodge on
955:
890:
792:
755:
713:
658:
535:
496:
24:This is an accepted version of this page
2230:
2003:
1514:Prodromus Systematis Mammalium et Avium
1406:
1404:
1073:
425:United States Fish and Wildlife Service
14:
2896:
2036:
1958:349th Air Mobility Wing Public Affairs
1839:. Camera Q Productions. Archived from
1262:
873:Bears were not directly harmed by the
813:
356:. It is one of the largest recognized
2736:
2735:
2636:Peninsular giant bear (provisionally
2466:Kamchatkan brown bear (provisionally
2317:
1917:
1837:"A guide to brown bear country (DVD)"
1745:
1743:
1542:"Unit 8 brown bear management report"
1468:
1340:
1148:
1146:
760:Two adult Alaskan Brown bears at the
2944:Endemic mammals of the United States
2493:Himalayan brown bear (provisionally
2127:
2024:"Kodiak Bear Archives - Page 2 of 4"
1401:
1241:
1239:
1237:
62:
2904:IUCN Red List least concern species
2674:Stickeen brown bear (provisionally
2540:Marsican brown bear (provisionally
2072:Bears: Their Biology and Management
2010:US Fish & Wildlife Publications
1802:Bears: Their Biology and Management
1324:"Kodiak Bears - Bears Of The World"
1269:Bears: Their Biology and Management
1051:"IUCN Brown Bear subspecies status"
762:Alaska Wildlife Conservation Center
48:
2949:Taxa named by Clinton Hart Merriam
2596:California grizzly bear (formerly
1740:
1601:. University of Idaho, Moscow, U.S
1143:
938:
912:Genetic diversity and endangerment
421:Alaska Department of Fish and Game
49:
2960:
2520:Tibetan blue bear (provisionally
2511:Syrian brown bear (provisionally
2502:Ussuri brown bear (provisionally
2281:
2243:. Native Village of Afognak. 2009
1954:"Ready Airman repels bear attack"
1234:
862:Terror Lake hydroelectric project
709:
2549:Cantabrian brown bear (formerly
2299:
2287:
1501:News, Published February 3, 2011
1357:. April 23, 1993. Archived from
324:Kodiak bear range within Alaska
180:
67:
2709:
2606:Mexican grizzly bear (formerly
2175:
2149:
2063:
2030:
2016:
1997:
1945:
1936:
1877:
1855:
1828:
1789:
1768:
1711:
1674:
1613:
1521:
1433:
1391:Kodiak National Wildlife Refuge
1154:"Polar bear, (Ursus maritimus)"
1015:
1006:
951:
704:Kodiak National Wildlife Refuge
588:
1884:Aho, Karen (6 November 1999).
1683:"Cub Adoption by Brown Bears,
1316:
1303:
1256:
1043:
933:U.S. Fish and Wildlife Service
858:environmental impact statement
634:Bears on Kodiak are naturally
483:mainland American Grizzly bear
430:
348:, inhabits the islands of the
13:
1:
2934:Kodiak Island Borough, Alaska
2004:Johnson, Ancel (1982-01-01).
1691:The Canadian Field-Naturalist
1036:
984:The bear is important to the
800:
629:
2909:Carnivorans of North America
2684:Ungava brown bear (formerly
1952:Penn, Brady (5 April 2022).
7:
2654:Sitka brown bear (formerly
1920:Kodiak Island and its bears
1687:, on Kodiak Island, Alaska"
1469:LaFee, Scott (2008-05-29).
782:National Geographic Society
751:
540:Kodiak bear's skull at the
435:
403:classifies the brown bear (
56:Alaska Peninsula brown bear
10:
2965:
2047:University of Alaska Press
1750:dfg.webmaster@alaska.gov.
1511:Illiger, J. K. W. (1811).
1263:Barnes, Victor G. (1994).
968:for a day of bear viewing.
679:
645:
53:
2939:Mammals described in 1896
2776:Ursus arctos middendorffi
2746:Ursus arctos middendorffi
2744:
2704:
2666:
2618:
2569:
2532:
2484:Gobi bear (provisionally
2458:
2409:
2355:
2307:Ursus arctos middendorffi
1985:Van Daele, L. J. (2003),
1685:Ursus arctos middendorffi
1074:Merriam, C. Hart (1896).
