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of six months when they have the same nutritional needs as adults. Wolf packs travel constantly in search of prey, covering roughly 9% of their territory per day, on average 25 km/d (16 mi/d). The core of their territory is on average 35 km (14 sq mi) where they spend 50% of their time. Prey density tends to be much higher on the territory's periphery. Wolves tend to avoid hunting on the fringes of their range to avoid fatal confrontations with neighbouring packs. The smallest territory on record was held by a pack of six wolves in northeastern
Minnesota, which occupied an estimated 33 km (13 sq mi), while the largest was held by an Alaskan pack of ten wolves encompassing 6,272 km (2,422 sq mi). Wolf packs are typically settled, and usually leave their accustomed ranges only during severe food shortages. Territorial fights are among the principal causes of wolf mortality, one study concluding that 14â65% of wolf deaths in Minnesota and the
2484: oz) at birth and begin to see after nine to 12 days. The milk canines erupt after one month. Pups first leave the den after three weeks. At one-and-a-half months of age, they are agile enough to flee from danger. Mother wolves do not leave the den for the first few weeks, relying on the fathers to provide food for them and their young. Pups begin to eat solid food at the age of three to four weeks. They have a fast growth rate during their first four months of life: during this period, a pup's weight can increase nearly 30 times. Wolf pups begin play-fighting at the age of three weeks, though unlike young coyotes and foxes, their bites are gentle and controlled. Actual fights to establish hierarchy usually occur at five to eight weeks of age. This is in contrast to young coyotes and foxes, which may begin fighting even before the onset of play behaviour. By autumn, the pups are mature enough to accompany the adults on hunts for large prey.
3369:. Wolves apparently develop the "furious" phase of rabies to a very high degree. This, coupled with their size and strength, makes rabid wolves perhaps the most dangerous of rabid animals. Bites from rabid wolves are 15 times more dangerous than those of rabid dogs. Rabid wolves usually act alone, travelling large distances and often biting large numbers of people and domestic animals. Most rabid wolf attacks occur in the spring and autumn periods. Unlike with predatory attacks, the victims of rabid wolves are not eaten, and the attacks generally occur only on a single day. The victims are chosen at random, though most cases involve adult men. During the fifty years up to 2002, there were eight fatal attacks in Europe and Russia, and more than two hundred in southern Asia.
1303:
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Alberto; Davy, Tom; Fedorov, Sergey; Fiore, Ivana; Fulton, Deirdre; GermonprĂ©, Mietje; Haile, James; Irving-Pease, Evan K.; Jamieson, Alexandra; Janssens, Luc; Kirillova, Irina; Horwitz, Liora Kolska; Kuzmanovic-CvetkoviÄ, Julka; Kuzmin, Yaroslav; Losey, Robert J.; Dizdar, Daria LoĆŸnjak; Mashkour, Marjan; Novak, Mario; Onar, Vedat; Orton, David; Pasaric, Maja; Radivojevic, Miljana; Rajkovic, Dragana; Roberts, Benjamin; Ryan, Hannah; Sablin, Mikhail; Shidlovskiy, Fedor; Stojanovic, Ivana; Tagliacozzo, Antonio; Trantalidou, Katerina; UllĂ©n, Inga; Villaluenga, Aritza; Wapnish, Paula; Dobney, Keith; Götherström, Anders; Linderholm, Anna; DalĂ©n, Love; Pinhasi, Ron; Larson, Greger; Skoglund, Pontus (2020).
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from their natal (birth) packs. A new pack is usually founded by an unrelated dispersing male and female, travelling together in search of an area devoid of other hostile packs. Wolf packs rarely adopt other wolves into their fold and typically kill them. In the rare cases where other wolves are adopted, the adoptee is almost invariably an immature animal of one to three years old, and unlikely to compete for breeding rights with the mated pair. This usually occurs between the months of
February and May. Adoptee males may mate with an available pack female and then form their own pack. In some cases, a lone wolf is adopted into a pack to replace a deceased breeder.
883:
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903:
812:
748:
716:
684:
780:
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1413:, a sloping back, and a heavily muscled neck. The wolf's legs are moderately longer than those of other canids, which enables the animal to move swiftly, and to overcome the deep snow that covers most of its geographical range in winter. The ears are relatively small and triangular. The wolf's head is large and heavy, with a wide forehead, strong jaws and a long, blunt muzzle. The skull is 230â280 mm (9â11 in) in length and 130â150 mm (5â6 in) in width. The teeth are heavy and large, making them better suited to crushing bone than those of other canids, though they are not as specialized as those found in
2780:, wolf populations have retained much of their historical range despite Soviet-era large scale extermination campaigns. Their numbers range from 1,500 in Georgia, to 20,000 in Kazakhstan and up to 45,000 in Russia. In Russia, the wolf is regarded as a pest because of its attacks on livestock, and wolf management means controlling their numbers by destroying them throughout the year. Russian history over the past century shows that reduced hunting leads to an abundance of wolves. The Russian government has continued to pay bounties for wolves and annual harvests of 20â30% do not appear to significantly affect their numbers.
2424:
badgers or marmots. An appropriated den is often widened and partly remade. On rare occasions, female wolves dig burrows themselves, which are usually small and short with one to three openings. The den is usually constructed not more than 500 m (550 yd) away from a water source. It typically faces southwards where it can be better warmed by sunlight exposure, and the snow can thaw more quickly. Resting places, play areas for the pups, and food remains are commonly found around wolf dens. The odor of urine and rotting food emanating from the denning area often attracts scavenging birds like
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1999:
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exterminating wolves has been found. Some nations help offset economic losses to wolves through compensation programs or state insurance. Domesticated animals are easy prey for wolves, as they have been bred under constant human protection, and are thus unable to defend themselves very well. Wolves typically resort to attacking livestock when wild prey is depleted. In
Eurasia, a large part of the diet of some wolf populations consists of livestock, while such incidents are rare in North America, where healthy populations of wild prey have been largely restored.
2397:, mated pairs usually remaining together for life. Should one of the pair die, another mate is found quickly. With wolves in the wild, inbreeding does not occur where outbreeding is possible. Wolves become mature at the age of two years and sexually mature from the age of three years. The age of first breeding in wolves depends largely on environmental factors: when food is plentiful, or when wolf populations are heavily managed, wolves can rear pups at younger ages to better exploit abundant resources. Females are capable of producing pups every year, one
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303:
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ignoring nearby humans. Wolf attacks on dogs may occur both in house yards and in forests. Wolf attacks on hunting dogs are considered a major problem in
Scandinavia and Wisconsin. Although the number of dogs killed each year by wolves is relatively low, it induces a fear of wolves entering villages and farmyards to prey on them. In many cultures, dogs are seen as family members, or at least working team members, and losing one can lead to strong emotional responses such as demanding more liberal hunting regulations.
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159:
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1530:. Most of the undercoat and some guard hairs are shed in spring and grow back in autumn. The longest hairs occur on the back, particularly on the front quarters and neck. Especially long hairs grow on the shoulders and almost form a crest on the upper part of the neck. The hairs on the cheeks are elongated and form tufts. The ears are covered in short hairs and project from the fur. Short, elastic and closely adjacent hairs are present on the limbs from the
3353:. Also, wolves with pups experience greater food stresses during this period. The majority of victims of predatory wolf attacks are children under the age of 18 and, in the rare cases where adults are killed, the victims are almost always women. Indian wolves have a history of preying on children, a phenomenon called "child-lifting". They may be taken primarily in the spring and summer periods during the evening hours, and often within human settlements.
117:
3329:, in which wolves gradually lose their fear of humans. The victims are repeatedly bitten on the head and face, and are then dragged off and consumed unless the wolves are driven off. Such attacks typically occur only locally and do not stop until the wolves involved are eliminated. Predatory attacks can occur at any time of the year, with a peak in the JuneâAugust period, when the chances of people entering forested areas (for livestock
1641:
tips on the hairs along the back form a broad stripe, with black hair tips on the shoulders, upper chest and rear of the body. The sides of the body, tail, and outer limbs are a pale dirty ochreous colour, while the inner sides of the limbs, belly, and groin are white. Apart from those wolves which are pure white or black, these tones vary little across geographical areas, although the patterns of these colours vary between individuals.
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80:
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their territory when hunting, using the same trails for extended periods. Wolves are nocturnal predators. During the winter, a pack will commence hunting in the twilight of early evening and will hunt all night, traveling tens of kilometres. Sometimes hunting large prey occurs during the day. During the summer, wolves generally tend to hunt individually, ambushing their prey and rarely giving pursuit.
1965:). This is more difficult during winter. Wolves in packs usually dominate cougars and can steal their kills or even kill them, while one-to-one encounters tend to be dominated by the cat, who likewise will kill wolves. Wolves more broadly affect cougar population dynamics and distribution by dominating territory and prey opportunities and disrupting the feline's behaviour. Wolf and
3074:" is derived. Some of his other fables concentrate on maintaining the trust between shepherds and guard dogs in their vigilance against wolves, as well as anxieties over the close relationship between wolves and dogs. Although Aesop used wolves to warn, criticize and moralize about human behaviour, his portrayals added to the wolf's image as a deceitful and dangerous animal. The
1984:, usually in disputes over carcasses. Striped hyenas feed extensively on wolf-killed carcasses in areas where the two species interact. One-to-one, hyenas dominate wolves, and may prey on them, but wolf packs can drive off single or outnumbered hyenas. There is at least one case in Israel of a hyena associating and cooperating with a wolf pack.
1538:. The winter fur is highly resistant to the cold. Wolves in northern climates can rest comfortably in open areas at â40 °C (â40 °F) by placing their muzzles between the rear legs and covering their faces with their tail. Wolf fur provides better insulation than dog fur and does not collect ice when warm breath is condensed against it.
3199:. His portrayal of wolves has been praised posthumously by wolf biologists for his depiction of them: rather than being villainous or gluttonous, as was common in wolf portrayals at the time of the book's publication, they are shown as living in amiable family groups and drawing on the experience of infirm but experienced elder pack members.
3271:(North America). The number of animals killed in single attacks varies according to species: most attacks on cattle and horses result in one death, while turkeys, sheep and domestic reindeer may be killed in surplus. Wolves mainly attack livestock when the animals are grazing, though they occasionally break into fenced enclosures.
3292:
behaviours, such as barking, social greeting, invitation to play or aggression. The historical use of shepherd dogs across
Eurasia has been effective against wolf predation, especially when confining sheep in the presence of several livestock guardian dogs. Shepherd dogs are sometimes killed by wolves.
4335:
Bergström, Anders; Stanton, David W. G.; Taron, Ulrike H.; Frantz, Laurent; Sinding, Mikkel-Holger S.; Ersmark, Erik; Pfrengle, Saskia; Cassatt-Johnstone, Molly; Lebrasseur, Ophélie; Girdland-Flink, Linus; Fernandes, Daniel M.; Ollivier, Morgane; Speidel, Leo; Gopalakrishnan, Shyam; Westbury, Michael
3487:. Although closely related to domestic dogs, wolves do not show the same tractability as dogs in living alongside humans, being generally less responsive to human commands and more likely to act aggressively. A person is more likely to be fatally mauled by a pet wolf or wolf-dog hybrid than by a dog.
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unaided. The size of a wolf hunting pack is related to the number of pups that survived the previous winter, adult survival, and the rate of dispersing wolves leaving the pack. The optimal pack size for hunting elk is four wolves, and for bison a large pack size is more successful. Wolves move around
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and generally establish territories far larger than they require to survive assuring a steady supply of prey. Territory size depends largely on the amount of prey available and the age of the pack's pups. They tend to increase in size in areas with low prey populations, or when the pups reach the age
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for rabies is eight to 21 days, and results in the host becoming agitated, deserting its pack, and travelling up to 80 km (50 mi) a day, thus increasing the risk of infecting other wolves. Although canine distemper is lethal in dogs, it has not been recorded to kill wolves, except in Canada
1776:
In North
America, the wolf's diet is dominated by wild large hoofed mammals (ungulates) and medium-sized mammals. In Asia and Europe, their diet is dominated by wild medium-sized hoofed mammals and domestic species. The wolf depends on wild species, and if these are not readily available, as in Asia,
4404:
Loog, Liisa; Thalmann, Olaf; Sinding, Mikkel-Holger S.; Schuenemann, Verena J.; Perri, Angela; Germonpré, Mietje; Bocherens, Herve; Witt, Kelsey E.; Samaniego
Castruita, Jose A.; Velasco, Marcela S.; LundstrĂžm, Inge K. C.; Wales, Nathan; Sonet, Gontran; Frantz, Laurent; Schroeder, Hannes (May 2020).
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Wolves kill dogs on occasion, and some wolf populations rely on dogs as an important food source. In
Croatia, wolves kill more dogs than sheep, and wolves in Russia appear to limit stray dog populations. Wolves may display unusually bold behaviour when attacking dogs accompanied by people, sometimes
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Livestock depredation has been one of the primary reasons for hunting wolves and can pose a severe problem for wolf conservation. As well as causing economic losses, the threat of wolf predation causes great stress on livestock producers, and no foolproof solution of preventing such attacks short of
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In the Middle East, only Israel and Oman give wolves explicit legal protection. Israel has protected its wolves since 1954 and has maintained a moderately sized population of 150 through effective enforcement of conservation policies. These wolves have moved into neighboring countries. Approximately
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When hunting large gregarious prey, wolves will try to isolate an individual from its group. If successful, a wolf pack can bring down game that will feed it for days, but one error in judgement can lead to serious injury or death. Most large prey have developed defensive adaptations and behaviours.
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point where the pads come in contact with ice and snow. In warm climates, the fur is coarser and scarcer than in northern wolves. Female wolves tend to have smoother furred limbs than males and generally develop the smoothest overall coats as they age. Older wolves generally have more white hairs on
1360:
compared to the wolf is proposed to be due to the coyote retaining more of the mitochondrial genome of this unidentified canid. Similarly, a museum specimen of a wolf from southern China collected in 1963 showed a genome that was 12â14% admixed from this unknown canid. In North
America, some coyotes
1172:) is widely accepted. Among the oldest fossils of the modern grey wolf is from Ponte Galeria in Italy, dating to 406,500â±â2,400 years ago. Remains from Cripple Creek Sump in Alaska may be considerably older, around 1 million years old, though differentiating between the remains of modern wolves and
585:
in their names. Examples include
Wulfhere ("Wolf Army"), Cynewulf ("Royal Wolf"), CÄnwulf ("Bold Wolf"), Wulfheard ("Wolf-hard"), Earnwulf ("Eagle Wolf"), WulfstÄn ("Wolf Stone") ĂĂ°elwulf ("Noble Wolf"), Wolfhroc ("Wolf-Frock"), Wolfhetan ("Wolf Hide"), Scrutolf ("Garb Wolf"), Wolfgang ("Wolf Gait")
2993:
creation myth, the wolf was the first animal brought to Earth. When humans killed it, they were punished with death, destruction and the loss of immortality. For the Pawnee, Sirius is the "wolf star" and its disappearance and reappearance signified the wolf moving to and from the spirit world. Both
2537:
The wolf must give chase and gain on its fleeing prey, slow it down by biting through thick hair and hide, and then disable it enough to begin feeding. Wolves may wound large prey and then lie around resting for hours before killing it when it is weaker due to blood loss, thereby lessening the risk
1941:
are much rarer because of differences in habitat preferences. Wolves have been recorded on numerous occasions actively seeking out American black bears in their dens and killing them without eating them. Unlike brown bears, American black bears frequently lose against wolves in disputes over kills.
1904:
Wolves typically dominate other canid species in areas where they both occur. In North America, incidents of wolves killing coyotes are common, particularly in winter, when coyotes feed on wolf kills. Wolves may attack coyote den sites, digging out and killing their pups, though rarely eating them.
1640:
ochreous (orange/red/brown) colours with light gray. The muzzle is pale ochreous gray, and the area of the lips, cheeks, chin, and throat is white. The top of the head, forehead, under and between the eyes, and between the eyes and ears is gray with a reddish film. The neck is ochreous. Long, black
1340:
showed admixture with Middle Eastern gray wolves and dogs. There is evidence of gene flow between golden jackals and Middle Eastern wolves, less so with European and Asian wolves, and least with North American wolves. This indicates the golden jackal ancestry found in North American wolves may have
4522:
Gopalakrishnan, Shyam; Sinding, Mikkel-Holger S.; Ramos-Madrigal, JazmĂn; Niemann, Jonas; Samaniego Castruita, Jose A.; Vieira, Filipe G.; CarĂže, Christian; Montero, Marc de Manuel; Kuderna, Lukas; Serres, Aitor; GonzĂĄlez-Basallote, VĂctor Manuel; Liu, Yan-Hu; Wang, Guo-Dong; Marques-Bonet, Tomas;
3321:
The fear of wolves has been pervasive in many societies, though humans are not part of the wolf's natural prey. How wolves react to humans depends largely on their prior experience with people: wolves lacking any negative experience of humans, or which are food-conditioned, may show little fear of
3291:
Dogs that are employed to guard sheep help to mitigate humanâwolf conflicts, and are often proposed as one of the non-lethal tools in the conservation of wolves. Shepherd dogs are not particularly aggressive, but they can disrupt potential wolf predation by displaying what is to the wolf ambiguous
3283:
carnivores did not mention any research on competition between dogs and wolves. Competition would favour the wolf, which is known to kill dogs; however, wolves usually live in pairs or in small packs in areas with high human persecution, giving them a disadvantage when facing large groups of dogs.
2755:
by 1770, in Central Europe by 1899, in France by the 1930s, and in much of Scandinavia by the early 1970s. They continued to survive in parts of Finland, Eastern Europe and Southern Europe. Since 1980, European wolves have rebounded and expanded into parts of their former range. The decline of the
2596:
The global wild wolf population in 2003 was estimated at 300,000. Wolf population declines have been arrested since the 1970s. This has fostered recolonization and reintroduction in parts of its former range as a result of legal protection, changes in land use, and rural human population shifts to
2337:
to assemble the pack usually before and after hunts, to pass on an alarm particularly at a den site, to locate each other during a storm, while crossing unfamiliar territory, and to communicate across great distances. Wolf howls can under certain conditions be heard over areas of up to 130 km
2245:
and competition within the pack for food. The distance travelled by dispersing wolves varies widely; some stay in the vicinity of the parental group, while other individuals may travel great distances of upwards of 206 km (128 mi), 390 km (240 mi), and 670 km (420 mi)
2232:
accompanied by their offspring. The average pack size in North America is eight wolves and 5.5 in Europe. The average pack across Eurasia consists of a family of eight wolves (two adults, juveniles, and yearlings), or sometimes two or three such families, with examples of exceptionally large packs
1813:
were among the most frequently consumed prey of North American wolves. Wolves can digest their meal in a few hours and can feed several times in one day, making quick use of large quantities of meat. A well-fed wolf stores fat under the skin, around the heart, intestines, kidneys, and bone marrow,
1501:
25 kg (55 lb). Females in any given wolf population typically weigh 2.3â4.5 kg (5â10 lb) less than males. Wolves weighing over 54 kg (119 lb) are uncommon, though exceptionally large individuals have been recorded in Alaska and Canada. In central Russia, exceptionally
4583:
Sinding, Mikkel-Holger S.; Gopalakrishan, Shyam; Vieira, Filipe G.; Samaniego Castruita, Jose A.; Raundrup, Katrine; Heide JÞrgensen, Mads Peter; Meldgaard, Morten; Petersen, Bent; Sicheritz-Ponten, Thomas; Mikkelsen, Johan Brus; Marquard-Petersen, Ulf; Dietz, Rune; Sonne, Christian; Dalén, Love;
3726:
Koepfli, Klaus-Peter; Pollinger, John; Godinho, Raquel; Robinson, Jacqueline; Lea, Amanda; Hendricks, Sarah; Schweizer, Rena M.; Thalmann, Olaf; Silva, Pedro; Fan, Zhenxin; Yurchenko, Andrey A.; Dobrynin, Pavel; Makunin, Alexey; Cahill, James A.; Shapiro, Beth; Ălvares, Francisco; Brito, JosĂ© C.;
1909:
relied on coyotes to accompany him and warn him of danger. Though they fed from his kills, he never allowed them to approach him. Interactions have been observed in Eurasia between wolves and golden jackals, the latter's numbers being comparatively small in areas with high wolf densities. Wolves
2700:
states of Minnesota, Wisconsin and Michigan where wolves number over 4,000 as of 2018. Wolves also occupy much of the northern Rocky Mountains region and the northwest, with a total population over 3,000 as of the 2020s. In Mexico and parts of the southwestern United States, the Mexican and U.S.
2557:
Once prey is brought down, wolves begin to feed excitedly, ripping and tugging at the carcass in all directions, and bolting down large chunks of it. The breeding pair typically monopolizes food to continue producing pups. When food is scarce, this is done at the expense of other family members,
2362:
and anal gland scents. This is more effective at advertising territory than howling and is often used in combination with scratch marks. Wolves increase their rate of scent marking when they encounter the marks of wolves from other packs. Lone wolves will rarely mark, but newly bonded pairs will
4799:
Bergström, Anders; Frantz, Laurent; Schmidt, Ryan; Ersmark, Erik; Lebrasseur, Ophelie; Girdland-Flink, Linus; Lin, Audrey T.; StorĂ„, Jan; Sjögren, Karl-Göran; Anthony, David; Antipina, Ekaterina; Amiri, Sarieh; Bar-Oz, Guy; Bazaliiskii, Vladimir I.; BulatoviÄ, Jelena; Brown, Dorcas; Carmagnini,
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lasts 62â75 days with pups usually being born in the spring months or early summer in very cold places such as on the tundra. Young females give birth to four to five young, and older females from six to eight young and up to 14. Their mortality rate is 60â80%. Newborn wolf pups look similar to
2423:
Dens are usually constructed for pups during the summer period. When building dens, females make use of natural shelters like fissures in rocks, cliffs overhanging riverbanks and holes thickly covered by vegetation. Sometimes, the den is the appropriated burrow of smaller animals such as foxes,
6127:
Miquelle, D. G.; Stephens, P. A.; Smirnov, E. N.; Goodrich, J. M.; Zaumyslova, O. J.; Myslenkov, A. E. (2005). "Tigers and Wolves in the Russian Far East: Competitive Exclusion, Functional Redundancy and Conservation Implications". In Ray, J. C.; Berger, J.; Redford, K. H.; Steneck, R. (eds.).
4230:
Fan, Zhenxin; Silva, Pedro; Gronau, Ilan; Wang, Shuoguo; Armero, Aitor Serres; Schweizer, Rena M.; Ramirez, Oscar; Pollinger, John; Galaverni, Marco; Ortega Del-Vecchyo, Diego; Du, Lianming; Zhang, Wenping; Zhang, Zhihe; Xing, Jinchuan; VilĂ , Carles; Marques-Bonet, Tomas; Godinho, Raquel; Yue,
2363:
scent mark the most. These marks are generally left every 240 m (260 yd) throughout the territory on regular travelways and junctions. Such markers can last for two to three weeks, and are typically placed near rocks, boulders, trees, or the skeletons of large animals. Raised leg
2166:). Most fluke species reside in the wolf's intestines. Tapeworms are commonly found in wolves, which they get though their prey, and generally cause little harm in wolves, though this depends on the number and size of the parasites, and the sensitivity of the host. Symptoms often include
1497:. The mean body mass of the wolf is 40 kg (88 lb), the smallest specimen recorded at 12 kg (26 lb) and the largest at 79.4 kg (175 lb). On average, European wolves weigh 38.5 kg (85 lb), North American wolves 36 kg (79 lb), and Indian and
1892:. In Eurasian areas with dense human activity, many wolf populations are forced to subsist largely on livestock and garbage. As prey in North America continue to occupy suitable habitats with low human density, North American wolves eat livestock and garbage only in dire circumstances.
1511:
1936:
typically dominate wolf packs in disputes over carcasses, while wolf packs mostly prevail against bears when defending their den sites. Both species kill each other's young. Wolves eat the brown bears they kill, while brown bears seem to eat only young wolves. Wolf interactions with
3356:
Cases of rabid wolves are low when compared to other species, as wolves do not serve as primary reservoirs of the disease, but can be infected by animals such as dogs, jackals and foxes. Incidents of rabies in wolves are very rare in North America, though numerous in the eastern
4692:
Iacolina, Laura; Scandura, Massimo; Gazzola, Andrea; Cappai, Nadia; Capitani, Claudia; Mattioli, Luca; Vercillo, Francesca; Apollonio, Marco (2010). "Y-chromosome microsatellite variation in Italian wolves: A contribution to the study of wolf-dog hybridization patterns".
3213:
and taught in several schools decades after its publication. Although credited with having changed popular perceptions on wolves by portraying them as loving, cooperative and noble, it has been criticized for its idealization of wolves and its factual inaccuracies.
4523:
Mirarab, Siavash; Fernandes, Carlos; Gaubert, Philippe; Koepfli, Klaus-Peter; Budd, Jane; Rueness, Eli Knispel; Heide-JĂžrgensen, Mads Peter; Petersen, Bent; Sicheritz-Ponten, Thomas; Bachmann, Lutz; Wiig, Ăystein; Hansen, Anders J.; Gilbert, M. Thomas P. (2018).
2346:
and whines. Wolves do not bark as loudly or continuously as dogs do in confrontations, rather barking a few times and then retreating from a perceived danger. Aggressive or self-assertive wolves are characterized by their slow and deliberate movements, high body
1210:. This suggests they either often processed carcasses, or that they competed with other carnivores and needed to consume their prey quickly. The frequency and location of tooth fractures in these wolves indicates they were habitual bone crackers like the modern
2965:
and guards the heavenly gate. In China, the wolf was traditionally associated with greed and cruelty and wolf epithets were used to describe negative behaviours such as cruelty ("wolf's heart"), mistrust ("wolf's look") and lechery ("wolf-sex"). In both
3456:
and large falcons, though this practice is declining, as experienced falconers are becoming few in number. Shooting wolves from aircraft is highly effective, due to increased visibility and direct lines of fire. Several types of dog, including the
8447:
1421:
have a flat chewing surface, but not to the same extent as the coyote, whose diet contains more vegetable matter. Females tend to have narrower muzzles and foreheads, thinner necks, slightly shorter legs, and less massive shoulders than males.
1657:
in North America acquired their colour from wolf-dog admixture after the first arrival of dogs across the Bering Strait 12,000 to 14,000 years ago. Research into the inheritance of white colour from dogs into wolves has yet to be undertaken.
1652:
of Alberta and British Columbia, the coat colour is predominantly black, some being blue-gray and some with silver and black. Differences in coat colour between sexes is absent in Eurasia; females tend to have redder tones in North America.
2672:
people can hunt wolves without restrictions, but others must acquire licenses for the hunting and trapping seasons. As many as 4,000 wolves may be harvested in Canada each year. The wolf is a protected species in national parks under the
1384:
was discovered through mitochondrial DNA analysis of jackals living in the Caucasus Mountains and in Bulgaria. In 2021, a genetic study found that the dog's similarity to the extant gray wolf was the result of substantial dog-into-wolf
5291:
Anderson, T. M.; Vonholdt, B. M.; Candille, S. I.; Musiani, M.; Greco, C.; Stahler, D. R.; Smith, D. W.; Padhukasahasram, B.; Randi, E.; Leonard, J. A.; Bustamante, C. D.; Ostrander, E. A.; Tang, H.; Wayne, R. K.; Barsh, G. S. (2009).
1697:
Wolves occur across Eurasia and North America. However, deliberate human persecution because of livestock predation and fear of attacks on humans has reduced the wolf's range to about one-third of its historic range; the wolf is now
3440:
poles carrying a human scent. This method relies heavily on the wolf's fear of human scents, though it can lose its effectiveness when wolves become accustomed to the odor. Some hunters can lure wolves by imitating their calls. In
1283:
and Chinese indigenous breeds are basal members of the domestic dog clade. The divergence time for wolves in Europe, the Middle East, and Asia is estimated to be fairly recent at around 1,600 years ago. Among New World wolves, the
3408:
said wolves are difficult to hunt because of their elusiveness, sharp senses, high endurance, and ability to quickly incapacitate and kill hunting dogs. Historic methods included killing of spring-born litters in their dens,
3115:
Cycle, where he stands for the low nobility, whilst his adversary, Reynard the fox, represents the peasant hero. Isengrim is forever the victim of Reynard's wit and cruelty, often dying at the end of each story. The tale of
5498:
Newsome, Thomas M.; Boitani, Luigi; Chapron, Guillaume; Ciucci, Paolo; Dickman, Christopher R.; Dellinger, Justin A.; López-Bao, José V.; Peterson, Rolf O.; Shores, Carolyn R.; Wirsing, Aaron J.; Ripple, William J. (2016).
