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Tam Pa Ling Cave

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The timing of modern human migration from Africa to East Asia is not known with certainty; because bone is poorly preserved in tropical climates, human fossils from the region are rare. Recent discoveries in China, the Philippines, Sri Lanka and Australia had previously established that archaic human
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Demeter, Fabrice; Shackelford, Laura; Westaway, Kira; Duringer, Philippe; Bacon, Anne-Marie; Ponche, Jean-Luc; Wu, Xiujie; Sayavongkhamdy, Thongsa; Zhao, Jian-Xin; Barnes, Lani; Boyon, Marc; Sichanthongtip, Phonephanh; Sénégas, Frank; Karpoff, Anne-Marie; Patole-Edoumba, Elise; Coppens, Yves; Braga,
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specimen was therefore considered a major discovery because it filled in a 60,000-year gap in the fossil record, demonstrating the presence of modern humans in Southeast Asia from at least 60,000 BP. Additionally, as Tam Pa Ling lies a thousand miles inland, the finds challenged previous assumptions
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Shackelford, Laura; Demeter, Fabrice; Westaway, Kira; Duringer, Philippe; Ponche, Jean-Luc; Sayavongkhamdy, Thongsa; Zhao, Jian-Xin; Barnes, Lani; Boyon, Marc; Sichanthongtip, Phonephanh; Sénégas, Frank; Patole-Edoumba, Elise; Coppens, Yves; Dumoncel, Jean; Bacon, Anne-Marie (2017). "Additional
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that humans migrated out of Africa by following coastlines. They suggest that the migration may also have proceeded along river valleys, which served as natural corridors through the continent.
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F. Demeter et al., "Early Modern Humans and Morphological Variation in Southeast Asia: Fossil Evidence from Tam Pa Ling, Laos", PLOS One, 7 April 2015, doi: 10.1371/journal.pone.0121193.
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Excavations at the eastern end of the cave's main gallery, at the base of the sloped entrance, were conducted by a team of American, French and Laotian researchers starting in 2009.
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exhibits a mix of archaic and anatomically modern human traits, exhibiting modern human features such as having a developed chin but not having a robust mandibular corpus; however,
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fossils were present between 125,000 and 100,000 BP, and those of modern humans from about 40,000 BP. The discovery of the fully modern
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Laura Shackleford, Fabrice Demeter and the team. In April 2016 they were returned to Laos, and are now housed in a new building of the
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Tam Pa Ling has a single, south-facing opening and descends 65 m (213 ft) to its main gallery. It is part of a network of
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periods. The main gallery measures 30 m (98 ft) from north to south and 40 m (130 ft) from east to west.
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also retains archaic human features such as having a broad anterior mandibular arch. Luminescence dating of the
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evidence for early modern human morphological diversity in Southeast Asia at Tam Pa Ling, Laos".
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The Emergence of Modern Human Behaviour: The Pleistocene Occupation Sites of South Africa
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features. As of 2017, it provides the earliest skeletal evidence for the presence of
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of the sediments established a minimum age of 51,000 to 46,000 years, and direct
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Cave of Altamira and Paleolithic Cave Art of Northern Spain World Heritage Site
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sediment layer provides a date range from around 70,000 to 48,000 years old.
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In 2013, researchers recovered the partial mandible of a third fossil find,
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The fossils were temporarily removed to the United States for study by
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and small overall size, with modern human traits like a developed
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mandible was found lower down in the same stratigraphic unit as
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Rock art of the Iberian Mediterranean Basin World Heritage Site
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Caves and Ice Age Art in the Swabian Jura World Heritage Site
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of the fossils indicated a maximum age of 63,000 years.
6080: 524: 845:"Oldest bones from modern humans in Asia discovered" 620:Marwick, Ben; Bouasisengpaseuth, Bounheung (2017). 1406:(archaic homo sapiens, anatomically modern humans) 622:"The History and Practice of Archaeology in Laos" 6116: 626:Handbook of East and Southeast Asian Archaeology 869:. Research News Bureau ILLINOIS. April 13, 2016 666:Proceedings of the National Academy of Sciences 749:"Ancient Human Fossils from Tam Pa Ling, Laos" 428:, and represents a mature adult that combines 341:The first fossil find, a hominin skull dubbed 6003: 1881: 912: 720:"Finding a Home for the Bones of Tam Pa Ling" 4918: 1391: 892:Anatomically modern human in Southeast Asia 6010: 5996: 1896:Prehistoric cave sites, rock shelters and 1888: 1874: 919: 905: 798: 788: 687: 677: 4663:Cradle of Humankind, World Heritage Site 1811:Human evolutionary developmental biology 867:"Bringing home the bones of Tam Pa Ling" 726:. University of Illinois Urbana-Campaign 411: 49: 717: 656: 391:. It was identified as belonging to an 305: 16:Cave in Laos with archaic human fossils 6117: 5991: 5593: 4917: 4369: 3523: 1931: 1869: 1596:Evolutionary models of human drug use 900: 322:beds that were laid down between the 1836: 742: 740: 926: 825:. Public Broadcasting Service (PBS) 13: 751:. Anthropology Net. April 11, 2015 601:. Anthropology Net. April 11, 2015 270:have been discovered in the cave: 239:Fabrice Demeter, Laura Shackleford 14: 6161: 3051:Mala Balanica and Velika Balanica 2214:Vézère Valley World Heritage Site 2009:Neolithic flint mines of Spiennes 885: 763: 737: 87:260 km (160 mi) NNE of 6102: 6090: 1847: 1835: 1824: 1823: 512:Early hominids in Southeast Asia 383:, as well as the right and left 48: 41: 25: 859: 837: 579:. Live Science. August 20, 2012 466: 815: 711: 650: 613: 591: 569: 1: 657:Demeter, F. (July 23, 2012). 