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Poales

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and thus form a large proportion of the forage of grazing livestock. Possibly due to pastoral nostalgia or simply a desire for open areas for play, they dominate most Western yards as lawns, which consume vast sums of money in upkeep (artificial grazing—mowing—for aesthetics and to keep the allergenic flowers suppressed, irrigation, and fertilizer). Many Bromeliaceae are used as ornamental plants (and one, the pineapple, is internationally grown in the tropics for fruit). Many wetland species of sedges, rushes, grasses, and cattails are important habitat plants for waterfowl, are used in weaving chair seats, and (especially cattails) were important pre-agricultural food sources for man. Two sedges, chufa (
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as well as bamboos (mostly used structurally, like wood, but somewhat as vegetables), and a few "seasonings" like sugarcane and lemongrass. Graminoids, especially the grasses, are typically dominant in open (low moisture but not yet arid, or also fire climax) habitats like prairie/steppe and savannah
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Barrett, Craig F.; Baker, William J.; Comer, Jason R.; Conran, John G.; Lahmeyer, Sean C.; Leebens-Mack, James H.; Li, Jeff; Lim, Gwynne S.; Mayfield-Jones, Dustin R.; Perez, Leticia; Medina, Jesus; Pires, J. Chris; Santos, Cristian; Wm. Stevenson, Dennis; Zomlefer, Wendy B.; Davis, Jerrold I.
572:. A phylogenetic analysis resolved most relationships within the order but found weak support for the monophyly of the cyperid clade. The relationship between Centrolepidaceae and Restoniaceae within the restiid clade remains unclear; the first may actually be embedded in the latter. 1487:
The Poales are the most economically important order of monocots and possibly the most important order of plants in general. Within the order, by far the most important family economically is the family of grasses (Poaceae, syn. Gramineae), which includes the starch staples
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Briggs, Barbara G.; Marchant, Adam D.; Perkins, Andrew J. (2014). "Phylogeny of the restiid clade (Poales) and implications for the classification of Anarthriaceae, Centrolepidaceae and Australian Restionaceae".
435:(1998) adopted the same placement of the order, although it used the spelling "commelinoids". It did not include the Bromeliaceae and Mayaceae, but had the additional families 529:
period about 66 million years ago, though some studies (e.g., Bremer, 2002) suggest the origin of the group may extend to nearly 115 million years ago, likely in
635: 553: 1735: 2934: 2746: 1014: 698: 569: 557: 2785: 847: 561: 1587:"Plastid genomes reveal support for deep phylogenetic relationships and extensive rate variation among palms and other commelinid monocots" 1876: 315:, which possess very reduced, one-flowered inflorescences). The flowers of many species are wind pollinated; the seeds usually contain 2720: 1924: 2759: 2764: 1777: 1543:"An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG III" 1474: 482: 1456: 540:
The phylogenetic position of Poales within the commelinids was difficult to resolve, but an analysis using complete
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D.J. Mabberley. 2000. The Plant-Book, 2nd ed. Cambridge University Press, Cambridge, UK, 858 p.
2608: 1353: 290: 2821: 1797: 2813: 2751: 2694: 8: 2883: 2575: 1909: 1752: 1511: 128: 1854: 277:. Sixteen plant families are currently recognized by botanists to be part of Poales. 2808: 2681: 2569: 2384: 1773: 1757: 1703: 1660: 1616: 1608: 1559: 1542: 1517: 1266: 498: 212: 2888: 2686: 2646: 2581: 2166: 1995: 1965: 1793: 1747: 1695: 1650: 1598: 1564: 1554: 1380: 1376: 1289: 1285: 1210: 1049: 506: 451: 407: 2660: 1768:
Judd, W. S., C. S. Campbell, E. A. Kellogg, P. F. Stevens, M. J. Donoghue (2002).
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Linder, H. Peter; Rudall, Paula J. (2005). "Evolutionary History of the Poales".
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did not include an order named Poales, assigning these families to the orders
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pp. 276–292 (Poales). Sinauer Associates, Sunderland, Massachusetts.
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Bouchenak-Khelladi, Yanis; Muasya, A. Muthama; Linder, H. Peter (2014).