2701:
governments collaborated from 1977 to 1980 in capturing all Mexican wolves remaining in the wild to prevent their extinction and established captive breeding programs for reintroduction. As of 2024, the reintroduced
1547:
the tip of the tail, along the nose, and on the forehead. Winter fur is retained longest by lactating females, although with some hair loss around their teats. Hair length on the middle of the back is 60â70 mm (
3250:
The majority of losses occur during the summer grazing period, untended livestock in remote pastures being the most vulnerable to wolf predation. The most frequently targeted livestock species are sheep (Europe),
4280:
Hennelly, Lauren M.; Habib, Bilal; Modi, Shrushti; Rueness, Eli K.; Gaubert, Philippe; Sacks, Benjamin N. (2021). "Ancient divergence of Indian and Tibetan wolves revealed by recombination-aware phylogenomics".
2237:
levels in wolves rise significantly when a pack member dies, indicating the presence of stress. During times of prey abundance caused by calving or migration, different wolf packs may join together temporarily.
8541:
Espuno, Nathalie; Lequette, Benoit; Poulle, Marie-Lazarine; Migot, Pierre; Lebreton, Jean-Dominique (2004). "Heterogeneous response to preventive sheep husbandry during wolf recolonization of the French Alps".
2772:
as of 2007. In a study of wolf conservation in Sweden, it was found that there was little opposition between the policies of the European Union and those of the Swedish officials implementing domestic policy.
2089:. Although lyme disease can debilitate individual wolves, it does not appear to significantly affect wolf populations. Leptospirosis can be contracted through contact with infected prey or urine, and can cause
2618:
2725:
2677:. In Alaska, 7,000â11,000 wolves are found on 85% of the state's 1,517,733 km (586,000 sq mi) area. Wolves may be hunted or trapped with a license; around 1,200 wolves are harvested annually.
2332:
Wolves communicate using vocalizations, body postures, scent, touch, and taste. The phases of the moon have no effect on wolf vocalization, and despite popular belief, wolves do not howl at the Moon. Wolves
2597:
cities. Competition with humans for livestock and game species, concerns over the danger posed by wolves to people, and habitat fragmentation pose a continued threat to the wolf. Despite these threats, the
1865:. They also eat grass, which may provide some vitamins, but is most likely used mainly to induce vomiting to rid themselves of intestinal parasites or long guard hairs. They are known to eat the berries of
1541:
In cold climates, the wolf can reduce the flow of blood near its skin to conserve body heat. The warmth of the foot pads is regulated independently from the rest of the body and is maintained at just above
8454:
2756:
traditional pastoral and rural economies seems to have ended the need to exterminate the wolf in parts of Europe. As of 2016, estimates of wolf numbers include: 4,000 in the Balkans, 3,460â3,849 in the
2684:, wolf declines were caused by the expansion of agriculture, the decimation of the wolf's main prey species like the American bison, and extermination campaigns. Wolves were given protection under the
1437:
Adult wolves measure 105â160 cm (41â63 in) in length and 80â85 cm (31â33 in) at shoulder height. The tail measures 29â50 cm (11â20 in) in length, the ears 90â110 mm (
2224:. Its populations consist of packs and lone wolves, most lone wolves being temporarily alone while they disperse from packs to form their own or join another one. The wolf's basic social unit is the
8348:
Vanak, A. T.; Dickman, C. R.; Silva-Rodriguez, E. A.; Butler, J. R. A.; Ritchie, E. G. (2014). "Top-dogs and under-dogs: competition between dogs and sympatric carnivores". In Gompper, M. E. (ed.).
5228:
Gipson, Philip S.; Bangs, Edward E.; Bailey, Theodore N.; Boyd, Diane K.; Cluff, H. Dean; Smith, Douglas W.; Jiminez, Michael D. (2002). "Color Patterns among Wolves in Western North America".
1905:
There are no records of coyotes killing wolves, though coyotes may chase wolves if they outnumber them. According to a press release by the U.S. Department of Agriculture in 1921, the infamous
1409:, and is further distinguished from coyotes and jackals by a broader snout, shorter ears, a shorter torso and a longer tail. It is slender and powerfully built, with a large, deeply descending
499:
on humans are rare because wolves are relatively few, live away from people, and have developed a fear of humans because of their experiences with hunters, farmers, ranchers, and shepherds.
2294:
1706:
and Japan. In modern times, the wolf occurs mostly in wilderness and remote areas. The wolf can be found between sea level and 3,000 m (9,800 ft). Wolves live in forests, inland
8608:
2799:. The wolf also appears to be widespread in Iran. Turkey has an estimated population of about 7,000 wolves. Outside of Turkey, wolf populations in the Middle East may total 1,000â2,000.
1765:
that can be divided into large size 240â650 kg (530â1,430 lb) and medium size 23â130 kg (51â287 lb), and have a body mass similar to that of the combined mass of the
7642:"Identifying suitable habitat and corridors for Indian Grey Wolf (Canis lupus pallipes) in Chotta Nagpur Plateau and Lower Gangetic Planes: A species with differential management needs"
2854:
has 2,000. 2017 evidence suggests that wolves range across all of mainland China. Wolves have been historically persecuted in China but have been legally protected since 1998. The last
2523:, geese, and hares, there is no risk to the wolf. Although people often believe wolves can easily overcome any of their prey, their success rate in hunting hoofed prey is usually low.
2312:
1961:
along portions of the Rocky Mountains and adjacent mountain ranges. Wolves and cougars typically avoid encountering each other by hunting at different elevations for different prey (
7563:
2444:. During pregnancy, female wolves remain in a den located away from the peripheral zone of their territories, where violent encounters with other packs are less likely to occur.
2519:
Wolves have been killed while attempting to bring down bison, elk, moose, muskoxen, and even by one of their smallest hoofed prey, the white-tailed deer. With smaller prey like
2506:
Single wolves or mated pairs typically have higher success rates in hunting than do large packs; single wolves have occasionally been observed to kill large prey such as moose,
2292:
2310:
7346:
1240:
from them 200,000 years ago. Other wolves appear to share most of their common ancestry much more recently, within the last 23,000 years (around the peak and the end of the
8091:"Territoriality and interpack aggression in wolves: Shaping a social carnivore's life history. Rudyard Kipling's Law of the Jungle Meets Yellowstone's Law of the Mountains"
1845:, and large insects when available. Wolves in some areas may consume fish and even marine life. Wolves also consume some plant material. In Europe, they eat apples, pears,
4635:
Wang, Guo-Dong; Zhang, Ming; Wang, Xuan; Yang, Melinda A.; Cao, Peng; Liu, Feng; Lu, Heng; Feng, Xiaotian; Skoglund, Pontus; Wang, Lu; Fu, Qiaomei; Zhang, Ya-Ping (2019).
1632:
A wolf's coat colour is determined by its guard hairs. Wolves usually have some hairs that are white, brown, gray and black. The coat of the Eurasian wolf is a mixture of
1726:, deserts, and rocky peaks on mountains. Habitat use by wolves depends on the abundance of prey, snow conditions, livestock densities, road densities, human presence and
8287:
6249:
Dinets, Vladimir; Eligulashvili, Beniamin (2016). "Striped Hyaenas (Hyaena hyaena) in Grey Wolf (Canis lupus) packs: Cooperation, commensalism or singular aberration?".
7512:
4731:
Kopaliani, N.; Shakarashvili, M.; Gurielidze, Z.; Qurkhuli, T.; Tarkhnishvili, D. (2014). "Gene Flow between Wolf and Shepherd Dog Populations in Georgia (Caucasus)".
6607:
3933:; Kitchener, Andrew C.; Koepfli, Klaus-Peter; Krofel, Miha; Moehlman, Patricia D.; Senn, Helen; Sillero-Zubiri, Claudio; Viranta, Suvi; Werhahn, Geraldine (2019).
2842:. As of 2019, it is estimated that there are around 2,000â3,000 Indian wolves in the country. In East Asia, Mongolia's population numbers 10,000â20,000. In China,
1336:, and gray wolves. The African wolf is a descendant of a genetically admixed canid of 72% wolf and 28% Ethiopian wolf ancestry. One African wolf from the Egyptian
2293:
491:
societies. Although the fear of wolves exists in many human societies, the majority of recorded attacks on people have been attributed to animals suffering from
3393:
1773:. The variation in diet between wolves living on different continents is based on the variety of hoofed mammals and of available smaller and domesticated prey.
8733:
8200:
Creel, S.; Fox, J. E.; Hardy, A.; Sands, J.; Garrott, B.; Peterson, R. O. (2002). "Snowmobile activity and glucocorticoid stress responses in wolves and elk".
3977:
Iurino, Dawid A.; Mecozzi, Beniamino; Iannucci, Alessio; Moscarella, Alfio; Strani, Flavia; Bona, Fabio; Gaeta, Mario; Sardella, Raffaele (February 25, 2022).
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540:
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people. Although wolves may react aggressively when provoked, such attacks are mostly limited to quick bites on extremities, and the attacks are not pressed.
3128:
is portrayed as a villain capable of imitating human speech and disguising itself with human clothing. The character has been interpreted as an allegorical
15864:
6230:
Nayak, S.; Shah, S.; Borah, J. (2015). "Going for the kill: an observation of wolf-hyaena interaction in Kailadevi Wildlife Sanctuary, Rajasthan, India".
2182:. Wolves can carry over 30 roundworm species, though most roundworm infections appear benign, depending on the number of worms and the age of the host.
7211:
2693:
1202:. It is proposed that these features were specialized adaptations for the processing of carcass and bone associated with the hunting and scavenging of
993:. Linnaeus considered the dog to be a separate species from the wolf because of its "cauda recurvata" (upturning tail) which is not found in any other
8683:
5583:
383:
species by its less pointed ears and muzzle, as well as a shorter torso and a longer tail. The wolf is nonetheless related closely enough to smaller
4042:
Tedford, Richard H.; Wang, Xiaoming; Taylor, Beryl E. (2009). "Phylogenetic Systematics of the North American Fossil Caninae (Carnivora: Canidae)".
3727:
Geffen, Eli; Leonard, Jennifer A.; Helgen, Kristofer M.; Johnson, Warren E.; o'Brien, Stephen J.; Van Valkenburgh, Blaire; Wayne, Robert K. (2015).
7472:
6861:
2689:
2668:
In Canada, 50,000â60,000 wolves live in 80% of their historical range, making Canada an important stronghold for the species. Under Canadian law,
15624:
7371:
14204:
9020:
8589:
8448:"Wolf attacks on dogs in Scandinavia 1995â2005âWill wolves in Scandinavia go extinct if dog owners are allowed to kill a wolf attacking a dog?"
2598:
476:
5867:
1648:, wolves in the Canadian arctic being white and those in southern Canada, the U.S., and Mexico being predominantly gray. In some areas of the
399:
with them. The wolf's fur is usually mottled white, brown, gray, and black, although subspecies in the arctic region may be nearly all white.
15715:
15663:
14275:
3640:
Systema naturĂŠ per regna tria naturĂŠ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I
3503:
Populations of Bhutan, India, Nepal, and Pakistan are included in Appendix I. Excludes domesticated form and dingo, which are referenced as
1896:
is not uncommon in wolves during harsh winters, when packs often attack weak or injured wolves and may eat the bodies of dead pack members.
15844:
5005:
Therrien, F. O. (2005). "Mandibular force profiles of extant carnivorans and implications for the feeding behaviour of extinct predators".
3086:
is said to have used wolves as illustrations of the dangers his followers, whom he represents as sheep, would face should they follow him.
7559:
7182:
6145:
6208:
2877:
7151:
2367:
is considered to be one of the most important forms of scent communication in the wolf, making up 60â80% of all scent marks observed.
1769:
members. The wolf specializes in preying on the vulnerable individuals of large prey, with a pack of 15 able to bring down an adult
15585:
14878:
7417:
The wolf in Eurasiaâa regional approach to the conservation and management of a top-predator in Central Asia and the South Caucasus
2591:
7342:
7313:
3935:"Old World Canis spp. with taxonomic ambiguity: Workshop conclusions and recommendations. CIBIO. VairĂŁo, Portugal, 28â30 May 2019"
15637:
8679:
Hunting the grisly and other sketches; an account of the big game of the United States and its chase with horse, hound, and rifle
5900:
1206:. Compared with modern wolves, some Pleistocene wolves showed an increase in tooth breakage similar to that seen in the extinct
7597:
Yadvendradev, V. Jhala; Giles, Robert H. Jr. (991). "The Status and Conservation of the Wolf in Gujarat and Rajasthan, India".
5579:"Dietary reconstruction and evidence of prey shifting in Pleistocene and recent gray wolves (Canis lupus) from Yukon Territory"
8164:
6899:
Thurber, J. M.; Peterson, R. O. (1993). "Effects of Population Density and Pack Size on the Foraging Ecology of Gray Wolves".
4092:
Thalmann, Olaf; Perri, Angela R. (2018). "Paleogenomic Inferences of Dog Domestication". In Lindqvist, C.; Rajora, O. (eds.).
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6202:
5479:
5168:
5138:
4109:
3854:
15642:
7508:
2621:(CITES), meaning international trade in the species (including parts and derivatives) is regulated. However, populations of
1957:, which may feed on smaller prey where wolves are present and may be suppressed by large wolf populations. Wolves encounter
15889:
5577:
Landry, Zoe; Kim, Sora; Trayler, Robin B.; Gilbert, Marisa; Zazula, Grant; Southon, John; Fraser, Danielle (June 1, 2021).
4887:
6498:
JÄdrzejewski, W. O.; Schmidt, K.; Theuerkauf, J. R.; JÄdrzejewska, B. A.; Kowalczyk, R. (2007). "Territory size of wolves
1946:, and will chase off those that attempt to scavenge from their kills. Wolverines escape from wolves in caves or up trees.
369:, though gray wolves, as popularly understood, only comprise naturally-occurring wild subspecies. The wolf is the largest
15899:
9374:
5723:
6563:
3453:
15874:
15536:
9013:
8974:
8357:
7929:
6871:
5785:
5756:
3474:
3139:
2658:
1380:, are first generation hybrids. Although mating between golden jackals and wolves has never been observed, evidence of
15859:
15808:
10744:
8525:
6642:
6467:
6139:
6003:
5076:
2720:
Europe, excluding Russia, Belarus and Ukraine, has 17,000 wolves in more than 28 countries. In many countries of the
2255:
2241:
Offspring typically stay in the pack for 10â54 months before dispersing. Triggers for dispersal include the onset of
1993:
1190:. Many Late Pleistocene wolf populations had more robust skulls and teeth than modern wolves, often with a shortened
410:
15554:
8744:
8141:
4768:"Unregulated hunting and genetic recovery from a severe population decline: The cautionary case of Bulgarian wolves"
3836:
1220:
studies suggest modern wolves and dogs descend from a common ancestral wolf population. A 2021 study found that the
15879:
15598:
8453:. Examensarbete, Institutionen för ekologi, Grimsö forskningsstation. Sveriges Lantbruksuniversitet. Archived from
4178:"Ancient Wolf Genome Reveals an Early Divergence of Domestic Dog Ancestors and Admixture into High-Latitude Breeds"
2582:
are eaten once they are exposed, followed by the muscles. A wolf can eat 15â19% of its body weight in one sitting.
5679:
Gable, T. D.; Windels, S. K.; Homkes, A. T. (2018). "Do wolves hunt freshwater fish in spring as a food source?".
4125:
Freedman, Adam H.; Gronau, Ilan; Schweizer, Rena M.; Ortega-Del Vecchyo, Diego; Han, Eunjung; et al. (2014).
15720:
15567:
11088:
7373:
Status, management and distribution of large carnivoresâbear, lynx, wolf & wolverineâin Europe December 2012.
3979:"A Middle Pleistocene wolf from central Italy provides insights on the first occurrence of Canis lupus in Europe"
2688:(ESA) of 1973, and have since returned to parts of their former range thanks to both natural recolonizations and
2493:
959:
417:
273:
8555:
8497:
8480:
1702:(locally extinct) from much of its range in Western Europe, the United States and Mexico, and completely in the
1263:
A 2016 genomic study suggests that Old World and New World wolves split around 12,500 years ago followed by the
1107:/Species Survival Commission's Canid Specialist Group considered the New Guinea singing dog and the dingo to be
460:, and garbage. Single wolves or mated pairs typically have higher success rates in hunting than do large packs.
7297:
3816:
2456:
pups. They are born blind and deaf and are covered in short soft grayish-brown fur. They weigh 300â500 g (
3436:
A popular method of wolf hunting in Russia involves trapping a pack within a small area by encircling it with
2914:
The wolf is a common motif in the mythologies and cosmologies of peoples throughout its historical range. The
15821:
15702:
15372:
13306:
9006:
8891:
4766:
Moura, A. E.; Tsingarska, E.; DÄ
browski, M. J.; Czarnomska, S. D.; JÄdrzejewska, B. A.; Pilot, M. G. (2013).
3930:
3066:, playing on the concerns of Ancient Greece's settled, sheep-herding world. His most famous is the fable of "
2744:
IV). There is extensive legal protection in many European countries, although there are national exceptions.
8388:
Lescureux, Nicolas; Linnell, John D. C. (2014). "Warring brothers: The complex interactions between wolves (
7204:
1493: in). The size and weight of the modern wolf increases proportionally with latitude in accordance with
1302:
15904:
14871:
9246:
2406:
44:
8677:
2558:
especially non-pups. Wolves typically commence feeding by gorging on the larger internal organs, like the
2204:
1372:
originated from dog ancestry, which indicates female wolves will breed with male dogs in the wild. In the
947:
Cladogram and divergence of the gray wolf (including the domestic dog) among its closest extant relatives
15676:
15474:
13297:
10500:
8941:
8916:
7479:
5825:
Klein, D. R. (1995). "The introduction, increase, and demise of wolves on Coronation Island, Alaska". In
5154:
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3659:
3333:
or berry and mushroom picking) increase. Cases of non-rabid wolf attacks in winter have been recorded in
2697:
2025:
1019:
444:
on the onset of sexual maturity and in response to competition for food within the pack. Wolves are also
15479:
3124:, is considered to have further contributed to the wolf's negative reputation in the Western world. The
3070:", which is directed at those who knowingly raise false alarms, and from which the idiomatic phrase "to
1271:
may have been the ancestor of the dog, with the dog's similarity to the extant wolf being the result of
479:(IUCN). Wolves have a long history of interactions with humans, having been despised and hunted in most
15854:
14762:
14418:
9264:
9228:
6031:
Jimenez, Michael D.; Asher, Valpa J.; Bergman, Carita; Bangs, Edward E.; Woodruff, Susannah P. (2008).
3163:
2935:
2899:
1817:
Nonetheless, wolves are not fussy eaters. Smaller-sized animals that may supplement their diet include
1267:
of the lineage that led to dogs from other Old World wolves around 11,100â12,300 years ago. An extinct
1133:
31:
15456:
12784:
15730:
15577:
15332:
11623:
9033:
4407:"Ancient DNA suggests modern wolves trace their origin to a Late Pleistocene expansion from Beringia"
2674:
2003:
1625:
1065:
1006:
355:
289:
7401:
5433:
Sorkin, Boris (2008). "A biomechanical constraint on body mass in terrestrial mammalian predators".
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15849:
14681:
14576:
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11605:
11333:
11300:
11276:
11097:
10349:
5860:
3934:
3582:
3054:
3018:
3013:
The concept of people turning into wolves, and the inverse, has been present in many cultures. One
2681:
1425:
533:
96:
15694:
7398:
The Wolf Dilemma: Following the Practices of Several Actors in Swedish Large Carnivore Management.
3929:
Alvares, Francisco; Bogdanowicz, Wieslaw; Campbell, Liz A.D.; Godinho, Rachel; Hatlauf, Jennifer;
2355:, while submissive ones carry their bodies low, flatten their fur, and lower their ears and tail.
158:
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15337:
14864:
14343:
13324:
13055:
12748:
12683:
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11641:
11114:
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9367:
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6365:
Molnar, B.; Fattebert, J.; Palme, R.; Ciucci, P.; Betschart, B.; Smith, D. W.; Diehl, P. (2015).
5578:
2851:
2748:
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523:
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882:
843:
15782:
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13620:
13602:
13430:
13124:
13115:
12757:
11495:
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8820:
Mammals of the Soviet Union Vol. II Part 1a, Sirenia and Carnivora (Sea cows; Wolves and Bears)
3844:
3382:
3117:
3094:
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2734:
Council Directive 92/43/EEC on the Conservation of Natural Habitats and of Wild Fauna and Flora
2685:
1377:
1049:
902:
811:
471:
The global wild wolf population was estimated to be 300,000 in 2003 and is considered to be of
17:
7314:"Mexican Wolf Population Grows for Eighth Consecutive Year | U.S. Fish & Wildlife Service"
7174:
6630:
6459:
6453:
6449:
6129:
5746:
5126:
2888:
747:
715:
683:
448:, and fights over territory are among the principal causes of mortality. The wolf is mainly a
15777:
15429:
15137:
15044:
14956:
14744:
13999:
13981:
13972:
13963:
13448:
13439:
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11581:
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11512:
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5995:
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5469:
5158:
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3134:
2046:
1878:
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1253:
1233:
1203:
1127:
964:
779:
663:
472:
253:
35:
15769:
12766:
12662:
8089:
Cassidy, K; Smith, D. W.; Mech, L. D.; MacNulty, D. R.; Stahler, D. R.; Metz, M. C. (2006).
3808:
3802:
3798:
3729:"Genome-wide Evidence Reveals that African and Eurasian Golden Jackals Are Distinct Species"
3142:". The hunting of wolves, and their attacks on humans and livestock, feature prominently in
3041:
would turn into wolves by donning wolf skins and would kill people and raid graveyards. The
483:
communities because of their attacks on livestock, while conversely being respected in some
15803:
15510:
15438:
15029:
14887:
14585:
14486:
14445:
14159:
13670:
13398:
12724:
11789:
11539:
11530:
11486:
11441:
11064:
10570:
10305:
10131:
9282:
8405:
8209:
7653:
7606:
7143:
6727:
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6511:
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5688:
5592:
5442:
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4536:
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4349:
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4189:
3990:
3740:
3635:
3305:
2757:
2386:
2348:
1747:
1241:
15655:
2609:
due to its relatively widespread range and stable population. The species is listed under
1510:
1397:
1138:
8:
15884:
15312:
14825:
14771:
14543:
13990:
13876:
13858:
13703:
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13457:
12707:
12653:
12402:
12144:
12135:
11399:
10672:
10654:
10509:
10358:
9861:
9337:
7367:
4488:
4471:
3832:
2777:
2662:
2432:. Though they mostly avoid areas within human sight, wolves have been known to nest near
2394:
2376:
2285:
2078:
1938:
1353:
1318:
1229:
1076:
1027:
898:
878:
106:
11895:
8409:
8213:
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7610:
7237:"Wolf Numbers in Minnesota, Wisconsin and Michigan (excluding Isle Royale)â1976 to 2015"
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6579:
6515:
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5309:
4706:
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10735:
10627:
10527:
10467:
9852:
9788:
9465:
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9291:
8559:
8288:"These Non-Lethal Methods Encouraged by Science Can Keep Wolves From Killing Livestock"
8225:
7858:
7800:
7773:
7737:
7728:
7712:
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7641:
7640:
Sharma, Lalit Kumar; Mukherjee, Tanoy; Saren, Phakir Chandra; Chandra, Kailash (2019).
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6527:
6401:
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5326:
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5237:
4879:
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4826:
4801:
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4585:
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4524:
4501:
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4406:
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4257:
4232:
4153:
4126:
4069:
4019:
3978:
3906:
3879:
3700:
3405:
3209:
is widely considered to be the most popular book on wolves, having been adapted into a
3182:
3143:
2883:
2733:
2267:
2250:
2163:
2094:
2058:
2033:
1962:
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445:
425:
153:
60:
5893:
1099:, as modern wolves are not closely related to the population of wolves that was first
15764:
15487:
14519:
14436:
14216:
13712:
13679:
12603:
12110:
11832:
11614:
11468:
11342:
11226:
10579:
10409:
10367:
10314:
10094:
10076:
9834:
9300:
9148:
8945:
8920:
8895:
8870:
8845:
8824:
8818:
8803:
8784:
8598:
8521:
8353:
8221:
8090:
7925:
7850:
7805:
7742:
7681:
7415:
7293:
7059:
6867:
6735:
6638:
6591:
6523:
6463:
6406:
6198:
6135:
6109:
5999:
5781:
5752:
5608:
5527:
5475:
5454:
5372:
5331:
5164:
5134:
5072:
4843:
4831:
4748:
4674:
4637:"Genomic Approaches Reveal an Endemic Subpopulation of Gray Wolves in Southern China"
4617:
4562:
4493:
4452:
4434:
4383:
4365:
4318:
4306:
4262:
4207:
4158:
4105:
4024:
4006:
3911:
3860:
3850:
3812:
3758:
3559:
3310:
3063:
3030:
2958:
2931:
2903:
2796:
2769:
2765:
2752:
2531:
2303:
2175:
2171:
2134:
2037:
2021:
1802:
1672:
1272:
1061:
1053:
1040:
934:
15443:
8563:
8229:
7862:
6980:"A proposed ethogram of large-carnivore predatory behavior, exemplified by the wolf"
6928:
6603:
6531:
6502:: Linking local (BiaĆowieĆŒa Primeval Forest, Poland) and Holarctic-scale patterns".
6270:
5708:
5536:
5474:. Fauna Series 7. Fauna of the National Parks of the United States. pp. 75â76.
5419:
4505:
4073:
3078:
uses an image of a wolf lying with a lamb in a utopian vision of the future. In the
2209:
1777:
the wolf is more reliant on domestic species. Across Eurasia, wolves prey mostly on
1575: in), and the guard hairs on the shoulders generally do not exceed 90 mm (
15362:
15269:
14798:
14631:
13909:
13339:
12629:
12411:
12161:
11351:
11250:
11034:
11025:
10609:
10400:
10391:
10185:
10176:
9734:
8551:
8492:
8413:
8217:
8183:
7840:
7795:
7785:
7732:
7724:
7671:
7661:
7614:
7452:
7421:
6991:
6908:
6583:
6519:
6396:
6386:
6258:
6099:
6089:
6056:
5826:
5696:
5600:
5522:
5512:
5450:
5407:
5384:
5362:
5321:
5313:
5014:
4869:
4821:
4813:
4779:
4740:
4710:
4664:
4656:
4607:
4597:
4552:
4544:
4483:
4442:
4426:
4373:
4357:
4298:
4252:
4244:
4197:
4148:
4138:
4097:
4059:
4051:
4014:
3998:
3901:
3891:
3748:
3577:
3528:
but not colloquial definition of 'wolf', and thus not in the scope of this article.
3398:
3252:
3186:
involves a wolf being captured for eating a duck, but is spared and sent to a zoo.
3155:
3121:
3003:
2983:
2927:
2907:
2448:
2398:
2017:
1974:
1970:
1645:
1535:
1345:
1187:
1012:
839:
566:
484:
15603:
15492:
6262:
5833:. Canadian Circumpolar Institute, Occasional Publication No. 35. pp. 275â280.
3388:
3007:
302:
15894:
15738:
15292:
15212:
15074:
14816:
14807:
14735:
14690:
14352:
14302:
14293:
14168:
14114:
14058:
13927:
13900:
13793:
13611:
13570:
13535:
13280:
12343:
12056:
12023:
12014:
11521:
11259:
10708:
10158:
10085:
10043:
9779:
9584:
9518:
9495:
9237:
8935:
8910:
8885:
8860:
8839:
8774:
8513:
8417:
7666:
7258:
6587:
6391:
6076:
Elbroch, L. M.; Lendrum, P. E.; Newsby, J.; Quigley, H.; Thompson, D. J. (2015).
5700:
5604:
4714:
4602:
4143:
3896:
3694:
3642:(in Latin) (10 ed.). HolmiĂŠ (Stockholm): Laurentius Salvius. pp. 39â40.
3480:
3268:
3195:
3190:
3129:
3033:
and involves people willingly turning into wolves to attack and kill others. The
2939:
2894:
2747:
Wolves have been persecuted in Europe for centuries, having been exterminated in
2453:
2381:
2359:
2242:
1998:
1649:
1337:
1280:
1268:
488:
15795:
15707:
15668:
7425:
5060:
4861:
4584:
Bachmann, Lutz; Wiig, Ăystein; Hansen, Anders J.; Gilbert, M. Thomas P. (2018).
3099:
1738:
518:
15562:
15549:
15348:
15039:
14936:
14789:
14708:
14510:
14257:
14150:
14123:
13176:
12793:
12393:
12257:
11940:
11477:
11459:
11390:
11001:
10992:
10594:
10555:
9509:
9421:
9309:
9139:
9130:
8866:
7055:
4660:
4361:
4002:
3342:
3205:
2947:
2923:
2915:
2839:
2827:
2721:
2713:
2619:
Convention on International Trade in Endangered Species of Wild Fauna and Flora
2550:, thus causing the animal to die within a few seconds to a minute. With small,
2547:
2225:
2029:
1966:
1882:
1221:
1168:
743:
433:
67:
8998:
6996:
6979:
6629:
Mech, L. David; Adams, L. G.; Meier, T. J.; Burch, J. W.; Dale, B. W. (2003).
6094:
5294:"Molecular and Evolutionary History of Melanism in North American Gray Wolves"
5018:
4784:
4767:
4548:
4202:
4177:
4176:
Skoglund, Pontus; Ersmark, Erik; Palkopoulou, Eleftheria; Dalén, Love (2015).