655:Reproduction and survival
501:Adult in a zoo in Germany
337:Ursus arctos middendorffi
323:
314:
300:Ursus arctos middendorffi
296:
289:
177:Scientific classification
175:
153:
144:
135:
126:
121:
1918:Dodge, Harry B. (2004).
1863:"Alaska's bears webpage"
1590:Van Daele, L.J. (2007).
1127:"Kodiak Bear Fact Sheet"
999:
772:Interactions with people
686:subpolar oceanic climate
594:Distribution and density
475:
423:however, along with the
31:latest accepted revision
2914:Endemic fauna of Alaska
2039:Into brown bear country
1835:Quinth, Stefan (2006).
1642:10.1023/A:1011019031766
1495:Biggest Bear Ever Found
1493:Dell'Amore, C. (2011):
1475:San Diego Union Tribune
1328:www.bearsoftheworld.net
960:A float plane based in
492:
385:California grizzly bear
381:Ursus arctos horribilis
282:U. a. middendorffi
2645:Kodiak bear (formerly
2215:. 2017. Archived from
969:
929:Endangered Species Act
900:
875:Exxon Valdez oil spill
848:Changes in land status
797:
789:History and management
764:
722:
664:
545:
531:Bismarck, North Dakota
502:
450:A. Th. von Middendorff
2628:brown bear (formerly
2266:. Alutiiqlanguage.org
2189:on September 23, 2006
2115:on September 22, 2006
1681:Barnes, V.G. (1993).
1599:Doctoral dissertation
1547:. In C. Brown (ed.).
1411:Wood, Gerald (1983).
980:Cultural significance
959:
894:
796:
759:
717:
663:Mother bear with cubs
662:
636:active during the day
539:
500:
360:or population of the
340:), also known as the
138:Kodiak Island, Alaska
2296:at Wikimedia Commons
1890:Anchorage Daily News
1843:on February 10, 2009
1622:Evolutionary Ecology
1348:"Mammalian Species:
1093:sp. nov. Kadiak Bear
487:Eurasian brown bears
2444:Eurasian brown bear
2430:Steppe brown bear (
2037:Troyer, W. (2005).
1756:www.adfg.alaska.gov
1731:on October 5, 2006.
1634:2000EvEco..14...39M
1499:National Geographic
814:Commercial harvests
147:Conservation status
87:editing the article
21:Page version status
2676:U. a. stikeenensis
2647:U. a. middendorffi
2598:U. a. californicus
2163:on 19 October 2011
1735:all plan documents
1397:on April 15, 2012.
1211:Gland, Switzerland
1186:2015-02-17 at the
1091:Ursus middendorffi
970:
925:endangered species
901:
798:
765:
723:
665:
600:Kodiak Archipelago
573:Boone and Crockett
546:
525:that lived at the
503:
446:Ursus middendorffi
389:U. a. californicus
383:) and the extinct
350:Kodiak Archipelago
346:Alaskan brown bear
27:
2891:
2890:
2738:Taxon identifiers
2729:
2728:
2700:
2699:
2696:
2695:
2565:
2564:
2561:
2560:
2495:U. a. isabellinus
2468:U. a. beringianus
2292:Media related to
2043:Fairbanks, Alaska
1991:Anchorage, Alaska
1426:978-0-85112-235-9
462:genetic diversity
342:Kodiak brown bear
328:
327:
170:
115:
114:
107:
80:layout guidelines
39:10 September 2024
18:
2956:
2884:
2883:
2871:
2870:
2858:
2857:
2845:
2844:
2832:
2831:
2819:
2818:
2806:
2805:
2793:
2792:
2780:
2779:
2778:
2765:
2764:
2763:
2733:
2732:
2720:
2713:
2686:U. a. ungavaesis
2616:
2615:
2585:U. a. horribilis
2576:
2575:
2551:U. a. pyrenaicus
2542:U. a. marsicanus
2456:
2455:
2440:
2439:
2338:
2331:
2324:
2315:
2314:
2304:Data related to
2303:
2291:
2275:
2274:
2272:
2271:
2258:
2252:
2251:
2249:
2248:
2242:
2234:
2228:
2227:
2225:
2224:
2205:
2199:
2198:
2196:
2194:
2179:
2173:
2172:
2170:
2168:
2153:
2147:
2146:
2144:
2142:
2131:
2125:
2124:
2122:
2120:
2105:
2096:
2095:
2067:
2061:
2060:
2034:
2028:
2027:
2026:. 11 March 2018.