2487: 2464: 2389: 2342: 2337: 2317: 2279: 2195: 2078: 2017: 2009: 1639:"A revised evolutionary history of Poales: origins and diversification" 1569: 1333: 1310: 875: 786: 718: 526: 490: 432: 424: 387: 355: 274: 266: 86: 51: 1655: 1638: 1603: 1586: 2738: 2707: 2517: 2497: 2454: 2444: 2161: 2022: 1953: 801: 663: 510: 459: 418: 381: 270: 91: 35: 2725: 2602: 1426: 2847: 2625: 2512: 2421: 2416: 2404: 2352: 2322: 2246: 2143: 2109: 2084: 2027: 1838: 1699: 1521:) are still at least locally important wetland starchy root crops. 475: 467: 81: 76: 61: 56: 46: 2712: 2556: 2479: 2459: 2309: 2299: 2289: 2284: 2171: 1318: 938: 392: 332: 262: 96: 71: 1736:"Gondwanan Evolution of the Grass Alliance of Families (Poales)" 2522: 2256: 2234: 2054: 1826: 1497: 1489: 534: 525:
The earliest fossils attributed to the Poales date to the late
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are typically small, enclosed by bracts, and arranged in
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Plant Systematics: A Phylogenetic Approach, 2nd edition.
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the name was used for an order placed in the subclass
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The four most species-rich families in the order are:
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Annual Review of Ecology, Evolution, and Systematics
1678: 1389:. The order consisted of one family only (1981): 2916: 1577: 1515:, also a significant weed) and water chestnut ( 1288:it consisted of only the single family. In the 261:, and includes families of plants such as the 1870: 1336:now assigns the plants involved to the order 1783: 1536: 1534: 1292:it is used for an order (placed in subclass 544:found support for Poales as sister group of 1455:. Unsourced material may be challenged and 1877: 1863: 520: 107: 16:Order of monocotyledonous flowering plants 1751: 1654: 1602: 1568: 1558: 1531: 1475:Learn how and when to remove this message 339:, and accepts the following 16 families: 1798:10.1146/annurev.ecolsys.36.102403.135635 1643:Botanical Journal of the Linnean Society 1547:Botanical Journal of the Linnean Society 1347: 1260: 284: 1806:Flora of the Southeastern United States 2935:Extant Maastrichtian first appearances 2917: 1733: 1409:now assigns these plants to the order 309:(except in three species of the genus 2607: 2606: 1858: 533:. The earliest known fossils include 501:(first edition, first volume, 1926), 2876:3a59907c-ae2c-40a6-9690-8a500742581f 2840:459CD035-D50B-4F3F-B6B7-514D6338A850 2801:09a2388f-3f6e-4f47-8319-0ee09f72e127 1453:adding citations to reliable sources 1420: 1251: 1541:Angiosperm Phylogeny Group (2009). 