3753:
3728:
3105:
Isengrim the wolf, a character first appearing in the 12th-century Latin poem
2986:, wolves are depicted as inhabitants of graveyards and destroyers of corpses.
2726:
Berne Convention on the Conservation of European Wildlife and Natural Habitats
1180:(which first appeared around 1.4 million years ago) as an early subspecies of
1064:(mtDNA) study as one of the guides in forming his decision, and listed the 38
15838:
15650:
15207:
15132:
15113:
14986:
14753:
14649:
14622:
14248:
13946:
13891:
13413:
13357:
13098:
12739:
12194:
11756:
10921:
10888:
10690:
10681:
10618:
10518:
10476:
10328:
9921:
9255:
9170:
8981:
5612:
4582:
4438:
4369:
4010:
3840:
3568:
3358:
3168:
3151:
3079:
2990:
2855:
2823:
2761:
2606:
2602:
2193:
2070:
2009:
1981:
1973:, where tigers significantly depress wolf numbers, sometimes to the point of
1954:
1766:
1703:
1418:
1333:
1211:
1115:
1100:
1092:
1088:
1080:
954:
775:
453:
441:
420:
as demonstrated by its physical adaptations to tackling large prey, its more
392:
351:
269:
126:
121:
88:
11378:
11190:
7829:, 1839) and Comparison with Representative Wolf and Domestic Dog Haplotypes"
7774:"Local villagers' perceptions of wolves in Jiuzhaigou County, western China"
6061:
6032:
5317:
4817:
4744:
3864:
3071:
2653:
1310:
15541:
15414:
15357:
15284:
15202:
15167:
15122:
15094:
15014:
14991:
14931:
14780:
14726:
14427:
14195:
14141:
13918:
13817:
13372:
13348:
13167:
13158:
12978:
12945:
12868:
12829:
12239:
12221:
12083:
11955:
11414:
10815:
10778:
10663:
10645:
10058:
10004:
9930:
9608:
9352:
9273:
9210:
9112:
7854:
7809:
7746:
7685:
6595:
6410:
6113:
5376:
5335:
4835:
4752:
4678:
4621:
4566:
4497:
4456:
4387:
4310:
4266:
4211:
4162:
4028:
3915:
3762:
3604:
3525:
3378:
3366:
3338:
3200:
3125:
2843:
2702:
2543:
2498:
2433:
2416:
2343:
2179:
2167:
2066:
1690:
1682:
1637:
1515:
1498:
1369:
1329:
1297:
1285:
1237:
1153:
711:
396:
8987:
6739:
5894:"Conservation Action Plan for the golden jackal Canis aureus L. in Greece"
4586:"Population genomics of grey wolves and wolf-like canids in North America"
4521:
4248:
4101:
3777:
2645:
which prohibits commercial international trade in wild-sourced specimens.
1348:
of an extinct unidentified canid. This canid was genetically close to the
1341:
occurred before the divergence of the Eurasian and North American wolves.
15743:
15689:
15611:
15423:
15303:
15049:
14966:
14394:
13585:
13502:
13475:
13224:
13138:
13070:
12996:
12987:
12954:
12912:
12877:
12594:
12487:
12319:
11859:
11771:
11679:
11324:
10897:
10717:
10382:
10296:
10279:
10256:
10219:
10103:
10067:
9912:
9701:
9556:
9435:
9201:
9179:
9121:
9103:
8125:
7825:"Mitochondrial DNA Analysis of the Japanese Wolf (Canis Lupus Hodophilax
7457:
7440:
6497:
6367:"Environmental and intrinsic correlates of stress in free-ranging wolves"
4124:
3880:"How Much Is That in Dog Years? The Advent of Canine Population Genomics"
3422:
3362:
3326:
3301:
3239:
3147:
3042:
3038:
2831:
2610:
2082:
2062:
2042:
1906:
1893:
1826:
1758:
1620:
1225:
1096:
1072:
1024:
570:
514:
508:
496:
480:
414:
15502:
15466:
8126:"Never Cry Wolf: Science, Sentiment, and the Literary Rehabilitation of
5367:
5350:
4730:
3654:
2638:
15590:
15274:
15192:
15172:
14946:
14603:
13849:
13780:
13767:
13734:
13520:
13258:
13018:
12566:
12543:
12429:
12420:
12369:
12334:
12310:
12074:
11973:
11922:
11868:
11850:
11823:
11730:
11680:
11079:
11052:
10980:
10963:
10726:
10699:
10545:
10449:
10433:
10269:
10149:
9989:
9825:
9692:
9484:
9452:
9436:
8993:
8645:
Rajpurohit, K.S. (1999). "Child lifting: Wolves in Hazaribagh, India".
8518:
The Domestic Dog: Its Evolution, Behaviour and Interactions With People
7845:
7824:
7626:
6920:
5241:
4883:
3804:
The Domestic Dog: Its Evolution, Behaviour and Interactions with People
3484:
3442:
3426:
3418:
3414:
3159:
3107:
3014:
2554:
prey, wolves leap in a high arc and immobilize it with their forepaws.
2324:
2229:
2214:
1933:
1919:
1915:
1846:
1810:
1727:
1699:
1677:
1654:
1527:
1195:
437:
370:
8776:
Wolf Almanac, New and Revised: A Celebration Of Wolves And Their World
7790:
6458:. Vol. 2 (4th ed.). Johns Hopkins University Press. p.
6078:"Recolonizing wolves influence the realized niche of resident cougars"
5517:
5500:
4430:
4302:
4233:"Worldwide patterns of genomic variation and admixture in gray wolves"
2526:
981:
he listed the doglike carnivores including domestic dogs, wolves, and
440:
accompanied by their offspring. Offspring may leave to form their own
15616:
15523:
15251:
14978:
14534:
14469:
14082:
14022:
13808:
13629:
13244:
13236:
13185:
13035:
12921:
12903:
12617:
12451:
12284:
12126:
12092:
11904:
11596:
10870:
10340:
9816:
9384:
9063:
4064:
3234:
2999:
2995:
2835:
2815:
2806:
are important strongholds for wolves. The wolf has been protected in
2538:
of injury to themselves. With medium-sized prey, such as roe deer or
2364:
2221:
2159:
2151:
2143:
2113:
2102:
2074:
2050:
1943:
1925:
1862:
1858:
1830:
1806:
1790:
1786:
1762:
1715:
1711:
1543:
1523:
1386:
1325:
1257:
1252:. While some sources have suggested that this was a consequence of a
1207:
1176:
is difficult and ambiguous, with some authors choosing to include C.
1108:
562:). The name "gray wolf" refers to the grayish colour of the species.
449:
421:
210:
170:
15385:
7009:
7007:
6912:
6819:
6817:
5866:. US Department of Agriculture Division of Publications. p. 2.
5069:
Wild Mammals of North America: Biology, Management, and Conservation
4976:
4974:
4972:
4970:
4968:
4966:
4964:
4874:
3674:
A. Lehrman (1987). "Anatolian Cognates of the PIE Word for 'Wolf'".
2132:. The most harmful to wolves, particularly pups, is the mange mite (
15528:
15408:
15264:
15197:
15079:
14912:
14906:
14900:
14231:
13694:
13511:
13487:
12848:
12698:
12644:
12580:
12575:
12528:
12460:
12378:
12248:
11997:
11988:
11886:
11841:
11721:
11554:
11291:
11200:
11169:
11129:
11016:
10948:
10856:
10847:
10167:
9963:
9716:
9599:
9566:
9415:
9403:
9327:
9051:
7031:
6168:
5662:
5660:
5658:
5656:
5411:
4962:
4960:
4958:
4956:
4954:
4952:
4950:
4948:
4946:
4944:
3450:
3446:
3430:
3410:
3280:
3223:
3026:
2971:
2967:
2847:
2803:
2634:
2339:
2234:
2155:
2147:
2106:
1874:
1798:
1782:
1743:
1502:
large males can reach a weight of 69â79 kg (152â174 lb).
1410:
1249:
1199:
461:
190:
7125:
7123:
6169:"Hyenas around the cities. The case of Kaftarkhoun (Kashan- Iran)"
4765:
4055:
3849:(3rd ed.). Johns Hopkins University Press. pp. 575â577.
3089:
2910:, Italy, 13th century AD. (The twins are a 15th-century addition.)
2412:
1980:
In Israel, Palestine, Central Asia and India wolves may encounter
79:
15816:
15756:
15222:
15187:
15182:
15159:
15147:
15084:
15066:
15054:
15024:
15006:
14996:
14941:
14923:
14856:
14613:
14460:
14334:
14049:
13561:
12891:
12357:
11913:
11877:
11744:
11712:
11696:
11209:
10912:
10491:
9972:
9879:
9797:
9725:
9667:
9644:
9537:
9069:
8347:
7402:
https://www.diva-portal.org/smash/get/diva2:821120/FULLTEXT01.pdf
7004:
6814:
6787:
6348:
6346:
6321:
6319:
6294:
6292:
5842:
5840:
4525:"Interspecific Gene Flow Shaped the Evolution of the Genus Canis"
3928:
3846:
Mammal Species of the World: A Taxonomic and Geographic Reference
3350:
3346:
3334:
3330:
3112:
2819:
2811:
2571:
2441:
2402:
2352:
2334:
2086:
1950:
1911:
1889:
1723:
1707:
1406:
1362:
1314:
1306:
1276:
1245:
1186:
Considerable morphological diversity existed among wolves by the
457:
429:
374:
347:
220:
7527:
6804:
6802:
6753:
6751:
6749:
6545:
6543:
6541:
5653:
5248:
5071:(2 ed.). Johns Hopkins University Press. pp. 482â510.
4941:
4902:
4127:"Genome Sequencing Highlights the Dynamic Early History of Dogs"
3557:
2810:
since 1972. The Indian wolf is distributed across the states of
2783:
15515:
15451:
15323:
15227:
15034:
15019:
14961:
14717:
14664:
14409:
14097:
13831:
12551:
12212:
12047:
11804:
11241:
10013:
9939:
9888:
9870:
9843:
9764:
9626:
9409:
9397:
9057:
9045:
8934:
Mech, L. David; Smith, Douglas W.; MacNulty, Daniel R. (2015).
8626:
7924:. Continuum International Publishing Group. pp. 405, 437.
7379:
7120:
7096:
7019:
6935:
6880:
6126:
5290:
5105:
5103:
5067:. In Feldhamer, G. A.; Thompson, B. C.; Chapman, J. A. (eds.).
4919:
4917:
4175:
3976:
3725:
3462:
3458:
3437:
3264:
3226:
levels in instances such as snowmobiling near their territory.
3177:
3176:
both feature scenes in which wolves are hunted with hounds and
3034:
2962:
2943:
2919:
2788:
2622:
2579:
2575:
2520:
2511:
2425:
2098:
2054:
2013:
1958:
1866:
1854:
1834:
1833:
and their eggs. When such foods are insufficient, they prey on
1818:
1751:
1719:
1357:
1217:
982:
679:
582:
578:
492:
465:
388:
200:
180:
30:"Grey Wolf" and "Gray Wolf" redirect here. For other uses, see
8245:
8243:
8241:
8239:
7874:
7872:
6718:
Peters, R. P.; Mech, L. D. (1975). "Scent-marking in wolves".
6651:
6343:
6316:
6289:
6277:
6012:
5942:
5930:
5837:
5558:
5398:
Earle, M (1987). "A flexible body mass in social carnivores".
4403:
3955:
3279:
A review of the studies on the competitive effects of dogs on
3222:
Human presence appears to stress wolves, as seen by increased
2142:, unlike foxes. Endoparasites known to infect wolves include:
2057:, is largely survivable in wolves, but can be lethal to pups.
1344:
The common ancestor of the coyote and the wolf admixed with a
15790:
15237:
15217:
15177:
15127:
15102:
14699:
14558:
14501:
14374:
14177:
14034:
13661:
13643:
13266:
12936:
12185:
12170:
10831:
10122:
9617:
9076:
8715:
8713:
8657:
7823:
Ishiguro, Naotaka; Inoshima, Yasuo; Shigehara, Nobuo (2009).
6959:
6947:
6841:
6799:
6746:
6538:
5971:
5969:
5643:
5641:
5548:
5546:
4798:
4691:
3260:
3083:
3075:
3059:
2979:
2975:
2807:
2630:
2626:
2563:
2559:
2551:
2539:
2507:
2429:
2139:
2125:
2090:
1922:, usually in disputes over carcasses, sometimes eating them.
1850:
1838:
1778:
1770:
1633:
1531:
1414:
1349:
1057:
994:
978:
969:
807:
547:
403:
379:
366:
343:
230:
143:
137:
8937:
Wolves on the Hunt: The Behavior of Wolves Hunting Wild Prey
8887:
The Wolf: The Ecology and Behaviour of an Endangered Species
7509:"Conservation in conflict: Advancement and the Arabian wolf"
6978:
MacNulty, Daniel; Mech, L. David; Smith, Douglas W. (2007).
6429:
5260:
5100:
4914:
2978:
Hinduism, the wolf is a symbol of the night and the daytime
2409:
begin in the second half of winter and lasts for two weeks.
1522:
The wolf has very dense and fluffy winter fur, with a short
1465: in) in height, and the hind feet are 220â250 mm (
1034:
36 wild subspecies, and proposed two additional subspecies:
15142:
14073:
12969:
12294:
9903:
9635:
9493:
8556:
10.2193/0091-7648(2004)032[1195:HRTPSH]2.0.CO;2
8498:
10.1641/0006-3568(2006)056[0253:TFTEWN]2.0.CO;2
8317:
8305:
8255:
8236:
8105:
7922:
The Continuum Encyclopedia of Animal Symbolism in World Art
7869:
7692:
7639:
7578:
7343:"Status of large carnivore populations in Europe 2012â2016"
7284:
Beyond Wolves: The Politics of Wolf Recovery and Management
7108:
6775:
6075:
5497:
4338:"Grey wolf genomic history reveals a dual ancestry of dogs"
4334:
3256:
3045:
believed wolves were once men and viewed them as brothers.
3022:
2951:
2567:
2437:
2129:
2121:
2117:
1842:
1822:
1104:
8710:
8481:"Tools for the Edge: What's New for Conserving Carnivores"
8058:
7084:
6829:
6699:
6476:
6417:
6364:
6331:
5966:
5954:
5918:
5794:
5638:
5543:
5025:
2802:
In southern Asia, the northern regions of Afghanistan and
14951:
12203:
11812:
10140:
9157:
8540:
8431:
Boitani, L. (1983). "Wolf and dog competition in Italy".
7822:
7560:"Turkey's Wolves Are Texting Their Travels to Scientists"
6763:
5133:(2 ed.). Oxford University Press. pp. 110â112.
4472:"Deciphering the Origin of Dogs: From Fossils to Genomes"
4096:. Population Genomics. Springer, Cham. pp. 273â306.
3553:
3551:
3549:
3547:
3545:
3521:
2926:
connected the wolf with their god of war and agriculture
2041:
and Alaska. The canine parvovirus, which causes death by
1794:
974:
362:
8965:
8088:
8034:
8010:
6030:
4279:
1356:
from the other canid species. The basal position of the
6687:
6304:
5576:
3242:, an escaped wolf shot for killing livestock in England
1095:
techniques reveal that the modern wolf and the dog are
8591:
The Fear of Wolves: A Review of Wolf Attacks on Humans
8070:
7896:
5831:
Ecology and conservation of wolves in a changing world
5206:
5184:
5182:
5180:
5090:
5088:
3707:
3542:
3325:
Predatory attacks may be preceded by a long period of
2871:
2419:
pups stimulating their mother to regurgitate some food
1929:
A wolf, a bear, coyotes and ravens compete over a kill
8698:
8329:
8046:
8022:
7998:
7986:
7974:
7962:
7950:
7938:
7884:
7753:
7539:
6191:
Mills, M. G. L.; Mills, Gus; Hofer, Heribert (1998).
6131:
Large Carnivores and the Conservation of Biodiversity
5227:
4517:
4515:
1256:, other studies have suggested that this a result of
13034:
10430:
8267:
7141:
7072:
6675:
6663:
6493:
6491:
3025:
as punishment for his evil deeds. The legend of the
1899:
1644:
In North America, the coat colours of wolves follow
1405:
The wolf is the largest extant member of the family
1313:, Poland. Left: product of a male wolf and a female
556:
538:
526:
6628:
6248:
6194:
Hyaenas: status survey and conservation action plan
5806:
5626:
5272:
5194:
5177:
5085:
3558:Boitani, L.; Phillips, M. & Jhala, Y. (2023) .
3006:symbol for clans of the Pacific Northwest like the
310:Global wolf range based on IUCN's 2023 assessment.
8511:
8383:
8381:
8379:
8377:
8375:
8373:
8371:
8369:
8199:
7281:
5678:
4986:
4929:
4512:
4229:
4044:Bulletin of the American Museum of Natural History
2694:repopulation of wolves in Midwestern United States
2338:(50 sq mi). Other vocalizations include
1793:. In North America, important range-wide prey are
8933:
7037:
7025:
7013:
6977:
6941:
6488:
5584:Palaeogeography, Palaeoclimatology, Palaeoecology
4634:
4041:
3790:
2205:Pack (canine) § Pack behavior in grey wolves
1809:. Prior to their extirpation from North America,
522:, which is itself thought to be derived from the
15836:
10830:
8387:
7596:
7551:
7259:"How many wild wolves are in the United States?"
7205:"Wolf Recovery under the Endangered Species Act"
5987:
4802:"Origins and genetic legacy of prehistoric dogs"
4726:
4724:
4695:Mammalian BiologyâZeitschrift fĂŒr SĂ€ugetierkunde
3465:, have been specifically bred for wolf hunting.
3002:the "wolf trail". The wolf is also an important
2724:, the wolf is strictly protected under the 1979
43:"Wolves" redirects here. For football club, see
9028:
8638:
8520:. University Press, Cambridge. pp. 21â47.
8366:
7713:"Reconsidering the distribution of gray wolves"
7407:
7229:
7210:. US Fish and Wildlife Service. February 2007.
7172:
6898:
6190:
6166:
5751:. University of California Press. p. 144.
5748:The Last Wild Wolves: Ghosts of the Rain Forest
5153:
4087:
4085:
4083:
2974:, the wolf is ridden by gods of protection. In
1114:, and therefore should not be assessed for the
1091:that Linnaeus studied in Sweden. Studies using
8505:
7502:
7500:
7478:. Wolves and Humans Foundation. Archived from
7337:
7335:
6637:. University of Minnesota Press. p. 163.
6229:
5738:
5493:
5491:
5120:
5118:
4578:
4576:
3825:
3693:Young, Stanley P.; Goldman, Edward A. (1944).
3629:
3627:
3625:
3583:10.2305/IUCN.UK.2023-1.RLTS.T3746A247624660.en
3468:
1888:In times of scarcity, wolves will readily eat
1291:
477:International Union for Conservation of Nature
14872:
11694:
9535:
9368:
9014:
8909:Mech, L. David; Boitani, Luigi, eds. (2003).
8816:
8800:Wolves in Russia: Anxiety throughout the ages
8663:
8597:. Norsk Institutt for Naturforskning (NINA).
8156:
7915:
7913:
7911:
7816:
7590:
6971:
6965:
6953:
6823:
6793:
6757:
6447:
5988:Sunquist, Melvin E.; Sunquist, Fiona (2002).
5899:. World Wildlife Fund Greece. pp. 1â47.
5885:
5666:
5266:
5254:
5223:
5221:
5147:
5109:
5054:
5052:
5050:
5048:
5046:
5044:
5042:
5040:
4980:
4923:
4908:
4792:
4721:
4469:
4091:
3877:
3796:
9382:
8424:
7473:"The Wolf in Russiaâsituations and problems"
7441:"Hunting as Sustainable Wildlife Management"
7306:
7135:
6892:
6120:
5981:
5715:
5284:
5058:
4998:
4685:
4463:
4273:
4169:
4118:
4080:
3719:
3692:
3132:. Villainous wolf characters also appear in
2861:
2112:Wolves are often infested with a variety of
1829:and smaller carnivores. They frequently eat
953:In 1758, the Swedish botanist and zoologist
15865:Extant Middle Pleistocene first appearances
8912:Wolves: Behaviour, Ecology and Conservation
8908:
8731:
8725:
8632:
8583:
8581:
8579:
8577:
8575:
8573:
8534:
8472:
8352:. Oxford University Press. pp. 69â93.
8350:Free-Ranging Dogs and Wildlife Conservation
8323:
8311:
8261:
8249:
8111:
7878:
7704:
7698:
7584:
7557:
7533:
7497:
7432:
7385:
7332:
7251:
7166:
7144:"Schedule 3 (section 26) Protected Species"
7129:
7114:
7102:
7090:
6886:
6847:
6835:
6808:
6781:
6769:
6711:
6705:
6657:
6564:"Wolf-Pack Buffer Zones as Prey Reservoirs"
6549:
6482:
6435:
6423:
6352:
6337:
6325:
6298:
6283:
6160:
6069:
6024:
6018:
5975:
5960:
5948:
5936:
5924:
5846:
5800:
5647:
5564:
5552:
5488:
5342:
5115:
5031:
4855:
4853:
4573:
4035:
3961:
3922:
3622:
3189:Wolves are among the central characters of
2233:consisting of up to 42 wolves being known.
1814:particularly during the autumn and winter.
1666:
1429:A wolf skeleton housed in the Wolf Museum,
14879:
14865:
9555:
9375:
9361:
9021:
9007:
8669:
8644:
8341:
7908:
7413:
7288:. University of Minnesota Press. pp.
6717:
6358:
6242:
5852:
5765:
5744:
5426:
5218:
5037:
4470:Freedman, Adam H; Wayne, Robert K (2017).
4225:
4223:
4221:
3871:
3831:
3673:
3372:
3317:(1833) by François Grenier de Saint-Martin
3048:
2878:Wolves in folklore, religion and mythology
2585:
1589: in), but can reach 110â130 mm (
377:, and is further distinguished from other
301:
115:
78:
13778:
10267:
9450:
8682:. G. P. Putnam's sons. pp. 179â207.
8675:
8496:
7844:
7799:
7789:
7771:
7736:
7675:
7665:
7456:
6995:
6622:
6555:
6441:
6400:
6390:
6223:
6184:
6103:
6093:
6060:
5891:
5818:
5672:
5526:
5516:
5391:
5366:
5325:
4873:
4825:
4783:
4759:
4668:
4611:
4601:
4556:
4487:
4446:
4377:
4256:
4201:
4152:
4142:
4063:
4018:
3905:
3895:
3752:
3581:
3229:
2705:population numbers over 250 individuals.
2358:Scent marking involves urine, feces, and
2138:), though they rarely develop full-blown
597:Canine phylogeny with ages of divergence
12124:
8570:
8445:
8082:
7558:ĆekercioÄlu, ĂaÄan (December 15, 2013).
7273:
7197:
7052:The Wolf: His Place in the Natural World
6853:
5721:
5501:"Food habits of the world's grey wolves"
5461:
5004:
4850:
4628:
3392:
3309:
3274:
3233:
3088:
2887:
2782:
2712:
2690:reintroductions in Yellowstone and Idaho
2652:
2592:List of gray wolf populations by country
2525:
2497:
2411:
2380:
2208:
1997:
1969:interactions are well-documented in the
1942:Wolves also dominate and sometimes kill
1924:
1885:, grain crops, and the shoots of reeds.
1737:
1676:
1619:
1509:
1424:
1396:
1361:and wolves show varying degrees of past
1352:and evolved after the divergence of the
1301:
1137:
11198:
9961:
9482:
8587:
8430:
7919:
7772:Xu, Yu; Yang, Biao; Dou, Liang (2015).
7765:
7633:
7465:
7185:from the original on September 30, 2019
6863:The Dog: Its Domestication and Behavior
5994:. University of Chicago Press. p.
5348:
5163:. Oxford University Press. p. 45.
5131:Dog Behaviour, Evolution, and Cognition
5124:
4218:
3807:. Cambridge University Press. pp.
3633:
1389:, with little evidence of the reverse.
1317:; right: from a female wolf and a male
456:as well as smaller animals, livestock,
14:
15837:
14291:
12527:
8858:
8817:Heptner, V. G.; Naumov, N. P. (1998).
8797:
8719:
8478:
8335:
8193:
8162:
8117:
8076:
8064:
8040:
8016:
7992:
7980:
7944:
7902:
7710:
7506:
7438:
7349:from the original on September 2, 2019
7142:Government of Canada (July 29, 2019).
7043:
6448:Nowak, R. M.; Paradiso, J. L. (1983).
6310:
5858:
5771:
5432:
3713:
3686:
3667:
2930:, and believed their city's founders,
2487:
15390:
15389:
14860:
13765:
13222:
12526:
11678:
10813:
10254:
9434:
9356:
9002:
8837:
8772:
8704:
8512:Coppinger, R.; Schneider, R. (1995).
8439:
8144:from the original on October 12, 2013
8123:
8052:
8028:
8004:
7968:
7956:
7890:
7759:
7545:
7515:from the original on November 7, 2019
7049:
6693:
6681:
6669:
5906:from the original on December 9, 2017
5829:; Fritts, S. H.; Seip, D. R. (eds.).
5824:
5397:
5278:
5212:
5200:
5188:
5094:
4399:
4397:
4330:
4328:
3972:
3970:
1949:Wolves may interact and compete with
15822:2F15C6CE-6C7A-4AF8-AE71-087151318E0E
15731:309809e8-1a21-45a0-80a0-1f13fee46c52
15568:3610bdd7-7432-4d43-9f6e-6684deb82987
8883:
8285:
8273:
7566:from the original on October 6, 2019
7173:State of Alaska (October 29, 2019).
7078:
6561:
5812:
5632:
5622:– via Elsevier Science Direct.
5467:
4992:
4935:
4859:
4489:10.1146/annurev-animal-022114-110937
3315:Country children surprised by a wolf
3295:
2768:, and around 2,500 in the northwest
1103:. In 2019, a workshop hosted by the
985:. He classified the domestic dog as
361:have been recognized, including the
15845:IUCN Red List least concern species
10745:South African small-spotted genet (
7420:. Fauna & Flora International.
7279:
7217:from the original on August 3, 2019
6859:
6045:, in Montana, Alberta, and Wyoming"
5780:. Chartwell Crestline. p. 53.
4476:Annual Review of Animal Biosciences
3878:Larson, G.; Bradley, D. G. (2014).
3696:The Wolves of North America, Part I
3569:IUCN Red List of Threatened Species
3413:with dogs (usually combinations of
3146:, and are included in the works of
2872:In folklore, religion and mythology
2198:
1044:(Meyer, 1793). Wozencraft included
327:
24:
14886:
8686:from the original on June 24, 2015
8286:Levy, Max G. (December 11, 2020).
7729:10.24272/j.issn.2095-8137.2017.021
7619:10.1111/j.1523-1739.1991.tb00354.x
7179:Alaska Department of Fish and Game
7154:from the original on April 9, 2019
6631:"Ch.8-The Denali Wolf-Prey System"
6610:from the original on July 24, 2018
6167:Monchot, H.; Mashkour, H. (2010).
6134:. Island Press. pp. 179â207.
5873:from the original on July 24, 2019
5059:Paquet, P.; Carbyn, L. W. (2003).
4890:from the original on July 31, 2019
4394:
4325:
3967:
3775:
3652:
3475:Wolves as pets and working animals
3140:The Wolf and the Seven Young Goats
3062:featured wolves in several of his
2922:, the god of light and order. The
2659:Sevilleta National Wildlife Refuge
2323:Problems playing these files? See
2274:
1761:, the wolf feeds predominantly on
1194:, a pronounced development of the
513:The English "wolf" stems from the
25:
15916:
11089:Broad-striped Malagasy mongoose (
8959:
8614:from the original on May 17, 2014
6211:from the original on May 16, 2016
6148:from the original on June 3, 2016
5859:Merrit, Dixon (January 7, 1921).
5722:Woodford, Riley (November 2004).
4231:Bisong; Wayne, Robert K. (2016).
3203:'s largely fictional 1963 memoir
3037:have traditionally believed that
3021:being transformed into a wolf by
2858:was captured and killed in 1905.
2256:Denali National Park and Preserve
1994:Parasites and pathogens of wolves
1900:Interactions with other predators
1288:diverged around 5,400 years ago.
10814:
8823:. Science Publishers, Inc. USA.
8734:"Can You Turn a Wolf into a Dog"
8279:
8222:10.1046/j.1523-1739.2002.00554.x
7391:
7361:
7239:. U.S. Fish and Wildlife Service
6524:10.1111/j.0906-7590.2007.04826.x
6232:Canid Biology & Conservation
5455:10.1111/j.1502-3931.2007.00091.x
2648:
2546:, severing nerve tracks and the
2308:
2290:
2261:
1685:in a mountainous habitat in the
1376:, ten percent of dogs including
1368:In more recent times, some male
1060:. Wozencraft referred to a 1999
901:
881:
842:
810:
778:
746:
714:
682:
662:
586:and Wolfdregil ("Wolf Runner").