2020:
2014:
2013:
2001:
1995:
1994:
1982:
1969:
1968:
1966:
1964:
1949:
1943:
1940:
1934:
1933:
1915:
1906:
1905:
1903:
1901:
1896:on July 23, 2011
1892:. Archived from
1881:
1875:
1874:
1872:
1870:
1859:
1853:
1852:
1850:
1848:
1832:
1826:
1825:
1793:
1787:
1786:
1784:
1782:
1772:
1766:
1765:
1763:
1762:
1747:
1738:
1732:
1730:
1723:
1715:
1709:
1708:
1706:
1704:10.5962/p.357155
1678:
1672:
1665:
1654:
1653:
1617:
1611:
1610:
1608:
1606:
1596:
1587:
1568:
1567:
1565:
1563:
1557:
1546:
1537:
1528:
1525:
1519:
1518:
1508:
1502:
1491:
1485:
1484:
1482:
1481:
1471:"Seeds of doubt"
1466:
1460:
1459:
1457:
1456:
1447:. Archived from
1437:
1431:
1430:
1418:
1408:
1399:
1398:
1393:. Archived from
1383:
1374:
1373:
1371:
1369:
1364:on March 4, 2016
1363:
1356:
1344:
1338:
1337:
1335:
1334:
1320:
1314:
1307:
1301:
1300:
1260:
1254:
1253:
1251:
1243:
1232:
1231:
1230:
1229:
1223:
1217:, archived from
1209:, vol. 44,
1208:
1197:
1191:
1178:
1173:
1171:
1165:
1158:
1150:
1141:
1140:
1138:
1137:
1131:
1123:
1096:
1095:
1071:
1062:
1061:
1059:
1057:
1047:
1030:
1019:
1013:
1010:
966:Raspberry Island
827:the population.
569:zygomatic arches
507:island gigantism
344:, sometimes the
319:
302:
185:
184:
164:
159:
158:
131:
119:
118:
110:
103:
99:
96:
90:
71:
70:
63:
2964:
2963:
2959:
2958:
2957:
2955:
2954:
2953:
2919:Fauna of Alaska
2894:
2893:
2892:
2887:
2879:
2876:Observation.org
2874:
2866:
2861:
2853:
2848:
2840:
2835:
2827:
2822:
2814:
2809:
2801:
2796:
2788:
2783:
2774:
2773:
2768:
2759:
2758:
2753:
2740:
2730:
2725:
2724:
2723:
2714:
2710:
2692:
2662:
2656:U. a. sitkensis
2614:
2583:
2581:
2557:
2528:
2522:U. a. pruinosus
2486:U. a. gobiensis
2447:
2445:
2438:
2422:U. a. crowtheri
2405:
2351:
2342:
2284:
2279:
2278:
2269:
2267:
2260:
2259:
2255:
2246:
2244:
2240:
2236:
2235:
2231:
2222:
2220:
2207:
2206:
2202:
2192:
2190:
2181:
2180:
2176:
2166:
2164:
2155:
2154:
2150:
2140:
2138:
2133:
2132:
2128:
2118:
2116:
2107:
2106:
2099:
2084:10.2307/3872907
2068:
2064:
2057:
2035:
2031:
2022:
2021:
2017:
2002:
1998:
1983:
1972:
1962:
1960:
1950:
1946:
1941:
1937:
1930:
1916:
1909:
1899:
1897:
1882:
1878:
