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Wallowing

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78: 336:) spends a large part of its day wallowing. When mud holes are unavailable, the rhino will deepen puddles with its feet and horns. One 20-month study of wallowing behaviour found they will visit no more than three wallows at any given time. After two to 12 weeks using a particular wallow, the rhino will abandon it. Typically, the rhino will wallow around midday for two to three hours at a time before foraging for food. Although in zoos the Sumatran rhino has been observed wallowing less than 45 minutes a day, the study of wild animals found 80–300 minutes per day spent in wallows. Captive individuals deprived of adequate wallowing have quickly developed broken and inflamed skins, 293: 355: 301:
after wallowing, the wet mud provides a cooling, and probably protecting, layer on the body. When pigs enter a wallow, they normally dig and root in the mud before entering with the fore-body first. They then wriggle the body back and forth, and rub their faces in the mud so all of the body surface is covered. Before they leave the wallow, they often shake their heads and body, often finishing with rubbing against a tree or a stone next to the wallow. When indoors and hot, domestic pigs often attempt to wallow on wet floor surfaces and in the dunging areas.
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Although temperature regulation seems to be the main motivation for wallowing in pigs, they will still wallow in colder weather. While many have suggested that pigs wallow in mud because of a lack of sweat glands, pigs and other wallowing animals may have not evolved functional sweat glands because
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Pigs lack functional sweat glands and are almost incapable of panting. To thermoregulate, they rely on wallowing in water or mud to cool the body. Adult pigs under natural or free-range conditions can often be seen to wallow when air temperature exceeds 20 °C. Mud is the preferred substrate;
640: 65:. Wallowing is often combined with other behaviours to fulfil its purpose; for example, elephants will often blow dirt over themselves after wallowing to create a thicker "coating", or pigs will allow the mud to dry before rubbing themselves on a tree or rock to remove 379:), for example, do not wallow. Even within the red deer species, there is variation between sub-species and breeds in wallowing behaviour. For example, although wapiti do wallow, they and crossbreds are less inclined to wallow than European red deer. 362:
Many deer perform wallowing, creating wallow sites in wet depressions in the ground, eventually forming quite large sites (2–3 m across and up to 1 m deep). However, it has been claimed that only some species of deer wallow;
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Pal, B.C. and Bhattacharyya, A., (1986). Wallowing behavior and wallows used by great Indian one-horned Rhinoceros at Garumara and Jaldapara wildlife sanctuaries, West Bengal, India. Proc. Zoologic. Soc., Calcutta, 35:
316:. It has been argued that wallowing behaviour and the desire to be in shallow, murky water could have been a step to the evolution of whales and other marine mammals from land-dwelling mammals. 725: 755: 112:
Many functions of wallowing have been proposed although not all have been tested by rigorous scientific investigation. Proposed functions include:
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Rhythm of daily activity and behavior of European bison in the Bialowieze Forest in the period without snow cover
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Ng, J., S.C.; Zainal-Zahari, Z.; Nordin, A. (2001). "Wallows and wallow utilization of the Sumatran rhinoceros (
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Wallowing behavior of American bison (Bos bison) in Tallgrass Prairie: An examination of alternate explanations
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Vestergaard, K.S. and Bjerg, B., (1996). Wallowing behavior in fattening pigs, In: I.J.H. Duncan,
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Review of wallowing in pigs: Description of the behaviour and its motivational basis
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Cabon-Racyznska, R.M., Krasinska, Z.A., Krasinska, J.M. and Wojcik, J.M. (1987).
603: 581:(1982). Adaptive behavioral strategies, In: J.W. Thomas and D.E. Toweill, (Eds). 1345: 1285: 1270: 1255: 1240: 1209: 1162: 828: 729: 658: 940:) in a natural enclosure in Sungai Dusun Wildlife Reserve, Selangor, Malaysia". 1310: 1265: 1260: 1142: 1050: 578: 215: 203: 168: 81: 1445: 1315: 1305: 1300: 1295: 1224: 1157: 1065: 894: 618: 270: 180: 120: 66: 693: 595: 1340: 1290: 1250: 1219: 1172: 707:
Behavior of the endangered tamaraw (Bubalus mindorensis Heude) in captivity
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Zhi-Tao, L., Jan-Hua, D., Yan-Ling, S., Zhi-Gao, Z. and Qiong, Z. (2007).
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Tick defense strategies in bison: the role of grooming and hair coat