428:, including individual or group
157:
13223:
10255:
8844:. J. M. Dent and Sons Limited.
8766:
7507:Fisher, A. (January 29, 2019).
7038:Mech, Smith & MacNulty 2015
7026:Mech, Smith & MacNulty 2015
7014:Mech, Smith & MacNulty 2015
6942:Mech, Smith & MacNulty 2015
5724:"Alaska's Salmon-Eating Wolves"
5570:
3942:IUCN/SSC Canid Specialist Group
3769:
2942:includes the feared giant wolf
2494:Hunting behavior of gray wolves
2370:
1857:. In North America, wolves eat
1757:Like all land mammals that are
1324:In the distant past, there was
1148:, the wolf's immediate ancestor
13766:
9247:Northern Rocky Mountain wolf (
8732:Tucker, P.; Weide, B. (1998).
8396:) in a conservation context".
8134:The Canadian Historical Review
5861:"World's Greatest Animal Dead"
3797:Clutton-Brock, Juliet (1995).
3646:
3597:
3514:
3497:
3397:Carcasses of hunted wolves in
3111:, is a major character in the
2982:must escape from its jaws. In
1392:
1166:(which in turn descended from
546:may also be the source of the
13:
1:
10501:Malabar large-spotted civet (
8966:The International Wolf Center
8892:University of Minnesota Press
6455:Walker's Mammals of the World
6263:10.1080/09397140.2016.1144292
6049:The Canadian Field-Naturalist
3535:
2866:
2696:has been concentrated in the
1987:
1000:
14763:Indonesian mountain weasel (
14419:North American river otter (
13307:Humboldt's hog-nosed skunk (
10361:Macrogalidia musschenbroekii
9265:Alexander Archipelago wolf (
9229:Vancouver Coastal Sea wolf (
8971:Staying Safe in Wolf Country
8676:Roosevelt, Theodore (1909).
8418:10.1016/j.biocon.2014.01.032
7667:10.1371/journal.pone.0215019
6866:. Garland STPM. p. 33.
6588:10.1126/science.198.4314.320
6392:10.1371/journal.pone.0137378
5892:Giannatos, G. (April 2004).
5701:10.1016/j.mambio.2018.03.007
5605:10.1016/j.palaeo.2021.110368
5351:"Wolf of a different colour"
4715:10.1016/j.mambio.2010.02.004
4603:10.1371/journal.pgen.1007745
4144:10.1371/journal.pgen.1004016
3897:10.1371/journal.pgen.1004093
3217:
3120:", first written in 1697 by
2898:, sculpture of the mythical
2401:annually being the average.
2385:Korean wolves mating in the
2185:
1275:between the two. The dingo,
1121:
502:
45:Wolverhampton Wanderers F.C.
7:
15890:Taxa named by Carl Linnaeus
11624:Somalian slender mongoose (
8942:University of Chicago Press
8917:University of Chicago Press
8514:"Evolution of working dogs"
7426:10.13140/RG.2.2.10128.20480
7414:Goldthorpe, Gareth (2016).
7404:Retrieved 30 September 2023
3801:. In Serpell, James (ed.).
3783:Online Etymology Dictionary
3660:Online Etymology Dictionary
3469:As pets and working animals
3449:, wolves are traditionally
3389:Human uses of hunted wolves
2795:300â600 wolves inhabit the
2061:carried by wolves include:
2026:infectious canine hepatitis
2012:carried by wolves include:
1309:in the wild animal park at
1292:Admixture with other canids
1156:descent of the extant wolf
1020:Mammal Species of the World
973:is the Latin word meaning "
589:
565:Since pre-Christian times,
464:and parasites, notably the
10:
15921:
15900:Habitats Directive species
14577:Asian small-clawed otter (
13316:American hog-nosed skunk (
13298:Molina's hog-nosed skunk (
11606:Angolan slender mongoose (
11301:Ethiopian dwarf mongoose (
10350:Small-toothed palm civet (
8588:Linnell, J. D. C. (2002).
8182:(3): 11â13. Archived from
7261:. Wolf Conservation Center
6251:Zoology in the Middle East
5778:Wolves: Spirit of the Wild
4661:10.1016/j.isci.2019.09.008
4362:10.1038/s41586-022-04824-9
4003:10.1038/s41598-022-06812-5
3472:
3386:
3376:
3299:
3164:Leonid Pavlovich Sabaneyev
3052:
2881:
2875:
2708:
2661:in New Mexico, as part of
2589:
2502:Wolves pursuing a bull elk
2491:
2374:
2265:
2258:were due to other wolves.
2202:
2191:
1991:
1670:
1661:
1295:
1134:Origin of the domestic dog
1131:
1125:
1010:
1004:
557:
539:
506:
424:, and its highly advanced
42:
32:Grey Wolf (disambiguation)
29:
15875:Mammals described in 1758
15398:
15373:Grizzlyâpolar bear hybrid
15368:Grizzlyâblack bear hybrid
15346:
15321:
15301:
15283:
15250:
15158:
15111:
15093:
15065:
15005:
14977:
14922:
14894:
14662:
14611:
14602:
14556:
14532:
14499:
14482:
14458:
14407:
14390:
14373:
14344:Saharan striped polecat (
14332:
14315:
14274:
14229:
14212:
14203:
14194:
14095:
14071:
14047:
14030:
14021:
13944:
13889:
13847:
13830:
13806:
13789:
13774:
13761:
13733:
13692:
13659:
13642:
13583:
13533:
13500:
13483:
13474:
13411:
13370:
13337:
13325:Striped hog-nosed skunk (
13278:
13257:
13235:
13231:
13218:
13154:
13137:
13096:
13068:
13056:Mediterranean monk seal (
13051:
13017:
12967:
12934:
12899:
12890:
12864:
12847:
12828:
12785:Juan FernĂĄndez fur seal (
12749:South American fur seal (
12737:
12720:
12696:
12684:South American sea lion (
12679:
12642:
12625:
12616:
12592:
12564:
12542:
12538:
12522:
12486:
12459:
12450:
12391:
12365:
12356:
12332:
12308:
12293:
12282:
12183:
12157:
12109:
12066:South American gray fox (
12036:
12010:
11986:
11969:
11954:
11939:
11802:
11769:
11752:
11743:
11710:
11690:
11674:
11642:Common slender mongoose (
11594:
11552:
11508:
11412:
11386:
11377:
11322:
11289:
11272:
11239:
11222:
11189:
11168:
11127:
11115:Narrow-striped mongoose (
11110:
11077:
11060:
11051:
11014:
10988:
10979:
10962:
10946:
10910:
10884:
10869:
10845:
10826:
10822:
10809:
10777:
10773:
10637:Crested servaline genet (
10592:
10553:
10544:
10489:
10463:
10448:
10426:
10380:
10336:
10327:
10278:
10263:
10250:
10218:
10120:
10056:
10039:
10030:
10002:
9985:
9952:
9901:
9814:
9777:
9760:
9751:
9714:
9690:
9681:
9665:
9597:
9564:
9551:
9531:
9478:
9461:
9446:
9430:
9392:
9192:
9094:
9040:
8664:Heptner & Naumov 1998
8544:Wildlife Society Bulletin
7920:Werness, Hope B. (2007).
6997:10.1644/06-MAMM-A-119R1.1
6966:Heptner & Naumov 1998
6954:Heptner & Naumov 1998
6824:Heptner & Naumov 1998
6794:Heptner & Naumov 1998
6758:Heptner & Naumov 1998
6095:10.1186/s40555-015-0122-y
5726:. Wildlifenews.alaska.gov
5667:Heptner & Naumov 1998
5471:The Wolves of Isle Royale
5267:Heptner & Naumov 1998
5255:Heptner & Naumov 1998
5230:Wildlife Society Bulletin
5110:Heptner & Naumov 1998
5019:10.1017/S0952836905007430
4981:Heptner & Naumov 1998
4924:Heptner & Naumov 1998
4909:Heptner & Naumov 1998
4785:10.1007/s10592-013-0547-y
4549:10.1016/j.cub.2018.08.041
4203:10.1016/j.cub.2015.04.019
3754:10.1016/j.cub.2015.06.060
2906:, from the legend of the
2862:Relationships with humans
2675:Canada National Parks Act
2004:Yellowstone National Park
1883:European black nightshade
1505:
1382:jackal-wolf hybridization
1028:W. Christopher Wozencraft
1007:Subspecies of Canis lupus
896:
876:
869:
837:
805:
773:
741:
709:
677:
657:
650:
643:
636:
629:
622:
615:
608:
309:
300:
285:
280:
259:
252:
154:Scientific classification
152:
135:
113:
104:
86:
77:
53:
15860:Mammals of North America
14700:Stoat/Beringian ermine (
14568:African clawless otter (
14320:African striped weasel (
14133:Yellow-throated marten (
14009:Formosan ferret-badger (
13713:Western mountain coati (
13704:Eastern mountain coati (
13621:Western lowland olingo (
13603:Eastern lowland olingo (
13431:Southern spotted skunk (
13125:Southern elephant seal (
13116:Northern elephant seal (
12906:Pagophilus groenlandicus
11683:("dog-like" carnivorans)
11496:Stripe-necked mongoose (
11117:Mungotictis decemlineata
9992:Herpailurus yagouaroundi
9439:("cat-like" carnivorans)
9220:British Columbian wolf (
8838:Lopez, Barry H. (1978).
8802:. Detselig Enterprises.
8781:Rowman & Littlefield
8516:. In Serpell, J. (ed.).
8479:Shivik, John A. (2006).
7175:"Wolf Hunting in Alaska"
6197:. IUCN. pp. 24â25.
3843:; Reeder, D. M. (eds.).
3490:
3255:(northern Scandinavia),
3166:, and others. Tolstoy's
3055:List of fictional wolves
2846:has roughly 650 wolves,
2682:contiguous United States
2657:Captive Mexican wolf at
2530:Two wolves feeding on a
2116:exoparasites, including
1667:Distribution and habitat
1017:In the third edition of
534:Proto-Indo-European root
452:and feeds on large wild
15880:Pleistocene carnivorans
14745:Yellow-bellied weasel (
14000:Burmese ferret-badger (
13982:Chinese ferret-badger (
13973:Bornean ferret badger (
13964:Vietnam ferret-badger (
13449:Eastern spotted skunk (
13440:Western spotted skunk (
13170:Leptonychotes weddellii
12812:Subantarctic fur seal (
12758:Australasian fur seal (
11582:Black-footed mongoose (
11564:Bushy-tailed mongoose (
11513:White-tailed mongoose (
11451:Indian brown mongoose (
11433:Short-tailed mongoose (
11424:Small Indian mongoose (
11361:Flat-headed kusimanse (
11334:Alexander's kusimanse (
11310:Common dwarf mongoose (
11277:Pousargues's mongoose (
11139:Brown-tailed mongoose (
11098:Grandidier's mongoose (
10352:Arctogalidia trivirgata
9576:Sunda clouded leopard (
9319:Alaskan Interior wolf (
8884:Mech, L. David (1981).
8633:Mech & Boitani 2003
8398:Biological Conservation
8324:Mech & Boitani 2003
8312:Mech & Boitani 2003
8262:Mech & Boitani 2003
8250:Mech & Boitani 2003
8112:Mech & Boitani 2003
7879:Mech & Boitani 2003
7711:Larson, Greger (2017).
7699:Mech & Boitani 2003
7585:Mech & Boitani 2003
7562:. National Geographic.
7534:Mech & Boitani 2003
7439:Baskin, Leonid (2016).
7386:Mech & Boitani 2003
7345:. European Commission.
7130:Mech & Boitani 2003
7115:Mech & Boitani 2003
7103:Mech & Boitani 2003
7091:Mech & Boitani 2003
6887:Mech & Boitani 2003
6848:Mech & Boitani 2003
6836:Mech & Boitani 2003
6809:Mech & Boitani 2003
6782:Mech & Boitani 2003
6770:Mech & Boitani 2003
6706:Mech & Boitani 2003
6658:Mech & Boitani 2003
6550:Mech & Boitani 2003
6483:Mech & Boitani 2003
6436:Mech & Boitani 2003
6424:Mech & Boitani 2003
6353:Mech & Boitani 2003
6338:Mech & Boitani 2003
6326:Mech & Boitani 2003
6299:Mech & Boitani 2003
6284:Mech & Boitani 2003
6062:10.22621/cfn.v122i1.550
6019:Mech & Boitani 2003
5976:Mech & Boitani 2003
5961:Mech & Boitani 2003
5949:Mech & Boitani 2003
5937:Mech & Boitani 2003
5925:Mech & Boitani 2003
5847:Mech & Boitani 2003
5801:Mech & Boitani 2003
5745:McAllister, I. (2007).
5648:Mech & Boitani 2003
5565:Mech & Boitani 2003
5553:Mech & Boitani 2003
5468:Mech, L. David (1966).
5349:Hedrick, P. W. (2009).
5318:10.1126/science.1165448
5160:Encyclopedia of Mammals
5032:Mech & Boitani 2003
4860:Mech, L. David (1974).
4818:10.1126/science.aba9572
3962:Mech & Boitani 2003
3373:Human hunting of wolves
3238:A 1905 postcard of the
3049:In fable and literature
3029:has been widespread in
2918:associated wolves with
2601:classifies the wolf as
2586:Status and conservation
1733:
1636:(yellow to orange) and
1378:livestock guardian dogs
1260:homogenising ancestry.
1023:published in 2005, the
528:
517:
15233:Congolese spotted lion
14586:Congo clawless otter (
14489:Hydrictis maculicollis
14487:Spotted-necked otter (
14446:Southern river otter (
14397:Pteronura brasiliensis
14160:European pine marten (
13671:South American coati (
13399:Palawan stink badger (
12725:New Zealand sea lion (
11790:Japanese raccoon dog (
11487:Crab-eating mongoose (
11442:Indian grey mongoose (
11065:Ring-tailed vontsira (
10876:(bone-crushing hyenas)
10571:Central African oyan (
10132:Chinese mountain cat (
9283:Mackenzie River wolf (
8859:Marvin, Garry (2012).
8433:Acta Zoologica Fennica
8163:Grooms, Steve (2008).
6041:, and a Grizzly Bear,
5991:Wild cats of the world
5772:Fuller, T. K. (2019).
3931:Jhala, Yadvendradev V.
3799:"2-Origins of the dog"
3634:LinnĂŠus, Carl (1758).
3505:Canus lupus familiaris
3483:are sometimes kept as
3402:
3383:Wolf hunting with dogs
3318:
3243:
3230:Predation on livestock
3118:Little Red Riding Hood
3102:
3095:Little Red Riding Hood
3068:The Boy Who Cried Wolf
2961:, the wolf represents
2911:
2791:
2717:
2686:Endangered Species Act
2665:
2534:
2503:
2420:
2390:
2279:
2217:
2006:
1930:
1754:
1694:
1629:
1519:
1434:
1402:
1321:
1149:
1050:New Guinea singing dog
402:Of all members of the
15778:Paleobiology Database
15333:Polecatâferret hybrid
14826:Back-striped weasel (
14772:Black-footed ferret (
14544:Smooth-coated otter (
14322:Poecilogale albinucha
13991:Javan ferret-badger (
13877:Sumatran hog badger (
13859:Northern hog badger (
13553:Crab-eating raccoon (
13458:Pygmy spotted skunk (
13179:Lobodon carcinophagus
12915:Histriophoca fasciata
12708:Australian sea lion (
12654:California sea lion (
12403:American black bear (
12145:Black-backed jackal (
12136:Side-striped jackal (
11976:Chrysocyon brachyurus
11400:Long-nosed mongoose (
10673:Rusty-spotted genet (
10510:Large-spotted civet (
10359:Sulawesi palm civet (
10288:Owston's palm civet (
9862:Andean mountain cat (
9338:Alaskan tundra wolf (
8798:Graves, Will (2007).
8773:Busch, R. H. (2007).
8446:Backeryd, J. (2007).
8165:"The Mixed Legacy of
7605:(4). Wiley: 476â483.
7485:on September 23, 2007
7400:Ramsey, Morag (2015)
6037:, Killed by Cougars,
5774:"Ch3-What wolves eat"
5528:10536/DRO/DU:30085823
5157:; Norris, S. (2001).
4772:Conservation Genetics
4745:10.1093/jhered/esu014
4249:10.1101/gr.197517.115
4102:10.1007/13836_2018_27
3576:: e.T3746A247624660.
3396:
3313:
3275:Competition with dogs
3237:
3135:The Three Little Pigs
3092:
2891:
2786:
2764:, 1,100â2,400 in the
2760:, 1,700â2,240 in the
2716:
2656:
2590:Further information:
2529:
2501:
2415:
2384:
2278:
2213:Indian wolves at the
2212:
2047:electrolyte imbalance
2001:
1928:
1741:
1680:
1655:Black-coloured wolves
1623:
1513:
1431:Abruzzo National Park
1428:
1401:A North American wolf
1400:
1305:
1269:Late Pleistocene wolf
1254:population bottleneck
1204:Pleistocene megafauna
1141:
1132:Further information:
1128:Evolution of the wolf
1071:under the biological
1038:(Linnaeus, 1758) and
1011:Further information:
965:binomial nomenclature
468:, may infect wolves.
395:, to produce fertile
387:species, such as the
373:member of the family
334:), also known as the
36:Wolf (disambiguation)
15563:Fauna Europaea (new)
14641:Long-tailed weasel (
14219:Lyncodon patagonicus
13868:Greater hog badger (
13390:Sunda stink badger (
13080:Hawaiian monk seal (
12803:Guadalupe fur seal (
12776:Antarctic fur seal (
12767:GalĂĄpagos fur seal (
12663:GalĂĄpagos sea lion (
11781:Common raccoon dog (
11633:Cape gray mongoose (
11573:Jackson's mongoose (
11533:Paracynictis selousi
11524:Cynictis penicillata
11148:Durrell's vontsira (
10736:Giant forest genet (
10528:Large Indian civet (
10468:Small Indian civet (
10077:Sunda leopard cat (
9853:Southern tiger cat (
9789:African golden cat (
9466:African palm civet (
9292:Baffin Island wolf (
8292:Smithsonian Magazine
8202:Conservation Biology
7599:Conservation Biology
7536:, pp. 323, 327.
7458:10.3106/041.041.0402
7148:Justice Laws Website
7058:. pp. 217â218.
7050:Zimen, Erik (1981).
6984:Journal of Mammalogy
6901:Journal of Mammalogy
6635:The Wolves of Denali
6562:Mech, L. D. (1977).
6173:Journal of Taphonomy
5125:Miklosi, A. (2015).
4535:(21): 3441â3449.e5.
4336:V. (July 14, 2022).
3605:"Appendices | CITES"
3524:are included in the
3306:List of wolf attacks
2934:, were suckled by a
2758:Carpathian Mountains
2387:Tama Zoological Park
2174:, irritation of the
1975:localized extinction
1939:American black bears
1748:Denali National Park
1626:La BoissiÚre-du-Doré
1244:), originating from
1242:Last Glacial Maximum
1236:to other wolves and
1142:Life restoration of
550:word for the animal
426:expressive behaviour
15905:Paraphyletic groups
15338:Polecatâmink hybrid
14684:M. aistoodonnivalis
14520:Hairy-nosed otter (
14437:Neotropical otter (
14217:Patagonian weasel (
13680:White-nosed coati (
12880:Cystophora cristata
12871:Erignathus barbatus
12604:Northern fur seal (
12381:Helarctos malayanus
12017:Atelocynus microtis
11724:U. cinereoargenteus
11615:Egyptian mongoose (
11542:Rhynchogale melleri
11540:Meller's mongoose (
11531:Selous's mongoose (
11515:Ichneumia albicauda
11469:Collared mongoose (
11343:Angolan kusimanse (
11227:Liberian mongoose (
10580:West African oyan (
10479:Civettictis civetta
10410:Golden palm civet (
10368:Masked palm civet (
10343:Arctictis binturong
10317:Hemigalus derbyanus
10315:Banded palm civet (
10306:Hose's palm civet (
10095:Rusty-spotted cat (
9334:(taxonomy disputed)
9301:Northwestern wolf (
8750:on December 8, 2015
8722:, pp. 121â140.
8635:, pp. 300â304.
8410:2014BCons.171..232L
8214:2002ConBi..16..809C
8095:Yellowstone Science
8067:, pp. 21, 123.
7717:Zoological Research
7658:2019PLoSO..1415019S
7611:1991ConBi...5..476J
7511:. Middle East Eye.
7388:, pp. 318â320.
7368:European Commission
7280:Nie, M. A. (2003).
7132:, pp. 321â324.
7105:, pp. 122â125.
6889:, pp. 119â121.
6860:Fox, M. W. (1978).
6826:, pp. 249â254.
6796:, pp. 234â238.
6732:1975AmSci..63..628P
6580:1977Sci...198..320M
6516:2007Ecogr..30...66J
6450:"Carnivora;Canidae"
6383:2015PLoSO..1037378M
6355:, pp. 2â3, 28.
6328:, pp. 202â208.
6301:, pp. 211â213.
6286:, pp. 208â211.
6021:, pp. 264â265.
5951:, pp. 263â264.
5939:, pp. 261â263.
5849:, pp. 266â268.
5693:2018MamBi..91...30G
5669:, pp. 213â231.
5597:2021PPP...57110368L
5567:, pp. 109â110.
5447:2008Letha..41..333S
5400:American Naturalist
5368:10.1038/hdy.2009.77
5310:2009Sci...323.1339A
5304:(5919): 1339â1343.
5257:, pp. 168â169.
4983:, pp. 164â270.
4911:, pp. 129â132.
4733:Journal of Heredity
4707:2010MamBi..75..341I
4653:2019iSci...20..110W
4541:2018CBio...28E3441G
4423:2020MolEc..29.1596L
4354:2022Natur.607..313B
4295:2021MolEc..30.6687H
4194:2015CBio...25.1515S
3995:2022NatSR..12.2882I
3964:, pp. 239â245.
3745:2015CBio...25.2158K
3703:, Inc. p. 390.
3520:Domestic and feral
2787:A wolf in southern
2778:former Soviet Union
2488:Hunting and feeding
2454:German Shepherd Dog
2377:Canine reproduction
2164:thorny-headed worms
2079:bovine tuberculosis
1354:African hunting dog
1319:West Siberian Laika
1198:muscle, and robust
1077:nominate subspecies
899:Black-backed jackal
879:Side-striped jackal
409:, the wolf is most
354:. More than thirty
107:Conservation status
15138:Lord Morton's mare
14799:European polecat (
14632:Colombian weasel (
14282:(African polecats)
13910:Caucasian badger (
13838:(Eurasian badgers)
13288:(hog-nosed skunks)
13025:("southern seals")
12924:Halichoerus grypus
12855:("northern seals")
12727:Phocarctos hookeri
12632:Eumetopias jubatus
12630:Steller sea lion (
12412:Asian black bear (
12162:African wild dog (
12116:(wolf-like canids)
11393:Atilax paludinosus
11352:Common kusimanse (
11279:Dologale dybowskii
11251:Gambian mongoose (
11035:Western falanouc (
11026:Eastern falanouc (
10995:Cryptoprocta ferox
10900:Parahyaena brunnea
10655:Johnston's genet (
10610:Abyssinian genet (
10562:(African linsangs)
10470:Viverricula indica
10401:Brown palm civet (
10392:Asian palm civet (
10299:Cynogale bennettii
10290:Chrotogale owstoni
10186:European wildcat (
10177:Black-footed cat (
9767:Leptailurus serval
9735:Asian golden cat (
9501:(Asiatic linsangs)
9303:C. l. occidentalis
8176:International Wolf
8124:Jones, K. (2001).
7846:10.2108/zsj.26.765
7833:Zoological Science
6720:American Scientist
6660:, pp. 66â103.
6082:Zoological Studies
5127:"Ch. 5.5.2âWolves"
5007:Journal of Zoology
3983:Scientific Reports
3701:Dover Publications
3425:), poisoning with
3406:Theodore Roosevelt
3403:
3319:
3244:
3183:Peter and the Wolf
3144:Russian literature
3103:
2954:'s faithful pets.
2912:
2902:feeding the twins
2884:Wolves in heraldry
2792:
2718:
2666:
2535:
2504:
2421:
2391:
2280:
2268:Wolf communication
2218:
2172:allergic reactions
2059:Bacterial diseases
2034:canine coronavirus
2007:
1963:niche partitioning
1931:
1871:lily of the valley
1755:
1742:A wolf carrying a
1718:(including Arctic
1695:
1630:
1520:
1435:
1403:
1374:Caucasus Mountains
1322:
1226:Indian plains wolf
1150:
1145:Canis mosbachensis
989:, and the wolf as
977:", and under this
61:Middle Pleistocene
15855:Mammals of Europe
15832:
15831:
15765:Open Tree of Life
15392:Taxon identifiers
15383:
15382:
15045:Jackalâdog hybrid
14957:Sheepâgoat hybrid
14854:
14853:
14850:
14849:
14846:
14845:
14842:
14841:
14838:
14837:
14817:Siberian weasel (
14808:American ermine (
14736:Japanese weasel (
14691:Mountain weasel (
14675:
14598:
14597:
14384:
14369:
14368:
14365:
14364:
14353:Striped polecat (
14303:Marbled polecat (
14285:
14270:
14269:
14242:
14190:
14189:
14169:Japanese marten (
14115:American marten (
14108:
13966:M. cucphuongensis
13957:
13940:
13939:
13928:European badger (
13901:Japanese badger (
13841:
13794:American badger (
13757:
13756:
13753:
13752:
13729:
13728:
13725:
13724:
13653:
13612:Northern olingo (
13596:
13571:Cozumel raccoon (
13546:
13470:
13469:
13424:
13383:
13291:
13272:
13214:
13213:
13210:
13209:
13206:
13205:
13202:
13201:
13198:
13197:
13188:Ommatophoca rossi
13161:Hydrurga leptonyx
13148:
13145:(Antarctic seals)
13118:M. angustirostris
13109:
13092:
13091:
13058:Monachus monachus
13045:
13028:
13013:
13012:
13009:
13008:
12858:
12841:
12824:
12823:
12686:Otaria flavescens
12675:
12674:
12586:
12554:Odobenus rosmarus
12518:
12517:
12514:
12513:
12510:
12509:
12506:
12505:
12469:
12446:
12445:
12442:
12441:
12344:Spectacled bear (
12302:
12278:
12277:
12274:
12273:
12270:
12269:
12118:
12105:
12104:
12024:Crab-eating fox (
12015:Short-eared dog (
11963:
11948:
11935:
11934:
11817:
11759:Otocyon megalotis
11703:
11670:
11669:
11666:
11665:
11662:
11661:
11658:
11657:
11654:
11653:
11522:Yellow mongoose (
11373:
11372:
11260:Banded mongoose (
11229:Liberiictus kuhni
11183:
11164:
11163:
11160:
11159:
11047:
11046:
10973:
10942:
10941:
10938:
10937:
10934:
10933:
10878:
10839:
10805:
10804:
10801:
10800:
10797:
10796:
10769:
10768:
10765:
10764:
10761:
10760:
10757:
10756:
10709:Servaline genet (
10628:Bourlon's genet (
10603:
10564:
10540:
10539:
10422:
10421:
10394:P. hermaphroditus
10246:
10245:
10242:
10241:
10238:
10237:
10214:
10213:
10210:
10209:
10206:
10205:
10202:
10201:
10198:
10197:
10159:African wildcat (
10116:
10115:
10086:Flat-headed cat (
10026:
10025:
9810:
9809:
9747:
9746:
9661:
9660:
9657:
9656:
9585:Clouded leopard (
9544:
9519:Spotted linsang (
9503:
9468:Nandinia binotata
9350:
9349:
9321:C. l. pambasileus
9258:C. l. labradorius
9238:Hudson Bay wolf (
9222:C. l. columbianus
8983:Death of a Legend
8951:978-0-226-25514-9
8926:978-0-226-51696-7
8901:978-0-8166-1026-6
8876:978-1-86189-879-1
8851:978-0-7432-4936-2
8841:Of Wolves and Men
8830:978-1-886106-81-9
8809:978-1-55059-332-7
8790:978-1-59921-069-8
8604:978-82-426-1292-2
8189:on June 21, 2010.
8043:, pp. 64â70.
8019:, pp. 38â45.
7791:10.7717/peerj.982
7065:978-0-285-62411-5
6850:, pp. 46â49.
6811:, pp. 42â46.
6696:, pp. 39â41.
6574:(4314): 320â321.
6552:, pp. 19â26.
6438:, pp. 12â13.
6313:, pp. 77â85.
6204:978-2-8317-0442-5
5681:Mammalian Biology
5518:10.1111/mam.12067
5481:978-1-4102-0249-9
5215:, pp. 19â20.
5170:978-0-7607-1969-5
5140:978-0-19-104572-1
5061:"Ch23: Gray wolf
4866:Mammalian Species
4812:(6516): 557â564.
4431:10.1111/mec.15329
4411:Molecular Ecology
4348:(7918): 313â320.
