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Pseudosuchia

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crocodiles than to birds" (Gauthier and Padian), "Extant crocodiles and all extinct archosaurs that are closer to crocodiles than they are to birds" (Gauthier, 1986), and more recently "the most inclusive clade within Archosauria that includes Crocodylia but not Aves" (Senter, 2005). As a branch-based clade, Pseudosuchia is the
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Pseudosuchia was defined as a stem-based clade in 1985. It includes crocodiles and all archosaurs more closely related to crocodiles than to birds. A second clade with a similar definition, Crocodylotarsi, was named in 1988, possibly as a replacement for Pseudosuchia. The name Pseudosuchia, meaning
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placement of phytosaurs. In 2011, Sterling J. Nesbitt found phytosaurs to be the sister taxon of Archosauria, and therefore not crocodile-line archosaurs. Because phytosaurs are included in the definition of Crurotarsi, crurotarsans are not solely crocodile-line archosaurs, but also bird-line
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for all archosaurs more closely related to crocodilians than to birds. This made the name Pseudosuchia somewhat ironic because true crocodiles (i.e. members of Crocodylia) were now included in the group. Phylogenetic definitions of Pseudosuchia include "Crocodiles and all archosaurs closer to
686:'s 2011 analysis places one crurotarsan group, Phytosauria, outside Pseudosuchia. Since the definition of Crurotarsi relies on phytosaurs, their placement outside Pseudosuchia (and thus Archosauria) means that the clade Crurotarsi includes both pseudosuchians and avemetatarsalians. 1666:, the retention of phytosaurs within Pseudosuchia, and a close relation between aetosaurs and crocodylomorphs, replicate the results of older studies. However, the findings of Nesbitt (2011) have been more widely supported by pseudosuchian-focused analyses published since 2011. 678:. It is not equivalent to Pseudosuchia, which by definition must include all crocodilian-line archosaurs. For many years, Pseudosuchia and Crurotarsi have been considered partial synonyms because the latter clade encompasses all crocodilian-line archosaurs in most 804:
progressed, the Protosuchia gave rise to more typically crocodile-like forms. While dinosaurs were the dominant animals on land, the crocodiles flourished in rivers, swamps, and the oceans, with far greater diversity than they have today. With the
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Contrary to popular belief, crocodilians differ significantly from their ancestors and distant relatives, as Pseudosuchia contains a staggering diversity of reptiles with many different lifestyles. Early pseudosuchians were successful in the
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Ezcurra, Martín D.; Fiorelli, Lucas E.; Martinelli, Agustín G.; Rocher, Sebastián; von Baczko, M. Belén; Ezpeleta, Miguel; Taborda, Jeremías R.A.; Hechenleitner, E. Martín; Trotteyn, M. Jimena; Desojo, Julia B. (11 September 2017).
837:("true crocodiles") as well. Crocodylotarsi may have been named to remove confusion, but as a stem-based clade, it is synonymous with Pseudosuchia. Because Pseudosuchia was named first, it has precedence. A third group, 328:
and all archosaurs more closely related to crocodilians than to birds. Pseudosuchians are also informally known as "crocodilian-line archosaurs". Despite Pseudosuchia meaning "false crocodiles", the name is a
524:. The clade Pseudosuchia as originally defined could still be used to identify crocodilian-line archosaurs, and since many recent studies support Nesbitt's findings, Pseudosuchia is again commonly used. 846:
archosaurs and phytosaurs. Under this phylogeny, Crurotarsi includes phytosaurs, crocodiles, pterosaurs, and dinosaurs, while Pseudosuchia still contains only crocodile-line archosaurs. Below is a
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Gauthier, J. A.; Padian, K. (1985). "Phylogenetic, functional, and aerodynamic analyses of the origin of birds and their flight". In Hecht, M. K.; Ostrom, J. H.; Viohl, G.; Wellnhofer, P. (eds.).
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Interestingly, the Mesozoic range of cranial disparity is higher than the Triassic one, suggesting crocodylomorphs attained a high degree of diversification compared to Triassic pseudosuchians.
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The following cladogram is from a slightly older study, Brusatte, Benton, Desojo and Langer (2010). Bold terminal taxa are collapsed. Several results of the study, such as the retention of a
833:"false crocodiles", has been used for over a century, and traditionally included only non-crocodilians, but when defined as a clade, Pseudosuchia came to include the group 710:', although achieving it a different way. The body is often protected by two or more rows of armored plates. Many crurotarsans reached lengths of three meters or more. 841:, traditionally included the same archosaurs as Pseudosuchia, but as a node-based clade it is not synonymous. The scope of Crurotarsi has recently been changed by the 1788: 1082: 702:; the snout is narrow and tends to be elongated, the neck is short and strong, and the limb posture ranges from a typical reptilian sprawl to an erect stance like 3003: 593:
dinosaur-like bipeds — to be typical pseudosuchians. These small forms were assumed to be the ancestors of all later archosaurs. The name Pseudosuchia became a
505:(evolutionary grouping) of archosaurs encompassing most reptiles previously identified as pseudosuchians. By this time, Pseudosuchia had also been defined as a 2928:; Julia B. Desojo; Max C. Langer. 2010. The higher-level phylogeny of Archosauria (Tetrapoda: Diapsida). Journal of Systematic Palaeontology, 8: 1, 3 — 47pp. 2867:"Rapid Initial Morphospace Expansion and Delayed Morphological Disparity Peak in the First 100 Million Years of the Archosauromorph Evolutionary Radiation" 516:
proposed that Crurotarsi, as it was then defined, must include not only crocodilian-line archosaurs, but all other archosaurs including birds, non-avian
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and all of its descendants. Benton and Clark also named a group called Crocodylotarsi, which includes most taxa now considered pseudosuchians.
2974: 2961: 789:), the latter being the ancestors of modern-day crocodiles. A study published in 2010 postulates that there is significant evidence that 3052: 3031: 384:. As a result, Crurotarsi could be a much broader clade than Pseudosuchia. Other recent studies support a more traditional phylogeny. 806: 2806:"Compound-specific carbon isotopes from Earth's largest flood basalt eruptions directly linked to the end-Triassic mass extinction" 465:
periods, although only a single subset of crocodylomorphs, the Crocodilia, survive to the present day. Living crocodilians include
809:, the dinosaurs became extinct, with the exception of the birds, while the crocodilians continued with little change. Today, the 4742: 774: 670:
to supplant Pseudosuchia. However, Sereno defined Crurotarsi as a node-based clade, relying on the inclusion of groups such as
577:. Zittel's aetosaurs were placed in their own suborder, Aetosauria. Colbert considered small lightly built archosaurs, such as 450: 2804:
Whiteside, Jessica H.; Olsen, Paul E.; Eglinton, Timothy; Brookfield, Michael E.; Sambrotto, Raymond N. (22 March 2010).
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Butler, Richard J.; Brusatte, Stephen L.; Reich, Mike; Nesbitt, Sterling J.; Schoch, Rainer R.; Hornung, Jahn J. (2011).
2995: 5927: 5323: 609:, two enigmatic Triassic reptiles that bear little resemblance to archosaurs, have been regarded as pseudosuchians. 5904: 546:) that were superficially crocodilian-like, but were not actually crocodilian. Hence the name "false crocodiles". 555: 6787: 793:
changed the climate, causing a mass extinction that wiped out the dinosaurs' main competitors. This allowed the
3024: 636:. Avemetatarsalians are bird-line archosaurs, including pterosaurs and dinosaurs (the latter including birds). 5919: 457:, survived the major extinction. Crocodylomorphs themselves evolved a diverse array of lifestyles during the 372:, was not closely related to other traditional "crurotarsans", at least compared to "bird-line archosaurs" ( 2531:
from the latest Early Triassic of Germany and the timing and biogeography of the early archosaur radiation"
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were small, light-footed predators. A few groups acquired herbivorous diets, such as the heavily armored
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Senter, P. (2005). "Phylogenetic taxonomy and the names of the major archosaurian (Reptilia) clades".
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niches. Their heyday was the Late Triassic, during which time their ranks included erect-limbed
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into which all thecodonts that did not fit in the other three suborders could be placed. Even
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was originally given to a group of superficially crocodile-like prehistoric reptiles from the
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Gauthier, J. (1986). "Saurischian monophyly and the origin of birds". In Padian, K. (ed.).
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modified from Nesbitt (2011) showing the new changes (bold terminal taxa are collapsed).
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Brochu, C.A. (1997). "Synonymy, redundancy, and the name of the crocodile stem-group".
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period, but fell out of use in the late 20th century, especially after the name
2694: 2678: 2060: 1908: 1180: 1155: 1118: 619:(1985) and Gauthier (1986) became the first to establish the name Pseudosuchia in a 6630: 6499: 6482: 6259: 5431: 4833: 3780: 3767: 3701: 3586: 3492: 3378: 3339: 3298: 3258: 3207: 3190: 3177: 3140: 3040: 2929: 2886: 2837: 2827: 2790: 2786: 2722: 2714: 2664: 2654: 2560: 2550: 2289: 1945: 1475: 1278: 857: 683: 644: 612: 560: 513: 345: 3006:
archives, for comments critical of applying "Pseudosuchia" in a cladistic context.
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Many of these Triassic pseudosuchian groups went extinct at or before the
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to succeed them as the dominant terrestrial carnivores and herbivores.
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caused the extinction of all the pseudosuchians with the exception of
6723: 6653: 6051: 4955: 4790: 4623: 4487: 4252: 4229: 4077: 4019: 3936: 3607: 3559: 3385: 2954:. Memoirs California Academy of Sciences. Vol. 8. pp. 1–55. 2669: 847: 810: 755: 747: 735: 695: 521: 470: 466: 381: 369: 205: 162: 96: 40: 6851: 6874: 6619: 6269: 6179: 5980: 5559: 5196: 5144: 4775: 3064: 3009: 2595: 2358: 1416: 1329: 834: 801: 794: 759: 743: 739: 723: 703: 656: 590: 537: 517: 494: 458: 423: 389: 361: 350: 330: 182: 86: 81: 66: 61: 51: 33: 2659: 2640: 6909: 5844: 5530: 5242: 5115: 3079: 2035: 1212: 542: 192: 101: 76: 2865:
Foth, Christian; Sookias, Roland B.; Ezcurra, Martín D. (2021).
2803: 1692: 912: 698:. The skull is often massively built, especially in contrast to 694:
Pseudosuchia is one of the two primary "daughter" clades of the
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as true crocodilians are now defined as a subset of the group.