1868:
1866:
1861:
1860:
1856:
1846:
1844:
1833:
1829:
1814:10.2307/3872753
1794:
1790:
1780:
1778:
1774:
1773:
1769:
1760:
1758:
1748:
1741:
1728:
1721:
1717:
1716:
1712:
1679:
1675:
1666:
1657:
1618:
1614:
1604:
1602:
1594:
1588:
1571:
1561:
1559:
1555:
1544:
1538:
1531:
1526:
1522:
1509:
1505:
1492:
1488:
1479:
1477:
1467:
1463:
1454:
1452:
1439:
1438:
1434:
1427:
1409:
1402:
1385:
1384:
1377:
1367:
1365:
1361:
1354:
1346:
1345:
1341:
1332:
1330:
1322:
1321:
1317:
1308:
1304:
1281:10.2307/3872685
1261:
1257:
1249:
1245:
1244:
1235:
1227:
1225:
1221:
1206:
1198:
1194:
1188:Wayback Machine
1169:
1167:
1166:on 11 July 2008
1163:
1156:
1152:
1151:
1144:
1135:
1133:
1129:
1125:
1124:
1099:
1072:
1065:
1055:
1053:
1049:
1048:
1044:
1039:
1034:
1033:
1020:
1016:
1011:
1007:
1002:
982:
954:
941:
939:Hunt-management
914:
889:
850:
833:
816:
803:
791:
774:
754:
712:
682:
657:
648:
632:
596:
591:
495:
478:
438:
433:
310:
304:
298:
285:
271:
179:
171:
160:
156:
149:
111:
100:
94:
91:
85:Please help by
84:
72:
68:
59:
52:
47:
46:
45:
44:
43:
42:
26:
12:
11:
5:
2962:
2952:
2951:
2946:
2941:
2936:
2931:
2926:
2921:
2916:
2911:
2906:
2889:
2888:
2886:
2885:
2872:
2859:
2846:
2833:
2820:
2807:
2794:
2781:
2766:
2750:
2748:
2742:
2741:
2727:
2726:
2722:
2721:
2707:
2706:
2705:
2702:
2701:
2698:
2697:
2694:
2693:
2691:
2690:
2680:
2670:
2668:
2664:
2663:
2661:
2660:
2651:
2642:
2633:
2622:
2620:
2613:
2612:
2602:
2591:
2589:
2582:(provisionally
2573:
2567:
2566:
2563:
2562:
2559:
2558:
2556:
2555:
2546:
2536:
2534:
2530:
2529:
2527:
2526:
2517:
2513:U. a. syriacus
2508:
2504:U. a. lasiotus
2499:
2490:
2481:
2477:U. a. collaris
2472:
2462:
2460:
2453:
2446:(provisionally
2437:
2436:
2426:
2415:
2413:
2407:
2406:
2404:
2403:
2395:
2387:
2381:
2375:
2369:
2363:
2356:
2353:
2352:
2350:or populations
2341:
2340:
2333:
2326:
2318:
2312:
2311:
2310:at Wikispecies
2297:
2283:
2282:External links
2280:
2277:
2276:
2253:
2229:
2213:Alutiiq Museum
2200:
2174:
2148:
2135:"Ursus arctos"
2126:
2097:
2062:
2055:
2029:
2015:
1996:
1970:
1944:
1935:
1928:
1907:
1876:
1854:
1827:
1788:
1767:
1739:
1710:
1697:(3): 365β367.