517: 6125:Archaeological sites in Laos 823:"The Woman from Tam Pa Ling" 790:10.1371/journal.pone.0121193 628:. Springer New York: 89–95. 555:10.1016/j.quaint.2016.12.002 145:1,170 m (3,839 ft) 7: 5594: 3239:Other caves with decoration 2321:Other caves with decoration 2243:Les Eyzies-de-Tayac-Sireuil 1919:Caves containing pictograms 1854:Evolutionary biology Portal 634:10.1007/978-1-4939-6521-2_8 495: 10: 6166: 6145:Peopling of Southeast Asia 6140:Paleoanthropological sites 4370: 3223:Other World Heritage Sites 2310:Other World Heritage Sites 1932: 1909:Paleoanthropological sites 847:. CBSNews. August 20, 2012 432:features such as a robust 333: 274:, a skull belonging to an 31:South entrance to the cave 6052: 6023: 5959: 5935: 5919: 5903: 5885: 5864: 5846: 5830: 5802: 5606: 5602: 5589: 5325: 5309: 5268: 5212: 5196: 5180: 5162: 5121: 5080: 5059: 5038: 4992: 4966: 4950: 4934: 4930: 4913: 4889: 4868: 4852: 4826: 4655: 4629: 4613: 4592: 4576: 4530: 4509: 4493: 4472: 4451: 4435: 4419: 4398: 4382: 4378: 4365: 4336: 4315: 4289: 4277: 4251: 4220: 4184: 4166: 4150: 4134: 4118: 4102: 4056: 4035: 4019: 3968: 3887: 3866: 3810: 3779: 3733: 3717: 3691: 3630: 3614: 3568: 3552: 3536: 3532: 3519: 3410: 3394: 3373: 3116: 3085: 3059: 3028: 2982: 2951: 2900: 2879: 2863: 2845: 2829: 2813: 2797: 2779: 2716: 2695: 2634: 2606: 2530: 2206: 2190: 2149: 2133: 2092: 2056: 2035: 1980: 1944: 1940: 1927: 1904: 1819: 1799:Evolutionary anthropology 1776: 1750: 1703: 1641: 1560: 1519: 1512: 1458: 1382: 1305: 1230: 1184: 1177: 1160: 1120: 1056: 1027: 991: 982: 949: 934: 393:anatomically modern human 276:anatomically modern human 235: 227: 222: 214: 200: 186: 181: 173: 165: 157: 149: 141: 104: 94: 76: 68: 36: 24: 5887:Northern Mariana Islands 3524: 543:Quaternary International 1705:Origin of modern humans 679:10.1073/pnas.1208104109 407:mainland Southeast Asia 387:and a largely complete 169:40 m (130 ft) 6034:Diepkloof Rock Shelter 4411:Manyana Rock Paintings 2462:La Chapelle-aux-Saints 771:José (April 7, 2015). 417: 359:uranium-thorium dating 177:30 m (98 ft) 126:20.20861°N 103.40972°E 4974:Actun Tunichil Muknal 1713:Recent African origin 951:Last common ancestors 415: 314:caves, formed by the 286:with both modern and 228:Excavation dates 69:Alternative name 6097:Evolutionary biology 4548:Ifri N'Amr Ou Moussa 3488:Red Lady of Paviland 2380:Mayrières supérieure 1728:Behavioral modernity 1718:Multiregional origin 1498:archaic Homo sapiens 1493:Homo heidelbergensis 1438:Red Deer Cave people 724:Illinois News Bureau 718:Shackelford, Laura. 306:Location and geology 215:Associated with 6130:Archaeology of Laos 6070:Klasies River Caves 6026:World Heritage Site 5363:Colorado Millennial 4001:Shiraho Saonetabaru 2805:Bukuroshja e Fjetur 1365:H. neanderthalensis 1285:H. e. tautavelensis 672:(36): 14375–14380. 486:Lao National Museum 482:paleoanthropologist 397:Sub-Saharan African 355:luminescence dating 324:Upper Carboniferous 252:) is a cave in the 250:Cave of the Monkeys 131:20.20861; 103.40972 122: /  72:Cave of the Monkeys 21: 5181:Dominican Republic 4942:Cueva de las Manos 3838:Huto and Kamarband 3248:Barranc del Migdia 3134:Caves in Cantabria 2558:Hohlenstein-Stadel 2365:Grotte de Gabillou 2350:La Chaire a Calvin 1552:Self-domestication 1343:H. heidelbergensis 1292:H. e. yuanmouensis 1257:H. e. lantianensis 984:Australopithecines 418: 254:Annamite Mountains 209:Middle Paleolithic 99:Annamite Mountains 81:Houaphanh Province 19: 6078: 6077: 5985: 5984: 5955: 5954: 5951: 5950: 5585: 5584: 5581: 5580: 5098:Piedras del Tunjo 5030:Toca da Tira Peia 4909: 4908: 4905: 4904: 4361: 4360: 4357: 4356: 3515: 3514: 3511: 3510: 3151:Hornos de la Peña 1863: 1862: 1804:Paleoanthropology 1746: 1745: 1723:Archaic admixture 1601:Stoned ape theory 1537:Endurance running 1454: 1453: 1450: 1449: 1446: 1445: 1301: 1300: 1264:H. e. nankinensis 1220:H. tsaichangensis 1156: 1155: 643:978-1-4939-6519-9 434:mandibular corpus 256:in north-eastern 243: 242: 205:Upper Paleolithic 6157: 6107: 6106: 6105: 6095: 6094: 6086: 6012: 6005: 5998: 5989: 5988: 5904:Papua New Guinea 5604: 5603: 5591: 5590: 5543:Stanfield-Worley 5518:Rockhouse Cliffs 4932: 4931: 4915: 4914: 4783:Howieson's Poort 4480:Enkapune Ya Muto 4380: 4379: 4367: 4366: 3833:Eshkaft-e Siahoo 3534: 3533: 3521: 3520: 3288:Peñas de Cabrera 2588:Kleine Feldhofer 2548:Geissenklösterle 2400:Roc-aux-Sorciers 1942: 1941: 1929: 1928: 1890: 1883: 1876: 1867: 1866: 1851: 1839: 1838: 1827: 1826: 1763:Human prehistory 1738:Recent evolution 1733:Early migrations 1675:Thermoregulation 1576:Expensive tissue 1547:Sexual selection 1517: 1516: 1389: 1388: 1271:H. e. pekinensis 1182: 1181: 1175: 1174: 1090:A. bahrelghazali 1059:Australopithecus 989: 988: 959:Chimpanzee–human 947: 946: 921: 914: 907: 898: 897: 879: 878: 876: 874: 863: 857: 856: 854: 852: 841: 835: 834: 832: 830: 819: 813: 812: 802: 792: 767: 761: 760: 758: 756: 744: 735: 734: 732: 731: 715: 709: 708: 706: 704: 691: 681: 663: 654: 648: 647: 617: 611: 610: 608: 606: 595: 589: 588: 586: 584: 573: 567: 566: 537: 161:Pa Hang Mountain 137: 136: 134: 133: 132: 127: 123: 120: 119: 118: 115: 52: 51: 45: 29: 22: 20:Tam Pa Ling Cave 18: 6165: 6164: 6160: 