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definition: "all taxa the least inclusive clade containing
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The Dinosaur Book: The ruling reptiles and their relatives
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Origin, decline, and re-use of the clade name Pseudosuchia
2602:. Vol. 1. Oxford: Clarendon Press. pp. 295–338. 446:
were both bipedal and herbivorous, with toothless beaks.
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Colbert, Edwin Harris; Knight, Charles Robert (1951).
746:(late Middle Triassic) they dominated the terrestrial 6691: 6582: 6566: 6556: 6545: 6535: 6525: 6515: 6505: 6495: 6432: 6401: 6320: 6305: 6295: 6285: 6275: 6265: 6255: 6245: 6197: 6160: 6123: 6108: 6097: 6087: 6077: 6067: 6057: 6047: 6037: 6027: 6016: 6006: 5996: 5986: 5976: 5749: 5734: 5724: 5714: 5704: 5694: 5683: 5673: 5662: 5651: 5641: 5631: 5621: 5611: 5601: 5591: 5581: 5571: 5382: 5367: 5223: 5155: 5095: 5080: 5069: 5059: 5048: 5037: 4967: 4951: 4941: 4931: 4608: 4598: 4588: 4578: 4568: 4558: 4548: 4538: 4528: 4518: 4508: 4483: 4473: 4463: 4453: 4443: 4433: 4423: 4282: 4271: 4226: 4185: 4148: 4133: 4123: 4113: 4103: 4093: 4083: 4073: 3978: 3948: 3873: 3858: 3848: 3838: 3828: 3817: 3807: 3797: 3670: 3644: 3618: 3603: 3593: 3582: 3572: 3310: 3294: 3284: 3274: 3264: 3254: 3244: 3234: 3223: 3213: 3203: 2374: 2349: 2317: 2280: 2221: 2201: 2168: 2141: 2097: 2051: 2026: 1936: 1899: 1862: 1818: 1747: 1715: 1683: 1589:(crocodilian‑lineage of archosaurs)  1466: 1439: 1407: 1375: 1343: 1306: 1269: 1228: 1203: 1171: 1146: 1109: 1038: 999: 967: 935: 903: 871: 258: 249: 239: 821:are the surviving representatives of this lineage. 5352: 2864: 2646:Bulletin of the American Museum of Natural History 589:— both of which were at the time reconstructed as 360:(Laurenti, 1768)." However, a major 2011 study of 6937: 320:, "crocodile") is one of two major divisions of 3778: 3188: 2952:The Origin of Birds and the Evolution of Flight 2910:. International Archaeopteryx Conference 1984. 770:crocodilians, and a few other assorted groups. 5957: 2918: 2905: 414:. Ornithosuchids were large scavengers, while 4736: 3025: 2914:: Freunde des Jura-Museums. pp. 185–197. 2612: 2600:Phylogeny and Classification of the Tetrapods 509:, but it was not widely embraced until 2011. 4912: 2590: 4743: 4729: 3032: 3018: 340:Pseudosuchia is potentially equivalent to 112: 4805: 3112: 2890: 2841: 2831: 2726: 2668: 2658: 2619:. New York, NY: McGraw-Hill. p. 153. 2564: 2554: 392:period. They included giant, quadrupedal 2949: 734:Pseudosuchians appeared during the late 717: 639:A different definition was suggested by 536:in 1887–1890 to include three taxa (two 344:even though the latter has a different, 6951:Extant Early Triassic first appearances 5340: 2764: 2762: 2638: 2634: 2632: 2630: 2628: 2626: 6938: 2958: 2768: 2586: 2584: 569:, Pseudosuchia constitutes one of the 6856: 6855: 5944: 5339: 4899: 4762: 4724: 4405: 3765: 3175: 3051: 3013: 501:was established in 1990 to label the 3766: 3176: 3039: 2759: 2623: 549:In mid-20th century textbooks, like 532:The name Pseudosuchia was coined by 527: 426:, and several were bipedal, such as 5945: 2899: 2581: 632:of another branch-based clade, the 13: 4406: 2993:Re: Even more last papers for 2005 2771:Journal of Vertebrate Paleontology 451:Triassic–Jurassic extinction event 364:archosaur relations proposed that 14: 6962: 2981: 722:Mounted skeleton of the Triassic 6836: 6835: 6786: 6779: 6772: 5926: 5918: 5911: 5903: 5322: 5315: 5307: 4900: 4750: 4705: 4704: 4388: 4382: 4375: 3747: 3739: 3732: 2381: 2357: 2325: 2288: 2229: 2177: 2105: 2059: 2034: 1967: 1944: 1907: 1870: 1826: 1787: 1753: 1723: 1691: 1497: 1474: 1415: 1383: 1351: 1314: 1277: 1236: 1211: 1179: 1154: 1117: 1081: 1046: 1007: 975: 943: 911: 879: 149: 44: 2598:. In Benton, Michael J. (ed.). 2858: 2797: 2791:10.1080/02724634.1997.10010992 2699:Nature Ecology & Evolution 2685: 2606: 2518: 689: 1: 2511: 3002:and follow up posts, on the 2975:Stem Archosauria—TaxonSearch 2556:10.1371/journal.pone.0025693 1080:(bird-lineage of archosaurs) 827: 713: 566:Evolution of the Vertebrates 7: 2988:Taxon Search – Pseudosuchia 573:of the now-abandoned order 10: 6967: 2942: 2871:Frontiers in Earth Science 2594:; Clark, James M. (1988). 453:. However, one group, the 6864: 6800: 6769: 6722: 6690: 6681: 6629: 6593: 6581: 6491: 6480: 6431: 6400: 6319: 6241: 6237: 6226: 6196: 6159: 6122: 5972: 5968: 5953: 5940: 5900: 5870: 5748: 5567: 5558: 5520: 5493: 5430: 5393: 5381: 5363: 5348: 5335: 5304: 5274: 5222: 5195: 5166: 5154: 5094: 5033: 5022: 4966: 4927: 4923: 4908: 4895: 4859: 4841: 4832: 4814: 4801: 4758: 4700: 4651: 4622: 4497: 4416: 4412: 4401: 4372: 4325: 4296: 4260: 4251: 4225: 4214: 4184: 4147: 4069: 4065: 4054: 3977: 3947: 3872: 3793: 3789: 3774: 3761: 3730: 3700: 3669: 3643: 3617: 3568: 3557: 3438: 3410: 3338: 3322: 3308: 3199: 3184: 3171: 3139: 3121: 3108: 3047: 2934:10.1080/14772010903537732 2892:10.3389/feart.2021.723973 2719:10.1038/s41559-017-0305-5 2527:"The sail-backed reptile 2372: 2347: 2340: 2315: 2308: 2278: 2271: 2219: 2199: 2192: 2166: 2159: 2139: 2132: 2125: 2095: 2088: 2049: 2024: 2017: 2010: 1959: 1934: 1927: 1897: 1890: 1860: 1853: 1846: 1816: 1809: 1780: 1773: 1745: 1738: 1713: 1706: 1681: 1674: 1489: 1464: 1457: 1437: 1430: 1405: 1398: 1373: 1366: 1341: 1334: 1304: 1297: 1267: 1260: 1226: 1201: 1194: 1169: 1144: 1137: 1107: 1100: 1073: 1066: 1036: 1029: 997: 990: 965: 958: 933: 926: 901: 894: 869: 862: 807:end-Cretaceous extinction 314: 303: 281: 274: 236: 231: 146:Scientific classification 144: 120: 111: 23: 286:Benton & Clark, 1988 2908:The Beginnings of Birds 2833:10.1073/pnas.1001706107 775:end-Triassic extinction 623:context, using it as a 556:Vertebrate Paleontology 512:In 2011 paleontologist 5399:Benggwigwishingasuchus 2639:Nesbitt, S.J. (2011). 762:, the large predatory 731: 534:Karl Alfred von Zittel 6918:Paleobiology Database 3004:Dinosaur Mailing List 2998:12 April 2016 at the 2924:Stephen L. Brusatte; 721: 643:and Clark, 1988: the 551:Alfred Sherwood Romer 351:Rutiodon carolinensis 2973:Sereno, P. C. 2005. 