1673:
1655:
1612:
1569:
1529:
1520:
1503:
1486:
1461:
1441:"Kodiak Bears"
1432:
1425:
1400:
1375:
1339:
1315:
1302:
1255:
1233:
1192:
1142:
1097:
1063:
1041:
1040:
1038:
1035:
1032:
1031:
1027:marine mammals
1014:
1004:
1003:
1001:
998:
981:
978:
953:
950:
940:
937:
913:
910:
888:
885:
880:Shuyak Islands
867:Kizhuyak River
849:
846:
832:
829:
825:overharvesting
815:
812:
802:
799:
790:
787:
773:
770:
753:
750:
742:mountain goats
711:
710:Feeding habits
708:
700:Afognak Island
681:
678:
656:
653:
647:
644:
631:
628:
595:
592:
590:
587:
565:sagittal crest
494:
491:
477:
474:
437:
434:
432:
429:
411:" in terms of
326:
325:
321:
320:
312:
311:
305:
294:
293:
291:Trinomial name
287:
286:
279:
277:
273:
272:
268:U. arctos
265:
263:
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133:
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113:
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75:
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66:
50:
28:
22:
19:
17:
9:
6:
4:
3:
2:
2961:
2950:
2947:
2945:
2942:
2940:
2937:
2935:
2932:
2930:
2929:Grizzly bears
2927:
2925:
2922:
2920:
2917:
2915:
2912:
2910:
2907:
2905:
2902:
2901:
2899:
2882:
2877:
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2856:
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2689:
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2609:
2608:U. a. nelsoni
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2432:U. a. priscus
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1181:Overview page
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409:least concern
406:
405:Ursus arctos)
402:
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378:
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371:
370:grizzly bears
367:
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352:in southwest
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162:Least Concern
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95:February 2021
88:
82:
81:
76:This article
74:
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32:
25:
20:
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2584:
2580:Grizzly bear
2550:
2541:
2521:
2512:
2503:
2494:
2485:
2476:
2467:
2449:U. a. arctos
2448:
2431:
2421:
2420:Atlas bear (
2399:
2391:
2306:
2268:. Retrieved
2262:
2256:
2245:. Retrieved
2232:
2221:. Retrieved
2217:the original
2203:
2191:. Retrieved
2187:the original
2177:
2165:. Retrieved
2161:the original
2151:
2139:. Retrieved
2129:
2117:. Retrieved
2113:the original
2075:
2071:
2065:
2038:
2032:
2018:
2009:
1999:
1986:
1961:. Retrieved
1957:
1947:
1938:
1919:
1898:. Retrieved
1894:the original
1889:
1879:
1867:. Retrieved
1857:
1845:. Retrieved
1841:the original
1830:
1805:
1801:
1791:
1779:. Retrieved
1770:
1759:. Retrieved
1755:
1726:the original
1713:
1694:
1690:
1684:
1676:
1668:
1628:(1): 39β60.
1625:
1621:
1615:
1603:. Retrieved
1598:
1560:. Retrieved
1553:the original
1548:
1523:
1513:
1506:
1489:
1478:. Retrieved
1474:
1464:
1453:. Retrieved
1449:the original
1444:
1435:
1414:
1395:the original
1366:. Retrieved
1359:the original
1350:Ursus arctos
1349:
1342:
1331:. Retrieved
1327:
1318:
1305:
1272:
1268:
1258:
1226:, retrieved
1219:the original
1202:
1195:
1175:
1168:. Retrieved
1161:the original
1134:. Retrieved
1090:
1089:
1085:
1079:
1054:. Retrieved
1045:
1017:
1008:
993:
988:people. Its
983:
974:Frazer River
971:
952:Bear-viewing
946:
942:
915:
906:
902:
872:
855:
851:
838:
834:
817:
804:
779:
775:
766:
733:elderberries
724:
718:Bear with a
691:Sitka spruce
683:
674:
666:
649:
633:
597:
589:Life history
580:
578:
554:
547:
515:
504:
479:
454:Ursus arctos
453:
445:
442:C.H. Merriam
439:
413:endangerment
404:
397:conservation
393:
388:
380:
377:grizzly bear
374:
345:
341:
336:
335:
331:
329:
299:
297:
281:
280:
276:Subspecies:
266:
254:
122:Kodiak bear
116:
101:
92:
77:
60:
38:
29:This is the
23:
2824:iNaturalist
2770:Wikispecies
2630:U. a. dalli
2626:Dall Island
2294:Kodiak Bear
1963:31 December
1781:1 September
670:cannibalism
550:polar bears
440:Taxonomist
431:Description
332:Kodiak bear
2898:Categories
2717:polar bear
2638:U. a. gyas
2348:subspecies
2345:Brown bear
2270:2017-07-21
2247:2017-07-21
2223:2017-07-21
2078:: 93β103.