4303:10.1111/mec.16127
4289:(24): 6687â6700.
4283:Molecular Ecology
4188:(11): 1515â1519.
4111:978-3-030-04752-8
3856:978-0-8018-8221-0
3837:"Order Carnivora"
3833:Wozencraft, W. C.
3776:Harper, Douglas.
3716:, pp. 74â75.
3653:Harper, Douglas.
3509:Canus lupus dingo
3296:Attacks on humans
3253:domestic reindeer
3031:European folklore
2959:Chinese astronomy
2932:Romulus and Remus
2904:Romulus and Remus
2797:Arabian Peninsula
2770:Iberian peninsula
2766:Italian Peninsula
2732:II) and the 1992
2544:biting the throat
2542:, wolves kill by
2532:white-tailed deer
2313:
2295:
2176:intestinal mucosa
2135:Sarcoptes scabiei
2038:incubation period
2036:. In wolves, the
2022:canine parvovirus
1803:white-tailed deer
1673:Wolf distribution
1536:calcaneal tendons
1526:and long, coarse
1363:genetic admixture
1273:genetic admixture
1160:from the earlier
1062:mitochondrial DNA
1054:taxonomic synonym
957:published in his
951:
950:
941:
940:
926:
925:
914:
913:
855:
854:
823:
822:
791:
790:
759:
758:
727:
726:
695:
694:
314:
313:
147:
130:
93:Canis lupus lupus
27:Species of canine
16:(Redirected from
15912:
15825:
15824:
15812:
15811:
15799:
15798:
15786:
15785:
15773:
15772:
15760:
15759:
15747:
15746:
15734:
15733:
15724:
15723:
15711:
15710:
15708:NBNSYS0000005184
15698:
15697:
15685:
15684:
15672:
15671:
15659:
15658:
15646:
15645:
15633:
15632:
15620:
15619:
15607:
15606:
15594:
15593:
15581:
15580:
15571:
15570:
15558:
15557:
15545:
15544:
15532:
15531:
15519:
15518:
15506:
15505:
15496:
15495:
15483:
15482:
15470:
15469:
15460:
15459:
15447:
15446:
15434:
15433:
15432:
15419:
15418:
15417:
15387:
15386:
15363:ABC Islands bear
15293:Macropod hybrids
15270:Hybrid orangutan
15030:Dingoâdog hybrid
14881:
14874:
14867:
14858:
14857:
14790:Malayan weasel (
14709:Steppe polecat (
14682:Sichuan weasel (
14674:
14673:
14669:
14609:
14608:
14546:L. perspicillata
14511:Eurasian otter (
14388:
14387:
14383:
14382:
14378:
14289:
14288:
14284:
14283:
14279:
14258:Greater grison (
14241:
14240:
14236:
14210:
14209:
14201:
14200:
14151:Nilgiri marten (
14124:Pacific marten (
14107:
14106:
14102:
14028:
14027:
14011:M. subaurantiaca
13956:
13955:
13954:(ferret-badgers)
13951:
13845:
13844:
13840:
13839:
13835:
13787:
13786:
13776:
13775:
13763:
13762:
13657:
13656:
13652:
13651:
13647:
13595:
13594:
13590:
13545:
13544:
13540:
13481:
13480:
13423:
13422:
13421:(spotted skunks)
13418:
13382:
13381:
13377:
13290:
13289:
13285:
13276:
13275:
13271:
13270:
13262:
13233:
13232:
13220:
13219:
13177:Crabeater seal (
13152:
13151:
13147:
13146:
13142:
13108:
13107:
13106:(elephant seals)
13103:
13082:N. schauinslandi
13049:
13048:
13044:
13043:
13039:
13032:
13031:
13027:
13026:
13022:
12897:
12896:
12862:
12861:
12857:
12856:
12852:
12845:
12844:
12840:
12839:
12833:
12794:Brown fur seal (
12769:A. galapagoensis
12656:Z. californianus
12623:
12622:
12590:
12589:
12585:
12584:
12570:
12540:
12539:
12532:
12524:
12523:
12468:
12467:
12463:
12457:
12456:
12372:Melursus ursinus
12363:
12362:
12306:
12305:
12301:
12298:
12291:
12290:
12258:Ethiopian wolf (
12122:
12121:
12117:
12114:
11967:
11966:
11962:
11959:
11952:
11951:
11947:
11944:
11833:Blanford's fox (
11816:
11809:
11750:
11749:
11708:
11707:
11702:
11699:
11692:
11691:
11684:
11676:
11675:
11478:Ruddy mongoose (
11460:Javan mongoose (
11391:Marsh mongoose (
11384:
11383:
11363:C. platycephalus
11196:
11195:
11187:
11186:
11182:
11179:
11177:
11058:
11057:
11002:Malagasy civet (
10986:
10985:
10977:
10976:
10972:
10967:
10960:
10959:
10955:
10882:
10881:
10877:
10874:
10843:
10842:
10838:
10835:
10828:
10827:
10824:
10823:
10811:
10810:
10775:
10774:
10602:
10599:
10563:
10560:
10551:
10550:
10461:
10460:
10457:
10446:
10445:
10441:
10428:
10427:
10334:
10333:
10276:
10275:
10265:
10264:
10252:
10251:
10046:Otocolobus manul
10037:
10036:
9959:
9958:
9835:Geoffroy's cat (
9758:
9757:
9688:
9687:
9679:
9678:
9674:
9562:
9561:
9553:
9552:
9549:
9548:
9543:
9540:
9533:
9532:
9510:Banded linsang (
9502:
9499:
9491:
9490:
9480:
9479:
9459:
9458:
9448:
9447:
9440:
9432:
9431:
9377:
9370:
9363:
9354:
9353:
9310:Greenland wolf (
9240:C. l. hudsonicus
9160:C. l. familiaris
9140:Himalayan wolf (
9131:Mongolian wolf (
9124:C. l. campestris
9023:
9016:
9009:
9000:
8999:
8955:
8930:
8905:
8880:
8855:
8834:
8813:
8794:
8760:
8759:
8757:
8755:
8749:
8743:. Archived from
8738:
8729:
8723:
8717:
8708:
8702:
8696:
8695:
8693:
8691:
8673:
8667:
8661:
8655:
8654:
8642:
8636:
8630:
8624:
8623:
8621:
8619:
8613:
8596:
8585:
8568:
8567:
8550:(4): 1195â1208.
8538:
8532:
8531:
8509:
8503:
8502:
8500:
8476:
8470:
8469:
8467:
8465:
8460:on July 17, 2019
8459:
8452:
8443:
8437:
8436:
8428:
8422:
8421:
8394:Canis familiaris
8385:
8364:
8363:
8345:
8339:
8333:
8327:
8321:
8315:
8309:
8303:
8302:
8300:
8298:
8283:
8277:
8271:
8265:
8259:
8253:
8247:
8234:
8233:
8197:
8191:
8190:
8188:
8173:
8160:
8154:
8153:
8151:
8149:
8121:
8115:
8109:
8103:
8102:
8086:
8080:
8074:
8068:
8062:
8056:
8050:
8044:
8038:
8032:
8026:
8020:
8014:
8008:
8002:
7996:
7990:
7984:
7978:
7972:
7966:
7960:
7954:
7948:
7942:
7936:
7935:
7917:
7906:
7900:
7894:
7888:
7882:
7876:
7867:
7866:
7848:
7820:
7814:
7813:
7803:
7793:
7769:
7763:
7757:
7751:
7750:
7740:
7708:
7702:
7696:
7690:
7689:
7679:
7669:
7637:
7631:
7630:
7594:
7588:
7582:
7576:
7575:
7573:
7571:
7555:
7549:
7543:
7537:
7531:
7525:
7524:
7522:
7520:
7504:
7495:
7494:
7492:
7490:
7484:
7477:
7469:
7463:
7462:
7460:
7436:
7430:
7429:
7411:
7405:
7395:
7389:
7383:
7377:
7365:
7359:
7358:
7356:
7354:
7339:
7330:
7329:
7327:
7325:
7310:
7304:
7303:
7287:
7277:
7271:
7270:
7268:
7266:
7255:
7249:
7248:
7246:
7244:
7233:
7227:
7226:
7224:
7222:
7216:
7209:
7201:
7195:
7194:
7192:
7190:
7170:
7164:
7163:
7161:
7159:
7139:
7133:
7127:
7118:
7112:
7106:
7100:
7094:
7088:
7082:
7076:
7070:
7069:
7047:
7041:
7040:, p. 82â89.
7035:
7029:
7023:
7017:
7011:
7002:
7001:
6999:
6975:
6969:
6963:
6957:
6951:
6945:
6939:
6933:
6932:
6896:
6890:
6884:
6878:
6877:
6857:
6851:
6845:
6839:
6833:
6827:
6821:
6812:
6806:
6797:
6791:
6785:
6779:
6773:
6767:
6761:
6755:
6744:
6743:
6715:
6709:
6703:
6697:
6691:
6685:
6679:
6673:
6667:
6661:
6655:
6649:
6648:
6626:
6620:
6619:
6617:
6615:
6559:
6553:
6547:
6536:
6535:
6495:
6486:
6480:
6474:
6473:
6445:
6439:
6433:
6427:
6421:
6415:
6414:
6404:
6394:
6362:
6356:
6350:
6341:
6335:
6329:
6323:
6314:
6308:
6302:
6296:
6287:
6281:
6275:
6274:
6246:
6240:
6239:
6227:
6221:
6220:
6218:
6216:
6188:
6182:
6180:
6164:
6158:
6157:
6155:
6153:
6124:
6118:
6117:
6107:
6097:
6073:
6067:
6066:
6064:
6028:
6022:
6016:
6010:
6009:
5985:
5979:
5973:
5964:
5958:
5952:
5946:
5940:
5934:
5928:
5922:
5916:
5915:
5913:
5911:
5905:
5898:
5889:
5883:
5882:
5880:
5878:
5872:
5865:
5856:
5850:
5844:
5835:
5834:
5822:
5816:
5810:
5804:
5798:
5792:
5791:
5769:
5763:
5762:
5742:
5736:
5735:
5733:
5731:
5719:
5713:
5712:
5676:
5670:
5664:
5651:
5645:
5636:
5630:
5624:
5623:
5621:
5619:
5574:
5568:
5562:
5556:
5550:
5541:
5540:
5530:
5520:
5495:
5486:
5485:
5465:
5459:
5458:
5430:
5424:
5423:
5395:
5389:
5388:
5370:
5346:
5340:
5339:
5329:
5288:
5282:
5276:
5270:
5264:
5258:
5252:
5246:
5245:
5225:
5216:
5210:
5204:
5198:
5192:
5186:
5175:
5174:
5155:Macdonald, D. W.
5151:
5145:
5144:
5122:
5113:
5107:
5098:
5092:
5083:
5082:
5056:
5035:
5029:
5023:
5022:
5002:
4996:
4990:
4984:
4978:
4939:
4933:
4927:
4921:
4912:
4906:
4900:
4899:
4897:
4895:
4877:
4857:
4848:
4847:
4829:
4796:
4790:
4789:
4787:
4763:
4757:
4756:
4728:
4719:
4718:
4689:
4683:
4682:
4672:
4632:
4626:
4625:
4615:
4605:
4596:(11): e1007745.
4580:
4571:
4570:
4560:
4519:
4510:
4509:
4491:
4467:
4461:
4460:
4450:
4417:(9): 1596â1610.
4401:
4392:
4391:
4381:
4332:
4323:
4322:
4277:
4271:
4270:
4260:
4227:
4216:
4215:
4205:
4173:
4167:
4166:
4156:
4146:
4122:
4116:
4115:
4089:
4078:
4077:
4067:
4039:
4033:
4032:
4022:
3974:
3965:
3959:
3953:
3952:
3950:
3948:
3939:
3926:
3920:
3919:
3909:
3899:
3875:
3869:
3868:
3829:
3823:
3822:
3794:
3788:
3787:
3773:
3767:
3766:
3756:
3739:(#16): 2158â65.
3723:
3717:
3711:
3705:
3704:
3690:
3684:
3683:
3671:
3665:
3664:
3650:
3644:
3643:
3631:
3620:
3619:
3617:
3615:
3601:
3595:
3594:
3592:
3590:
3585:
3555:
3529:
3518:
3512:
3501:
3481:wolf-dog hybrids
3399:Volgograd Oblast
3156:Nikolay Nekrasov
3122:Charles Perrault
2984:Tantric Buddhism
2908:founding of Rome
2743:
2739:
2731:
2642:
2614:
2483:
2482:
2478:
2475:
2469:
2468:
2464:
2461:
2449:gestation period
2315:
2314:
2297:
2296:
2277:
2228:consisting of a
2199:Social structure
2018:canine distemper
2002:Wolf with mange
1971:Russian Far East
1616:
1615:
1611:
1608:
1602:
1601:
1597:
1594:
1588:
1587:
1583:
1580:
1574:
1573:
1569:
1566:
1560:
1559:
1555:
1552:
1518:, northern India
1492:
1491:
1487:
1484:
1478:
1477:
1473:
1470:
1464:
1463:
1459:
1456:
1450:
1449:
1445:
1442:
1346:ghost population
1307:Wolfâdog hybrids
1188:Late Pleistocene
1112:Canis familiaris
1013:Pleistocene wolf
987:Canis familiaris
905:
885:
872:
871:
846:
840:African wild dog
814:
782:
750:
718:
686:
666:
653:
652:
646:
645:
639:
638:
632:
631:
625:
624:
618:
617:
611:
610:
604:
603:
594:
593:
567:Germanic peoples
560:
559:
544:
543:
531:
436:consisting of a
434:nuclear families
432:. It travels in
329:
305:
295:
265:
162:
161:
141:
124:
119:
118:
99:in Bardu, Norway
82:
71:
64:
57:Temporal range:
51:
50:
21:
15920:
15919:
15915:
15914:
15913:
15911:
15910:
15909:
15870:Holarctic fauna
15850:Mammals of Asia
15835:
15834:
15833:
15828:
15820:
15815:
15807:
15802:
15794:
15789:
15781:
15776:
15768:
15763:
15755:
15750:
15742:
15739:Observation.org
15737:
15729:
15727:
15719:
15714:
15706:
15701:
15693:
15688:
15680:
15675:
15667:
15662:
15654:
15649:
15641:
15636:
15628:
15623:
15615:
15610:
15602:
15597:
15589:
15584:
15576:
15574:
15566:
15561:
15553:
15548:
15540:
15535:
15527:
15522:
15514:
15509:
15501:
15499:
15491:
15486:
15478:
15473:
15465:
15463:
15455:
15450:
15442:
15437:
15428:
15427:
15422:
15413:
15412:
15407:
15394:
15384:
15379:
15342:
15317:
15297:
15279:
15246:
15213:Panthera hybrid
15154:
15107:
15103:Elephant hybrid
15089:
15075:Clymene dolphin
15061:
15001:
14973:
14918:
14917:
14890:
14885:
14855:
14834:
14810:M. richardsonii
14754:European mink (
14671:
14670:
14668:
14658:
14650:American mink (
14623:Amazon weasel (
14594:
14552:
14528:
14495:
14478:
14454:
14403:
14380:
14379:
14377:
14361:
14328:
14311:
14281:
14280:
14278:
14266:
14249:Lesser grison (
14238:
14237:
14235:
14225:
14186:
14104:
14103:
14101:
14091:
14067:
14043:
14017:
13953:
13952:
13950:
13936:
13885:
13837:
13836:
13834:
13826:
13802:
13783:
13770:
13749:
13721:
13688:
13649:
13648:
13646:
13638:
13592:
13591:
13589:
13579:
13542:
13541:
13539:
13529:
13496:
13466:
13433:S. angustifrons
13420:
13419:
13417:
13407:
13380:(stink badgers)
13379:
13378:
13376:
13366:
13358:Striped skunk (
13333:
13327:C. semistriatus
13287:
13286:
13284:
13264:
13263:
13261:
13253:
13247:Ailurus fulgens
13227:
13194:
13144:
13143:
13141:
13133:
13105:
13104:
13102:
13088:
13064:
13041:
13040:
13038:
13024:
13023:
13021:
13005:
12963:
12930:
12886:
12854:
12853:
12851:
12837:
12835:
12834:
12832:
12820:
12733:
12716:
12692:
12671:
12638:
12612:
12579:
12574:
12572:
12571:
12569:
12565:
12560:
12534:
12530:
12502:
12482:
12465:
12464:
12438:
12387:
12352:
12328:
12299:
12297:
12287:
12266:
12195:Golden jackal (
12179:
12153:
12115:
12113:
12101:
12032:
12026:Cerdocyon thous
12006:
11982:
11960:
11958:
11945:
11943:
11931:
11896:RĂŒppell's fox (
11810:
11808:
11798:
11783:N. procyonoides
11765:
11757:Bat-eared fox (
11739:
11704:
11700:
11686:
11682:
11650:
11635:H. pulverulenta
11590:
11548:
11504:
11471:U. semitorquata
11426:U. auropunctata
11408:
11369:
11318:
11285:
11268:
11235:
11218:
11180:
11178:
11173:
11172:
11156:
11123:
11106:
11073:
11067:Galidia elegans
11043:
11010:
10970:
10968:
10966:
10956:
10951:
10930:
10922:Spotted hyena (
10906:
10889:Striped hyena (
10875:
10873:
10865:
10836:
10834:
10818:
10793:
10753:
10691:Aquatic genet (
10682:Pardine genet (
10619:Angolan genet (
10600:
10598:
10588:
10573:P. richardsonii
10561:
10559:
10536:
10519:Malayan civet (
10485:
10477:African civet (
10453:
10452:
10442:
10437:
10436:
10432:
10418:
10376:
10323:
10308:Diplogale hosei
10272:
10259:
10234:
10194:
10112:
10052:
10032:
10022:
9998:
9981:
9954:
9948:
9922:Eurasian lynx (
9897:
9806:
9773:
9753:
9743:
9710:
9683:
9675:
9670:
9653:
9593:
9545:
9541:
9527:
9500:
9487:
9474:
9455:
9442:
9438:
9426:
9388:
9381:
9351:
9346:
9340:C. l. tundrarum
9285:C. l. mackenzii
9256:Labrador wolf (
9249:C. l. irremotus
9231:C. l. crassodon
9194:
9188:
9171:Eurasian wolf (
9096:
9090:
9036:
9027:
8989:Cry of the Wild
8962:
8952:
8927:
8902:
8877:
8852:
8831:
8810:
8791:
8769:
8764:
8763:
8753:
8751:
8747:
8736:
8730:
8726:
8718:
8711:
8703:
8699:
8689:
8687:
8674:
8670:
8662:
8658:
8643:
8639:
8631:
8627:
8617:
8615:
8611:
8605:
8594:
8586:
8571:
8539:
8535:
8528:
8510:
8506:
8477:
8473:
8463:
8461:
8457:
8450:
8444:
8440:
8435:(174): 259â264.
8429:
8425:
8386:
8367:
8360:
8346:
8342:
8334:
8330:
8322:
8318:
8310:
8306:
8296:
8294:
8284:
8280:
8272:
8268:
8260:
8256:
8248:
8237:
8198:
8194:
8186:
8171:
8161:
8157:
8147:
8145:
8122:
8118:
8110:
8106:
8087:
8083:
8075:
8071:
8063:
8059:
8051:
8047:
8039:
8035:
8027:
8023:
8015:
8011:
8003:
7999:
7991:
7987:
7979:
7975:
7967:
7963:
7955:
7951:
7943:
7939:
7932:
7918:
7909:
7901:
7897:
7889:
7885:
7877:
7870:
7821:
7817:
7770:
7766:
7758:
7754:
7709:
7705:
7697:
7693:
7652:(4): e0215019.
7638:
7634:
7595:
7591:
7583:
7579:
7569:
7567:
7556:
7552:
7544:
7540:
7532:
7528:
7518:
7516:
7505:
7498:
7488:
7486:
7482:
7475:
7471:
7470:
7466:
7437:
7433:
7412:
7408:
7396:
7392:
7384:
7380:
7366:
7362:
7352:
7350:
7341:
7340:
7333:
7323:
7321:
7320:. March 5, 2024
7312:
7311:
7307:
7300:
7278:
7274:
7264:
7262:
7257:
7256:
7252:
7242:
7240:
7235:
7234:
7230:
7220:
7218:
7214:
7207:
7203:
7202:
7198:
7188:
7186:
7171:
7167:
7157:
7155:
7140:
7136:
7128:
7121:
7113:
7109:
7101:
7097:
7089:
7085:
7077:
7073:
7066:
7048:
7044:
7036:
7032:
7024:
7020:
7016:, pp. 1â3.
7012:
7005:
6976:
6972:
6964:
6960:
6952:
6948:
6940:
6936:
6913:10.2307/1382426
6897:
6893:
6885:
6881:
6874:
6858:
6854:
6846:
6842:
6834:
6830:
6822:
6815:
6807:
6800:
6792:
6788:
6780:
6776:
6768:
6764:
6756:
6747:
6716:
6712:
6704:
6700:
6692:
6688:
6680:
6676:
6668:
6664:
6656:
6652:
6645:
6627:
6623:
6613:
6611:
6560:
6556:
6548:
6539:
6496:
6489:
6481:
6477:
6470:
6446:
6442:
6434:
6430:
6426:, pp. 1â2.
6422:
6418:
6377:(9). e0137378.
6363:
6359:
6351:
6344:
6336:
6332:
6324:
6317:
6309:
6305:
6297:
6290:
6282:
6278:
6247:
6243:
6228:
6224:
6214:
6212:
6205:
6189:
6185:
6165:
6161:
6151:
6149:
6142:
6125:
6121:
6074:
6070:
6029:
6025:
6017:
6013:
6006:
5986:
5982:
5974:
5967:
5959:
5955:
5947:
5943:
5935:
5931:
5923:
5919:
5909:
5907:
5903:
5896:
5890:
5886:
5876:
5874:
5870:
5863:
5857:
5853:
5845:
5838:
5823:
5819:
5811:
5807:
5799:
5795:
5788:
5770:
5766:
5759:
5743:
5739:
5729:
5727:
5720:
5716:
5677:
5673:
5665:
5654:
5646:
5639:
5631:
5627:
5617:
5615:
5575:
5571:
5563:
5559:
5551:
5544:
5496:
5489:
5482:
5466:
5462:
5431:
5427:
5396:
5392:
5347:
5343:
5289:
5285:
5277:
5273:
5265:
5261:
5253:
5249:
5226:
5219:
5211:
5207:
5199:
5195:
5187:
5178:
5171:
5152:
5148:
5141:
5123:
5116:
5108:
5101:
5093:
5086:
5079:
5057:
5038:
5030:
5026:
5003:
4999:
4991:
4987:
4979:
4942:
4934:
4930:
4922:
4915:
4907:
4903:
4893:
4891:
4875:10.2307/3503924
4858:
4851:
4797:
4793:
4764:
4760:
4729:
4722:
4690:
4686:
4633:
4629:
4581:
4574:
4529:Current Biology
4520:
4513:
4468:
4464:
4402:
4395:
4333:
4326:
4278:
4274:
4237:Genome Research
4228:
4219:
4182:Current Biology
4174:
4170:
4137:(1). e1004016.
4123:
4119:
4112:
4090:
4081:
4040:
4036:
3975:
3968:
3960:
3956:
3946:
3944:
3937:
3927:
3923:
3890:(1): e1004093.