649:last common ancestor 357:Crocodylus niloticus 354:(Emmons, 1856), and 5355:Paracrocodylomorpha 5342:Paracrocodylomorpha 5276:Paracrocodylomorpha 2883:2021FrEaS...9..763F 2824:2010PNAS..107.6721W 2783:1997JVPal..17..448B 2711:2017NatEE...1.1477E 2547:2011PLoSO...625693B 1886:Paracrocodylomorpha 1293:Paracrocodylomorpha 728:Saurosuchus galilei 324:, including living 6072:Pseudhesperosuchus 4674:Ichthyosauromorpha 4664:Drepanosauromorpha 2592:Benton, Michael J. 791:volcanic eruptions 766:, the small agile 732: 674:, Aetosauria, and 666:erected the clade 625:branch-based taxon 6933: 6932: 6858:Taxon identifiers 6849: 6848: 6796: 6795: 6767: 6766: 6763: 6762: 6759: 6758: 6755: 6754: 6751: 6750: 6677: 6676: 6586:Shartegosuchoidea 6550:Pachycheilosuchus 6476: 6475: 6222: 6221: 6062:Phyllodontosuchus 5936: 5935: 5898: 5897: 5894: 5893: 5890: 5889: 5554: 5553: 5331: 5330: 5302: 5301: 5298: 5297: 5294: 5293: 5218: 5217: 5018: 5017: 4891: 4890: 4887: 4886: 4883: 4882: 4879: 4878: 4718: 4717: 4696: 4695: 4692: 4691: 4397: 4396: 4370: 4369: 4366: 4365: 4362: 4361: 4358: 4357: 4354: 4353: 4210: 4209: 4050: 4049: 3757: 3756: 3728: 3727: 3724: 3723: 3720: 3719: 3553: 3552: 3543:Sharovipterygidae 3412:Sharovipterygidae 3356:Dinocephalosaurus 3167: 3166: 3163: 3162: 3159: 3158: 2926:Michael J. Benton 2705:(10): 1477–1483. 2508: 2507: 2499: 2498: 2490: 2489: 2481: 2480: 2472: 2471: 2458: 2457: 2449: 2448: 2440: 2439: 2431: 2430: 2422: 2421: 2413: 2412: 2404: 2403: 2395: 2394: 2260: 2259: 2251: 2250: 2242: 2241: 2072: 2071: 1999: 1998: 1990: 1989: 1981: 1980: 1751:Proterochampsidae 1656: 1655: 1647: 1646: 1638: 1637: 1629: 1628: 1620: 1619: 1611: 1610: 1602: 1601: 1593: 1592: 1583: 1582: 1574: 1573: 1565: 1564: 1556: 1555: 1547: 1546: 1538: 1537: 1529: 1528: 1520: 1519: 1511: 1510: 1249: 1248: 595:wastebasket taxon 528:Taxonomic history 374:Avemetatarsalians 293: 292: 287: 227: 16:Clade of reptiles 6958: 6926: 6925: 6913: 6912: 6900: 6899: 6898: 6885: 6884: 6883: 6853: 6852: 6839: 6838: 6791: 6790: 6783: 6777: 6776: 6693: 6688: 6687: 6631:Shartegosuchidae 6591: 6590: 6584: 6568: 6558: 6547: 6537: 6527: 6517: 6507: 6500:Dianchungosaurus 6497: 6489: 6488: 6483:Mesoeucrocodylia 6434: 6403: 6322: 6307: 6297: 6287: 6277: 6267: 6260:Eopneumatosuchus 6257: 6247: 6239: 6238: 6235: 6234: 6199: 6162: 6125: 6110: 6099: 6089: 6079: 6069: 6059: 6049: 6039: 6029: 6018: 6008: 5998: 5988: 5978: 5970: 5969: 5966: 5965: 5955: 5954: 5942: 5941: 5931: 5930: 5923: 5922: 5916: 5915: 5908: 5907: 5751: 5736: 5726: 5716: 5706: 5696: 5685: 5675: 5664: 5653: 5643: 5633: 5623: 5613: 5603: 5593: 5583: 5573: 5565: 5564: 5432:Ctenosauriscidae 5391: 5390: 5384: 5369: 5361: 5360: 5350: 5349: 5337: 5336: 5326: 5320: 5319: 5312: 5311: 5225: 5164: 5163: 5157: 5097: 5082: 5071: 5061: 5050: 5039: 5031: 5030: 4969: 4953: 4943: 4933: 4925: 4924: 4921: 4920: 4910: 4909: 4897: 4896: 4849:Archosauromorpha 4839: 4838: 4834:Archosauromorpha 4812: 4811: 4803: 4802: 4796: 4795: 4760: 4759: 4745: 4738: 4731: 4722: 4721: 4708: 4707: 4652:Possible members 4610: 4600: 4590: 4580: 4570: 4560: 4550: 4540: 4530: 4520: 4510: 4485: 4475: 4465: 4455: 4445: 4435: 4425: 4414: 4413: 4403: 4402: 4392: 4386: 4380: 4379: 4284: 4273: 4258: 4257: 4228: 4223: 4222: 4187: 4150: 4135: 4125: 4115: 4105: 4095: 4085: 4075: 4067: 4066: 4063: 4062: 3980: 3950: 3875: 3860: 3850: 3840: 3830: 3819: 3809: 3799: 3791: 3790: 3787: 3786: 3781:Archosauriformes 3776: 3775: 3768:Archosauriformes 3763: 3762: 3752: 3751: 3744: 3743: 3736: 3702:Archosauriformes 3672: 3646: 3620: 3605: 3595: 3587:Cryptovaranoides 3584: 3574: 3566: 3565: 3493:Langobardisaurus 3379:Protanystropheus 3340:Trachelosauridae 3320: 3319: 3312: 3299:Vritramimosaurus 3296: 3286: 3276: 3266: 3259:Mecistotrachelos 3256: 3246: 3236: 3225: 3215: 3208:Aenigmastropheus 3205: 3197: 3196: 3191:Archosauromorpha 3186: 3185: 3178:Archosauromorpha 3173: 3172: 3141:Archosauromorpha 3119: 3118: 3110: 3109: 3103: 3102: 3049: 3048: 3041:Archosauromorpha 3034: 3027: 3020: 3011: 3010: 2970: 2955: 2936: 2922: 2916: 2915: 2903: 2897: 2896: 2894: 2862: 2856: 2855: 2845: 2835: 2801: 2795: 2794: 2766: 2757: 2756: 2730: 2689: 2683: 2682: 2672: 2662: 2636: 2621: 2620: 2610: 2604: 2603: 2588: 2579: 2578: 2568: 2558: 2522: 2385: 2376: 2361: 2351: 2343: 2342: 2329: 2319: 2311: 2310: 2292: 2282: 2274: 2273: 2233: 2223: 2203: 2195: 2194: 2181: 2170: 2162: 2161: 2143: 2135: 2134: 2128: 2127: 2109: 2099: 2091: 2090: 2063: 2053: 2038: 2028: 2020: 2019: 2013: 2012: 1972: 1971: 1948: 1938: 1930: 1929: 1911: 1901: 1893: 1892: 1874: 1864: 1856: 1855: 1849: 1848: 1830: 1820: 1812: 1811: 1791: 1776: 1775: 1757: 1749: 1741: 1740: 1727: 1717: 1709: 1708: 1695: 1685: 1677: 1676: 1670: 1669: 1502: 1501: 1478: 1468: 1460: 1459: 1441: 1433: 1432: 1419: 1409: 1401: 1400: 1387: 1377: 1369: 1368: 1355: 1345: 1337: 1336: 1318: 1308: 1300: 1299: 1281: 1271: 1263: 1262: 1240: 1230: 1215: 1205: 1197: 1196: 1183: 1173: 1158: 1148: 1140: 1139: 1121: 1111: 1103: 1102: 1085: 1069: 1068: 1050: 1040: 1032: 1031: 1011: 1001: 993: 992: 979: 969: 961: 960: 947: 937: 929: 928: 915: 905: 897: 896: 883: 873: 865: 864: 858:Archosauriformes 854: 853: 684:Sterling Nesbitt 645:node-based taxon 561:Edwin H. Colbert 514:Sterling Nesbitt 319: 317: 308: 306: 285: 260: 251: 241: 222: 154: 153: 116: 106: 43: 27:Temporal range: 21: 20: 6966: 6965: 6961: 6960: 6959: 6957: 6956: 6955: 6936: 6935: 6934: 6929: 6921: 6916: 6908: 6903: 6894: 6893: 6888: 6879: 6878: 6873: 6860: 6850: 6845: 6792: 6785: 6771: 6747: 6718: 6673: 6625: 6606:Shantungosuchus 6577: 6485: 6472: 6427: 6396: 6361:Hemiprotosuchus 6315: 6231: 6229:Crocodyliformes 6218: 6192: 6155: 6150:Terrestrisuchus 6127:Saltoposuchidae 6118: 6102:Syntomiprosopus 5962: 5960:Crocodylomorpha 5949: 5947:Crocodylomorpha 5932: 5925: 5917: 5910: 5902: 5886: 5872:Crocodylomorpha 5866: 5860:Vytshegdosuchus 5852:Vjushkovisaurus 5744: 5550: 5516: 5504:Dolichobrachium 5489: 5426: 5377: 5357: 5344: 5327: 5314: 5306: 5290: 5270: 5227:Gracilisuchidae 5214: 5191: 5159:Aetosauriformes 5150: 5090: 5027: 5014: 4971:Ornithosuchidae 4962: 4917: 4904: 4875: 4855: 4828: 4797: 4765: 4764: 4754: 4749: 4719: 4714: 4688: 4669:Kuehneosauridae 4647: 4618: 4603:Vjushkovisaurus 4493: 4468:Protecovasaurus 4408: 4393: 4374: 4350: 4321: 4309:Avemetatarsalia 4298:Avemetatarsalia 4292: 4247: 4219: 4206: 4200:Proterochampsia 4189:Proterochampsia 4180: 4143: 4059: 4046: 3991:Bharitalasuchus 3982:Erythrosuchidae 3973: 3943: 3877:Proterosuchidae 3868: 3863:Syntomiprosopus 3783: 3770: 3753: 3746: 3738: 3716: 3696: 3665: 3639: 3613: 3562: 3549: 3521:Sclerostropheus 3456:Augustaburiania 3440:Tanystropheidae 3434: 3406: 3393:Sclerostropheus 3334: 3316: 3304: 3228:Elachistosuchus 3193: 3180: 3155: 3135: 3104: 3054: 3053: 3043: 3038: 3000:Wayback Machine 2984: 2979: 2945: 2940: 2939: 2923: 2919: 2904: 2900: 2863: 2859: 2802: 2798: 2767: 2760: 2690: 2686: 2637: 2624: 2611: 2607: 2589: 2582: 2523: 2519: 2514: 2509: 2500: 2491: 2482: 2473: 2459: 2450: 2441: 2432: 2423: 2414: 2405: 2396: 2261: 2252: 2243: 2073: 2056:Ornithosuchidae 2000: 1991: 1982: 1966: 1963:Crocodylomorpha 1784:Avemetatarsalia 1657: 1648: 1639: 1630: 1621: 1612: 1603: 1594: 1584: 1575: 1566: 1557: 1548: 1539: 1530: 1521: 1512: 1496: 1493:Crocodylomorpha 1250: 1114:Ornithosuchidae 