44:
66:
178:, which are distinct from satellite viruses in that they do not encode all of their own structural proteins but require proteins from their helper viruses in order to assemble. Additional common features include a rod-like structure, an RNA-binding "delta antigen" encoded in the genome, and animal hosts. Furthermore, the size range of the genomes of these viruses is between around 1547–1735nt, they encode a
649:
1093:
Paraskevopoulou S, Pirzer F, Goldmann N, Schmid J, Corman VM, Gottula LT, Schroeder S, Rasche A, Muth D, Drexler JF, Heni AC, Eibner GJ, Page RA, Jones TC, MĂĽller MA, Sommer S, Glebe D, Drosten C (July 2020).
1224:
1217:
1210:
186:, and their coding capacity only involves one conserved protein. Most lineages of this realm are poorly understood, the notable exception being the genus
157:
1883:
1780:
746:
Hepojoki J, Hetzel U, Paraskevopoulou S, Drosten C, Harrach B, Zerbini M, Koonin EV, Krupovic M, Dolja V, Kuhn JH (6 December 2020).
1347:
1713:
264:
was abolished and replaced with eight new species, and the taxonomy developed to reach its current form, detailed below.
1770:
1888:
1775:
1698:
183:
1686:
365:
17:
1042:
Wille M, Netter HJ, Littlejohn M, Yuen L, Shi M, Eden JS, Klaassen M, Holmes EC, Hurt AC (December 2018).
65:
1202:
662:
399:
1757:
1703:
1792:
913:
747:
626:
586:
1836:
1916:
1874:
993:
Hetzel U, Szirovicza L, Smura T, Prähauser B, Vapalahti O, Kipar A, Hepojoki J (April 2019).
1802:
1111:
800:
60:
8:
1377:
944:
Chang WS, Pettersson JH, Le Lay C, Shi M, Lo N, Wille M, Eden JS, Holmes EC (July 2019).
455:
281:, with no intermediate taxa between realm and family. This family contains eight genera.
214:
201:
has an unclear evolutionary origin. It has been proposed that they may have derived from
179:
145:
1115:
804:
1671:
1593:
1186:
1161:
1134:
1095:
1070:
1043:
1019:
994:
970:
945:
890:
855:
831:
788:
246:
remained unassigned to any higher taxa until 2018, when the ICTV mistakenly classified
1177:
1649:
1565:
1467:
1237:
1191:
1139:
1075:
1024:
975:
895:
877:
836:
818:
789:"Viroids and Viroid-like Circular RNAs: Do They Descend from Primordial Replicators?"
439:
337:
160:
TaxoProp 2020.012D, the realm is named after the presence of genomic and antigenomic
1911:
1859:
1181:
1173:
1129:
1119:
1065:
1055:
1014:
1006:
965:
957:
885:
867:
826:
808:
598:
289:
141:
1509:
1096:"Mammalian deltavirus without hepadnavirus coinfection in the neotropical rodent
719:
415:
393:
381:
325:
206:
43:
1558:
1523:
1516:
1495:
1474:
1451:
1331:
1104:
Proceedings of the
National Academy of Sciences of the United States of America
856:"Hepatitis Delta Virus (HDV) and Delta-Like Agents: Insights Into Their Origin"
654:
481:
466:
296:
872:
1905:
1797:
1725:
1618:
1586:
1572:
1551:
1537:
1488:
1481:
1400:
1273:
881:
822:
614:
609:
433:
427:
353:
1124:
1814:
1747:
1611:
1579:
1544:
1530:
1502:
1428:
1407:
1143:
1079:
1028:
979:
899:
840:
493:
175:
153:
1195:
1010:
1868:
1676:
813:
191:
961:
285:
1807:
1754:
1341:
1254:
581:
476:
376:
348:
166:
1232:
1060:
1414:
1393:
1295:
667:
320:
252:
202:
1830:
745:
1853:
1787:
1732:
1319:
1303:
1268:
1092:
699:. International Committee on Taxonomy of Viruses (ICTV). March 2021
450:
410:
161:
1819:
1737:
1718:
1278:
1263:
946:"Novel hepatitis D-like agents in vertebrates and invertebrates"
1765:
1708:
1693:
1643:
1373:
696:
672:
234:
of this realm to receive acceptance by the ICTV is the species
210:
992:
1742:
1369:
1313:
1308:
231:
149:
77:
648:
1681:
1285:
1233:
853:
995:"Identification of a Novel Deltavirus in Boa Constrictors"
1086:
1041:
765:. International Committee on Taxonomy of Viruses (ICTV)
854:
Netter HJ, Barrios MH, Littlejohn M, Yuen LK (2021).