13: 1753:10.1111/j.0014-3820.2002.tb01451.x 331:(2009) accepts the order within a 14: 2946: 1819: 2587: 2586: 2555: 1837: 1825: 1670: 1560:10.1111/j.1095-8339.2009.00996.x 1425: 1236: 1219: 1202: 1185: 1168: 132: 39: 1727: 1343: 1298:) and circumscribed as (1981): 439:(now included in Thurniaceae), 1714: 280: 1: 1524: 1284:(update, of 1964) and in the 1375:. The name was published by 1256: 1159:Eriocaulaceae: 1,150 species 1141: 497:(update of 1964) and in the 7: 1276:was a name for an order of 1156:Bromeliaceae: 3,170 species 515:Bentham & Hooker system 322: 10: 2951: 487:descriptive botanical name 2615: 2564: 2553: 2478: 2430: 2412: 2403: 2380: 2369: 2308: 2255: 2242: 2233: 2220: 2209: 2191: 2180: 2152: 2141: 2100: 2050: 2041: 2008: 2004: 1986: 1975: 1949: 1938: 1898: 1153:Cyperaceae: 5,500 species 1062: 1047: 1040: 1025: 1018: 973: 958: 951: 936: 929: 914: 907: 897: 873: 858: 851: 841: 799: 784: 777: 762: 755: 731: 716: 709: 702: 692: 685: 661: 646: 639: 629: 602: 587: 580: 238: 231: 224: 219: 129:Scientific classification 127: 115: 106: 23: 443:(now in Typhaceae), and 395:(Grasses, Bamboo, Reeds) 31:- Recent (but see text) 1416: 1150:Poaceae: 12,070 species 521:Evolution and phylogeny 509:(last revised 1935) or 1357: 1354:Eriocaulon decangulare 1270: 517:(third volume, 1883). 298: 291:Billbergia pyramidalis 2822:Paleobiology Database 1804:Small, J. K. (1903). 1351: 1264: 450:The morphology-based 288: 1834:at Wikimedia Commons 1449:improve this section 636:Bromeliad clade 1393:order Eriocaulales 1164:Diversity of Poales 904:Graminid clade 239:About 1,050 genera 1734:Bremer, K (2002). 1512:Cyperus esculentus 1358: 1280:. As used in the 1271: 1015:Restiid clade 699:Cyperid clade 299: 2930:Angiosperm orders 2912: 2911: 2809:Open Tree of Life 2609:Taxon identifiers 2600: 2599: 2570:Basal angiosperms 2551: 2550: 2547: 2546: 2543: 2542: 2539: 2538: 2535: 2534: 2531: 2530: 2385:Berberidopsidales 2365: 2364: 2361: 2360: 2137: 2136: 2133: 2132: 1830:Media related to 1656:10.1111/boj.12160 1604:10.1111/nph.13617 1518:Eleocharis dulcis 1485: 1484: 1477: 1267:Cyperus javanicus 1252:Historic taxonomy 1138: 1137: 1129: 1128: 1120: 1119: 1111: 1110: 1102: 1101: 1093: 1092: 1084: 1083: 1075: 1074: 1004: 1003: 995: 994: 986: 985: 886: 885: 830: 829: 821: 820: 812: 811: 744: 743: 674: 673: 615: 614: 499:Hutchinson system 243: 242: 215: 121:Triticum aestivum 2942: 2905: 2904: 2892: 2891: 2879: 2878: 2866: 2865: 2856: 2855: 2843: 2842: 2830: 2829: 2817: 2816: 2804: 2803: 2794: 2793: 2781: 2780: 2778:NHMSYS0021060390 2768: 2767: 2755: 2754: 2742: 2741: 2729: 2728: 2716: 2715: 2703: 2702: 2690: 2689: 2677: 2676: 2664: 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2262: 2260: 2258: 2254: 2248: 2245: 2244: 2241: 2238: 2236: 2232: 2226: 2223: 2222: 2219: 2216: 2213: 2208: 2202: 2199: 2197: 2194: 2193: 2190: 2187: 2184: 2183:Core eudicots 2179: 2173: 2170: 2168: 2165: 2163: 2160: 2158: 2155: 2154: 2151: 2148: 2145: 2140: 2126: 2123: 2121: 2118: 2116: 2113: 2111: 2108: 2107: 2105: 2103: 2099: 2092: 2089: 2086: 2083: 2080: 2077: 2074: 2071: 2068: 2067:Petrosaviales 2065: 2062: 2059: 2056: 2053: 2052: 2049: 2046: 2044: 2040: 2034: 2031: 2029: 2026: 2024: 2021: 2019: 2016: 