1077:Avemetatarsalia 972:Proterochampsia 876:Proterosuchidae 830: 787:Crocodylomorpha 783:Crocodyliformes 716: 692: 676:Crocodylomorpha 634:Avemetatarsalia 530: 487: 455:crocodylomorphs 438:. The bizarre, 315: 304: 284:Crocodylotarsi 262:Ornithosuchidae 221: 148: 107: 105: 104: 99: 94: 89: 84: 79: 74: 69: 64: 59: 54: 49: 38: 37: 28: 25: 17: 12: 11: 5: 6964: 6954: 6953: 6948: 6946:Pseudosuchians 6931: 6930: 6928: 6927: 6914: 6901: 6886: 6870: 6868: 6862: 6861: 6847: 6846: 6844: 6843: 6833: 6828: 6823: 6818: 6813: 6811:Prestosuchidae 6808: 6801: 6798: 6797: 6794: 6793: 6770: 6768: 6765: 6764: 6761: 6760: 6757: 6756: 6753: 6752: 6749: 6748: 6746: 6745: 6744: 6743: 6737: 6728: 6726: 6720: 6719: 6717: 6716: 6715: 6714: 6708: 6699: 6697: 6685: 6679: 6678: 6675: 6674: 6672: 6671: 6668:Shartegosuchus 6664: 6657: 6650: 6643: 6635: 6633: 6627: 6626: 6624: 6623: 6616: 6613:Sichuanosuchus 6609: 6602: 6594: 6588: 6579: 6578: 6576: 6575: 6571:Thalattosuchia 6564: 6554: 6543: 6533: 6530:Lusitanisuchus 6523: 6513: 6503: 6492: 6486: 6481: 6478: 6477: 6474: 6473: 6471: 6470: 6463: 6455: 6448: 6440: 6438: 6429: 6428: 6426: 6425: 6417: 6409: 6407: 6405:Notochampsidae 6398: 6397: 6395: 6394: 6387: 6383:Pedeticosaurus 6379: 6371: 6364: 6357: 6354:Erythrochampsa 6350: 6343: 6336: 6333:Coloradisuchus 6328: 6326: 6317: 6316: 6314: 6313: 6303: 6293: 6283: 6273: 6263: 6253: 6242: 6232: 6227: 6224: 6223: 6220: 6219: 6217: 6216: 6215: 6214: 6212:Thalattosuchia 6205: 6203: 6201:Thalattosuchia 6194: 6193: 6191: 6190: 6183: 6176: 6168: 6166: 6157: 6156: 6154: 6153: 6146: 6139: 6131: 6129: 6120: 6119: 6117: 6116: 6106: 6095: 6085: 6082:Redondavenator 6075: 6065: 6055: 6045: 6035: 6025: 6014: 6004: 5994: 5991:Dibothrosuchus 5984: 5973: 5963: 5958: 5951: 5950: 5938: 5937: 5934: 5933: 5901: 5899: 5896: 5895: 5892: 5891: 5888: 5887: 5885: 5884: 5883: 5882: 5876: 5874: 5868: 5867: 5865: 5864: 5856: 5848: 5841: 5833: 5826: 5818: 5811: 5804: 5796: 5789: 5782: 5774: 5766: 5757: 5755: 5746: 5745: 5743: 5742: 5732: 5722: 5712: 5702: 5692: 5681: 5671: 5667:Incertovenator 5660: 5649: 5639: 5629: 5619: 5609: 5599: 5589: 5579: 5568: 5562: 5556: 5555: 5552: 5551: 5549: 5548: 5541: 5534: 5526: 5524: 5518: 5517: 5515: 5514: 5507: 5499: 5497: 5491: 5490: 5488: 5487: 5480: 5472: 5469:Hypselorhachis 5465: 5462:Ctenosauriscus 5458: 5455:Bystrowisuchus 5451: 5444: 5436: 5434: 5428: 5427: 5425: 5424: 5417: 5409: 5402: 5394: 5388: 5379: 5378: 5376: 5375: 5364: 5358: 5353: 5346: 5345: 5333: 5332: 5329: 5328: 5305: 5303: 5300: 5299: 5296: 5295: 5292: 5291: 5289: 5288: 5287: 5286: 5280: 5278: 5272: 5271: 5269: 5268: 5261: 5254: 5247: 5239: 5231: 5229: 5220: 5219: 5216: 5215: 5213: 5212: 5211: 5210: 5201: 5199: 5193: 5192: 5190: 5189: 5186:Revueltosaurus 5182: 5175: 5167: 5161: 5152: 5151: 5149: 5148: 5141: 5134: 5127: 5120: 5112: 5103: 5101: 5099:Erpetosuchidae 5092: 5091: 5089: 5088: 5078: 5067: 5057: 5046: 5034: 5028: 5023: 5020: 5019: 5016: 5015: 5013: 5012: 5009:Venaticosuchus 5005: 4998: 4991: 4984: 4975: 4973: 4964: 4963: 4961: 4960: 4949: 4939: 4936:Dasygnathoides 4928: 4918: 4913: 4906: 4905: 4893: 4892: 4889: 4888: 4885: 4884: 4881: 4880: 4877: 4876: 4874: 4873: 4872: 4871: 4865: 4863: 4857: 4856: 4854: 4853: 4852: 4851: 4842: 4836: 4830: 4829: 4827: 4826: 4825: 4824: 4815: 4809: 4799: 4798: 4794: 4793: 4784: 4778: 4772: 4756: 4755: 4748: 4747: 4740: 4733: 4725: 4716: 4715: 4713: 4712: 4701: 4698: 4697: 4694: 4693: 4690: 4689: 4687: 4686: 4684:Thalattosauria 4681: 4676: 4671: 4666: 4661: 4655: 4653: 4649: 4648: 4646: 4645: 4640: 4635: 4629: 4627: 4620: 4619: 4617: 4616: 4606: 4596: 4586: 4576: 4566: 4556: 4546: 4536: 4526: 4516: 4505: 4503: 4495: 4494: 4492: 4491: 4481: 4471: 4461: 4451: 4441: 4431: 4420: 4418: 4410: 4409: 4407:Related topics 4399: 4398: 4395: 4394: 4373: 4371: 4368: 4367: 4364: 4363: 4360: 4359: 4356: 4355: 4352: 4351: 4349: 4348: 4347: 4346: 4340: 4331: 4329: 4323: 4322: 4320: 4319: 4318: 4317: 4311: 4302: 4300: 4294: 4293: 4291: 4290: 4280: 4268: 4266: 4263:Incertae sedis 4255: 4249: 4248: 4246: 4245: 4244: 4243: 4234: 4232: 4220: 4215: 4212: 4211: 4208: 4207: 4205: 4204: 4203: 4202: 4193: 4191: 4182: 4181: 4179: 4178: 4171: 4168:Halazhaisuchus 4164: 4156: 4154: 4145: 4144: 4142: 4141: 4131: 4121: 4118:Marcianosuchus 4111: 4101: 4091: 4081: 4070: 4060: 4055: 4052: 4051: 4048: 4047: 4045: 4044: 4037: 4030: 4027:Guchengosuchus 4023: 4016: 4008: 4001: 3994: 3986: 3984: 3975: 3974: 3972: 3971: 3964: 3956: 3954: 3945: 3944: 3942: 3941: 3933: 3925: 3918: 3911: 3904: 3897: 3893:Chasmatosuchus 3889: 3881: 3879: 3870: 3869: 3867: 3866: 3856: 3846: 3836: 3826: 3815: 3805: 3794: 3784: 3779: 3772: 3771: 3759: 3758: 3755: 3754: 3731: 3729: 3726: 3725: 3722: 3721: 3718: 3717: 3715: 3714: 3713: 3712: 3706: 3704: 3698: 3697: 3695: 3694: 3687: 3679: 3677: 3667: 3666: 3664: 3663: 3662: 3661: 3652: 3650: 3641: 3640: 3638: 3637: 3636: 3635: 3633:Allokotosauria 3626: 3624: 3622:Allokotosauria 3615: 3614: 3612: 3611: 3601: 3598:Tasmaniosaurus 3591: 3580: 3569: 3563: 3558: 3555: 3554: 3551: 3550: 3548: 3547: 3539: 3532: 3525: 3517: 3510: 3503: 3496: 3489: 3486:Gwyneddosaurus 3482: 3474: 3466: 3459: 3452: 3444: 3442: 3436: 3435: 3433: 3432: 3425: 3417: 3415: 3408: 3407: 3405: 3404: 3401:Trachelosaurus 3397: 3389: 3382: 3375: 3367: 3359: 3352: 3344: 3342: 3336: 3335: 3333: 3332: 3323: 3317: 3309: 3306: 3305: 3303: 3302: 3292: 3282: 3272: 3269:Prolacertoides 3262: 3252: 3249:Malutinisuchus 3242: 3232: 3221: 3211: 3200: 3194: 3189: 3182: 3181: 3169: 3168: 3165: 3164: 3161: 3160: 3157: 3156: 3154: 3153: 3152: 3151: 3145: 3143: 3137: 3136: 3134: 3133: 3132: 3131: 3122: 3116: 3106: 3105: 3101: 3100: 3091: 3082: 3073: 3067: 3061: 3045: 3044: 3037: 3036: 3029: 3022: 3014: 3008: 3007: 2990: 2983: 2982:External links 2980: 2978: 2977: 2971: 2956: 2946: 2944: 2941: 2938: 2937: 2917: 2898: 2857: 2818:(15): 6721–5. 2796: 2777:(2): 448–449. 2758: 2684: 2622: 2605: 2580: 2541:(10): e25693. 