644:
288:in various languages. The family name comes from
1903:
284:The names of all genera in this realm allude to
148:nucleic acids — infectious agents that resemble
1162:"Progress towards a higher taxonomy of viruses"
760:and seven new genera for a total of 15 species"
1218:
1044:"A Divergent Hepatitis D-Like Agent in Birds"
943:
939:
937:
935:
933:
260:returned to its original position. In 2020,
1781:Reverse transcriptase-related cellular gene
1225:
1211:
42:
1185:
1133:
1123:
1069:
1059:
1018:
986:
969:
930:
889:
871:
830:
812:
786:
250:within the then newly established realm,
225:
1035:
782:
780:
256:. In 2019, this error was rectified and
1762:Retroelements not elsewhere classified
14:
1904:
741:
739:
691:
689:
687:
1835:
1834:
1206:
777:
1159:
914:"Taxonomy History – Taxonomy – ICTV"
1714:Integrative and conjugative element
736:
724:University of California, Riverside
684:
24:
1153:
302:This taxonomy is shown hereafter:
190:, comprising the causal agents of
25:
1928:
1771:Diversity-generating retroelement
295:"triangle," another reference to
267:
209:) or a viroid-like element (i.e.
152:, but cannot replicate without a
1776:Telomerase reverse transcriptase
1348:Microbes with highly unusual DNA
787:Lee BD, Koonin EV (2022-01-12).
717:
647:
64:
217:) with capsid protein capture.
1699:Defective interfering particle
906:
847:
711:
697:"Virus Taxonomy: 2020 Release"
361:Dagazvirus schedorhinotermitis
184:hepatitis delta virus ribozyme
13:
1:
1687:Clonally transmissible cancer
1178:10.1016/S0923-2516(06)80059-2
678:
366:Schedorhinotermes intermedius
124:
752:) including one new family (
7:
1240:, and comparable structures
640:
220:
10:
1933:
275:contains a single family,
1843:
1662:
1636:
1603:
1459:
1445:
1438:
1385:
1368:
1361:
1329:
1294:
1252:
1245:
873:10.3389/fmicb.2021.652962
860:Frontiers in Microbiology
663:List of higher virus taxa
400:Liasis mackloti savuensis
115:
110:
59:
50:
41:
34:
1704:Endogenous viral element
720:"Subcellular Life Forms"
1125:10.1073/pnas.2006750117
1098:Proechimys semispinosus
748:"Create one new realm (
627:Proechimys semispinosus
587:early Cyrillic alphabet
489:Deltavirus cameroonense
463:Deevirus actinopterygii
546:Deltavirus senegalense
226:Historical development
1011:10.1128/mBio.00014-19
557:Deltavirus taiwanense
262:Hepatitis delta virus
236:Hepatitis delta virus
1803:Transposable element
1793:Spiegelman's Monster
1166:Research in Virology
814:10.3390/life12010103
524:Deltavirus japanense
513:Deltavirus italiense
297:the Greek letter "Δ"
170:type. The agents in
61:Virus classification
1116:2020PNAS..11717977P
1110:(30): 17977–17983.
918:talk.ictvonline.org
805:2022Life...12..103L
535:Deltavirus peruense
180:hammerhead ribozyme
1672:Bio-like structure
1594:Tolecusatellitidae
756:) including genus
622:Thurisazvirus myis
594:Dobrovirus bufonis
568:Deltavirus togense
502:Deltavirus carense
1899:
1898:
1837:Taxon identifiers
1828:
1827:
1658:
1657:
1632:
1631:
1628:
1627:
1566:Portogloboviridae
1468:Alphasatellitidae
1362:Non-cellular life
1357:
1356:
1238:non-cellular life
1061:10.3390/v10120720
962:10.1093/ve/vez021
338:Cynops orientalis
333:Daazvirus cynopis
156:. Established in
133:
132:
16:(Redirected from
1924:
1892:
1891:
1879:
1878:
1877:
1864:
1863:
1862:
1832:
1831:
1443:
1442:
1383:
1382:
1366:
1365:
1250:
1249:
1227:
1220:
1213:
1204:
1203:
1199:
1189:
1160:Ward CW (1993).