3876:
3872:
3857:
3830:
3826:
3819:
3795:
3791:
3774:
3770:
3733:Current Biology
3724:
3720:
3712:
3708:
3691:
3687:
3672:
3668:
3651:
3647:
3632:
3623:
3613:
3611:
3603:
3602:
3598:
3588:
3586:
3556:
3543:
3538:
3533:
3532:
3519:
3515:
3502:
3498:
3493:
3477:
3471:
3391:
3385:
3377:Main articles:
3375:
3308:
3300:Main articles:
3298:
3277:
3232:
3220:
3196:The Jungle Book
3191:Rudyard Kipling
3130:sexual predator
3057:
3051:
2940:Norse mythology
2895:Capitoline Wolf
2886:
2880:
2874:
2869:
2864:
2850:has 10,000 and
2741:
2737:
2729:
2711:
2651:
2640:
2612:
2594:
2588:
2496:
2490:
2480:
2476:
2473:
2471:
2466:
2462:
2459:
2457:
2379:
2373:
2330:
2329:
2321:
2319:
2318:
2317:
2316:
2309:
2306:
2300:
2299:
2298:
2291:
2288:
2281:
2275:
2270:
2264:
2243:sexual maturity
2207:
2201:
2196:
2188:
1996:
1990:
1902:
1736:
1675:
1669:
1664:
1650:Rocky Mountains
1613:
1609:
1606:
1604:
1599:
1595:
1592:
1590:
1585:
1581:
1578:
1576:
1571:
1567:
1564:
1562:
1557:
1553:
1550:
1548:
1544:tissue-freezing
1508:
1495:Bergmann's rule
1489:
1485:
1482:
1480:
1475:
1471:
1468:
1466:
1461:
1457:
1454:
1452:
1447:
1443:
1440:
1438:
1395:
1338:Sinai Peninsula
1300:
1294:
1281:Tibetan Mastiff
1174:C. mosbachensis
1163:C. mosbachensis
1136:
1130:
1124:
1087:) based on the
1075:of "wolf", the
1015:
1009:
1003:
960:Systema Naturae
942:
927:
915:
856:
824:
792:
760:
728:
696:
592:
511:
505:
489:hunter-gatherer
287:
276:
267:
261:
248:
156:
148:
131:
120:
116:
109:
100:
73:
72:
65:
59:
55:
47:
39:
28:
23:
22:
15:
12:
11:
5:
15918:
15908:
15907:
15902:
15897:
15892:
15887:
15882:
15877:
15872:
15867:
15862:
15857:
15852:
15847:
15830:
15829:
15827:
15826:
15813:
15800:
15787:
15774:
15761:
15748:
15735:
15725:
15712:
15699:
15686:
15673:
15660:
15647:
15634:
15621:
15608:
15595:
15582:
15572:
15559:
15550:Fauna Europaea
15546:
15533:
15520:
15507:
15497:
15484:
15471:
15461:
15448:
15435:
15420:
15404:
15402:
15396:
15395:
15381:
15380:
15378:
15377:
15376:
15375:
15370:
15365:
15354:
15352:
15344:
15343:
15341:
15340:
15335:
15329:
15327:
15319:
15318:
15316:
15315:
15309:
15307:
15299:
15298:
15296:
15295:
15289:
15287:
15281:
15280:
15278:
15277:
15272:
15267:
15262:
15256:
15254:
15248:
15247:
15245:
15244:
15243:
15242:
15241:
15240:
15235:
15230:
15225:
15220:
15210:
15205:
15200:
15195:
15190:
15185:
15180:
15175:
15164:
15162:
15156:
15155:
15153:
15152:
15151:
15150:
15145:
15140:
15135:
15130:
15119:
15117:
15109:
15108:
15106:
15105:
15099:
15097:
15091:
15090:
15088:
15087:
15082:
15077:
15071:
15069:
15063:
15062:
15060:
15059:
15058:
15057:
15052:
15047:
15042:
15040:Eastern coyote
15037:
15032:
15027:
15022:
15011:
15009:
15003:
15002:
15000:
14999:
14994:
14989:
14983:
14981:
14975:
14974:
14972:
14971:
14970:
14969:
14964:
14959:
14954:
14949:
14944:
14939:
14937:American Breed
14928:
14926:
14920:
14919:
14916:
14915:
14909:
14903:
14896:
14895:
14892:
14891:
14888:Mammal hybrids
14884:
14883:
14876:
14869:
14861:
14852:
14851:
14848:
14847:
14844:
14843:
14840:
14839:
14836:
14835:
14833:
14832:
14828:M. strigidorsa
14823:
14814:
14805:
14796:
14787:
14781:Least weasel (
14778:
14769:
14760:
14751:
14742:
14733:
14727:Haida ermine (
14724:
14715:
14711:M. eversmannii
14706:
14697:
14688:
14678:
14676:
14660:
14659:
14657:
14656:
14647:
14638:
14629:
14619:
14617:
14606:
14600:
14599:
14596:
14595:
14593:
14592:
14583:
14574:
14564:
14562:
14554:
14553:
14551:
14550:
14540:
14538:
14530:
14529:
14527:
14526:
14517:
14507:
14505:
14497:
14496:
14494:
14493:
14483:
14480:
14479:
14477:
14476:
14466:
14464:
14456:
14455:
14453:
14452:
14443:
14439:L. longicaudis
14434:
14428:Marine otter (
14425:
14415:
14413:
14405:
14404:
14402:
14401:
14391:
14385:
14371:
14370:
14367:
14366:
14363:
14362:
14360:
14359:
14350:
14340:
14338:
14330:
14329:
14327:
14326:
14316:
14313:
14312:
14310:
14309:
14299:
14297:
14286:
14272:
14271:
14268:
14267:
14265:
14264:
14255:
14245:
14243:
14227:
14226:
14224:
14223:
14213:
14207:
14198:
14192:
14191:
14188:
14187:
14185:
14184:
14175:
14166:
14157:
14148:
14142:Beech marten (
14139:
14130:
14121:
14111:
14109:
14093:
14092:
14090:
14089:
14079:
14077:
14069:
14068:
14066:
14065:
14055:
14053:
14045:
14044:
14042:
14041:
14031:
14025:
14019:
14018:
14016:
14015:
14006:
13997:
13988:
13979:
13970:
13960:
13958:
13942:
13941:
13938:
13937:
13935:
13934:
13925:
13919:Asian badger (
13916:
13907:
13897:
13895:
13887:
13886:
13884:
13883:
13874:
13865:
13861:A. albogularis
13855:
13853:
13842:
13828:
13827:
13825:
13824:
13818:Honey badger (
13814:
13812:
13804:
13803:
13801:
13800:
13790:
13784:
13779:
13772:
13771:
13759:
13758:
13755:
13754:
13751:
13750:
13748:
13747:
13746:
13745:
13739:
13737:
13731:
13730:
13727:
13726:
13723:
13722:
13720:
13719:
13710:
13700:
13698:
13690:
13689:
13687:
13686:
13677:
13667:
13665:
13654:
13640:
13639:
13637:
13636:
13627:
13618:
13609:
13599:
13597:
13581:
13580:
13578:
13577:
13568:
13559:
13555:P. cancrivorus
13549:
13547:
13531:
13530:
13528:
13527:
13523:B. sumichrasti
13518:
13508:
13506:
13498:
13497:
13495:
13494:
13484:
13478:
13472:
13471:
13468:
13467:
13465:
13464:
13455:
13446:
13437:
13427:
13425:
13409:
13408:
13406:
13405:
13396:
13386:
13384:
13368:
13367:
13365:
13364:
13355:
13349:Hooded skunk (
13345:
13343:
13335:
13334:
13332:
13331:
13322:
13313:
13304:
13294:
13292:
13273:
13255:
13254:
13252:
13251:
13241:
13239:
13229:
13228:
13216:
13215:
13212:
13211:
13208:
13207:
13204:
13203:
13200:
13199:
13196:
13195:
13193:
13192:
13183:
13174:
13168:Weddell seal (
13165:
13159:Leopard seal (
13155:
13149:
13135:
13134:
13132:
13131:
13122:
13112:
13110:
13094:
13093:
13090:
13089:
13087:
13086:
13076:
13074:
13066:
13065:
13063:
13062:
13052:
13046:
13029:
13015:
13014:
13011:
13010:
13007:
13006:
13004:
13003:
12994:
12985:
12979:Caspian seal (
12975:
12973:
12965:
12964:
12962:
12961:
12952:
12946:Spotted seal (
12942:
12940:
12932:
12931:
12929:
12928:
12919:
12910:
12900:
12894:
12888:
12887:
12885:
12884:
12875:
12869:Bearded seal (
12865:
12859:
12842:
12838:or true seals)
12836:(earless seals
12826:
12825:
12822:
12821:
12819:
12818:
12809:
12800:
12791:
12782:
12773:
12764:
12755:
12745:
12743:
12735:
12734:
12732:
12731:
12721:
12718:
12717:
12715:
12714:
12704:
12702:
12694:
12693:
12691:
12690:
12680:
12677:
12676:
12673:
12672:
12670:
12669:
12660:
12650:
12648:
12640:
12639:
12637:
12636:
12626:
12620:
12614:
12613:
12611:
12610:
12600:
12598:
12587:
12562:
12561:
12559:
12558:
12548:
12546:
12536:
12535:
12520:
12519:
12516:
12515:
12512:
12511:
12508:
12507:
12504:
12503:
12501:
12500:
12499:
12498:
12492:
12490:
12484:
12483:
12481:
12480:
12479:
12478:
12472:
12470:
12454:
12448:
12447:
12444:
12443:
12440:
12439:
12437:
12436:
12427:
12418:
12409:
12399:
12397:
12389:
12388:
12386:
12385:
12376:
12366:
12360:
12354:
12353:
12351:
12350:
12340:
12338:
12330:
12329:
12327:
12326:
12322:A. melanoleuca
12316:
12314:
12303:
12288:
12283:
12280:
12279:
12276:
12275:
12272:
12271:
12268:
12267:
12265:
12264:
12255:
12246:
12240:Eastern wolf (
12237:
12228:
12222:African wolf (
12219:
12210:
12204:Domestic dog (
12201:
12191:
12189:
12181:
12180:
12178:
12177:
12168:
12158:
12155:
12154:
12152:
12151:
12142:
12132:
12130:
12119:
12107:
12106:
12103:
12102:
12100:
12099:
12090:
12084:Sechuran fox (
12081:
12077:L. gymnocercus
12072:
12063:
12057:Darwin's fox (
12054:
12044:
12042:
12034:
12033:
12031:
12030:
12021:
12011:
12008:
12007:
12005:
12004:
11994:
11992:
11984:
11983:
11981:
11980:
11970:
11964:
11949:
11937:
11936:
11933:
11932:
11930:
11929:
11920:
11911:
11902:
11893:
11884:
11875:
11866:
11857:
11848:
11839:
11830:
11826:V. bengalensis
11820:
11818:
11800:
11799:
11797:
11796:
11787:
11777:
11775:
11767:
11766:
11764:
11763:
11753:
11747:
11741:
11740:
11738:
11737:
11728:
11718:
11716:
11705:
11695:
11688:
11687:
11672:
11671:
11668:
11667:
11664:
11663:
11660:
11659:
11656:
11655:
11652:
11651:
11649:
11648:
11639:
11630:
11621:
11612:
11602:
11600:
11592:
11591:
11589:
11588:
11579:
11570:
11566:B. crassicauda
11560:
11558:
11550:
11549:
11547:
11546:
11537:
11528:
11519:
11509:
11506:
11505:
11503:
11502:
11493:
11484:
11475:
11466:
11457:
11448:
11439:
11430:
11420:
11418:
11410:
11409:
11407:
11406:
11397:
11387:
11381:
11375:
11374:
11371:
11370:
11368:
11367:
11358:
11349:
11340:
11330:
11328:
11320:
11319:
11317:
11316:
11307:
11297:
11295:
11287:
11286:
11284:
11283:
11273:
11270:
11269:
11267:
11266:
11257:
11247:
11245:
11237:
11236:
11234:
11233:
11223:
11220:
11219:
11217:
11216:
11206:
11204:
11193:
11184:
11166:
11165:
11162:
11161:
11158:
11157:
11155:
11154:
11145:
11135:
11133:
11125:
11124:
11122:
11121:
11111:
11108:
11107:
11105:
11104:
11100:G. grandidieri
11095:
11085:
11083:
11075:
11074:
11072:
11071:
11061:
11055:
11049:
11048:
11045:
11044:
11042:
11041:
11032:
11022:
11020:
11012:
11011:
11009:
11008:
10999:
10989:
10983:
10974:
10957:
10947:
10944:
10943:
10940:
10939:
10936:
10935:
10932:
10931:
10929:
10928:
10918:
10916:
10908:
10907:
10905:
10904:
10895:
10885:
10879:
10867:
10866:
10864:
10863:
10853:
10851:
10840:
10820:
10819:
10807:
10806:
10803:
10802:
10799:
10798:
10795:
10794:
10792:
10791:
10790:
10789:
10783:
10781:
10771:
10770:
10767:
10766:
10763:
10762:
10759:
10758:
10755:
10754:
10752:
10751:
10742:
10733:
10724:
10715:
10706:
10697:
10688:
10679:
10670:
10664:Letaba genet (
10661:
10652:
10646:Common genet (
10643:
10634:
10625:
10616:
10606:
10604:
10590:
10589:
10587:
10586:
10577:
10567:
10565:
10548:
10542:
10541:
10538:
10537:
10535:
10534:
10525:
10516:
10507:
10497:
10495:
10487:
10486:
10484:
10483:
10474:
10464:
10458:
10443:
10431:
10424:
10423:
10420:
10419:
10417:
10416:
10412:P. zeylonensis
10407:
10398:
10388:
10386:
10378:
10377:
10375:
10374:
10370:Paguma larvata
10365:
10356:
10347:
10337:
10331:
10325:
10324:
10322:
10321:
10312:
10303:
10294:
10284:
10282:
10273:
10268:
10261:
10260:
10248:
10247:
10244:
10243:
10240:
10239:
10236:
10235:
10233:
10232:
10231:
10230:
10224:
10222:
10216:
10215:
10212:
10211:
10208:
10207:
10204:
10203:
10200:
10199:
10196:
10195:
10193:
10192:
10183:
10174:
10165:
10156:
10147:
10141:Domestic cat (
10138:
10128:
10126:
10118:
10117:
10114:
10113:
10111:
10110:
10101:
10097:P. rubiginosus
10092:
10083:
10074:
10070:P. bengalensis
10064:
10062:
10054:
10053:
10051:
10050:
10044:Pallas's cat (
10040:
10034:
10028:
10027:
10024:
10023:
10021:
10020:
10010:
10008:
10000:
9999:
9997:
9996:
9986:
9983:
9982:
9980:
9979:
9969:
9967:
9956:
9950:
9949:
9947:
9946:
9937:
9931:Iberian lynx (
9928:
9919:
9909:
9907:
9899:
9898:
9896:
9895:
9886:
9877:
9868:
9859:
9850:
9841:
9832:
9822:
9820:
9812:
9811:
9808:
9807:
9805:
9804:
9795:
9785:
9783:
9775:
9774:
9772:
9771:
9761:
9755:
9749:
9748:
9745:
9744:
9742:
9741:
9732:
9722:
9720:
9712:
9711:
9709:
9708:
9698:
9696:
9685:
9676:
9666:
9663:
9662:
9659:
9658:
9655:
9654:
9652:
9651:
9642:
9633:
9624:
9615:
9609:Snow leopard (
9605:
9603:
9595:
9594:
9592:
9591:
9582:
9572:
9570:
9559:
9546:
9536:
9529:
9528:
9526:
9525:
9516:
9506:
9504:
9488:
9483:
9476:
9475:
9473:
9472:
9462:
9456:
9451:
9444:
9443:
9428:
9427:
9425:
9424:
9422:Laurasiatheria
9418:
9412:
9406:
9400:
9393:
9390:
9389:
9380:
9379:
9372:
9365:
9357:
9348:
9347:
9345:
9344:
9335:
9325:
9316:
9307:
9298:
9294:C. l. manningi
9289:
9280:
9274:Eastern wolf (
9271:
9262:
9253:
9244:
9235:
9226:
9217:
9211:Mexican wolf (
9208:
9198:
9196:
9190:
9189:
9187:
9186:
9182:C. l. pallipes
9177:
9168:
9158:Domestic dog (
9155:
9146:
9137:
9128:
9119:
9113:Arabian wolf (
9110:
9100:
9098:
9092:
9091:
9089:
9088:
9080:
9072:
9066:
9060:
9054:
9048:
9041:
9038:
9037:
9026:
9025:
9018:
9011:
9003:
8997:
8996:
8978:
8977:(January 2009)
8968:
8961:
8960:External links
8958:
8957:
8956:
8950:
8931:
8925:
8906:
8900:
8881:
8875:
8867:Reaktion Books
8856:
8850:
8835:
8829:
8814:
8808:
8795:
8789:
8779:(3 ed.).
8768:
8765:
8762:
8761:
8724:
8709:
8707:, p. 108.
8697:
8668:
8666:, p. 267.
8656:
8637:
8625:
8603:
8569:
8533:
8526:
8504:
8471:
8438:
8423:
8365:
8359:978-0199663217
8358:
8340:
8328:
8326:, p. 306.
8316:
8314:, p. 307.
8304:
8278:
8276:, p. 173.
8266:
8264:, p. 309.
8254:
8252:, p. 305.
8235:
8208:(3): 809â814.
8192:
8167:Never Cry Wolf
8155:
8116:
8114:, p. 294.
8104:
8081:
8079:, p. 162.
8069:
8057:
8055:, p. 263.
8045:
8033:
8031:, p. 259.
8021:
8009:
8007:, p. 123.
7997:
7985:
7973:
7971:, p. 110.
7961:
7959:, p. 133.
7949:
7937:
7931:978-0826419132
7930:
7907:
7905:, p. 128.
7895:
7893:, p. 210.
7883:
7881:, p. 292.
7868:
7839:(11): 765â70.
7815:
7764:
7762:, p. 232.
7752:
7723:(3): 115â116.
7703:
7701:, p. 328.
7691:
7632:
7589:
7587:, p. 327.
7577:
7550:
7548:, p. 231.
7538:
7526:
7496:
7464:
7451:(4): 173â180.
7431:
7406:
7390:
7378:
7360:
7331:
7305:
7298:
7272:
7250:
7228:
7196:
7165:
7134:
7119:
7117:, p. 230.
7107:
7095:
7083:
7081:, p. 185.
7071:
7064:
7056:Souvenir Press
7042:
7030:
7018:
7003:
6990:(3): 595â605.
6970:
6968:, p. 240.
6958:
6956:, p. 233.
6946:
6934:
6907:(4): 879â889.
6891:
6879:
6873:978-0894642029
6872:
6852:
6840:
6828:
6813:
6798:
6786:
6784:, p. 175.
6774:
6762:
6760:, p. 248.
6745:
6726:(6): 628â637.
6710:
6698:
6686:
6674:
6662:
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6643:
6621:
6554:
6537:
6487:
6475:
6468:
6440:
6428:
6416:
6357:
6342:
6340:, p. 164.
6330:
6315:
6303:
6288:
6276:
6241:
6222:
6203:
6183:
6159:
6140:
6119:
6068:
6033:"Gray Wolves,
6023:
6011:
6004:
5980:
5978:, p. 265.
5965:
5963:, p. 266.
5953:
5941:
5929:
5927:, p. 269.
5917:
5884:
5851:
5836:
5817:
5815:, p. 180.
5805:
5803:, p. 109.
5793:
5787:978-0785837381
5786:
5764:
5758:978-0520254732
5757:
5737:
5714:
5671:
5652:
5650:, p. 201.
5637:
5635:, p. 172.
5625:
5569:
5557:
5555:, p. 107.
5542:
5511:(4): 255â269.
5487:
5480:
5460:
5441:(4): 333â347.
5425:
5412:10.1086/284670
5406:(5): 755â760.
5390:
5361:(6): 435â436.
5341:
5283:
5271:
5269:, p. 168.
5259:
5247:
5236:(3): 821â830.
5217:
5205:
5193:
5176:
5169:
5146:
5139:
5114:
5112:, p. 174.
5099:
5084:
5077:
5036:
5034:, p. 112.
5024:
5013:(3): 249â270.
4997:
4985:
4940:
4928:
4926:, p. 166.
4913:
4901:
4849:
4791:
4778:(2): 405â417.
4758:
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4684:
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4393:
4324:
4272:
4243:(2): 163â173.
4217:
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3596:
3540:
3539:
3537:
3534:
3531:
3530:
3513:
3495:
3494:
3492:
3489:
3473:Main article:
3470:
3467:
3374:
3371:
3297:
3294:
3276:
3273:
3231:
3228:
3219:
3216:
3211:Hollywood film
3206:Never Cry Wolf
3180:. The musical
3172:and Chekhov's
3050:
3047:
2948:Geri and Freki
2924:Ancient Romans
2916:Ancient Greeks
2876:Main article:
2873:
2870:
2868:
2865:
2863:
2860:
2840:Andhra Pradesh
2828:Madhya Pradesh
2722:European Union
2710:
2707:
2663:reintroduction
2650:
2647:
2637:are listed in
2587:
2584:
2548:carotid artery
2492:Main article:
2489:
2486:
2372:
2369:
2320:
2307:
2302:
2301:
2289:
2286:Wolves howling
2284:
2283:
2282:
2273:
2272:
2271:
2266:Main article:
2263:
2260:
2226:nuclear family
2220:The wolf is a
2200:
2197:
2187:
2184:
2030:papillomatosis
2010:Viral diseases
1992:Main article:
1989:
1986:
1982:striped hyenas
1967:Siberian tiger
1953:, such as the
1901:
1898:
1735:
1732:
1671:Main article:
1668:
1665:
1663:
1660:
1624:Wolves in the
1507:
1504:
1499:Arabian wolves
1394:
1391:
1370:Italian wolves
1334:golden jackals
1330:African wolves
1296:Main article:
1293:
1290:
1228:are part of a
1222:Himalayan wolf
1126:Main article:
1123:
1120:
1066:subspecies of
1005:Main article:
1002:
999:
949:
948:
944:
943:
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789:
788:
785:
784:
772:
769:
768:
762:
761:
757:
756:
753:
752:
744:Ethiopian wolf
740:
737:
736:
730:
729:
725:
724:
721:
720:
708:
705:
704:
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598:
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588:
524:Proto-Germanic
504:
501:
454:hooved mammals
356:subspecies of
312:
311:
307:
306:
298:
297:
290:Subspecies of
283:
282:
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15216:
15215:
15214:
15211:
15209:
15208:Serengeti cat
15206:
15204:
15201:
15199:
15196:
15194:
15191:
15189:
15186:
15184:
15181:
15179:
15176:
15174:
15171:
15170:
15169:
15166:
15165:
15163:
15161:
15157:
15149:
15146:
15144:
15141:
15139:
15136:
15134:
15131:
15129:
15126:
15125:
15124:
15121:
15120:
15118:
15116:
15115:
15110:
15104:
15101:
15100:
15098:
15096:
15092:
15086:
15083:
15081:
15078:
15076:
15073:
15072:
15070:
15068:
15064:
15056:
15053:
15051:
15048:
15046:
15043:
15041:
15038:
15036:
15033:
15031:
15028:
15026:
15023:
15021:
15018:
15017:
15016:
15013:
15012:
15010:
15008:
15004:
14998:
14995:
14993:
14990:
14988:
14985:
14984:
14982:
14980:
14976:
14968:
14965:
14963:
14960:
14958:
14955:
14953:
14950:
14948:
14945:
14943:
14940:
14938:
14935:
14934:
14933:
14930:
14929:
14927:
14925:
14921:
14914:
14910:
14908:
14904:
14902:
14898:
14897:
14893:
14889:
14882:
14877:
14875:
14870:
14868:
14863:
14862:
14859:
14831:
14829:
14824:
14822:
14820:
14815:
14813:
14811:
14806:
14804:
14802:
14797:
14795:
14793:
14788:
14786:
14784:
14779:
14777:
14775:
14770:
14768:
14766:
14765:M. lutreolina
14761:
14759:
14757:
14752:
14750:
14748:
14743:
14741:
14739:
14734:
14732:
14730:
14725:
14723:
14721:
14716:
14714:
14712:
14707:
14705:
14703:
14698:
14696:
14694:
14689:
14687:
14685:
14680:
14679:
14677:
14667:
14666:
14661:
14655:
14653:
14648:
14646:
14644:
14639:
14637:
14635:
14630:
14628:
14626:
14621:
14620:
14618:
14616:
14615:
14610:
14607:
14605:
14601:
14591:
14589:
14584:
14582:
14580:
14575:
14573:
14571:
14566:
14565:
14563:
14561:
14560:
14555:
14549:
14547:
14542:
14541:
14539:
14537:
14536:
14531:
14525:
14523:
14518:
14516:
14514:
14509:
14508:
14506:
14504:
14503:
14498:
14492:
14490:
14485:
14484:
14481:
14475:
14473:
14468:
14467:
14465:
14463:
14462:
14457:
14451:
14449:
14444:
14442:
14440:
14435:
14433:
14431:
14426:
14424:
14422:
14421:L. canadensis
14417:
14416:
14414:
14412:
14411:
14406:
14400:
14398:
14395:Giant otter (
14393:
14392:
14389:
14386:
14376:
14372:
14358:
14356:
14351:
14349:
14347:
14342:
14341:
14339:
14337:
14336:
14331:
14325:
14323:
14318:
14317:
14314:
14308:
14306:
14301:
14300:
14298:
14296:
14295:
14290:
14287:
14277:
14273:
14263:
14261:
14256:
14254:
14252:
14247:
14246:
14244:
14234:
14233:
14228:
14222:
14220:
14215:
14214:
14211:
14208:
14206:
14202:
14199:
14197:
14193:
14183:
14181:
14176:
14174:
14172:
14167:
14165:
14163:
14158:
14156:
14154:
14153:M. gwatkinsii
14149:
14147:
14145:
14140:
14138:
14136:
14131:
14129:
14127:
14122:
14120:
14118:
14113:
14112:
14110:
14100:
14099:
14094:
14088:
14086:
14081:
14080:
14078:
14076:
14075:
14070:
14064:
14062:
14057:
14056:
14054:
14052:
14051:
14046:
14040:
14038:
14033:
14032:
14029:
14026:
14024:
14020:
14014:
14012:
14007:
14005:
14003:
13998:
13996:
13994:
13993:M. orientalis
13989:
13987:
13985:
13980:
13978:
13976:
13971:
13969:
13967:
13962:
13961:
13959:
13949:
13948:
13943:
13933:
13931:
13926:
13924:
13922:
13917:
13915:
13913:
13908:
13906:
13904:
13899:
13898:
13896:
13894:
13893:
13888:
13882:
13880:
13875:
13873:
13871:
13866:
13864:
13862:
13857:
13856:
13854:
13852:
13851:
13846:
13843:
13833:
13829:
13823:
13821:
13816:
13815:
13813:
13811:
13810:
13805:
13799:
13797:
13796:Taxidea taxus
13792:
13791:
13788:
13785:
13782:
13777:
13773:
13769:
13764:
13760:
13743:
13742:
13741:
13740:
13738:
13736:
13732:
13718:
13716:
13711:
13709:
13707:
13706:N. meridensis
13702:
13701:
13699:
13697:
13696:
13691:
13685:
13683:
13678:
13676:
13674:
13669:
13668:
13666:
13664:
13663:
13658:
13655:
13645:
13641:
13635:
13633:
13628:
13626:
13624:
13619:
13617:
13615:
13610:
13608:
13606:
13601:
13600:
13598:
13588:
13587:
13582:
13576:
13574:
13569:
13567:
13565:
13560:
13558:
13556:
13551:
13550:
13548:
13538:
13537:
13532:
13526:
13524:
13519:
13517:
13515:
13510:
13509:
13507:
13505:
13504:
13499:
13493:
13491:
13486:
13485:
13482:
13479:
13477:
13473:
13463:
13461:
13456:
13454:
13452:
13447:
13445:
13443:
13438:
13436:
13434:
13429:
13428:
13426:
13416:
13415:
13410:
13404:
13402:
13397:
13395:
13393:
13392:M. javanensis
13388:
13387:
13385:
13375:
13374:
13369:
13363:
13361:
13356:
13354:
13352:
13347:
13346:
13344:
13342:
13341:
13336:
13330:
13328:
13323:
13321:
13319:
13318:C. leuconotus
13314:
13312:
13310:
13309:C. humboldtii
13305:
13303:
13301:
13296:
13295:
13293:
13283:
13282:
13277:
13274:
13268:
13260:
13256:
13250:
13248:
13243:
13242:
13240:
13238:
13234:
13230:
13226:
13221:
13217:
13191:
13189:
13184:
13182:
13180:
13175:
13173:
13171:
13166:
13164:
13162:
13157:
13156:
13153:
13150:
13140:
13136:
13130:
13128:
13123:
13121:
13119:
13114:
13113:
13111:
13101:
13100:
13095:
13085:
13083:
13078:
13077:
13075:
13073:
13072:
13067:
13061:
13059:
13054:
13053:
13050:
13047:
13037:
13033:
13030:
13020:
13016:
13002:
13000:
12997:Baikal seal (
12995:
12993:
12991:
12988:Ringed seal (
12986:
12984:
12982:
12977:
12976:
12974:
12972:
12971:
12966:
12960:
12958:
12955:Harbor seal (
12953:
12951:
12949:
12944:
12943:
12941:
12939:
12938:
12933:
12927:
12925:
12920:
12918:
12916:
12913:Ribbon seal (
12911:
12909:
12907:
12902:
12901:
12898:
12895:
12893:
12889:
12883:
12881:
12878:Hooded seal (
12876:
12874:
12872:
12867:
12866:
12863:
12860:
12850:
12846:
12843:
12831:
12827:
12817:
12815:
12814:A. tropicalis
12810:
12808:
12806:
12801:
12799:
12797:
12792:
12790:
12788:
12783:
12781:
12779:
12774:
12772:
12770:
12765:
12763:
12761:
12756:
12754:
12752:
12747:
12746:
12744:
12742:
12741:
12740:Arctocephalus
12736:
12730:
12728:
12723:
12722:
12719:
12713:
12711:
12706:
12705:
12703:
12701:
12700:
12695:
12689:
12687:
12682:
12681:
12678:
12668:
12666:
12665:Z. wollebaeki
12661:
12659:
12657:
12652:
12651:
12649:
12647:
12646:
12641:
12635:
12633:
12628:
12627:
12624:
12621:
12619:
12615:
12609:
12607:
12602:
12601:
12599:
12597:
12596:
12591:
12588:
12582:
12577:
12573:(eared seals:
12568:
12563:
12557:
12555:
12550:
12549:
12547:
12545:
12541:
12537:
12533:
12525:
12521:
12496:
12495:
12494:
12493:
12491:
12489:
12485:
12476:
12475:
12474:
12473:
12471:
12462:
12458:
12455:
12453:
12449:
12435:
12433:
12428:
12426:
12424:
12419:
12417:
12415:
12414:U. thibetanus
12410:
12408:
12406:
12405:U. americanus
12401:
12400:
12398:
12396:
12395:
12390:
12384:
12382:
12377:
12375:
12373:
12368:
12367:
12364:
12361:
12359:
12355:
12349:
12347:
12342:
12341:
12339:
12337:
12336:
12331:
12325:
12323:
12320:Giant panda (
12318:
12317:
12315:
12313:
12312:
12307:
12304:
12296:
12292:
12289:
12286:
12281:
12263:
12261:
12256:
12254:
12252:
12247:
12245:
12243:
12238:
12236:
12234:
12229:
12227:
12225:
12220:
12218:
12216:
12211:
12209:
12207:
12206:C. familiaris
12202:
12200:
12198:
12193:
12192:
12190:
12188:
12187:
12182:
12176:
12174:
12169:
12167:
12165:
12164:Lycaon pictus
12160:
12159:
12156:
12150:
12148:
12143:
12141:
12139:
12134:
12133:
12131:
12129:
12128:
12123:
12120:
12112:
12108:
12098:
12096:
12091:
12089:
12087:
12082:
12080:
12078:
12073:
12071:
12069:
12064:
12062:
12060:
12055:
12053:
12051:
12046:
12045:
12043:
12041:
12040:
12035:
12029:
12027:
12022:
12020:
12018:
12013:
12012:
12009:
12003:
12001:
11996:
11995:
11993:
11991:
11990:
11985:
11979:
11977:
11972:
11971:
11968:
11965:
11957:
11953:
11950:
11942:
11938:
11928:
11926:
11921:
11919:
11917:
11912:
11910:
11908:
11903:
11901:
11899:
11894:
11892:
11890:
11885:
11883:
11881:
11876:
11874:
11872:
11867:
11865:
11863:
11860:Tibetan fox (
11858:
11856:
11854:
11849:
11847:
11845:
11840:
11838:
11836:
11831:
11829:
11827:
11822:
11821:
11819:
11814:
11807:
11806:
11801:
11795:
11793:
11792:N. viverrinus
11788:
11786:
11784:
11779:
11778:
11776:
11774:
11773:
11768:
11762:
11760:
11755:
11754:
11751:
11748:
11746:
11742:
11736:
11734:
11733:U. littoralis
11729:
11727:
11725:
11720:
11719:
11717:
11715:
11714:
11709:
11706:
11698:
11693:
11689:
11685:
11677:
11673:
11647:
11645:
11640:
11638:
11636:
11631:
11629:
11627:
11622:
11620:
11618:
11613:
11611:
11609:
11608:H. flavescens
11604:
11603:
11601:
11599:
11598:
11593:
11587:
11585:
11580:
11578:
11576:
11571:
11569:
11567:
11562:
11561:
11559:
11557:
11556:
11551:
11545:
11543:
11538:
11536:
11534:
11529:
11527:
11525:
11520:
11518:
11516:
11511:
11510:
11507:
11501:
11499:
11498:U. vitticolla
11494:
11492:
11490:
11485:
11483:
11481:
11476:
11474:
11472:
11467:
11465:
11463:
11458:
11456:
11454:
11449:
11447:
11445:
11440:
11438:
11436:
11431:
11429:
11427:
11422:
11421:
11419:
11417:
11416:
11411:
11405:
11403:
11402:Xenogale naso
11398:
11396:
11394:
11389:
11388:
11385:
11382:
11380:
11376:
11366:
11364:
11359:
11357:
11355:
11350:
11348:
11346:
11341:
11339:
11337:
11332:
11331:
11329:
11327:
11326:
11321:
11315:
11313:
11308:
11306:
11304:
11299:
11298:
11296:
11294:
11293:
11288:
11282:
11280:
11275:
11274:
11271:
11265:
11263:
11258:
11256:
11254:
11249:
11248:
11246:
11244:
11243:
11238:
11232:
11230:
11225:
11224:
11221:
11215:
11213:
11208:
11207:
11205:
11203:
11202:
11197:
11194:
11192:
11188:
11185:
11176:
11175:sensu stricto
11171:
11167:
11153:
11151:
11146:
11144:
11142:
11137:
11136:
11134:
11132:
11131:
11126:
11120:
11118:
11113:
11112:
11109:
11103:
11101:
11096:
11094:
11092:
11087:
11086:
11084:
11082:
11081:
11076:
11070:
11068:
11063:
11062:
11059:
11056:
11054:
11050:
11040:
11038:
11033:
11031:
11029:
11024:
11023:
11021:
11019:
11018:
11013:
11007:
11005:
11004:Fossa fossana
11000:
10998:
10996:
10991:
10990:
10987:
10984:
10982:
10978:
10975:
10965:
10961:
10958:
10954:
10950:
10945:
10927:
10925:
10920:
10919:
10917:
10915:
10914:
10909:
10903:
10901:
10898:Brown hyena (
10896:
10894:
10892:
10891:Hyaena hyaena
10887:
10886:
10883:
10880:
10872:
10868:
10862:
10860:
10855:
10854:
10852:
10850:
10849:
10844:
10841:
10833:
10829:
10825:
10821:
10817:
10812:
10808:
10787:
10786:
10785:
10784:
10782:
10780:
10776:
10772:
10750:
10748:
10743:
10741:
10739:
10734:
10732:
10730:
10725:
10723:
10721:
10718:Hausa genet (
10716:
10714:
10712:
10707:
10705:
10703:
10698:
10696:
10694:
10689:
10687:
10685:
10680:
10678:
10676:
10671:
10669:
10667:
10662:
10660:
10658:
10653:
10651:
10649:
10644:
10642:
10640:
10635:
10633:
10631:
10626:
10624:
10622:
10621:G. angolensis
10617:
10615:
10613:
10612:G. abyssinica
10608:
10607:
10605:
10597:
10596:
10591:
10585:
10583:
10578:
10576:
10574:
10569:
10568:
10566:
10558:
10557:
10552:
10549:
10547:
10543:
10533:
10531:
10526:
10524:
10522:
10521:V. tangalunga
10517:
10515:
10513:
10508:
10506:
10504:
10499:
10498:
10496:
10494:
10493:
10488:
10482:
10480:
10475:
10473:
10471:
10466:
10465:
10462:
10459:
10456:
10455:sensu stricto
10451:
10447:
10444:
10440:
10435:
10429:
10425:
10415:
10413:
10408:
10406:
10404:
10399:
10397:
10395:
10390:
10389:
10387:
10385:
10384:
10379:
10373:
10371:
10366:
10364:
10362:
10357:
10355:
10353:
10348:
10346:
10344:
10339:
10338:
10335:
10332:
10330:
10329:Paradoxurinae
10326:
10320:
10318:
10313:
10311:
10309:
10304:
10302:
10300:
10297:Otter civet (
10295:
10293:
10291:
10286:
10285:
10283:
10281:
10277:
10274:
10271:
10266:
10262:
10258:
10253:
10249:
10228:
10227:
10226:
10225:
10223:
10221:
10217:
10191:
10189:
10188:F. silvestris
10184:
10182:
10180:
10175:
10173:
10171:
10166:
10164:
10162:
10157:
10155:
10153:
10148:
10146:
10144:
10139:
10137:
10135:
10130:
10129:
10127:
10125:
10124:
10119:
10109:
10107:
10106:P. viverrinus
10104:Fishing cat (
10102:
10100:
10098:
10093:
10091:
10089:
10084:
10082:
10080:
10079:P. javanensis
10075:
10073:
10071:
10068:Leopard cat (
10066:
10065:
10063:
10061:
10060:
10055:
10049:
10047:
10042:
10041:
10038:
10035:
10029:
10019:
10017:
10012:
10011:
10009:
10007:
10006:
10001:
9995:
9993:
9988:
9987:
9984:
9978:
9976:
9971:
9970:
9968:
9966:
9965:
9960:
9957:
9951:
9945:
9943:
9938:
9936:
9934:
9929:
9927:
9925:
9920:
9918:
9916:
9915:L. canadensis
9913:Canada lynx (
9911:
9910:
9908:
9906:
9905:
9900:
9894:
9892:
9887:
9885:
9883:
9878:
9876:
9874:
9869:
9867:
9865:
9860:
9858:
9856:
9851:
9849:
9847:
9842:
9840:
9838:
9833:
9831:
9829:
9824:
9823:
9821:
9819:
9818:
9813:
9803:
9801:
9796:
9794:
9792:
9787:
9786:
9784:
9782:
9781:
9776:
9770:
9768:
9763:
9762:
9759:
9756:
9750:
9740:
9738:
9737:C. temminckii
9733:
9731:
9729:
9724:
9723:
9721:
9719:
9718:
9713:
9707:
9705:
9702:Marbled cat (
9700:
9699:
9697:
9695:
9694:
9689:
9686:
9680:
9677:
9673:
9672:sensu stricto
9669:
9664:
9650:
9648:
9643:
9641:
9639:
9634:
9632:
9630:
9625:
9623:
9621:
9616:
9614:
9612:
9607:
9606:
9604:
9602:
9601:
9596:
9590:
9588:
9583:
9581:
9579:
9574:
9573:
9571:
9569:
9568:
9563:
9560:
9558:
9554:
9550:
9547:
9539:
9534:
9530:
9524:
9522:
9521:P. pardicolor
9517:
9515:
9513:
9508:
9507:
9505:
9498:
9497:
9492:
9489:
9486:
9481:
9477:
9471:
9469:
9464:
9463:
9460:
9457:
9454:
9449:
9445:
9441:
9433:
9429:
9423:
9419:
9417:
9413:
9411:
9407:
9405:
9401:
9399:
9395:
9394:
9391:
9386:
9378:
9373:
9371:
9366:
9364:
9359:
9358:
9355:
9343:
9341:
9336:
9333:
9331:
9326:
9324:
9322:
9317:
9315:
9313:
9308:
9306:
9304:
9299:
9297:
9295:
9290:
9288:
9286:
9281:
9279:
9277:
9272:
9270:
9268:
9263:
9261:
9259:
9254:
9252:
9250:
9245:
9243:
9241:
9236:
9234:
9232:
9227:
9225:
9223:
9218:
9216:
9214:
9213:C. l. baileyi
9209:
9207:
9205:
9202:Arctic wolf (
9200:
9199:
9197:
9191:
9185:
9183:
9180:Indian wolf (
9178:
9176:
9174:
9169:
9167:
9165:
9164:C. familiaris
9161:
9156:
9154:
9152:
9147:
9145:
9143:
9138:
9136:
9134:
9129:
9127:
9125:
9122:Steppe wolf (
9120:
9118:
9116:
9111:
9109:
9107:
9104:Tundra wolf (
9102:
9101:
9099:
9093:
9087:
9086:
9081:
9079:
9078:
9073:
9071:
9067:
9065:
9061:
9059:
9055:
9053:
9049:
9047:
9043:
9042:
9039:
9035:
9032:
9024:
9019:
9017:
9012:
9010:
9005:
9004:
9001:
8995:
8991:
8990:
8985:
8984:
8979:
8976:
8972:
8969:
8967:
8964:
8963:
8953:
8947:
8943:
8939:
8938:
8932:
8928:
8922:
8918:
8914:
8913:
8907:
8903:
8897:
8893:
8889:
8888:
8882:
8878:
8872:
8868:
8864:
8863:
8857:
8853:
8847:
8843:
8842:
8836:
8832:
8826:
8822:
8821:
8815:
8811:
8805:
8801:
8796:
8792:
8786:
8782:
8778:
8777:
8771:
8770:
8746:
8742:
8735:
8728:
8721:
8716:
8714:
8706:
8701:
8685:
8681:
8680:
8672:
8665:
8660:
8652:
8648:
8641:
8634:
8629:
8610:
8606:
8600:
8593:
8592:
8584:
8582:
8580:
8578:
8576:
8574:
8565:
8561:
8557:
8553:
8549:
8545:
8537:
8529:
8527:9780521425377
8523:
8519:
8515:
8508:
8499:
8494:
8490:
8486:
8482:
8475:
8456:
8449:
8442:
8434:
8427:
8419:
8415:
8411:
8407:
8403:
8399:
8395:
8391:
8384:
8382:
8380:
8378:
8376:
8374:
8372:
8370:
8361:
8355:
8351:
8344:
8338:, p. 45.