2015: 2013: 2011: 2007: 2003: 1997: 1994: 1992: 1991:Chloranthales 1989: 1988: 1985: 1982: 1979: 1974: 1967: 1964: 1961: 1958: 1955: 1952: 1951: 1948: 1945: 1942: 1937: 1931: 1928: 1926: 1923: 1921: 1918: 1916: 1913: 1911: 1908: 1905: 1901: 1900: 1897: 1892: 1888: 1880: 1875: 1873: 1868: 1866: 1861: 1860: 1857: 1851: 1848: 1845: 1840: 1836: 1833: 1828: 1824: 1823: 1815: 1814:United States 1811: 1807: 1803: 1799: 1795: 1791: 1787: 1782: 1779: 1778:0-87893-403-0 1775: 1771: 1767: 1763: 1759: 1754: 1749: 1745: 1741: 1737: 1732: 1731: 1717: 1709: 1705: 1701: 1697: 1693: 1689: 1681: 1673: 1666: 1662: 1657: 1652: 1648: 1644: 1640: 1633: 1631: 1622: 1618: 1614: 1610: 1605: 1600: 1596: 1592: 1588: 1580: 1571: 1566: 1561: 1556: 1552: 1548: 1544: 1537: 1535: 1530: 1522: 1520: 1519: 1514: 1513: 1507: 1503: 1499: 1495: 1491: 1479: 1476: 1468: 1458: 1454: 1450: 1444: 1443: 1439: 1434:This section 1432: 1428: 1423: 1422: 1414: 1412: 1408: 1407:APG IV system 1399: 1398:Eriocaulaceae 1395: 1394: 1392: 1391: 1390: 1388: 1387: 1382: 1378: 1374: 1370: 1366: 1362: 1356: 1355: 1350: 1341: 1339: 1335: 1325: 1321: 1320: 1315: 1313: 1312: 1307: 1306: 1305: 1301: 1300: 1299: 1297: 1296: 1291: 1287: 1283: 1282:Engler system 1279: 1275: 1269: 1268: 1263: 1245: 1239: 1234: 1230: 1229: 1222: 1217: 1213: 1212: 1205: 1200: 1196: 1195: 1194:Carex demissa 1188: 1183: 1179: 1178: 1171: 1166: 1163: 1162: 1158: 1155: 1152: 1149: 1148: 1147: 1134: 1133: 1125: 1124: 1116: 1115: 1107: 1106: 1098: 1097: 1089: 1088: 1080: 1079: 1071: 1070: 1067: 1066: 1060: 1059: 1056: 1055: 1052: 1051: 1045: 1044: 1038: 1037: 1034: 1033: 1030: 1029: 1028:Anarthriaceae 1023: 1022: 1016: 1013: 1012: 1009: 1008: 1000: 999: 991: 990: 982: 981: 978: 977: 971: 970: 967: 966: 963: 962: 956: 955: 949: 948: 945: 944: 941: 940: 934: 933: 927: 926: 923: 922: 919: 918: 912: 911: 905: 902: 901: 895: 894: 891: 890: 882: 881: 878: 877: 871: 870: 867: 866: 863: 862: 861:Eriocaulaceae 856: 855: 849: 846: 845: 839: 838: 835: 834: 826: 825: 817: 816: 808: 807: 804: 803: 797: 796: 793: 792: 789: 788: 782: 781: 775: 774: 771: 770: 767: 766: 760: 759: 753: 752: 749: 748: 740: 739: 736: 735: 729: 728: 725: 724: 721: 720: 714: 713: 707: 706: 700: 697: 696: 690: 689: 683: 682: 679: 678: 670: 669: 666: 665: 659: 658: 655: 654: 651: 650: 644: 643: 637: 634: 633: 627: 624: 623: 620: 619: 611: 610: 607: 606: 600: 599: 596: 595: 592: 591: 585: 584: 578: 577: 573: 571: 567: 563: 559: 555: 551: 547: 543: 538: 536: 532: 531:South America 528: 518: 516: 512: 508: 504: 500: 496: 495:Engler system 492: 488: 484: 479: 477: 473: 469: 465: 461: 457: 453: 448: 446: 445:Hydatellaceae 442: 441:Sparganiaceae 438: 434: 426: 423: 420: 417: 415: 412: 409: 405: 402: 400: 397: 394: 391: 389: 386: 383: 380: 378: 375: 373: 370: 368: 367:Eriocaulaceae 365: 363: 360: 357: 354: 351: 348: 346: 345:Anarthriaceae 343: 342: 340: 338: 335:clade called 334: 330: 320: 318: 314: 313: 308: 304: 297: 293: 