6159: 6158: 6156: 6155: 6154: 6135:Limestone caves 6115: 6114: 6113: 6103: 6101: 6089: 6081: 6079: 6074: 6048: 6019: 6016: 5986: 5981: 5947: 5931: 5915: 5899: 5881: 5860: 5842: 5826: 5798: 5714:Mackintosh 90/1 5634:Beginner's Luck 5598: 5577: 5393:Dutchess Quarry 5333:Arnold Research 5321: 5305: 5264: 5208: 5192: 5176: 5158: 5117: 5076: 5055: 5034: 4988: 4984:Midnight Terror 4962: 4946: 4926: 4901: 4885: 4864: 4848: 4822: 4651: 4625: 4609: 4588: 4572: 4563:Kelif el Boroud 4526: 4505: 4489: 4468: 4447: 4431: 4415: 4394: 4374: 4353: 4332: 4311: 4285: 4273: 4247: 4216: 4180: 4162: 4146: 4130: 4114: 4098: 4052: 4031: 4015: 3964: 3883: 3862: 3806: 3775: 3729: 3713: 3687: 3626: 3610: 3564: 3548: 3528: 3507: 3406: 3390: 3369: 3268:los Murciélagos 3216:Roca dels Moros 3112: 3081: 3055: 3024: 2995:Chertovy Vorota 2978: 2947: 2896: 2875: 2859: 2847:North Macedonia 2841: 2825: 2809: 2793: 2775: 2712: 2691: 2630: 2602: 2526: 2289:Abri du Poisson 2254:Les Combarelles 2202: 2186: 2145: 2129: 2088: 2052: 2031: 2027:Trou de l'Abîme 1976: 1936: 1923: 1900: 1894: 1864: 1859: 1815: 1772: 1758:Human evolution 1742: 1699: 1643: 1637: 1616:Cooperative eye 1561:Specific models 1556: 1508: 1487:Homo antecessor 1442: 1378: 1372:H. rhodesiensis 1336:H. floresiensis 1297: 1278:H. e. soloensis 1250:H. e. georgicus 1226: 1190:H. gautengensis 1165: 1163: 1152: 1116: 1052: 1023: 978: 969:Orangutan–human 938: 930: 928:Human evolution 925: 888: 883: 882: 872: 870: 865: 864: 860: 850: 848: 843: 842: 838: 828: 826: 821: 820: 816: 783:(4): e0121193. 768: 764: 754: 752: 747: 745: 738: 729: 727: 716: 712: 702: 700: 661: 655: 651: 644: 618: 614: 604: 602: 597: 596: 592: 582: 580: 575: 574: 570: 538: 525: 520: 498: 469: 336: 308: 130: 128: 124: 121: 116: 113: 111: 109: 108: 64: 63: 62: 61: 55: 54: 53: 32: 17: 12: 11: 5: 6163: 6153: 6152: 6147: 6142: 6137: 6132: 6127: 6112: 6111: 6099: 6076: 6075: 6073: 6072: 6067: 6062: 6056: 6054: 6050: 6049: 6047: 6046: 6041: 6039:Pinnacle Point 6036: 6030: 6028: 6021: 6020: 6015: 6014: 6007: 6000: 5992: 5983: 5982: 5980: 5979: 5974: 5969: 5964: 5960: 5957: 5956: 5953: 5952: 5949: 5948: 5946: 5945: 5939: 5937: 5933: 5932: 5930: 5929: 5923: 5921: 5917: 5916: 5914: 5913: 5907: 5905: 5901: 5900: 5898: 5897: 5891: 5889: 5883: 5882: 5880: 5879: 5874: 5868: 5866: 5862: 5861: 5859: 5858: 5852: 5850: 5844: 5843: 5841: 5840: 5834: 5832: 5828: 5827: 5825: 5824: 5819: 5814: 5808: 5806: 5800: 5799: 5797: 5796: 5791: 5786: 5781: 5776: 5771: 5766: 5761: 5756: 5751: 5746: 5741: 5736: 5731: 5726: 5721: 5716: 5711: 5706: 5701: 5696: 5691: 5686: 5681: 5676: 5671: 5666: 5661: 5656: 5651: 5646: 5641: 5636: 5631: 5629:Beeton Shelter 5626: 5621: 5616: 5610: 5608: 5600: 5599: 5587: 5586: 5583: 5582: 5579: 5578: 5576: 5575: 5570: 5565: 5560: 5555: 5550: 5545: 5540: 5535: 5530: 5525: 5520: 5515: 5510: 5505: 5500: 5495: 5490: 5485: 5480: 5475: 5470: 5465: 5460: 5455: 5450: 5445: 5440: 5435: 5430: 5425: 5420: 5415: 5410: 5405: 5400: 5395: 5390: 5385: 5380: 5375: 5370: 5365: 5360: 5355: 5350: 5345: 5340: 5335: 5329: 5327: 5323: 5322: 5320: 5319: 5313: 5311: 5307: 5306: 5304: 5303: 5301:T'uqu T'uquyuq 5298: 5293: 5288: 5283: 5278: 5272: 5270: 5266: 5265: 5263: 5262: 5257: 5252: 5247: 5242: 5237: 5232: 5227: 5222: 5216: 5214: 5210: 5209: 5207: 5206: 5200: 5198: 5194: 5193: 5191: 5190: 5184: 5182: 5178: 5177: 5175: 5174: 5168: 5166: 5160: 5159: 5157: 5156: 5151: 5146: 5141: 5136: 5131: 5125: 5123: 5119: 5118: 5116: 5115: 5110: 5105: 5100: 5095: 5090: 5084: 5082: 5078: 5077: 5075: 5074: 5069: 5063: 5061: 5057: 5056: 5054: 5053: 5048: 5042: 5040: 5036: 5035: 5033: 5032: 5027: 5022: 5017: 5012: 5007: 5002: 4996: 4994: 4990: 4989: 4987: 4986: 4981: 4976: 4970: 4968: 4964: 4963: 4961: 4960: 4954: 4952: 4948: 4947: 4945: 4944: 4938: 4936: 4928: 4927: 4911: 4910: 4907: 4906: 4903: 4902: 4900: 4899: 4893: 4891: 4887: 4886: 4884: 4883: 4878: 4872: 4870: 4866: 4865: 4863: 4862: 4856: 4854: 4850: 4849: 4847: 4846: 4841: 4836: 4830: 4828: 4824: 4823: 4821: 4820: 4815: 4810: 4805: 4803:Pinnacle Point 4800: 4795: 4790: 4785: 4780: 4775: 4770: 4765: 4760: 4755: 4750: 4745: 4741: 4740: 4735: 4730: 4725: 4720: 4715: 4710: 4705: 4700: 4695: 4690: 4685: 4680: 4675: 4670: 4665: 4659: 4657: 4653: 4652: 4650: 4649: 4644: 4639: 4633: 4631: 4627: 4626: 4624: 4623: 4617: 4615: 4611: 4610: 4608: 4607: 4605:The White Lady 4602: 4596: 4594: 4590: 4589: 4587: 4586: 4580: 4578: 4574: 4573: 4571: 4570: 4565: 4560: 4555: 4550: 4545: 4540: 4534: 4532: 4528: 4527: 4525: 4524: 4519: 4513: 4511: 4507: 4506: 4504: 4503: 4497: 4495: 4491: 4490: 4488: 4487: 4482: 4476: 4474: 4470: 4469: 4467: 4466: 4461: 4455: 4453: 4449: 4448: 4446: 4445: 4439: 4437: 4433: 4432: 4430: 4429: 4423: 4421: 4417: 4416: 4414: 4413: 4408: 4402: 4400: 4396: 4395: 4393: 4392: 4386: 4384: 4376: 4375: 4363: 4362: 4359: 4358: 4355: 4354: 4352: 4351: 4346: 4340: 4338: 4334: 4333: 4331: 4330: 4325: 4319: 4317: 4313: 4312: 4310: 4309: 4304: 4299: 4293: 4291: 4287: 4286: 4281: 4279: 4275: 4274: 4272: 4271: 4266: 4261: 4255: 4253: 4249: 4248: 4246: 4245: 4240: 4235: 4230: 4224: 4222: 4218: 4217: 4215: 4214: 4209: 4204: 4199: 4194: 4188: 4186: 4182: 4181: 4179: 4178: 4172: 4170: 4164: 4163: 4161: 4160: 4154: 4152: 4148: 4147: 4145: 4144: 4138: 4136: 4132: 4131: 4129: 4128: 4126:Khoit Tsenkher 4122: 4120: 4116: 4115: 4113: 4112: 4106: 4104: 4100: 4099: 4097: 4096: 4091: 4086: 4081: 4076: 4071: 4066: 4060: 4058: 4054: 4053: 4051: 4050: 4045: 4039: 4037: 4033: 4032: 4030: 4029: 4023: 4021: 4017: 4016: 4014: 4013: 4008: 4003: 3998: 3993: 3988: 3983: 3978: 3972: 3970: 3966: 3965: 3963: 3962: 3957: 3952: 3947: 3942: 3937: 3932: 3927: 3922: 3917: 3912: 3907: 3902: 3897: 3891: 3889: 3885: 3884: 3882: 3881: 3876: 3870: 3868: 3864: 3863: 3861: 3860: 3855: 3850: 3845: 3840: 3835: 3830: 3825: 3820: 3814: 3812: 3808: 3807: 3805: 3804: 3799: 3794: 3789: 3783: 3781: 3777: 3776: 3774: 3773: 3768: 3763: 3758: 3753: 3748: 3743: 3737: 3735: 3731: 3730: 3728: 3727: 3721: 3719: 3715: 3714: 3712: 3711: 