2529:Ctenosauriscus 2516: 2515: 2513: 2510: 2506: 2505: 2502: 2501: 2497: 2496: 2493: 2492: 2488: 2487: 2484: 2483: 2479: 2478: 2475: 2474: 2470: 2469: 2461: 2460: 2456: 2455: 2452: 2451: 2447: 2446: 2443: 2442: 2438: 2437: 2434: 2433: 2429: 2428: 2425: 2424: 2420: 2419: 2416: 2415: 2411: 2410: 2407: 2406: 2402: 2401: 2398: 2397: 2393: 2392: 2389: 2388: 2371: 2368: 2367: 2364: 2363: 2346: 2341: 2339: 2336: 2335: 2332: 2331: 2314: 2309: 2307: 2304:Prestosuchidae 2299: 2298: 2295: 2294: 2277: 2272: 2270: 2267: 2266: 2263: 2262: 2258: 2257: 2254: 2253: 2249: 2248: 2245: 2244: 2240: 2239: 2236: 2235: 2218: 2215: 2214: 2211: 2210: 2198: 2193: 2191: 2188: 2187: 2184: 2183: 2165: 2160: 2158: 2155: 2154: 2151: 2150: 2138: 2133: 2131: 2126: 2124: 2116: 2115: 2112: 2111: 2094: 2089: 2087: 2079: 2078: 2075: 2074: 2070: 2069: 2066: 2065: 2048: 2045: 2044: 2041: 2040: 2031:Revueltosaurus 2023: 2018: 2016: 2011: 2009: 2006: 2005: 2002: 2001: 1997: 1996: 1993: 1992: 1988: 1987: 1984: 1983: 1979: 1978: 1975: 1974: 1958: 1955: 1954: 1951: 1950: 1933: 1928: 1926: 1918: 1917: 1914: 1913: 1896: 1891: 1889: 1881: 1880: 1877: 1876: 1859: 1854: 1852: 1847: 1845: 1837: 1836: 1833: 1832: 1815: 1810: 1808: 1798: 1797: 1794: 1793: 1779: 1774: 1772: 1764: 1763: 1760: 1759: 1744: 1739: 1737: 1734: 1733: 1730: 1729: 1712: 1707: 1705: 1702: 1701: 1698: 1697: 1680: 1675: 1673: 1668: 1654: 1653: 1650: 1649: 1645: 1644: 1641: 1640: 1636: 1635: 1632: 1631: 1627: 1626: 1623: 1622: 1618: 1617: 1614: 1613: 1609: 1608: 1605: 1604: 1600: 1599: 1596: 1595: 1591: 1590: 1586: 1585: 1581: 1580: 1577: 1576: 1572: 1571: 1568: 1567: 1563: 1562: 1559: 1558: 1554: 1553: 1550: 1549: 1545: 1544: 1541: 1540: 1536: 1535: 1532: 1531: 1527: 1526: 1523: 1522: 1518: 1517: 1514: 1513: 1509: 1508: 1505: 1504: 1488: 1485: 1484: 1481: 1480: 1463: 1458: 1456: 1453: 1452: 1449: 1448: 1436: 1431: 1429: 1426: 1425: 1422: 1421: 1404: 1399: 1397: 1394: 1393: 1390: 1389: 1372: 1367: 1365: 1362: 1361: 1358: 1357: 1340: 1335: 1333: 1325: 1324: 1321: 1320: 1303: 1298: 1296: 1288: 1287: 1284: 1283: 1266: 1261: 1259: 1256: 1255: 1252: 1251: 1247: 1246: 1243: 1242: 1225: 1222: 1221: 1218: 1217: 1208:Revueltosaurus 1200: 1195: 1193: 1190: 1189: 1186: 1185: 1168: 1165: 1164: 1161: 1160: 1143: 1138: 1136: 1128: 1127: 1124: 1123: 1106: 1101: 1099: 1092: 1091: 1088: 1087: 1072: 1067: 1065: 1057: 1056: 1053: 1052: 1035: 1030: 1028: 1018: 1017: 1014: 1013: 996: 991: 989: 986: 985: 982: 981: 964: 959: 957: 954: 953: 950: 949: 932: 927: 925: 922: 921: 918: 917: 900: 895: 893: 890: 889: 886: 885: 868: 863: 861: 852: 829: 826: 715: 712: 691: 688: 647:including the 529: 526: 486: 483: 420:gracilisuchids 394:apex predators 309:, "false" and 291: 290: 289: 288: 279: 278: 272: 271: 270: 269: 264: 256: 247: 244:Dasygnathoides 234: 233: 229: 228: 216: 209: 208: 203: 196: 195: 190: 186: 185: 180: 176: 175: 170: 166: 165: 160: 156: 155: 142: 141: 118: 117: 109: 108: 100: 95: 90: 85: 80: 75: 70: 65: 60: 55: 50: 45: 30:Early Triassic 26: 24:Pseudosuchians 15: 9: 6: 4: 3: 2: 6963: 6952: 6949: 6947: 6944: 6943: 6941: 6924: 6919: 6915: 6911: 6906: 6902: 6897: 6891: 6887: 6882: 6876: 6872: 6871: 6869: 6867: 6863: 6859: 6854: 6842: 6834: 6832: 6829: 6827: 6824: 6822: 6819: 6817: 6814: 6812: 6809: 6807: 6803: 6802: 6799: 6789: 6784: 6782: 6775: 6742: 6738: 6736: 6732: 6731: 6730: 6729: 6727: 6725: 6721: 6713: 6709: 6707: 6703: 6702: 6701: 6700: 6698: 6696: 6689: 6686: 6684: 6680: 6670: 6669: 6665: 6663: 6662: 6658: 6656: 6655: 6651: 6649: 6648: 6647:Fruitachampsa 6644: 6642: 6641: 6637: 6636: 6634: 6632: 6628: 6622: 6621: 6617: 6615: 6614: 6610: 6608: 6607: 6603: 6601: 6600: 6596: 6595: 6592: 6589: 6587: 6580: 6573: 6572: 6565: 6563: 6562: 6561:Titanochampsa 6555: 6552: 6551: 6544: 6542: 6541: 6540:Neuquensuchus 6534: 6532: 6531: 6524: 6522: 6521: 6514: 6512: 6511: 6504: 6502: 6501: 6494: 6493: 6490: 6487: 6484: 6479: 6469: 6468: 6464: 6461: 6460: 6456: 6454: 6453: 6449: 6447: 6446: 6442: 6441: 6439: 6437: 6436:Gobiosuchidae 6430: 6423: 6422: 6418: 6416: 6415: 6411: 6410: 6408: 6406: 6399: 6393: 6392: 6388: 6385: 6384: 6380: 6377: 6376: 6372: 6370: 6369: 6365: 6363: 6362: 6358: 6356: 6355: 6351: 6349: 6348: 6344: 6342: 6341: 6337: 6335: 6334: 6330: 6329: 6327: 6325: 6324:Protosuchidae 6318: 6312: 6311: 6304: 6302: 6301: 6300:Stegomosuchus 6294: 6292: 6291: 6290:Platyognathus 6284: 6282: 6281: 6274: 6272: 6271: 6264: 6262: 6261: 6254: 6252: 6251: 6250:Entradasuchus 6244: 6243: 6240: 6236: 6233: 6230: 6225: 6213: 6209: 6208: 6207: 6206: 6204: 6202: 6195: 6189: 6188: 6187:Macelognathus 6184: 6182: 6181: 6177: 6175: 6174: 6170: 6169: 6167: 6165: 6158: 6152: 6151: 6147: 6145: 6144: 6143:Saltoposuchus 6140: 6138: 6137: 6136:Litargosuchus 6133: 6132: 6130: 6128: 6121: 6115: 6114: 6107: 6104: 6103: 6096: 6094: 6093: 6086: 6084: 6083: 6076: 6074: 6073: 6066: 6064: 6063: 6056: 6054: 6053: 6046: 6044: 6043: 6042:Kayentasuchus 6036: 6034: 6033: 6032:Junggarsuchus 6026: 6023: 6022: 6015: 6013: 6012: 6011:Hesperosuchus 6005: 6003: 6002: 6001:Dromicosuchus 5995: 5993: 5992: 5985: 5983: 5982: 5975: 5974: 5971: 5967: 5964: 5961: 5956: 5952: 5948: 5943: 5939: 5929: 5924: 5921: 5914: 5906: 5880: 5879: 5878: 5877: 5875: 5873: 5869: 5862: 5861: 5857: 5854: 5853: 5849: 5847: 5846: 5842: 5839: 5838: 5834: 5832: 5831: 5827: 5824: 5823: 5819: 5817: 5816: 5815:Scolotosuchus 5812: 5810: 5809: 5805: 5802: 5801: 5800:Procerosuchus 5797: 5795: 5794: 5790: 5788: 5787: 5783: 5780: 5779: 5775: 5772: 5771: 5767: 5764: 5763: 5759: 5758: 5756: 5754: 5747: 5741: 5740: 5733: 5731: 5730: 5729:Stagonosuchus 5723: 5721: 5720: 5713: 5711: 5710: 5709:Procerosuchus 5703: 5701: 5700: 5693: 5690: 5689: 5682: 5680: 5679: 5672: 5669: 5668: 5661: 5658: 5657: 5650: 5648: 5647: 5640: 5638: 5637: 5630: 5628: 5627: 5620: 5618: 5617: 5616:Decuriasuchus 5610: 5608: 5607: 5600: 5598: 5597: 5596:Batrachotomus 5590: 5588: 5587: 5580: 5578: 5577: 5570: 5569: 5566: 5563: 5561: 5557: 5547: 5546: 5542: 5540: 5539: 5535: 5533: 5532: 5528: 5527: 5525: 5523: 5522:Shuvosauridae 5519: 5513: 5512: 5508: 5506: 5505: 5501: 5500: 5498: 5496: 5492: 5486: 5485: 5481: 5478: 5477: 5473: 5471: 5470: 5466: 5464: 5463: 5459: 5457: 5456: 5452: 5450: 5449: 5445: 5443: 5442: 5441:Arizonasaurus 5438: 5437: 5435: 5433: 5429: 5423: 5422: 5418: 5415: 5414: 5413:Schultzsuchus 5410: 5408: 5407: 5403: 5401: 5400: 5396: 5395: 5392: 5389: 5387: 5386:Poposauroidea 5380: 5374: 5373: 5366: 5365: 5362: 5359: 5356: 5351: 5347: 5343: 5338: 5334: 5325: 5321: 5318: 5310: 5284: 5283: 5282: 5281: 5279: 5277: 5273: 5267: 5266: 5262: 5260: 5259: 5258:Turfanosuchus 5255: 5253: 5252: 5248: 5245: 5244: 5240: 5238: 5237: 5236:Gracilisuchus 5233: 5232: 5230: 5228: 5221: 5209: 5205: 5204: 5203: 5202: 5200: 5198: 5194: 5188: 5187: 5183: 5181: 5180: 5179:Euscolosuchus 5176: 5174: 5173: 5169: 5168: 5165: 5162: 5160: 5153: 5147: 5146: 5142: 5140: 5139: 5135: 5133: 5132: 5128: 5126: 5125: 5121: 5118: 5117: 5113: 5110: 5109: 5105: 5104: 5102: 5100: 5093: 5087: 5086: 5079: 5076: 5075: 5068: 5066: 5065: 5058: 5055: 5054: 5053:Koilamasuchus 5047: 5044: 5043: 5036: 5035: 5032: 5029: 5026: 5021: 5011: 5010: 5006: 5004: 5003: 4999: 4997: 4996: 4995:Ornithosuchus 4992: 4990: 4989: 4985: 4982: 4981: 4977: 4976: 4974: 4972: 4965: 4958: 4957: 4950: 4948: 4947: 4940: 4938: 4937: 4930: 4929: 4926: 4922: 4919: 4916: 4911: 4907: 4903: 4898: 4894: 4869: 4868: 4867: 4866: 4864: 4862: 4858: 4850: 4846: 4845: 4844: 4843: 4840: 4837: 4835: 4831: 4823: 4819: 4818: 4817: 4816: 4813: 4810: 4808: 