1761:2023-05-07
1480:2016-03-08
1455:2016-06-05
1333:2019-03-18
1228:2019-11-20
1136:2008-10-27
1056:August 29,
1037:References
801:Prehistory
630:Home range
624:Sitkalidak
527:Dakota Zoo
466:inbreeding
417:extinction
366:polar bear
362:brown bear
358:subspecies
136:A bear in
2924:Fur trade
2719:ancestry.
2667:In Canada
2659:; hybrid)
2619:In Alaska
2571:New World
2533:In Europe
2411:Old World
2397:Species:
2379:Carnivora
2359:Kingdom:
2263:Taquka'aq
1808:: 41β56.
1368:March 17,
1289:1936-0614
1275:: 63β73.
1088:: 69β71.
1023:Pinnipeds
1021:Treating
994:Taqukaβaq
640:nocturnal
616:Raspberry
470:parasites
458:Kamchatka
262:Species:
234:Carnivora
200:Kingdom:
194:Eukaryota
2855:14000982
2755:Wikidata
2383:Family:
2373:Mammalia
2367:Chordata
2365:Phylum:
2361:Animalia
2193:30 April
2167:22 March
2141:22 March
2119:30 April
1900:30 April
1869:30 April
1847:30 April
1605:30 April
1562:30 April
1445:bear.org
1184:Archived
1170:22 March
992:name is
921:Red List
860:for the
752:Behavior
561:calipers
523:specimen
436:Taxonomy
240:Family:
224:Mammalia
214:Chordata
210:Phylum:
204:Animalia
190:Domain:
167:IUCN 3.1
35:reviewed
2816:6163850
2803:1266086
2761:Q237260
2459:In Asia
2389:Genus:
2385:Ursidae
2377:Order:
2371:Class:
2092:3872907
1822:3872753
1650:1525994
1630:Bibcode
1297:3872685
990:Alutiiq
986:Alutiiq
931:of the
927:by the
841:Afognak
807:ice age
680:Habitat
646:Denning
608:Afognak
582:Grizzly
511:captive
307:Merriam
250:Genus:
244:Ursidae
230:Order:
220:Class:
165: (
2881:596363
2868:116961
2842:726993
2829:147090
2400:arctos
2209:"Bear"
2090:
2053:
1926:
1820:
1648:
1423:
1295:
1287:
962:Kodiak
897:Kodiak
728:salmon
720:salmon
696:Kodiak
620:Uganik
612:Shuyak
604:Kodiak
557:skulls
354:Alaska
309:, 1896
2790:5LPWZ
2392:Ursus
2241:(PDF)
2088:JSTOR
1818:JSTOR
1729:(PDF)
1722:(PDF)
1646:S2CID
1595:(PDF)
1556:(PDF)
1545:(PDF)
1362:(PDF)
1355:(PDF)
1293:JSTOR
1250:(PDF)
1222:(PDF)
1207:(PDF)
1164:(PDF)
1157:(PDF)
1130:(PDF)
1000:Notes
820:otter
476:Color
255:Ursus
140:, US
2863:NCBI
2837:ITIS
2811:GBIF
2195:2008
2169:2008
2143:2008
2121:2008
2051:ISBN
1965:2023
1924:ISBN
1902:2008
1871:2008
1849:2008
1783:2017
1607:2008
1564:2008
1421:ISBN
1370:2016
1285:ISSN
1215:IUCN
1172:2008
1058:2022
916:The
738:deer
518:wild
493:Size
485:and
401:IUCN
330:The
2850:MSW
2798:EoL
2785:CoL
2080:doi
1810:doi
1699:doi
1695:107
1638:doi
1277:doi
1025:as
746:elk
529:in
415:or
37:on
2900::
2878::
2865::
2852::
2839::
2826::
2813::
2800::
2787::
2772::
2757::
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2100:^
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2074:.
2049:.
2045::
2041:.
2008:.
1989:,
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1956:.
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1888:.
1816:.
1804:.
1800:.
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1100:^
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740:,
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2059:.
2012:.
1967:.
1932:.
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41:.
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