1148:
1147:
1137:
1127:
1090:
1084:
1083:
1073:
1063:
1039:
1033:
1032:
1022:
990:
984:
983:
973:
941:
928:
927:
925:
924:
910:
904:
903:
893:
875:
851:
845:
844:
834:
816:
784:
775:
774:
772:
770:
764:
743:
734:
733:
731:
730:
715:
709:
708:
706:
704:
693:
657:
652:
651:
599:Bufo gargarizans
456:English alphabet
207:retrotransposons
69:
68:
46:
32:
31:
27:Realm of viruses
21:
1932:
1931:
1927:
1926:
1925:
1923:
1922:
1921:
1902:
1901:
1900:
1895:
1887:
1882:
1873:
1872:
1867:
1858:
1857:
1852:
1839:
1829:
1824:
1664:
1654:
1624:
1599:
1510:Finnlakeviridae
1455:
1434:
1376:
1372:
1353:
1325:
1290:
1241:
1231:
1156:
1154:Further reading
1151:
1091:
1087:
1040:
1036:
991:
987:
950:Virus Evolution
942:
931:
922:
920:
912:
911:
907:
852:
848:
785:
778:
768:
766:
762:
744:
737:
728:
726:
716:
712:
702:
700:
695:
694:
685:
681:
653:
646:
643:
465:(host: various
440:A. superciliosa
423:Dalvirus anatis
416:Arabic alphabet
394:Boa constrictor
389:Daletvirus boae
382:Hebrew alphabet
326:Gothic alphabet
270:
228:
223:
129:
123:
122:
120:
118:
63:
28:
23:
22:
15:
12:
11:
5:
1930:
1920:
1919:
1914:
1897:
1896:
1894:
1893:
1880:
1865:
1849:
1847:
1841:
1840:
1826:
1825:
1823:
1822:
1817:
1812:
1811:
1810:
1800:
1795:
1790:
1785:
1784:
1783:
1778:
1773:
1768:
1760:
1752:
1751:
1750:
1740:
1735:
1730:
1721:
1716:
1711:
1706:
1701:
1696:
1691:
1690:
1689:
1684:
1674:
1668:
1666:
1660:
1659:
1656:
1655:
1653:
1652:
1647:
1640:
1638:
1634:
1633:
1630:
1629:
1626:
1625:
1623:
1622:
1615:
1607:
1605:
1601:
1600:
1598:
1597:
1590:
1583:
1576:
1569:
1562:
1559:Polydnaviridae
1555:
1548:
1541:
1534:
1527:
1524:Globuloviridae
1520:
1517:Fuselloviridae
1513:
1506:
1499:
1496:Bicaudaviridae
1492:
1485:
1478:
1475:Ampullaviridae
1471:
1463:
1461:
1457:
1456:
1452:Naldaviricetes
1449:
1447:
1440:
1436:
1435:
1433:
1432:
1425:
1418:
1411:
1404:
1397:
1389:
1387:
1380:
1363:
1359:
1358:
1355:
1354:
1352:
1351:
1345:
1337:
1335:
1332:Incertae sedis
1327:
1326:
1324:
1323:
1316:
1311:
1306:
1300:
1298:
1292:
1291:
1289:
1288:
1283:
1282:
1281:
1276:
1266:
1260:
1258:
1247:
1243:
1242:
1230:
1229:
1222:
1215:
1207:
1201:
1200:
1155:
1152:
1150:
1149:
1085:
1034:
985:
929:
905:
846:
776:
735:
710:
682:
680:
677:
676:
675:
670:
665:
659:
658:
655:Viruses portal
642:
639:
638:
637:
636:
635:
634:
633:
632:
631:
605:
604:
603:
577:
576:
575:
564:
553:
542:
