1035:-based mating system. Mating solicitations are made by both males and females, half of the time for each. To solicit mating, the mantled guereza will walk near its potential partner and make low-intensity mouth clicks or tough-smacks. During copulation, the males hold on the female's ankles and body. Most matings take place between individuals of the same group but copulations outside of the group have been recorded. In multi-male groups, more than one male may mate with the females. The gestation period lasts 158 days with a 16–22 month interbirth interval. The newborn guereza relies on its mother for support and must cling to her. As they grow older, infants can move on their own but keep returning to their mothers. The infants take up most of the attention in the groups. The other females in a group may handle an infant although the latter are only comfortable with their mothers. The males normally do not pay much attention to infants until they are four to five weeks old. Infants can eat solid food at about eight to nine weeks and by fifty weeks they are fully weaned and no longer need to hold on to their mothers.
1044:
communication and can regulate inter-group spacing without direct, physical contact while foraging. When one male starts roaring, neighboring males will start to roar as well. Often, the mantled guereza will respond to calls regardless of "caller identity," focusing more on the collective vocal displays and not the familiarity of the caller. There is variation in the roars of males which could signal the status of their group and fighting ability. With a roar, a male can advertise his body size; both actual and exaggerated. Other vocalizations are made as well. Males may snort, possibly as an alarm call. "Purrs" are made before group movements. Females and infants may "caw" when under mild distress. When in more serious distress, like if an infant is in danger, females and sub-adults will squeak or scream. "Tongue-clicking" is made during mild aggression. In addition to vocalizations, the mantled guereza communicates with several different body postures and movements, displaying of fringe fur, facial expressions, and touches.
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921:. While it mainly eats leaves and fruit, its diet is quite variable. It may eat bark, wood, seeds, flowers, petioles, lianas, aquatic-plants, arthropods, soil, and even concrete from buildings. The amount of each food item in its diet varies by area and time of year. Nutritional factors like protein, tannins, and sodium levels in leaves influence its food choices. It may even intermittently travel longer distances to access plants with higher levels of nutrition. Leaves usually make up over half of its diet, although fruits are occasionally eaten more depending on the season. When foraging for leaves, the mantled guereza prefers young ones over old. With fleshy fruits, the mantled guereza prefers to eat them unripe, which may serve to reduce competition with primates that eat ripe fruits. It consumes a number of plant species but only some make up most of its diet at a specific site.
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818:, which is mostly black, with long white fringes of silky hair—known as a mantle or ornamentation—along the sides of its body and tail. The bands that make up the mantle start at the shoulders and extend along the back until they connect at the lower torso. The tail is long and ends in a white tuft which varies in how much it covers the tail. These features vary in color among subspecies, for example the tail of
996:
992:), resources, and mates. During intergroup encounters, males can engage in direct or indirect mate defense, like defending a female's resources. It is the males that participate in agonistic inter-group encounters but female may do so as well. Aggressive encounters between groups usually involve chases, displays and vocalizations rather than physical contact.
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social structure. Female guerezas living in a group often have an egalitarian dominance style with no formalized rank relations. Relationships are relaxed and friendly with rare signals of dominance or subordinance. Physical aggression within the group is usually not harmful and rarely escalates into
905:
and rests for up to half the day. Foraging or travelling are the next most common activity. Sometime after dawn, mantled guereza groups leave their sleeping trees and will return to them at dusk. During the day, the mantled guereza has long rest periods in between periods of moving and feeding. Other
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related. They rarely disperse from their natal groups, except possibly when they break apart. Males on the other hand, usually leave when they become subadults or adults. They may start out being solitary and or in bachelor groups. They gain entry into a social group either by being on the periphery
829:
Infants are born with pink skin and white hair. The hair and skin darken as they age and by three to four months they attain adult coloration. Male usually gain their coloration before females. The male typically weighs 9.3 and 13.5 kilograms (21 and 30 lb) and the female weighs between 7.8 and
969:
The mantled guereza lives in stable social groups usually containing three to fifteen members. The groups usually contain one male, several females and juveniles. In some populations, groups containing several males are common. In multi-male groups, males tend to be aggressive with one another with
466:
from Asia and Africa. This subfamily is split into three groups, the colobus monkeys of Africa, of which the mantled guereza is a part, the langurs, or leaf monkeys, of Asia, and an "odd-nosed" group. The
African colobus monkeys are divided again by distinctions in color, behavior, and ecology. The
410:
The mantled guereza has many alternative common names including the guereza, the eastern black-and-white colobus, the magistrate colobus, or the
Abyssinian black-and-white colobus. The name "mantled" refers to its mantle, the long silky white fringes of hair that run along its body and "guereza" is
378:
of just over five months, infants are born with pink skin and white fur, which darkens to the adult coloration by three to four months. The mantled guereza is well known for its dawn chorus, the males' "roar" is a method of long-distance communication that reinforces territorial boundaries. It also
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The
Dodinga Hills guereza has not been recorded since the 1960s. Unlike most other primate species, the mantled guereza can survive habitat degradation and can even thrive in degraded forests. Sometimes, logging may increase the number of preferred food trees for the mantled guereza and it is more
987:
is an important part of mantled guereza interactions and mostly occurs between females. The adult males rarely groom in the groups. While not strictly territorial, mantled guereza groups can be aggressive towards each other. In some populations, groups may defend core areas (which exist as a small
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abundant in logged areas than unlogged ones. However, complete forest clearance causes dramatic declines in numbers. In Uganda complete forest clearings caused a decline of 50% over eight years. The mantled guereza is also threatened by hunting for meat and its skin. Mantled guereza meat sells as
924:
Like all colobi, the mantled guereza is able to digest leaves and other plant fibers with a large, multi-chambered stomach that contains bacteria in certain areas. Like most colobines, it prefers foods with high fiber content that can be easily extracted with its specialized stomach. The mantled
1043:
The most notable vocalization of the mantled guereza is the "roar", which is made mainly at night or dawn by males. The sound of a roar can be carried for up to a mile. It is normally the dominant male who roars when there are multiple males in the group. Roars are used for long-distance
970:
one being dominant. Some males may be expelled from these groups. Multi-male groups may contain father-son pairs or unrelated males. Males that are not part of groups either live solitarily or with other outside males in bachelor groups. The females keep the groups cohesive and they are
1068:
because "although locally threatened in parts of its range, this widespread species is not thought to be declining fast enough to place it in a higher category of threat." However, some of the subspecies are classified under different categories. The Mt
Uaraguess guereza,
882:
if given the choice. It is likely that the mantled guereza prefers these forests due to the increased number of food trees and the weaker chemical defenses of the species within. The mantled guereza is sometimes found in swamps as well as human-made habitats such as
998:
862:. It ranges from Nigeria and Cameroon in the west to Ethiopia, Kenya, Uganda, and northern Tanzania in the east. The mantled guereza lives in both deciduous and evergreen forests. It mainly inhabits forest and savannah woodlands and often extend into highland and
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9.2 kilograms (17 and 20 lb). The head and body length averages 61.5 centimetres (24.2 in) for males and 57.6 centimetres (22.7 in) for females. Like most colobi, the mantled guereza has a small thumb that is
669:
390:(IUCN) because it is widespread – although it is locally threatened in some areas, the decline is not great enough to list it in a higher category of threat. However, one subspecies found in Kenya is listed as
767:
320:. The species consists of several subspecies that differ in appearance. It has a distinctive appearance, which is alluded to in its name; the long white fringes of hair that run along each side of its black
874:, and upland forests. It is particularly common in forests close to rivers and lakes and at high elevations. It can be found in elevations as high as 3,300 metres (10,800 ft). This species prefers
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Harris, T. R.; Caillaud, D.; Chapman, C. A.; Vigilant, L. (2009). "Neither genetic nor observational data alone are sufficient for understanding sex-biased dispersal in a social-group-living species".