4804: 4800: 4792: 4788: 4785: 4783: 4779: 4777: 4773: 4771: 4767: 4766: 4761: 4757: 4753: 4746: 4741: 4739: 4734: 4732: 4727: 4726: 4723: 4711: 4703: 4702: 4699: 4685: 4682: 4680: 4679:Sauropterygia 4677: 4675: 4672: 4670: 4667: 4665: 4662: 4660: 4657: 4656: 4654: 4650: 4644: 4641: 4639: 4638:Protorosauria 4636: 4634: 4633:Proterosuchia 4631: 4630: 4628: 4625: 4621: 4615: 4614: 4607: 4605: 4604: 4597: 4595: 4594: 4587: 4585: 4584: 4577: 4575: 4574: 4567: 4565: 4564: 4557: 4555: 4554: 4547: 4545: 4544: 4537: 4535: 4534: 4527: 4525: 4524: 4517: 4515: 4514: 4507: 4506: 4504: 4502: 4501: 4496: 4490: 4489: 4482: 4480: 4479: 4472: 4470: 4469: 4462: 4460: 4459: 4452: 4450: 4449: 4442: 4440: 4439: 4438:Lucianosaurus 4432: 4430: 4429: 4422: 4421: 4419: 4415: 4411: 4404: 4400: 4391: 4387: 4385: 4378: 4345: 4341: 4339: 4335: 4334: 4333: 4332: 4330: 4328: 4324: 4316: 4312: 4310: 4306: 4305: 4304: 4303: 4301: 4299: 4295: 4289: 4288: 4281: 4278: 4277: 4276:Sikannisuchus 4270: 4269: 4267: 4265: 4264: 4259: 4256: 4254: 4250: 4242: 4238: 4237: 4236: 4235: 4233: 4231: 4224: 4221: 4218: 4213: 4201: 4197: 4196: 4195: 4194: 4192: 4190: 4183: 4177: 4176: 4172: 4170: 4169: 4165: 4163: 4162: 4158: 4157: 4155: 4153: 4152:Euparkeriidae 4146: 4140: 4139: 4132: 4130: 4129: 4128:Polymorphodon 4122: 4120: 4119: 4112: 4110: 4109: 4102: 4100: 4099: 4092: 4090: 4089: 4082: 4080: 4079: 4072: 4071: 4068: 4064: 4061: 4058: 4053: 4043: 4042: 4038: 4036: 4035: 4031: 4029: 4028: 4024: 4022: 4021: 4017: 4014: 4013: 4009: 4007: 4006: 4005:Erythrosuchus 4002: 4000: 3999: 3995: 3993: 3992: 3988: 3987: 3985: 3983: 3976: 3970: 3969: 3965: 3963: 3962: 3958: 3957: 3955: 3953: 3952:Protopyknosia 3946: 3939: 3938: 3934: 3931: 3930: 3926: 3924: 3923: 3922:Samsarasuchus 3919: 3917: 3916: 3915:Proterosuchus 3912: 3910: 3909: 3905: 3903: 3902: 3898: 3895: 3894: 3890: 3888: 3887: 3883: 3882: 3880: 3878: 3871: 3865: 3864: 3857: 3855: 3854: 3853:Sarmatosuchus 3847: 3845: 3844: 3843:Koilamasuchus 3837: 3835: 3834: 3827: 3824: 3823: 3816: 3814: 3813: 3806: 3804: 3803: 3796: 3795: 3792: 3788: 3785: 3782: 3777: 3773: 3769: 3764: 3760: 3750: 3745: 3742: 3735: 3710: 3709: 3708: 3707: 3705: 3703: 3699: 3693: 3692: 3688: 3686: 3685: 3681: 3680: 3678: 3675: 3674:Prolacertidae 3668: 3660: 3659:Rhynchosauria 3656: 3655: 3654: 3653: 3651: 3649: 3648:Rhynchosauria 3642: 3634: 3630: 3629: 3628: 3627: 3625: 3623: 3616: 3610: 3609: 3602: 3600: 3599: 3592: 3589: 3588: 3581: 3579: 3578: 3571: 3570: 3567: 3564: 3561: 3556: 3545: 3544: 3540: 3538: 3537: 3536:Tanytrachelos 3533: 3531: 3530: 3529:Tanystropheus 3526: 3523: 3522: 3518: 3516: 3515: 3511: 3509: 3508: 3504: 3502: 3501: 3497: 3495: 3494: 3490: 3488: 3487: 3483: 3480: 3479: 3478:Gracilicollum 3475: 3472: 3471: 3467: 3465: 3464: 3460: 3458: 3457: 3453: 3451: 3450: 3446: 3445: 3443: 3441: 3437: 3431: 3430: 3429:Sharovipteryx 3426: 3424: 3423: 3419: 3418: 3416: 3413: 3409: 3403: 3402: 3398: 3395: 3394: 3390: 3388: 3387: 3383: 3381: 3380: 3376: 3373: 3372: 3371:Gracilicollum 3368: 3365: 3364: 3360: 3358: 3357: 3353: 3351: 3350: 3346: 3345: 3343: 3341: 3337: 3330: 3329: 3328:Jesairosaurus 3325: 3324: 3321: 3318: 3315: 3307: 3301: 3300: 3293: 3291: 3290: 3283: 3281: 3280: 3279:Protorosaurus 3273: 3271: 3270: 3263: 3261: 3260: 3253: 3251: 3250: 3243: 3241: 3240: 3233: 3230: 3229: 3222: 3220: 3219: 3212: 3210: 3209: 3202: 3201: 3198: 3195: 3192: 3187: 3183: 3179: 3174: 3170: 3149: 3148: 3147: 3146: 3144: 3142: 3138: 3130: 3126: 3125: 3124: 3123: 3120: 3117: 3115: 3111: 3107: 3099: 3098:Archelosauria 3095: 3092: 3090: 3086: 3083: 3081: 3077: 3074: 3072: 3068: 3066: 3062: 3060: 3056: 3055: 3050: 3046: 3042: 3035: 3030: 3028: 3023: 3021: 3016: 3015: 3012: 3005: 3001: 2997: 2994: 2991: 2989: 2986: 2985: 2976: 2972: 2968: 2964: 2963: 2957: 2953: 2948: 2947: 2935: 2931: 2927: 2921: 2913: 2909: 2902: 2893: 2888: 2884: 2880: 2876: 2872: 2868: 2861: 2853: 2849: 2844: 2839: 2834: 2829: 2825: 2821: 2817: 2813: 2812: 2807: 2800: 2792: 2788: 2784: 2780: 2776: 2772: 2765: 2763: 2754: 2750: 2746: 2742: 2738: 2734: 2729: 2724: 2720: 2716: 2712: 2708: 2704: 2700: 2696: 2688: 2680: 2676: 2671: 2666: 2661: 2660:10.1206/352.1 2656: 2652: 2648: 2647: 2642: 2635: 2633: 2631: 2629: 2627: 2618: 2617: 2609: 2601: 2597: 2593: 2587: 2585: 2576: 2572: 2567: 2562: 2557: 2552: 2548: 2544: 2540: 2536: 2532: 2530: 2521: 2517: 2504: 2503: 2495: 2494: 2486: 2485: 2477: 2476: 2467: 2463: 2462: 2454: 2453: 2445: 2444: 2436: 2435: 2427: 2426: 2418: 2417: 2409: 2408: 2400: 2399: 2391: 2390: 2387: 2386: 2384: 2380: 2370: 2369: 2366: 2365: 2362: 2360: 2356: 2355: 2354:Batrachotomus 2345: 2344: 2338: 2337: 2334: 2333: 2330: 2328: 2324: 2323: 2313: 2312: 2305: 2301: 2300: 2297: 2296: 2293: 2291: 2287: 2286: 2276: 2275: 2269: 2268: 2265: 2264: 2256: 2255: 2247: 2246: 2238: 2237: 2234: 2232: 2228: 2227: 2217: 2216: 2213: 2212: 2209: 2208: 2207: 2206:Stagonosuchus 2197: 2196: 2190: 2189: 2186: 2185: 2182: 2180: 2176: 2174: 2164: 2163: 2157: 2156: 2153: 2152: 2149: 2148: 2147: 2137: 2136: 2130: 2129: 2122: 2121:Rauisuchoidea 2118: 2117: 2114: 2113: 2110: 2108: 2104: 2103: 2102:Poposauroidea 2093: 2092: 2085: 2081: 2080: 2077: 2076: 2068: 2067: 2064: 2062: 2058: 2057: 2047: 2046: 2043: 2042: 2039: 2037: 2033: 2032: 2022: 2021: 2015: 2014: 2008: 2007: 2004: 2003: 1995: 1994: 1986: 1985: 1977: 1976: 1973: 1970: 1965: 1964: 1957: 1956: 1953: 1952: 1949: 1947: 1943: 1942: 1932: 1931: 1924: 1920: 1919: 1916: 1915: 1912: 1910: 1906: 1905: 1904:Gracilisuchus 1895: 1894: 1887: 1883: 1882: 1879: 1878: 1875: 1873: 1869: 1868: 1858: 1857: 1851: 1850: 1843: 1839: 1838: 1835: 1834: 1831: 1829: 1825: 1824: 1814: 1813: 1806: 1805: 1800: 1799: 1796: 1795: 1792: 1790: 1786: 1785: 1778: 1777: 1770: 1766: 1765: 1762: 1761: 1758: 1756: 1752: 1743: 1742: 1736: 1735: 1732: 1731: 1728: 1726: 1722: 1721: 1711: 1710: 1704: 1703: 1700: 1699: 1696: 1694: 1690: 1689: 1688:Erythrosuchus 1679: 1678: 1672: 1671: 1667: 1665: 1662: 1652: 1651: 1643: 1642: 1634: 1633: 1625: 1624: 1616: 1615: 1607: 1606: 1598: 1597: 1588: 1587: 1579: 1578: 1570: 1569: 1561: 1560: 1552: 1551: 1543: 1542: 1534: 1533: 1525: 1524: 1516: 1515: 1507: 1506: 1503: 1500: 1495: 1494: 1487: 1486: 1483: 1482: 1479: 1477: 1473: 1472: 1462: 1461: 1455: 1454: 1451: 1450: 1447: 1446: 1445: 1435: 1434: 1428: 1427: 1424: 1423: 1420: 1418: 1414: 1413: 1412:Batrachotomus 1403: 1402: 1396: 1395: 1392: 1391: 1388: 1386: 1382: 1381: 1371: 1370: 1364: 1363: 1360: 1359: 1356: 1354: 1350: 1349: 1339: 1338: 1331: 1327: 1326: 1323: 1322: 1319: 1317: 1313: 1312: 1311:Poposauroidea 1302: 1301: 1294: 1290: 1289: 1286: 1285: 1282: 1280: 1276: 1275: 1265: 1264: 1258: 1257: 1254: 1253: 1245: 1244: 1241: 1239: 1235: 1234: 1224: 1223: 1220: 1219: 