3706: 3701: 3695: 3693: 3689: 3688: 3686: 3685: 3680: 3675: 3670: 3665: 3660: 3655: 3650: 3645: 3640: 3634: 3632: 3628: 3627: 3625: 3624: 3618: 3616: 3612: 3611: 3609: 3608: 3603: 3598: 3593: 3588: 3583: 3578: 3572: 3570: 3566: 3565: 3563: 3562: 3556: 3554: 3550: 3549: 3547: 3546: 3540: 3538: 3530: 3529: 3517: 3516: 3513: 3512: 3509: 3508: 3506: 3505: 3500: 3495: 3490: 3485: 3480: 3475: 3470: 3465: 3460: 3455: 3450: 3445: 3440: 3438:Creswell Crags 3435: 3430: 3425: 3420: 3418:Aveline's Hole 3414: 3412: 3411:United Kingdom 3408: 3407: 3405: 3404: 3398: 3396: 3392: 3391: 3389: 3388: 3383: 3377: 3375: 3371: 3370: 3368: 3367: 3362: 3360:Santa Catalina 3357: 3352: 3347: 3342: 3337: 3332: 3327: 3322: 3317: 3312: 3306: 3305: 3300: 3295: 3290: 3285: 3280: 3275: 3270: 3265: 3260: 3255: 3250: 3245: 3240: 3236: 3235: 3230: 3225: 3219: 3218: 3213: 3208: 3201: 3200: 3195: 3190: 3185: 3180: 3179: 3178: 3175: 3174: 3173: 3168: 3165: 3162: 3155:Monte Castillo 3152: 3149: 3144: 3141: 3131: 3126: 3120: 3118: 3114: 3113: 3111: 3110: 3105: 3100: 3095: 3089: 3087: 3083: 3082: 3080: 3079: 3074: 3069: 3063: 3061: 3057: 3056: 3054: 3053: 3048: 3043: 3038: 3036:Hadži-Prodan's 3032: 3030: 3026: 3025: 3023: 3022: 3017: 3012: 3007: 3002: 2997: 2992: 2986: 2984: 2980: 2979: 2977: 2976: 2971: 2966: 2961: 2955: 2953: 2949: 2948: 2946: 2945: 2940: 2935: 2930: 2925: 2920: 2915: 2910: 2904: 2902: 2898: 2897: 2895: 2894: 2889: 2883: 2881: 2877: 2876: 2874: 2873: 2867: 2865: 2861: 2860: 2858: 2857: 2851: 2849: 2843: 2842: 2840: 2839: 2833: 2831: 2827: 2826: 2824: 2823: 2817: 2815: 2811: 2810: 2808: 2807: 2801: 2799: 2795: 2794: 2792: 2791: 2785: 2783: 2777: 2776: 2774: 2773: 2768: 2763: 2758: 2751: 2746: 2741: 2736: 2731: 2726: 2720: 2718: 2714: 2713: 2711: 2710: 2705: 2699: 2697: 2693: 2692: 2690: 2689: 2684: 2679: 2674: 2669: 2664: 2659: 2654: 2649: 2644: 2638: 2636: 2632: 2631: 2629: 2628: 2623: 2618: 2612: 2610: 2604: 2603: 2601: 2600: 2595: 2590: 2585: 2580: 2575: 2571: 2570: 2565: 2560: 2555: 2550: 2545: 2540: 2534: 2532: 2528: 2527: 2525: 2524: 2519: 2514: 2509: 2504: 2499: 2494: 2489: 2484: 2479: 2474: 2469: 2464: 2459: 2456: 2451: 2446: 2443: 2440: 2437: 2434: 2429: 2424: 2418: 2417: 2412: 2407: 2402: 2397: 2392: 2387: 2382: 2377: 2372: 2367: 2362: 2357: 2352: 2347: 2342: 2337: 2332: 2327: 2322: 2318: 2317: 2312: 2306: 2305: 2300: 2295: 2290: 2287: 2282: 2279: 2274: 2273:Laugerie-Haute 2271: 2269:Laugerie-Basse 2266: 2261: 2256: 2251: 2250:Abri Chadourne 2248: 2245: 2240: 2239: 2238: 2233: 2227: 2222: 2219: 2216: 2210: 2208: 2204: 2203: 2201: 2200: 2194: 2192: 2188: 2187: 2185: 2184: 2179: 2174: 2169: 2164: 2159: 2153: 2151: 2150:Czech Republic 2147: 2146: 2144: 2143: 2137: 2135: 2131: 2130: 2128: 2127: 2122: 2120:Romuald's Cave 2117: 2112: 2107: 2102: 2096: 2094: 2090: 2089: 2087: 2086: 2081: 2076: 2071: 2066: 2060: 2058: 2054: 2053: 2051: 2050: 2045: 2039: 2037: 2033: 2032: 2030: 2029: 2024: 2019: 2014: 2011: 2006: 2001: 1996: 1991: 1988: 1984: 1982: 1978: 1977: 1975: 1974: 1969: 1964: 1959: 1954: 1948: 1946: 1938: 1937: 1925: 1924: 1922: 1921: 1916: 1914:Cave paintings 1911: 1905: 1902: 1901: 1898:cave paintings 1893: 1892: 1885: 1878: 1870: 1861: 1860: 1858: 1857: 1845: 1833: 1820: 1817: 1816: 1814: 1813: 1808: 1807: 1806: 1796: 1791: 1786: 1780: 1778: 1774: 1773: 1771: 1770: 1768:Human timeline 1765: 1760: 1754: 1752: 1748: 1747: 1744: 1743: 1741: 1740: 1735: 1730: 1725: 1720: 1715: 1709: 1707: 1701: 1700: 1698: 1697: 1692: 1687: 1682: 1677: 1672: 1667: 1662: 1657: 1652: 1646: 1644: 1639: 1638: 1636: 1635: 1634: 1633: 1628: 1620: 1619: 1618: 1613: 1605: 1604: 1603: 1598: 1593: 1591:Drunken monkey 1585: 1584: 1583: 1578: 1573: 1564: 1562: 1558: 1557: 1555: 1554: 1549: 1544: 1539: 1534: 1529: 1523: 1521: 1520:General models 1514: 1510: 1509: 1507: 1506: 1464: 1462: 1456: 1455: 1452: 1451: 1448: 1447: 1444: 1443: 1441: 1440: 1435: 1430: 1425: 1420: 1413: 1408: 1399: 1397: 1386: 1380: 1379: 1377: 1376: 1368: 1361: 1354: 1346: 1339: 1332: 1324: 1319: 1311: 1309: 1307:Archaic humans 1303: 1302: 1299: 1298: 1296: 1295: 1288: 1281: 1274: 1267: 1260: 1253: 1246: 1238: 1236: 1228: 1227: 1225: 1224: 1216: 1212:H. rudolfensis 1208: 1201: 1194: 1185: 1179: 1172: 1158: 1157: 1154: 1153: 1151: 1150: 1143: 1136: 1133:P. aethiopicus 1128: 1126: 1118: 1117: 1115: 1114: 1107: 1100: 1093: 1086: 1079: 1072: 1064: 1062: 1054: 1053: 1051: 1050: 1043: 1035: 1033: 1025: 1024: 1022: 1021: 1014: 1011:Sahelanthropus 1007: 1000: 997:Nakalipithecus 992: 986: 980: 979: 977: 976: 971: 966: 961: 955: 953: 944: 932: 931: 924: 923: 916: 909: 901: 895: 894: 887: 886:External links 884: 881: 880: 858: 836: 814: 762: 736: 710: 649: 642: 612: 590: 568: 522: 521: 519: 516: 515: 514: 509: 504: 497: 494: 468: 465: 395:with distinct 377:right parietal 335: 332: 307: 304: 296:Southeast Asia 241: 240: 237: 236:Archaeologists 233: 232: 229: 225: 224: 220: 219: 216: 212: 211: 202: 198: 197: 188: 184: 183: 179: 178: 175: 171: 170: 167: 163: 162: 159: 155: 154: 151: 147: 146: 143: 139: 138: 106: 102: 101: 96: 92: 91: 78: 74: 73: 70: 66: 65: 56: 47: 46: 40: 39: 38: 37: 34: 33: 30: 15: 9: 6: 4: 3: 2: 6162: 6151: 6150:Caves of Laos 6148: 6146: 6143: 6141: 6138: 6136: 6133: 6131: 6128: 6126: 6123: 6122: 6120: 6110: 6100: 6098: 6093: 6088: 6087: 6084: 6071: 6068: 6066: 6063: 6061: 6058: 6057: 6055: 6051: 6045: 6042: 6040: 6037: 6035: 6032: 6031: 6029: 6027: 6024:Parts of the 6022: 6013: 6008: 6006: 6001: 5999: 5994: 5993: 5990: 5978: 5975: 5973: 5972:Cave painting 5970: 5968: 5967:List of caves 5965: 5962: 5961: 5958: 5944: 5941: 5940: 5938: 5934: 5928: 5925: 5924: 5922: 5918: 5912: 5909: 5908: 5906: 5902: 5896: 5893: 5892: 5890: 5888: 5884: 5878: 5875: 5873: 5870: 5869: 5867: 5863: 5857: 5854: 5853: 5851: 5849: 5848:New Caledonia 5845: 5839: 5836: 5835: 5833: 5829: 5823: 5820: 5818: 5815: 5813: 5810: 5809: 5807: 5805: 5801: 5795: 5792: 5790: 5787: 5785: 5782: 5780: 5777: 5775: 5772: 5770: 5767: 5765: 5762: 5760: 5757: 5755: 5754:New Guinea II 5752: 