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The mantled guereza lives in social groups of three to fifteen individuals. These groups normally include a dominant male, several females, and the offspring of the females. It has a
646:
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Eves, H. E.; Ruggiero, R. G. (2000). "Socioeconomics and the sustainability of hunting in the forests of northern Congo (Brazzaville)". In
Robinson, J. G.; Bennett, E. L. (eds.).
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makes up 80% of the tail. The mantle color ranges from white to cream or yellow. Its face is framed by white hair and it has bushy cheek hairs. The thigh has a white stripe.
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351:. It is able to digest plant material with a high fibre content with its specialised stomach and may only eat from a few plant species at a time. It is preyed on by
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Because it can live in both dry and gallery forests and move on the ground, the mantled guereza is less threatened than many other colobine species. The
1509:
Catalogue of
Primates in the British Museum (Natural History) and Elsewhere in the British Isles, part III: Family Cercopithecidae, Subfamily Colobinae
2384:
4483:
4313:
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Lwanga, J. S. (2006). "Spatial distribution of primates in a mosaic of colonizing and old growth forest at Ngogo, Kibale
National Park, Uganda".
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among the subspecies. In some, the males have larger teeth than females, in others the reverse is true, and some have no significant difference.
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and in some areas it is more common in logged areas than unlogged ones. The mantled guereza is also threatened by hunting for
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4370:
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1325:
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4331:
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Hayes, V. J.; Freedman, L.; Oxnard, C. E. (1995). "The differential expression of dental sexual dimorphism in subspecies of
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527:, occurs from eastern Nigeria, Cameroon, and Gabon at the edge of its western range to South Sudan and Uganda, west of the
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2009:
Struhsaker, T. T.; Leakey, M. (1990). "Prey selectivity by crowned hawk-eagles on monkeys in the Kibale Forest, Uganda".
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Harris, T. R.; Monfort, S. L. (2006). "Mating behavior and endocrine profiles of wild black and white colobus monkeys (
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3000:
2810:"Long-term effects of logging on African primate communities: A 28-year comparison from Kibale National Park, Uganda"
1139:
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Harris, T. R. (2005). Roaring, intergroup aggression, and feeding competition in black and white colobus monkeys (
901:, but does sometimes descend on the ground to forage and travel, perhaps more so than most other colobines. It is
4528:
1814:
von Hippel, F. A. (1996). "Interactions between overlapping multimale groups of black and white colobus monkeys (
632:
The morphological difference between subspecies is most pronounced between the southeastern
Kilimanjaro guereza,
2840:
2526:
2253:
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Because of its low quality diet and the dispersed distribution of its food, the mantled guereza has a resident-
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or allogrooming mainly occurs between females and is an important social interaction in mantled guereza groups.
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2954:
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4513:
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4227:
2419:
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Dunbar, R. I. M. (1987). "Habitat quality, population dynamics, and group composition in colobus monkeys (
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close to rivers or lakes. Although previously thought to eat only leaves, it also eats seeds, fruits, and
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activities, including grooming, greeting, playing and being vigilant, are performed to a lesser extent.
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Dunbar, R. I. M. (1976). "Contrasts in social structure among black-and-white colobus monkey groups".
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makes other vocalization and uses body postures, movements, and facial expressions to communicate.
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1886:"Variation in diet and ranging of black and white colobus monkeys in Kibale National Park, Uganda"
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Chapman, C. A.; Naughton-Treves, L.; Lawes, M. J.; Wasserman, M. D.; Gillespie, T. R. (2007).
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are known as a mantle. Its face is framed with white hair and it has a large white tail tuft.
4404:
3913:
3805:
3360:
2866:"Population declines of Colobus in western Uganda and conservation value of forest fragments"
2258:): Evidence for resource defense mediated through males and a comparison with other primates"
1341:
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Infants are born with white fur and are always carried during the first months of their life.
846:
214:
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Chapman, C. A.; Balcomb, S. R.; Gillespie, T. R.; Skorupa, J. P.; Struhsaker, T. T. (2000).
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1953:"Influence of plant and soil chemistry on food selection, ranging patterns, and biomass of
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Despite its reputation as an exclusive leaf-eater, the mantled guereza is not an obligate
913:
The diet of the mantled guereza is predominantly leaves, often of only a few tree species.
822:
is gray until the white tail tuft which covers half of its length, while the tail tuft of
8:
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Ecology: Studies of Feeding and Ranging Behaviour in Lemurs, Monkeys and Apes
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2046:"Hunting attempt by chimpanzees on Abyssinian colobus at the Kalinzu Forest, Uganda"
2030:
1983:
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due to its small range and its risk from hunting, while the Dodinga Hills guereza,
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plantations, which may be frequented when the monkey has nutritional deficiencies.
875:
851:
803:
511:, is uncertain, although not listed by Groves in MSW, it is recognised in his 2007
463:
432:
395:
375:
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181:
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866:. It can be found in other forest habitats, both primary and secondary, such as
343:. It is an adaptable species that can cope with habitat disturbance and prefers
3940:
3850:
3814:
3737:
3566:
3532:
3517:
3202:
3184:
2531:) roars as a source of both honest and exaggerated information about body mass"
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492:
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2503:
1975:
1904:
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56:
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577:
476:
383:
84:
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2370:) at Kanyawara, Kibale National Park, Uganda. Ph.D. thesis. Yale University.
1642:
Dunbar, R. I. M.; Dunbar, E. P. (1974). "Ecology and population dynamics of
1149:
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for $ 4–9 US. The skins have been sold for fashion or in the tourist trade.
3877:
3193:
2448:
Oates, J. F. (1977). "The social life of a black-and-white colobus monkey,
2323:
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Oates, J. F. (1978). "Water-plant and soil consumption by guereza monkeys (
1707:
1463:
1422:
1121:
1052:
984:
930:
831:
500:
484:
352:
2276:
2079:"The alarm call system of two species of black-and-white colobus monkeys (
1667:
909:
4396:
4300:
4194:
3647:
3226:
2473:
2347:
Guereza dietary and behavioural patterns at the Entebbe Botanical Gardens
2148:
979:
606:
551:
472:
1343:
A Catalogue of the Mammalia in the Museum of the Hon. East-India Company
1134:(3rd ed.). Baltimore: Johns Hopkins University Press. p. 168.
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2230:
2022:
1757:
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989:
902:
885:
625:
391:
328:
161:
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Harris, T. R.; Fitch, W. T.; Goldstein, L. M.; Fashing, P. J. (2006).
2315:
2254:"Male and female strategies during intergroup encounters in guerezas (
1832:
10.1002/(SICI)1098-2345(1996)38:3<193::AID-AJP1>3.0.CO;2-U
1659:
1169:
450:
4305:
4274:
3770:
3103:
2302:): Toward an understanding of their life history and mating system".
2213:
Grunau, T.; Kuester, J. (2001). "Dominance style in female guerezas (
2105:
807:
640:. The intermediate subspecies show a gradual change between the two.
559:
512:
459:
348:
336:
111:
4156:
2863:
1749:
587:, occurs from western Kenya and Uganda south into northern Tanzania.
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2062:
2045:
1098:
918:
898:
742:
399:
309:
301:
289:
131:
1990:
30:
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3134:
1131:
Mammal Species of the World: A Taxonomic and Geographic Reference
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360:
297:
151:
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3128:
3116:
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Primates Of The World: Distribution, Abundance And Conservation
1222:
University of Michigan Museum of Zoology's Animal Diversity Web
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661:
565:
Djaffa Mountains guereza or Neumann's black-and-white colobus,
479:. There are three black-and-white colobi: the mantled guereza,
411:
the native name of the monkey in Ethiopia. The scientific name
313:
285:
171:
141:
121:
2807:
1165:
1163:
1161:
1159:
4417:
2169:
1946:
1944:
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subfamily, also known as the leaf-eating monkeys, a group of
305:
2588:
2586:
590:
Mt Uaraguess guereza or Percival's black-and-white colobus,
3975:
2757:
2657:
2524:
1156:
1061:
569:, found in the Ethiopian Highlands east of the Rift Valley.