531:
520:
509:
498:
482:Greek alphabet
472:
471:
470:
467:Actinopterygii
446:
445:
444:
406:
405:
404:
372:
371:
370:
344:
343:
342:
269:
268:Current status
266:
238:and its genus
227:
224:
222:
219:
174:are satellite
131:
130:
116:
113:
112:
108:
107:
100:
96:
95:
88:
81:
80:
75:
71:
70:
57:
56:
48:
47:
39:
38:
26:
9:
6:
4:
3:
2:
1929:
1918:
1915:
1913:
1910:
1909:
1907:
1890:
1885:
1881:
1876:
1870:
1866:
1861:
1855:
1851:
1850:
1848:
1846:
1842:
1838:
1833:
1821:
1818:
1816:
1813:
1809:
1806:
1805:
1804:
1801:
1799:
1798:Tandem repeat
1796:
1794:
1791:
1789:
1786:
1782:
1779:
1777:
1774:
1772:
1769:
1767:
1764:
1763:
1761:
1759:
1756:
1753:
1749:
1746:
1745:
1744:
1741:
1739:
1736:
1734:
1731:
1728:
1727:
1726:Nanobacterium
1722:
1720:
1717:
1715:
1712:
1710:
1707:
1705:
1702:
1700:
1697:
1695:
1692:
1688:
1685:
1683:
1680:
1679:
1678:
1675:
1673:
1670:
1669:
1667:
1661:
1651:
1648:
1645:
1642:
1641:
1639:
1635:
1621:
1620:
1619:Rhizidiovirus
1616:
1614:
1613:
1609:
1608:
1606:
1602:
1596:
1595:
1591:
1589:
1588:
1587:Thaspiviridae
1584:
1582:
1581:
1577:
1575:
1574:
1573:Pospiviroidae
1570:
1568:
1567:
1563:
1561:
1560:
1556:
1554:
1553:
1552:Plasmaviridae
1549:
1547:
1546:
1542:
1540:
1539:
1538:Halspiviridae
1535:
1533:
1532:
1528:
1526:
1525:
1521:
1519:
1518:
1514:
1512:
1511:
1507:
1505:
1504:
1500:
1498:
1497:
1493:
1491:
1490:
1489:Avsunviroidae
1486:
1484:
1483:
1482:Anelloviridae
1479:
1477:
1476:
1472:
1470:
1469:
1465:
1464:
1462:
1458:
1454:
1453:
1448:
1444:
1441:
1437:
1431:
1430:
1426:
1424:
1423:
1419:
1417:
1416:
1412:
1410:
1409:
1405:
1403:
1402:
1401:Duplodnaviria
1398:
1396:
1395:
1391:
1390:
1388:
1384:
1381:
1379:
1375:
1371:
1367:
1364:
1360:
1349:
1346:
1344:
1343:
1339:
1338:
1336:
1334:
1333:
1328:
1321:
1317:
1315:
1312:
1310:
1307:
1305:
1302:
1301:
1299:
1297:
1293:
1287:
1284:
1280:
1277:
1275:
1274:Mitochondrion
1272:
1271:
1270:
1267:
1265:
1262:
1261:
1259:
1256:
1251:
1248:
1246:Cellular life
1244:
1239:
1235:
1228:
1223:
1221:
1216:
1214:
1209:
1208:
1205:
1197:
1193:
1188:
1183:
1179:
1175:
1172:(6): 419–53.
1171:
1167:
1163:
1158:
1157:
1145:
1141:
1136:
1131:
1126:
1121:
1117:
1113:
1109:
1105:
1101:
1099:
1089:
1081:
1077:
1072:
1067:
1062:
1057:
1053:
1049:
1045:
1038:
1030:
1026:
1021:
1016:
1012:
1008:
1004:
1000:
996:
989:
981:
977:
972:
967:
963:
959:
956:(2): vez021.