8337:
8332:
8325:
8320:
8313:
8308:
8293:
8289:
8282:
8275:
8270:
8263:
8258:
8251:
8246:
8244:
8242:
8240:
8231:
8227:
8223:
8219:
8215:
8211:
8207:
8203:
8196:
8185:
8181:
8177:
8170:
8168:
8159:
8143:
8139:
8135:
8131:
8129:
8120:
8113:
8108:
8100:
8096:
8092:
8085:
8078:
8073:
8066:
8061:
8054:
8049:
8042:
8037:
8030:
8025:
8018:
8013:
8006:
8001:
7995:, p. 50.
7994:
7989:
7983:, p. 47.
7982:
7977:
7970:
7965:
7958:
7953:
7947:, p. 78.
7946:
7941:
7933:
7927:
7923:
7916:
7914:
7912:
7904:
7899:
7892:
7887:
7880:
7875:
7873:
7864:
7860:
7856:
7852:
7847:
7842:
7838:
7834:
7830:
7828:
7819:
7811:
7807:
7802:
7797:
7792:
7787:
7783:
7779:
7775:
7768:
7761:
7756:
7748:
7744:
7739:
7734:
7730:
7726:
7722:
7718:
7714:
7707:
7700:
7695:
7687:
7683:
7678:
7673:
7668:
7663:
7659:
7655:
7651:
7647:
7643:
7636:
7628:
7624:
7620:
7616:
7612:
7608:
7604:
7600:
7593:
7586:
7581:
7565:
7561:
7554:
7547:
7542:
7535:
7530:
7514:
7510:
7503:
7501:
7481:
7474:
7468:
7459:
7454:
7450:
7446:
7442:
7435:
7427:
7423:
7419:
7418:
7410:
7403:
7399:
7394:
7387:
7382:
7375:
7374:
7369:
7364:
7348:
7344:
7338:
7336:
7319:
7315:
7309:
7301:
7295:
7291:
7286:
7285:
7276:
7260:
7254:
7238:
7232:
7213:
7206:
7200:
7184:
7180:
7176:
7169:
7153:
7149:
7145:
7138:
7131:
7126:
7124:
7116:
7111:
7104:
7099:
7093:, p. 58.
7092:
7087:
7080:
7075:
7067:
7061:
7057:
7053:
7046:
7039:
7034:
7027:
7022:
7015:
7010:
7008:
6998:
6993:
6989:
6985:
6981:
6974:
6967:
6962:
6955:
6950:
6943:
6938:
6930:
6926:
6922:
6918:
6914:
6910:
6906:
6902:
6895:
6888:
6883:
6875:
6869:
6865:
6864:
6856:
6849:
6844:
6838:, p. 47.
6837:
6832:
6825:
6820:
6818:
6810:
6805:
6803:
6795:
6790:
6783:
6778:
6771:
6766:
6759:
6754:
6752:
6750:
6741:
6737:
6733:
6729:
6725:
6721:
6714:
6708:, p. 90.
6707:
6702:
6695:
6690:
6684:, p. 38.
6683:
6678:
6672:, p. 59.
6671:
6666:
6659:
6654:
6646:
6644:0-8166-2959-5
6640:
6636:
6632:
6625:
6609:
6605:
6601:
6597:
6593:
6589:
6585:
6581:
6577:
6573:
6569:
6565:
6558:
6551:
6546:
6544:
6542:
6533:
6529:
6525:
6521:
6517:
6513:
6509:
6505:
6501:
6494:
6492:
6485:, p. 38.
6484:
6479:
6471:
6469:9780801825255
6465:
6461:
6457:
6456:
6451:
6444:
6437:
6432:
6425:
6420:
6412:
6408:
6403:
6398:
6393:
6388:
6384:
6380:
6376:
6372:
6368:
6361:
6354:
6349:
6347:
6339:
6334:
6327:
6322:
6320:
6312:
6307:
6300:
6295:
6293:
6285:
6280:
6272:
6268:
6264:
6260:
6256:
6252:
6245:
6237:
6233:
6226:
6210:
6206:
6200:
6196:
6195:
6187:
6178:
6174:
6170:
6163:
6147:
6143:
6141:1-55963-080-9
6137:
6133:
6132:
6123:
6115:
6111:
6106:
6101:
6096:
6091:
6087:
6083:
6079:
6072:
6063:
6058:
6054:
6050:
6046:
6044:
6040:
6039:Puma concolor
6036:
6027:
6020:
6015:
6007:
6005:0-226-77999-8
6001:
5997:
5993:
5992:
5984:
5977:
5972:
5970:
5962:
5957:
5950:
5945:
5938:
5933:
5926:
5921:
5902:
5895:
5888:
5869:
5862:
5855:
5848:
5843:
5841:
5832:
5828:
5827:Carbyn, L. N.
5821:
5814:
5809:
5802:
5797:
5789:
5783:
5779:
5775:
5768:
5760:
5754:
5750:
5749:
5741:
5725:
5718:
5710:
5706:
5702:
5698:
5694:
5690:
5686:
5682:
5675:
5668:
5663:
5661:
5659:
5657:
5649:
5644:
5642:
5634:
5629:
5614:
5610:
5606:
5602:
5598:
5594:
5590:
5586:
5585:
5580:
5573:
5566:
5561:
5554:
5549:
5547:
5538:
5534:
5529:
5524:
5519:
5514:
5510:
5506:
5505:Mammal Review
5502:
5494:
5492:
5483:
5477:
5473:
5472:
5464:
5456:
5452:
5448:
5444:
5440:
5436:
5429:
5421:
5417:
5413:
5409:
5405:
5401:
5394:
5386:
5382:
5378:
5374:
5369:
5364:
5360:
5356:
5352:
5345:
5337:
5333:
5328:
5323:
5319:
5315:
5311:
5307:
5303:
5299:
5295:
5287:
5281:, p. 22.
5280:
5275:
5268:
5263:
5256:
5251:
5243:
5239:
5235:
5231:
5224:
5222:
5214:
5209:
5203:, p. 18.
5202:
5197:
5191:, p. 19.
5190:
5185:
5183:
5181:
5172:
5166:
5162:
5161:
5156:
5150:
5142:
5136:
5132:
5128:
5121:
5119:
5111:
5106:
5104:
5097:, p. 23.
5096:
5091:
5089:
5080:
5078:0-8018-7416-5
5074:
5070:
5066:
5064:
5055:
5053:
5051:
5049:
5047:
5045:
5043:
5041:
5033:
5028:
5020:
5016:
5012:
5008:
5001:
4995:, p. 14.
4994:
4989:
4982:
4977:
4975:
4973:
4971:
4969:
4967:
4965:
4963:
4961:
4959:
4957:
4955:
4953:
4951:
4949:
4947:
4945:
4938:, p. 13.
4937:
4932:
4925:
4920:
4918:
4910:
4905:
4889:
4885:
4881:
4876:
4871:
4867:
4863:
4862:"Canis lupus"
4856:
4854:
4845:
4841:
4837:
4833:
4828:
4823:
4819:
4815:
4811:
4807:
4803:
4795:
4786:
4781:
4777:
4773:
4769:
4762:
4754:
4750:
4746:
4742:
4739:(3): 345â53.
4738:
4734:
4727:
4725:
4716:
4712:
4708:
4704:
4700:
4696:
4688:
4680:
4676:
4671:
4666:
4662:
4658:
4654:
4650:
4646:
4642:
4638:
4631:
4623:
4619:
4614:
4609:
4604:
4599:
4595:
4591:
4590:PLOS Genetics
4587:
4579:
4577:
4568:
4564:
4559:
4554:
4550:
4546:
4542:
4538:
4534:
4530:
4526:
4518:
4516:
4507:
4503:
4499:
4495:
4490:
4485:
4481:
4477:
4473:
4466:
4458:
4454:
4449:
4444:
4440:
4436:
4432:
4428:
4424:
4420:
4416:
4412:
4408:
4400:
4398:
4389:
4385:
4380:
4375:
4371:
4367:
4363:
4359:
4355:
4351:
4347:
4343:
4339:
4331:
4329:
4320:
4316:
4312:
4308:
4304:
4300:
4296:
4292:
4288:
4284:
4276:
4268:
4264:
4259:
4254:
4250:
4246:
4242:
4238:
4234:
4226:
4224:
4222:
4213:
4209:
4204:
4199:
4195:
4191:
4187:
4183:
4179:
4172:
4164:
4160:
4155:
4150:
4145:
4140:
4136:
4132:
4131:PLOS Genetics
4128:
4121:
4113:
4107:
4103:
4099:
4095:
4094:Paleogenomics
4088:
4086:
4084:
4075:
4071:
4066:
4061:
4057:
4056:10.1206/574.1
4053:
4049:
4045:
4038:
4030:
4026:
4021:
4016:
4012:
4008:
4004:
4000:
3996:
3992:
3988:
3984:
3980:
3973:
3971:
3963:
3958:
3943:
3936:
3932:
3925:
3917:
3913:
3908:
3903:
3898:
3893:
3889:
3885:
3884:PLOS Genetics
3881:
3874:
3866:
3862:
3858:
3852:
3848:
3847:
3842:
3841:Wilson, D. E.
3838:
3834:
3828:
3820:
3814:
3810:
3806:
3805:
3800:
3793:
3785:
3784:
3779:
3772:
3764:
3760:
3755:
3750:
3746:
3742:
3738:
3734:
3730:
3722:
3715:
3710:
3702:
3698:
3697:
3689:
3681:
3677:
3670:
3662:
3661:
3656:
3649:
3641:
3637:
3636:"Canis Lupus"
3630:
3628:
3626:
3610:
3606:
3600:
3584:
3579:
3575:
3571:
3570:
3565:
3563:
3554:
3552:
3550:
3548:
3546:
3541:
3527:
3523:
3517:
3510:
3506:
3500:
3496:
3488:
3486:
3482:
3476:
3466:
3464:
3463:Kyrgyz Tajgan
3460:
3455:
3452:
3448:
3444:
3439:
3434:
3432:
3428:
3424:
3420:
3416:
3412:
3407:
3400:
3395:
3390:
3384:
3380:
3370:
3368:
3364:
3360:
3359:Mediterranean
3354:
3352:
3348:
3344:
3340:
3336:
3332:
3328:
3323:
3316:
3312:
3307:
3303:
3293:
3289:
3285:
3282:
3272:
3270:
3266:
3262:
3258:
3254:
3248:
3241:
3236:
3227:
3225:
3215:
3212:
3208:
3207:
3202:
3198:
3197:
3192:
3187:
3185:
3184:
3179:
3175:
3171:
3170:
3169:War and Peace
3165:
3161:
3157:
3153:
3152:Anton Chekhov
3149:
3145:
3141:
3137:
3136:
3131:
3127:
3123:
3119:
3114:
3110:
3109:
3101:
3097:
3096:
3091:
3087:
3085:
3081:
3080:New Testament
3077:
3073:
3069:
3065:
3061:
3056:
3046:
3044:
3040:
3036:
3032:
3028:
3024:
3020:
3016:
3011:
3009:
3008:KwakwakaÊŒwakw
3005:
3001:
2997:
2992:
2987:
2985:
2981:
2977:
2973:
2969:
2964:
2960:
2955:
2953:
2949:
2945:
2941:
2937:
2933:
2929:
2925:
2921:
2917:
2909:
2905:
2901:
2897:
2896:
2890:
2885:
2879:
2859:
2857:
2856:Japanese wolf
2853:
2849:
2845:
2841:
2837:
2833:
2829:
2825:
2824:Uttar Pradesh
2821:
2817:
2813:
2809:
2805:
2800:
2798:
2790:
2785:
2781:
2779:
2774:
2771:
2767:
2763:
2762:Baltic states
2759:
2754:
2750:
2749:Great Britain
2745:
2735:
2727:
2723:
2715:
2706:
2704:
2699:
2695:
2691:
2687:
2683:
2678:
2676:
2671:
2670:First Nations
2664:
2660:
2655:
2649:North America
2646:
2644:
2636:
2632:
2628:
2624:
2620:
2616:
2608:
2604:
2603:Least Concern
2600:
2593:
2583:
2581:
2577:
2573:
2569:
2565:
2561:
2555:
2553:
2549:
2545:
2541:
2533:
2528:
2524:
2522:
2516:
2513:
2509:
2500:
2495:
2485:
2455:
2450:
2445:
2443:
2439:
2435:
2431:
2427:
2418:
2414:
2410:
2408:
2404:
2400:
2396:
2388:
2383:
2378:
2368:
2366:
2361:
2356:
2354:
2350:
2345:
2341:
2336:
2328:
2326:
2305:
2287:
2269:
2262:Communication
2259:
2257:
2252:
2247:
2244:
2239:
2236:
2231:
2227:
2223:
2222:social animal
2216:
2211:
2206:
2195:
2194:Dog behaviour
2190:
2183:
2181:
2177:
2173:
2169:
2165:
2161:
2157:
2153:
2149:
2145:
2141:
2137:
2136:
2131:
2127:
2123:
2119:
2115:
2110:
2109:, and death.
2108:
2104:
2100:
2096:
2092:
2088:
2084:
2080:
2076:
2072:
2071:leptospirosis
2068:
2064:
2060:
2056:
2052:
2048:
2044:
2039:
2035:
2031:
2027:
2023:
2019:
2015:
2011:
2005:
2000:
1995:
1985:
1983:
1978:
1976:
1972:
1968:
1964:
1960:
1956:
1955:Eurasian lynx
1952:
1947:
1945:
1940:
1935:
1927:
1923:
1921:
1917:
1913:
1908:
1897:
1895:
1891:
1886:
1884:
1880:
1876:
1872:
1868:
1864:
1860:
1856:
1852:
1848:
1844:
1840:
1836:
1832:
1828:
1824:
1820:
1815:
1812:
1808:
1804:
1800:
1796:
1792:
1788:
1784:
1780:
1774:
1772:
1768:
1764:
1760:
1753:
1749:
1746:hindquarter,
1745:
1740:
1731:
1729:
1725:
1721:
1717:
1713:
1709:
1705:
1704:British Isles
1701:
1692:
1688:
1684:
1679:
1674:
1659:
1656:
1651:
1647:
1646:Gloger's rule
1642:
1639:
1635:
1627:
1622:
1618:
1545:
1539:
1537:
1533:
1529:
1525:
1517:
1512:
1503:
1500:
1496:
1432:
1427:
1423:
1420:
1416:
1412:
1408:
1399:
1390:
1388:
1383:
1379:
1375:
1371:
1366:
1364:
1359:
1355:
1351:
1347:
1342:
1339:
1335:
1331:
1327:
1320:
1316:
1312:
1308:
1304:
1299:
1289:
1287:
1282:
1278:
1274:
1270:
1266:
1261:
1259:
1255:
1251:
1247:
1243:
1239:
1235:
1231:
1227:
1223:
1219:
1215:
1213:
1212:spotted hyena
1209:
1205:
1201:
1197:
1193:
1189:
1184:
1183:
1179:
1175:
1171:
1170:
1165:
1164:
1159:
1155:
1147:
1146:
1140:
1135:
1129:
1119:
1117:
1116:IUCN Red List
1113:
1110:
1106:
1102:
1098:
1094:
1090:
1089:type specimen
1086:
1082:
1081:Eurasian wolf
1078:
1074:
1070:
1069:
1063:
1059:
1055:
1051:
1047:
1043:
1042:
1037:
1033:
1030:listed under
1029:
1026:
1022:
1021:
1014:
1008:
998:
996:
992:
988:
984:
980:
976:
972:
971:
966:
962:
961:
956:
955:Carl Linnaeus
946:
945:
937:
936:
931:
930:
922:
921:2.62 mya
919:
918:
910:
909:
906:
904:
900:
894:
893:
890:
889:
886:
884:
880:
874:
873:
867:
866:
863:
862:3.06 mya
860:
859:
851:
850:
847:
845:
841:
835:
834:
831:
830:2.74 mya
828:
827:
819:
818:
815:
813:
809:
803:
802:
799:
798:1.92 mya
796:
795:
787:
786:
783:
781:
777:
776:Golden jackal
771:
770:
767:
766:1.62 mya
764:
763:
755:
754:
751:
749:
745:
739:
738:
735:
734:1.32 mya
732:
731:
723:
722:
719:
717:
713:
707:
706:
703:
702:1.10 mya
700:
699:
691:
690:
687:
685:
681:
675:
674:
671:
670:
667:
665:
661:
655:
654:
648:
647:
641:
640:
634:
633:
627:
626:
620:
619:
613:
612:
606:
605:
601:
600:
596:
595:
587:
584:
580:
576:
572:
568:
563:
561:
553:
549:
545:
542:
535:
530:
525:
521:
520:
516:
510:
500:
498:
494:
490:
486:
482:
478:
474:
473:Least Concern
469:
467:
463:
459:
455:
451:
447:
443:
439:
435:
431:
427:
423:
422:social nature
419:
416:
412:
408:
405:
400:
398:
394:
393:golden jackal
390:
386:
382:
381:
376:
372:
368:
364:
360:
359:
353:
352:North America
349:
345:
342:, is a large
341:
337:
333:
325:
324:
319:
308:
304:
299:
296:
294:
293:
284:
279:
275:
271:
266:
264:
258:
255:
254:Binomial name
251:
247:
246:
245:C. lupus
241:
238:
237:
234:
233:
229:
226:
225:
222:
219:
216:
215:
212:
209:
206:
205:
202:
199:
196:
195:
192:
189:
186:
185:
182:
179:
176:
175:
172:
169:
166:
165:
160:
155:
151:
145:
139:
134:
128:
123:
122:Least Concern
112:
108:
103:
98:
94:
90:
89:Eurasian wolf
85:
81:
76:
69:
62:
52:
49:
46:
41:
37:
33:
19:
15399:
15358:Ursid hybrid
15347:
15322:
15313:Iron Age pig
15302:
15285:Macropodinae
15203:Savannah cat
15168:Felid hybrid
15123:Equid hybrid
15112:
15095:Elephantidae
15015:Canid hybrid
14932:Bovid hybrid
14827:
14818:
14809:
14800:
14791:
14782:
14773:
14764:
14755:
14746:
14737:
14728:
14719:
14710:
14701:
14692:
14683:
14663:
14651:
14642:
14633:
14624:
14612:
14587:
14578:
14569:
14557:
14545:
14533:
14522:L. sumatrana
14521:
14512:
14500:
14488:
14471:
14459:
14447:
14438:
14429:
14420:
14408:
14396:
14354:
14345:
14333:
14321:
14305:V. peregusna
14304:
14292:
14259:
14250:
14230:
14218:
14205:Lyncodontini
14196:Ictonychinae
14180:M. zibellina
14179:
14170:
14161:
14152:
14143:
14135:M. flavigula
14134:
14125:
14117:M. americana
14116:
14096:
14084:
14072:
14060:
14048:
14037:Eira barbara
14036:
14010:
14002:M. personata
14001:
13992:
13983:
13974:
13965:
13945:
13929:
13920:
13912:M. canescens
13911:
13902:
13890:
13878:
13869:
13860:
13848:
13819:
13807:
13795:
13714:
13705:
13693:
13681:
13672:
13660:
13631:
13622:
13613:
13604:
13584:
13572:
13563:
13554:
13534:
13522:
13521:Cacomistle (
13513:
13501:
13490:Potos flavus
13489:
13459:
13450:
13441:
13432:
13412:
13400:
13391:
13371:
13359:
13350:
13338:
13326:
13317:
13308:
13299:
13279:
13246:
13187:
13178:
13169:
13160:
13126:
13117:
13097:
13081:
13069:
13057:
13042:(monk seals)
12998:
12989:
12980:
12968:
12956:
12947:
12935:
12923:
12914:
12905:
12879:
12870:
12813:
12805:A. townsendi
12804:
12795:
12787:A. philippii
12786:
12777:
12768:
12759:
12751:A. australis
12750:
12738:
12726:
12709:
12697:
12685:
12664:
12655:
12643:
12631:
12605:
12593:
12553:
12432:U. maritimus
12431:
12430:Polar bear (
12422:
12421:Brown bear (
12413:
12404:
12392:
12380:
12371:
12370:Sloth bear (
12345:
12333:
12321:
12309:
12259:
12250:
12241:
12232:
12230:
12223:
12214:
12205:
12196:
12184:
12173:Cuon alpinus
12172:
12163:
12147:L. mesomelas
12146:
12137:
12125:
12094:
12085:
12076:
12075:Pampas fox (
12067:
12058:
12049:
12037:
12025:
12016:
12000:S. venaticus
11999:
11987:
11975:
11974:Maned wolf (
11956:Cerdocyonina
11924:
11923:Fennec fox (
11915:
11906:
11898:V. rueppelli
11897:
11888:
11879:
11870:
11869:Arctic fox (
11862:V. ferrilata
11861:
11852:
11851:Corsac fox (
11843:
11834:
11825:
11824:Bengal fox (
11803:
11791:
11782:
11770:
11758:
11732:
11731:Island fox (
11723:
11711:
11644:H. sanguinea
11643:
11634:
11625:
11617:H. ichneumon
11616:
11607:
11595:
11583:
11574:
11565:
11553:
11541:
11532:
11523:
11514:
11497:
11488:
11479:
11470:
11461:
11452:
11444:U. edwardsii
11443:
11435:U. brachyura
11434:
11425:
11413:
11401:
11392:
11362:
11353:
11344:
11336:C. alexandri
11335:
11323:
11311:
11302:
11290:
11278:
11261:
11253:M. gambianus
11252:
11240:
11228:
11212:S. suricatta
11211:
11199:
11174:
11149:
11140:
11128:
11116:
11099:
11090:
11078:
11066:
11036:
11027:
11015:
11003:
10994:
10971:carnivorans)
10952:
10923:
10911:
10899:
10890:
10858:
10846:
10816:Herpestoidea
10779:Herpestoidea
10746:
10738:G. victoriae
10737:
10728:
10727:Cape genet (
10719:
10711:G. servalina
10710:
10701:
10700:King genet (
10693:G. piscivora
10692:
10683:
10674:
10665:
10657:G. johnstoni
10656:
10647:
10638:
10629:
10620:
10611:
10593:
10582:P. leightoni
10581:
10572:
10554:
10529:
10520:
10512:V. megaspila
10511:
10503:V. civettina
10502:
10490:
10478:
10469:
10454:
10438:
10411:
10402:
10393:
10381:
10369:
10360:
10351:
10342:
10316:
10307:
10298:
10289:
10187:
10178:
10170:F. margarita
10169:
10160:
10151:
10150:Jungle cat (
10142:
10133:
10121:
10105:
10096:
10088:P. planiceps
10087:
10078:
10069:
10059:Prionailurus
10057:
10045:
10015:
10003:
9991:
9990:Jaguarundi (
9974:
9962:
9941:
9932:
9923:
9914:
9902:
9890:
9881:
9872:
9863:
9854:
9845:
9837:L. geoffroyi
9836:
9827:
9826:Pampas cat (
9815:
9799:
9790:
9778:
9766:
9736:
9727:
9715:
9704:P. marmorata
9703:
9691:
9671:
9646:
9637:
9628:
9619:
9610:
9598:
9586:
9577:
9565:
9520:
9511:
9494:
9467:
9420:Superorder:
9414:Infraclass:
9339:
9329:
9320:
9311:
9302:
9293:
9284:
9276:C. l. lycaon
9275:
9267:C. l. ligoni
9266:
9257:
9248:
9239:
9230:
9221:
9212:
9204:C. l. arctos
9203:
9181:
9172:
9163:
9159:
9150:
9142:C. l. chanco
9141:
9133:C. l. chanco
9132:
9123:
9114:
9105:
9084:
9083:
9075:
9030:
8988:
8982:
8936:
8911:
8886:
8861:
8840:
8819:
8799:
8775:
8767:Bibliography
8752:. Retrieved
8745:the original
8740:
8727:
8700:
8688:. Retrieved
8678:
8671:
8659:
8650:
8646:
8640:
8628:
8616:. Retrieved
8590:
8547:
8543:
8536:
8517:
8507:
8488:
8484:
8474:
8462:. Retrieved
8455:the original
8441:
8432:
8426:
8401:
8397:
8393:
8392:) and dogs (
8389:
8349:
8343:
8331:
8319:
8307:
8297:December 16,
8295:. Retrieved
8291:
8281:
8269:
8257:
8205:
8201:
8195:
8184:the original
8179:
8175:
8166:
8158:
8146:. Retrieved
8137:
8133:
8127:
8119:
8107:
8098:
8094:
8084:
8072:
8060:
8048:
8036:
8024:
8012:
8000:
7988:
7976:
7964:
7952:
7940:
7921:
7898:
7886:
7836:
7832:
7826:
7818:
7781:
7777:
7767:
7755:
7720:
7716:
7706:
7694:
7649:
7645:
7635:
7602:
7598:
7592:
7580:
7570:November 19,
7568:. Retrieved
7553:
7541:
7529:
7519:November 11,
7517:. Retrieved
7489:September 2,
7487:. Retrieved
7480:the original
7467:
7448:
7445:Mammal Study
7444:
7434:
7416:
7409:
7397:
7393:
7381:
7372:
7363:
7353:September 2,
7351:. Retrieved
7322:. Retrieved
7317:
7308:
7283:
7275:
7263:. Retrieved
7253:
7241:. Retrieved
7231:
7221:September 1,
7219:. Retrieved
7199:
7187:. Retrieved
7178:
7168:
7156:. Retrieved
7147:
7137:
7110:
7098:
7086:
7074:
7051:
7045:
7033:
7028:, p. 7.