528:
507:(MSW) (2005). The validity of the Djaffa Mountain guereza,
317:
3030:
Conclusions: the past, present and future of the colobines
2076:
1941:
1736:): a relationship with minerals and toxins in the diet?".
1524:
1522:
1520:
1518:
3057:
Images and movies of the Eastern black-and-white colobus
2583:
620:, found in Tanzania and Kenya in the forests surrounding
534:
Omo River guereza or Abyssinian black-and-white colobus,
443:
when three individuals were purchased from a dealer from
2993:
Colobine Monkeys: Their Ecology, Behaviour and Evolution
2707:
1951:
Fashing, P. J.; Dierenfeld, E. S.; Mowry, C. B. (2007).
1573:
1481:
Ackerman, D. E. (1991). "A study of the colobus monkey (
1301:
419:
meaning "mutilated" which refers to its lack of thumbs.
2485:
2483:
1534:
1515:
431:, a German naturalist and explorer, during his trip to
2077:
Schel, A. M.; Tranquilli, S.; ZuberbĂĽhler, K. (2009).
1950:
1297:. Vol. 10. Van Nostrand Reinhold Co. p. 464.
2385:"Maternal care and infant transfer in two species of
1780:
Niche separation of black-and-white colobus monkeys (
1307:
435:
between 1830 and 1834. He wrote about the species in
2480:
1879:
1877:
1875:
1427:"The taxonomic diversity of the Colobinae of Africa"
1056:
The mantled guereza is hunted for its skin and meat.
2002:
1637:
1635:
1633:
1631:
1629:
1627:
1540:
1170:de Jong, Y.A.; Butynski, T.M.; Oates, J.F. (2019).
519:(IUCN) assessors Gippolliti and Butynski in 2008.
3009:
2291:
1996:
1872:
454:Distribution map of subspecies of Mantled guereza
4450:
3010:Oates, J. F.; Davies, A. G.; Delson, E. (1994).
2953:Oates, J. F. (1977c). "The guereza and man". In
2928:. New York: Columbia University Press. pp.
2913:
2857:
2801:
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2653:
2651:
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2008:
1809:
1807:
1805:
1803:
1801:
1799:
1797:
1624:
1295:Grzimek's Animal Life Encyclopedia: Mammals I-IV
439:in 1835. It was first seen in Europe in 1890 in
284:. It is native to much of west central and east
4499:Mammals of the Democratic Republic of the Congo
2687:10.2305/IUCN.UK.2020-2.RLTS.T136880A17983306.en
2518:
2163:
1470:. Washington DC: Smithsonian Institution Press.
374:is initiated with vocal communication. After a
2965:. New York: Academic Press. pp. 419–467.
2924:Hunting for Sustainability in Tropical Forests
2751:
2634:10.2305/IUCN.UK.2019-3.RLTS.T40007A17983118.en
2598:
2378:
2376:
2206:
1308:Beolens, B.; Watkins, M.; Grayson, M. (2009).
1260:
858:The mantled guereza is distributed throughout
517:International Union for Conservation of Nature
388:International Union for Conservation of Nature
3080:
2995:. Cambridge, UK: Cambridge University Press.
2737:10.2305/IUCN.UK.2019-3.RLTS.T5150A17983175.en
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2360:
2358:
2356:
2297:
2212:
1883:
1859:
1857:
1794:
1773:
1771:
1769:
1767:
1681:
1679:
1677:
1502:
1500:
1458:
1456:
1417:
1415:
1350:
1195:10.2305/IUCN.UK.2019-3.RLTS.T5143A17944705.en
1015:
778:At the Amora Gedel Park in Awassa, Ethiopia
605:, occurs in Kenya on the Ngong Escarpment of
503:lists seven subspecies of mantled guereza in
437:Neue Wirbelthiere con Abyssinien, Saengthiere
2919:
1727:
1725:
1641:
1474:
427:The mantled guereza was first classified by
2990:
2373:
2245:
1868:. London: Academic Press. pp. 275–321.
1528:
1511:. London: British Museum (Natural History).
1381:
1235:
841:
761:At the Amora Gedel Park in Awassa, Ethiopia
422:
3087:
3073:
2991:Davies, A. G.; Oates, J. F., eds. (1994).
2946:
2604:
2556:
2436:
2353:
2338:
2120:
1854:
1813:
1764:
1674:
1585:
1497:
1453:
1412:
239:
73:
46:
29:
2785:
2735:
2685:
2632:
2070:
2061:
2037:
1722:
1394:(2nd ed.). Academic Press. pp.
1193:
850:Mantled guerezas are arboreal and prefer
680:At the Semliki Wildlife Reserve in Uganda
3021:The natural history of African colobines
1480:
1346:(Google eBook). Allen. 1851. p. 16.
1334:
1241:
1051:
1019:
994:
957:
908:
845:
797:
787:
636:, and the northwestern western guereza,
449:
4484:Mammals of the Central African Republic
2251:
1387:
1292:
783:
4451:
3027:
3018:
2952:
2758:Butynski, T.M.; de Jong, Y.A. (2020).
2658:Butynski, T.M.; de Jong, Y.A. (2020).
2592:
2562:
2344:
2126:
1884:Harris, T. R.; Chapman, C. A. (2007).
1863:
1777:
1685:
1591:
1579:
1506:
1462:
1421:
1286:
1120:
1047:
814:The mantled guereza has a distinctive
4161:
4160:
3068:
2447:
2043:
1790:(Ph.D.). City University of New York.
1731:
1116:
1114:
4494:Mammals of the Republic of the Congo
4436:1EC7CD29-B0D3-483F-A66A-9B4A895EAD3E
2873:International Journal of Primatology
1964:International Journal of Primatology
1598:International Journal of Primatology
1547:International Journal of Primatology
1356:
4459:IUCN Red List least concern species
4052:Black-and-white snub-nosed monkey (
2773:IUCN Red List of Threatened Species
2723:IUCN Red List of Threatened Species
2708:Fashing, P.J.; Oates, J.F. (2019).
2673:IUCN Red List of Threatened Species
2620:IUCN Red List of Threatened Species
2492:Behavioral Ecology and Sociobiology
2383:Horwich, R. H.; Manski, D. (1975).
2265:Behavioral Ecology and Sociobiology
2011:Behavioral Ecology and Sociobiology
1435:Journal of Anthropological Sciences
1215:
1181:IUCN Red List of Threatened Species
953:
925:guereza is mostly preyed on by the
13:
2466:10.1111/j.1439-0310.1977.tb01007.x
1818:) in the Kakamega Forest, Kenya".
1111:
941:is known to hunt the guereza. The
274:Abyssinian black-and-white colobus
14:
4560:
3040:
3012:The diversity of living colobines
2527:"Black and white colobus monkey (
2094:Journal of Comparative Psychology
897:The mantled guereza is primarily
601:Eastern black-and-white colobus,
382:The mantled guereza is listed as
2983:
2829:10.1046/j.1523-1739.2000.98592.x
2550:10.1111/j.1439-0310.2006.01247.x
2184:10.1111/j.1365-294x.2009.04139.x
1391:Primate Adaptation and Evolution
1369:(22). Bonnier Corporation: 393.
1312:The Eponym Dictionary of Mammals
1081:, the Djaffa Mountains guereza,
1038:
929:, but it is also eaten by other
870:(near fresh or brackish water),
766:
749:
725:
709:
685:
668:
645:
98:
3788:Black-crested Sumatran langur (
2454:Zeitschrift fĂĽr Tierpsychologie
2304:American Journal of Primatology
1997:Oates, Davies & Delson 1994
1820:American Journal of Primatology
788:
270:eastern black-and-white colobus
2349:(M.A.). University of Calgary.
1216:Kim, Kenneth (June 21, 2024).
1209:
1085:, and the Mau Forest guereza,
945:is another possible predator.