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Proc. Thirtieth International Cong. International Soc. Appl. Ethol.
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Social behavior of the American buffalo (Bison bison bison)
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Quantitative analysis of wallowing in a confined bison herd
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Deer of the World: Their Evolution, Behaviour, and Ecology
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White rhinoceroses (Ceratotherium simum) wallowing in mud.
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McMillan, B.R., Cottam, M.R. and Kaufman, D.W. (2000).
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Stackpole Books, Harrisburg, Pennsylvania. pp. 219-278
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wallowing was a part of their behavioural repertoire.
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male during the rut and its function in reproduction
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History, range, and home life of the northern bison
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The American Midland Naturalist, 144:X 506: 434: 350:Self-anointing in animals § Ungulates 820: 256:(preventing dehydration) – hippopotamus ( 705:Momongan, V.G. and Walde, G.I., (1993). 639:Mooring, M.S. and Samuel, W.M., (1998). 596:Wallowing behavior of Hainan Eld’s deer 353: 323: 291: 94: 76: 29: 929: 493:"Common Warthog Phacochoerus africanus" 14: 1444: 652: 319: 994: 880: 802:"Hippopotamus-Hippopotamus amphibius" 853: 157:Male-male conflict social behaviour 24: 915:"The ethology of domestic animals" 371:) particularly like to wallow but 25: 1463: 978: 744:. Ecology Monographs, 11: 347-412 1426: 1425: 287: 86:Phacochoerus africanus africanus 952: 907: 874: 847: 793: 780: 747: 734: 712: 699: 686: 673: 646: 633: 607: 588: 61:this is usually referred to as 1071:Bee learning and communication 987:- Video of elephants wallowing 761:. BBC Wildlife. Archived from 572: 559: 555:10.1016/j.applanim.2011.01.002 467: 450: 428: 406: 13: 1: 942:Journal of Wildlife and Parks 569:. Z. Tierpsychol., 24: 80-114 399: 328:Sumatran rhinoceros wallowing 27:Bathing in mud, water or snow 1020: 756:"Face to face with big nose" 696:. Acta Theriol., 32: 335-372 683:. Acta Theriol., 30: 149-156 72: 7: 959:Deer Industry New Zealand. 881:Scott, Julia (2014-05-22). 804:. About.com. Archived from 709:. Asia Life Sci. 2: 241-250 491:Maryland Zoo in Baltimore. 382: 153:– pigs, warthogs, elephants 10: 1468: 347: 226:Relief from biting insects 1421: 1375: 1354: 1233: 1128:Evolutionary neuroscience 1028: 856:"Elk (Cervus Canadensis)" 643:. Behaviour, 135: 693-718 1081:Behavioral endocrinology 938:Dicerorhinus Sumatrensis 334:Dicerorhinus Sumatrensis 218:, elephant seals (genus 141:Phacochoerus aethiopicus 1276:Irenäus Eibl-Eibesfeldt 1056:Animal sexual behaviour 827:Valerius Geist (1998). 679:Reinhardt, V., (1985). 565:Struhsaker, T. (1967). 343: 296:Domestic pigs wallowing 1215:Tool use by non-humans 1168:Philosophical ethology 1113:Comparative psychology 1061:Animal welfare science 786:McHugh, T.S., (1958). 617:. 2012. Archived from 460:and D.B. Haley (Eds). 359: 329: 297: 258:Hippopotamus amphibius 109: 92: 35: 740:Soper, J.D., (1941). 583:Elk of North America. 413:The Free Dictionary. 357: 348:Further information: 327: 295: 248:Play in young animals 244:), warthog, elephants 151:Providing a sunscreen 147:(family Elephantidae) 107: 80: 33: 1321:William Homan Thorpe 1086:Behavioural genetics 1046:Animal consciousness 1041:Animal communication 790:. Zoologica, 4: 1-40 598:Cervus eldi hainanus 332:The Sumatran rhino ( 133:Rhinoceros unicornis 18:Wallowing in animals 1076:Behavioural ecology 833:. Stackpole Books. 768:on January 12, 2012 754:Morris, D. (2005). 545:Bracke, M. (2011). 320:Sumatran rhinoceros 236:), American bison, 234:Bubalus mindorensis 192:), American bison ( 190:Ceratotherium simum 34:A buffalo wallowing 1405:Behavioral Ecology 1326:Nikolaas Tinbergen 1118:Emotion in animals 1096:Cognitive ethology 887:The New York Times 728:2016-02-25 at the 615:"White rhinoceros" 360: 330: 298: 110: 93: 69:stuck in the mud. 36: 1439: 1438: 1331:Jakob von Uexküll 1101:Comfort behaviour 840:978-0-8117-0496-0 808:on April 15, 2013 389:Personal grooming 129:Indian rhinoceros 105: 43:comfort behaviour 16:(Redirected from 1459: 1429: 1428: 1391:Animal Cognition 1384:Animal Behaviour 1336:Wolfgang Wickler 1036:Animal cognition 1015: 1008: 1001: 992: 991: 972: 971: 969: 967: 956: 950: 949: 933: 927: 926: 924: 922: 911: 905: 904: 902: 901: 878: 872: 871: 869: 867: 862:on June 17, 2011 858:. 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Index

Wallowing in animals

comfort behaviour
mud
water
snow
ethology
dust bathing
ectoparasites

Nolan warthog
Senegal
domestic pigs
Indian rhinoceros
warthogs
elephants
elk
European bison
ectoparasites
white rhinoceros
American bison
moulting
European bison
tamaraw
tapirs
Scent-marking

hippopotamus
whales

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