1216: 1214: 1210: 1209: 1199: 1198: 1192: 1191: 1188: 1187: 1184: 1182: 1178: 1177: 1176:Turfanosuchus 1167: 1166: 1163: 1162: 1159: 1157: 1153: 1152: 1151:Gracilisuchus 1142: 1141: 1134: 1130: 1129: 1126: 1125: 1122: 1120: 1116: 1115: 1105: 1104: 1098: 1094: 1093: 1090: 1089: 1086: 1084: 1079: 1078: 1071: 1070: 1063: 1059: 1058: 1055: 1054: 1051: 1049: 1045: 1044: 1034: 1033: 1026: 1025: 1020: 1019: 1016: 1015: 1012: 1010: 1006: 1005: 995: 994: 988: 987: 984: 983: 980: 978: 974: 973: 963: 962: 956: 955: 952: 951: 948: 946: 942: 941: 931: 930: 924: 923: 920: 919: 916: 914: 910: 909: 908:Erythrosuchus 899: 898: 892: 891: 888: 887: 884: 882: 878: 877: 867: 866: 859: 856: 855: 851: 849: 844: 840: 836: 825: 822: 820: 816: 812: 808: 803: 798: 796: 792: 788: 784: 780: 776: 771: 769: 768:sphenosuchian 765: 761: 757: 753: 749: 745: 741: 737: 730: 729: 725: 720: 711: 709: 705: 701: 697: 687: 685: 681: 677: 673: 669: 665: 660: 658: 654: 650: 646: 642: 637: 635: 631: 626: 622: 618: 614: 610: 608: 607: 602: 601: 600:Sharovipteryx 596: 592: 588: 587: 586:Hesperosuchus 582: 581: 580:Ornithosuchus 576: 572: 568: 567: 562: 558: 557: 552: 547: 545: 544: 539: 535: 525: 523: 519: 515: 510: 508: 504: 500: 496: 492: 482: 480: 476: 472: 468: 464: 460: 456: 452: 447: 445: 441: 437: 436: 431: 430: 425: 421: 417: 416:erpetosuchids 413: 412: 407: 406: 401: 400: 395: 391: 385: 383: 379: 375: 371: 367: 363: 359: 358: 353: 352: 347: 343: 339: 334: 332: 327: 323: 312: 301: 297: 283: 282: 280: 277: 273: 268: 265: 263: 257: 254: 248: 246: 245: 238: 237: 235: 230: 225: 220: 217: 214: 211: 210: 207: 204: 201: 198: 197: 194: 191: 188: 187: 184: 181: 178: 177: 174: 171: 168: 167: 164: 161: 158: 157: 152: 147: 143: 139: 135: 134: 133:Desmatosuchus 129: 125: 124: 119: 115: 110: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 48: 42: 35: 31: 22: 19: 6896:Pseudosuchia 6866:Pseudosuchia 6865: 6821:Sphenosuchia 6778: 6741:crocodilians 6712:notosuchians 6666: 6661:Nominosuchus 6659: 6652: 6645: 6638: 6618: 6611: 6604: 6599:Hadongsuchus 6597: 6569: 6559: 6548: 6538: 6528: 6518: 6508: 6498: 6465: 6459:Lisboasaurus 6457: 6450: 6445:Cassissuchus 6443: 6419: 6412: 6389: 6381: 6373: 6366: 6359: 6352: 6347:Edentosuchus 6345: 6338: 6331: 6310:Tagarosuchus 6308: 6298: 6288: 6280:Microchampsa 6278: 6268: 6258: 6248: 6185: 6178: 6173:Almadasuchus 6171: 6164:Hallopodidae 6148: 6141: 6134: 6111: 6100: 6092:Sphenosuchus 6090: 6080: 6070: 6060: 6050: 6040: 6030: 6019: 6009: 5999: 5989: 5979: 5909: 5858: 5850: 5843: 5835: 5830:Teratosaurus 5828: 5822:Scythosuchus 5820: 5813: 5806: 5798: 5791: 5786:Polonosuchus 5784: 5776: 5770:Energosuchus 5768: 5762:Arganasuchus 5760: 5753:Rauisuchidae 5739:Youngosuchus 5737: 5727: 5717: 5707: 5699:Prestosuchus 5697: 5686: 5678:Luperosuchus 5676: 5665: 5654: 5644: 5636:Fasolasuchus 5634: 5624: 5614: 5604: 5594: 5586:Arganasuchus 5584: 5574: 5543: 5536: 5529: 5509: 5502: 5495:Poposauridae 5482: 5476:Tsylmosuchus 5474: 5467: 5460: 5453: 5448:Bromsgroveia 5446: 5439: 5419: 5411: 5404: 5397: 5370: 5313: 5265:Yonghesuchus 5263: 5256: 5249: 5241: 5234: 5184: 5177: 5172:Acaenasuchus 5170: 5143: 5138:Parringtonia 5136: 5131:Pagosvenator 5129: 5124:Erpetosuchus 5122: 5114: 5106: 5085:Ticinosuchus 5083: 5072: 5062: 5051: 5040: 5007: 5000: 4993: 4988:Dynamosuchus 4986: 4980:Aenigmaspina 4978: 4954: 4944: 4934: 4915:Pseudosuchia 4914: 4902:Pseudosuchia 4901: 4861:Pseudosuchia 4860: 4786: 4752:Pseudosuchia 4751: 4659:Choristodera 4624:Paraphyletic 4611: 4601: 4591: 4581: 4571: 4561: 4551: 4541: 4531: 4521: 4511: 4500:Nomina dubia 4498: 4486: 4476: 4466: 4458:Palaeosaurus 4456: 4448:Tecovasaurus 4446: 4436: 4428:Crosbysaurus 4426: 4381: 4344:crocodilians 4338:Pseudosuchia 4327:Pseudosuchia 4326: 4285: 4274: 4261: 4173: 4166: 4159: 4136: 4126: 4116: 4108:Litorosuchus 4106: 4096: 4086: 4076: 4039: 4034:Shansisuchus 4032: 4025: 4018: 4010: 4003: 3996: 3989: 3966: 3959: 3935: 3929:Tsylmosuchus 3927: 3920: 3913: 3906: 3899: 3891: 3884: 3861: 3851: 3841: 3831: 3820: 3810: 3800: 3737: 3689: 3682: 3606: 3596: 3585: 3575: 3541: 3534: 3527: 3519: 3512: 3505: 3498: 3491: 3484: 3476: 3470:Fuyuansaurus 3468: 3461: 3454: 3447: 3427: 3420: 3399: 3391: 3384: 3377: 3369: 3363:Fuyuansaurus 3361: 3354: 3347: 3326: 3297: 3289:Rhombopholis 3287: 3277: 3267: 3257: 3247: 3237: 3226: 3218:Czatkowiella 3216: 3206: 3093: 3084: 3075: 2966: 2960: 2951: 2920: 2907: 2901: 2874: 2870: 2860: 2815: 2809: 2799: 2774: 2770: 2702: 2698: 2687: 2650: 2644: 2615: 2608: 2599: 2538: 2534: 2528: 2520: 2466:Pseudosuchia 2465: 2379:Prestosuchus 2377: 2373: 2352: 2348: 2320: 2316: 2285:Rauisuchidae 2283: 2279: 2226:Ticinosuchus 2224: 2220: 2204: 2200: 2173:Fasolasuchus 2171: 2167: 2146:Arganasuchus 2144: 2140: 2100: 2096: 2054: 2050: 2029: 2025: 1961: 1960: 1941:Erpetosuchus 1939: 1935: 1902: 1898: 1865: 1861: 1821: 1817: 1802: 1782: 1781: 1746: 1718: 1714: 1686: 1682: 1661:monophyletic 1658: 1491: 1490: 1471:Rauisuchidae 1469: 1465: 1444:Fasolasuchus 1442: 1438: 1410: 1406: 1378: 1374: 1348:Prestosuchus 1346: 1342: 1309: 1305: 1274:Ticinosuchus 1272: 1268: 1231: 1227: 1206: 1202: 1174: 1170: 1149: 1145: 1112: 1108: 1097:Pseudosuchia 1096: 1075: 1074: 1041: 1037: 1022: 1002: 998: 970: 966: 938: 934: 906: 902: 874: 870: 843:phylogenetic 831: 823: 799: 779:Sphenosuchia 772: 752:rauisuchians 733: 726: 700:ornithodires 693: 680:phylogenetic 661: 653:Rauisuchidae 638: 630:sister taxon 621:phylogenetic 611: 604: 598: 584: 578: 564: 554: 548: 541: 531: 511: 491:Pseudosuchia 490: 488: 448: 444:shuvosaurids 440:ornithomimid 433: 427: 411:Fasolasuchus 409: 405:Prestosuchus 403: 397: 386: 365: 355: 349: 335: 326:crocodilians 296:Pseudosuchia 295: 294: 242: 219:Pseudosuchia 218: 212: 199: 131: 128:Rauisuchidae 121: 18: 6890:Wikispecies 6826:Protosuchia 6706:Ziphosuchia 6695:Ziphosuchia 6640:Adzhosuchus 6510:Hsisosuchus 6467:Zaraasuchus 6452:Gobiosuchus 6421:Orthosuchus 6414:Notochampsa 6391:Protosuchus 6375:Orthosuchus 6368:Hoplosuchus 6340:Dianosuchus 6021:Heteropelta 5793:Postosuchus 5719:Saurosuchus 5688:Mandasuchus 5656:Heteropelta 5646:Heptasuchus 5576:Apatosuchus 5545:Sillosuchus 5538:Shuvosaurus 5484:Xilousuchus 5421:Qianosuchus 5372:Mambawakale 5251:Parvosuchus 5108:Archeopelta 5074:Nundasuchus 5064:Mandasuchus 5042:Heteropelta 5002:Riojasuchus 4956:Phytosauria 4946:Nundasuchus 4791:Archosauria 4643:Thecodontia 4613:Wangisuchus 4593:Microcnemus 4573:Fenhosuchus 4563:Exilisuchus 4553:Collilongus 4543:Blomosuchus 4523:Ankistrodon 4478:Uatchitodon 4253:Archosauria 4241:Phytosauria 4230:Phytosauria 4098:Heteropelta 4057:Eucrocopoda 4041:Uralosaurus 3998:Chalishevia 3908:Jaikosuchus 3886:Archosaurus 3802:Antarctanax 3577:Boreopricea 3507:Macrocnemus 3449:Amotosaurus 2728:11336/41466 2322:Saurosuchus 1823:Phytosauria 1769:Archosauria 1380:Saurosuchus 1062:Archosauria 1043:Phytosauria 756:herbivorous 696:Archosauria 690:Description 682:analyses. 