458:The mantled guereza is in the
355:and some mammals, such as the
1:
3842:East Sumatran banded langur (
3482:Northern plains gray langur (
2955:Rainier III, Prince of Monaco
2141:10.1016/s0003-3472(76)80102-9
1104:
720:At the Lake Naivasha in Kenya
4549:Taxa named by Eduard RĂĽppell
4504:Mammals of Equatorial Guinea
3253:Miss Waldron's red colobus (
975:or displacing a group male.
948:
538:, found in Ethiopia, in the
525:Colobus guereza occidentalis
405:
264:), also known simply as the
7:
4079:Myanmar snub-nosed monkey (
1483:Colobus guereza kikuyuensis
1363:The Popular Science Monthly
737:At the Amora Gedel Park in
505:Mammal Species of the World
10:
4565:
4070:Tonkin snub-nosed monkey (
4043:Golden snub-nosed monkey (
3833:Robinson's banded langur (
3756:Indochinese black langur (
3603:Selangor silvered langur (
3491:Black-footed gray langur (
3388:Ulindi River red colobus (
1957:in Kakamega Forest, Kenya"
1272:Merriam-Webster dictionary
1028:The mantled guereza has a
1016:Reproduction and parenting
892:
4544:Mammals described in 1835
4169:
4143:
4115:
4091:
4023:
3973:
3966:
3768:
3630:Indochinese grey langur (
3530:
3435:
3428:
3400:
3334:Niger Delta red colobus (
3224:
3156:
3149:
3111:
3095:Extant species of family
2885:10.1007/s10764-007-9142-8
2504:10.1007/s00265-011-1282-3
1976:10.1007/s10764-006-9096-2
1905:10.1007/s10329-006-0036-8
1700:10.1007/s10329-005-0173-5
1007:Two Mantled guereza in a
394:. It can survive well in
247:
238:
220:
213:
95:Scientific classification
93:
71:
62:
54:
45:
37:
28:
23:
4061:Gray snub-nosed monkey (
3824:Raffles' banded langur (
3797:Black-and-white langur (
3370:Oustalet's red colobus (
3325:Tana River red colobus (
3166:(Black and white colobi)
1242:Wolfheim, J. H. (1983).
1128:; Reeder, D. M. (eds.).
842:Distribution and habitat
794:3d model of the skeleton
658:Semliki Wildlife Reserve
441:Berlin Zoological Garden
423:Taxonomic classification
4469:Fauna of Central Africa
3914:Sabah grizzled langur (
3806:Black Sumatran langur (
3343:Bouvier's red colobus (
3280:Thollon's red colobus (
2780:: e.T136846A176219904.
2577:10.1163/156853972X00266
1529:Davies & Oates 1994
1388:Fleagle, J. G. (1998).
572:Dodinga Hills guereza,
327:The mantled guereza is
278:black-and-white colobus
4529:Mammals of South Sudan
3950:Natuna Island surili (
3869:White-fronted surili (
3860:White-thighed surili (
3639:Phayre's leaf monkey (
3316:Zanzibar red colobus (
3307:Udzungwa red colobus (
3271:Preuss's red colobus (
3028:Oates, J. F. (1994b).
2680:: e.T136880A17983306.
2345:Grimes, K. H. (2000).
1778:Bocian, C. M. (1997).
1507:Napier, P. H. (1985).
1442:: 7–34. Archived from
1057:
1025:
1012:
966:
914:
878:and selects them over
855:
811:
795:
469:black-and-white colobi
455:
248:Mantled guereza range
55:Female with infant at
4405:Paleobiology Database
3729:White-headed langur (
3684:Gee's golden langur (
3675:Shortridge's langur (
3509:Purple-faced langur (
3464:Kashmir gray langur (
3352:Semliki red colobus (
3298:Ugandan red colobus (
3244:Western red colobus (
2627:: e.T40007A17983118.
2277:10.1007/s002650100358
1788:) in the Ituri Forest
1487:Animal Keeper's Forum
1316:. JHU Press. p.
1055:
1023:
1006:
961:
912:
849:
834:. There is dentition
801:
793:
616:Kilimanjaro guereza,
515:review paper, and by
467:three genera are the
453:
4474:Fauna of East Africa
4033:(Snub-nosed monkeys)
4002:Black-shanked douc (
3923:Maroon leaf monkey (
3500:Tufted gray langur (
3379:Lomami red colobus (
3361:Lang's red colobus (
2963:Primate Conservation
2817:Conservation Biology
2730:: e.T5150A17983175.
2252:Fashing, P. (2001).
1293:Grzimek, B. (1972).
1188:: e.T5143A17944705.
784:Physical description
583:Mau Forest guereza,
4534:Mammals of Tanzania
4514:Mammals of Ethiopia
4479:Mammals of Cameroon
4127:Pig-tailed langur (
4011:Gray-shanked douc (
3747:Delacour's langur (
3621:Dusky leaf monkey (
3594:Tenasserim lutung (
3558:West Javan langur (
3549:East Javan langur (
3473:Tarai gray langur (
3455:Nepal gray langur (
3289:Foa's red colobus (
3262:Pennant's colobus (
3099:(Old World monkeys)
3032:. pp. 347–358.
2605:Mwenja, I. (2019).
2595:, pp. 347–358.
1648:Folia Primatologica
1048:Conservation status
415:derives from Greek
65:Conservation status
4103:Proboscis monkey (
3993:Red-shanked douc (
3702:François' langur (
3576:Germain's langur (
3023:. pp. 75–128.
2408:10.1007/BF02381799
2231:10.1007/BF02629621
2044:Ihobe, H. (2001).
2023:10.1007/bf00170902
1782:Colobus angolensis
1610:10.1007/bf02737386
1582:, pp. 75–128.
1559:10.1007/bf02696112
1359:"Two rare monkeys"
1089:are all listed as
1058:
1026:
1013:
967:
927:crowned hawk-eagle
915:
880:old-growth forests
856:
812:
796:
676:C. g. occidentalis
653:C. g. occidentalis
638:C. g. occidentalis
456:
402:and for its skin.
370:mating system and
4539:Mammals of Uganda
4524:Mammals of Rwanda
4446:
4445:
4163:Taxon identifiers
4154:
4153:
4139:
4138:
4036:
3986:
3962:
3961:
3932:Mentawai langur (
3905:Miller's langur (
3887:Thomas's langur (
3781:
3585:Annamese langur (
3448:
3424:
3423:
3237:
3212:Mantled guereza (
3169:
3100:
3059:(Colobus guereza)
3047:Primate Info Net
3014:. pp. 45–73.
2972:978-0-12-576150-5
2939:978-0-231-50492-8
2316:10.1002/ajp.20232
2172:Molecular Ecology
2081:Colobus polykomos
1999:, pp. 45–73.
1660:10.1159/000155600
1405:978-0-12-260341-9
1357:Heck, L. (1892).