955:
951:
947:
940:
938:
936:
934:
919:
915:
909:
901:
897:
892:
887:
883:
879:
874:
869:
865:
861:
857:
850:
842:
838:
833:
828:
824:
820:
815:
810:
806:
802:
798:
794:
790:
783:
781:
761:
759:
755:
754:Kolmioviridae
751:
742:
740:
725:
721:
714:
698:
692:
690:
688:
683:
674:
671:
669:
666:
664:
661:
660:
656:
650:
645:
629:
628:
623:
619:
618:
616:
615:Elder Futhark
612:
611:
610:Thurisazvirus
606:
601:
600:
595:
591:
590:
588:
584:
583:
578:
573:
569:
565:
562:
558:
554:
551:
547:
543:
540:
536:
532:
529:
525:
521:
518:
514:
510:
507:
503:
499:
496:
495:
490:
486:
485:
483:
479:
478:
473:
468:
464:
460:
459:
457:
453:
452:
447:
442:
441:
436:
435:
430:
429:
428:Anas gracilis
424:
420:
419:
417:
413:
412:
407:
402:
401:
396:
395:
390:
386:
385:
383:
379:
378:
373:
368:
367:
362:
358:
357:
355:
354:Elder Futhark
351:
350:
345:
340:
339:
334:
330:
329:
327:
323:
322:
317:
316:
315:
314:Kolmioviridae
311:
310:
309:
305:
304:
303:
300:
298:
294:
291:
287:
282:
280:
279:
278:Kolmioviridae
274:
265:
263:
259:
255:
254:
249:
245:
241:
237:
233:
218:
216:
212:
208:
205:(a family of
204:
200:
195:
193:
189:
185:
181:
177:
176:nucleic acids
173:
169:
168:
163:
159:
155:
151:
147:
143:
139:
138:
128:
127:
114:
109:
106:
105:
104:Kolmioviridae
101:
98:
97:
94:
93:
89:
86:
83:
82:
79:
76:
73:
72:
67:
62:
58:
54:
51:Diagram of a
49:
45:
40:
37:
36:Kolmioviridae
33:
30:
19:
18:Kolmioviridae
1917:Virus realms
1844:
1815:Transpoviron
1748:Fungal prion
1724:
1617:
1612:Dinodnavirus
1610:
1592:
1585:
1580:Spiraviridae
1578:
1571:
1564:
1557:
1550:
1545:Ovaliviridae
1543:
1536:
1531:Guttaviridae
1529:
1522:
1515:
1508:
1503:Clavaviridae
1501:
1494:
1487:
1480:
1473:
1466:
1450:
1429:Varidnaviria
1427:
1421:
1420:
1413:
1408:Monodnaviria
1406:
1399:
1392:
1340:
1330:
1169:
1165:
1107:
1103:
1097:
1088:
1051:
1047:
1037:
1002:
998:
988:
953:
949:
921:. Retrieved
917:
908:
863:
859:
849:
796:
792:
767:. Retrieved
757:
753:
749:
727:. Retrieved
723:
713:
701:. Retrieved
625:
621:
608:
597:
593:
580:
571:
567:
560:
556:
549:
545:
538:
534:
527:
523:
516:
512:
505:
501:
494:Homo sapiens
492:
488:
475:
462:
449:
438:
432:
426:
422:
409:
398:
392:
388:
375:
364:
360:
347:
336:
332:
319:
313:
307:
301:
292:
286:the letter D
283:
277:
276:
272:
271:
261:
257:
251:
247:
243:
239:
235:
229:
198:
196:
187:
171:
165:
154:helper virus
136:
135:
134:
125:
103:
102:
91:
90:
84:
74:(unranked):
52:
35:
29:
1875:Ribozyviria
1869:Wikispecies
1845:Ribozyviria
1758:microsphere
1677:Cancer cell
1422:Ribozyviria
1054:(12): 720.
750:Ribozyviria
434:A. castanea
308:Ribozyviria
273:Ribozyviria
242:, in 1993.
199:Ribozyviria
194:in humans.
192:hepatitis D
172:Ribozyviria
137:Ribozyviria
92:Ribozyviria
1906:Categories
1860:Q106951424
1808:Retroposon
1755:Proteinoid
1665:structures
1663:Comparable
1439:Unassigned
1342:Parakaryon
1255:Prokaryota
923:2021-10-08
799:(1): 103.