5750: 5747: 5745: 5742: 5740: 5737: 5735: 5732: 5730: 5729:Mannalargenna 5727: 5725: 5722: 5720: 5717: 5715: 5712: 5710: 5707: 5705: 5702: 5700: 5697: 5695: 5692: 5690: 5687: 5685: 5682: 5680: 5677: 5675: 5672: 5670: 5667: 5665: 5662: 5660: 5657: 5655: 5652: 5650: 5647: 5645: 5642: 5640: 5637: 5635: 5632: 5630: 5627: 5625: 5622: 5620: 5617: 5615: 5612: 5611: 5609: 5605: 5601: 5597: 5592: 5588: 5574: 5571: 5569: 5566: 5564: 5561: 5559: 5556: 5554: 5551: 5549: 5546: 5544: 5541: 5539: 5536: 5534: 5531: 5529: 5526: 5524: 5521: 5519: 5516: 5514: 5511: 5509: 5506: 5504: 5501: 5499: 5498:On Your Knees 5496: 5494: 5491: 5489: 5486: 5484: 5481: 5479: 5476: 5474: 5471: 5469: 5466: 5464: 5461: 5459: 5456: 5454: 5451: 5449: 5446: 5444: 5441: 5439: 5436: 5434: 5431: 5429: 5426: 5424: 5423:Hidden Valley 5421: 5419: 5416: 5414: 5411: 5409: 5406: 5404: 5401: 5399: 5396: 5394: 5391: 5389: 5386: 5384: 5381: 5379: 5376: 5374: 5371: 5369: 5366: 5364: 5361: 5359: 5356: 5354: 5351: 5349: 5346: 5344: 5341: 5339: 5336: 5334: 5331: 5330: 5328: 5326:United States 5324: 5318: 5315: 5314: 5312: 5308: 5302: 5299: 5297: 5294: 5292: 5289: 5287: 5284: 5282: 5279: 5277: 5274: 5273: 5271: 5267: 5261: 5258: 5256: 5253: 5251: 5248: 5246: 5245:Guilá Naquitz 5243: 5241: 5238: 5236: 5233: 5231: 5230:Dzibilchaltún 5228: 5226: 5223: 5221: 5218: 5217: 5215: 5211: 5205: 5202: 5201: 5199: 5195: 5189: 5186: 5185: 5183: 5179: 5173: 5170: 5169: 5167: 5165: 5161: 5155: 5152: 5150: 5147: 5145: 5142: 5140: 5137: 5135: 5132: 5130: 5127: 5126: 5124: 5120: 5114: 5111: 5109: 5106: 5104: 5101: 5099: 5096: 5094: 5091: 5089: 5086: 5085: 5083: 5079: 5073: 5070: 5068: 5065: 5064: 5062: 5058: 5052: 5049: 5047: 5044: 5043: 5041: 5037: 5031: 5028: 5026: 5023: 5021: 5018: 5016: 5013: 5011: 5010:Pedra Pintada 5008: 5006: 5003: 5001: 4998: 4997: 4995: 4991: 4985: 4982: 4980: 4977: 4975: 4972: 4971: 4969: 4965: 4959: 4956: 4955: 4953: 4949: 4943: 4940: 4939: 4937: 4933: 4929: 4925: 4924:South America 4921: 4916: 4912: 4898: 4895: 4894: 4892: 4888: 4882: 4879: 4877: 4874: 4873: 4871: 4867: 4861: 4858: 4857: 4855: 4851: 4845: 4842: 4840: 4837: 4835: 4832: 4831: 4829: 4825: 4819: 4816: 4814: 4811: 4809: 4806: 4804: 4801: 4799: 4796: 4794: 4791: 4789: 4788:Klasies River 4786: 4784: 4781: 4779: 4776: 4774: 4771: 4769: 4766: 4764: 4763:Byneskranskop 4761: 4759: 4756: 4754: 4751: 4749: 4746: 4743: 4742: 4739: 4736: 4734: 4731: 4729: 4726: 4724: 4721: 4719: 4716: 4714: 4711: 4709: 4706: 4704: 4701: 4699: 4696: 4694: 4691: 4689: 4686: 4684: 4681: 4679: 4676: 4674: 4671: 4669: 4666: 4664: 4661: 4660: 4658: 4654: 4648: 4645: 4643: 4640: 4638: 4635: 4634: 4632: 4628: 4622: 4619: 4618: 4616: 4612: 4606: 4603: 4601: 4598: 4597: 4595: 4591: 4585: 4582: 4581: 4579: 4575: 4569: 4566: 4564: 4561: 4559: 4556: 4554: 4553:Ifri Oudadane 4551: 4549: 4546: 4544: 4541: 4539: 4536: 4535: 4533: 4529: 4523: 4522:Uan Muhuggiag 4520: 4518: 4515: 4514: 4512: 4508: 4502: 4499: 4498: 4496: 4492: 4486: 4483: 4481: 4478: 4477: 4475: 4471: 4465: 4462: 4460: 4457: 4456: 4454: 4450: 4444: 4441: 4440: 4438: 4434: 4428: 4425: 4424: 4422: 4418: 4412: 4409: 4407: 4404: 4403: 4401: 4397: 4391: 4388: 4387: 4385: 4381: 4377: 4373: 4368: 4364: 4350: 4347: 4345: 4342: 4341: 4339: 4335: 4329: 4326: 4324: 4321: 4320: 4318: 4314: 4308: 4305: 4303: 4300: 4298: 4295: 4294: 4292: 4288: 4284: 4280: 4276: 4270: 4267: 4265: 4262: 4260: 4257: 4256: 4254: 4250: 4244: 4241: 4239: 4236: 4234: 4231: 4229: 4228:Batadombalena 4226: 4225: 4223: 4219: 4213: 4210: 4208: 4205: 4203: 4200: 4198: 4195: 4193: 4190: 4189: 4187: 4183: 4177: 4174: 4173: 4171: 4169: 4165: 4159: 4156: 4155: 4153: 4149: 4143: 4140: 4139: 4137: 4133: 4127: 4124: 4123: 4121: 4117: 4111: 4108: 4107: 4105: 4101: 4095: 4092: 4090: 4087: 4085: 4084:Ras Baalbek I 4082: 4080: 4077: 4075: 4072: 4070: 4067: 4065: 4062: 4061: 4059: 4055: 4049: 4046: 4044: 4041: 4040: 4038: 4034: 4028: 4025: 4024: 4022: 4018: 4012: 4009: 4007: 4004: 4002: 3999: 3997: 3994: 3992: 3989: 3987: 3984: 3982: 3979: 3977: 3974: 3973: 3971: 3967: 3961: 3958: 3956: 3953: 3951: 3948: 3946: 3943: 3941: 3938: 3936: 3933: 3931: 3928: 3926: 3923: 3921: 3920:Nahal Me'arot 3918: 3916: 3913: 3911: 3908: 3906: 3903: 3901: 3898: 3896: 3893: 3892: 3890: 3886: 3880: 3877: 3875: 3872: 3871: 3869: 3865: 3859: 3856: 3854: 3851: 3849: 3846: 3844: 3841: 3839: 3836: 3834: 3831: 3829: 3826: 3824: 3821: 3819: 3816: 3815: 3813: 3809: 3803: 3800: 3798: 3795: 3793: 3790: 3788: 3785: 3784: 3782: 3778: 3772: 3769: 3767: 3764: 3762: 3759: 3757: 3754: 3752: 3749: 3747: 3744: 3742: 3739: 3738: 3736: 3732: 3726: 3723: 3722: 3720: 3716: 3710: 3707: 3705: 3702: 3700: 3697: 3696: 3694: 3690: 3684: 3681: 3679: 3676: 3674: 3671: 3669: 3666: 3664: 3661: 3659: 3656: 3654: 3651: 3649: 3646: 3644: 3641: 3639: 3636: 3635: 3633: 3629: 3623: 3620: 3619: 3617: 3613: 3607: 3604: 3602: 3599: 3597: 3594: 3592: 3589: 3587: 3584: 3582: 3579: 3577: 3574: 3573: 3571: 3567: 3561: 3558: 3557: 3555: 3551: 3545: 3542: 3541: 3539: 3535: 3531: 3527: 3522: 3518: 3504: 3503:Windmill Hill 3501: 3499: 3496: 3494: 3491: 3489: 3486: 3484: 3481: 3479: 3476: 3474: 3471: 3469: 3466: 3464: 3461: 3459: 3456: 3454: 3451: 3449: 3448:Heathery Burn 3446: 3444: 3441: 3439: 3436: 3434: 3431: 3429: 3426: 3424: 3421: 3419: 3416: 3415: 3413: 3409: 3403: 3400: 3399: 3397: 3393: 3387: 3384: 3382: 3379: 3378: 3376: 3372: 3366: 3363: 3361: 3358: 3356: 3353: 3351: 3348: 3346: 3343: 3341: 3338: 3336: 3333: 3331: 3328: 3326: 3323: 3321: 3318: 3316: 3313: 3311: 3308: 3307: 3304: 3301: 3299: 3296: 3294: 3291: 3289: 3286: 3284: 3281: 3279: 3276: 3274: 3271: 3269: 3266: 3264: 3261: 3259: 3256: 3254: 3251: 3249: 3246: 3244: 3241: 3238: 3237: 3234: 3231: 3229: 3226: 3224: 3221: 3220: 3217: 3214: 3212: 3209: 3206: 3203: 3202: 3199: 3196: 3194: 3191: 3189: 3186: 3184: 3183:Tito Bustillo 3181: 3176: 3172: 3169: 3166: 3164:Las Chimeneas 3163: 3161: 3158: 3157: 3156: 3153: 3150: 3148: 3145: 3142: 3140: 3137: 3136: 3135: 3132: 3130: 3127: 3125: 3122: 3121: 3119: 