1327:978-0-8018-9533-3
1274:. Merriam-Webster
1253:978-3-7186-0190-5
1218:"Colobus guereza"
1004:
939:common chimpanzee
876:secondary forests
860:Equatorial Africa
852:secondary forests
836:sexual dimorphism
622:Mount Kilimanjaro
603:C. g. kikuyuensis
523:Western guereza,
464:Old World monkeys
357:common chimpanzee
341:evergreen forests
294:Equatorial Guinea
252:
251:
88:
16:Species of mammal
4556:
4519:Mammals of Kenya
4509:Mammals of Gabon
4439:
4438:
4426:
4425:
4413:
4412:
4400:
4399:
4387:
4386:
4374:
4373:
4361:
4360:
4348:
4347:
4335:
4334:
4322:
4321:
4309:
4308:
4296:
4295:
4283:
4282:
4270:
4269:
4257:
4256:
4244:
4243:
4231:
4230:
4218:
4217:
4205:
4204:
4203:
4190:
4189:
4188:
4158:
4157:
4035:
4034:
4030:
3985:
3984:
3980:
3971:
3970:
3941:Siberut langur (
3851:Sarawak surili (
3815:Mitered langur (
3780:
3779:
3775:
3738:Laotian langur (
3731:T. leucocephalus
3722:T. poliocephalus
3605:T. selangorensis
3567:Silvery lutung (
3518:Nilgiri langur (
3447:
3446:
3442:
3433:
3432:
3236:
3235:
3231:
3203:Ursine colobus (
3185:Angola colobus (
3168:
3167:
3163:
3154:
3153:
3098:
3097:Cercopithecidae
3089:
3082:
3075:
3066:
3065:
3033:
3024:
3015:
3006:
2977:
2976:
2950:
2944:
2943:
2927:
2917:
2911:
2910:
2908:
2907:
2901:
2895:. Archived from
2870:
2861:
2855:
2854:
2852:
2851:
2845:
2839:. Archived from
2814:
2805:
2799:
2798:
2796:
2794:
2789:
2755:
2749:
2748:
2746:
2744:
2739:
2705:
2699:
2698:
2696:
2694:
2689:
2655:
2646:
2645:
2643:
2641:
2636:
2602:
2596:
2590:
2581:
2580:
2560:
2554:
2553:
2535:
2522:
2516:
2515:
2487:
2478:
2477:
2445:
2434:
2433:
2431:
2430:
2424:
2418:. Archived from
2393:
2380:
2371:
2364:
2351:
2350:
2342:
2336:
2335:
2295:
2289:
2288:
2262:
2249:
2243:
2242:
2217:RĂĽppell 1835)".
2210:
2204:
2203:
2178:(8): 1777–1790.
2167:
2161:
2160:
2129:Animal Behaviour
2124:
2118:
2117:
2106:10.1037/a0014280
2091:
2074:
2068:
2067:
2065:
2041:
2035:
2034:
2006:
2000:
1994:
1988:
1987:
1961:
1948:
1939:
1938:
1936:
1935:
1929:
1923:. Archived from
1890:
1881:
1870:
1869:
1861:
1852:
1851:
1811:
1792:
1791:
1775:
1762:
1761:
1729:
1720:
1719:
1683:
1672:
1671:
1654:(3–4): 188–208.
1639:
1622:
1621:
1589:
1583:
1577:
1571:
1570:
1538:
1532:
1526:
1513:
1512:
1504:
1495:
1494:
1478:
1472:
1471:
1468:Primate Taxonomy
1460:
1451:
1450:
1448:
1431:
1419:
1410:
1409:
1385:
1379:
1378:
1354:
1348:
1347:
1338:
1332:
1331:
1315:
1305:
1299:
1298:
1290:
1284:
1283:
1281:
1279:
1264:
1258:
1257:
1239:
1233:
1232:
1230:
1228:
1213:
1207:
1206:
1204:
1202:
1197:
1167:
1154:
1153:
1118:
1005:
954:Social structure
935:Verreaux's eagle
804:Museum Wiesbaden
792:
773:juvenile female
770:
753:
729:
713:
689:
672:
649:
396:degraded forests
376:gestation period
345:secondary forest
335:, found in both
282:Old World monkey
243:
226:
103:
102:
82:
77:
76:
50:
40:Henry Doorly Zoo
33:
24:Mantled guereza
21:
20:
4564:
4563:
4559:
4558:
4557:
4555:
4554:
4553:
4489:Mammals of Chad
4449:
4448:
4447:
4442:
4434:
4429:
4421:
4416:
4408:
4403:
4395:
4392:Observation.org
4390:
4382:
4377:
4369:
4364:
4356:
4351:
4343:
4338:
4330:
4325:
4317:
4312:
4304:
4299:
4291:
4286:
4278:
4273:
4265:
4260:
4252:
4247:
4239:
4234:
4228:colobus-guereza
4226:
4221:
4215:Colobus_guereza
4213:
4208:
4201:Colobus guereza
4199:
4198:
4193:
4184:
4183:
4178:
4171:Colobus guereza
4165:
4155:
4150:
4135:
4111:
4087:
4032:
4031:
4029:
4019:
3982:
3981:
3979:
3967:Odd-nosed group
3958:
3896:Hose's langur (
3777:
3776:
3774:
3764:
3720:Cat Ba langur (
3711:Hatinh langur (
3666:Capped langur (
3526:
3444:
3443:
3441:
3420:
3412:Olive colobus (
3396:
3354:P. semlikiensis
3327:P. rufomitratus
3300:P. tephrosceles
3233:
3232:
3230:
3220:
3176:Black colobus (
3165:
3164:
3162:
3145:
3107:
3093:
3049:Colobus guereza
3043:
3038:
3019:Oates (1994a).
3003:
2986:
2981:
2980:
2973:
2951:
2947:
2940:
2918:
2914:
2905:
2903:
2899:
2868:
2862:
2858:
2849:
2847:
2843:
2812:
2806:
2802:
2792:
2790:
2762:Colobus guereza
2756:
2752:
2742:
2740:
2712:Colobus guereza
2706:
2702:
2692:
2690:
2662:Colobus guereza
2656:
2649:
2639:
2637:
2609:Colobus guereza
2603:
2599:
2591:
2584:
2561:
2557:
2533:
2529:Colobus guereza
2523:
2519:
2488:
2481:
2450:Colobus guereza
2446:
2437:
2428:
2426:
2422:
2391:
2381:
2374:
2368:Colobus guereza
2365:
2354:
2343:
2339:
2300:Colobus guereza
2296:
2292:
2260:
2256:Colobus guereza
2250:
2246:
2215:Colobus guereza
2211:
2207:
2168:
2164:
2125:
2121:
2089:
2085:Colobus guereza
2075:
2071:
2050:Pan Africa News
2042:
2038:
2007:
2003:
1995:
1991:
1959:
1955:Colobus guereza
1949:
1942:
1933:
1931:
1927:
1888:
1882:
1873:
1862:
1855:
1816:Colobus guereza
1812:
1795:
1776:
1765:
1750:10.2307/2387676
1734:Colobus guereza
1730:
1723:
1684:
1675:
1644:Colobus guereza
1640:
1625:
1594:Colobus guereza
1590:
1586:
1578:
1574:
1543:Colobus guereza
1539:
1535:
1527:
1516:
1505:
1498:
1479:
1475:
1461:
1454:
1446:
1429:
1420:
1413:
1406:
1386:
1382:
1355:
1351:
1340:
1339:
1335:
1328:
1306:
1302:
1291:
1287:
1277:
1275:
1266:
1265:
1261:
1254:
1240:
1236:
1226:
1224:
1214:
1210:
1200:
1198:
1174:Colobus guereza
1168:
1157:
1142:
1119:
1112:
1107:
1087:C. g. matschiei
1073:, is listed as
1071:C. g. percivali
1050:
1041:
1018:
995:
963:Social grooming
956:
951:
895:
864:montane forests
844:
802:A skull at the
786:
779:
777:
771:
762:
760:
754:
745:
736:
730:
721:
719:
717:C. g. matschiei
714:
705:
696:
693:C. g. matschiei
690:
681:
679:
673:
664:
655:
650:
594:, found in the
592:C. g. percivali
585:C. g. matschiei
580:in South Sudan.