758:Deltavirus
729:2020-06-04
679:References
613:(from the
585:(from the
582:Dobrovirus
572:H. sapiens
561:H. sapiens
550:H. sapiens
539:H. sapiens
528:H. sapiens
517:H. sapiens
506:H. sapiens
480:(from the
477:Deltavirus
454:(from the
414:(from the
380:(from the
377:Daletvirus
352:(from the
349:Dagazvirus
324:(from the
258:Deltavirus
248:Deltavirus
244:Deltavirus
240:Deltavirus
230:The first
215:satellites
203:retrozymes
197:The realm
188:Deltavirus
167:Deltavirus
121:15 species
53:Deltavirus
1415:Riboviria
1394:Adnaviria
1378:Satellite
1296:Eukaryota
882:1664-302X
823:2075-1729
668:Retrozyme
620:Species:
592:Species:
566:Species:
555:Species:
544:Species:
533:Species:
522:Species:
511:Species:
500:Species:
487:Species:
461:Species:
421:Species:
387:Species:
359:Species:
331:Species:
321:Daazvirus
299:(delta).
253:Riboviria
162:ribozymes
146:satellite
1854:Wikidata
1788:Ribozyme
1733:Phagemid
1460:Families
1320:Protista
1304:Animalia
1269:Bacteria
1144:32651267
1080:30562970
1029:30940697
980:31321078
900:34234753
866:: 1448.
841:35054497
718:Baez J.
641:See also
451:Deevirus
411:Dalvirus
312:Family:
221:Taxonomy
126:See text
119:8 genera
117:1 family
111:Subtaxa
99:Family:
1912:Viruses
1889:2842242
1820:Xenobot
1738:Plasmid
1719:Jeewanu
1650:Obelisk
1446:Classes
1314:Plantae
1279:Plastid
1264:Archaea
1196:8140287
1187:7135741
1135:7395443
1112:Bibcode
1071:6315422
1048:Viruses
1020:6445931
971:6628682
891:8256844
832:8781251
801:Bibcode
624:(host:
607:Genus:
596:(host:
579:Genus:
570:(host:
559:(host:
548:(host:
537:(host:
526:(host:
515:(host:
504:(host:
491:(host:
474:Genus:
448:Genus:
425:(host:
408:Genus:
391:(host:
374:Genus:
363:(host:
346:Genus:
335:(host:
318:Genus:
306:Realm:
290:Finnish
211:viroids
164:of the
150:viruses
55:virion
1766:Retron
1709:Fosmid
1694:Cosmid
1644:Nanobe
1604:Genera
1386:Realms
1374:Viroid
1194:
1184:
1142:
1132:
1078:
1068:
1027:
1017:
978:
968:
898:
888:
880:
839:
829:
821:
769:27 May
763:(docx)
703:10 May
673:Viroid
293:kolmio
1743:Prion
1637:Other
1370:Virus
1309:Fungi
1005:(2).
469:spp.)
182:or a
142:realm
140:is a
85:Realm
78:Virus
1884:NCBI
1682:HeLa
1286:LUCA
1234:Life
1192:PMID
1140:PMID
1076:PMID
1025:PMID
999:mBio
976:PMID
896:PMID
878:ISSN
837:PMID
819:ISSN
793:Life
771:2021
705:2021
232:taxa
213:and
158:ICTV
1646:(?)
1350:(?)
1182:PMC
1174:doi
1170:144
1130:PMC
1120:doi
1108:117
1066:PMC
1056:doi
1015:PMC
1007:doi
966:PMC
958:doi
886:PMC
868:doi
827:PMC
809:doi
144:of
1908::
1886::
1871::
1856::
1236:,
1190:.
1180:.
1168:.
1164:.
1138:.
1128:.
1118:.
1106:.
1102:.
1074:.
1064:.
1052:10
1050:.
1046:.
1023:.
1013:.
1003:10
1001:.
997:.
974:.
964:.
952:.
948:.
932:^
916:.
894:.
884:.
876:.
864:12
862:.
858:.
835:.
825:.
817:.
807:.
797:12
795:.
791:.
779:^
738:^
722:.
686:^
617:)
589:)
484:)
458:)
437:,
431:,
418:)
397:,
384:)
356:)
328:)
87::
1729:"
1723:"
1322:'
1318:'
1257:"
1253:"
1226:e
1219:t
1212:v
1198:.
1176::
1146:.
1122::
1114::
1100:"
1082:.
1058::
1031:.
1009::
982:.
960::
954:5
926:.
902:.
870::
843:.
811::
803::
773:.
732:.
707:.
630:)
602:)
574:)
563:)
552:)
541:)
530:)
519:)
508:)
497:)
443:)
403:)
369:)
341:)
20:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.