672:Phytosauria 664:Paul Sereno 606:Longisquama 575:Thecodontia 435:Postosuchus 399:Saurosuchus 370:Phytosauria 322:Archosauria 253:Phytosauria 206:Archosauria 123:Postosuchus 39:248–0  6940:Categories 6831:Mesosuchia 6816:Bathyotica 6806:Rauisuchia 6804:See also: 6683:Metasuchia 6520:Induszalim 6113:Trialestes 5881:see below↓ 5837:Tikisuchus 5808:Rauisuchus 5778:Jushatyria 5626:Etjosuchus 5606:Dagasuchus 5511:Poposaurus 5406:Lotosaurus 5285:see below↓ 5208:Aetosauria 5197:Aetosauria 4870:see below↓ 4822:Sauropsida 4807:Sauropsida 4782:Sauropsida 4583:Megacnemus 4513:Albisaurus 4417:Tooth taxa 4217:Crurotarsi 4175:Osmolskina 4161:Euparkeria 4138:Vancleavea 4088:Dorosuchus 4012:Fugusuchus 3968:Triopticus 3961:Kranosaura 3901:Gamosaurus 3833:Kalisuchus 3822:Eorasaurus 3812:Cuyosuchus 3711:see below↓ 3691:Prolacerta 3684:Kadimakara 3514:Raibliania 3500:Luxisaurus 3463:Cosesaurus 3349:Austronaga 3314:Tanysauria 3239:Elessaurus 3150:see below↓ 3129:Sauropsida 3114:Sauropsida 3071:Sauropsida 2512:References 2084:Rauisuchia 1923:Bathyotica 1867:Aetosauria 1804:Crurotarsi 1720:Euparkeria 1664:Rauisuchia 1233:Aetosauria 1024:Crurotarsi 1004:Euparkeria 940:Vancleavea 839:Crurotarsi 815:alligators 811:crocodiles 742:); by the 668:Crurotarsi 522:pterosaurs 499:Crurotarsi 471:alligators 467:crocodiles 463:Cretaceous 429:Poposaurus 378:pterosaurs 376:) such as 368:'s group, 346:node-based 342:Crurotarsi 232:Subgroups 138:Aetosauria 6739:includes 6735:Neosuchia 6724:Neosuchia 6710:includes 6654:Kyasuchus 6052:Parrishia 4768:Kingdom: 4488:Zanclodon 4342:includes 4313:includes 4078:Asperoris 4020:Garjainia 3937:Vonhuenia 3608:Teyujagua 3560:Crocopoda 3386:Pectodens 3057:Kingdom: 2969:(2): 1–7. 2962:PaleoBios 2912:Eichstätt 2753:256707805 2737:2397-334X 2670:2246/6112 2653:: 1–292. 848:cladogram 828:Phylogeny 795:dinosaurs 764:poposaurs 760:aetosaurs 748:carnivore 736:Olenekian 724:loricatan 714:Evolution 704:dinosaurs 662:In 1990, 657:aetosaurs 571:suborders 538:aetosaurs 518:dinosaurs 489:The name 479:gavialids 424:aetosaurs 382:dinosaurs 318:(souchos) 307:(pseudos) 169:Kingdom: 163:Eukaryota 6910:42334946 6881:Q2742285 6875:Wikidata 6841:Category 6620:Zosuchus 6270:Kemkemia 6180:Hallopus 5981:Carnufex 5560:Loricata 5145:Tarjadia 4776:Chordata 4774:Phylum: 4770:Animalia 4710:Category 3080:Diapsida 3065:Chordata 3063:Phylum: 3059:Animalia 2996:Archived 2852:20308590 2745:29185518 2679:83493714 2575:22022431 2535:PLOS ONE 1330:Loricata 835:Eusuchia 819:gharials 802:Mesozoic 758:armored 744:Ladinian 740:Triassic 613:Gauthier 591:theropod 495:Triassic 459:Jurassic 396:such as 390:Triassic 366:Rutiodon 362:Triassic 331:misnomer 276:Synonyms 193:Reptilia 183:Chordata 179:Phylum: 173:Animalia 159:Domain: 5845:Vivaron 5531:Effigia 5243:Maehary 5116:Dyoplax 4780:Class: 3069:Class: 2943:Sources 2879:Bibcode 2877:: 763. 2843:2872409 2820:Bibcode 2779:Bibcode 2707:Bibcode 2566:3194824 2543:Bibcode 2468:  2306:  2123:  2086:  1925:  1888:  1844:  1807:  1771:  1332:  1295:  1135:  1064:  1027:  860:  800:As the 738:(early 708:mammals 543:Dyoplax 475:caimans 189:Class: 34:Present 5025:Suchia 4626:groups 4533:Avipes 3422:Ozimek 3089:Sauria 2850:  2840:  2751:  2743:  2735:  2677:  2573:  2563:  2464:  2302:  2119:  2082:  1921:  1884:  1842:Suchia 1840:  1801:  1767:  1328:  1291:  1133:Suchia 1131:  1095:  1060:  1021:  817:, and 785:(both 641:Benton 617:Padian 520:, and 477:, and 442:-like 408:, and 316:σούχος 305:ψεύδος 298:(from 267:Suchia 226:, 1887 224:Zittel 130:) and 6923:93706 4787:Clade 4315:birds 3094:Clade 3085:Clade 3076:Clade 2749:S2CID 2675:S2CID 706:' or 507:clade 503:clade 338:clade 311:Greek 300:Greek 213:Clade 200:Clade 6733:see 6704:see 6210:see 5206:see 4847:see 4820:see 4336:see 4307:see 4287:Smok 4239:see 4198:see 3657:see 3631:see 3127:see 2848:PMID 2811:PNAS 2741:PMID 2733:ISSN 2571:PMID 781:and 773:The 655:and 615:and 603:and 583:and 559:and 540:and 461:and 432:and 418:and 380:and 336:The 47:PreꞒ 6905:EoL 2930:doi 2887:doi 2838:PMC 2828:doi 2816:107 2787:doi 2723:hdl 2715:doi 2665:hdl 2655:doi 2651:352 2561:PMC 2551:doi 651:of 563:'s 553:'s 6942:: 6920:: 6907:: 6892:: 6877:: 4789:: 3096:: 3087:: 3078:: 2967:25 2965:. 2885:. 2873:. 2869:. 2846:. 2836:. 2826:. 2814:. 2808:. 2785:. 2775:17 2773:. 2761:^ 2747:. 2739:. 2731:. 2721:. 2713:. 2701:. 2697:. 2673:. 2663:. 2649:. 2643:. 2625:^ 2583:^ 2569:. 2559:. 2549:. 2537:. 2533:. 813:, 754:, 481:. 473:, 469:, 402:, 313:: 302:: 215:: 202:: 140:) 97:Pg 41:Ma 36:, 6692:† 6583:† 6574:? 6567:† 6557:† 6553:? 6546:† 6536:† 6526:† 6516:† 6506:† 6496:† 6462:? 6433:† 6424:? 6402:† 6386:? 6378:? 6321:† 6306:† 6296:† 6286:† 6276:† 6266:† 6256:† 6246:† 6198:† 6161:† 6124:† 6109:† 6105:? 6098:† 6088:† 6078:† 6068:† 6058:† 6048:† 6038:† 6028:† 6024:? 6017:† 6007:† 5997:† 5987:† 5977:† 5863:? 5855:? 5840:? 5825:? 5803:? 5781:? 5773:? 5765:? 5750:† 5735:† 5725:† 5715:† 5705:† 5695:† 5691:? 5684:† 5674:† 5670:? 5663:† 5659:? 5652:† 5642:† 5632:† 5622:† 5612:† 5602:† 5592:† 5582:† 5572:† 5479:? 5416:? 5383:† 5368:† 5246:? 5224:† 5156:† 5119:? 5111:? 5096:† 5081:† 5077:? 5070:† 5060:† 5056:? 5049:† 5045:? 5038:† 4983:? 4968:† 4959:? 4952:† 4942:† 4932:† 4744:e 4737:t 4730:v 4609:† 4599:† 4589:† 4579:† 4569:† 4559:† 4549:† 4539:† 4529:† 4519:† 4509:† 4484:† 4474:† 4464:† 4454:† 4444:† 4434:† 4424:† 4283:† 4279:? 4272:† 4227:† 4186:† 4149:† 4134:† 4124:† 4114:† 4104:† 4094:† 4084:† 4074:† 4015:? 3979:† 3949:† 3940:? 3932:? 3896:? 3874:† 3859:† 3849:† 3839:† 3829:† 3825:? 3818:† 3808:† 3798:† 3676:? 3671:† 3645:† 3619:† 3604:† 3594:† 3590:? 3583:† 3573:† 3546:? 3524:? 3481:? 3473:? 3414:? 3396:? 3374:? 3366:? 3331:? 3311:† 3295:† 3285:† 3275:† 3265:† 3255:† 3245:† 3235:† 3231:? 3224:† 3214:† 3204:† 3033:e 3026:t 3019:v 2932:: 2895:. 2889:: 2881:: 2875:9 2854:. 2830:: 2822:: 2793:. 2789:: 2781:: 2755:. 2725:: 2717:: 2709:: 2703:1 2681:. 2667:: 2657:: 2577:. 2553:: 2545:: 2539:6 2375:† 2350:† 2318:† 2281:† 2222:† 2202:† 2175:' 2169:† 2142:† 2098:† 2052:† 2027:† 1937:† 1900:† 1863:† 1819:† 1748:† 1716:† 1684:† 1467:† 1440:† 1408:† 1376:† 1344:† 1307:† 1270:† 1229:† 1204:† 1172:† 1147:† 1110:† 1039:† 1000:† 968:† 936:† 904:† 872:† 259:† 255:? 250:† 240:† 136:( 126:( 102:N 92:K 87:J 82:T 77:P 72:C 67:D 62:S 57:O 52:Ꞓ 32:–

Index

Early Triassic
Present
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Postosuchus
Rauisuchidae
Desmatosuchus
Aetosauria
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Reptilia
Archosauria
Pseudosuchia
Zittel
Dasygnathoides

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