576:, found in the
481:Colobus guereza
425:
408:
261:Colobus guereza
256:mantled guereza
234:
228:
224:Colobus guereza
222:
209:
206:C. guereza
182:Cercopithecidae
97:
89:
78:
74:
67:
17:
12:
11:
5:
4562:
4552:
4551:
4546:
4541:
4536:
4531:
4526:
4521:
4516:
4511:
4506:
4501:
4496:
4491:
4486:
4481:
4476:
4471:
4466:
4461:
4444:
4443:
4441:
4440:
4427:
4414:
4401:
4388:
4375:
4362:
4349:
4336:
4323:
4310:
4297:
4284:
4271:
4258:
4245:
4232:
4219:
4206:
4191:
4175:
4173:
4167:
4166:
4152:
4151:
4144:
4141:
4140:
4137:
4136:
4134:
4133:
4123:
4121:
4113:
4112:
4110:
4109:
4099:
4097:
4089:
4088:
4086:
4085:
4076:
4067:
4058:
4049:
4039:
4037:
4021:
4020:
4018:
4017:
4008:
3999:
3989:
3987:
3968:
3964:
3963:
3960:
3959:
3957:
3956:
3947:
3938:
3929:
3920:
3911:
3902:
3893:
3884:
3878:Javan surili (
3875:
3866:
3857:
3853:P. chrysomelas
3848:
3839:
3830:
3821:
3812:
3803:
3794:
3784:
3782:
3766:
3765:
3763:
3762:
3753:
3744:
3735:
3726:
3717:
3713:T. hatinhensis
3708:
3691:
3690:
3681:
3677:T. shortridgei
3672:
3655:
3654:
3645:
3636:
3632:T. crepusculus
3627:
3610:
3609:
3600:
3591:
3582:
3573:
3564:
3555:
3538:
3536:
3533:Trachypithecus
3528:
3527:
3525:
3524:
3515:
3506:
3497:
3488:
3479:
3470:
3461:
3457:S. schistaceus
3451:
3449:
3445:(Gray langurs)
3430:
3426:
3425:
3422:
3421:
3419:
3418:
3408:
3406:
3398:
3397:
3395:
3394:
3385:
3381:P. parmentieri
3376:
3367:
3358:
3349:
3340:
3331:
3322:
3313:
3304:
3295:
3286:
3277:
3268:
3259:
3250:
3240:
3238:
3222:
3221:
3219:
3218:
3209:
3200:
3194:King colobus (
3191:
3182:
3172:
3170:
3151:
3147:
3146:
3144:
3143:
3137:
3131:
3125:
3119:
3112:
3109:
3108:
3092:
3091:
3084:
3077:
3069:
3063:
3062:
3053:
3042:
3041:External links
3039:
3037:
3036:
3035:
3034:
3025:
3016:
3001:
2987:
2985:
2982:
2979:
2978:
2971:
2945:
2938:
2912:
2856:
2800:
2750:
2700:
2647:
2597:
2582:
2571:(3): 175–197.
2555:
2517:
2479:
2435:
2372:
2352:
2337:
2310:(4): 383–396.
2290:
2271:(3): 219–230.
2244:
2205:
2162:
2119:
2100:(2): 136–150.
2069:
2063:10.5134/143397
2036:
2017:(6): 435–443.
2001:
1989:
1940:
1899:(3): 208–221.
1871:
1853:
1826:(3): 193–209.
1793:
1763:
1744:(4): 241–253.
1721:
1694:(3): 230–238.
1673:
1646:in Ethiopia".
1623:
1604:(4): 299–329.
1584:
1572:
1553:(6): 971–996.
1533:
1514:
1496:
1473:
1452:
1449:on 2013-04-19.
1411:
1404:
1380:
1349:
1333:
1326:
1300:
1285:
1259:
1252:
1234:
1208:
1155:
1140:
1109:
1108:
1106:
1103:
1091:Data Deficient
1083:C. g. gallarum
1079:C. g. dodingae
1049:
1046:
1040:
1037:
1017:
1014:
955:
952:
950:
947:
894:
891:
843:
840:
824:C. g. caudatus
785:
782:
781:
780:
772:
765:
763:
755:
748:
746:
731:
724:
722:
715:
708:
706:
691:
684:
682:
674:
667:
665:
651:
644:
634:C. g. caudatus
630:
629:
618:C. g. caudatus
614:
611:Aberdare Range
599:
596:Matthews Range
588:
581:
574:C. g. dodingae
570:
567:C. g. gallarum
563:
532:
509:C. g. gallarum
493:Angola colobus
429:Eduard RĂĽppell
424:
421:
407:
404:
250:
249:
245:
244:
236:
235:
229:
218:
217:
211:
210:
203:
201:
197:
196:
189:
185:
184:
179:
175:
174:
169:
165:
164:
159:
155:
154:
149:
145:
144:
139:
135:
134:
129:
125:
124:
119:
115:
114:
109:
105:
104:
91:
90:
72:
69:
68:
63:
60:
59:
52:
51:
43:
42:
35:
34:
26:
25:
15:
9:
6:
4:
3:
2:
4561:
4550:
4547:
4545:
4542:
4540:
4537:
4535:
4532:
4530:
4527:
4525:
4522:
4520:
4517:
4515:
4512:
4510:
4507:
4505:
4502:
4500:
4497:
4495:
4492:
4490:
4487:
4485:
4482:
4480:
4477:
4475:
4472:
4470:
4467:
4465:
4462:
4460:
4457:
4456:
4454:
4437:
4432:
4428:
4424:
4419:
4415:
4411:
4406:
4402:
4398:
4393:
4389:
4385:
4380:
4376:
4372:
4367:
4363:
4359:
4354:
4350:
4346:
4341:
4337:
4333:
4328:
4324:
4320:
4315:
4311:
4307:
4302:
4298:
4294:
4289:
4285:
4281:
4276:
4272:
4268:
4263:
4259:
4255:
4250:
4246:
4242:
4237:
4233:
4229:
4224:
4220:
4216:
4211:
4207:
4202:
4196:
4192:
4187:
4181:
4177:
4176:
4174:
4172:
4168:
4164:
4159:
4149:
4148:
4142:
4132:
4130:
4125:
4124:
4122:
4120:
4119:
4114:
4108:
4106:
4101:
4100:
4098:
4096:
4095:
4090:
4084:
4082:
4077:
4075:
4073:
4068:
4066:
4064:
4059:
4057:
4055:
4050:
4048:
4046:
4041:
4040:
4038:
4028:
4027:
4026:Rhinopithecus
4022:
4016:
4014:
4009:
4007:
4005:
4000:
3998:
3996:
3991:
3990:
3988:
3978:
3977:
3972:
3969:
3965:
3955:
3953:
3948:
3946:
3944:
3939:
3937:
3935:
3934:P. potenziani
3930:
3928:
3926:
3921:
3919:
3917:
3912:
3910:
3908:
3903:
3901:
3899:
3894:
3892:
3890:
3885:
3883:
3881:
3876:
3874:
3872:
3867:
3865:
3863:
3858:
3856:
3854:
3849:
3847:
3845:
3840:
3838:
3836:
3831:
3829:
3827:
3822:
3820:
3818:
3813:
3811:
3809:
3804:
3802:
3800:
3795:
3793:
3791:
3790:P. melalophos
3786:
3785:
3783:
3773:
3772:
3767:
3761:
3759:
3754:
3752:
3750:
3745:
3743:
3741:
3736:
3734:
3732:
3727:
3725:
3723:
3718:
3716:
3714:
3709:
3707:
3705:
3699:
3697:
3693:
3692:
3689:
3687:
3682:
3680:
3678:
3673:
3671:
3669:
3663:
3661:
3657:
3656:
3653:
3651:
3648:Popa langur (
3646:
3644:
3642:
3637:
3635:
3633:
3628:
3626:
3624:
3618:
3616:
3612:
3611:
3608:
3606:
3601:
3599:
3597:
3592:
3590:
3588:
3583:
3581:
3579:
3574:
3572:
3570:
3565:
3563:
3561:
3556:
3554:
3552:
3546:
3544:
3540:
3539:
3537:
3535:
3534:
3529:
3523:
3521:
3516:
3514:
3512:
3507:
3505:
3503:
3498:
3496:
3494:
3493:S. hypoleucos
3489:
3487:
3485:
3480:
3478:
3476:
3471:
3469:
3467:
3462:
3460:
3458:
3453:
3452:
3450:
3440:
3439:
3438:Semnopithecus
3434:
3431:
3427:
3417:
3415:
3410:
3409:
3407:
3405:
3404:
3399:
3393:
3391:
3390:P. lulindicus
3386:
3384:
3382:
3377:
3375:
3373:
3368:
3366:
3364:
3359:
3357:
3355:
3350:
3348:
3346:
3341:
3339:
3337:
3332:
3330:
3328:
3323:
3321:
3319:
3314:
3312:
3310:
3309:P. gordonorum
3305:
3303:
3301:
3296:
3294:
3292:
3287:
3285:
3283:
3278:
3276:
3274:
3269:
3267:
3265:
3260:
3258:
3256:
3251:
3249:
3247:
3242:
3241:
3239:
3229:
3228:
3223:
3217:
3215:
3210:
3208:
3206:
3205:C. vellerosus
3201:
3199:
3197:
3192:
3190:
3188:
3187:C. angolensis
3183:
3181:
3179:
3174:
3173:
3171:
3161:
3160:
3155:
3152:
3150:African group
3148:
3142:
3138:
3136:
3132:
3130:
3126:
3124:
3120:
3118:
3114:
3113:
3110:
3105:
3101:
3090:
3085:
3083:
3078:
3076:
3071:
3070:
3067:
3061:
3060:
3054:
3052:
3050:
3045:
3044:
3031:
3026:
3022:
3017:
3013:
3008:
3007:
3004:
3002:9780521331531
2998:
2994:
2989:
2988:
2984:Cited sources
2974:
2968:
2964:
2960:
2959:Bourne, G. H.