3115: 3109: 3106: 3104: 3101: 3099: 3096: 3094: 3091: 3090: 3088: 3084: 3078: 3075: 3073: 3070: 3068: 3065: 3064: 3062: 3058: 3052: 3049: 3047: 3044: 3042: 3039: 3037: 3034: 3033: 3031: 3027: 3021: 3018: 3016: 3013: 3011: 3008: 3006: 3003: 3001: 2998: 2996: 2993: 2991: 2990:Akhshtyrskaya 2988: 2987: 2985: 2981: 2975: 2972: 2970: 2967: 2965: 2962: 2960: 2957: 2956: 2954: 2950: 2944: 2941: 2939: 2938:Pego do Diabo 2936: 2934: 2931: 2929: 2926: 2924: 2921: 2919: 2916: 2914: 2911: 2909: 2906: 2905: 2903: 2899: 2893: 2890: 2888: 2885: 2884: 2882: 2878: 2872: 2869: 2868: 2866: 2862: 2856: 2853: 2852: 2850: 2848: 2844: 2838: 2835: 2834: 2832: 2828: 2822: 2819: 2818: 2816: 2812: 2806: 2803: 2802: 2800: 2796: 2790: 2787: 2786: 2784: 2782: 2778: 2772: 2769: 2767: 2764: 2762: 2759: 2757: 2756: 2752: 2750: 2747: 2745: 2742: 2740: 2737: 2735: 2732: 2730: 2729:Arene Candide 2727: 2725: 2722: 2721: 2719: 2715: 2709: 2706: 2704: 2701: 2700: 2698: 2694: 2688: 2685: 2683: 2680: 2678: 2675: 2673: 2670: 2668: 2665: 2663: 2660: 2658: 2655: 2653: 2650: 2648: 2645: 2643: 2640: 2639: 2637: 2633: 2627: 2624: 2622: 2619: 2617: 2616:Devil's Tower 2614: 2613: 2611: 2609: 2605: 2599: 2596: 2594: 2591: 2589: 2586: 2584: 2581: 2579: 2576: 2573: 2572: 2569: 2566: 2564: 2561: 2559: 2556: 2554: 2551: 2549: 2546: 2544: 2541: 2539: 2536: 2535: 2533: 2529: 2523: 2520: 2518: 2515: 2513: 2510: 2508: 2505: 2503: 2500: 2498: 2495: 2493: 2490: 2488: 2485: 2483: 2480: 2478: 2475: 2473: 2470: 2468: 2465: 2463: 2460: 2457: 2455: 2452: 2450: 2447: 2444: 2441: 2438: 2435: 2433: 2430: 2428: 2425: 2423: 2420: 2419: 2416: 2413: 2411: 2408: 2406: 2403: 2401: 2398: 2396: 2393: 2391: 2390:Pair-non-Pair 2388: 2386: 2383: 2381: 2378: 2376: 2375:Le Mas-d'Azil 2373: 2371: 2368: 2366: 2363: 2361: 2358: 2356: 2353: 2351: 2348: 2346: 2343: 2341: 2338: 2336: 2333: 2331: 2328: 2326: 2325:Arcy-sur-Cure 2323: 2320: 2319: 2316: 2313: 2311: 2308: 2307: 2304: 2301: 2299: 2296: 2294: 2291: 2288: 2286: 2283: 2280: 2278: 2275: 2272: 2270: 2267: 2265: 2264:Font-de-Gaume 2262: 2260: 2257: 2255: 2252: 2249: 2246: 2244: 2241: 2237: 2234: 2232:Abri Castanet 2231: 2230: 2229:Castel Merle 2228: 2226: 2223: 2220: 2217: 2215: 2212: 2211: 2209: 2205: 2199: 2196: 2195: 2193: 2189: 2183: 2180: 2178: 2175: 2173: 2170: 2168: 2165: 2163: 2160: 2158: 2155: 2154: 2152: 2148: 2142: 2139: 2138: 2136: 2132: 2126: 2123: 2121: 2118: 2116: 2113: 2111: 2108: 2106: 2103: 2101: 2098: 2097: 2095: 2091: 2085: 2082: 2080: 2077: 2075: 2072: 2070: 2067: 2065: 2062: 2061: 2059: 2055: 2049: 2046: 2044: 2041: 2040: 2038: 2034: 2028: 2025: 2023: 2020: 2018: 2015: 2012: 2010: 2007: 2005: 2002: 2000: 1997: 1995: 1992: 1989: 1986: 1985: 1983: 1979: 1973: 1970: 1968: 1965: 1963: 1960: 1958: 1955: 1953: 1950: 1949: 1947: 1943: 1939: 1935: 1930: 1926: 1920: 1917: 1915: 1912: 1910: 1907: 1906: 1903: 1899: 1891: 1886: 1884: 1879: 1877: 1872: 1871: 1868: 1856: 1855: 1850: 1846: 1844: 1843: 1834: 1832: 1831: 1822: 1821: 1818: 1812: 1809: 1805: 1802: 1801: 1800: 1797: 1795: 1792: 1790: 1787: 1785: 1782: 1781: 1779: 1775: 1769: 1766: 1764: 1761: 1759: 1756: 1755: 1753: 1749: 1739: 1736: 1734: 1731: 1729: 1726: 1724: 1721: 1719: 1716: 1714: 1711: 1710: 1708: 1706: 1702: 1696: 1693: 1691: 1688: 1686: 1683: 1681: 1678: 1676: 1673: 1671: 1668: 1666: 1663: 1661: 1658: 1656: 1653: 1651: 1648: 1647: 1645: 1640: 1632: 1629: 1627: 1624: 1623: 1622:Life history 1621: 1617: 1614: 1612: 1609: 1608: 1606: 1602: 1599: 1597: 1594: 1592: 1589: 1588: 1586: 1582: 1579: 1577: 1574: 1572: 1569: 1568: 1566: 1565: 1563: 1559: 1553: 1550: 1548: 1545: 1543: 1540: 1538: 1535: 1533: 1530: 1528: 1525: 1524: 1522: 1518: 1515: 1511: 1505: 1504: 1499: 1495: 1494: 1489: 1488: 1483: 1482: 1477: 1476: 1475:Homo ergaster 1471: 1470: 1466: 1465: 1463: 1461: 1457: 1439: 1436: 1434: 1431: 1429: 1426: 1424: 1421: 1419: 1418: 1414: 1412: 1409: 1407: 1405: 1404:H. s. sapiens 1401: 1400: 1398: 1396: 1395: 1390: 1387: 1385: 1384:Modern humans 1381: 1374: 1373: 1369: 1367: 1366: 1362: 1360: 1359: 1358:H. luzonensis 1355: 1352: 1351: 1347: 1345: 1344: 1340: 1338: 1337: 1333: 1330: 1329: 1325: 1323: 1320: 1318: 1317: 1316:H. antecessor 1313: 1312: 1310: 1308: 1304: 1294: 1293: 1289: 1287: 1286: 1282: 1280: 1279: 1275: 1273: 1272: 1268: 1266: 1265: 1261: 1259: 1258: 1254: 1252: 1251: 1247: 1245: 1244: 1243:H. e. erectus 1240: 1239: 1237: 1235: 1234: 1229: 1222: 1221: 1217: 1214: 1213: 1209: 1207: 1206: 1202: 1200: 1199: 1195: 1192: 1191: 1187: 1186: 1183: 1180: 1176: 1173: 1171: 1169: 1159: 1149: 1148: 1144: 1142: 1141: 1137: 1135: 1134: 1130: 1129: 1127: 1125: 1124: 1119: 1113: 1112: 1108: 1106: 1105: 1101: 1099: 1098: 1097:A. deyiremeda 1094: 1092: 1091: 1087: 1085: 1084: 1080: 1078: 1077: 1073: 1071: 1070: 1066: 1065: 1063: 1061: 1060: 1055: 1049: 1048: 1044: 1042: 1041: 1037: 1036: 1034: 1032: 1031: 1026: 1020: 1019: 1018:Kenyanthropus 1015: 1013: 1012: 1008: 1006: 1005: 1001: 999: 998: 994: 993: 990: 987: 985: 981: 975: 972: 970: 967: 965: 964:Gorilla–human 962: 960: 957: 956: 954: 952: 948: 945: 942: 937: 933: 929: 922: 917: 915: 910: 908: 903: 902: 899: 893: 890: 889: 868: 862: 846: 840: 824: 818: 810: 806: 801: 796: 791: 786: 782: 778: 774: 766: 750: 743: 741: 725: 721: 714: 699: 695: 690: 685: 680: 675: 671: 667: 660: 653: 645: 639: 635: 631: 627: 623: 616: 600: 599:"Tam Pa Ling" 594: 578: 572: 564: 560: 556: 552: 548: 544: 536: 534: 532: 530: 528: 523: 513: 510: 508: 505: 503: 500: 499: 493: 491: 487: 483: 478: 475: 464: 462: 458: 454: 450: 446: 441: 439: 435: 431: 430:archaic human 427: 423: 416:TPL2 mandible 414: 410: 408: 404: 403: 398: 394: 390: 386: 382: 381:temporal bone 378: 374: 370: 367:includes the 366: 362: 360: 356: 352: 348: 344: 339: 331: 329: 325: 321: 317: 313: 303: 301: 297: 293: 289: 285: 281: 277: 273: 269: 266: 261: 259: 255: 251: 247: 238: 234: 230: 226: 221: 217: 213: 210: 206: 203: 199: 196: 192: 189: 185: 180: 176: 172: 168: 164: 160: 156: 152: 148: 144: 140: 135: 107: 103: 100: 97: 93: 90: 86: 82: 79: 