7021:
6987:
6983:
6973:
6961:
6949:
6944:, p. 4.
6937:
6904:
6900:
6894:
6882:
6862:
6855:
6843:
6831:
6789:
6777:
6772:, p. 5.
6765:
6723:
6719:
6713:
6701:
6689:
6677:
6665:
6653:
6634:
6624:
6612:. Retrieved
6571:
6567:
6557:
6510:(1): 66â76.
6507:
6503:
6499:
6478:
6454:
6443:
6431:
6419:
6374:
6370:
6360:
6333:
6306:
6279:
6254:
6250:
6244:
6235:
6231:
6225:
6215:November 22,
6213:. Retrieved
6193:
6186:
6176:
6172:
6162:
6152:November 22,
6150:. Retrieved
6130:
6122:
6085:
6081:
6071:
6052:
6048:
6043:Ursus arctos
6042:
6038:
6034:
6026:
6014:
5990:
5983:
5956:
5944:
5932:
5920:
5908:. Retrieved
5887:
5875:. Retrieved
5854:
5830:
5820:
5808:
5796:
5777:
5767:
5747:
5740:
5728:. Retrieved
5717:
5684:
5680:
5674:
5628:
5616:. Retrieved
5588:
5582:
5572:
5560:
5508:
5504:
5470:
5463:
5438:
5434:
5428:
5403:
5399:
5393:
5358:
5354:
5344:
5301:
5297:
5286:
5274:
5262:
5250:
5233:
5229:
5208:
5196:
5159:
5149:
5130:
5068:
5062:
5027:
5010:
5006:
5000:
4988:
4931:
4904:
4892:. Retrieved
4865:
4809:
4805:
4794:
4775:
4771:
4761:
4736:
4732:
4698:
4694:
4687:
4644:
4640:
4630:
4593:
4589:
4532:
4528:
4479:
4475:
4465:
4414:
4410:
4345:
4341:
4286:
4282:
4275:
4240:
4236:
4185:
4181:
4171:
4134:
4130:
4120:
4093:
4047:
4043:
4037:
3986:
3982:
3957:
3945:. Retrieved
3941:
3924:
3887:
3883:
3873:
3845:
3827:
3803:
3792:
3781:
3771:
3736:
3732:
3721:
3709:
3699:. New York,
3695:
3688:
3679:
3675:
3669:
3658:
3648:
3639:
3612:. Retrieved
3608:
3599:
3587:. Retrieved
3573:
3567:
3561:
3526:phylogenetic
3516:
3508:
3504:
3499:
3478:
3451:hunted using
3435:
3423:Fox Terriers
3404:
3379:Wolf hunting
3367:Central Asia
3355:
3324:
3320:
3314:
3290:
3286:
3278:
3263:(Mongolia),
3249:
3245:
3221:
3204:
3201:Farley Mowat
3194:
3188:
3181:
3173:
3167:
3133:
3126:Big Bad Wolf
3106:
3104:
3100:Gustave Doré
3093:
3058:
3012:
2988:
2956:
2913:
2893:
2844:Heilongjiang
2801:
2793:
2775:
2751:by 1684, in
2746:
2719:
2703:Mexican wolf
2679:
2667:
2595:
2574:lining. The
2556:
2536:
2517:
2505:
2446:
2422:
2417:Iberian wolf
2392:
2371:Reproduction
2357:
2331:
2322:
2304:Rallying cry
2248:
2240:
2219:
2189:
2180:malnutrition
2170:, toxic and
2168:constipation
2133:
2111:
2097:, vomiting,
2067:Lyme disease
2008:
1979:
1948:
1932:
1920:corsac foxes
1903:
1887:
1867:mountain-ash
1827:insectivores
1816:
1775:
1759:pack hunters
1756:
1696:
1691:Sassoferrato
1683:Italian wolf
1643:
1631:
1540:
1534:down to the
1521:
1516:Spiti Valley
1436:
1404:
1367:
1343:
1323:
1298:Canid hybrid
1286:Mexican wolf
1262:
1216:
1185:
1181:
1178:mosbachensis
1177:
1173:
1167:
1161:
1157:
1154:phylogenetic
1151:
1143:
1111:
1101:domesticated
1093:paleogenomic
1084:
1067:
1045:
1039:
1035:
1031:
1018:
1016:
990:
986:
968:
958:
952:
932:
920:
897:
877:
861:
838:
829:
806:
797:
774:
765:
742:
733:
712:African wolf
710:
701:
678:
659:
658:
574:
571:Anglo-Saxons
569:such as the
564:
555:
551:
537:
512:
497:Wolf attacks
470:
466:rabies virus
418:game hunting
406:
401:
384:
378:
357:
339:
335:
331:
322:
321:
317:
315:
291:
286:
262:
260:
244:
243:
231:
92:
48:
40:
15690:NatureServe
15612:iNaturalist
15457:canis-lupus
15444:Canis_lupus
15430:Canis lupus
15424:Wikispecies
15400:Canis lupus
15260:Denisova 11
15050:Sulimov dog
14819:M. sibirica
14801:M. putorius
14774:M. nigripes
14756:M. lutreola
14729:M. haidarum
14625:N. africana
14588:A. congicus
14579:A. cinereus
14570:A. capensis
14470:Sea otter (
14448:L. provocax
14355:I. striatus
14276:Ictonychini
14171:M. melampus
14083:Wolverine (
14061:P. pennanti
13984:M. moschata
13975:M. everetti
13921:M. leucurus
13879:A. hoevenii
13870:A. collaris
13820:M. capensis
13715:N. olivacea
13630:Olinguito (
13586:Bassaricyon
13573:P. pygmaeus
13503:Bassariscus
13476:Procyonidae
13451:S. putorius
13442:S. gracilis
13360:M. mephitis
13351:M. macroura
13245:Red panda (
13225:Musteloidea
13186:Ross seal (
13139:Lobodontini
13071:Neomonachus
12999:P. sibirica
12957:P. vitulina
12922:Grey seal (
12904:Harp seal (
12796:A. pusillus
12760:A. forsteri
12595:Callorhinus
12529:Pinnipedia
12488:Musteloidea
12260:C. simensis
12224:C. lupaster
12093:Hoary fox (
12086:L. sechurae
12059:L. fulvipes
12050:L. culpaeus
11946:(true dogs)
11905:Swift fox (
11880:V. macrotis
11772:Nyctereutes
11681:Caniformia
11626:H. ochracea
11584:B. nigripes
11575:B. jacksoni
11462:U. javanica
11379:Herpestinae
11354:C. obscurus
11345:C. ansorgei
11325:Crossarchus
11181:(mongooses)
11170:Herpestidae
11150:S. durrelli
11141:S. concolor
11091:G. fasciata
11028:E. goudotii
10949:Herpestidae
10859:P. cristata
10720:G. thierryi
10675:G. maculata
10639:G. cristata
10630:G. bourloni
10383:Paradoxurus
10341:Binturong (
10280:Hemigalinae
10257:Viverroidea
10220:Viverroidea
10179:F. nigripes
10031:Leopard cat
10016:P. concolor
9933:L. pardinus
9882:L. tigrinus
9873:L. pardalis
9864:L. jacobita
9855:L. guttulus
9828:L. colocola
9587:N. nebulosa
9557:Pantherinae
9437:Feliformia
9330:C. l. rufus
9312:C. l. orion
9173:C. l. lupus
9151:C. l. dingo
9115:C. l. arabs
9106:C. l. albus
8741:Wild Sentry
8720:Graves 2007
8404:: 232â245.
8390:Canis lupus
8336:Graves 2007
8128:Canis Lupus
8101:(1): 37â41.
8077:Marvin 2012
8065:Graves 2007
8041:Marvin 2012
8017:Marvin 2012
7993:Marvin 2012
7981:Marvin 2012
7945:Marvin 2012
7903:Marvin 2012
7318:www.fws.gov
7189:October 30,
7158:October 30,
6614:January 10,
6500:Canis lupus
6311:Graves 2007
6238:(7): 27â29.
6179:(1): 17â32.
6088:(41): e41.
6035:Canis lupus
5910:October 29,
5065:and allies"
5063:Canis lupus
4868:(37): 1â6.
4647:: 110â118.
4482:: 281â307.
3989:(1): 2882.
3714:Marvin 2012
3676:Die Sprache
3614:January 14,
3562:Canis lupus
3485:exotic pets
3479:Wolves and
3419:Bloodhounds
3415:sighthounds
3363:Middle East
3327:habituation
3302:Wolf attack
3240:Hexham wolf
3148:Leo Tolstoy
2994:Pawnee and
2832:Maharashtra
2698:Great Lakes
2393:Wolves are
2351:and raised
2251:territorial
2249:Wolves are
2083:listeriosis
2063:brucellosis
2043:dehydration
1934:Brown bears
1907:Custer Wolf
1894:Cannibalism
1863:raspberries
1859:blueberries
1811:wild horses
1628:Zoo, France
1617: in).
1528:guard hairs
1393:Description
1169:C. etruscus
1097:sister taxa
1085:C. l. lupus
1073:common name
1025:mammalogist
991:Canis lupus
515:Old English
509:Wolf (name)
446:territorial
415:cooperative
411:specialized
358:Canis lupus
323:Canis lupus
292:Canis lupus
281:Subspecies
263:Canis lupus
140:Appendix II
66:(400,000â0
15885:Scavengers
15839:Categories
15275:Koolakamba
15193:Kellas cat
15173:Bengal cat
14947:Dwarf Lulu
14792:M. nudipes
14783:M. nivalis
14747:M. kathiah
14702:M. erminea
14693:M. altaica
14643:N. frenata
14634:N. felipei
14604:Mustelinae
14346:I. libycus
14260:G. vittata
14126:M. caurina
13903:M. anakuma
13781:Mustelidae
13768:Mustelidae
13744:see belowâ
13735:Mustelidae
13632:B. neblina
13543:(raccoons)
13514:B. astutus
13512:Ringtail (
13488:Kinkajou (
13460:S. pygmaea
13401:M. marchei
13259:Mephitidae
13127:M. leonina
13019:Monachinae
12990:P. hispida
12981:P. caspica
12778:A. gazella
12710:N. cinerea
12606:C. ursinus
12544:Odobenidae
12497:see belowâ
12477:see belowâ
12461:Pinnipedia
12379:Sun bear (
12346:T. ornatus
12335:Tremarctos
12311:Ailuropoda
12249:Red wolf (
12215:C. latrans
12138:L. adustus
12095:L. vetulus
12068:L. griseus
11998:Bush dog (
11889:V. pallida
11887:Pale fox (
11871:V. lagopus
11842:Cape fox (
11722:Gray fox (
11480:U. smithii
11312:H. parvula
11303:H. hirtula
11191:Mungotinae
11080:Galidictis
11053:Galidiinae
10981:Euplerinae
10964:Eupleridae
10953:sensu lato
10924:C. crocuta
10857:Aardwolf (
10788:see belowâ
10729:G. tigrina
10702:G. poensis
10684:G. pardina
10666:G. letabae
10648:G. genetta
10546:Genettinae
10530:V. zibetha
10450:Viverrinae
10439:sensu lato
10434:Viverrinae
10403:P. jerdoni
10270:Viverridae
10229:see belowâ
10168:Sand cat (
9975:A. jubatus
9800:C. caracal
9693:Pardofelis
9512:P. linsang
9453:Feliformia
9328:Red wolf (
9195:subspecies
9097:subspecies
9034:subspecies
8994:Bill Mason
8705:Lopez 1978
8653:: 162â166.
8618:August 16,
8491:(3): 253.
8485:BioScience
8053:Lopez 1978
8029:Lopez 1978
8005:Lopez 1978
7969:Busch 2007
7957:Lopez 1978
7891:Lopez 1978
7760:Busch 2007
7546:Busch 2007
7299:0816639787
6694:Lopez 1978
6682:Lopez 1978
6670:Busch 2007
5591:: 110368.
5279:Lopez 1978
5213:Lopez 1978
5201:Lopez 1978
5189:Lopez 1978
5095:Lopez 1978
3818:0521415292
3536:References
3443:Kazakhstan
3427:strychnine
3387:See also:
3160:Ivan Bunin
3108:Ysengrimus
3053:See also:
3015:Greek myth
2882:See also:
2867:In culture
2395:monogamous
2375:See also:
2325:media help
2230:mated pair
2215:Mysore Zoo
2203:See also:
2192:See also:
2160:roundworms
2144:protozoans
1988:Infections
1944:wolverines
1910:also kill
1879:cowberries
1875:bilberries
1849:, melons,
1728:topography
1716:grasslands
1712:shrublands
1700:extirpated
1311:KadzidĆowo
1265:divergence
1196:temporalis
1079:being the
1046:hallstromi
1036:familiaris
1001:Subspecies
933:3.50
507:See also:
438:mated pair
346:native to
97:Polar Park
15252:Hominidae
14979:Camelidae
14738:M. itatsi
14672:(weasels)
14535:Lutrogale
14472:E. lutris
14430:L. felina
14239:(grisons)
14162:M. martes
14105:(martens)
14023:Guloninae
13809:Mellivora
13682:N. narica
13623:B. medius
13614:B. gabbii
13605:B. alleni
13593:(olingos)
13562:Raccoon (
13414:Spilogale
13300:C. chinga
13281:Conepatus
13237:Ailuridae
13036:Monachini
12948:P. largha
12618:Otariinae
12581:sea lions
12576:fur seals
12567:Otariidae
12452:Mustelida
12423:U. arctos
12285:Arctoidea
12242:C. lycaon
12197:C. aureus
12127:Lupulella
12039:Lycalopex
11916:V. vulpes
11914:Red fox (
11878:Kit fox (
11853:V. corsac
11597:Herpestes
11210:Meerkat (
10969:(Malagasy
10871:Hyaeninae
10832:Hyaenidae
10747:G. felina
10161:F. lybica
9973:Cheetah (
9891:L. wiedii
9880:Oncilla (
9846:L. guigna
9817:Leopardus
9798:Caracal (
9791:C. aurata
9726:Bay cat (
9647:P. pardus
9645:Leopard (
9620:P. tigris
9578:N. diardi
9496:Prionodon
9396:Kingdom:
9385:Carnivora
9193:New World
9095:Old World
9082:Species:
9064:Carnivora
9044:Kingdom:
9031:gray wolf
8754:March 21,
8274:Mech 1981
7243:March 23,
7079:Mech 1981
6504:Ecography
6257:: 85â87.
5813:Mech 1981
5687:: 30â33.
5633:Mech 1981
5618:April 23,
5613:0031-0182
4993:Mech 1981
4936:Mech 1981
4844:225956269
4439:0962-1083
4370:0028-0836
4319:237147842
4065:2246/5999
4050:: 1â218.
4011:2045-2322
3609:cites.org
3345:oblasts,
3281:sympatric
3259:(India),
3218:Conflicts
3017:tells of
3000:Milky Way
2998:call the
2996:Blackfoot
2836:Karnataka
2816:Rajasthan
2728:(Appendix
2552:mouselike
2434:domiciles
2365:urination
2360:preputial
2186:Behaviour
2156:tapeworms
2148:helminths
2114:arthropod
2103:hematuria
2075:tularemia
2053:shock or
2051:endotoxic
1831:waterfowl
1807:mule deer
1797:, moose,
1791:wild boar
1763:ungulates
1687:Apennines
1524:undercoat
1387:gene flow
1326:gene flow
1258:gene flow
1208:dire wolf
1200:premolars
1182:C. lupus.
1122:Evolution
660:Gray wolf
503:Etymology
462:Pathogens
450:carnivore
340:grey wolf
336:gray wolf
239:Species:
211:Carnivora
177:Kingdom:
171:Eukaryota
63:â present
15791:Species+
15695:2.105212
15682:14000738
15630:11407661
15464:BioLib:
15409:Wikidata
15265:Humanzee
15198:Pumapard
15080:Wholphin
14913:Mammalia
14907:Chordata
14901:Animalia
14899:Kingdom
14718:Ferret (
14652:N. vison
14513:L. lutra
14381:(otters)
14375:Lutrinae
14232:Galictis
14144:M. foina
14059:Fisher (
13947:Melogale
13930:M. meles
13850:Arctonyx
13695:Nasuella
13673:N. nasua
13650:(coatis)
13564:P. lotor
13340:Mephitis
13099:Mirounga
12849:Phocinae
12830:Phocidae
12699:Neophoca
12645:Zalophus
12552:Walrus (
12251:C. rufus
12233:C. lupus
12213:Coyote (
12048:Culpeo (
11989:Speothos
11925:V. zerda
11907:V. velox
11844:V. chama
11555:Bdeogale
11453:U. fusca
11292:Helogale
11262:M. mungo
11201:Suricata
11130:Salanoia
11037:E. major
11017:Eupleres
10848:Proteles
10837:(hyenas)
10601:(genets)
10152:F. chaus
10143:F. catus
10134:F. bieti
10014:Cougar (
9964:Acinonyx
9942:L. rufus
9940:Bobcat (
9889:Margay (
9871:Ocelot (
9844:Kodkod (
9765:Serval (
9728:C. badia
9717:Catopuma
9627:Jaguar (
9611:P. uncia
9600:Panthera
9567:Neofelis
9485:Feloidea
9416:Eutheria
9410:Mammalia
9404:Chordata
9402:Phylum:
9398:Animalia
9068:Family:
9058:Mammalia
9052:Chordata
9050:Phylum:
9046:Animalia
8684:Archived
8609:Archived
8564:86058778
8464:July 17,
8230:84878446
8148:July 28,
8142:Archived
7863:27005517
7855:19877836
7827:Temminck
7810:26082870
7784:: e982.
7747:28585433
7686:30969994
7646:PLOS ONE
7564:Archived
7513:Archived
7376:page 50.
7347:Archived
7324:March 6,
7212:Archived
7183:Archived
7152:Archived
6929:52063038
6608:Archived
6604:22125487
6596:17770508
6532:62800394
6411:26398784
6371:PLOS ONE
6271:85957777
6209:Archived
6146:Archived
6114:31966128
5901:Archived
5877:July 26,
5868:Archived
5730:July 25,
5709:91073874
5537:31174275
5420:85236511
5377:19603061
5355:Heredity
5336:19197024
4894:July 30,
4888:Archived
4836:33122379
4753:24622972
4679:31563851
4641:iScience
4622:30419012
4567:30344120
4506:26721918
4498:27912242
4457:31840921
4388:35768506
4311:34398980
4267:26680994
4212:26004765
4163:24453982
4074:83594819
4029:35217686
3947:March 6,
3916:24453989
3865:62265494
3835:(2005).
3778:"canine"
3763:26234211
3682:: 13â18.
3447:Mongolia
3431:trapping
3411:coursing
3401:, Russia
3224:cortisol
3174:Peasants
3098:(1883),
3072:cry wolf
3043:Dena'ina
3027:werewolf
2972:Buddhism
2968:Hinduism
2936:she-wolf
2900:she-wolf
2848:Xinjiang
2804:Pakistan
2639:Appendix
2635:Pakistan
2611:Appendix
2607:Red List
2512:muskoxen
2442:railways
2436:, paved
2235:Cortisol
2095:anorexia
1855:cherries
1787:roe deer
1783:red deer
1724:pastures
1708:wetlands
1634:ochreous
1514:Wolf in
1411:rib cage
1328:between
1250:Beringia
1232:that is
1224:and the
1158:C. lupus
1068:C. lupus
1056:for the
1032:C. lupus
590:Taxonomy
573:took on
485:agrarian
481:pastoral
391:and the
270:Linnaeus
217:Family:
201:Mammalia
191:Chordata
187:Phylum:
181:Animalia
167:Domain:
127:IUCN 3.1
15817:ZooBank
15757:1506689
15669:1005943
15591:5219173
15324:Mustela
15223:Liliger
15188:Chausie
15183:Caraval
15160:Felidae
15148:Zebroid
15085:Narluga
15067:Cetacea
15055:Wolfdog
15025:Coywolf
15007:Canidae
14997:Huarizo
14942:Beefalo
14924:Bovidae
14905:Phylum
14720:M. furo
14665:Mustela
14614:Neogale
14461:Enhydra
14335:Ictonyx
14294:Vormela
14251:G. cuja
14178:Sable (
14085:G. gulo
14050:Pekania
14035:Tayra (
13832:Melinae
13644:Nasuina
13536:Procyon
12892:Phocini
12531:(seals)
12466:(seals)
12358:Ursinae
12300:(bears)
12295:Ursidae
12171:Dhole (
11961:(zorro)
11835:V. cana
11745:Vulpini
11713:Urocyon
11697:Canidae
11489:U. urva
10993:Fossa (
10913:Crocuta
10595:Genetta
10492:Viverra
10033:lineage
9955:lineage
9924:L. lynx
9780:Caracal
9754:lineage
9752:Caracal
9684:lineage
9682:Bay cat
9668:Felinae
9629:P. onca
9618:Tiger (
9538:Felidae
9408:Class:
9387:species
9383:Extant
9149:Dingo (
9074:Genus:
9070:Canidae
9062:Order:
9056:Class:
9029:Extant
8690:May 14,
8406:Bibcode
8210:Bibcode
7801:4465947
7738:5460078
7677:6457547
7654:Bibcode
7627:2386069
7607:Bibcode
7265:May 10,
6921:1382426
6740:1200478
6728:Bibcode
6576:Bibcode
6568:Science
6512:Bibcode
6402:4580640
6379:Bibcode
6105:6661435
5689:Bibcode
5593:Bibcode
5443:Bibcode
5435:Lethaia
5385:5228987
5327:2903542
5306:Bibcode
5298:Science
5242:3784236
4884:3503924
4827:7116352
4806:Science
4703:Bibcode
4670:6817678
4649:Bibcode
4613:6231604
4558:6224481
4537:Bibcode
4448:7317801
4419:Bibcode
4379:9279150
4350:Bibcode
4291:Bibcode
4258:4728369
4190:Bibcode
4154:3894170
4020:8881584
3991:Bibcode
3907:3894154
3741:Bibcode
3589:June 2,
3351:Ukraine
3347:Karelia
3343:Irkutsk
3335:Belarus
3331:grazing
3269:turkeys
3178:Borzois
3113:Reynard
3039:witches
2989:In the
2820:Haryana
2812:Gujarat
2776:In the
2753:Ireland
2709:Eurasia
2680:In the
2617:of the
2605:on its
2576:kidneys
2572:stomach
2479:⁄
2465:⁄
2426:magpies
2403:Oestrus
2389:, Japan
2353:hackles
2349:posture
2107:icterus
2087:anthrax
1959:cougars
1890:carrion
1851:berries
1835:lizards
1819:rodents
1799:caribou
1744:caribou
1693:, Italy
1662:Ecology
1612:⁄
1598:⁄
1584:⁄
1570:⁄
1556:⁄
1488:⁄
1474:⁄
1460:⁄
1446:⁄
1433:, Italy
1407:Canidae
1315:spaniel
1277:Basenji
1246:Siberia
1230:lineage
1218:Genomic
983:jackals
541:wÄșÌ„kÊ·os
475:by the
458:carrion
430:howling
397:hybrids
375:Canidae
348:Eurasia
227:Genus:
221:Canidae
207:Order:
197:Class:
142: (
125: (
15895:Wolves
15770:247341
15728:NZOR:
15643:180596
15575:FEIS:
15555:305289
15529:CANILU
15516:328607
15500:ECOS:
15452:ARKive
15415:Q18498
15228:Marozi
15035:Dogxim
15020:Coydog
14967:Ć»ubroĆ
14962:Yakalo
14911:Class
14410:Lontra
14098:Martes
13373:Mydaus
13267:skunks
12231:Wolf (
12111:Canina
11941:Canini
11805:Vulpes
11701:(dogs)
11242:Mungos
10556:Poiana
9638:P. leo
9636:Lion (
9542:(cats)
8980:Watch
8948:
8923:
8898:
8873:
8848:
8827:
8806:
8787:
8601:
8562:
8524:
8356:
8228:
7928:
7861:
7853:
7808:
7798:
7745:
7735:
7684:
7674:
7625:
7296:
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2481:4
2477:3
2474:+
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2467:2
2463:1
2460:+
2327:.
2150:(
1614:8
1610:1
1607:+
1605:5
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1600:8
1596:3
1593:+
1591:4
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1083:(
536:*
320:(
146:)
129:)
91:(
70:)
38:.
20:)
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