2956:
2949:
2941:
2935:
2931:
2926:
2925:
2916:
2902:on 2014-02-22
2898:
2894:
2890:
2886:
2882:
2878:
2874:
2867:
2860:
2846:on 2021-03-03
2842:
2838:
2834:
2830:
2826:
2822:
2818:
2811:
2804:
2788:
2783:
2779:
2775:
2774:
2769:
2767:
2763:
2754:
2738:
2733:
2729:
2725:
2724:
2719:
2717:
2713:
2704:
2688:
2683:
2679:
2675:
2674:
2669:
2667:
2663:
2654:
2652:
2635:
2630:
2626:
2622:
2621:
2616:
2614:
2610:
2601:
2594:
2589:
2587:
2578:
2574:
2570:
2566:
2559:
2551:
2547:
2543:
2539:
2532:
2530:
2521:
2513:
2509:
2505:
2501:
2497:
2493:
2486:
2484:
2475:
2471:
2467:
2463:
2459:
2455:
2451:
2444:
2442:
2440:
2425:on 2015-09-23
2421:
2417:
2413:
2409:
2405:
2401:
2397:
2390:
2388:
2379:
2377:
2369:
2363:
2361:
2359:
2357:
2348:
2341:
2333:
2329:
2325:
2321:
2317:
2313:
2309:
2305:
2301:
2294:
2286:
2282:
2278:
2274:
2270:
2266:
2259:
2257:
2248:
2240:
2236:
2232:
2228:
2224:
2220:
2216:
2209:
2201:
2197:
2193:
2189:
2185:
2181:
2177:
2173:
2166:
2158:
2154:
2150:
2146:
2142:
2138:
2134:
2130:
2123:
2115:
2111:
2107:
2103:
2099:
2095:
2088:
2086:
2082:
2073:
2064:
2059:
2055:
2051:
2047:
2040:
2032:
2028:
2024:
2020:
2016:
2012:
2005:
1998:
1993:
1985:
1981:
1977:
1973:
1969:
1965:
1958:
1956:
1947:
1945:
1930:on 2016-03-05
1926:
1922:
1918:
1914:
1910:
1906:
1902:
1898:
1894:
1887:
1880:
1878:
1876:
1867:
1860:
1858:
1849:
1845:
1841:
1837:
1833:
1829:
1825:
1821:
1817:
1810:
1808:
1806:
1804:
1802:
1800:
1798:
1789:
1785:
1781:
1774:
1772:
1770:
1768:
1759:
1755:
1751:
1747:
1743:
1739:
1735:
1728:
1726:
1717:
1713:
1709:
1705:
1701:
1697:
1693:
1689:
1682:
1680:
1678:
1669:
1665:
1661:
1657:
1653:
1649:
1645:
1638:
1636:
1634:
1632:
1630:
1628:
1619:
1615:
1611:
1607:
1603:
1599:
1595:
1588:
1581:
1576:
1568:
1564:
1560:
1556:
1552:
1548:
1544:
1537:
1530:
1525:
1523:
1521:
1519:
1510:
1503:
1501:
1493:(4): 164–171.
1492:
1488:
1484:
1477:
1469:
1465:
1459:
1457:
1445:
1441:
1437:
1436:
1428:
1424:
1418:
1416:
1407:
1401:
1397:
1393:
1392:
1384:
1376:
1372:
1368:
1364:
1360:
1353:
1345:
1344:
1337:
1329:
1323:
1319:
1314:
1313:
1304:
1296:
1289:
1273:
1269:
1263:
1255:
1249:
1246:. Routledge.
1245:
1238:
1223:
1219:
1212:
1196:
1191:
1187:
1183:
1182:
1177:
1175:
1166:
1164:
1162:
1160:
1151:
1147:
1143:
1141:0-801-88221-4
1137:
1133:
1132:
1127:
1126:Wilson, D. E.
1123:
1122:Groves, C. P.
1117:
1115:
1110:
1102:
1100:
1094:
1092:
1088:
1084:
1080:
1076:
1072:
1067:
1066:Least Concern
1063:
1054:
1045:
1039:Communication
1036:
1034:
1031:
1022:
1010:
993:
991:
986:
981:
976:
973:
972:matrilineally
964:
960:
946:
944:
940:
936:
932:
931:birds of prey
928:
922:
920:
911:
907:
904:
900:
890:
888:
887:
881:
877:
873:
869:
865:
861:
853:
848:
839:
837:
833:
827:
825:
821:
820:C. g. guereza
817:
809:
805:
800:
791:
776:
775:C. g. guereza
769:
764:
759:
758:C. g. guereza
752:
747:
744:
740:
735:
734:C. g. guereza
728:
723:
718:
712:
707:
704:
700:
699:Lake Naivasha
695:with juvenile
694:
688:
683:
678:with juvenile
677:
671:
666:
663:
659:
654:
648:
643:
642:
641:
639:
635:
627:
623:
619:
615:
612:
608:
604:
600:
597:
593:
589:
586:
582:
579:
578:Didinga Hills
575:
571:
568:
564:
561:
558:, and in the
557:
553:
549:
545:
541:
537:
536:C. g. guereza
533:
530:
526:
522:
521:
520:
518:
514:
510:
506:
502:
498:
497:C. angolensis
494:
490:
486:
482:
478:
474:
470:
465:
461:
452:
448:
446:
442:
438:
434:
430:
420:
418:
414:
403:
401:
397:
393:
389:
385:
384:Least Concern
380:
377:
373:
369:
364:
362:
358:
354:
353:birds of prey
350:
346:
342:
338:
334:
330:
325:
323:
319:
315:
311:
307:
303:
299:
295:
291:
287:
283:
279:
275:
271:
267:
263:
262:
257:
246:
242:
237:
232:
227:
225:
219:
216:
215:Binomial name
212:
208:
207:
202:
199:
198:
195:
194:
190:
187:
186:
183:
180:
177:
176:
173:
170:
167:
166:
163:
160:
157:
156:
153:
150:
147:
146:
143:
140:
137:
136:
133:
130:
127:
126:
123:
120:
117:
116:
113:
110:
107:
106:
101:
96:
92:
86:
81:
80:Least Concern
70:
66:
61:
58:
53:
49:
44:
41:
36:
32:
27:
22:
19:
4170:
4145:
4128:
4116:
4104:
4092:
4080:
4072:R. avunculus
4071:
4062:
4053:
4045:R. roxellana
4044:
4024:
4012:
4003:
3994:
3974:
3951:
3942:
3933:
3925:P. rubicunda
3924:
3915:
3906:
3897:
3888:
3879:
3870:
3862:P. siamensis
3861:
3852:
3843:
3835:P. robinsoni
3834:
3826:P. femoralis
3825:
3816:
3808:P. sumatrana
3807:
3798:
3789:
3769:
3757:
3749:T. delacouri
3748:
3739:
3730:
3721:
3712:
3704:T. francoisi
3703:
3696:T. francoisi
3695:
3694:
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3676:
3667:
3659:
3658:
3649:
3640:
3631:
3622:
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3587:T. margarita
3586:
3577:
3569:T. cristatus
3568:
3560:T. mauritius
3559:
3550:
3543:T. cristatus
3542:
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3429:Langur group
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3372:P. oustaleti
3371:
3362:
3353:
3344:
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3326:
3317:
3308:
3299:
3290:
3281:
3272:
3264:P. pennantii
3263:
3255:P. waldronae
3254:
3245:
3234:(Red colobi)
3227:Piliocolobus
3225:
3213:
3211:
3204:
3196:C. polykomos
3195:
3186:
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983:a conflict.