75: 71: 67: 60: 44: 35: 28: 23: 6109:Paleontology 5779:Wargata Mina 5739:Murrawijinie 5674:Eagles Reach 5669:Devil's Lair 5573:Wilson Butte 5358:Cherry Creek 5348:Bull Thistle 5093:Chiribiquete 5051:Charlie Lake 4979:Barton Creek 4793:Melkhoutboom 4733:Sterkfontein 4723:Plovers Lake 4656:South Africa 4558:Jebel Irhoud 4543:Ifri N'Ammar 4278:Turkmenistan 4243:Hunugalagala 4047: 4027:Iraq ed-Dubb 3792:Jeriji Saléh 3453:High Pasture 3010:Shulgan-Tash 2933:Pedra Furada 2754: 2593:Lichtenstein 2583:Brillenhöhle 2512:Le Regourdou 2472:La Ferrassie 2467:Combe Grenal 2410:Trois Frères 2345:Fontéchevade 2298:La Madeleine 1952:Drachenhöhle 1852: 1840: 1828: 1695:Gender roles 1690:Intelligence 1503:Homo sapiens 1501: 1497: 1491: 1485: 1481:Homo erectus 1479: 1473: 1469:Homo habilis 1467: 1432: 1428:Manot people 1417:H. s. idaltu 1415: 1411:Jebel Irhoud 1403: 1394:Homo sapiens 1392: 1370: 1363: 1356: 1348: 1341: 1334: 1326: 1314: 1290: 1283: 1276: 1269: 1262: 1255: 1248: 1241: 1233:Homo erectus 1231: 1218: 1210: 1203: 1196: 1188: 1178:Proto-humans 1167: 1164:proto-humans 1145: 1138: 1131: 1123:Paranthropus 1121: 1109: 1102: 1095: 1088: 1083:A. anamensis 1081: 1076:A. africanus 1074: 1069:A. afarensis 1067: 1057: 1045: 1038: 1030:Ardipithecus 1028: 1016: 1009: 1002: 995: 974:Gibbon–human 871:. Retrieved 861: 849:. Retrieved 839: 827:. Retrieved 817: 780: 776: 765: 753:. Retrieved 728:. Retrieved 723: 713: 701:. Retrieved 669: 665: 652: 625: 615: 603:. Retrieved 593: 581:. Retrieved 571: 546: 542: 479: 473: 470: 467:Significance 460: 456: 452: 448: 444: 442: 425: 421: 419: 402:Homo sapiens 400: 364: 363: 346: 342: 340: 337: 309: 291: 290:traits; and 279: 271: 262: 249: 245: 244: 231:2008-ongoing 218:Paleo-humans 158:Part of 57:location in 6065:Border Cave 6044:Sibudu Cave 5865:New Zealand 5689:Gwion Gwion 5558:Trail Creek 5483:Meadowcroft 5443:Last Supper 5373:Daugherty's 5235:Las Flechas 5020:Rei do Mato 4744:Other caves 4728:Rising Star 4703:Makapansgat 4668:Bolt's Farm 4485:Njoro River 4328:Teshik-Tash 4323:Obi-Rakhmat 4307:Yarımburgaz 4207:Lapuz Lapuz 4185:Philippines 4094:Nachcharini 4089:Ras El Kelb 4048:Tam Pa Ling 4043:Tham An Mah 3925:Nahal Hemar 3673:Zhoukoudian 3648:Guanyindong 3622:Laang Spean 3544:Darra-e Kur 3537:Afghanistan 3483:Portbraddon 3473:Marble Arch 3386:Wildkirchli 3374:Switzerland 3310:Other caves 3283:Ojo Guareña 3263:Maltravieso 3258:Los Casares 3233:Siega Verde 3198:Los Aviones 3193:Santimamiñe 3167:Las Monedas 3160:El Castillo 3067:Čertova pec 3015:Mezmaiskaya 2923:Lagar Velho 2855:Cocev Kamen 2734:Balzi Rossi 2574:Other caves 2563:Sirgenstein 2492:Le Moustier 2482:Fontbrégoua 2442:Bonne-Femme 2422:Other caves 2141:Aetokremnos 2084:Utroba Cave 1990:Claminforge 1987:Belle-Roche 1626:Grandmother 1581:Shore-based 1542:Aquatic ape 1433:Tam Pa Ling 1328:H. ergaster 1147:P. robustus 507:Laang Spean 502:Tham An Mah 351:Radiocarbon 316:dissolution 246:Tam Pa Ling 129: / 117:103°24′35″E 105:Coordinates 83:, northern 6119:Categories 5977:Speleology 5794:Wellington 5749:Naracoorte 5734:Mudgegonga 5719:Madjedbebe 5684:Gabarnmung 5624:Ballawinne 5614:Abrakurrie 5513:Pictograph 5291:Qillqatani 5286:Pikimachay 5281:Guitarrero 5276:Chivateros 5255:Oxtotitlán 5108:Tequendama 4818:Wonderwerk 4798:Nelson Bay 4778:Elands Bay 4738:Swartkrans 4683:Gladysvale 4630:Somaliland 4577:Mozambique 4517:Haua Fteah 4316:Uzbekistan 3981:Minatogawa 3976:Kosegasawa 3930:Nahal Oren 3874:Hazar Merd 3843:Qaleh Bozi 3828:Do-Ashkaft 3725:Satsurblia 3692:East Timor 3569:Azerbaijan 3458:Kendrick's 3423:Bontnewydd 3345:Don Gaspar 3298:Praileaitz 3253:Las Caldas 3171:La Pasiega 3098:Divje Babe 3020:Okladnikov 3005:Ignatievka 2928:Pala Pinta 2837:Għar Dalam 2814:Luxembourg 2789:St Brelade 2771:Villabruna 2682:Stravomyti 2672:Megalakkos 2657:Eileithyia 2642:Alepotrypa 2553:Hohle Fels 2439:Balauzière 2395:Pech Merle 2303:Rouffignac 2277:La Micoque 2259:Cro-Magnon 2218:Bara Bahau 2162:Býčí skála 2110:Vela Spila 2069:Devetashka 2064:Bacho Kiro 1665:Skin color 1650:Bipedalism 1611:Killer ape 1423:Cro-Magnon 1322:Denisovans 1198:H. habilis 1162:Humans and 1047:A. ramidus 1040:A. kadabba 873:August 21, 851:August 21, 829:August 21, 755:August 21, 730:2016-08-21 703:August 23, 605:August 21, 583:August 21, 549:: 93–106. 518:References 371:, partial 298:by 60,000 223:Site notes 114:20°12′31″N 5943:Nanumanga 5927:Falemauga 5838:Makauwahi 5607:Australia 5563:Trinchera 5438:La Grange 5408:Gatecliff 5403:Franktown 5398:Fort Rock 5378:Dry Creek 5296:Toquepala 5260:Sac Actun 5250:Ox Bel Ha 5240:Frightful 5225:Coxcatlan 5204:Long Mile 4958:Quadiriki 4935:Argentina 4773:Diepkloof 4758:Boomplaas 4713:Minnaar's 4698:Kromdraai 4647:Laas Geel 4642:Dhaymoole 4637:Dhambalin 4600:Apollo 11 4501:Liphofung 4427:Shum Laka 4390:Gueldaman 4344:Con Moong 4221:Sri Lanka 4168:Palestine 4142:Padah-Lin 4079:Ksar Akil 4011:Yamashita 3996:Pinza-Abu 3802:Pettakere 3797:Liang Bua 3780:Indonesia 3766:Pahargarh 3751:Bhimbetka 3709:Lene Hara 3699:Jerimalai 3678:Zengpiyan 3668:Yuchanyan 3468:Long Hole 3365:del Valle 3340:dels Bous 3293:la Pileta 3228:Atapuerca 3143:Covalanas 3041:Pešturina 2969:Muierilor 2959:Coliboaia 2871:Svarthola 2687:Theopetra 2677:Petralona 2662:Franchthi 2652:Asphendou 2608:Gibraltar 2578:Baumann's 2568:Vogelherd 2543:Bockstein 2522:Vallonnet 2517:Rochereil 2507:Raymonden 2497:Noisetier 2449:Bruniquel 2370:Marsoulas 2360:Lombrives 2355:La Marche 2281:La Mouthe 2247:Abri Audi 2225:Cap Blanc 2167:Koněprusy 2157:Amatérská 2125:Veternica 2074:Kozarnika 2048:Ledenjača 1972:Tischofer 1962:Lurgrotte 1784:Theorists 1751:Timelines 1631:Patriarch 1607:Behavior 1532:Gathering 1460:Ancestors 1205:H. naledi 1140:P. boisei 1111:A. sediba 563:1040-6182 490:Vientiane 389:dentition 373:occipital 320:limestone 191:limestone 89:Vientiane 5784:Warratyi 5769:Tarragal 5759:Nunamira 5744:Murujuga 5709:Kutikina 5704:Koonalda 5699:Koongine 5659:Cliefden 5654:Cave Bay 5553:Tomaquag 5538:Sisyphus 5468:Mantle's 5458:Lovelock 5453:LoDaisKa 5433:Humboldt 