292: 287: 278: 276: 272: 268: 264: 260: 256: 252: 248: 237: 234: 230: 227: 223: 218: 214: 209: 206: 203: 202: 199: 196: 193: 190: 189: 186: 183: 180: 177: 176: 173: 170: 167: 164: 163: 160: 159:Tracheophytes 157: 154: 151: 150: 147: 144: 141: 140: 135: 130: 126: 122: 118: 114: 110: 105: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 48: 43: 37: 30: 22: 19: 2616: 2440:Aquifoliales 2333:Picramniales 2295:Cucurbitales 2275:Malpighiales 2225:Saxifragales 2157:Ranunculales 2125:Zingiberales 2120:Commelinales 2114: 2073:Dioscoreales 1954:Amborellales 1930:Angiospermae 1805: 1789: 1785: 1769: 1743: 1739: 1728:Bibliography 1716: 1694:(1): 24–46. 1691: 1687: 1680: 1646: 1642: 1594: 1590: 1579: 1550: 1546: 1516: 1510: 1486: 1471: 1462: 1447:Please help 1435: 1410: 1404: 1386:Commelinidae 1384: 1361:Eriocaulales 1360: 1359: 1352: 1344:Eriocaulales 1337: 1331: 1323: 1317: 1309: 1303: 1295:Commelinidae 1293: 1273: 1272: 1265: 1226: 1214:, Xyridaceae 1209: 1197:, Cyperaceae 1192: 1175: 1145: 1065:Restionaceae 1063: 1048: 1026: 974: 959: 937: 915: 874: 859: 800: 785: 763: 732: 717: 662: 649:Bromeliaceae 647: 625: 605:Zingiberales 603: 590:Commelinales 588: 550:Zingiberales 546:Commelinales 539: 537:and fruits. 524: 480: 464:Hydatellales 449: 431:The earlier 430: 404:Restionaceae 352:(Bromeliads) 350:Bromeliaceae 326: 310: 300: 296:Bromeliaceae 289: 249:are a large 246: 244: 225: 207: 191: 178: 165: 152: 120: 117:Common wheat 18: 2734:iNaturalist 2641:Wikispecies 2508:Gentianales 2503:Boraginales 2432:Campanulids 2348:Brassicales 2270:Celastrales 2212:Superrosids 2201:Dilleniales 2102:Commelinids 2091:Asparagales 2061:Alismatales 2033:Magnoliales 1960:Nymphaeales 1941:Angiosperms 1902:Supergroup 1792:: 107–124. 1649:(1): 4–16. 1570:10654/18083 1180:, Typhaceae 765:Thurniaceae 734:Rapateaceae 503:Glumiflorae 472:Restionales 456:Bromeliales 437:Prioniaceae 414:Thurniaceae 406:(including 399:Rapateaceae 337:commelinids 281:Description 198:Commelinids 172:Angiosperms 2919:Categories 2576:Alismatids 2488:Icacinales 2465:Dipsacales 2390:Santalales 2343:Huerteales 2338:Sapindales 2318:Geraniales 2280:Oxalidales 2196:Gunnerales 2079:Pandanales 2018:Canellales 2010:Magnoliids 1885:Orders of 1525:References 1465:March 2019 1334:APG system 1311:Cyperaceae 876:Xyridaceae 787:Cyperaceae 719:Mayacaceae 527:Cretaceous 491:Graminales 433:APG system 425:Xyridaceae 421:(Cattails) 388:Mayacaceae 356:Cyperaceae 294:of family 267:bromeliads 34:66–0  2523:Vahliales 2518:Solanales 2498:Garryales 2455:Bruniales 2445:Asterales 2162:Proteales 2023:Piperales 1910:Bryophyta 1740:Evolution 1708:0040-0262 1665:0024-4074 1613:0028-646X 1436:does not 1379:. In the 1324:Gramineae 1304:Cyperales 1274:Cyperales 1257:Cyperales 1246:, Poaceae 1142:Diversity 802:Juncaceae 664:Typhaceae 554:bromeliad 511:Glumaceae 481:In early 460:Cyperales 419:Typhaceae 382:Juncaceae 233:Diversity 226:See text 220:Families 142:Kingdom: 2871:VicFlora 2860:VASCAN: 2853:50181956 2848:Tropicos 2626:Wikidata 2592:Category 2582:Lilioids 2513:Lamiales 2422:Ericales 2417:Cornales 2405:Asterids 2353:Malvales 2323:Myrtales 2144:Eudicots 2110:Arecales 2085:Liliales 2055:Acorales 2043:Monocots 2028:Laurales 1810:New York 1762:12206239 1621:26350789 1585:(2015). 