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546:down to the
542:west of the
535:
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496:
489:C. polykomos
488:
485:king colobus
480:
477:olive colobi
457:
436:
426:
416:
412:
409:
381:
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288:, including
280:, a type of
273:
269:
265:
260:
259:
255:
253:
223:
221:
205:
204:
192:
168:Infraorder:
38:Male at the
18:
4301:iNaturalist
4195:Wikispecies
4129:S. concolor
4105:N. larvatus
4081:R. strykeri
4063:R. brelichi
4004:P. nigripes
3907:P. canicrus
3871:P. frontata
3668:T. pileatus
3660:T. pileatus
3623:T. obscurus
3615:T. obscurus
3578:T. germaini
3484:S. entellus
3345:P. bouvieri
3282:P. tholloni
3102:(subfamily
2823:: 207–217.
2793:11 November
2743:11 November
2693:11 November
2640:11 November
2593:Oates 1994b
2460:(1): 1–60.
1580:Oates 1994a
1201:11 November
980:egalitarian
609:and in the
607:Mount Kenya
552:Awash River
544:Rift Valley
447:, Eritrea.
172:Simiiformes
57:MĂĽnster Zoo
4453:Categories
4013:P. cinerea
3995:P. nemaeus
3952:P. natunae
3889:P. thomasi
3844:P. percura
3817:P. mitrata
3799:P. bicolor
3641:T. phayrei
3551:T. auratus
3511:S. vetulus
3403:Procolobus
3273:P. preussi
3214:C. guereza
3178:C. satanas
3141:Haplorhini
3139:Suborder:
2906:2015-09-02
2879:(3): 513.
2850:2015-09-04
2544:(9): 911.
2498:(3): 361.
2429:2015-09-02
2225:(4): 301.
1970:(3): 673.
1934:2015-09-02
1786:C. guereza
1738:Biotropica
1464:Groves, C.
1423:Groves, C.
1105:References
1075:Endangered
1030:polygynous
990:home range
886:Eucalyptus
626:Mount Meru
491:, and the
475:, and the
473:red colobi
392:Endangered
372:copulation
368:polygynous
349:arthropods
162:Haplorhini
158:Suborder:
3976:Pygathrix
3943:P. siberu
3916:P. sabana
3880:P. comata
3778:(Surilis)
3771:Presbytis
3758:T. ebenus
3740:T. laotum
3596:T. barbei
3520:S. johnii
3475:S. hector
3336:P. epieni
3318:P. kirkii
3246:P. badius
3115:Kingdom:
3104:Colobinae
3055:ARKive –
3051:Factsheet
2766:matschiei
2613:percivali
2565:Behaviour
2402:: 49–73.
1848:210133123
1375:0161-7370
1268:"Guereza"
949:Behaviour
832:vestigial
810:, Germany
808:Wiesbaden
598:in Kenya.
560:Blue Nile
556:Omo River
540:highlands
513:Colobinae
460:Colobinae
433:Abyssinia
406:Etymology
337:deciduous
272:, or the
200:Species:
118:Kingdom:
112:Eukaryota
4418:Species+
4371:12100605
4319:10857961
4180:Wikidata
4147:Category
4054:R. bieti
3898:P. hosei
3502:S. priam
3414:P. verus
3363:P. langi
3135:Primates
3129:Mammalia
3123:Chordata
3121:Phylum:
3117:Animalia
2961:(eds.).
2893:20186034
2716:gallarum
2666:dodingae
2538:Ethology
2416:45582808
2396:Primates
2389:monkeys"
2324:16534807
2285:12964572
2239:43214796
2219:Primates
2200:26345361
2192:19302351
2157:20145070
2114:19450021
2031:21024373
1984:25984104
1913:17429575
1893:Primates
1840:31918478
1708:16479411
1688:Primates
1567:23753155
1466:(2001).
1425:(2007).
1227:June 21,
1150:62265494
1124:(2005).
1099:bushmeat
1009:Japanese
933:such as
919:folivore
899:arboreal
868:riparian
743:Ethiopia
400:bushmeat
359:and the
333:arboreal
310:Tanzania
302:Ethiopia
290:Cameroon
178:Family:
152:Primates
142:Mammalia
132:Chordata
128:Phylum:
122:Animalia
108:Domain:
85:IUCN 3.1
4464:Colobus
4431:ZooBank
4358:1000637
4293:5963045
4186:Q628239
4094:Nasalis
3983:(Doucs)
3686:T. geei
3650:T. popa
3466:S. ajax
3291:P. foai
3159:Colobus
3133:Order:
3127:Class:
2837:6827577
2512:7149453
2387:Colobus
2332:7198092
1921:6986510
1758:2387676
1716:8734069
1668:4214737
1618:2006267
1396:207–209
1278:2 April
943:leopard
903:diurnal
893:Ecology
872:gallery
697:At the
656:At the
550:of the
548:reaches
445:Massawa
417:kolobus
413:Colobus
386:by the
361:leopard
329:diurnal
298:Nigeria
276:, is a
266:guereza
231:RĂĽppell
193:Colobus
188:Genus:
148:Order:
138:Class:
83: (
4410:231533
4332:573036
4280:CLBUGU
4267:128444
4223:ARKive
4118:Simias
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937:. The
816:pelage
739:Awassa
662:Uganda
562:gorge.
554:, the
501:Groves
483:, the
471:, the
314:Uganda
286:Africa
268:, the
233:, 1835
4397:79442
4384:33548
4314:IRMNG
4306:43527
4254:5ZN3P
4241:26339
3698:group
3662:group
3617:group
3545:group
2900:(PDF)
2889:S2CID
2869:(PDF)
2844:(PDF)
2833:S2CID
2813:(PDF)
2764:ssp.
2714:ssp.
2664:ssp.
2611:ssp.
2534:(PDF)
2508:S2CID
2423:(PDF)
2412:S2CID
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2281:S2CID
2261:(PDF)
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2196:S2CID
2153:S2CID
2090:(PDF)
2027:S2CID
1980:S2CID
1960:(PDF)
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1917:S2CID
1889:(PDF)
1844:S2CID
1754:JSTOR
1712:S2CID
1614:S2CID
1563:S2CID
1447:(PDF)
1430:(PDF)
1033:harem
756:male
732:male
703:Kenya
322:trunk
306:Kenya
4423:3758
4379:NCBI
4345:5143
4340:IUCN
4327:ITIS
4288:GBIF
4275:EPPO
4236:BOLD
2997:ISBN
2967:ISBN
2934:ISBN
2795:2021
2778:2020
2745:2021
2728:2019
2695:2021
2678:2020
2642:2021
2625:2019
2470:PMID
2320:PMID
2188:PMID
2145:PMID
2110:PMID
2083:and
1909:PMID
1836:PMID
1784:and
1704:PMID
1664:PMID
1596:)".
1485:)".
1400:ISBN
1371:ISSN
1322:ISBN
1280:2013
1248:ISBN
1229:2024
1203:2021
1186:2019
1146:OCLC
1136:ISBN
1062:IUCN
1011:zoo.
624:and
529:Nile
339:and
331:and
318:Chad
316:and
254:The
4366:MSW
4353:MDD
4262:EoL
4249:CoL
4210:ADW
2930:427
2881:doi
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