5317:Werehpai 5310:Suriname 5220:Chan Hol 5139:Centella 5129:Ambrosio 5081:Colombia 5046:Bluefish 4890:Zimbabwe 4827:Tanzania 4813:Stadsaal 4718:Motsetsi 4688:Gondolin 4678:Drimolen 4673:Cooper's 4568:Taforalt 4538:Hercules 4464:Swimmers 4436:DR Congo 4420:Cameroon 4399:Botswana 4269:Tham Lod 4259:Pha Taem 4252:Thailand 4233:Belilena 4151:Pakistan 4119:Mongolia 4103:Malaysia 4064:Antelias 4006:Tobayama 3960:Zuttiyeh 3879:Shanidar 3741:Ambadevi 3658:Tianyuan 3638:Baishiya 3615:Cambodia 3355:El Mirón 3350:Guanches 3325:Armintxe 3243:Bacinete 3188:Altxerri 3177:El Pendo 3147:La Garma 3129:Altamira 3086:Slovenia 3077:Jasovská 3060:Slovakia 3046:Risovača 3000:Denisova 2964:Cuciulat 2918:Furninha 2913:Escoural 2901:Portugal 2892:Paradise 2887:Obłazowa 2821:Loschbur 2761:Paglicci 2744:Corbeddu 2626:Vanguard 2621:Gorham's 2502:La Quina 2445:Bouillon 2432:Aurignac 2236:Reverdit 2221:Bernifal 2100:Grapčeva 2057:Bulgaria 2017:Scladina 2004:Naulette 1967:Salzofen 1830:Category 1685:Language 1655:Skeleton 1350:H. longi 1104:A. garhi 941:Hominins 936:Taxonomy 809:25849125 777:PLOS ONE 698:22908291 496:See also 385:maxillae 284:mandible 187:Material 142:Altitude 77:Location 6083:Portals 6060:Blombos 5895:Chugai' 5877:Ruakuri 5812:Gadao's 5789:Warreen 5724:Mammoth 5694:Jenolan 5639:Blanche 5619:Acheron 5596:Oceania 5568:Ventana 5548:Tainter 5523:Russell 5508:Pendejo 5503:Paisley 5463:Mammoth 5383:Durango 5343:Bonfire 5197:Jamaica 5164:Curaçao 5103:Sáchica 5088:El Abra 5072:Milodón 5015:Peruaçu 5005:Maquiné 4897:Bambata 4876:Kalemba 4748:Blombos 4693:Haasgat 4614:Nigeria 4593:Namibia 4531:Morocco 4494:Lesotho 4406:Tsodilo 4383:Algeria 4337:Vietnam 4297:Belbaşı 4283:Dzhebel 4238:Fa Hien 4197:Kalanay 4158:Sanghao 4135:Myanmar 4074:Kaukaba 4057:Lebanon 3945:Raqefet 3915:Misliya 3900:HaYonim 3848:Warwasi 3823:Darband 3818:Bisitun 3787:Harimau 3761:Gudiyam 3756:Edakkal 3718:Georgia 3663:Xianren 3591:Damjili 3586:Buzeyir 3560:Areni-1 3553:Armenia 3478:Oldbury 3443:Gough's 3428:Cathole 3402:Verteba 3395:Ukraine 2952:Romania 2943:Salemas 2908:Aroeira 2739:Cavallo 2724:Addaura 2703:Baradla 2696:Hungary 2647:Apidima 2531:Germany 2487:Lazaret 2415:Villars 2335:Cosquer 2315:Chauvet 2293:Lascaux 2191:Finland 2115:Vindija 2105:Krapina 2093:Croatia 2013:Ramioul 1981:Belgium 1957:Gudenus 1945:Austria 1842:Commons 1794:Fossils 1660:Muscles 1571:Cooking 1527:Hunting 1004:Orrorin 800:4388508 689:3437904 369:frontal 334:Fossils 328:Permian 288:archaic 268:fossils 265:hominin 201:Periods 182:History 5936:Tuvalu 5872:Moncks 5856:Pindai 5831:Hawaii 5822:Talagi 5817:Mahlac 5679:Fossil 5664:Cloggs 5649:Burrup 5528:Sandia 5473:Marmes 5418:Hidden 5413:Graham 5368:Danger 5353:Burnet 5213:Mexico 5188:Pomier 5149:Patana 5134:Calero 5113:Tibitó 5039:Canada 5000:Gentio 4993:Brazil 4967:Belize 4881:Mumbwa 4869:Zambia 4853:Uganda 4834:Kondoa 4808:Sibudu 4753:Border 4708:Malapa 4584:Ngalue 4459:Beasts 4443:Matupi 4372:Africa 4302:Karain 4290:Turkey 4264:Spirit 4202:Minori 4192:Callao 4176:Shuqba 4020:Jordan 3991:Ōzakai 3986:Muroya 3935:Qafzeh 3905:Kebara 3888:Israel 3858:Yafteh 3853:Wezmeh 3683:Zhiren 3596:Tağlar 3498:Thor's 3433:Coygan 3381:Bichon 3335:Bedmar 3303:Sidrón 3139:Chufín 3072:Domica 3029:Serbia 2983:Russia 2880:Poland 2864:Norway 2798:Kosovo 2781:Jersey 2766:Romito 2708:Szelim 2667:Kleidi 2635:Greece 2340:Cussac 2330:Gargas 2285:Pataud 2207:France 2177:Mladeč 2134:Cyprus 2079:Magura 2043:Badanj 2036:Bosnia 1934:Europe 1777:Others 1680:Speech 1642:Topics 1587:Drugs 1513:Models 807:  797:  696:  686:  640:  561:  379:, and 263:Three 166:Length 95:Region 6053:Other 5920:Samoa 5774:Ubirr 5764:ORS 7 5533:Shoup 5493:Mummy 5488:Modoc 5478:Martz 5428:Hogup 5338:Baker 5154:Pluma 5060:Chile 5025:Santo 4951:Aruba 4920:North 4860:Nyero 4844:Mumba 4768:Cango 4510:Libya 4473:Kenya 4452:Egypt 4349:Ngườm 4212:Tabon 4069:Jeita 3969:Japan 3955:Tabun 3950:Skhul 3940:Qesem 3910:Manot 3771:Tikla 3746:Belum 3734:India 3704:Laili 3653:Luobi 3643:Fuyan 3631:China 3606:Gazma 3581:Azykh 3576:Allar 3463:Kents 3330:Axlor 3320:Antón 3315:Ángel 3273:Nerja 3211:Araña 3117:Spain 3108:Potok 3103:Pekel 3093:Betal 2830:Malta 2755:Felci 2717:Italy 2598:Ofnet 2458:Cauna 2454:Calès 2427:Arago 2405:Renne 2385:Niaux 2182:Šipka 2172:Kůlna 1999:Goyet 1994:Engis 1789:Books 1567:Diet 662:(PDF) 312:karst 195:karst 174:Width 5963:Cave 5911:Kilu 5804:Guam 5644:Bone 5448:Levi 5388:Dust 5269:Peru 5172:Hato 5144:Cura 5122:Cuba 5067:Fell 4922:and 4839:Bahi 4110:Niah 4036:Laos 3895:Amud 3867:Iraq 3811:Iran 3526:Asia 3493:Sand 3278:Niño 2974:Oase 2749:Deer 2477:Fées 2198:Wolf 1670:Hair 1490:) → 1168:Homo 875:2016 853:2016 831:2016 805:PMID 757:2016 705:2016 694:PMID 638:ISBN 607:2016 585:2016 559:ISSN 474:TPL1 461:TPL3 457:TPL3 453:TPL3 449:TPL2 445:TPL3 438:chin 426:TPL1 422:TPL2 420:The 365:TPL1 353:and 347:TPL2 343:TPL1 326:and 292:TPL3 282:, a 280:TPL2 272:TPL1 258:Laos 153:Cave 150:Type 85:Laos 59:Laos 4621:Rop 3601:Zar 2436:Azé 2022:Spy 1484:(→ 1375:(?) 1353:(?) 1331:(?) 1223:(?) 1215:(?) 1193:(?) 795:PMC 785:doi 684:PMC 674:doi 670:109 630:doi 551:doi 547:466 488:in 405:in 318:of 6121:: 1500:→ 1496:→ 1478:/ 1472:→ 803:. 793:. 781:10 779:. 775:. 739:^ 722:. 692:. 682:. 668:. 664:. 636:. 624:. 557:. 545:. 526:^ 492:. 451:, 440:. 409:. 375:, 302:. 300:BP 278:; 207:, 193:, 6085:: 6011:e 6004:t 5997:v 3207:) 1889:e 1882:t 1875:v 1170:) 1166:( 943:) 939:( 920:e 913:t 906:v 877:. 855:. 833:. 811:. 787:: 759:. 733:. 707:. 676:: 646:. 632:: 609:. 587:. 565:. 553:: 248:(

Index

Tam Pa Ling Cave Laos
Tam Pa Ling cave Laos
Laos
Houaphanh Province
Laos
Vientiane
Annamite Mountains
20°12′31″N 103°24′35″E / 20.20861°N 103.40972°E / 20.20861; 103.40972
limestone
karst
Upper Paleolithic
Middle Paleolithic
Annamite Mountains
Laos
hominin
fossils
anatomically modern human
mandible
archaic
Southeast Asia
BP
karst
dissolution
limestone
Upper Carboniferous
Permian
Radiocarbon
luminescence dating
uranium-thorium dating
frontal

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