566:graminid 489:such as 476:Typhales 468:Juncales 384:(Rushes) 358:(Sedges) 323:Taxonomy 185:Monocots 2480:Lamiids 2460:Apiales 2310:Malvids 2300:Fagales 2290:Rosales 2285:Fabales 2247:Vitales 2172:Buxales 1457:removed 1442:sources 1396:family 1367:for an 1319:Poaceae 1316:family 1308:family 939:Poaceae 558:cyperid 513:in the 505:in the 493:in the 483:systems 393:Poaceae 333:monocot 303:flowers 263:grasses 257:in the 204:Order: 146:Plantae 2925:Poales 2902:234035 2827:182987 2814:921871 2798:NZOR: 2765:846620 2661:214973 2647:Poales 2632:Q28502 2617:Poales 2257:Fabids 2235:Rosids 2115:Poales 1891:APG IV 1844:Poales 1832:Poales 1808:, 48. 1776:  1760:  1706:  1663:  1619:  1611:  1504:, and 1498:millet 1490:barley 1411:Poales 1338:Poales 1302:order 626:Poales 568:, and 535:pollen 317:starch 312:Mayaca 275:sedges 271:rushes 247:Poales 208:Poales 24:Poales 2897:WoRMS 2835:Plazi 2791:38820 2752:12583 2747:IRMNG 2739:47162 2713:1POAO 2687:627VM 1904:Plant 1850:APWeb 1688:Taxon 1506:wheat 1494:maize 1369:order 1363:is a 562:xyrid 548:plus 251:order 213:Small 192:Clade 179:Clade 166:Clade 153:Clade 2786:NCBI 2760:ITIS 2726:1369 2721:GBIF 2708:EPPO 2700:4075 2669:BOLD 2656:APNI 1774:ISBN 1758:PMID 1704:ISSN 1661:ISSN 1617:PMID 1609:ISSN 1502:rice 1440:any 1438:cite 1417:Uses 1405:The 1332:The 1322:(or 474:and 327:The 301:The 273:and 245:The 42:PreꞒ 2884:WFO 2773:NBN 2695:EoL 2682:CoL 2674:200 1794:doi 1748:doi 1696:doi 1651:doi 1647:175 1599:doi 1595:209 1565:hdl 1555:doi 1551:161 1451:by 1371:of 253:of 2921:: 2899:: 2886:: 2873:: 2863:55 2850:: 2837:: 2824:: 2811:: 2788:: 2775:: 2762:: 2749:: 2736:: 2723:: 2710:: 2697:: 2684:: 2671:: 2658:: 2643:: 2628:: 1812:, 1790:36 1788:. 1756:. 1744:56 1742:. 1738:. 1702:. 1692:63 1690:. 1659:. 1645:. 1641:. 1629:^ 1615:. 1607:. 1593:. 1589:. 1563:. 1549:. 1545:. 1533:^ 1500:, 1496:, 1492:, 1413:. 1340:. 564:, 560:, 556:, 478:. 470:, 466:, 462:, 458:, 319:. 269:, 265:, 194:: 181:: 168:: 155:: 123:) 92:Pg 36:Ma 2580:³ 2574:² 2568:¹ 2093:³ 2087:³ 2081:³ 2075:³ 2069:³ 2063:² 2057:² 1968:¹ 1962:¹ 1956:¹ 1906:: 1893:) 1889:( 1878:e 1871:t 1864:v 1800:. 1796:: 1780:. 1764:. 1750:: 1710:. 1698:: 1667:. 1653:: 1623:. 1601:: 1573:. 1567:: 1557:: 1478:) 1472:( 1467:) 1463:( 1459:. 1445:. 1326:) 410:) 119:( 97:N 87:K 82:J 77:T 72:P 67:C 62:D 57:S 52:O 47:Ꞓ

Index

Late Cretaceous
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Common wheat
Scientific classification
Edit this classification
Plantae
Tracheophytes
Angiosperms
Monocots
Commelinids
Poales
Small
Diversity
order
flowering plants
monocotyledons
grasses

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